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ProQR Therapeutics N.V.

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FY2016 Annual Report · ProQR Therapeutics N.V.
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ANNUAL
REPORT 
2016

Acting in the interest of patients

ProQR Therapeutics N.V. 

T  : +31 88 166 7000 

W : www.proqr.com 

E  : info@proqr.com

Headquarters Leiden:

Zernikedreef 9, 2333 CK Leiden,

the Netherlands

Office Palo Alto:

543 Bryant Street, Palo Alto,

CA 94301, USA

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ANNUAL REPORT 2016

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ANNUAL RE PORT 2016

FINDING TREATMENTS FOR RARE GENETIC DISEASES TO 

CHANGE 
THE 
LIVES OF 
PATIENTS

ProQR develops drugs to improve the lives of patients around the world.  

Our scientific staff pioneer in the development of RNA medicines that fight 

rare genetic diseases. Almost five years after we founded ProQR, it has 

moved far beyond the status of being a very promising start-up company. 

And with the positive outcome of our proof-of-concept clinical trial in  

CF patients we see the first results of our efforts, which encourage and 

inspire us to keep pushing the envelope. 

This is a 
company 
in which 
NEW IDEAS 
emerge on  
a daily base

At ProQR, we are in the business 

of developing RNA medicines for 

New solutions, real medicines
It is this constant thinking of the 

patients that suffer from severe 

need of patients that keeps us at 

genetic rare diseases. The business 

ProQR on our toes. Our approach 

itself is not what makes us try hard-

leads to results, thanks to the appli-

er every day. What is? 

cation of highly specific and elegant 

RNA approaches that have been de-

It is the belief that our company can 

veloped building on the science that 

truly make a difference in the lives 

was done over the last 20 years. 

of patients. We are convinced that, 

Our in-house research team in what 

when we target all our knowledge, 

we call ‘the innovation unit’, has 

creative thinking and perseverance 

discovered many new molecules 

to the same goal, we can truly make 

that we aim to develop to become 

a meaningful impact on the lives of 

real medicines. 

patients and their loved ones.

Programs and ideas
To this end, we started our lead 

The RNA technologies have helped 

us build an extensive and what we 

believe is a valuable pipeline. Over 

program in CF that entered the 

time it will give us the opportunity 

clinical trial phase in 2015. But 

to help patients that are in need of 

from the start, we have strived to 

new therapies and build a sustain-

go beyond that. We now have two 

able independent business.

other programs in development for 

inherited blindness and a debili-

tating skin disease. And we won’t 

Progress in 2016
During 2016 we made true progress. 

stop there; we have numerous 

programs in various stages of early 

First of all, we announced the  

development. This is a company in 

encouraging results from our  

which new ideas emerge on a daily 

proof-of-concept clinical trial in  

basis. Ideas get challenged, some 

CF patients. Furthermore, we estab-

rejected, some selected for further 

lished a new group that focuses on 

investigation. Some make it to 

inherited blindness; the group now 

projects that one day may become 

works on the development of sev-

programs and – who knows – real 

eral products, the most advanced 

medicines that can help patients. 

one is for LCA 10 (Leber’s congenital 

amaurosis Type 10).

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ANNUAL REPORT 2016

A bright 
future 
beckons in 
which DNA 
errors can  
be targeted  
at the  
RNA LEVEL

Also, we brought our first program 

milestones and significant successes  

Valerio, the founding CEO of the very 

for debilitating skin diseases from 

in the next few years. To help our 

successful Dutch biotech Crucell. 

the idea phase into development. 

company grow stronger in this area 

in 2016, James Shannon, former 

Where will our quest bring ProQR?  

As said, we won’t stop there. We are 

Chief Medical Officer at GSK, joined 

A bright future beckons in which DNA 

moving on, beyond the phase of 

the supervisory board. We already 

defects can be targeted at the RNA 

‘promising’. We have grown into a 

had a very strong team including 

level. We will put the latest advance-

team that can bring ProQR to real  

people like Henri Termeer, long-

ments to use, to changes the lives of 

term CEO at Genzyme and Dinko 

patients and their loved ones. 

RESEARCH AND DEVELOPMENT PIPELINE

QRX-704
Huntington’s
disease

QRX-411
Usher syndrome

QRX-504
Fuchs (FECD)

QRX-421
Usher syndrome

QR-313

Dystrophic EB
~2,000 patients

QRX-021
Undisclosed
CF target

QRX-604
Friedreich's 
ataxia

QRX-323
Dystrophic EB

QRX-203
Amyloid beta
related disorders 

QR-110

LCA10
~2,000 patients

QR-010

cystic fibrosis
>65,000 patients

Innovation

Pre-clin
development

Phase 1 &
clinical PoC

Pivotal
studies

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ANNUA L REPORT 2016

140

ProQRians

Average age

37yrs

Gender

44%

56%

Nationalities

30

 
 
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ANNUAL RE PORT 2016

ProQR – WHO WE ARE, WHAT WE DO

IT’S IN 
OUR RNA

ProQR is building. Initially, the company was founded to beat cystic 

fibrosis in just one child. Today, ProQR’s quest leads far beyond, as its 

mission is to develop treatments for all patients with rare genetic diseases. 

With a pipeline across 5 different therapeutics area’s progressing well,  

the company’s activities gain momentum. ProQR is proud to have 

expanded its one-vessel enterprise into a fleet. 

Yes, we  
aim high,  
as we are 
not afraid to 
THINK BIG

the CF program that led to the gen-

and gene therapy approaches. This 

– we are moving fast in building a 

eration of strong biomarker data in 

is a truly powerful technology, as it 

sustainable future for the company.

patients with CF early in the develop-

offers ProQR’s scientists – and, in a 

ment process. 

Pipeline built on RNA
Based on the company’s strong 

later stage: doctors – the potential 

to treat severe genetic diseases, 

Infrastructure for growth
As it all starts with people, ProQR 

including Leber’s congenital  

builds its infrastructure and its 

amaurosis, Usher syndrome, Fuchs 

development power with a strong 

foundation a diversified pipeline has 

endothelial corneal dystrophy,  

focus on finding the ProQRians with 

been constructed with programs 

dystrophic epidermolysis bullosa 

the professionalism and spirit. With 

that target diseases in which a signif-

and Amyloid beta related disorders.

our 140 strong staff we are proud 

icant impact can be made to  

individual patient’s lives. All of  

ProQR’s programs are based on 

Aim high, think big
At ProQR, we aim to play a deci-

of our team culture, based on the 

shared belief that, together, we can 

make a difference. 

innovative and promising RNA 

sive role in all of these rare genetic 

technologies including one licensed 

diseases. Yes, we aim high, as we 

Also, ProQR has built its facilities 

from Massachusetts General Hos-

are not afraid to think big. It is in our 

to make this happen. In 2016 the 

pital in Boston, USA. An obvious 

RNA! Our founder and CEO Daniel 

new company headquarters were 

choice since an RNA molecule can 

de Boer has a track record in the 

opened, according to the ‘science-

be designed specifically for a certain 

IT industry with its ever-ambitious 

at-our-core’ model. At the new  

defect that causes the disease. 

timelines; he is therefore not afraid 

ProQR location, the labs are the 

to challenge the pharma & drug  

core of the building.

ProQR is building on the over 20 

development paradigm to keep a 

years of experience in the RNA field. 

high pace of development. 

And, last but not least, ProQR fur-

With the different technologies 

ther builds its relationship with both 

developed a wide range of genetic 

ProQR’s deep-rooted urge to 

patients and medical professionals. 

disease can now be targeted at 

generate tangible and meaningful 

As the company strongly believes  

the underlying cause, the mutated 

progress has led to some unconven-

in patient-centric development,  

mRNA that leads to a defective 

tional steps, like listing the company 

a patient and medical community 

protein and disease symptoms. 

on Nasdaq only two years after its 

engagement (PMCE) team has been 

With several approved products, 

foundation. The same motto applies 

installed in 2016. It shows ProQR’s 

ProQR’s strong development in 2016 

built methodically. The founding 

were set up to drive the most prom-

RNA is becoming an established 

to starting a proof-of-concept study, 

dedication to maintaining close rela-

underlines the company’s remark-

team put together a strong manage-

ising science to patients in need. 

approach. Our programs take this 

showing us very early in the devel-

tionships with the communities we 

able execution power, based on the 

ment team as well as a supervisory 

to the next level and aim to restore 

opment process the drug activity 

serve. The support ProQR received 

firm belief that there is a future in 

board with exceptional biotech ex-

The execution power was also 

the function of these defective 

of our lead molecule for CF. The 

from them is crucial in reaching our 

targeting DNA defects at the RNA 

perts and a team of motivated RNA 

extended to the scientific level of 

proteins without changing the DNA 

decision to invest in the discovery of 

goals – in 2016, in 2017 and in the 

level. Since ProQRs inception, the 

and CF experts. Collaborations with 

ProQR. An aggressive and innovative 

of the cells. This has advantages 

new programs and technologies also 

years to come – finding treatments 

foundation of the company has been 

top-notch academic collaborators 

development plan was executed for 

over for example a small molecule, 

shows this typical quality of ProQR 

for severe rare genetic diseases. 

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ANNUAL RE PORT 2016

This is the story of twin sisters Myrthe and Rosanne, 29, living in 

Amsterdam. Not long ago, Myrthe has successfully finished a 150-kilometre 

skating race. Rosanne, currently working on a three-month project in South 

Africa as part of a job that involves lots of international travel, ran the half 

marathon of Amsterdam. Twin sisters, in great shape? Certainly, but there  

is one minor detail: both Myrthe and Rosanne suffer from cystic fibrosis,  

also known as CF.

“We despe-
rately hope 
for A REAL 
TREATMENT, 
one that 
changes 
lives”

Myrthe

Two women, almost in their thirties. 

familiar with occasional health 

After studying Business Administra-

setbacks, leading to a decline in 

tion, Rosanne found employment 

pulmonary function. 

with an international consulting 

firm. Her sister Myrthe studied  

Myrthe and Rosanne both found 

Economics and currently works as  

jobs with an employer that is very 

a business controller for a large 

sympathetic to their situation. 

retail organisation. 

Rosanne: “I did not beat around 

the bush during the job interview, 

So far, nothing out of the ordinary. 

stating that CF is a serious disease. 

However, both start their working 

I know how to control it, I said, and 

day on similar routines, familiar 

how to stay healthy. My boss is 

to anyone with CF. “I switch on the 

very understanding. After I was in 

nebulizer”, Rosanne explains, “to 

hospital for two months following 

get my daily shots of medication in 

an infection, we decided that a four-

about 40 minutes, including antibi-

day contract would better suit me.  

otics. It takes about 30 minutes to 

I work on Monday, Tuesday, Thurs-

get ready for a new working day, 

day and Friday. On Wednesday, 

eating and applying make-up not 

I rest and recharge. I think I have 

included.” Her sister adds that the 

found my ideal work-life balance.”

evening routine is slightly shorter. 

“We both have to take pills with 

pancreatic enzymes whenever 

Ticking clock
Meanwhile, there is this nasty 

we eat any fatty foods to support 

disease, CF. Myrthe: “From a young 

digestion. It requires quite a num-

age I knew that my health would 

ber of pills to get us through the 

deteriorate over time. For many 

day, especially since our doctors 

years, I thought I was to lose this 

insist we consume at least 3,000 to 

battle eventually. My lung function 

3,500 calories per day.” Rosanne, 

declined and I even got lung haem-

giggling: “Fortunately, we both like 

orrhages.” To make things worse, 

fastfood, haha!”

Health setbacks
Three months after they were born 

both Rosanne and Myrthe were 

diagnosed with Cystic Fibrosis  

Related Diabetes. Myrthe: “We 

accept it as it is; we take the insulin 

Rosanne was hospitalized with 

and go on with our lives.”

severe lung infections. Despite the 

diagnosis of CF, they had a fairly 

normal childhood. Both became 

PUSHING 
THE BOUN-
DARIES OF 
CYSTIC 
FIBROSIS

 Rosanne (left) and Myrthe (right)

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ANNUAL RE PORT 2016

CF BASICS 

Cystic Fibrosis

Other 
mutations

65,000

CF patients with 
F508del mutation
worldwide

Lungs and 
other organs

Median age of death is 
30 years or less

Genetic

No therapy 
available

Fortunately, the big picture of CF 

Myrthe: “Any advancement in  

since this year also as treasurer) of 

has changed for the better. Over 

science may help stop the clock 

the Skate4AIR project. “This year, 

the years, scientific advancements 

from ticking for a longer period 

my aim was to skate the 150-kilo-

and new treatments have resulted 

– or even indefinitely. When we 

meter race. Not only did  

in a much higher life expectancy for 

were born, life expectancy was 

I finish well with 167 kilometres,  

CF patients. Myrthe and Rosanne 

low at about 20. During our lives, 

I also reached my goal of collecting 

have also benefitted from the trend 

scientific developments raised life 

€10,000 to support CF research! 

– their long function is now stable 

expectancy, but we hope for more. 

In fact, I raised over €16,000!” 

at around 70 percent. Rosanne:  

All we can do it lead a sensible life, 

Rosanne has adopted a similarly 

“Without successful research, 

with as much sports as possible.”

sport-minded lifestyle. She ran  

the clock would be ticking louder 

every day for us, counting back 

from 70, 60 and 50 to, eventually, 

Living like athletes
Hence, both Rosanne and Myrthe 

the half marathon of Amsterdam  

1 hour and 59 minutes. A formi-

dable achievement.

zero. Today, I am happy with my 

live like athletes to stay in shape.  

“By staying 
in shape, 
we try to 
BUY TIME” 

Rosanne

situation. Fortunately, medical care 

that area, having a child one day 

for CF patients in Johannesburg 

definitely is an option.”

is quite advanced, and there is no 

need to worry.”

The future
What about the future? How do 

The twins come across as opti-

mistic, smiling, confident women. 

“We’re not there yet”, Rosanne 

insists. “We are still waiting for the 

these CF patients deal with relation-

real breakthrough. Current drugs 

ships, possibly even with the idea 

do not work for every patient. 

of one day becoming a mother? 

There is still much research need-

Rosanne: “We both have been in 

ed to provide a better future for 

relationships, in which we chose 

CF patients. We don’t know how 

Rosanne left for Johannesburg 

to be honest and open about our 

our disease will develop later on. 

to work in a three-month project 

situation. Our doctors told us that 

By staying in shape, we try to buy 

lung function as it is. I will try to do 

in 2014, Myrthe skated a 100- 

Pushing boundaries is what the 

for one of her employer’s clients. 

childbirth is possible for CF patients 

time.” Myrthe: “We desperately 

everything in my power to stay at 

kilometer race on the Weissensee 

twins keeps doing, in different ways. 

“Though I am a CF patient, I am 

with a lung function of over 50 

hope for a real treatment, one that 

this level.” 

in Austria, as a participant (and 

One day after our conversation, 

confident about my current health 

percent. Since we both do well in 

changes lives.” 

PAGE  XI
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ANNUAL REPORT 2016

WORKING 
ON THE 
“POTENTIAL 
TO CHANGE  
THE LIFE  
OF CF- 
PATIENTS”

Nicolas Lamontagne, QR‑010 captain 
at ProQR, about clinical trials

In just four years, ProQR has advanced QR-010 – an innovative inhaled 

therapy for cystic fibrosis based on RNA – from an idea to two clinical trials. 

To Nicolas Lamontagne, QR-010 captain at ProQR, the clinical trial results 

are a clear step forward. “Most companies that develop drugs have to go 

from multiple failures to a positive clinical trial result. We obtained a 

positive first trial very quickly. In the industry, that is an amazing result.”

PAGE XI I
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ANNUAL RE PORT 2016

ABOUT QR-010 & CF

NPD – an important signal
“This NPD test”, Nicolas Lamontagne 

show activity. In the cells of CF pa-

tients, it does what it should be  

says with a smile, “was developed 

doing. This is the best result we 

Cystic fibrosis (CF) is one of the 

more than thirty years ago as a 

could have hoped for, as failure in 

genetic diseases that ProQR is 

diagnostics tool. It was a quick way 

phase 1 is very common in the in-

hoping to find a treatment for. 

to show that an important protein is 

dustry.” Nicolas adds that the results 

QR‑010, the molecule that ProQR 

not working, indicating the patient 

for the safety and tolerability study 

has discovered and is developing, 

may have CF. We used the tool the 

are expected to become available 

specifically targets the most 

other way around, to show that a 

later in 2017. “We are excited. I can’t 

common mutation in cystic fibrosis, 

treatment has a positive effect on 

wait to see the results.”

called the F508del mutation, 

this protein function. Hence, the 

affecting ~65,000 CF patients in  

NPD test provided an important 

the western world.  

signal of the therapeutic potential of 

More and bigger studies
Next on the menu will be more and 

QR-010 in people with cystic fibrosis. 

bigger – phase 2 – studies in 2018, 

QR‑010 targets the disease at the 

It is a first but promising signal of 

with more patients, focusing on the 

RNA level with the aim to restore 

drug activity. As we all know, drug 

safety and efficacy of the molecule, 

the function of the defective CFTR 

development is a long process. The 

over longer periods. “We plan for 

protein that causes CF. The goal of 

NPD trial is a clear ‘go ahead’ signal.”

success!”, Nicolas smiles. Thinking 

ProQR’s QR‑010 is to address the 

this over, he adds: “This company 

underlying defect and stop the 

Nicolas adds that ‘quick’ does not 

has a way of operating that I have 

progression of cystic fibrosis. 

necessarily mean ‘easy’. “The test  

not seen before. We don’t lose time 

is viable, but difficult to perform.  

in lengthy discussions, but focus on 

We took no chances – we wanted  

key actions leading to results. I tell 

a top-quality trial. To get it right –  

my team all the time – ‘there is no 

of QR-010 in approximately 64 CF 

‘fit to purpose’ - we decided to  

time to waste’. There are patients 

patients, by inhalation in the lung. 

partner with the lead five doctors  

waiting for treatment. There is a 

Nicolas Lamontagne: “The study 

in this area.”

is on-going, we expect to see final 

results in mid-2017. So far, the data 

seem to indicate that QR-010 is 

Reassuring outcomes
What does the good news really 

safe and well tolerated.” 

mean? The outcomes are, in Lamon-

sense of urgency at ProQR – we are 

always seeking opportunities to 

move faster and to bring this drug  

to patients in less time!”

tagne’s words, “reassuring”. “It helps 

There is still a lot more to know 

The second trial, performed with 

to reassure people that we are work-

about how the drug works., Nicolas 

the nasal potential difference 

ing on something that has a poten-

insists. “We know the clinical mode 

method, better known as NPD, 

tial to change the life of CF patients. 

of action very well - people are sick 

aimed to find evidence of efficacy. 

The sooner we know that, the better 

because their CFTR protein does 

NPD is a well-accepted diagnostic 

it is for all people involved in this 

not work. We are looking further 

From ProQR’s founding in 2012, the 

translated to actual cystic fibrosis 

test that has been used before to 

huge effort. It builds confidence that 

into the molecular mode of action, 

company has embraced the RNA 

(CF) patients? The first clinical tests 

confirm the diagnosis of cystic  

QR-010 is actually going to work.”

e.g. what QR-010 does in the cell to 

technology that was discovered 

of ProQR’s molecule QR-010 seem 

fibrosis and more recently to 

make CFTR work better. The NPD 

by a scientist from Massachusetts 

to indicate it does.

assess the therapeutic benefit of 

Nicolas Lamontagne is eager to put 

study tells us that QR-010 does what 

General Hospital. The technology 

that was specifically designed for 

CF is based on the finding that the 

The trials: safety and efficacy
In 2015, ProQR’s QR-010 molecule, 

investigational agents in clinical 

the results in perspective. “Does the 

patients need. It improves their 

trials. In laboratory models of CF, 

drug reach the target? Yes. Does 

CFTR function.”

QR-010 has shown the ability to 

it engage the target? Yes. Does it 

function of a defective protein can 

based on RNA technology, has 

restore NPD toward normal levels. 

produce the effect you want to see? 

The ProQR’s QR-010 captain is 

be restored through an interven-

been put to the test in humans 

In this clinical study the effect has 

Yes. Those three answers made the 

convinced that ProQR is betting on 

tion at the RNA level. The big ques-

for the first time. The first clini-

been repeated in humans, to find 

results positive – it proves that the 

the RNA technology for the right 

tion is then: can the effects that 

cal (phase 1b) trial of QR-010 is 

similar signs of efficacy. 

fundamentals are there.” Possibly, 

reasons. “The results strengthen 

were shown in the laboratory be 

assessing the safety and tolerability 

QR-010 works? “The tests clearly 

my belief in an RNA approach. 

 
PAGE  XIII
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ANNUAL REPORT 2016

“The results 
strengthen 
MY BELIEF 
in the RNA 
approach”

Other companies have decided to 

follow our track in developing RNA 

Bringing QR‑010 to the finish line
With all the knowledge and ex-

technologies to treat CF and other 

perience – and creativity – that 

diseases. But we are definitely  

was involved in finding the mole-

operating in the forefront.”

cule, other qualities are essential 

to get this therapy to patients. 

Nicolas is convinced that, with 

“Determination is what it takes”, 

ProQR maturing, the company has 

says Nicolas. “There are dozens of 

definitely passed the age of a start-

challenges ahead. Fortunately, we 

up company. “Testing our drug on 

have many different qualities – like 

patients involves new responsibili-

scientific excellence – and a diverse 

ties and a different way of thinking. 

team at ProQR. We aim to do 

This is the time to prove that ProQR 

things right, fast and well from the 

is patient-centric, and that we care 

beginning.. Bringing QR-010 to the 

about their future and well-being.”

finish line is what we will do.” 

ABOUT NASAL 
POTENTIAL  
DIFFERENCE 

(homozygous). This indicates that the 

CFTR channels where more active 

after treatment with QR‑010. This 

effect was not observed in patients 

The CFTR protein functions as a  

with only one copy of the F508del 

salt channel in – for example –  

mutation (compound heterozygous).  

the cells in lungs and the nose. 

When someone has CF, this CFTR 

Determining the fact that QR‑010 

protein isn’t there or doesn’t 

restored the function of CFTR is 

function properly. Therefore, salts 

important because CF patients feel 

cannot move in and out of the cells 

better when the CFTR channels in 

the way they normally do. Doctors 

their lungs are more active. That 

can measure how well the channels 

is what QR‑010 needs to do! It was 

work as the movement of salts 

shown that QR‑010 does this in 

creates an electric current that can 

the noses of F508del homozygous 

be picked up with certain equipment 

patients and this is important 

(electrodes) in the nose. This 

because it’s known that the lung 

measurement procedure is called the 

cells respond very similar to cells in 

nasal potential difference (NPD) test. 

the nose. 

In ProQR’s clinical trial, investigators 

Nicolas Lamontagne, QR‑010 

used the NPD test to study whether 

captain at ProQR adds: “We are 

QR‑010 can improve the function of 

doing another study where we 

the CFTR channels in CF patients 

administered the drug in the 

with the F508del mutation. They 

lungs of 64 F508del homozygous 

measured NPD before and after 

CF patients with QR‑010 to prove 

treating the insides of CF patient’s 

that this is safe. Secondly, we will 

noses with QR‑010. After treatment, 

look at whether patients have 

the electric current was more normal 

any benefit from inhaling QR‑010 

in the noses of patients that have 

into their lungs. Results from this 

two copies of the F508del mutation 

study are expected this year.”` 

 
 
 
PAGE  XV
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PAGE XV I
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ANNUAL RE PORT 2016

“TOP-NOTCH

SCIENCE THAT
LEADS TO A
TREATMENT”

Patricia Biasutto, 
QR‑110 captain at ProQR

2017 may become a very exciting year for Patricia Biasutto, QR-110 captain. 

The work of her department, aimed at developing a novel drug for Leber’s 

congenital amaurosis Type 10 (LCA 10) patients, has progressed steadily.  

The team is expecting to start a first clinical trial in patients this year.

Patricia Biasutto, born and raised  

centres we have made tremendous 

in Venezuela and now heading  

progress and in 2017 we will take 

“A clinical  
trial SAVED 
MY LIFE. 
That event 
strengthens 
my personal 
motivation”

Until now, every one of these steps, 

ProQR was founded, Patricia was 

every new test produces another 

among the first to join. “There were 

‘green light’. “Seeing the molecule 

only five people working here.  

perform well in the ‘optic cup’ mod-

At the time, I was working in Paris. 

el is truly exciting. It is the sensation 

Taking the job interview by Skype 

that scientists, like me, flourish on”. 

did not feel right, so I volunteered 

to pay for my roundtrip to come 

Patricia calls the process of becom-

over to Leiden to have it in person. 

ing aware of the molecule’s poten-

Looking back at it, it was the right 

tial “a very fulfilling experience” and 

decision. It was like love at first 

“an amazing feeling”. “But as much 

sight with the team. There was this 

as I celebrate it, the life-changing 

immediate connection with the 

moment has yet to come. That will 

founder and CEO, Daniel de Boer, 

have to come when the molecule 

who’s initial motivation to start this 

actually proves to be effective in a 

business was to find a cure for his 

patient. We took the first step – or 

son’s disease, CF. In my four years 

For LCA 10 we had no good models 

hurdle. There is a green light, but 

at ProQR, I never once doubted  

to test this in and you cannot just 

we need more of them.”

my decision.”

take an eye from a patient! We 

worked together with the Uni-

versity College London (UCL) that 

A personal angle
In Patricia Biasutto’s mind, there 

What defines this connection, this 

common goal? “It is the shared 

recently developed this amazing 

is no doubt that the molecule will 

awareness that, in this profession 

technique where you can take a 

reach the all-important phase of 

and at this company, we truly have 

bit of skin from an LCA 10 patient 

clinical trials. Her optimism is purely 

the potential to change patients’ 

and turn the cells into stem cells 

scientifically-based, but there is 

lives. This is not just a job. I am 

and basically grow retinas in the 

also a more personal angle to this. 

thrilled that soon, we may be able 

laboratory. We were one of the 

“In short, a clinical trial saved my 

to offer patients the opportunity 

first companies to use this 3D 

life. That life event strengthens my 

that saved and shaped me. To 

organoid model and with it show 

personal motivation to get QR-110 

anyone else, clinical testing is just 

that QR-110 does exactly what we 

to the clinical trial phase ASAP.” 

a technical term. To me, it is much 

hoped it does in the closest repre-

more than that. It is hope.”

sentation of a patient’s retina.”

She explains: “When I was a child, 

The key to patient benefit
In the laboratory, the team found 

I suffered from cancer. It was a se-

vere, life-threatening type of cancer. 

“It is all about people”
ProQR may not be the largest bio-

My future was very uncertain, the 

tech company in the world today, 

ProQR’s QR-110 research and  

development to the next phase: 

confirmation that the QR-110 mole-

doctors had very limited options to 

not by far. To Patricia, the size is 

development team in Leiden  

clinical trials.”

cule restores the RNA sequence of 

treat the cancer which unfortunately 

irrelevant, as even the smallest of 

(The Netherlands), has multiple 

reasons for being optimistic. 

Obviously, as a biotech scientist 

Change the lives of patients
In search for the one molecule that 

the CEP290 protein that has a very 

did not work to stop the disease. 

biotech firms can achieve great 

important role in keeping photo- 

At that time, a new combination of 

results. “It is all about people, their 

receptor cells (rods and cones) alive 

drugs just reached the clinical trial 

knowledge and their experience. 

looking for a treatment for LCA 10, 

may change the lives of the 2,000 

in the retina. In LCA 10 patients, this 

phase. My very caring and com-

It is about putting the right teams 

she cannot confirm that there is a 

patients (see text box) that suffer 

protein is not formed. Hence, the 

mitted doctors with my parent’s 

together, to find the right entrepre-

breakthrough. Not yet. 

from LCA 10, Biasutto and her col-

photoreceptor cells in the retina 

support managed to get me on the 

neurial and scientific chemistry”. 

“We have definitely found some-

as “the amazing innovative model 

see. “Restoring production of this 

I am here today.”

leagues applied what she describes 

die and – with that – the ability to 

program. That is the reason why  

thing. ProQR’s team has developed 

of ‘optic cups’.” She explains, in a 

a molecule that shows promising 

short lecture: “Before we can test 

results in laboratory tests. In col-

our molecule in patients we first 

protein may be the key to keep the 

retinas intact and prevent blind-

ness. The next, exciting step will be 

A scientific connection
It was this event that eventually 

out boundaries, where scientists 

feel they can unleash their creativ-

laboration with various academic 

need to find out whether it works.  

clinical testing in patients.”

brought Patricia to ProQR. When 

ity. “It can only happen when you 

Patricia insists that innovation can 

only be born in environments with-

PAGE  XVII
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ANNUAL REPORT 2016

OPTIC CUP

have recreated this special tissue in 

These stem cells were then grown 

the laboratory. In order to do this we 

in presence of special chemicals that 

Human retina is a highly complex 

took skin cells from a LCA 10 patient 

trick them into developing retina 

tissue that helps us to see things. 

and reprogrammed them into stem 

like structure called optic cup.

In order to properly test QR‑110 we 

cells by expressing some key genes. 

PAGE  XVIII
Ma ga zine
ANNUAL RE PORT 2016

LCA 10 BASICS 

Leber’s congenital amaurosis Type 10

Other 
mutations

LCA 2 due to 
RPE65 mutations

~2,000

patients with  
p.Cys998X in CEP290
in Western world 

Eye

Lose sight in first 
years of life

Genetic

No therapy 
available

feel free to challenge your co-work-

ing RNA technologies – tested or 

patient-centric approach. “We 

There are over 300 genetic diseases that can cause 

In a healthy eye, the outer segment of light detecting 

ers in a professional way and 

not – to genetic diseases. You need 

want QR-110 to be administered in 

blindness and we have chosen to make these 

cells (rods and cones) in the retina are constantly 

when you know your efforts are 

strong partnerships with the best 

the most comfortable and effec-

important targets for our quest for treatments. Our 

renewed, a process in which the CEP290 protein plays a 

valued. I think ProQR does perform 

academic centres, their research 

tive way. To ensure this we are 

lead program is for Leber’s congenital amaurosis (LCA), 

major part. It facilitates the transport of new ‘building 

exceptionally well in this area – it is 

and their tools. I guess that when 

always looking to communicate 

a rare genetic eye disease characterized by progressive 

materials’ to the outer segments to keep them alive. Due 

a particularly nourishing environ-

a company has all that – and this 

with patient groups to tap into 

loss of vision. Depending on the genetic mutation 

to a defect in the DNA of LCA 10 patients, this crucial 

ment, especially when you need to 

applies to ProQR – there is a great 

their expertise. There is also a 

patients usually become completely blind during the 

protein is not produced. As a result, the light detecting 

work long hours and weekends to 

chance of success.”

practical aspect in designing a trial 

time between birth and the age of 8. As such, LCA is 

cells are not maintained and the cells die, leading to poor 

find solutions to tough problems.”

Creativity and perseverance
To Patricia Biasutto, success in 

Next steps
What are the next steps to bring 

QR-110 to patients? “In our  

and providing information that 

is accessible for people who are 

visually impaired.” 

biotech research comes from 

upcoming trial we aim to make 

For LCA 10 patients, 2017 could be 

the leading cause of genetic blindness in children.  

vision and – eventually – blindness.

We also develop programs for other inherited 

blindness called Usher syndrome and Fuchs 

ProQR is developing a novel RNA therapy for LCA 10 

endothelial corneal dystrophy.

patients, called QR‑110. The molecule is specifically 

designed to exclude the p.Cys998X mutation that is 

mixing creativity and persever-

sure our molecule is safe, but the 

a year of promise. “And of hope”, 

The key player in LCA 10 – the most severe form of LCA, 

present in the CEP290 mRNA of some LCA 10 patients. 

ance – and a little bit of luck. “You 

trial will also tell us much about 

Patricia adds. What is the ultimate 

affecting approximately 2,000 patients in the Western 

This mRNA is the ‘blueprint’ of the CEP290 protein. By 

need to have a vision of what you 

the efficacy.” 

want to achieve, as you need some 

dream? “Repairing sight for all  

patients with inherited blindness.  

world – is the CEP290 protein. This protein helps the 

fixing this blueprint a normal healthy CEP290 protein 

development of cilia in the photoreceptor cells (rods and 

will be formed that is expected to have normal function. 

sense of direction. You need to be 

Patricia stresses the importance of 

Or at least to stop the progression of 

cones) in the retina. A non-functioning CEP290 protein 

The goal of QR-110 is to stop the progression of or 

curious to find new ways of apply-

involving patients, as part ProQR’s 

these diseases in young patients.” 

causes subsequent retinal degeneration in  

LCA 10 patients. 

potentially even reverse some of the effects of LCA 10 
caused by the p.Cys998X mutation.  

PAGE  XIX
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ANNUAL REPORT 2016

PAGE XX
Ma ga zine
ANNUA L REPORT 2016

DYSTROPHIC EPIDERMOLYSIS BULLOSA

A LIFE
OF
 PAIN
AND
ITCH

Prof. dr. Jonkman, Dermatologist 
Universitair Medisch Centrum Groningen (UMCG)

Skin diseases are one of ProQR’s key therapeutic areas for RNA-based 

therapies. The lead program for dystrophic epidermolysis bullosa (DEB) 

focuses on a rare genetic disease that causes blisters, itching and severe 

pains. “Patients lead a life of pain and itch”, says Prof. dr. Marcel F. Jonkman, 

one of the top EB experts in The Netherlands. “The disease’s serious 

complications often result in skin cancer and, eventually, death.”

“I am often 
amazed by 
the POSITIVE 
ENERGY  
and MENTAL 
RESILIENCE 
that RDEB 
patients 
show”

Prof. dr. Jonkman

Prof. dr. Jonkman is the leading 

large body surfaces, as well as  

dermatologist at the UMCG Uni-

inside the body amongst many 

versity Medical Center Groningen 

other symptoms.” 

in The Netherlands. His depart-

ment runs The Netherlands’ Cen-

In daily life, any bump or rubbing 

ter for Blistering Diseases. Of the 

against the skin causes blisters 

500 patients that visit the center 

filled with fluid. Drs. J. Duipmans: 

regularly, recessive dystrophic EB 

“The blisters heal slowly and cause 

(RDEB) patients suffer the most 

scars in weeks and months. We 

severe symptoms. Many of them 

help RDEB patients deal with the 

don’t reach the age of 40. Although 

pain and the itching; these symp-

the disease itself is not life threat-

tomatic treatments are aimed at 

ening, the damages caused often 

reducing the pain and itch and 

are. Prof. dr. Jonkman: “Most RDEB 

help the wounds heal. We teach 

patients die of squamous cell 

patients and their families how to 

carcinoma, a form of skin cancer. 

deal with infection.” 

Other patients may die of heart 

and kidney problems.”

Missing adhesion molecule
Technically, the problem is the 

Recurring problem
“Caregivers know the drill. The 

recurring everyday problem of 

the RDEB patient is the wound 

absence of an adhesion molecule 

dressing changes that can take 

– type VII collagen – that functions 

many hours, sometimes using a 

as a velcro in the skin, sticking 

disinfecting chloride bath. To most 

the skin layers together. Healthy 

patients, this procedure comes 

people have this molecule that 

with excruciating pain, as dress-

grows adherence structures, but 

ings tend to stick to the wounds. 

in RDEB, this molecule is missing. 

The patients live a life of pain and 

This absence causes a wide range 

of itch that disturbs their rest at 

of effects. “Patients have a skin so 

night, leading to more blisters and 

fragile that even minor trauma like 

more pain and itch.”

rubbing may cause blistering. The 

skin disadheres in its lower layers”, 

Prof. dr. Jonkman explains.

Blisters and scars
“What we see among RDEB  

Beyond the skin 
But RDEB reaches far beyond 

problems of the skin. The damag-

ing effects of RDEB like the fusing 

of fingers and toes eventually lead 

patients”, says Nurse practitioner 

to loss of mobility. Prof. dr. Jonk-

drs. J. Duipmans, “is blisters over 

man: “To treat range of symptoms 

 
PAGE  XXI
Magazi ne
ANNUAL REPORT 2016

“It is a 
LIFE-LONG 
NIGHTMARE 
for patients 
and their 
caretakers”

Nurse practitioner 
drs. J. Duipmans

in RDEB, we work closely together 

with many specialists in the UMCG. 

Major effect on lives
Drs. J. Duipmans: “I must admit 

Under the coordination of myself 

that novice – and even professional 

and the nurse, we work together in 

– nurses sometimes find it hard to 

a team of 20 scientific disciplines, 

witness the pain and anxiety that 

from pediatricians, surgeons, 

comes with RDEB, specifically among 

orthopedists, gastroenterologists, 

young children. No matter how well 

dieticians and plastic surgeons to 

we take care of the wounds and 

rehabilitation doctors, physio- 

blisters, there will always be new 

therapists and others. To make 

ones. This skin disease has a major 

the periodical checkups as com-

effect on the lives of patients as well 

fortable as possible, RDEB patients 

as of their families. Witnessing your 

can be seen by these specialists in 

child grow up in pain, with limited 

one single visit. For any issue that 

opportunities in school and work 

cannot wait until the next visit to 

and in developing relationships 

Groningen, we offer consulting 

is tough for a parent – and to the 

via video-conferencing and other 

people that love and care about him 

means of communication.” 

or her. It is a lifelong nightmare for 

PAGE  XXII
Ma ga zine
ANNUAL RE PORT 2016

DEB BASICS 

Dystrophic Epidermolysis Bullosa

Other 
mutations

~2,000

DEB patients with exon 73 
mutation (in COL7A1) in 
Western world

Skin & 
other organs

Blistering of
skin from birth 

Genetic

No therapy 
available

QR-313 FOR DEB

skin layers together. Absence of 

aim to restore the anchoring fibrils 

anchoring fibrils make the skin very 

in patients that have a mutation in 

DEB patients miss functional struc‑

fragile. It blisters with the slightest 

exon 73 of the COL7A1 gene. The 

tures in their skin called anchoring 

friction, for example from wearing 

goal is to increase the quality of life 

patients and their caretakers.”

fibrils (see illustration) that are im‑

clothes, resulting in wounds that 

for patients by normalizing wound 

Prof. dr. Jonkman expresses a 

strong hope for a treatment for 

“Having said that, I am often 

RDEB. “RDEB is a very rare disease 

amazed and touched by the posi-

but probably the most severe 

tive energy and mental resilience 

disease in dermatology. The 

that RDEB patients show. These  

multi-organ involvement makes 

patients deserve a cure or a treat-

it, as they say, ‘the worst disease 

ment that improves their quality 

you never heard of’. I am keeping 

of life or at least makes their lives 

my fingers crossed for a treatment 

bearable. Ask any patient and he 

that reduces the blister formation 

or she will say that reducing the 

and the itching, being the primary 

formation of blister by half would 

problems of RDEB patients.” 

immensely change their lives.” 

portant for binding the different 

do not easily heal. With QR‑313 we 

healing and reducing blistering.

Anchoring
fibrils

Interstitial
collagen fibers

Keratinocytes

Healthy skin

DEB skin

DEB skin + QR-313

PAGE  XXIII
PAGE  XXII I
Magazi ne
Magazi ne
ANNUAL REPORT 2016
ANNUAL REPORT 2016

“CREATIVITY, 
PERSISTENCE 
AND DRIVE.  
IT ALL COMES  
TOGETHER  
IN ProQR”

Smital Shah, CFO ProQR

Smital Shah, Chief Financial Officer at ProQR, is excited about how the 

company has progressed. With three programs in full development and  

one with clinical data, ProQR has moved into the next phase of its existence. 

“In under five years, ProQR has evolved into a strong, emerging biotech 

company with great potential.”

Smital Shah has a 13-year track 

record of management and leader-

Clear progress in programs
After ProQR embarked on its jour-

ship in biopharma companies and 

ney less than five years ago with its 

PAGE XXIV
Ma ga zine
ANNUAL RE PORT 2016

“We will 
soon have 
a SECOND 
PROGRAM  
in clinical 
trials”

analysts on a regular basis. “They 

Before Smital Shah joined ProQR 

understand and acknowledge we 

in 2014, she was in awe when the 

are making good progress. However, 

facts about ProQR were presented 

I don’t think that what we believe is 

to her. “ProQR wanted to make a 

phenomenal data on QR-010  

difference, but I was impressed by 

– presented in October 2016 – is  

how they executed this. Two years 

reflected in the ProQR stock price 

before that, the founder – with no 

yet. With the strong clinical data 

biotech background – had ‘an idea’. 

along with our promising pipeline, 

In such a short time, ProQR had 

in my mind, this company is under-

advanced this idea into something 

valued; but biotech is an industry 

viable, with the help of visionary 

where intrinsic value is not always 

scientists and proven biotech  

reflected at all times, so we believe 

entrepreneurs like Dinko Valerio 

that this may change over time.

and Henri Termeer. I was impressed 

environment of knowledge sharing 

and saw that determination, in com-

but maintain the laser focus that 

Shah insists that ProQR has shed off 

bination with professionalism and 

comes with small dedicated teams.”

its initial label of ‘start-up company’ 

knowledge, can lead to great things.

Three promising tracks –  
more concepts developing
ProQR is financially in good shape 

with a single idea. “Biotech happens 

in a series of jumps. Most companies 

are born out of a good idea, and 

Innovation incentives
Smital Shah smiles at the idea of 

then it’s about executing and achiev-

ProQR being a Dutch company. 

and is making great progress in its 

ing proof-of-concept on that idea. 

“From a fiscal point of view, having 

programs. Shah: “What distinguishes 

We have matured and accomplished 

The Netherlands as a base makes 

ProQR from most other biotech 

this for QR-010 in patients and are 

good sense, as this country offers 

companies is that in its very short 

now evolving to the next stage.  

many tax advantages and innova-

history, it has not just one but  

We will soon have a second program 

tion incentives for a company like 

three promising tracks and more  

in clinical trials and possibly a third 

ours. Also, the low cost of operating 

concepts in early stage of develop-

next year. Ultimately we are look-

a business makes The Netherlands 

ment, along with the strong drive to 

ing forward and building towards 

very attractive, as compared to the 

make a difference for patients with 

developing ProQR as a commercial 

U.S. Every time I fly over to Amster-

severe genetic rare diseases. That is 

company in the future and truly have 

dam from Palo Alto, I notice how 

pretty unique.”

an impact on patient lives.”

tall people are, haha, but also their 

According to ProQR’s CFO, knowing 

that after QR-010, the QR-110 & 

Building on the success of RNA
To investors, Shah pitches ProQR as 

easy, up front and refreshing way 

of expressing opinions and ideas.  

That, in combination with the  

investment banking, with particular 

QR-010 program for cystic fibro-

QR-313 programs came from the 

a strong, patient-focused company 

access to biotech talents that  

experience in financial strategy and 

sis, the contours of our final goal 

company’s in-house innovation unit 

that is successful in applying RNA 

ProQR has in this market, is an 

capital markets. Looking back on 

are beginning to show. Two more 

is an important trust-builder among 

therapeutics. “Interfering in a disease 

important advantage.” 

her first two and a half years with 

programs have entered the devel-

ProQR’s several stakeholders – im-

at the RNA level has come a long way 

ProQR, she says: “This is a compa-

opment stage. QR-110 for Leber’s 

portantly, the patients, the scientific 

with 5 approved RNA products that 

“It is part of ProQR’s DNA, just like 

ny that has moved from exploring 

congenital amaurosis Type 10 and 

community, our employees as well 

have come before us. We have built 

qualities such as creativity, per-

one ‘promising’ idea to executing 

QR-313 for dystrophic epidermoly-

as others like investors and busi-

on this success and believe that  

sistence and drive. They all come 

on a global clinical program from 

sis bullosa.

that idea and then advancing  

ness partners. “This shows we are 

ProQR is a front runner in the next 

together in ProQR, in this unique 

serious about targeting unmet med-

wave of advancement in RNA thera-

way of looking at the world and at 

other programs in the pipeline 

With the CF program as its shining 

ical needs in rare genetic diseases.”

peutics. We are grateful that so much 

the job at hand – being prepared to 

in just 4 short years. With three 

example of what you can do with a 

programs in development we are 

strong focus, ProQR wants to create 

heading in the right direction,  

the same dynamics for the other 

Strong data, promising pipeline
Being the CFO of ProQR, Smital 

us. We are building on their evidence 

this one goal.” 

and results, to achieve our own goals 

work has been done by others before 

give everything you have to get to 

with more to come.” 

programs. “We wanted to create an 

Shah meets investors and industry 

in our patient-focused strategy.”

ANNUAL
REPORT 
2016

ANNUAL

REPORT 

2016

Acting in the interest of patients

PAGE 1 / 91 
Table of Contents 
ANNUAL REPORT 2016 

Table of Contents 

Message from the CEO 

Key Figures 

Management Board 

Supervisory Board 

Management Board Report 

Supervisory Board Report 

Corporate Governance 

Risk Management 

Financial Statements 2016 

  2 

  3 

  4 

  5 

  5 

  32 

  35 

 45 

 47 

 
 
 
 
 
 
PAGE 2 / 91 
Message from the CEO 
ANNUAL REPORT 2016 

Message from the CEO 

2016 was an important year for ProQR in which we made significant progress on our 

mission to bring life changing therapies to patients in need. We advanced our pipeline of 

RNA therapies targeting severe genetic rare diseases and further enforced our strong team 

of dedicated ProQRians to bring our company past exciting milestones in the upcoming 

years. First of all we completed the first clinical trial of OR-010 in patients with cystic fibrosis, 

a major achievement for a young company like ProQR. The results from this early trial told 

us that QR-010 has potential to become a meaningful therapy for patients. A second clinical 

trial is ongoing. The first part of the study was completed and showed that QR-010 was safe 

and well tolerated. During 2016 the FDA granted the QR-010 program Fast Track 

designation. These important events have encouraged us and the medical community in 

the further development of QR-010 and our quest to help CF patients.  

The second leg of our company is developing therapies for genetic blindness. The first 

program in this is called QR-110 and targets subtype 10 of the most common genetic 

blindness in children, Leber’s congenital amaurosis (LCA). We are currently doing all the 

preparations to start a clinical trial of QR-110 in the first half of 2017. During several 

scientific conferences in 2016 we have published very promising pre-clinical results showing 

this second molecule can target and restore the underlying defect causing LCA 10. In the 

same year both the FDA and the EMA have granted QR-110 Orphan Drug designation. 

During 2016 we initiated a first development program for a debilitating skin disease, an 

area with a high unmet medical need. Our first investigational product in this area is QR-

313, and is targeting a disorder called dystrophic epidermolysis bullosa. We are planning to 

rapidly move the program through the first stages of development in 2017 to start and 

complete a clinical trial in 2018. 

The advancement of this third development program shows the exceptional productivity of 

the innovation unit since its inception in 2014. The goal of the innovation unit is to expand 

our pipeline by discovering promising new therapeutics and RNA technologies. During our 

first Research & Development day in 2016 we revealed some of the other programs that 

are being pursued in the innovation unit including additional programs for inherited 

blindness, Usher syndrome and Fuchs endothelial corneal dystrophy (FECD), and a program 

for beta-amyloid related disorders. 

We could not have achieved any of these results without the support from the patients and 

medical research teams that participated in our clinical trials, our academic partners, our 

shareholders, and of course the tireless efforts of all ProQRians. Thank you for making this 

another meaningful year. 

Daniel de Boer 

 
 
 
PAGE 3 / 91 
Key Figures 
ANNUAL REPORT 2016 

Key Figures 

dsssds  

2016 

2015 

Result from continued operations (in € 1,000) 

Net revenue 

Other income 

Research and development costs 

General and administrative costs 

Operating result 

Net result 

Balance sheet information (in € 1,000) 

Non-current assets 

Current assets 

Total assets 

Shareholders’ equity 

Non-current liabilities 

Current liabilities 

Cash flows (in € 1,000) 

Net cash used in operating activities 

Net cash used in investing activities 

Net cash generated by financing activities 

Ratio’s (in %) 

Current ratio 

Solvency 

Figures per share 

Weighted average number of shares outstanding 

Basic and diluted earnings per share (in €) 

Cash flow per share (in €) 

Employees 

Average number of staff for the period 

-- 

1,828 

(31,923) 

(9,478) 

(39,573) 

(39,103) 

3,528 

62,015 

65,543 

53,136 

5,697 

6,710 

(34,221) 

(2,539) 

357 

-- 

3,235 

(23,401) 

(6,837) 

(27,003) 

(20,832) 

2,340 

97,769 

100,109 

89,799 

4,824 

5,486 

(24,232) 

(1,324) 

1,620 

9.2 

81.1 

17.8 

89.7 

23,346,507 

23,343,262 

(1.68) 

(1.56) 

(0.89) 

(1.03) 

133.4 

86.1 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
PAGE 4 / 91 
Management Board 
ANNUAL REPORT 2016 

Management Board 

We have a two-tier board structure consisting of our Management Board (raad van bestuur) and a separate 

Supervisory Board (raad van commissarissen). The Management Board operates under the chairmanship of 

the Chief Executive Officer and shares responsibility for the deployment of ProQR’s strategy and policies, and 

the achievement of its objectives and results.  

Under Dutch Law, the Management Board has ultimate responsibility for the management and external 

reporting of the Company and is answerable to shareholders at the General Meeting of Shareholders. 

Pursuant to the two-tier corporate structure, the Management Board is accountable for its performance to a 

separate and independent Supervisory Board. 

The following table sets out information with respect to each of our Management Board members, their 

respective ages and their positions at the Company as of the date of this annual report. 

Name 

Gender 

Date of Birth 

Position 

Date of  
Appointment 

Term 
expires 

Daniel de Boer 

Male 

April 12, 1983 

Chief Executive Officer 

February 21, 2012 

René Beukema 

Male  March 26, 1964 

Chief Corporate 
Development Officer and 
General Counsel 

April 17, 2014 

2018 

2018 

The following sets forth biographical information regarding our management board members. 

Daniel de Boer is our founding Chief Executive Officer and has served as such since our incorporation in 

February 2012. Mr. de Boer is a passionate and driven entrepreneur and advocate for CF patients, and has 

assembled an experienced team of successful biotech executives as co-founders and early investors. As a 

serial entrepreneur in IT, he founded and led a number of tech companies through phases of growth, 

initiating development and launch of several IT related products in several European countries. Prior to 

founding ProQR, Mr. de Boer served as a founder and Chief Executive Officer of RNA Systems, founder and 

Chief Executive Officer of PC Basic, and founder and Chief Executive Officer of Running IT. Mr. de Boer is 

responsible for the overall strategy and general business in the company. 

René Beukema has served as our Chief Corporate Development Officer and General Counsel since April 2014. 

Mr. Beukema joined us in September 2013 and is a seasoned in-house corporate lawyer in the Dutch 

biotechnology arena. Prior to joining us, Mr. Beukema served as general counsel and corporate secretary of 

Crucell N.V. for twelve years, following his experience as a senior legal counsel at GE Capital / TIP Europe and 

legal counsel at TNT Express Worldwide. Mr. Beukema was also a venture partner of Aescap Venture, a life 

sciences venture capital firm. Mr. Beukema is the co-founder and advisor of Mytomorrows, a Dutch life 

sciences company, and a member of the VU Medical Cancer Center children in Amsterdam. He holds a post-

doctoral degree in corporate law from the University of Nijmegen in co-operation with the Dutch Association 

of In-house Counsel (Nederlands Genootschap van Bedrijfsjuristen) and a Master’s degree in Dutch law from 

the University of Amsterdam.  

 
 
 
 
 
 
 
 
 
 
 
 
PAGE 5 / 91 
Supervisory Board 
ANNUAL REPORT 2016 

Supervisory Board 

The Supervisory Board supervises the policies of the Management Board and the general course of affairs of 

ProQR and advises the Management Board thereon. The Supervisory Board, in the two-tier corporate 

structure under Dutch law, is a separate and independent corporate body. 

The following table sets forth information with respect to each of our supervisory board members and their 

respective ages as of the date of this annual report. The terms of office of all our supervisory board members 

expire according to a rotation schedule drawn up by our supervisory board. 

Our supervisory board is currently composed of the following members, all of whom are independent under 

applicable NASDAQ standards and all of whom, with the exception of Mr. Antoine Papiernik are independent 

under the Dutch Corporate Governance Code (DCGC): 

Name 

Gender  Nationality 

Date of Birth 

Position  Date of Appointment 

Term expires 

Dinko Valerio 

Male 

NL 

August 3, 1956 

Chairman 

January 1, 2014 

Alison Lawton 

Female 

US  September 26, 1961 

Member 

September 17, 2014 

Antoine Papiernik  

Henri Termeer 

James Shannon 

Paul Baart 

Male 

Male 

Male 

Male 

FR 

NL 

GB 

NL 

July 21, 1966 

Member 

January 1, 2014 

February 28, 1946 

Member 

January 1, 2014 

June 5, 1956 

Member 

November 9, 1950 

Member 

June 21, 2016 

June 10, 2015 

2020 

2018 

2017 

2020 

2020 

2019 

The following sets forth biographical information regarding our supervisory board members.  

Dinko Valerio is one of our founders and currently serves as the chairman of our supervisory board. Mr. 

Valerio has served on our supervisory board since January 2014. Mr. Valerio is a scientist and an experienced 

biotech entrepreneur with experience in both public and private companies as CEO and board member. Mr. 

Valerio is founder and former CEO of Crucell N.V., a Dutch biotech company, and founder and general 

partner of Aescap Venture, a life sciences venture capital firm. In 1999, Mr. Valerio was one of the founders of 

Galapagos Genomics N.V., a spinout from Crucell N.V. which develops novel mode of action medicines. 

Adding to his corporate experience, Mr. Valerio is a professor in the field of gene therapy of the 

hematopoietic system at the University of Leiden. He received his Master of Science degree in Biology from 

the University of Amsterdam in 1982 and completed his Ph.D. in Molecular Genetics with Honors at the 

University of Leiden in 1986. Mr. Valerio also was a visiting scientific specialist at Genentech Inc., San 

Francisco in 1985 and a postdoctoral fellow at the Salk Institute, San Diego from 1986 to 1987. He is an 

author on more than 100 articles in peer-reviewed journals and an inventor on 11 patent-families. 

Alison Lawton has served on our supervisory board since September 2014. Ms. Lawton is currently the Chief 

Operating officer of Aura Biosciences Inc. From January 2013 to January 2014, Ms. Lawton served as Chief 

Operating Officer of OvaScience, Inc., a public life sciences company. From 1991 to 2013, Ms. Lawton worked 

at various positions of increasing responsibility at Genzyme Corporation, or Genzyme, and subsequently at 

Sanofi-Aventis, following its 2011 acquisition of Genzyme, each a global biopharmaceutical company. Ms. 

Lawton served as head of Genzyme Biosurgery, where she was responsible for Genzyme’s global orthopedics, 

surgical and cell therapy and regenerative medicine businesses. Prior to that, Ms. Lawton oversaw Global 

Market Access at Genzyme, which included Regulatory Affairs, Global Health Outcomes and Strategic Pricing, 

Global Public Policy, and Global Product Safety & Risk Management. Before joining Genzyme, Ms. Lawton 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
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worked for seven years in the United Kingdom at Parke-Davis, a pharmaceutical company. Ms. Lawton serves 

on the board of directors of Verastem, Inc., a public biopharmaceutical company. In 2016 she joined the 

board of directors of CoLucid Pharmaceuticals. She also served on the board of directors of Cubist 

Pharmaceuticals for three years until its acquisition by Merck &Co., Inc. in 2015. She is member of the 

Corporate Advisory Board of X4 Pharmaceuticals. She is past President and Chair of the Board of Regulatory 

Affairs Professional Society and past FDA Advisory Committee member for Cell and Gene Therapy 

Committee. She earned her BSc in Pharmacology, with honors, from King’s College London. 

Antoine Papiernik has served on our supervisory board since January 2014. Mr. Papiernik is managing partner 

at Sofinnova Partners, which he joined in 1997. Mr. Papiernik has been an initial investor and active board 

member in public companies like Actelion, Addex, Auris Medical, Orexo, NovusPharma (then sold to CTI), 

Movetis (then sold to Shire), Mainstay, Pixium and Stentys, which went public respectively on the Zurich Stock 

Exchange, the NASDAQ Global Market, the Stockholm Stock Exchange, the Milan Nuovo Mercato, the Belgium 

Stock Exchange, the Dublin Stock Exchange and EuroNext Paris, in Cotherix (initially NASDAQ listed, then sold 

to Actelion), Core Valve (sold to Medtronic), Fovea (sold to Sanofi Aventis) and Ethical Oncology Science (EOS, 

sold to ClovisOncology). Mr. Papiernik has also invested in and is a board member of private companies MD 

Start, ReCor, Shockwave Medical and Reflexion Medical, Gecko Biomedical and Rgenix. Mr. Papiernik has an 

MBA degree from the Wharton School of Business, University of Pennsylvania.  

Henri Termeer is vice chairman and has served on our supervisory board since January 2014. From October 

1983 to June 2011, Mr. Termeer served as chairman, president and chief executive officer of Genzyme 

Corporation. For ten years prior to joining Genzyme, Mr. Termeer worked for Baxter International 

Laboratories, Inc., a manufacturer of human health care products. Mr. Termeer resigned from Genzyme in 

June 2011 following the acquisition of Genzyme by Sanofi. Widely acknowledged for his contributions to the 

biotechnology industry and health care field, Mr. Termeer is active in the areas of humanitarian assistance, 

policy issues, and innovation in providing access to health care. He is a member of the board of each of 

Massachusetts General Hospital and Partners HealthCare and a member of the board of fellows of Harvard 

Medical School. Mr. Termeer is also a member of the board of the Massachusetts Institute of Technology and 

serves on its Executive Committee and a board member of the Biotechnology Industry Organization (BIO). He 

is a board member of the New England Healthcare Institute, a nonprofit, applied research health policy 

organization he was instrumental in founding and on the boards of Boston Ballet, Museum of Science, WGBH 

and Project Hope. Mr. Termeer is also currently a board member of Abiomed Inc., Aveo Pharmaceuticals, 

Moderna Therapeutics and was a board member of Allergan, Inc. from 2014 through its acquisition by Actavis 

in March 2015. Mr. Termeer was chairman of the Federal Reserve Bank of Boston’s board of directors from 

2010-2011. Mr. Termeer studied economics at the Economische Hogeschool (Erasmus University, the 

Netherlands) and earned an MBA from the Darden School at the University of Virginia. 

James Shannon, MD has served on our Supervisory Board since June 2016. Mr. Shannon has had an extensive 

career in drug development and pharma. From 2012 until his retirement in 2015, Mr. Shannon was Chief 

Medical Officer at GlaxoSmithKline. Prior to that he was Global Head of Pharma Development at Novartis and 

Senior Vice-President, Clinical Development at Sterling Winthrop Pharmaceuticals. He held board positions at 

companies including Biotie, Circassia, Crucell, Endocyte, MannKind and Cerimon Pharmaceuticals. He 

received his undergraduate and postgraduate degrees at Queen's University of Belfast and is a Member of 

the Royal College of Physicians (UK). Mr. Shannon currently holds board positions at Mannkind Corp (USA), 

myTomonows (NL) and lmmodulon (UK). 

 
 
 
 
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Paul Baart has served on our supervisory board since June 2015. Mr. Baart made his career in public 

accounting in both the Netherlands and the USA. At PwC the Netherlands he served on the management 

board and the supervisory board. He was also a member of the global board of PwC International. He has 

served many large (listed) and international clients in various industries. He held professional qualifications 

both in the Netherlands and in the USA. He was chairman of Royal NIVRA, the Dutch Institute of Registered 

Accountants (now NBA), member of the Dutch Council on Annual Reporting (RJ) and supervisory board 

member of Nyenrode Business University. Present roles include outside member Enterprise Chamber 

Amsterdam Court of Appeal (Ondernemingskamer) and chairman Supervisory Board Grant Thornton the 

Netherlands. He studied business economics at the Vrije Universiteit in Amsterdam, where he also passed the 

Registeraccountantsexam. 

 
 
 
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Management Board Report 

The Company 

ProQR Therapeutics N.V., or “ProQR” or the “Company”, is dedicated to changing lives through the creation of 

transformative RNA medicines for the treatment of severe genetic rare diseases (sometimes called orphan 

diseases) such as cystic fibrosis, Leber’s congenital amaurosis Type 10 and dystrophic epidermolysis bullosa. 

Based on our unique proprietary RNA platform technologies we are growing our pipeline with patients and 

loved ones in mind. 

We were incorporated in the Netherlands, on February 21, 2012 and reorganized from a private company 

with limited liability to a public company with limited liability on September 23, 2014. Legal demerger of our 

Company was effectuated as per June 30, 2015. Our Company has its statutory seat in Leiden, the 

Netherlands. The address of its headquarters and registered office is Zernikedreef 9, 2333 CK Leiden, the 

Netherlands.  

Since September 18, 2014, our ordinary shares have been listed on the NASDAQ Global Market under the 

ticker symbol PRQR. 

Operations 

We are an innovative biopharmaceutical company that discovers and develops RNA therapeutics for a better 

future for patients with severe genetic rare diseases. Our team of dedicated ProQRians works closely with 

patient groups and our academic partners and this has led to many successes in the short life-span of the 

company. Our lead program targets the most common mutation in CF and because of an aggressive and 

innovative development strategy positive data from a biomarker study in CF patients was presented in 2016. 

A second clinical trial in CF patients is ongoing and is expected to read out in mid-2017. Our first program 

targeting an inherited blindness is expected to enter clinical trials during the first half of 2017 and more 

programs in this therapeutic area are being advanced. The third area that we focus on are debilitating skin 

diseases, for which we are developing several programs. The first of these is expected to enter clinical trials in 

2018. Our diversified pipeline that we presented during a Research & Development Day in 2016 is the result 

of our in-house discovery engine that we call the innovation unit. This group has been very productive in 

discovering new programs based upon our toolbox of RNA technologies that we have in-licensed or 

developed in-house. We currently have discovery programs including programs in Usher syndrome, Fuchs 

endothelial corneal dystrophy (FECD), Huntington’s disease, amyloid beta related disorders and Friedreich’s 

ataxia. We believe our strong team, excellent partners and our extensive pipeline will lead to a sustainable 

future for our company and to accomplish our quest to make a meaningful impact on the lives of patients in 

need. 

QR-010 and Cystic Fibrosis (CF) 

CF is a genetic disease that causes early morbidity and mortality. CF currently has no cure. The median age of 

death for CF patients is 30 years or less, and more than 90% of CF patients die from respiratory failure. To 

date, all but two of the therapies approved to treat CF patients are designed to treat the symptoms of CF 

rather than address the underlying cause. CF is caused by mutations in the gene that encodes for a protein 

called cystic fibrosis transmembrane conductance regulator, or CFTR. Although there are more than 1,900 

different genetic mutations that cause CF, the F508del mutation that we are targeting is the most prevalent 

and is present in approximately 85% of all CF patients in the Western world and approximately 65,000 

patients worldwide. In CF patients, this mutated gene and the resulting defective protein lead to the 

dysfunction of multiple organ systems, including the lungs, pancreas and gastrointestinal tract. In the lung 

 
 
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airways, absence of functional CFTR protein leads to unusually thick, sticky mucus that clogs the lungs and 

increases vulnerability to chronic, lung-damaging infections.  

Our lead product candidate in the CF space, QR-010, a first-in-class RNA-based oligonucleotide, is designed to 

address the underlying cause of the disease by targeting the mRNA defect encoded by the F508del mutation 

in the CFTR gene of CF patients and restoring CFTR function. QR-010 is designed to be self-administered 

through a small, handheld aerosol delivery device, or nebulizer, in the form of a mist inhaled into the lungs. In 

pre-clinical studies we have shown this method could allow maximum exposure of QR-010 to the primary 

target organ, the lung, as well as significant exposure to other affected organs through systemic absorption 

into the blood. Based on our extensive pre-clinical studies on safety, delivery and efficacy in relevant cell and 

animal models we started two global clinical studies of QR-010 in 2015. During the North American Cystic 

Fibrosis conference (NACFC) from October 26 – 29, 2016 we presented positive results from PQ-010-002, a 

proof-of-concept trial demonstrating that QR-010 restores CFTR function in the nasal linings of patients that 

are homozygous (who carry two allelic copies) of the F508del mutation. CFTR is the protein channel that is 

defective in patients with CF, and presence or absence of function of CFTR can be measured by an important 

biomarker called the nasal potential difference, or NPD, assay. Following four weeks of topical therapy, QR-

010 improved the CFTR-mediated total chloride response, a direct measure of CFTR function. This was 

confirmed by the restoration of other indicators of CFTR function, such as the sodium channel activity. In 

subjects that were compound heterozygous (who carry one copy of the F508del mutation and one other 

disease causing mutation), no meaningful difference was measured. QR-010 was observed to be well-

tolerated in all subjects. 

Besides the completed NPD trial, we are running a second clinical trial. This Phase 1b clinical trial, which we 

refer to as PQ-010-001, is a randomized, double-blind, placebo-controlled, 28-day dose-escalation trial that is 

being conducted in 27 sites in North America and Europe. The primary endpoint of the trial is to evaluate the 

safety, tolerability and pharmacokinetics, of single and multiple ascending doses of inhaled QR-010 in 

approximately 64 CF patients carrying two copies (homozygotes) of the F508del mutation. This trial will also 

assess a number of exploratory efficacy endpoints, although the trial is not powered for statistical 

significance on these endpoints. The single dose portion of the trial has been completed. No dose-limiting 

toxicity was observed up to the highest dose tested. We expect to report top-line data from the full trial in 

mid-2017. 

In July 2016, QR-010 received Fast Track designation from the FDA for the treatment of patients with CF due 

to the F508del mutation. Fast Track designation is granted by the FDA to drugs that are under development 

for serious conditions and have the potential to fulfill an unmet medical need. It was established with the 

intention to bring promising drugs to patients sooner by facilitating the development with more frequent FDA 

interactions and expediting the review process.  

QR-010 has been granted orphan drug designation in the United States and the European Union for CF. 

Generally, if a product with an orphan drug designation subsequently receives the first marketing approval 

for the indication for which it has such designation, the product is entitled to a period of market exclusivity. 

This exclusivity precludes the U.S. Food and Drug Administration, or FDA, or the European Medicines Agency, 

or EMA, as applicable, from approving another marketing application for the same or, in the European Union, 

a similar drug for the same indication for that time period, unless the later product is clinically superior. 

Orphan drug designation does not convey any advantage in, or shorten the duration of, the regulatory review 

and approval process.  

QR-110 and Leber’s Congenital Amaurosis Type 10 (LCA 10) 

LCA is the most common genetic cause of blindness in childhood. LCA is caused by a genetic defect in 19 or 

more associated genes. Classification of LCA is based on the disease causing gene. The most frequently 

 
 
 
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mutated LCA gene in LCA patients in North America and Europe is CEP290 that is associated with LCA Type 10 

(LCA 10). The most common mutation is the p.Cys998X (also known as c.2991+1655A>G) in the CEP290 

(Centrosomal protein of 290 kDa) gene. Although diagnosis rates vary, based on our estimations we believe 

this mutation occurs in approximately 2,000 patients in the Western world. Most patients affected by this 

mutation lose sight in the first few years of life. There is currently no disease modifying therapy available on 

the market or being tested in clinical development for this specific subtype of the disease. In LCA 10 patients, 

this mutation leads to significant decrease of active CEP290 protein in the photoreceptor cells in the retina in 

the eye. The absence of this essential protein causes blindness. 

Our lead product candidate in the LCA 10 space, QR-110, a first-in-class oligonucleotide, is designed to treat 

the disease by repairing the underlying cause in the mRNA, which results in the production of wild-type 

CEP290 protein. The p.Cys998X mutation is a substitution of one nucleotide in the pre-mRNA that leads to a 

defective mRNA and non-functional CEP290 protein. QR-110 is designed to bind to the mutated location in 

the pre-mRNA, thereby leading to normally spliced or wild-type mRNA, which could produce wild-type or 

normal protein. QR-110 is designed to be administered through intravitreal injections in the eye. We believe 

the activity in pre-clinical models of LCA 10 provides support for the clinical development and therapeutic 

potential of QR-110. In studies conducted with QR-110 using relevant pre-clinical LCA 10 models, QR-110 was 

observed to restore CEP290 wild-type mRNA and protein levels. It was observed that QR-110 restored CEP290 

mRNA and protein levels in primary LCA 10 fibroblasts from patients that are homozygous for the p.Cys998X 

mutation to approximately 100% of wild-type and to approximately 50% of wild-type in cells from compound 

heterozygous patients. It was also observed that QR-110 reaches the correct layer of the retina (the outer 

nuclear layer) after administration by intravitreal injections. In a 3D optic cup organoid model QR-110 showed 

restoration of CEP290 wild-type mRNA in a dose dependent manner. In the first half of 2017 we intend to 

start our first clinical trial directly in LCA 10 patients. There is recent precedent for an accelerated 

development path in another LCA mutation, and we believe this accelerated development pathway can also 

be applied to QR-110. 

QR-110 has been granted orphan drug designation in the United States and the European Union for LCA.  

QR-313 and Dystrophic Epidermolysis Bullosa (DEB) 

Dystrophic epidermolysis bullosa (DEB) is a genetic orphan disease of the skin and other mucosal 

membranes. The hallmark of the disease is severe blistering and poorly healing wounds that result from 

minimal pressure.  Patients with the recessive form of DEB (RDEB) have a limited life expectancy and low 

quality of life. Patients with the dominant form (DDEB) have variable expression of the disease but is also 

associated with significant morbidity. There is currently no treatment for DEB. Aggressive and costly palliative 

care provided to these patients does not address the underlying cause of the disease. DEB is caused by 

mutations in the COL7A1 gene which leads to an absence of functional collagen type VII (C7) protein which is 

essential for formation of anchoring fibrils that link the epidermis to the dermis.  

We are developing a single-stranded oligonucleotide, QR-313, for patients with DEB caused by mutations in a 

specific part of the COL7A1 gene called exon 73. QR-313 is designed to exclude exon 73 from the COL7A1 

mRNA. Skipping of exon 73 leads to an mRNA that is translated into a truncated, but functional COL7A1 

protein that is able to form anchoring fibrils that should improve the stability of the skin. There are multiple 

mutations associated with DEB, several of which lie within exon 73.   

QR-313 is being formulated in a hydrogel that will be applied topically to existing wounds in patients with 

DEB.  QR-313 is designed to restore functional COL7A1 protein with the aim to facilitate wound healing and 

protect against future blistering. In pre-clinical models skipping of exon 73 by QR-313 has been observed in a 

3D human full thickness skin model. If this exon skipping approach is proven to be of benefit for DEB 

patients, there may be other COL7A1 mutations that can be targeted with an approach similar to QR-313. 

 
 
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Innovation pipeline 

The innovation unit is our internal discovery engine, which we use to discover additional molecules through 

internal research and external collaborators. These pipeline programs are based on our multiple RNA 

technologies that were discovered internally or in-licensed. We have a rigorous evaluation process in 

identifying programs that are ready to leave the innovation unit and move into development. The criteria 

include established genetic causality, ability to deliver to the target organ, intellectual property protection, 

strong pre-clinical proof of concept, and a high unmet need. Our early stage programs are in various stages 

of discovery and target different severe genetic disorders where we believe our technologies have the 

potential to make a life altering impact for affected patients. These include programs for Usher syndrome 

and Fuchs endothelial corneal dystrophy (FECD) both areas of ophthalmology with high unmet medical need. 

We further have programs in our central nervous system, or CNS, franchise for Huntington’s disease and 

amyloid beta related disorders. In our neuromuscular franchise we have a program for Friedreich’s ataxia.  

Our Strategy  

We are dedicated to improving the lives of patients and their loved ones through the development of RNA-

therapies for severe genetic orphan diseases. We have an initial focus on patients with CF, LCA 10 and DEB. 

Key elements of our strategy include:  

 

Develop drugs for patients in need. Through our patient-centric approach we work to develop best-in-

class therapies and to advance the understanding of conditions that we target. As RNA Therapies have 

become an established modality we are translating new applications in a pipeline of products for 

patients suffering from rare diseases. We believe this strategy enables us to build a sustainable 

independent business. 

 

Independently develop and commercialize QR-010 for the treatment of CF. Our lead product candidate 

in the CF space, QR-010, has generated compelling data in pre-clinical studies and a first clinical trial in 

CF patients. We believe these data support the potential of QR-010 as a disease-modifying therapy for 

CF patients. We are currently running a second global clinical trial of QR-010 that will enroll 

approximately 64 CF patients with two copies of the F508del mutation. Top-line data is expected in mid-

2017. We are studying applications of RNA technologies for CF mutations other than F508del. We intend 

to commercialize QR-010 ourselves, if approved, and retain all commercial rights in major markets. 

 

Rapidly advance our ophthalmology franchise, including QR-110 for the treatment of LCA 10. We 

recognize the great opportunity for oligonucleotides in the ophthalmology space and therefore have 

established an ophthalmology franchise that now has one program in development and several in the 

discovery pipeline. These include LCA 10, Usher syndrome and Fuchs endothelial corneal dystrophy 

(FECD). We are developing QR-110 to treat patients with the most common mutation causing LCA, the 

leading genetic cause of blindness in childhood. We conducted further pre-clinical studies during 2016 

and expect to start our first clinical trial in LCA 10 patients in the first half of 2017. 

 

Utilize our proprietary RNA technologies and know-how to develop additional product candidates 

targeting genetic diseases with high unmet medical need. We are developing a product pipeline 

targeting severe genetic diseases that have significant unmet need and are caused by mutations that we 

believe can be treated with our RNA technologies. We are currently working on approximately 100 

potential target indications in several therapeutic areas and have organized our discovery effort in 

franchises such as respiratory, ophthalmology, dermatology, CNS and neuromuscular disorders.  

 

Leverage our pipeline through considering out-licensing, spinouts or collaborative partnerships. We 

plan to continue to advance the programs and technologies in our discovery pipeline and ensure that 

these programs have the potential to make an impact for patients in these areas of unmet need, we will 

consider strategic alternatives that include spinouts, out-licensing or collaborative partnerships with 

pharmaceutical companies. These partnerships may provide us with further validation of our approach, 

funding to advance our product candidates and access to development, manufacturing and commercial 

expertise and capabilities.  

 
 
 
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Our RNA Technologies  

Genes are the specific sequences of DNA that provide the blueprint used by the human body’s cells to make 

proteins, which are enzymes or other molecules in cells that serve a functional purpose. Each gene consists of 

a specific sequence of nucleotides that leads to the production of a specific protein. The gene’s coding DNA 

sequence is transcribed into mRNA, which is subsequently translated into the specific protein. A mutation, or 

defect, in a specific gene can result in the transcription of abnormal mRNA, which then can produce a 

defective, misfolded or truncated protein that is unable to carry out its normal function.  

In the maturing RNA therapeutics space and the developments in understanding their potential, we have 

gathered a toolbox of different novel RNA technologies with which we believe we target defective mRNA in 

order to restore protein functionality.  Our goal to restore translation of functional proteins is unlike other 

approaches in the RNA therapeutics field, such as RNAi and antisense that use RNA molecules to 

downregulate genes. Our molecules are single-stranded RNA-based oligonucleotides that are chemically 

modified (phosphorothioate backbone and 2’ O-methyl modifications) so that no vector or envelope is 

needed for delivery. We believe these RNA approaches will allow us to develop novel therapies for genetic 

disorders that are currently untreatable or have limited effective treatment options.  

Our product candidates 

In selection of discovery programs to bring into our development pipeline we apply a rigorous process of 

review by a committee of internal and external scientific experts and thought leaders to all key aspects of a 

program. Among others we look at the following criteria: 

 

 

High unmet medical need 

A pre-clinical proof-of-concept that shows strong promise for translation to the clinic 

  Well understood relationship between the genetic defect and the disease manifestations 

 

 

Feasibility of delivery to target organ(s) 

Strong IP position and initial freedom to operate established 

We believe our current pipeline represents a mix of high-value indications where we can make a big impact to 

the lives of patients. 

Program pipeline 

 
 
 
 
 
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Patient Centric Approach 

ProQR aims to develop best-in-class therapies as well as to improve patient care through awareness, 

education, and advancing the understanding of conditions that we target. In order to achieve this goal, ProQR 

strives to integrate the patient voice into our decision-making throughout the drug development process. 

Because we believe that a patient-centric strategy is crucial to our success, we have established the Patient 

and Medical Community Engagement (PMCE) team. This dedicated team’s purpose is to listen to and 

represent the patient voice internally as well as to collaborate externally with the communities we serve. 

Cystic Fibrosis  

Cystic Fibrosis Background  

CF is the most common fatal inherited disease in the Western world and affects an approximately 65,000 

patients worldwide. There is no cure for CF. CF patients require lifelong treatment with multiple daily 

medications, frequent hospitalizations and ultimately lung transplants, which can extend life for five years on 

average. The quality of life for CF patients is compromised by approximately four hours of self-care per day 

and frequent outpatient doctor visits and hospitalizations.  

CF is caused by mutations in a gene that encodes the CFTR protein. The CFTR protein channel regulates the 

movement, or efflux, of specific ions in and out of the cells of organs like the lungs, pancreas and 

gastrointestinal tract. Through regulation of these ions, the amount of salts in the fluid both inside and 

outside the cell remains balanced. In CF patients, however, the CFTR protein is defective and cannot perform 

its normal function of transporting ions across the cell membrane, and this results in an environment 

characterized by thick mucus in vital organs such as the lung, the pancreas and the gastrointestinal tract. The 

figure below illustrates a defective CFTR protein hampering the efflux of chloride in lung epithelial cells.  

Chloride ion flow by wild-type CFTR and F508del CFTR 

Wild-type 

F508del 

The lack of functional CFTR in CF 

patients is particularly problematic in 

the lungs, where the build-up of 

thick mucus obstructs parts of the 

lung, allows bacteria to grow 

unfettered and impairs the 

functionality of the local immune 

system. Of all the manifestations of 

CF, lung disease is the most critical 

and is characterized by a 

combination of airway obstruction, 

infection and inflammation such that 

more than 90% of all CF patients die 

of respiratory failure. In the 

pancreas, the buildup of mucus 

prevents the release of digestive 

enzymes that help the body break down food and absorb important nutrients. In the GI tract, the thick mucus 

leads to impaired ability to absorb nutrients. The median age of death for all CF patients is 30 years or less.  

According to the medical literature, restoration of as little as approximately 15% of wild-type CFTR function in 

CF patients should result in a therapeutic benefit.  

Cystic Fibrosis Genetics  

CF is an autosomal recessive disease. A normal, healthy gene has two alleles that encode for a given protein. 

In an autosomal recessive disease such as CF, a patient has a mutation in both alleles. Non-affected carriers 

have a mutation in only one of the alleles. CF patients have a defect in the gene encoding for the CFTR 

 
 
 
 
 
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protein, and they either have two copies of the same mutation, referred to as homozygotes, or one copy of 

two different mutations, referred to as compound heterozygotes. Although over 1,900 CF-causing gene 

mutations have been identified, approximately 85% of CF patients in the Western world are affected by the 

F508del mutation. Of which approximately 45% are homozygous for the F508del mutation and approximately 

40% are compound heterozygous for the F508del mutation. 

In the F508del mutation, the genetic defect is a deletion of three of the coding base pairs, or nucleotides, in 

the CFTR gene that results in the transcription of defective mRNA, which results in the production of CFTR 

protein that is misfolded and can neither migrate to its normal location on the surface of epithelial cells nor 

perform its normal function. 

Cystic Fibrosis Incidence and Diagnosis  

CF affects one out of 3,500 live births in the United States and one out of 2,500 live births in Western Europe. 

Many individuals are also non-affected carriers of a mutated CFTR gene. Carrier results across ethnic groups 

in the United States are well established, and reports from the American College of Obstetricians and 

Gynecologists indicate rates of one out of 25 in non-Hispanic Caucasians, one out of 58 in Hispanic 

Caucasians, one out of 61 in African Americans, and one out of 94 in Asian Americans. While the life 

expectancy of CF patients has improved over the last three decades, the median age of death is still only 30 

years or less in the U.S.  

Most CF patients in the United States, the European Union, Canada and Australia are now diagnosed at birth 

through newborn screening, and more than 75% of CF patients are diagnosed by the age of two. CF can be 

diagnosed by conducting a Nasal Potential Difference, or NPD, test that measures CFTR activity in the nose, or 

a pilocarpine iontophoretic sweat chloride test, which measures the amount of salt in a person’s sweat. A 

genetic test is also often used to confirm a CF diagnosis and/or identify the disease-causing mutations. In a 

genetic test, a blood sample or cells from the inside of the cheek are taken and sent to a laboratory that 

specializes in genetic testing.  

Approaches to the Treatment of Cystic Fibrosis  

Treatment overview  

There is no cure for CF, and to date, all but one of the therapies approved to treat CF patients have been 

designed to treat the symptoms rather than address the underlying cause. CF patients require lifelong 

treatment with multiple daily medications, frequent hospitalizations and ultimately lung transplant, which is 

life-extending but not curative. In the United States, the average CF patient incurs approximately $50,000 per 

year in expenses for outpatient medications and services and substantial additional costs for frequent 

hospitalizations. As the median age of death for CF patients is 30 years or less, this results in an average 

lifetime cost per CF patient in the U.S. of $1,350,000 in outpatient expenses alone. CF patients who can be 

treated with Vertex’s Kalydeco or Orkambi have additional annual costs of approximately $300,000. 

Standards of care are generally similar across Western European nations.  

Palliative treatments  

The current standard of care for CF patients includes palliative treatment to manage the symptoms of the 

disease. In CF patients, the thick mucus that builds up in the lungs and other vital organs such as the 

pancreas and gastrointestinal tract hampers mucus clearance and leads to airway obstruction and difficulty 

absorbing nutrients, leading to poor growth and development. Primary treatment options include inhaled 

therapies such as rhDNase, marketed as Pulmozyme, which thins the mucus in the lungs, as well as 

pancreatic enzyme replacement therapy, which improves absorption of nutrients. Due to the proliferation of 

bacteria on the mucus build-up, CF patients often develop chronic lung infections that require inhaled 

antibiotics treatments, such as TOBI or Cayston, to suppress the infections. CF patients also take a number of 

other prescribed and over-the-counter medications to alleviate the symptoms of CF and reduce 

 
 
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complications, including bronchodilators, inhaled corticosteroids, and ursodeoxycholic acid for biliary tree 

dysfunction. 

Potentiators for certain non-F508del mutations  

For a subset of patients who suffer from the G551D and other gating mutations of the CFTR gene, Vertex 

Pharmaceuticals has developed a so-called “potentiator” molecule marketed under the trade name Kalydeco 

(ivacaftor). This product was approved by the FDA in 2012 to treat patients with the G551D CFTR mutation 

and, in 2014, the label was expanded to include eight additional gating mutations. In 2015, the label was 

further expanded to include a total of ten gating mutations and children as young as two years old. Vertex 

has estimated that approximately 2,400 CF patients in the U.S., Europe and Australia have a G551D or a non-

G551D gating mutation. Gating mutations are characterized by the presence of CFTR at the cell surface that 

does not open and close the ion channels properly. Kalydeco is believed to keep the ion channels open for 

longer. For this population of CF patients, medication costs are approximately $300,000 per year for Kalydeco 

prescriptions. Kalydeco is an exciting development as it provides a proof of concept that it is possible to 

target the defective CFTR protein that causes CF and improve key symptoms of the disease.  

The F508del mutation affects approximately 85% of CF patients in the Western world. Unlike the “gating” 

mutations, F508del is a “processing” mutation, and as such, CFTR with the F508del mutation is not expressed 

at the cell surface and cannot be potentiated by small potentiating molecules like Kalydeco.  

Potentiator/corrector combination for F508del mutations  

For patients aged 6 years and above and homozygous for the F508del mutation, Vertex Pharmaceuticals has 

received regulatory approval for Orkambi. Orkambi is a fixed-dose combination of lumacaftor and ivacaftor 

(Kalydeco). Lumacaftor is a new molecular entity also referred to as a CFTR “corrector” that is purported to 

work by stabilizing and promoting the folding of the defective F508del CFTR and thereby increasing the 

likelihood that the CFTR channel will be found at the cell membrane. Kalydeco purportedly potentiates the 

activity of CFTR channel at the cell surface. We believe the clinical benefit of Orkambi for many homozygous 

F508del patients is not commensurate with the benefit demonstrated by Kalydeco in the G551D population, 

but is comparable to some of the symptom relief medications approved for use with CF. Approximately 

12,000 US patients could be treated with Orkambi at an estimated annual cost of approximately $260,000 in 

addition to the cost of standard of care. We believe these studies validate that F508del CFTR is a treatable 

target and indicate there is need for more efficacious therapies. 

Gene therapy  

Gene therapy is a process in which a functional gene is introduced into a cell to override the effects of a 

defective gene. The CFTR gene was first identified in 1989. Since that time, several academic consortia and 

drug-development companies have attempted to develop gene therapies targeting mutations in the CFTR 

gene. These companies aimed to permanently correct the CFTR gene at the DNA level by delivering full-length 

CFTR genes to lung epithelial cells to express wild-type CFTR protein. However, these programs encountered 

limitations faced by gene therapy in general as well as limitations specific to the CFTR gene. These barriers 

included safety concerns, challenges in delivery of the gene therapy constructs to target cells in the lungs, 

challenges of both delivery and incorporation into the genome given the size and complexity of the CFTR 

gene, and immunologic responses to the gene therapy vectors. The most advanced effort in gene therapy for 

CF is with an academic consortium in the U.K. In 2015, the Gene Therapy Consortium presented results of a 

136-patient trial using a CFTR gene delivered in a liposome envelope. While the trial showed no overall 

efficacy, specific subgroups did show a modest benefit in lung function compared to the placebo group. The 

Gene Therapy Consortium has announced that they will conduct a follow-up trial of gene therapy in the 

future but that a different vector will be needed for delivery of the gene. 

 
 
 
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Our RNA Approach  

QR-010 is a first-in-class RNA oligonucleotide designed to address the underlying cause of the disease by 

targeting the mRNA in CF patients that have the F508del mutation. QR-010 is designed to bind to the 

defective CFTR mRNA and restore CFTR function. We believe we are currently the only company pursuing this 

novel approach for CF patients.  

QR-010 for Treatment of CF  

We are developing QR-010 as an inhaled treatment for CF patients. QR-010 is a single stranded RNA 

oligonucleotide designed to restore CFTR function in CF patients with the F508del gene mutation. QR-010 is 

33 nucleotides long and is designed to bind to the CFTR mRNA sequences that are adjacent to the deleted 

F508del region of the mRNA.  

The figure represents an F508del mutated cell treated with QR-010, which would be expected to result in 

restoration of chloride efflux.  

Chloride ion flow: restoration 
through QR-010 treatment 

F508del + QR-010 

Clinical Development for QR-010  

We conducted two clinical trials of QR-010 in parallel. Study 

PQ-010-002 is a proof-of-concept trial evaluating topical 

administration of QR-010 and its effect on the nasal potential 

difference (NPD), a biomarker of CFTR function. This trial 

opened for enrollment in September 2015 and was completed 

in September 2016. Study PQ-010-001 is a Phase 1b safety and 

tolerability trial. This trial opened for enrollment in June 2015 

and is currently enrolling.  

PQ-010-002 Proof-of Concept NPD study  

The NPD assay is a standard test for detection and 

quantification of CFTR function in the airways in CF patients. 

The NPD test is a well-accepted diagnostic tool and has been 

used in multiple therapeutic intervention trials to demonstrate 

the restoration of CFTR mediated ion transport in pre-clinical 

animal models and in CF patients. Our trial was designed to 

investigate the ability of QR-010 to restore CFTR function in 

patients. Restoration of CFTR function has been observed in 

pre-clinical NPD studies using mouse models. The primary outcome measure was to determine the effect of 

topical administration of QR-010 to the nasal mucosa on the restoration of CFTR-mediated chloride transport 

as measured by NPD in CF patients with the F508del CFTR mutation. Secondary endpoints included maximal 

basal potential difference reflecting sodium channel activity. Nasal administration is not the intended route of 

administration for QR-010. However, the nasal epithelium is the most accessible site for measuring CFTR 

function in humans and provides a human model of epithelial cell uptake and restoration of CFTR function. 

All subjects were adults over 18 years old with CF either homozygous for the F508del mutation or compound 

heterozygotes with one copy of the F508del mutation and one copy of another disease causing mutation. The 

trial was conducted in five sites in the U.S., France and Belgium. QR-010 was administered intranasal 5 mg in 

each nostril 3 times weekly for 4 weeks (12 doses). The NPD measurements were done at baseline, after 6 

doses (Day 15), after 11 doses (Day 26) and 21 days after the last dose (Day 47). 

Topline results were reported at the North American CF Conference (Sermet-Gaudulus, Pediatr Pulmonol 

2016 Suppl 45:485) in October 2016. In the per-protocol population of subjects homozygous for the F508del 

mutation meeting the pre-specified inclusion criteria (n=7), the average change from baseline in NPD at day 

26 was statistically significant, -4.1 mV (p=0.0389). This finding was supported by a change in sodium channel 

 
 
 
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activity (specifically, a measure called max basal potential difference, or PD) and other sensitivity analyses of 

the NPD measurements, all pointing to strong evidence of restoration of CFTR activity. In subjects compound 

heterozygous for the F508del mutation, the average change from baseline in NPD was not significantly 

different at day 26. A responder analysis of individual subjects assessing the impact of the second mutation is 

currently ongoing. QR-010 administered via the intranasal route was observed to be well tolerated.  

QR-010 improves CFTR function in F508del homozygous  
CF patients as measured by NPD 

N= 7 (per protocol population). Parameter = within subject change from baseline in Cl-free+iso. Average 
both nostrils. Baseline = average of 2 most recent pre-dose assessments. P = one-sided 5% paired t-test. 

We observed from the results of this trial that QR-010 improved CFTR function in homozygous F508del 

patients as evidenced by both the increase in CFTR activity measured in the CFTR-mediated total chloride 

response and the decrease in sodium channel activity as measured by the max basal potential difference. 

The magnitude of the change observed in this trial is similar to that published for other commercially 

available treatment in CF patients with the G551D mutation and superior to data published for lumacaftor in 

patients with the F508del mutation. 

Putting QR-010 clinical NPD results in perspective 

 
 
 
 
 
 
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PQ-010-001 Phase 1b Safety and Tolerability Trial 

This clinical trial with QR-010 is a Phase 1b, randomized, double-blind, placebo-controlled, 28-day dose-

escalation trial to evaluate the safety, tolerability and pharmacokinetics of QR-010. The trial includes CF 

patients that are homozygous for F508del and age 18 years and above. The trial is being conducted at 27 

sites in North America and select EU countries and will enroll approximately 64 patients. The trial consists of 

4 cohorts of ascending single dosed and 4 cohorts of ascending multiples doses (12 doses over 4 weeks). In 

each cohort, randomization is 3:1, meaning that 6 patients will receive QR-010 and 2 patients will receive 

placebo.  

QR-010 is given as a nebulized solution to the lower airways after chest physiotherapy, which is a standard 

procedure used with other currently administered inhaled medications. This method of drug administration 

is common in CF patients. The primary outcome measures are to characterize safety, tolerability and 

pharmacokinetics of QR-010 in CF patients. Pharmacokinetics will be assessed in serum, urine and sputum. 

These measurements will allow us to establish the safety for QR-010 as well as give indications of uptake into 

the lung and systemic circulation in order to provide PK/PD information to design our future trials. We are 

also assessing exploratory efficacy outcome measures, including lung functionality, chloride levels in sweat, 

weight gain and other quality-of-life measures specific to CF. In October 2016, we reported that the single 

dose portion of the trial consisting of 4 cohorts has been completed. No dose-limiting toxicity was observed 

up to the highest dose tested. The dose escalating multiple-dose trial (12 doses administered over 4 weeks) is 

currently enrolling cohort 7 and topline results are expected to be available in mid-2017. Further update on 

enrollment will be provided at the European Cystic Fibrosis Conference (ECFS). 

PQ-010-003 Phase 2 Trial 

PQ-001-003 is currently planned as a Phase 2 multicenter, randomized, double-blind, placebo-controlled 12-

week trial to evaluate the safety, efficacy, and pharmacokinetics of QR-010 in cystic fibrosis subjects with the 

F508del mutation. The trial will be conducted at clinical centers in North America, EU and possibly other 

countries. We anticipate to begin recruitment for this trial in 2018.  

Inhaled administration of QR-010 

To achieve broad distribution to CF-affected organs, we deliver QR-010 through inhalation by means of a 

small handheld nebulizer, a method of drug delivery used to administer medication in the form of a mist 

inhaled into the lungs. On October 8, 2014 we entered into an agreement with PARI Pharma GmbH, pursuant 

to which the Company is granted an exclusive license to the use of PARI’s eFlow technology for the 

administration of oligonucleotide-based drugs in the F508del mutation in cystic fibrosis. The nebulizer device 

rapidly and efficiently processes a therapeutic agent through the microscopic holes of a mesh and creates a 

mist to provide rapid and consistent delivery to the lungs. Commercially-available nebulizers are currently 

used for other CF therapies and in other clinical studies involving inhaled oligonucleotides. 

 
 
 
 
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Bio-distribution of QR-010 to organs in wild-type  
and CF-like lung phenotype 

Pre-clinical evidence for QR-010  

As shown in the figure, after 

orotracheal delivery of QR-010 to the 

lungs of wild-type mice and mice 

specifically engineered to have a CF-

like lung phenotype, called the 

betaENaC overexpressing mouse, we 

observed significant exposure of QR-

010 to the lungs as well as to other CF-

affected organs with no significant 

difference between wild-type and 

betaENaC overexpressing mice. We 

believe this beneficial bio-distribution 

pattern may potentially allow us to treat not just the lung but also other organs affected by CF and shows that 

the thick mucus layer that is present in the lungs of CF patients is unlikely to be a barrier for uptake of QR-

010. We believe this method will allow maximum exposure of QR-010 to the primary target organ, the lung, as 

well as significant exposure to other affected organs through systemic absorption into the blood. 

We have conducted extensive in vitro and in vivo pre-clinical studies that support the development and 

therapeutic potential of QR-010. QR-010 has been shown to increase the function of CFTR as demonstrated 

by enhancing chloride efflux in vitro and in vivo models that carry the same mutation as F508del patients. In 

vitro QR-010 demonstrated improved chloride ion efflux in a fluorescent chloride ion indicator, or MQAE, 

assay and in a well-accepted model, the Ussing Chamber assay using human bronchial epithelial cells with 

the F508del mutation. Most notably, and distinct from other molecules in development for CFTR mutation 

specific molecules, in two independent in vivo activity assays in F508del-CFTR mice that are similar to human 

diagnostic tests, QR-010 restored CFTR function up to wild-type levels. The first was a study of Nasal Potential 

Difference, or NPD, in F508del-CFTR mice in which QR-010 restored NPD in response to specific stimuli to 

normal levels. The second was a saliva secretion assay, a mouse equivalent of the sweat chloride test, in 

which QR-010 restored saliva secretion to normal levels in female mice.  

Research Grants 

In August 2014, we entered into an agreement with Cystic Fibrosis Foundation Therapeutics, Inc., or CFFT, a 

subsidiary of the Cystic Fibrosis Foundation, pursuant to which CFFT agreed to provide us with up to $ 3 

million to support the clinical development of QR-010. In 2015, the Company and its academic partners 

received a grant from the European Union under the Horizon 2020 research and innovation programme 

under grant agreement No. 633545. The maximum amount of € 6.0 million was granted to support the 

clinical development of QR-010. In 2016, ProQR also received additional tranches totaling €0.4 million under 

the Innovation credit program or “Innovatiekrediet” by the Dutch government, through its agency RVO 

(previously: “AgentschapNL”) of the Ministry of Economic Affairs, for the cystic fibrosis development program. 

 
 
 
 
 
 
 
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Leber’s Congenital Amaurosis  

LCA Background 

LCA is the most common genetic cause of blindness in childhood. We believe that the p.Cys998X mutation 

(also known as c.2991+1655A>G) in the CEP290 (Centrosomal protein of 290 kDa) gene is the most prevalent 

mutation which generally accounts for the most severe disease phenotype (LCA 10). Patients affected by this 

mutation typically lose sight in the first years of life. In LCA 10 patients, this mutation leads to significant 

decrease in CEP290 protein within the photoreceptor cells in the retina. Clinical features of CEP290-mediated 

LCA include loss of vision, involuntary eye movement or nystagmus, abnormalities of pupil reactions and no 

detectable photoreceptor electrical signals on electroretinography (ERG).  

LCA Genetics 

The p.Cys998X mutation is a single nucleotide substitution in the CEP290 gene that creates a new splice site, 

also called a cryptic splice site, between exon 26 and 27. During the splicing of the pre-mRNA this causes a 

part of the intron, or pseudoexon, to be included in the mRNA. The pseudoexon contains a premature stop 

Representation of the p.Cys998X  
mutation causing LCA 10 

Wild-type 

Leber’s congenital amaurosis type 10 

codon thus the mRNA is not translated into the 

full length CEP290 protein. The CEP290 protein is 

involved in the formation and stability of the 

connecting cilium in photoreceptor cells, which 

facilitates the transport of proteins from the inner 

segment to the outer segment of the cell. When 

CEP290 is absent, there is a disturbance in normal 

protein transport to the outer segments which 

provokes the shortening of the outer segment   

and its inability to perform its light transducing 

function.  

LCA Prevalence and Diagnosis  

LCA is caused by a genetic defect in 20 or more 

associated genes. The most common mutation is 

the p.Cys998X in the CEP290 gene causing LCA 10. 

Although diagnosis rates vary, our estimations 

indicate this mutation to occur in approximately 

2,000 patients in the Western world. 

Patients are initially diagnosed through the 

presence of clinical symptoms. Nystagmus, rapid 

involuntary movements of the eyes, tends to be the first symptom visible as well as oculo-digital signs 

comprising eye poking, pressing, and rubbing, in the most severe cases; vision impairment or blindness 

becomes obvious as age increases. After ophthalmological examination, LCA is diagnosed. A genetic 

screening including all known mutations causing LCA is performed to confirm the diagnosis and determine 

the type of LCA in order to give the patient the most accurate prognosis possible (approximately 30% of all 

patients carry a mutation that has not been described to date). 

Approaches for the Treatment of LCA 10 

There are currently no disease modifying treatments approved or potential treatments in clinical trials for 

patients with p.Cys998X associated LCA 10, a form of LCA. There are other approaches in pre-clinical 

development for the p.Cys998X mutation that target the disease at the DNA level. The eye is highly suitable 

for oligonucleotide therapies as it is a contained organ with physical cellular barriers (i.e. efficient blood-

retina barrier and lack of efferent lymphatics) which strongly limits the free entry and exit of cells and larger 

molecules in and out of the eye therefore limiting the systemic exposure of locally administered therapies.  

 
 
 
 
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QR-110 for the treatment of LCA 10 

Our lead product candidate in the LCA 10 space, QR-110, is a first-in-class single stranded RNA 

oligonucleotide of 17 nucleotides long. It is designed to treat the disease by binding to the pre-mRNA and 

thereby silencing the cryptic splice site caused by the p.Cys998X mutation. The splicing machinery can thus 

splice the pre-mRNA correctly resulting in normal mRNA and we expect the production of full-length 

functional wild-type CEP290 protein. The intended route of delivery is through intravitreal injection. 

QR-110 for LCA 10, splice correction for  
p.Cys998X CEP290 mRNA 

QR-110 binds to pre-mRNA and silences the cryptic splice site leading to 
production of normal mRNA 

Clinical Development for QR-110 

We believe the activity seen in our 

pre-clinical models of LCA 10 

provides strong support for the 

clinical development and 

therapeutic potential of QR-110. 

Currently, we are finalizing pre-

clinical good laboratory practice, or 

GLP, safety studies and other work 

to start our first clinical trial in LCA 

10 patients. 

We expect to commence clinical 

development of QR-110 in the first 

half of 2017 with the initiation of a 

Phase 1b/2 clinical trial. This trial is an open label trial evaluating multiple doses of QR-110 at different dose 

levels. Eligible subjects will be LCA 10 patients that are homozygous or compound heterozygous for the 

p.Cys998X mutation. The primary objective will be to evaluate the safety and tolerability of QR-110 

administered via intravitreal injection in subjects with LCA 10 due to the p.Cys998X mutation. Secondary 

objectives will include the assessment of pharmacokinetics and efficacy as assessed by specialized 

ophthalmic tests. 

Pre-clinical evidence for QR-110 

We have conducted in vitro and in vivo pre-clinical studies that we believe support the clinical development to 

explore the therapeutic potential of QR-110.  

QR-110 assessment in patient fibroblasts 

Since QR-110 targets the splicing process, the most direct measurable outcome of activity is the profiling and 

quantification of CEP290 transcripts (wild-type and mutant) and protein before and after treatment. In pre-

clinical studies to date, QR-110 has demonstrated restoration of CEP290 wild-type (correctly spliced) mRNA 

and protein in cultured fibroblast cells of LCA 10 patients homozygous and compound heterozygous for the 

p.Cys998X mutation.  

 
 
 
 
 
 
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Homozygous cells (p.Cys998X/p.Cys998X; LFB-3) (figure A) 

Compound heterozygous cells (p.Cys998X/p.Lys1575X; LFB-6) (figure B) 

Effect of QR-110 at the mRNA and protein level in fibroblast cells from LCA 10 patients that are A) homozygous and B) 
compound heterozygous for the p.Cys998X mutation. Normalized wild-type and mutant CEP290 mRNA expression 
(copies/ng) after transfection of LCA 10 fibroblasts with QR-110, analyzed with one-step ddPCR. For protein data (Western 
Blot), expression is shown relative to control cells without the mutation. Error bars show mean with SEM. *p<0.05, 
**p<0.01, ***p<0.001, vs. mock treated cells, Student’s t-test. 

The figure above summarizes the observations from our pre-clinical data that treatment with QR-110 may be 

able to increase the expression of wild-type CEP290 mRNA and protein in fibroblast cells from LCA 10 patient 

that are homozygous for the p.Cys998X mutation. Furthermore, we observed that treatment with QR-110 

resulted in a decrease in levels of mutant mRNA (figure A, left and center). The mRNA and protein profile 

restoration trend is also observed in LCA 10 fibroblasts that are compound heterozygous for the p.Cys998X 

mutation (figure B, left and center).  

Changes in the mRNA profile are supported by a wild-type CEP290 protein increase illustrated by Western 

blot. Results demonstrate that in LCA 10 fibroblasts that are homozygous for the p.Cys998X mutation, in vitro 

treatment with QR-110 restored CEP290 protein levels to that of control cells (figure A, right panel). In LCA 10 

fibroblast that are compound heterozygous for the p.Cys998X mutation, QR-110 treatment in vitro restored 

CEP290 protein levels to ~50% of control cells (figure B, right panel). This is expected since in these compound 

heterozygous cells only one mutated allele carries the p.Cys998X mutation and therefore only one allele can 

be targeted by QR-110 treatment. It is worth it to point out that patients that are heterozygous for the 

p.Cys998X mutation, with one normal allele and one allele carrying the p.Cys998X mutation, are 

asymptomatic. This indicates that correction of one diseased allele could be enough to prevent or stop 

progression of the disease. 

QR-110 activity in optic cup model 

Optic cups are a retinal organoid model derived from fibroblasts of a LCA 10 patient harvested through skin 

biopsies. The cells are reprogrammed into induced pluripotent stem cells, or iPSC, and later differentiated 

into retinal pigmented epithelium cells and neural retinal cells, also known as three-dimensional optic cups. 

 
 
 
 
 
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Optic cups constitute a convenient and clinically relevant model system to thoroughly study the mechanisms 

of inherited retinal degeneration since, unlike the classic cell models, these 3D organoids simulate the 

disease phenotype and provide an appropriate cellular model with the genetic mutations in genomic context.  

The clinical and molecular relevance of the optic cup model, coupled with the absence of an animal model, 

makes the optic cup the best model in which to simulate the mechanisms of CEP290 LCA and effectively test 

the potential of QR-110.  

LCA 10 patient derived optic cups were exposed to QR-110. First, we observed from the results that QR-110 is 

able to enter the cells without use of any transfection agents. Second, QR-110 elicited a dose-dependent 

restoration of CEP290 wild-type mRNA expression. And third, increased CEP290 mRNA expression was also 

associated with a commensurate decrease in mutant CEP290 mRNA.  

Generation of LCA 10 patient iPSC-derived optic cups 

QR-110 increases wild-type CEP290 mRNA levels in a  
dose-dependent manner in LCA 10 optic cups 

LCA 10 homozygous optic cups 

LCA 10 p.Cys998X homozygous patient fibroblasts were reprogrammed into iPSC which were 
differentiated into optic cups for 96 days and treated with different amounts of QR-110 for 
another 28 days (Parfitt et al. 2016) and analyzed using end-point PCR. 

 
 
 
 
 
 
 
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Retinal Distribution of QR-110 

Labelled QR-110 (green) administered via intravitreal injection into wild-type mice eyes. We demonstrated 

that QR-110 enters the target cells of the retina, including the photoreceptor cells. QR-110 was detected 60 

days (the maximum time point tested) following a single injection. 

QR-110 reaches target cells after intravitreal injection 

Retinal distribution of 6FAM-labelled QR-110 following single intravitreal injection of  
100 µg in wild-type mice. 

Dystrophic Epidermolysis Bullosa 

DEB Background 

Epidermolysis bullosa (EB) is a rare genetic disorder, primarily manifesting as a debilitating disease of the skin 

and mucosal membranes. It is characterized by mechanical fragility of epithelial tissues, blister formation, 

scarring and, in some subtypes, involvement of multiple other organs. EB is classified into four main 

subtypes, namely EB simplex (EBS), junctional EB (JEB), dystrophic epidermolysis bullosa (DEB), and Kindler 

Syndrome (KS). The four main EB subtypes are distinguished by the level of the skin at which blisters develop. 

In DEB the outer layer of the skin, the epidermis, separates from the inner layer, the dermis. This separation 

renders the skin fragile and causes severe blistering and scarring. All mucosal membranes are affected in 

DEB, therefore blistering is not limited to the skin, but is also present in the mouth, esophagus and 

downstream intestines. 

DEB is usually a chronic, seriously debilitating disease with a shortened life expectancy due to malnutrition, 

infections, and malignancies. 

DEB Genetics 

The disease is caused by mutations in the COL7A1 gene. This gene is responsible for the production of a 

protein called collagen type VII (also referred to as C7), which is a major component of the anchoring fibril 

located below the basement membrane in the upper dermis that normally links the epidermis and the 

dermis together. DEB causing mutations occur more often in certain parts of the gene. One of those parts is 

exon 73. 

DEB Prevalence and Diagnosis 

DEB is a genetic disease that in some cases is inherited as an autosomal dominant (DDEB) and in others as an 

autosomal recessive trait (RDEB). The prevalence of DEB could differ across countries due to founder effects 

and differences in ethnic composition. While spatial variations, compounded with the scarcity of available 

data, make accurate calculations difficult, the estimated number of DEB patients in the western world is 

approximately 6,000 of which approximately 2,000 have mutations in exon 73. 

 
 
 
 
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Diagnostic testing for DEB is based the identification of the level of skin cleavage via immunofluorescence 

antigen mapping with C7 specific antibodies and/or determination of anchoring fibrils using transmission 

electron microscopy on, preferably, newly formed blisters. 

Approaches for the Treatment of DEB 

Currently no disease modifying treatment is available for DEB. Palliative treatment is the only treatment 

available for DEB patients and constitutes a time-consuming daily activity. Palliative treatment primarily 

consists of 1) treatment of (new) blisters by puncturing and draining blisters to prevent further spread from 

fluid pressure, 2) wound management to prevent infections, 3) prevention of skin trauma to avoid new blister 

formation, and 4) pain and itch relief.  

QR-313 for the treatment of DEB 

QR-313 is designed to specifically target mutations in exon 73 of the COL7A1 gene. QR-313 binds to a specific 

sequence in the COL7A1 pre-mRNA, thereby excluding exon 73 from the mature mRNA. This leads to a 

shortened version of the C7 protein that is functional in the formation of anchoring fibrils.  

Because of the exon skipping approach, QR-313 is not specific to a single mutation but instead targets any 

mutation contained in exon 73. 

Functional C7 protein: Restoration through QR-313 Treatment 

Schematic shows pathway for generation of C7 protein in the healthy and disease situations (left and center diagrams, 
respectively). Hybridization of QR-313 to a specific sequence in COL7A1 pre-mRNA results in the exclusion of exon 73 from 
the mRNA, which leads to the production of a truncated but still functional C7 protein (right diagram).  

Pre-clinical evidence for QR-313 

Clinical development of QR-313 focusses on topical delivery in the wounded skin of patients, with the aim of 

improved wound healing and reduced skin fragility. Therefore we aim to formulate QR-313 into a hydrogel for 

wound application that can be incorporated in the standard of care of patients. 

Activity of QR-313 after topical application on human skin equivalents 

In order to investigate topical delivery and exon skip potential of QR-313 we used Human Skin Equivalents, or 

HSEs. HSEs are an often used model to mimic the human skin. They are composed of both a dermal layer 

containing fibroblasts and an epidermal layer containing keratinocytes. The keratinocytes are fully 

differentiated to form all the different layers in the epidermis, including the stratum corneum. The culturing 

of HSEs is done at the air-liquid interface and therefore mimics the human situation. Moreover, by removing 

the epidermis from a portion of the skin equivalent, the blister phenotype of DEB can be modeled.  

 
 
 
 
 
 
 
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Cy5-labeled QR-313 in a hydrogel formulation was used in HSEs where a DEB blister was mimicked by 

removal of part of the epidermis. The figure below shows that diffusion of QR-313 into the dermis was 

observed both in the middle of the blister (blister bed) and at the edge of the blister (blister edge). QR-313 is 

not able to penetrate intact HSEs, and ex vivo skin (not shown).  

Delivery of Cy5-labeled QR-313 formulated in a hydrogel 

After 24 hour treatment (indicated at left of picture rows), HSE pieces 
(indicated at top of picture columns) were analyzed. QR-313 is 
depicted in red, nuclei are depicted in blue. White dotted line 
represents border between epidermis and dermis.  

QR-313 induced Skipping of Exon 73 in C7 mRNA 

Splicing products of COL7A1 mRNA either untreated or following 
treatment with Cy5-labeled QR-313 formulated in a hydrogel. RNA was 
isolated from treated HSEs (indicated at top of columns) and RT-PCR 
analysis was performed. The different COL7A1 mRNA products were 
analyzed for length. The 350 bp fragment represents the wild-type, full 
length amplicon including exon 73 mRNA while the 150 bp nucleotide 
fragment represents the modified Δ73 mRNA product. 

To examine the ability of QR-313 to 

induce exon skipping in dermal 

fibroblasts, we separated the 

epidermis from the dermis from the 

24-hours incubated HSEs. RNA 

isolation was performed and analyzed 

for exclusion of exon 73 using RT-PCR. 

Exon 73 exclusion from the COL7A1 

mRNA was observed in dermal cells 

treated with QR-313 formulated in a 

hydrogel. This shows that upon local 

application QR-313 is active in this 

model that mimics blistered EB skin. 

Other Research and Development  

Our internal discovery engine that we 

call the innovation unit, is a dedicated 

group in our company that focuses on 

the discovery and early development 

of RNA therapeutics in genetic 

indications with a high unmet medical 

need. Leveraging our experience with 

RNA therapeutics, we are screening for 

therapeutic molecules that can be used 

to treat severe genetic disorders 

beyond CF, LCA 10 and DEB. We have 

built a diverse toolbox of RNA 

technologies that we believe can 

address underlying genetic defects in a 

novel way. We have grouped the 

different programs in franchises by 

therapeutic area so that we can 

leverage our expertise in the different 

fields and create synergies between 

programs. We have identified five therapeutic areas that show high potential for RNA based oligonucleotides: 

respiratory, ophthalmology, dermatology, CNS and neuromuscular disorders. We go through a thorough 

selection process prior to advancing programs into development, and consider criteria including: high unmet 

medical need, a pre-clinical proof-of-concept that shows strong promise for translation to the clinic, well 

understood relationship between the genetic defect and disease manifestations, feasibility of delivery to 

target organ(s), and strong IP position and initial freedom to operate.   

Respiratory 

Besides our program for CF caused by the F508del mutation we are working on other CFTR mutations that 

can potentially be treated using our RNA technologies. We could potentially target an additional 5% of the CF 

population with these programs.  

 
 
 
 
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Ophthalmology 

Our ophthalmology group was founded on the basis that the eye is a well validated target for 

oligonucleotides. Given the long half-life of these molecules and the lower risk of systemic exposure, we 

believe oligonucleotide based therapies have the potential to be an important class of drugs for ophthalmic 

indications. Besides our LCA 10 program that we intend to take into clinical development in 2017, we have 

several discovery stage programs, including two programs targeting Usher syndrome and a program 

targeting Fuchs endothelial corneal dystrophy (FECD). 

Usher syndrome is a genetic orphan disease that is the leading cause of combined deafness and blindness. 

Usher syndrome type II is most commonly caused by mutations in the USH2A gene. Patients with this 

syndrome generally progress to a stage in which they have severely limited central vision and moderate to 

severe deafness. The moderate to severe deafness that patients experience with this subtype of the disease 

is manageable with cochlear implants. However, there are currently no available treatment options for the 

vision loss associated with this disease. The disease is caused by a genetic defect that results in the lack of a 

functional USH2A protein. Similar to CEP290, this protein is responsible for the maintenance of the 

connecting cilium in photoreceptor cells and lack of a fully functional USH2A protein results in reduced 

protein trafficking to the photoreceptor outer segments with a consequent impact on photo-transduction. 

With our two programs, called QRX-411 & QRX-421, we aim to target genetic alterations in the USH2A gene 

that lead to this vision loss. QRX-411 targets the frequent deep-intronic c.7595-2144A>G mutation that causes 

the inclusion of a pseudoexon in the mRNA disrupting the function of the protein. QRX-411 is designed to 

target the pre-mRNA and restore a wild-type sequence in the mRNA leading to wild-type mRNA and 

functional USH2A protein. QRX-421 targets mutations in exon 13 of the USH2A gene by skipping exon 13 

from the mRNA restoring the reading frame and producing a truncated but functional protein. Both QRX-411 

and QRX-421 are single-stranded RNA oligonucleotides intended to be administered by intravitreal injections. 

FECD is a common inherited condition characterized by the dysfunction and degeneration of the corneal 

endothelium. The disease segregates into early-onset and age-related FECD that are caused by different 

mutations. Early signs of FECD are the presence of corneal guttae and a large proportion of patients over 40 

years old have evident corneal guttae. A portion of these patients develop advanced disease with corneal 

edema and corneal clouding. These symptoms can worsen leading to complete vision loss and the 

requirement for surgical intervention and a corneal transplant. There are currently no other treatment 

options for any form of FECD patients with vision loss, apart from corneal transplantation. However, 

transplantation has several limitations, including the availability of donors, risk of rejection, the inherent risk 

of an invasive procedure and is only available to patients with advanced FECD. The majority of age-related 

FECD is caused by a repeat expansion mutation in the TCF4 gene. Such expansions result in toxic RNA species 

which aggregate as nuclear foci and sequester important splicing proteins rendering the cell devoid of the 

splicing proteins for other important genes. The impact of acquired splicing defects in these other genes are 

thought to result in corneal endothelial dysfunction and Fuchs. Our program, called QRX-504, is a single-

stranded RNA oligonucleotide that aims to prevent the buildup of RNA-protein foci that cause the corneal 

dystrophy in patients with expansion repeat mutation in the TCF4 gene. 

Dermatology 

Our product candidate, QR-313 targeting DEB caused by mutations in exon 73 has been moved into clinical 

development. If the exon skipping approach is proven to be of benefit for DEB patients, there may be other 

COL7A1 mutations that can be targeted with an exon skipping approach similar to QR-313. 

CNS 

In our CNS group we are working on product candidates for several disease targets, including a wide range of 

neuronal and cerebrovascular related amyloid beta disorders and Huntington’s disease, or HD.  

 
 
 
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Amyloid beta, or Aβ is a highly toxic and aggregate-prone family of peptides that are appears crucially 

involved in Alzheimer’s disease, or AD, cerebral amyloid angiopathy, or CAA, and its familial form hereditary 

cerebral hemorrhage with amyloidosis of the Dutch type (HCHWA-D). We are developing QRX-203, a RNA 

modulation therapy for Aβ-induced amyloidosis. Using antisense oligonucleotide mediated exon skipping, we 

believe QRX-203 may prevent the translation of the amyloid region into its precursor protein APP. We believe 

this approach renders the release and aggregation of Aβ impossible and may ultimately prevent the onset 

and/or or slow the progression of disease. 

HD is an inherited progressive neurodegenerative disease, and one of the most common genetic disorders, 

with symptoms including involuntary movements, incoordination, impaired speech, cognitive decline, and 

depression. Individuals with HD have shortened life expectancy, and there is currently no disease-modifying 

treatment available. The disease is caused by an expanded repeat of CAG nucleotides in the Huntingtin, or 

HTT, gene, resulting in a mutated Huntingtin protein. When the mutated protein is present in the cells, small 

polyglutamine-containing protein fragments are formed. These fragments stick to each other, and 

accumulate in nerve cells, interfering with normal cellular functions, eventually leading to cell death. QRX-704 

is an oligonucleotide based approach that is aimed to modify the HTT mRNA to prevent the formation of the 

toxic fragments, while the Huntingtin protein remains functional. 

Neuromuscular 

Friedreich's ataxia, or FA, is the most common inherited ataxia that causes progressive damage to the 

nervous system. The disease is caused by GAA repeat expansion mutations in the gene that codes for the 

Frataxin protein. The expanded repeat mutations cause silencing of the gene leading to decreased levels of 

the Frataxin protein. Symptoms range from muscle weakness and speech problems to heart disease. With 

only palliative treatments available, most patients are wheelchair bound within 10–15 years after diagnosis 

and do not live beyond early adulthood. Frataxin is an essential mitochondrial protein involved in the 

regulation of energy production in cells and enzymes that contain an iron-sulfur cluster. We have identified a 

potential treatment, called QRX-604, with the aim to increase Frataxin levels. 

Human resources 

At ProQR we have set ourselves the immense task of developing drugs that will potentially transform the lives 

of patients suffering from severe genetic diseases like cystic fibrosis, Leber’s congenital amaurosis, and 

epidermolysis bullosa. To make this happen we demand the utmost of ourselves. We actively create a caring 

atmosphere filled with fun and joy, in which we love to work and maintain productive and happy lives. At 

ProQR we foster empowerment, self development, creativity and a sense of community.  

We are a supportive, ingenious and persistent team that does things different. We're passionate and driven 

to change the lives of patients and their loved ones. 

Corporate social responsibility 

It is required by regulatory authorities to demonstrate the safety and efficacy of a new drug in both animals 

and humans, before the authorities can approve the new product and will provide Marketing Authorization.  

ProQR attaches great importance to the welfare of animals and humans participating in our pre-clinical and 

clinical studies for reasons of ethics, quality, reliability and applicability of scientific studies. For conducting 

high quality (scientific) animal research, animal welfare is a prerequisite. By actively pursuing the 3R principles 

(Reduce, Refine and Replace), we are committed to minimalizing the number of animals needed, minimizing 

discomfort and pain of animals used and to using alternatives to animal research whenever possible in 

research and in the obligatory animal studies. All our current studies are approved by the (institutional or 

national) animal care and use committees. 

 
 
 
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Our aim is to monitor continually that animal experiments will be performed only if there are no viable or 

legal alternatives. Additionally, case by case, it will be evaluated if advances can be made in study designs 

(such as by ex-vivo studies or by conduction of small pilot studies first), or by using new technologies to 

achieve adequate statistical power without increasing the number of animals, combining studies, and 

improving use of TK data to optimize dose selection. 

External collaborators contracted for the execution of our in-vivo pre-clinical studies (contract research 

organizations, CROs) are selected based on their expertise, quality and accreditations for laboratory animal 

care and welfare. CRO facilities are audited in person prior contracting. The housing, husbandry and animal 

welfare must comply with the highest international standards. Personnel responsible for housing, husbandry 

and care of the animals must have received adequate and relevant documented education.  

We strive for welfare improvements to be implemented in CRO policies. An important achievement in 2014 

was that on our request our preferred CRO has replaced the housing which was compliant with their national 

legislations and installed new group housings with significantly more living space that to a larger extent take 

in consideration the physiological and behavioral needs of the laboratory animals concerned. This will also 

contribute to higher welfare standards in the studies for other (future) clients.  

In 2015 we became part of an interdisciplinary consortium with Utrecht University (Faculty of Veterinary 

medicine and Ethics Institute), Radboud University (Medical Center, SYRCLE) and another private company, 

partly financed by The Netherlands Organization for Scientific Research, Responsible Innovation grant. The 

project proposes a more integrated approach towards innovation in the field of animal testing and focuses 

on translational strategies. ProQR is involved in the part of the project that aims to deliver step stones for 

practical guidelines to build robust translational strategies, to design innovative experiments (including 

animal models) for cystic fibrosis and develop a translational strategy for CF as a showcase.   

Main financial developments 

Financial position 

In 2016, we successfully expanded our operating activities. Operating costs went up significantly while our 

liquidity and solvency went down. ProQR’s cash and cash equivalents at December 31, 2016 amounted to  

€ 59,200,000 compared to € 94,865,000 at December 31, 2015. During the year 2015, operating cash used 

amounted to € 34,221,000, compared to € 24,232,000 in 2015. Shareholders’ equity decreased to  

€ 53,136,000. 

As at December 31, 2016, we had non-current liabilities of € 5,697,000, which fully consisted of borrowings 

from a government body. 

Income statement 

We have generated losses since our formation in February 2012. For the years ended December 31, 2015 and 

2016, we incurred net losses of € 20,832,000 and € 39,103,000, respectively. As at December 31, 2016, we had 

an accumulated deficit of € 75,733,000. We expect to continue incurring losses for the foreseeable future as 

we continue our pre-clinical studies of our product candidates, continue clinical development of our product 

candidate QR-010, advance QR-110 and QR-313 into clinical development, increase investments in our other 

research programs, apply for marketing approval of our product candidates and, if approved, build a sales 

and marketing infrastructure for the commercialization of our product candidates. To date, we have not 

generated any revenues from royalties or product sales. Based on our current plans, we do not expect to 

generate royalty or product revenues for the foreseeable future. 

Other income is incidental by nature. In August 2014, we entered into an agreement with Cystic Fibrosis 

Foundation Therapeutics, Inc., or CFFT, a subsidiary of the Cystic Fibrosis Foundation, pursuant to which CFFT 

 
 
 
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ANNUAL REPORT 2016 

agreed to provide us with up to $ 3 million to support the clinical development of QR-010. In 2015, the QR-010 

project has received funding of € 6 million from the European Union’s Horizon 2020 research and innovation 

programme under grant agreement No 633545. Other income amounted to € 1,828,000 in 2016, compared 

to € 3,235,000 in 2015. We expect to continue generating other income from CFFT and Horizon 2020 in 2017. 

Research and development costs increased to € 31,923,000 from € 23,401,000 in 2015. These research and 

development costs comprise allocated employee costs including share-based payments, the costs of 

materials and laboratory consumables, outsourced activities, license and intellectual property costs and other 

allocated costs. These costs were primarily related to our product candidates, QR-010, QR-110 and QR-313, 

and our innovation unit. Our research and development expense is highly dependent on the development 

phases of our product candidates and is expected to continue to increase at a moderate level, although it 

fluctuates significantly from period to period. 

The increase in expenses was primarily due to the advancement of our pipeline, which included clinical 

development of QR-010, preclinical development of QR-110 and QR-313 and progress of our innovation 

programs in ophthalmology, neuromuscular and central nervous system (CNS) diseases. The variances in 

research and development costs between the years ended December 31, 2016 and 2015 are mainly due to:  

 

 

 

 

 

 

 

costs we incurred on clinical trials for QR-010; 

increased staff costs as a result of increased staff working on (pre-)clinical development of our product 

candidates and the growth of our innovation unit. The number of full-time equivalent employees 

working on research and development increased from 72 at December 31, 2015 to 100 at December 31, 

2016;  

increased costs for externally conducted studies, including various in vivo studies, proof of concept 

studies and dose ranging and toxicity studies conducted in connection with the development of our 

product candidates;  

costs for the production of QR-010 and QR-110 compounds, including the costs of GMP batches in 

preparation of our clinical studies;  

increased laboratory costs including purchases of compounds and laboratory materials used by the 

research and development staff in proportion to the increase in the number of employees, and 

increased costs for the use of laboratories;  

increased project-related consultancy costs, including regulatory and intellectual property support; and  

increased share-based compensation, reflecting grants of share options to research and development 

staff made after we adopted our Option Plan in September 2013.  

General and administrative costs increased to € 9,478,000 in 2016 from € 6,837,000 in 2015. These general 

and administrative costs comprise employee costs, office costs, general consultancy costs and other costs. As 

a public company, we face increased legal, accounting, administrative and other costs and expenses. The 

increase was primarily related to:  

 

 

 

 

increased staff costs associated with the increase of our general and administrative staff from 27 full-

time equivalent employees at December 31, 2015 to 33 full-time equivalent employees at December 31, 

2016;  

increased office and general costs, including office rent, information technology and communication 

costs, travel costs and office consumables, as well as costs to improve our internal control environment;  

increased costs for legal support, accounting and other consultancy costs; and 

increased share-based compensation, reflecting grants of share options to non-research and 

development staff made after we adopted our Option Plan in September 2013. 

 
 
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In 2016 share-based compensation amounted to € 2,454,000, compared to € 1,212,000 in 2015. Net financial 

income amounted to € 470,000, compared to € 6,171,000 in 2015. Financial income mainly results from 

foreign exchange differences on cash denominated in U.S. dollars and can fluctuate significantly.  

Outlook 

In 2017, we continue to invest in our organization, while we continue our pre-clinical studies and clinical 

development of our product candidates and increase investments in our other research programs. Our goal 

is to realise this at our current operational level. A significant increase in headcount is not expected. We 

believe we have sufficient cash to fund these expenses and to prepare the Company for future growth. Given 

the development stage of the Company, we do not anticipate revenues in the foreseeable future. 

Leiden, March 31, 2017 

On behalf of the Management Board, 

Daniel de Boer 

CEO 

 
 
 
 
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Supervisory Board Report 
ANNUAL REPORT 2016 

Supervisory Board Report 

ProQR Therapeutics has chosen for its governance structure to be a so-called two-tier system. In such a 

setting the Supervisory Board supervises and advises the Management Board in performing their 

management tasks and setting the strategy of the Company. The Supervisory Board as well as its individual 

members act in the interests of ProQR, its business and development and all its stakeholders. 

In 2016, James Shannon was formally appointed as a Board member. James is a seasoned pharma executive 

with ample know-how in drug development from his previous positions as Chief Medical Officer at Glaxo 

SmithKline and Global Head Pharma Development at Novartis. The Supervisory Board and its sub-

committees held frequent and productive interactions with the Executive Board. Where appropriate, decision 

taking was endorsed by the Supervisory Board and matters of both short term as well as long term strategic 

importance were discussed in a constructive and transparent manner. 

Below is a more specific description of the Supervisory Board’s activities during the financial year 2016 and 

other relevant information on its functioning. 

Activities of the Supervisory Board 

The Supervisory Board and the Board of Directors met 5 times during 2016, and have held various additional 

informal meetings and telephone conferences, both collectively and individually. During these meetings, the 

progress of the various projects, the main risks of the business, the funding and the strategic direction of the 

Company were discussed. In addition, a two-day off-site was held during which the long-term strategy of the 

company was discussed. The Supervisory Board meetings were very well attended (100%) and the 

Committees reported back on their activities to the full Supervisory Board on a regular basis. 

Committees of the Supervisory Board 

We have an audit committee, a compensation committee and a nominating and corporate governance 

committee. We have adopted a charter for each of these committees.  

Compensation Committee 

The Compensation Committee has met 2 times in 2016. 

Compensation report 2016 

In September 2014, the supervisory board adopted our Compensation Policy. This Compensation Policy also 

applied to the financial year 2016 and will apply to subsequent years. Attraction and retention of world class 

talent is a prerequisite for the success of ProQR and competitive compensation plays a vital role in our ability 

to achieve this. The Compensation Committee have elected to offer compensation for all employees including 

the Management Board into a fixed annual salary and a variable, performance related, short and long term 
incentive element. The Compensation Policy is designed based on the following principles: 

 

Three compensation pillars consisting of: 

 

 

 

Annual Base Salary; 

Short Term Incentive (annual cash bonus); 

Long Term Incentive (Stock Option Plan); 

 

Flexibility: The Compensation Policy should provide flexibility to allow the Supervisory Board, acting on 

the recommendation of the Compensation Committee, to reward the Management Board in a fair and 

equitable manner; 

 
 
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Supervisory Board Report 
ANNUAL REPORT 2016 

 

 

 

 

 

 

This Compensation Policy should drive the right kind of management behaviour, discourage unjustified 

risk taking and minimise any gaming opportunity; 

This Compensation Policy should pay for performance, considering not only the measurable financial 

performance of / or milestones achieved by the Company, but also, where appropriate, the efforts 

made by the Management Board, individually and as a group, in managing the Company. For the 

variable components, the Compensation Committee performs an analysis of the possible outcomes 

under different scenarios; 

Design of the Compensation Policy shall be based on current legislation applicable in the Netherlands; 

This Compensation Policy shall foster alignment of interests with shareholders;  

The pension of the Management Board shall be based on the defined contribution system; and 

Pay differentials and position within the Company are considered and evaluated regularly.   

Annual Base Salary 

The Compensation Committee reviewed the annual base salary of the Management Board taking into 

consideration the Compensation Reference Group as contained in the Compensation Policy. Based on this 

review the annual base salary levels for 2016 have been set at € 285,000 for the CEO, Daniel de Boer and at  

€ 255,000 for the chief corporate development officer and general counsel, René Beukema. 

Short Term Incentive  

The Compensation Committee reviewed the performance of the Company during 2016 in comparison to the 

objectives and reviewed the achievements of the members of the Management Board versus their personal 

objectives.  

Based on the recommendation of the Compensation Committee, the Supervisory Board decided early 2017 

that the CEO Daniel de Boer has achieved 88.5% and the chief corporate development officer and general 

counsel, René Beukema has achieved 88.5% of the objectives that had been set to determine their individual 

bonus awards for the year 2016. For 2016 the individual bonuses have been set at € 130,927 for Daniel de 

Boer and € 76,430 for René Beukema. These bonuses will be paid in cash in the first quarter of 2017.  

Long Term Incentive 

Based on the recommendation of the Compensation Committee, the Supervisory Board decided to grant 

stock options in 2016 to the CEO, Daniel de Boer and the chief corporate development officer and general 

counsel, René Beukema. Based on this decision stock options with an exercise price of $ 7.23 have been 

granted with respect to 129,727 shares to the CEO, Daniel de Boer and 50,608 shares to the chief corporate 

development officer and general counsel, René Beukema.  

Pensions 

The pension contributions paid during 2016 amount to € 7,050 for the CEO, Daniel de Boer and € 12,926 for 

the chief corporate development officer and general counsel, René Beukema. 

Supervisory board remuneration 

In June 2016, our shareholders approved an amended compensation policy whereby members of our 

supervisory board will receive board fees of € 25,000 per year and the chairperson will receive board fees of  

€ 30,000 per year. In addition, each board committee chairperson will receive € 5,000 per year for service on 

such committee (except for the chairperson for the nominating committee who will receive € 3,000), and each 

other member of a board committee will receive € 3,000 per year for service on such committee. On top of 

that, several supervisory board members were granted options as set out in Note 23 to the financial 

statements or $ 55,000 in cash. 

 
 
 
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Nominating and Corporate Governance Committee 

The chairman of the Nominating and Corporate Governance Committee elected to involve the entire 

Supervisory Board in the selection process of additional Supervisory Board members. Hence no formal 

nomination committee meeting was held. Based on discussions held, it was concluded that the Supervisory 

Board is complete. Currently, no new nominations are considered necessary. 

Audit Committee 

The audit committee met 5 times in 2016. Main topics addressed were the quarterly results, financial risk 

management, compliance and SOx implementation, the audit plan and management letter of the external 

auditor, cash management and corporate governance. 

The audit committee also reviewed ProQR’s annual financial statements, including non-financial information, 

prior to publication thereof. These financial statements for 2016 have been audited and provided with an 

unqualified opinion by our external auditor, Deloitte Accountants B.V., and were extensively discussed with 

the auditors in the meetings of the Supervisory Board, Audit Committee and Management Board on March 

27, 2017. The Supervisory Board is of the opinion that the Financial Statements 2016 meet all requirements 

and recommends that the Annual General Meeting of Shareholders adopts the financial statements and the 

appropriation of net result proposed by the Management Board. 

The Company’s external auditor attended all Audit Committee meetings. The Audit Committee evaluates the 

performance of Deloitte as independent external auditor annually. Due to the limited size of the Company, it 

was concluded that there was currently no need to appoint an internal auditor. 

The Supervisory Board is responsible for the quality of its own performance and it discusses, once a year on 

its own, without the members of the Management Board present, both its own functioning and that of the 

individual members, and the functioning of the Management Board and that of its individual members. The 

Supervisory Board discussed its composition and competencies and concluded no changes are necessary 

based on this review. We feel the additional efforts of all staff at ProQR form a strong foundation for the 

success and growth of the Company and all milestones reached this past year. Therefore, we would like to 

express our thanks to the members of the Management Board, senior management and all other employees 

for their contribution and performance during the year. We thank our shareholders for their continued 

support. 

Leiden, March 31, 2017 

On behalf of the Supervisory Board, 

Dinko Valerio 

Chairman 

 
 
 
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Corporate Governance 
ANNUAL REPORT 2016 

Corporate Governance 

ProQR values the importance of complying with Corporate Governance regulations. At the same time, the 

Board of Directors is of the opinion that certain deviations from the provisions of the Dutch Corporate 

Governance Code 2008 (“DCGC” or “the Code”) are justified, in view of our activities, our size and the specific 

circumstances in which we operate. In such cases, which are mentioned in this corporate governance 

statement, we apply the “comply or explain” principle.  

Deviations from certain aspects of the Code, when deemed necessary in the interests of the Company, will be 

disclosed in the Annual Report. Deviations are due to our Company being listed in the United States with 

most of our investors being outside of the Netherlands, as well as to the international business focus of our 

Company. As a Company listed on NASDAQ, we comply with NASDAQ’s corporate governance listing 

standards, except for instances where we follow our home country’s corporate governance practices in lieu of 

certain NASDAQ’s standards as explained below, as NASDAQ investors are more familiar with NASDAQ’s rules 

than with the Code. 

In this report, the Company addresses its overall corporate governance structure and states to what extent 

and how it applies the principles and best practice provisions of the Code. This report also includes the 

information which the Company is required to disclose pursuant to the Dutch governmental decree on Article 

10 Takeover Directive and the governmental decree on Corporate Governance.  

Substantial changes in the Company’s corporate governance structure and in the Company’s compliance with 

the DCGC, if any, will be submitted to the General Meeting of Shareholders for discussion under a separate 

agenda item. The Supervisory Board and the Management Board, which are responsible for the corporate 

governance structure of the Company, are of the opinion that the principles and best practice provisions of 

the DCGC that are addressed to the management board and the supervisory board, interpreted and 

implemented in line with the best practices followed by the Company, are being applied.  

The full text of the DCGC can be found at the website of the Monitoring Commission Corporate Governance 

Code (www.commissiecorporategovernance.nl) and for an overview of our conformity with the Code the 

following documents are available at our website (www.ProQR.com): audit committee charter, compensation 

committee charter, nominating and corporate governance committee charter and our code of business 

conduct and ethics. 

Management Board 

The Management Board’s role is setting and achieving the operational and financial objectives of the 

company in order to pursue the long-term success of ProQR. The Board does so by assuming the authority 

and responsibilities assigned to it by Dutch corporate law and by combining expertise and experience with 

entrepreneurial leadership. The management Board operates under the supervision of the supervisory 

board. The management board is required to:  

 

 

 

keep the supervisory board informed in a timely manner in order to allow the supervisory board to 

carry out its responsibilities;  

consult with the supervisory board on important matters; and  

submit important decisions to the supervisory board for its approval.  

 
 
 
 
 
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Corporate Governance 
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Our management board may perform all acts necessary or useful for achieving our corporate purposes, 

other than those acts that are prohibited by law or by our articles of association. The management board as a 

whole and any management board member individually, are authorized to represent us in dealings with third 

parties.  

Under our articles of association, the number of management board members is determined by the 

supervisory board, and the management board must consist of at least one member. The supervisory board 

elects a CEO from among the members of the management board.  

Members of the management board are appointed by the general meeting of shareholders upon a binding 

nomination of the supervisory board. Our general meeting of shareholders may at all times deprive such a 

nomination of its binding character by a resolution passed by at least two-thirds of the votes cast 

representing more than 50% of our issued share capital, following which our supervisory board shall draw up 

a new binding nomination.  

Our management board rules provide that, unless the resolution appointing a management board member 

provides otherwise, members of our management board will serve for a maximum term of four years. Our 

articles of association provide that the management board members must retire periodically in accordance 

with a rotation schedule adopted by the management board. A management board member who retires in 

accordance with the rotation schedule may be reappointed immediately for a term of not more than four 

years at a time.  

Supervisory Board  

Our supervisory board is responsible for the supervision of the activities of our management board and our 

Company’s general affairs and business. Our supervisory board may, also on its own initiative, provide the 

management board with advice and may request any information from the management board that it deems 

appropriate. In performing its duties, the supervisory board is required to act in the interests of our Company 

(including its stakeholders) and its associated business as a whole. The members of the supervisory board 

are not authorized to represent us in dealings with third parties.  

Pursuant to Dutch law, members of the supervisory board must be natural persons. Under our articles of 

association, the number of supervisory board members is determined by our supervisory board itself, 

provided there will be at least three supervisory board members. Our articles of association provide that 

members of the supervisory board are appointed by the general meeting of shareholders upon a binding 

nomination by the supervisory board. Our general meeting of shareholders may at all times deprive such a 

nomination of its binding character by a resolution passed by at least two-thirds of the votes cast 

representing more than 50% of our issued share capital, following which our supervisory board shall draw up 

a new binding nomination.  

Our supervisory board rules provide that members of our supervisory board will serve for a maximum 

duration of three terms of four years. Our articles of association provide that the supervisory board members 

must retire periodically in accordance with a rotation schedule adopted by the supervisory board. A 

supervisory board member who retires in accordance with the rotation schedule can be reappointed 

immediately. The supervisory board appoints a chairman from among its members.  

With the exception of Antoine Papiernik, each member of our supervisory board has been and remains fully 

independent within the meaning of best practice provision III.2.2 of the DCGC. Mr. Papiernik is affiliated with 

Sofinnova which holds 11.9 % of our shares and is therefore not independent within the meaning of best 

practice provision III.2.2.f of the Code. We feel this deviation is justified by his specific knowledge and 

 
 
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Corporate Governance 
ANNUAL REPORT 2016 

experience of our business. Based on the above, we comply with best practice provision III.2.1 of the DCGC, 

according to which not more than one supervisory board member is allowed not to be independent. 

Under our articles of association, the general meeting of shareholders may suspend or remove supervisory 

board members at any time. A resolution of our general meeting of shareholders to suspend or remove a 

supervisory board member may be passed by a simple majority of the votes cast, provided that the 

resolution is based on a proposal by our supervisory board. In the absence of a proposal by our supervisory 

board, a resolution of our general meeting of shareholders to suspend or remove a supervisory board 

member shall require a majority of at least two-thirds of the votes cast representing more than 50% of our 

issued share capital.  

In a meeting of the supervisory board, each supervisory board member is entitled to cast one vote. A 

supervisory board member may grant a written proxy to another supervisory board member to represent 

him at a meeting of the supervisory board. All resolutions by our supervisory board are adopted by a simple 

majority of the votes cast unless our supervisory board rules provide otherwise. In case of a tie in any vote of 

the supervisory board, the chairman of the supervisory board shall have the casting vote. Our supervisory 

board may also adopt resolutions outside a meeting, provided that such resolutions are adopted in writing, 

all supervisory board members are familiar with the resolution to be passed and provided that no 

supervisory board member objects to such decision-making process. 

Committees of the Supervisory Board  

We have an audit committee, a compensation committee and a nominating and corporate governance 

committee. We have adopted a charter for each of these committees.  

Audit Committee  

Our audit committee consists of Paul Baart (chairman), Alison Lawton and James Shannon. James Shannon 

was appointed at our AGM on June 21, 2016. Until that date, Antoine Papiernik was member of the audit 

committee. Each member satisfies the independence requirements of the NASDAQ listing standards / Rule 

10A-3(b)(1) under the Exchange Act, and each member meets the criteria for independence set forth in best 

practice III.2.2 of the DCGC. Paul Baart qualifies as an “audit committee financial expert,” as defined by the 

SEC in Item 16A: “Audit Committee Financial Expert” and as determined by our supervisory board. The audit 

committee oversees our accounting and financial reporting processes and the audits of our financial 

statements. The audit committee is responsible for, among other things:  

 

 

 

 

 

 

 

 

the operation of the internal risk management and control systems, including supervision of the 

enforcement of relevant primary and secondary legislation, and supervising the operation of codes of 

conduct; 

the provision of financial information by the company (choice of accounting policies, application and 

assessment of the effects of new rules, information about the handling of estimated items in the 

financial statements, forecasts, work of internal and external auditors, etc.); 

compliance with recommendations and observations of internal and external auditors; 

reviewing the need for an internal audit function; 

the policy of the company on tax planning; 

relations with the external auditor, including, in particular, his independence, remuneration and any 

non-audit services for the company; 

the financing of the company; and 

the applications of information and communication technology. 

 
 
 
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Corporate Governance 
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Compensation Committee  

Our compensation committee consists of James Shannon (chairman), Dinko Valerio and Alison Lawton. Each 

member satisfies the independence requirements of the NASDAQ listing standards / Rule 10A-3(b)(1) under 

the Exchange Act, and each member meets the criteria for independence set forth in best practice III.2.2 of 

the DCGC. The compensation committee assists our supervisory board in reviewing and approving or 

recommending our compensation structure, including all forms of compensation relating to our supervisory 

board members, our management board members and our officers. Members of our management board 

may not be present at any compensation committee meeting while their compensation is deliberated. 

Subject to and in accordance with the terms of the compensation policy approved by our general meeting of 

shareholders from time to time, as required by Dutch law, the compensation committee is responsible for, 

among other things:  

  making a proposal to the supervisory board for the remuneration policy to be pursued; 

  making a proposal for the remuneration of the individual members of the management board, for 

adoption by the supervisory board; such proposal shall, in any event, deal with: (i) the remuneration 

structure and (ii) the amount of the fixed remuneration, the shares and/or options to be granted and/or 

other variable remuneration components, pension rights, redundancy pay and other forms of 

compensation to be awarded, as well as the performance criteria and their application; and 

 

preparing the remuneration report as referred to in best practice provision II.2.12. 

Our supervisory board may also delegate certain tasks and powers under our Option Plan to the 

compensation committee.  

Nominating and Corporate Governance Committee  

Our nominating and corporate governance committee consists of Dinko Valerio (chairman), Henri Termeer 

and Paul Baart. Each member satisfies the independence requirements of the NASDAQ listing standards as 

well as the criteria for independence set forth in best practice III.2.2 of the DCGC. The nominating and 

corporate governance committee assists our supervisory board in selecting individuals qualified to become 

our supervisory board members and management board members and in determining the composition of 

the management board, supervisory board and its committees and our officers. The nominating and 

corporate governance committee is responsible for, among other things:  

 

 

 

drawing up selection criteria and appointment procedures for supervisory board members and 

management board members; 

periodically assessing the size and composition of the supervisory board and the management board, 

and making a proposal for a composition profile of the supervisory board; 

periodically assessing the functioning of individual supervisory board members and management board 

members, and reporting on this to the supervisory board; 

  making proposals for appointments and reappointments; and 

 

supervising the policy of the management board on the selection criteria and appointment procedures 

for senior management. 

Insurance and Indemnification of Management Board and Supervisory Board Members  

Under Dutch law, management board members, supervisory board members and certain other 

representatives may be held liable for damages in the event of improper or negligent performance of their 

duties. They may be held jointly and severally liable for damages to the Company for infringement of the 

articles of association or of certain provisions of the Dutch Civil Code. They may also be liable towards third 

parties for infringement of certain provisions of the Dutch Civil Code. In certain circumstances they may also 

incur additional specific civil and criminal liabilities.  

 
 
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Corporate Governance 
ANNUAL REPORT 2016 

Our articles of association provide that we will indemnify our management board members, supervisory 

board members, former management board members and former supervisory board members (each an 

“Indemnified Person”) against (i) any financial losses or damages incurred by such Indemnified Person and 

(ii) any expense reasonably paid or incurred by such Indemnified Person in connection with any threatened, 

pending or completed suit, claim, action or legal proceedings, whether civil, criminal, administrative or 

investigative and whether formal or informal, in which he becomes involved, to the extent this relates to his 

position with the Company, in each case to the fullest extent permitted by applicable law. No indemnification 

shall be given to an Indemnified Person (a) if a Dutch court has established, without possibility for appeal, 

that the acts or omissions of such Indemnified Person that led to the financial losses, damages, suit, claim, 

action or legal proceedings result from either an improper performance of his duties as an officer of the 

Company or an unlawful or illegal act and (b) to the extent that his financial losses, damages and expenses 

are covered by an insurance and the insurer has settled these financial losses, damages and expenses (or has 

indicated that it would do so). Our supervisory board may stipulate additional terms, conditions and 

restrictions in relation to such indemnification.  

Board composition and diversity 

Our management board comprised two persons in 2015, both of whom are male. Our supervisory board has 

five male members and one female member. As a Company, we support diversity of culture, gender and age 

in our Company. Our current management board and supervisory board members were selected based on 

the required profile and talent and abilities of the members without positive or negative bias on gender, 

culture or age. In the future, this will continue to be our basis for selection of new board members. 

Controls and procedures 

Our managing board and our chief financial officer, after evaluating the effectiveness of our disclosure 

controls and procedures (as defined in Rule 13a-15(e) under the Exchange Act) as of December 31, 2015, have 

concluded that based on the evaluation of these controls and procedures required by Rule 13a-15(b) of the 

Exchange Act, our disclosure controls and procedures were effective. The internal risk management and 

control systems provide reasonable assurance that the financial reporting does not contain any errors of 

material importance and that the risk management and control systems worked properly in the year under 

review. 

Risk factors and the risk management approach, as well as the sensitivity of our results to external factors 

and variables are described in more detail in ”Risk Management”. Our internal control system has been 

discussed with the Audit Committee and the external auditors. 

In view of the requirements of the U.S. Securities Exchange Act, procedures are in place to enable the CEO 

(chief executive officer) and the CFO (chief financial officer) to provide certifications with respect to the 

Annual Report on Form 20F. 

General Meeting of Shareholders 

General meetings of shareholders are held in Leiden, Oegstgeest, Leidschendam, Katwijk, Noordwijk, 

Wassenaar, Amsterdam, Rotterdam, The Hague, or Schiphol Airport (municipality of Haarlemmermeer) 

(Schiphol Airport), the Netherlands. All shareholders and others entitled to attend general meetings of 

shareholders are authorized to attend the general meeting of shareholders, to address the meeting and, in 

so far as they have such right, to vote, either in person or by proxy.  

Annually, at least one general meeting of shareholders shall be held, within six months after the end of our 

financial year. A general meeting of shareholders shall also be held within three months after our 

management board has considered it to be likely that the Company’s equity has decreased to an amount  

 
 
 
 
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Corporate Governance 
ANNUAL REPORT 2016 

equal to or lower than half of its paid up and called up capital. If the management board and supervisory 

board have failed to ensure that such general meetings of shareholders as referred to in the preceding 

sentences are held in a timely fashion, each shareholder and other person entitled to attend shareholders’ 

meetings may be authorized by the Dutch court to convene the general meeting of shareholders. 

Our management board and our supervisory board may convene additional extraordinary general meetings 

of shareholders whenever they so decide. Pursuant to Dutch law, one or more shareholders and/or others 

entitled to attend general meetings of shareholders, alone or jointly representing at least ten percent of our 

issued share capital may on their application, be authorized by the Dutch court to convene a general meeting 

of shareholders. The Dutch court will disallow the application if it does not appear to it that the applicants 

have previously requested that the management board or supervisory board convenes a shareholders’ 

meeting and neither the management board nor the supervisory board has taken the necessary steps so that 

the shareholders’ meeting could be held within six weeks after the request.  

General meetings of shareholders are convened by a notice which includes an agenda stating the items to be 

discussed. For the annual general meeting of shareholders the agenda will include, among other things, the 

adoption of our annual accounts, the appropriation of our profits or losses, discharge of the members of the 

Management Board for their management, discharge of the members of the Supervisory Board for their 

supervision on the management and proposals relating to the composition and filling of any vacancies of the 

management board or supervisory board. In addition, the agenda for a general meeting of shareholders 

includes such items as have been included therein by our management board or our supervisory board. 

Pursuant to Dutch law, one or more shareholders and/or others entitled to attend general meetings of 

shareholders, alone or jointly representing at least 3% of the issued share capital have the right to request 

the inclusion of additional items on the agenda of shareholders’ meetings. Such requests must be made in 

writing, substantiated, or by a proposal for a resolution and received by us no later than the sixtieth day 

before the day the relevant general meeting is held. No resolutions will be adopted on items other than those 

which have been included in the agenda.  

We will give notice of each general meeting of shareholders by publication on our website and, to the extent 

required by applicable law, in a Dutch daily newspaper with national distribution, and in any other manner 

that we may be required to follow in order to comply with Dutch law, applicable stock exchange and SEC 

requirements. We will observe the statutory minimum convening notice period for a general meeting of 

shareholders.  

Pursuant to our articles of association, our management board may determine a record date 

(registratiedatum) of 28 calendar days prior to a general meeting of shareholders to establish which 

shareholders and others with meeting rights are entitled to attend and, if applicable, vote in the general 

meeting of shareholders. The record date, if any, and the manner in which shareholders can register and 

exercise their rights will be set out in the convocation notice of the general meeting. Our articles of 

association provide that a shareholder must notify the Company in writing of his identity and his intention to 

attend (or be represented at) the general meeting of shareholders, such notice to be received by us ultimately 

on the seventh day prior to the general meeting. If this requirement is not complied with or if upon direction 

of the Company to that effect no proper identification is provided by any person wishing to enter the general 

meeting of shareholders, the chairman of the general meeting of shareholders may, in his sole discretion, 

refuse entry to the shareholder or his proxy holder.  

Pursuant to our articles of association, our general meeting of shareholders is chaired by the chairman of our 

supervisory board. If the chairman of our supervisory board is absent and has not charged another person to 

chair the meeting in his place, the supervisory board members present at the meeting shall appoint one of  

 
 
 
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Corporate Governance 
ANNUAL REPORT 2016 

them to be chairman. If no supervisory board members are present at the general meeting of shareholders, 

the general meeting of shareholders will be chaired by our CEO or, if our CEO is absent, another managing 

board member present at the meeting and, if none of them is present, the general meeting shall appoint its 

own chairman. The person who should chair the meeting may appoint another person in his stead.  

The chairman of the general meeting may decide at his discretion to admit other persons to the meeting. The 

chairman of the general meeting shall appoint another person present at the shareholders’ meeting to act as 

secretary and to minute the proceedings at the meeting. The chairman of the general meeting may instruct a 

civil law notary to draw up a notarial report of the proceedings at the Company’s expense, in which case no 

minutes need to be taken. The chairman of the general meeting is authorized to eject any person from the 

general meeting of shareholders if the chairman considers that person to disrupt the orderly proceedings. 

The general meeting of shareholders shall be conducted in the English language. 

Voting Rights and Quorum Requirements  

In accordance with Dutch law and our articles of association, each issued ordinary share and preferred share 

confers the right on the holder thereof to cast one vote at the general meeting of shareholders. The voting 

rights attached to any shares held by us or our direct or indirect subsidiaries are suspended as long as they 

are held in treasury. Dutch law does not permit cumulative voting for the election of management board 

members or supervisory board members.  

Voting rights may be exercised by shareholders or by a duly appointed proxy holder (the written proxy being 

acceptable to the chairman of the general meeting of shareholders) of a shareholder, which proxy holder 

need not be a shareholder. Our articles of association do not limit the number of shares that may be voted by 

a single shareholder.  

Under our articles of association, blank votes, abstentions and invalid votes shall not be counted as votes 

cast. Further, shares in respect of which a blank or invalid vote has been cast and shares in respect of which 

the person with meeting rights who is present or represented at the meeting has abstained from voting are 

counted when determining the part of the issued share capital that is present or represented at a general 

meeting of shareholders. The chairman of the general meeting shall determine the manner of voting and 

whether voting may take place by acclamation.  

In accordance with Dutch law and generally accepted business practices, our articles of association do not 

provide quorum requirements generally applicable to general meetings of shareholders. To this extent, our 

practice varies from the requirement of NASDAQ Listing Rule 5620(c), which requires an issuer to provide in 

its bylaws for a generally applicable quorum, and that such quorum may not be less than one-third of the 

outstanding voting shares.  

Resolutions of the general meeting of shareholders are adopted by a simple majority of votes cast without 

quorum requirement, except where Dutch law or our articles of association provide for a special majority 

and/or quorum in relation to specified resolutions.  

Anti-takeover provisions 

We have adopted several provisions that may have the effect of making a takeover of our Company more 

difficult or less attractive, including: 

 

granting a perpetual and repeatedly exercisable call option to a protection foundation, which confers 

upon the protection foundation the right to acquire, under certain conditions, the number of preferred 

shares in the capital of the Company. The issuance of such preferred shares will occur upon the 

protection foundation’s exercise of the call option and will not require shareholder consent; 

 
 
 
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Corporate Governance 
ANNUAL REPORT 2016 

 

 

 

the staggered four-year terms of our supervisory board members, as a result of which only 

approximately one-fourth of our supervisory board members will be subject to election in any one year; 

a provision that our management board members and supervisory board members may only be 

appointed upon a binding nomination by our supervisory board, which can be set aside by a two-thirds 

majority of our shareholders representing more than half of our issued share capital;  

a provision that our management board members and supervisory board members may only be 

removed by our general meeting of shareholders by a two-thirds majority of votes cast representing 

more than 50% of our issued share capital (unless the removal was proposed by the supervisory board); 

and  

 

a requirement that certain matters, including an amendment of our articles of association, may only be 

brought to our shareholders for a vote upon a proposal by our management board that has been 

approved by our supervisory board.  

Deviations from the Dutch Corporate Governance Code 

The Code contains a “comply-or-explain” principle, offering the possibility to deviate from the Code as long as 

any such deviations are explained. We acknowledge the importance of good corporate governance. However, 

at this stage, we do not comply with all the provisions of the DCGC for specific reasons. The main deviations 

from best practice provisions are listed below. 

 

Pursuant to the best practice provisions II.2.4 and II.2.5 of the DCGC, options granted to our 

management board members should not be exercisable during the first three years after the date of 

grant; shares granted to our management board members for no financial consideration should be 

retained by them for a period of at least five years or until they cease to hold office, whichever is the 

shorter period; and the number of options and/or shares granted to our management board members 

should be dependent on the achievement of pre-determined performance criteria. We do not intend to 

comply with all of the above requirements as we believe it is in the best interest of the company to 

attract and retain highly skilled management board members on conditions based on market practice, 

as we believe these are.  

 

Pursuant to best practice provision II.2.8 the remuneration of the management board in the event of 

dismissal may not exceed one year’s salary. The management services agreements with our 

management board members provide for a lump-sum equal to 24 months of the individual’s monthly 

gross fixed salary. Based on the risk profile of the Company and to be able to attract highly skilled 

management, we assumed this period to be appropriate. 

 

Best practice provision III.7.1 prohibits the granting of shares or rights to shares to members of the 

supervisory board as compensation. It is common practice for companies listed on the NASDAQ Global 

Market to grant shares to the members of the supervisory board as compensation, in order to align the 

interests of the members of the supervisory board with our interests and those of our shareholders, 

and we have granted and expect to grant options to acquire ordinary shares to some of our supervisory 

board members.  

 

Pursuant to best practice provision III.7.2, any shares held by supervisory board members are long-term 

investments. We do not request our supervisory board members to comply with this provision. We 

believe it is in the best interest of the Company not to apply this provision in order to be able to attract 

and retain highly skilled supervisory board members on internationally competitive terms. 

 

Best practice provision IV.1.1 provides that the general meeting of shareholders may pass a resolution 

to cancel the binding nature of a nomination for the appointment of a member of the management 

board or of the supervisory board or a resolution to dismiss such member by an absolute majority of 

the votes cast. It may be provided that such majority should represent a given proportion of the issued 

capital, but this proportion may not exceed one third. In addition, best practice IV.1.1. provides that if 

such proportion of the share capital is not represented at the meeting, but an absolute majority of the 

votes cast is in favor of a resolution to cancel the binding nature of the nomination, a new general 

 
 
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Corporate Governance 
ANNUAL REPORT 2016 

meeting of shareholders will be convened where the resolution may be adopted by absolute majority, 

regardless of the proportion of the share capital represented at the meeting. Our articles of association 

provide that these resolutions can only be adopted with at least a 2/3 majority which must represent 

more than 50% of our issued capital, and that no such second meeting will be convened, because we 

believe that the decision to overrule a nomination by the management board or the supervisory board 

for the appointment or dismissal of a member of our management board or of our supervisory board 

must be widely supported by our shareholders.  

 

Best practice provision IV.3.1 stipulates that meetings with analysts, presentations to analysts, 

presentations to investors and institutional investors and press conferences must be announced in 

advance on the Company’s website and by means of press releases. Provision must be made for all 

shareholders to follow these meetings and presentations in real time, for example by means of 

webcasting or telephone. After the meetings, the presentations must be posted on the Company’s 

website. We believe that enabling shareholders to follow in real time all the meetings with analysts, 

presentations to analysts and presentations to investors, would create an excessive burden on our 

resources and therefore, we do not intend to comply with all of the above requirements.  

 

Best practice provision IV.3.13 stipulates that an outline policy on bilateral contacts with the 

shareholders shall be formulated and published on the Company’s website. The Company has not 

formulated such policy as it believes this is already covered by our regular process for public disclosure 

of information. 

Summary of significant corporate governance differences from NASDAQ Listing Standards 

Our ordinary shares are listed on NASDAQ. The Sarbanes-Oxley Act of 2002, as well as related rules 

subsequently implemented by the SEC, requires foreign private issuers, including our Company, to comply 

with various corporate governance practices. As a foreign private issuer, subject to certain exceptions, the 

NASDAQ listing standards permit a foreign private issuer to follow its home country practice in lieu of the 

NASDAQ listing standards. Our corporate governance practices differ in certain respects from those that U.S. 

companies must adopt in order to maintain a NASDAQ listing. The home country practices followed by our 

Company in lieu of NASDAQ rules are described below:  

  We do not intend to follow NASDAQ’s quorum requirements applicable to meetings of shareholders. In 

accordance with Dutch law and generally accepted business practice, our articles of association do not 

provide quorum requirements generally applicable to general meetings of shareholders.  

  We do not intend to follow NASDAQ’s requirements regarding the provision of proxy statements for 

general meetings of shareholders. Dutch law does not have a regulatory regime for the solicitation of 

proxies and the solicitation of proxies is not a generally accepted business practice in the Netherlands. 

We do intend to provide shareholders with an agenda and other relevant documents for the general 

meeting of shareholders and shareholders will be entitled to give proxies and voting instructions to us 

and/or third parties. 

We intend to take all actions necessary for us to maintain compliance as a foreign private issuer under the 

applicable corporate governance requirements of the Sarbanes-Oxley Act of 2002, the rules adopted by the 

SEC and NASDAQ’s listing standards.  

New Dutch Corporate Governance Code 

A new DCGC has been issued by the DCGC Monitoring Committee as per December 8, 2016 and will replace 

the DCGC 2008 from financial year 2017 and onwards. During 2017 the DCGC will be formally approved by 

the Dutch Authoritities and incorporated in Dutch law.  

 
 
 
 
 
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Corporate Governance 
ANNUAL REPORT 2016 

The amendments will focus on the following topics: 

  More focus on long term value creation by management board and supervisory board; 

 

 

 

 

Importance of risk management and strengthening of internal control; 

Company culture should be an explicit part of the corporate governance structure; 

A number of provisions with respect to remuneration will be revised; 

Deviations should be more carefully explained. The new code will include a number of requirements for 

reporting of deviations. 

Based on our initial assessment, we expect no major impact on ProQR Corporate Governance. 

 
 
 
PAGE 45 / 91 
Risk Management 
ANNUAL REPORT 2016 

Risk Management 

Our business is subject to numerous risks and uncertainties. In the table below, we focus on the key risks and 

uncertainties the Company currently faces. For the avoidance of doubt, this does not mean that the risks 

which were previously signaled and not described here are no longer relevant. For a complete understanding 

of the risks that we face you should also read the full list of risks and uncertainties as disclosed in item 3.D 

Risk Factors of the annual report on Form 20-F. Some of these risks and uncertainties are outside the control 

of the Company, others may be influenced or mitigated. In 2015, we have implemented a Risk & Control 

framework, based on the COSO 2013 internal control framework, for enhancing our control environment as 

well as compliance with the U.S. SEC’s Sarbanes Oxley (SOx) Act of 2002, which we are required to as a 

company listed on the NASDAQ.  As part of the SOx implementation program, our Risk & Control framework 

was further enhanced in 2016, focusing on IT and entity level controls. Improvement of our Risk & Control 

framework is an ongoing effort for the Company. 

Our main risks are those that threaten the achievement of the Company’s corporate objectives, including 

compliance. If any of these risks actually occurs, our business, prospects, operating results and financial 

condition could suffer materially. These risks include, but are not limited to, the following:  

Risk related to 

Risk area 

Development and 
Regulatory Approval of 
our Product Candidates 

Our products will not be able to 
demonstrate safety and efficacy 
in the preclinical studies and 
clinical trials that are needed to 
obtain product approval. 

Expected impact upon 
materialization 

The Company will be unable to 
commercialize the product and 
therefore generate revenues. 

The regulatory approval process 
is lengthy, time-consuming and 
unpredictable and products 
developed may ultimately not 
lead to regulatory approval of 
the product. 

Failure to comply with the 
requirements in the regulatory 
process could result in delays, 
suspension, refusals and 
withdrawal of approvals as well 
as fines. 

We may not able to maintain 
orphan product status for QR-
010 and QR-110 or obtain such 
status for QR-313. 

We may not benefit from 
rewards including fee 
reductions and market 
exclusivity. Sales could be 
impacted if other products are 
granted authorization for the 
same indications as QR-010 and 
QR-110. 

Risk-mitigating actions 

This is an inherent risk with drug 
development as the safety and 
efficacy of products can only be 
assessed when these studies 
are conducted. However, the 
Company has multiple products 
in the pipeline and therefore is 
diversified. The Company also 
monitors the progress of the 
programs and aims to make 
decisions that mitigate safety 
and efficacy related risks. 

Although the Company 
monitors the regulatory 
landscape and engages with the 
authorities when it deems that 
necessary, this is an inherent 
risk in biotech drug 
development and therefore has 
limited mitigation abilities. 

We take orphan drug 
requirements into consideration 
in the design of our clinical 
development plans. 

We may be precluded from 
obtaining marketing 
authorization for our products 
when our competitors have 

We may encounter delays in 
marketing our products for a 
significant period of time. 

We take orphan drug 
requirements into consideration 
in the design of our clinical 
development plans. 

 
 
 
 
 
 
 
 
 
 
PAGE 46 / 91 
Risk Management 
ANNUAL REPORT 2016 

obtained market exclusivity 
before we do. 

Risk related to 

Risk area 

Capital Needs and 
Financial Position 

The Company depends largely 
on equity financing and 
financing through third party 
collaboration agreements and 
government subsidies.  

Dependence on Third 
Parties 

Intellectual Property 

The Company relies upon third-
party contractors and service 
providers for the execution of 
several aspects of its preclinical 
and clinical development 
programs, which include CRO’s, 
third party manufacturers and 
other service providers. 

The Company is highly 
dependent on its portfolio of 
patents and other intellectual 
property, proprietary 
information and knowhow and 
its ability to protect and enforce 
these assets.  

The Company is subject to the 
risk of infringing third party 
intellectual property rights. 

Commercialization of Our 
Product Candidates 

We face competition from 
entities that have developed or 
may develop product 
candidates for our target 
indications.  

Expected impact upon 
materialization 

Risk-mitigating actions 

Volatility of the Company’s 
share price, failure to deliver 
under collaboration agreements 
and/or the reevaluation or 
withdrawal of government 
subsidies may have a negative 
impact on the Company's ability 
to obtain future financing. 

The ability of third party 
financing is dependent on 
external factors and is therefore 
not entirely in the Company’s 
control. The Company monitors 
the market conditions for 
opportunities to add additional 
capital. 

Failure of third parties to 
provide services of a suitable 
quality and within acceptable 
timeframes may cause delay or 
failure of the Company's 
development programs. 

The Company reviews and 
monitors the activities of the 
third parties. These include 
setting contractual deliverables, 
quality assurance audits and 
performance reports, among 
other activities. 

Inadequate intellectual property 
protection or enforcement may 
impede the Company’s ability to 
compete effectively. If the 
Company is not able to protect 
its trade secrets, know-how or 
other proprietary information, 
the value of its technology and 
product candidates could be 
significantly diminished. 
Intellectual property rights 
conflicts may result in costly 
litigation and could result in the 
Company having to pay 
substantial damages or limit the 
Company’s ability to 
commercialize its product 
candidates. 

If our competitors develop 
technologies or product 
candidates more rapidly than 
we do or their technologies, 
including delivery technologies, 
are more effective, our ability to 
develop and successfully 
commercialize our product 
candidates may be adversely 
affected. 

The Company files and 
prosecutes patent applications 
to protect its products and 
technologies to the best of its 
knowledge and with assistance 
from internal and external 
counsel. Prior to disclosing any 
confidential information to third 
parties, the Company maintains 
strict confidentiality standards 
and agreements for 
collaborating parties.  

Competition is an inherent risk 
for any industry including drug 
development. Through our IP 
strategy and orphan drug 
designation application, we 
attempt to have data exclusivity 
for our products. Development 
in other companies is essentially 
out of our control but we 
monitor the competitive 
landscape and incorporate that 
into our business strategy. 

In addition to the above key risks, the Company’s activities expose it to a variety of financial risks: market risk 

(including currency risk, interest rate risk and price risk), credit risk and liquidity risk. Unfavorable exchange 

rate developments and historically low interest rates may impact the financial income of the Company. The 

Company has a cash management policy in place to minimize potential adverse effects resulting from 

unpredictability of financial markets on the Company’s financial performance. 

 
 
 
 
 
 
 
 
 
 
 
PAGE 47 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

Financial Statements 2016 

Consolidated statement of financial position at December 31, 2016 

ASSETS 

Non-current assets 

Intangible assets 

Property, plant and equipment 

Current assets 

Social securities and other taxes 

Prepayments and other receivables 

Cash and cash equivalents 

TOTAL ASSETS 

EQUITY 

Shareholders' equity 

Share capital 

Share premium reserve 

Equity settled employee benefits reserve 

Translation reserve 

Accumulated deficit 

LIABILITIES 

Non-current liabilities 

Finance lease liabilities 

Borrowings 

Current liabilities 

Finance lease liabilities 

Trade payables 

Social securities and other taxes 

Pension premiums 

Deferred income 

Other current liabilities 

Note 

December 31, 
2016 

December 31, 
2015 

€ 1,000 

€ 1,000 

7 

8 

9 

10 

11 

12 

13 

14 

90 

3,438 

3,528 

395 

2,420 

59,200 

62,015 

141 

2,199 

2,340 

956 

1,948 

94,865 

97,769 

65,543 

100,109 

934 

123,597 

4,353 

(15) 

(75,733) 

53,136 

-- 

5,697 

5,697 

-- 

328 

312 

13 

-- 

6,057 

6,710 

934 

123,595 

1,899 

1 

(36,630) 

89,799 

-- 

4,824 

4,824 

15 

885 

235 

16 

144 

4,191 

5,486 

TOTAL EQUITY AND LIABILITIES 

65,543 

100,109 

The accompanying notes are an integral part of these financial statements. 

 
 
 
 
 
 
 
 
 
 
 
  
  
 
 
 
 
 
 
 
 
 
 
  
 
 
 
 
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
 
 
 
 
 
PAGE 48 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

Consolidated statement of profit or loss and comprehensive income for the year ended 
December 31, 2016 

Other income 

Research and development costs 

General and administrative costs 

Total operating costs 

Operating result 

Financial income and expense 

Result before corporate income taxes 

Corporate income taxes 

Note 

2016 

€ 1,000 

2015 

€ 1,000 

15 

16 

18 

19 

1,828 

3,235 

(31,923) 

(9,478) 

(23,401) 

(6,837) 

(41,401) 

(30,238) 

(39,573) 

470 

(27,003) 

6,171 

(39,103) 

(20,832) 

-- 

-- 

Result for the year (attributable to equity holders of the Company) 

(39,103) 

(20,832) 

Other comprehensive income 

Items that will never be reclassified to profit or loss 

Items that are or may be reclassified to profit or loss 

Foreign operations – foreign currency translation differences 

(16) 

1 

Total comprehensive income for the year 
(attributable to equity holders of the Company) 

(39,119) 

(20,831) 

Share information 

20 

Weighted average number of shares outstanding1 

23,346,507 

23,343,262 

Earnings per share attributable to the equity holders  
of the Company (expressed in Euro per share) 

Basic earnings per share1 

Diluted earnings per share1 

The accompanying notes are an integral part of these financial statements. 

(1.68) 

(1.68) 

(0.89) 

(0.89) 

1 Basic and diluted earnings are equal due to the anti-dilutive nature of the options outstanding since the Company is loss-
making. 

 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
 
 
 
  
 
 
 
 
  
 
 
 
 
  
 
 
 
 
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
 
 
 
  
 
 
 
 
 
 
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
                                                               
 
 
PAGE 49 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

Consolidated statement of changes in equity for the year ended December 31, 2016 

Share 
Capital 

Share 
Premium 

Equity 
Settled 
Employee 
Benefit 
Reserve 

Translation 
Reserve 

Accumulated 
Deficit 

Total 
Equity 

€ 1,000 

€ 1,000 

€ 1,000 

€ 1,000 

€ 1,000 

€ 1,000 

Balance at January 1, 2015 

934 

123,581 

687 

Result for the year 

Other comprehensive income 

Recognition of share-based 
payments 

Share options exercised 

-- 

-- 

-- 

0 

-- 

-- 

-- 

14 

-- 

-- 

1,212 

-- 

Balance at December 31, 2015 

934 

123,595 

1,899 

Result for the year 

Other comprehensive income 

Recognition of share-based 
payments 

Share options exercised 

-- 

-- 

-- 

0 

-- 

-- 

-- 

2 

-- 

-- 

2,454 

-- 

-- 

-- 

1 

-- 

-- 

1 

-- 

(16) 

-- 

-- 

(15,798) 

109,404 

(20,832) 

(20,832) 

-- 

-- 

-- 

1 

1,212 

14 

(36,630) 

89,799 

(39,103) 

(39,103) 

-- 

-- 

-- 

(16) 

2,454 

2 

Balance at December 31, 2016 

934 

123,597 

4,353 

(15) 

(75,733) 

53,136 

The accompanying notes are an integral part of these financial statements. 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
 
 
 
 
 
 
 
 
PAGE 50 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

Consolidated statement of cash flows for the year ended December 31, 2016 

Cash flow from operating activities 

Result for the year 

Adjustments for: 

— Depreciation 

— Share-based compensation 

— Financial income and expense 

Changes in working capital 

Cash used in operations 

Corporate income tax paid 

Interest received/(paid) 

Note 

2016 

€ 1,000 

2015 

€ 1,000 

(39,119) 

(20,831) 

7, 8    

12 

18 

1,245 

2,454 

(470) 

1,433 

(34,457) 

-- 

236 

480 

1,212 

(6,171) 

637 

(24,673) 

-- 

441 

Net cash used in operating activities 

(34,221) 

(24,232) 

Cash flow from investing activities 

Purchases of intangible assets 

Purchases of property, plant and equipment 

-- 

(2,539) 

(28) 

(1,296) 

Net cash used in investing activities 

(2,539) 

(1,324) 

Cash flow from financing activities 

Proceeds from exercise of share options 

Proceeds from borrowings 

Redemption of financial lease 

Net cash generated by financing activities 

Net increase/(decrease) in cash and cash equivalents 

Currency effect cash and cash equivalents 

Cash and cash equivalents at the beginning of the year 

Cash and cash equivalents at the end of the year 

The accompanying notes are an integral part of these financial statements. 

13 

13 

11 

11 

2 

370 

(15) 

357 

14 

1,640 

(34) 

1,620 

(36,403) 

(23,936) 

738 

94,865 

6,065 

112,736 

59,200 

94,865 

 
 
  
 
 
 
 
 
 
 
 
  
 
 
 
 
  
 
 
 
 
 
  
 
 
 
 
 
  
 
 
 
 
  
 
 
 
 
 
 
 
 
  
 
 
 
 
  
 
 
 
 
 
 
 
  
 
 
 
 
  
 
 
 
 
  
 
 
 
 
  
 
 
 
 
 
PAGE 51 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

Notes to the consolidated financial statements for the year ended December 31, 2016 

1. General Information  

ProQR Therapeutics N.V., or “ProQR” or the “Company”, is a development stage company domiciled in the 

Netherlands that primarily focuses on the development and commercialization of novel therapeutic 

medicines.  

Since September 18, 2014, the Company’s ordinary shares are listed on the NASDAQ Global Market under 

ticker symbol PRQR. 

The Company was incorporated in the Netherlands, on February 21, 2012 (Chamber of Commerce no. 

54600790) and was reorganized from a private company with limited liability to a public company with limited 

liability on September 23, 2014. The Company has its statutory seat in Leiden, the Netherlands. The address 

of its headquarters and registered office is Zernikedreef 9, 2333 CK Leiden, the Netherlands. 

At December 31, 2016, ProQR Therapeutics N.V. is the ultimate parent company of the following entities: 

 

 

 

 

 

 

 

 

 

 

ProQR Therapeutics Holding B.V. (the Netherlands, 100%); 

ProQR Therapeutics I B.V. (the Netherlands, 100%); 

ProQR Therapeutics II B.V. (the Netherlands, 100%); 

ProQR Therapeutics III B.V. (the Netherlands, 100%); 

ProQR Therapeutics IV B.V. (the Netherlands, 100%); 

ProQR Therapeutics VI B.V. (the Netherlands, 100%); 

ProQR Therapeutics VII B.V. (the Netherlands, 100%); 

ProQR Therapeutics VIII B.V. (the Netherlands, 100%); 

ProQR Therapeutics IX B.V. (the Netherlands, 100%); 

ProQR Therapeutics I Inc. (United States, 100%). 

As used in these consolidated financial statements, unless the context indicates otherwise, all references to 

“ProQR”, the “Company” or the “Group” refer to ProQR Therapeutics N.V. including its subsidiaries. 

2. Basis of preparation 

(a) Statement of compliance  
These consolidated financial statements have been prepared in accordance with International Financial 

Reporting Standards, or IFRS, as adopted by the European Union (“EU”). 

With reference to the income statement of the Company, use has been made of the exemption pursuant to 

Section 402 of Book 2 of the Netherlands Civil Code. 

(b) Basis of measurement  
The financial statements have been prepared on the historical cost basis except for financial instruments and 

share-based payment obligations which have been based on fair value. Historical cost is generally based on 

the fair value of the consideration given in exchange for assets.  

(c) Functional and presentation currency 
These consolidated financial statements are presented in euro, which is the Company’s functional currency. 

All amounts have been rounded to the nearest thousand, unless otherwise indicated. 

 
 
 
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Financial Statements 2016 
ANNUAL REPORT 2016 

(d) Going Concern  

The management board of ProQR has, upon preparing and finalizing the 2016 financial statements, assessed 

the Company’s ability to fund its operations for a period of at least one year after the date of signing these 

financial statements.  

The management board of the Company is confident about the continuity of the Company based on its 

existing funding, taking into account the Company’s current cash position and the projected cash flows based 

on the activities under execution on the basis of ProQR’s business plan and budget. 

(e) Use of estimates and judgements  

In preparing these consolidated financial statements, management has made judgements, estimates and 

assumptions that affect the application of accounting policies and the reported amounts of assets, liabilities, 

income and expenses. Actual results may differ from these estimates.  

Estimates and underlying assumptions are reviewed on an ongoing basis. Revisions to accounting estimates 

are recognized in the period in which the estimate is revised if the revision affects only that period or in the 

period of the revision and future periods if the revision affects both current and future periods.  

Information about assumptions and estimation uncertainties that may have a significant risk of resulting in a 

material adjustment is included below. 

(i) Share-based payments  

Share options granted to employees and consultants are measured at the fair value of the equity instruments 

granted. Fair value is determined through the use of an option-pricing model considering, among others, the 

following variables:  

 

 

 

 

 

 

the exercise price of the option;  

the expected life of the option;  

the current value of the underlying shares;  

the expected volatility of the share price;  

the dividends expected on the shares; and  

the risk-free interest rate for the life of the option.  

For the Company’s share option plans, management’s judgment is that the Black-Scholes valuation method is 

the most appropriate for determining the fair value of the Company’s share options.  

Initially, the Company’s ordinary shares were not publicly traded and consequently the Company needed to 

estimate the fair value of its share and the expected volatility of that value. The expected volatility of all 

options granted was therefore based on the average historical volatility of the Company’s peers over a period 

that agrees with the expected option life. All assumptions and estimates are further discussed in Note 12(d) 

to the financial statements. The value of the underlying shares was determined on the basis of the prior sale 

of company stock method. As such, the Company has benchmarked the value per share to external 

transactions of Company shares and external financing rounds.  

For options granted from the moment of listing, the Company uses the closing price of the ordinary shares 

on the previous business day as exercise price of the options granted. 

The result of the share option valuations and the related compensation expense is dependent on the model 

and input parameters used. Even though Management considers the fair values reasonable and defensible 

 
 
PAGE 53 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

based on the methodologies applied and the information available, others might derive a different fair value 

for the Company’s share options. 

(ii) Corporate income taxes  

The Company recognizes deferred tax assets arising from unused tax losses or tax credits only to the extent 

that the Company has sufficient taxable temporary differences or there is convincing evidence that sufficient 

taxable profit will be available against which the unused tax losses or unused tax credits can be utilized. 

Management’s judgment is that such convincing evidence is currently not sufficiently available and a deferred 

tax asset is therefore only recognized to the extent that the Company has sufficient taxable temporary 

differences.  

(iii) Grant income  

Grants (to be) received are reflected in the balance sheet as other receivables or deferred income. At each 

balance sheet date, for grants approved, the Company estimates the associated costs incurred, the level of 

service performed and the progress of the associated projects. Based on this analysis grant income is 

recognized. 

(iv) Research and development expenditures  

Research expenditures are currently not capitalized but are reflected in the income statement because the 

criteria for capitalization are not met. At each balance sheet date, the Company estimates the level of service 

performed by the vendors and the associated costs incurred for the services performed.  

Although we do not expect the estimates to be materially different from amounts actually incurred, the 

understanding of the status and timing of services performed relative to the actual status and timing of 

services performed may vary and could result in reporting amounts that are too high or too low in any 

particular period.  

(f) Changes in accounting policies  

The financial statements have been prepared on the basis of International Financial Reporting Standards 

(“IFRS”), as issued by the International Accounting Standards Board (“IASB”). New Standards and 

Interpretations, which became effective as of January 1, 2016, did not have a material impact on our financial 

statements.  

3. Significant Accounting Policies  

The Company has consistently applied the following accounting policies to all periods presented in these 

consolidated financial statements. 

(a) Basis of consolidation 
(i) Subsidiaries 

Subsidiaries are entities controlled by the Group. The Group controls an entity when it is exposed to, or has 

rights to, variable returns from its involvement with the entity and has the ability to affect those returns 

through its power over the entity. The financial statements of subsidiaries are included in the consolidated 

financial statements from the date on which control commences until the date on which control ceases. 

(ii) Loss of control 

When the Group loses control over a subsidiary, it derecognises the assets and liabilities of the subsidiary, 

and any non-controlling interests and other components of equity. Any resulting gain or loss is recognised in 

profit or loss. Any interest retained in the former subsidiary is measured at fair value when control is lost.  

 
 
 
PAGE 54 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

(iii) Transactions eliminated on consolidation 

Intra-group balances and transactions, and any unrealised income and expenses arising from intra-group 

transactions, are eliminated. Unrealised gains arising from transactions with equity-accounted investees are 

eliminated against the investment to the extent of the Group’s interest in the investee. Unrealised losses are 

eliminated in the same way as unrealised gains, but only to the extent that there is no evidence of 

impairment. 

(b) Classes of financial instruments  
Financial instruments are both primary financial instruments, such as receivables and payables, and financial 

derivatives. For primary financial instruments, reference is made to the treatment per the corresponding 

balance sheet item.  

Financial derivatives are valued at fair value. Upon first recognition, financial derivatives are recognized at fair 

value and then revalued as at balance sheet date.  

(c) Foreign currencies  
(i) Foreign currency transactions 

Foreign currency transactions are translated into the functional currency using the exchange rates prevailing 

at the dates of the transactions.  

Monetary assets and liabilities denominated in foreign currencies are translated into the functional currency 

at the exchange rate at the reporting date. Non-monetary assets and liabilities denominated in foreign 

currencies that are measured at fair value are translated into the functional currency at the exchange rate 

when the fair value was determined. Foreign currency differences are generally recognized in profit or loss. 

Non-monetary items that are measured based on historical cost in a foreign currency are not translated. 

(ii) Foreign operations 

The assets and liabilities of foreign operations are translated into euro at exchange rates at the reporting 

date. The income and expenses of foreign operations are translated into euros at the exchange rates at the 

dates of the transactions. Foreign currency differences are recognized in OCI and accumulated in the 

translation reserve, except to the extent that the translation difference is allocated to NCI. 

(d) Recognition of other income  
Other income includes amounts earned from third parties and are recognized when earned in accordance 

with the substance and under the terms of the related agreements and when it is probable that the economic 

benefits associated with the transaction will flow to the entity and the amount of the income can be 

measured reliably. The grants are recognized in other income in the same period in which the related R&D 

costs are recognized. 

(e) Government grants—WBSO  

The WBSO (“afdrachtvermindering speur- en ontwikkelingswerk”) is a Dutch fiscal facility that provides 

subsidies to companies, knowledge centers and self-employed people who perform research and 

development activities (as defined in the WBSO Act). Under this Act, a contribution is paid towards the labor 

costs of employees directly involved in research and development. The contribution is in the form of a 

reduction of payroll taxes and social security contributions. Subsidies relating to labor costs are deferred and 

recognized in the income statement as negative labor costs over the period necessary to match them with 

the labor costs that they are intended to compensate.  

 
 
PAGE 55 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

(f) Employee benefits  
(i) Short-term employee benefits 

Short-term employee benefits are expensed as the related service is provided. A liability is recognized for the 

amount expected to be paid if the Group has a present legal or constructive obligation to pay this amount as 

a result of past service provided by the employee and the obligation can be estimated reliably. 

(ii) Share-based payment transactions  

The grant-date fair value of equity-settled share-based payment awards granted to employees is generally 

recognized as an expense, with a corresponding increase in equity, over the vesting period of the awards. The 

amount recognized as an expense is adjusted to reflect the number of awards for which the related service 

and non-market performance conditions are expected to be met, such that the amount ultimately recognized 

is based on the number of awards that meet the related service and non-market performance conditions at 

the vesting date. For share-based payment awards with non-vesting conditions, the grant-date fair value of 

the share-based payment is measured to reflect such conditions and there is no true-up for differences 

between expected and actual outcomes. 

(iii) Pension obligations  

The Company operates defined contribution pension plans for all employees funded through payments to 

insurance companies. The Company has no legal or constructive obligation to pay further contributions once 

the contributions have been paid. The contributions are recognized as employee benefit expense when they 

are due. Prepaid contributions are recognized as an asset to the extent that a cash refund or a reduction in 

the future payments is available.  

(g) Taxation  
Income tax expense represents the sum of the tax currently payable and deferred tax. It is recognized in 

profit or loss except to the extent that it relates to a business combination, or items recognized directly in 

equity or in OCI. 

(i) Current tax  

The tax currently payable is based on taxable profit for the year. Taxable profit differs from profit as reported 

in the income statement because of items of income or expense that are taxable or deductible in other years 

and items that are never taxable or deductible. The Company’s liability for current tax is calculated using tax 

rates that have been enacted or substantively enacted by the end of the reporting period. 

(ii) Deferred tax  

Deferred tax is recognized on differences between the carrying amounts of assets and liabilities in the 

financial statements and the corresponding tax bases used in the computation of taxable profit.  

The carrying amount of deferred tax assets is reviewed at the end of each reporting period and reduced to 

the extent that it is no longer probable that sufficient taxable profits will be available to allow all or part of the 

asset to be recovered. Since the Company does not expect to be profitable in the foreseeable future, its 

deferred tax assets are valued at nil.  

Deferred tax assets and liabilities are measured at the tax rates that are expected to apply in the period in 

which the liability is settled or the asset realized, based on tax rates (and tax laws) that have been enacted or 

substantively enacted by the end of the reporting period. The measurement of deferred tax liabilities and 

assets reflects the tax consequences that would follow from the manner in which the Company expects, at 

the end of the reporting period, to recover or settle the carrying amount of its assets and liabilities.  

 
 
 
PAGE 56 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

(h) Intangible assets  
(i) Licenses  

Acquired patents have a finite useful life and are carried at cost less accumulated amortization and 

impairment losses. Amortization is calculated using the straight-line method to allocate the cost of patents 

over their estimated useful lives (generally 10 years unless a patent expires prior to that date). Amortization 

begins when an asset is available for its intended use.  

(ii) Research and development  

Research expenditures are recognized as expenses as incurred. Costs incurred on development projects are 

recognized as intangible assets as of the date that it can be established that it is probable that future 

economic benefits that are attributable to the asset will flow to the Company considering its commercial and 

technological feasibility, generally when filed for regulatory approval for commercial production, and when 

costs can be measured reliably. Given the current stage of the development of the Company’s products no 

development expenditures have yet been capitalized.  

Registration costs for patents are part of the expenditures for the research and development project. 

Therefore, registration costs for patents are expensed as incurred as long as the research and development 

project concerned does not yet meet the criteria for capitalization.  

(iii) Other intangible assets 

Other intangible assets, including software, that are acquired by the Company and have finite useful lives are 

measured at cost less accumulated amortization and accumulated impairment losses. 

(iv) Amortization 

Amortization is calculated to write off the cost of intangible assets less their estimated residual values using 

the straight-line method over their estimated useful lives, and is recognized in profit or loss.  

The estimated useful lives for current and comparative periods are as follows: 

 

software:    

3 years. 

Amortization methods, useful lives and residual values are reviewed at each reporting date and adjusted if 

appropriate. 

(i) Property, plant and equipment 
(i) Recognition and measurement 

Items of property, plant and equipment are measured at cost less accumulated depreciation and any 

accumulated impairment losses. If significant parts of an item of property, plant and equipment have 

different useful lives, then they are accounted for as separate items (major components) of property, plant 

and equipment. Any gain or loss on disposal of an item of property, plant and equipment is recognized in 

profit or loss. 

(ii) Depreciation 

Depreciation is calculated to write off the cost of items of property, plant and equipment less their estimated 

residual values using the straight-line method over their estimated useful lives, and is recognized in profit or 

loss. Leased assets are depreciated over the shorter of the lease term and their useful lives unless it is 

reasonably certain that the Company will obtain ownership by the end of the lease term.  

 
 
 
 
 
 
 
PAGE 57 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

The estimated useful lives of property, plant and equipment for current and comparative periods are as 

follows: 

 

 

 

leasehold improvements:    

5 - 10 years. 

laboratory equipment:  

5 years. 

other:  

3 - 5 years. 

Depreciation methods, useful lives and residual values are reviewed at each reporting date and adjusted if 

appropriate. 

(j) Impairment of tangible and intangible assets  
At the end of each reporting period, the Company reviews the carrying amounts of its tangible and intangible 

assets to determine whether there is any indication that those assets have suffered an impairment loss. If 

any such indication exists, the recoverable amount of the asset is estimated in order to determine the extent 

of the impairment loss (if any). Where it is not possible to estimate the recoverable amount of an individual 

asset, the Company estimates the recoverable amount of the cash-generating unit to which the asset 

belongs. Where a reasonable and consistent basis of allocation can be identified, corporate assets are also 

allocated to individual cash-generating units, or otherwise they are allocated to the smallest group of cash-

generating units for which a reasonable and consistent allocation basis can be identified.  

Intangible assets with indefinite useful lives and intangible assets not yet available for use are tested for 

impairment at least annually, and whenever there is an indication that the asset may be impaired.  

The recoverable amount is the higher of fair value less costs of disposal and value in use. In assessing value in 

use, the estimated future cash flows are discounted to their present value using a pre-tax discount rate that 

reflects current market assessments of the time value of money and the risks specific to the asset for which 

the estimates of future cash flows have not been adjusted.  

If the recoverable amount of an asset (or cash-generating unit) is estimated to be less than its carrying 

amount, the carrying amount of the asset (or cash-generating unit) is reduced to its recoverable amount. An 

impairment loss is recognized immediately in profit or loss, unless the relevant asset is carried at a revalued 

amount, in which case the impairment loss is treated as a revaluation decrease.  

Where an impairment loss subsequently reverses, the carrying amount of the asset (or cash-generating unit) 

is increased to the revised estimate of its recoverable amount, but so that the increased carrying amount 

does not exceed the carrying amount that would have been determined had no impairment loss been 

recognized for the asset (or cash-generating unit) in prior years. A reversal of an impairment loss is 

recognized immediately in profit or loss, unless the relevant asset is carried at a revalued amount, in which 

case the reversal of the impairment loss is treated as a revaluation increase.  

(k) Financial assets  
All financial assets are recognized and derecognized on the trade date where the purchase or sale of a 

financial asset is under a contract whose terms require delivery of the financial asset within the timeframe 

established by the market concerned, and are initially measured at fair value, plus transaction costs, except 

for those financial assets classified as at fair value through profit or loss, which are initially measured at fair 

value.  

 
 
 
 
 
 
 
 
PAGE 58 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

(i) Loans and receivables  

Trade receivables, loans and other receivables that have fixed or determinable payments that are not quoted 

in an active market are classified as “loans and receivables”. Loans and receivables are measured at 

amortized cost using the effective interest method, less any impairment.  

An allowance for doubtful accounts is established when there is objective evidence that the Company will not 

be able to collect all amounts due according to the original terms of receivables. Significant financial 

difficulties of the debtor, probability that the debtor will enter into bankruptcy or financial reorganization, and 

default or delinquency in payments are considered indicators that the trade receivable is impaired. Loans 

and receivables are included in ‘current assets’, except for maturities greater than 12 months after the 

balance sheet date, which are classified as ‘non-current assets’.  

For all financial assets, the fair value approximates its carrying value.  

(l) Cash and cash equivalents  
Cash and cash equivalents include cash on hand and all highly liquid investments with original maturities of 

three months or less that are convertible to a known amount of cash and bear an insignificant risk of change 

in value.  

(m) Financial liabilities and equity instruments  
Debt and equity instruments are classified as either financial liabilities or as equity in accordance with the 

substance of the contractual arrangement.  

(i) Equity instruments  

An equity instrument is any contract that evidences a residual interest in the assets of an entity after 

deducting all of its liabilities. Equity instruments issued by the Company are recognized at the proceeds 

received, net of direct issue costs.  

(ii) Other financial liabilities  

Other financial liabilities, including borrowings, are initially measured at fair value, net of transaction costs 

incurred, and are subsequently measured at amortized cost using the effective interest method, with interest 

expense recognized on an effective yield basis.  

The effective interest method is a method of calculating the amortized cost of a financial liability and of 

allocating interest expense over the relevant period. The effective interest rate is the rate that exactly 

discounts estimated future cash payments through the expected life of the financial liability, or, where 

appropriate, a shorter period.  

Borrowings and other financial liabilities are classified as ‘non-current liabilities,’ other than liabilities with 

maturities up to one year, which are classified as “current liabilities”.  

The Company derecognizes financial liabilities when the liability is discharged, cancelled or expired. For all 

financial liabilities, the fair value approximates its carrying amount.  

(n) Leases 
(i) Determining whether an arrangement contains a lease 

At inception of an arrangement, the Company determines whether such an arrangement is or contains a 

lease. 

 
 
PAGE 59 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

At inception or on reassessment of an arrangement that contains a lease, the Company separates payments 

and other consideration required by such an arrangement into those for the lease and those for other 

elements on the basis of their relative fair values. If the Company concludes for a finance lease that it is 

impracticable to separate the payments reliably, then an asset and a liability are recognized at an amount 

equal to the fair value of the underlying asset. Subsequently, the liability is reduced as payments are made 

and an imputed finance cost on the liability is recognized using the Company’s incremental borrowing rate. 

(ii) Leased assets 

Assets held by the Company under leases that transfer to the Company substantially all of the risks and 

rewards of ownership are classified as finance leases. The leased assets are measured initially at an amount 

equal to the lower of their fair value and the present value of the minimum lease payments. Subsequent to 

initial recognition, the assets are accounted for in accordance with the accounting policy applicable to that 

asset.  

Assets held under other leases are classified as operating leases and are not recognized in the Company’s 

statement of financial position. 

(iii) Lease payments 

Payments made under operating leases are recognized in profit or loss on a straight-line basis over the term 

of the lease. Lease incentives received are recognized as an integral part of the total lease expense, over the 

term of the lease. 

Minimum lease payments made under finance leases are apportioned between the finance expense and the 

reduction of the outstanding liability. The finance expense is allocated to each period during the lease term 

so as to produce a constant periodic rate of interest on the remaining balance of the liability. 

4. New standards and interpretations not yet adopted 

A number of new standards, amendments to standards and interpretations are effective for annual periods 

beginning after 1 January 2017, and have not been applied in preparing these consolidated financial 

statements. Those which may be relevant to the Group are set out below. The Group does not plan to adopt 

these standards early. 

IFRS 9 Financial Instruments 

IFRS 9, published in July 2014, replaces the existing guidance in IAS 39 Financial Instruments: Recognition and 

Measurement. IFRS 9 includes revised guidance on the classification and measurement of financial 

instruments, including a new expected credit loss model for calculating impairment on financial assets, and 

the new general hedge accounting requirements. It also carries forward the guidance on recognition and 

derecognition of financial instruments from IAS 39. 

IFRS 9 is effective for annual reporting periods beginning on or after 1 January 2018, with early adoption 

permitted. 

IFRS 15 Revenue from Contracts with Customers 

IFRS 15 establishes a comprehensive framework for determining whether, how much and when revenue is 

recognised. It replaces existing revenue recognition guidance, including IAS 18 Revenue, IAS 11 Construction 

Contracts and IFRIC 13 Customer Loyalty Programmes. 

IFRS 15 is effective for annual reporting periods beginning on or after 1 January 2018, with early adoption 

permitted. 

 
 
 
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Financial Statements 2016 
ANNUAL REPORT 2016 

IFRS 16 Leases 

IFRS 16 specifies how a company will recognise, measure, present and disclose leases. The standard provides 

a single lessee accounting model, requiring lessees to recognise assets and liabilities for all leases unless the 

lease term is 12 months or less or the underlying asset has a low value. Lessors continue to classify leases as 

operating or finance, with IFRS 16’s approach to lessor accounting substantially unchanged from its 

predecessor, IAS 17. 

IFRS 16 is effective for annual reporting periods beginning on or after 1 January 2019, with early adoption 

permitted. 

The adoption of these Standards and Interpretations are not expected to have a material effect on the 

financial statements.  

5. Financial Risk Management  

5.1. Financial risk factors 
The Company’s activities expose it to a variety of financial risks: market risk (including currency risk, interest 

rate risk and price risk), credit risk and liquidity risk. The Company’s overall financial risk management seeks 

to minimize potential adverse effects resulting from unpredictability of financial markets on the Company’s 

financial performance.  

Financial risk management is carried out by the finance department. The finance department identifies and 

evaluates financial risks and proposes mitigating actions if deemed appropriate.  

(a) Market risk  

Market risk is the risk that changes in market prices – such as foreign exchange rates, interest rates and 

equity prices – will affect the Company’s income or the value of its holdings of financial instruments. The 

objective of market risk management is to manage and control market risk exposures within acceptable 

parameters, while optimizing the return. 

Foreign exchange risk 

Foreign exchange risk arises from future commercial transactions and recognized assets and liabilities in 

foreign currencies, primarily with respect to the U.S. Dollar. The Company has an exposure associated with 

the time delay between entering into a contract, budget or forecast and the realization thereof. The Company 

operates a foreign exchange policy to manage the foreign exchange risk against the functional currency 

based on the Company’s cash balances and the projected future spend per major currency. 

At December 31, 2016 there was a net liability in U.S. Dollars of € 2.4 million (2015: € 1.1 million). Foreign 

currency denominated receivables and trade payables are short term in nature (generally 30 to 45 days). As a 

result foreign exchange rate movements on receivables and trade payables, during the years presented had 

an immaterial effect on the financial statements.  

At year-end, a substantial amount of our cash balances are denominated in U.S. Dollars. This amount reflects 

our current expectation of future expenditure in U.S. dollars. 

A reasonably possible strengthening (weakening) of the U.S. Dollar by 10% against all other currencies at 

December 31, 2016 would have affected the measurement of our cash balances denominated in a U.S. Dollar 

and affected equity and profit or loss by € 2.5 million (2015: € 5.2 million). The analysis assumes that all other 

variables, in particular interest rates, remain constant. 

 
 
PAGE 61 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

Price risk 

The market prices for the production of preclinical and clinical materials and services as well as external 

contracted research may vary over time. Currently, the commercial prices of any of the Company’s product 

candidates is uncertain. When the development products near the regulatory approval date or potential 

regulatory approval date, the uncertainty of the potential sales price decreases. The Company is not exposed 

to commodity price risk.  

Furthermore the Company does not hold investments classified as available-for-sale or at fair value through 

profit or loss, therefore are not exposed to equity securities price risk.  

Cash flow and fair value Interest rate risk  

The Company’s exposure to interest rate risks is limited due to the use of loans with fixed rates. The 

Company has one loan with a fixed interest, amounting to € 5,697,000 at December 31, 2016 (2015: € 

4,824,000). Details on the interest rates and maturities of these loans are provided in Note 13.  

(b) Credit risk  

Credit risk represents the risk of financial loss caused by default of the counterparty. The Company has no 

large receivables balances with external parties. The Company’s principal financial assets are cash and cash 

equivalents which are placed at ABN Amro, Rabobank and Wells Fargo. Our cash management policy is 

focused on preserving capital, providing liquidity for operations and optimizing yield while accepting limited 

risk (Short-term credit ratings must be rated  A-1/P-1/F1 at a minimum by at least one of the Nationally 

Recognized Statistical Rating Organizations (NRSROs) specifically Moody’s, Standard & Poor’s or Fitch. Long-

term credit rating must be rated A- or A3 at a minimum by at least one NRSRO). 

At December 31, 2016 and December 31, 2015, substantially all of our cash and cash equivalents were placed 

at two large institutions, Rabobank and ABN Amro. In 2016, this also included Wells Fargo. All institutions are 

highly rated (ratings of Aa2, A1 and A2 for Rabobank, ABN Amro and Wells Fargo respectively) with sufficient 

capital adequacy and liquidity metrics. 

There are no financial assets past due date or impaired. No credit limits were exceeded during the reporting 

period. 

(c) Liquidity risk  

Liquidity risk represents the risk that an entity will encounter difficulty in meeting obligations associated with 

its financial liabilities. Prudent liquidity risk management implies ensuring sufficient availability of cash 

resources for funding of operations and planning to raise cash if and when needed, either through issue of 

shares or through credit facilities. Management monitors rolling forecasts of the Company’s liquidity reserve 

on the basis of expected cash flow.  

The table below analyzes ProQR’s undiscounted liabilities into relevant maturity groupings based on the 

remaining period at year-end until the contractual maturity date:  

dsssds  

At December 31, 2016 

Borrowings  

Trade payables and other payables   

Less than  
1 year 

Between  
1 and 2 years 

Between  
2 and 5 years 

€ 1,000 

€ 1,000 

€ 1,000 

-- 

6,710 

6,710 

1,839 

-- 

1,839 

4,860 

-- 

4,860 

Over 
5 years 

€ 1,000 

-- 

-- 

-- 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
PAGE 62 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

At December 31, 2015 

Borrowings  

Finance lease liabilities   

Trade payables and other payables   

Less than  
1 year 

Between  
1 and 2 years 

Between  
2 and 5 years 

€ 1,000 

€ 1,000 

€ 1,000 

-- 

15 

5,471 

5,486 

1,691 

4,712 

-- 

-- 

-- 

-- 

1,691 

4,712 

Over 
5 years 

€ 1,000 

-- 

-- 

-- 

-- 

5.2. Capital risk management 
The Company's objectives when managing capital are to safeguard the Company's ability to continue as a 

going concern in order to provide returns for shareholders, benefits for other stakeholders and to maintain 

an optimal capital structure to reduce the cost of capital. 

In order to maintain or adjust the capital structure, the Company may adjust the amount of dividends paid to 

shareholders (although at this time the Company does not have retained earnings and is therefore currently 

unable to pay dividends), return capital to shareholders, issue new shares or sell assets to reduce debt. 

The total amount of equity as recorded on the balance sheet is managed as capital by the Company. 

5.3. Fair value measurement 
For financial instruments that are measured on the balance sheet at fair value, IFRS 13 requires disclosure of 

fair value measurements by level of the following fair value measurement hierarchy: 

 

 

 

quoted prices (unadjusted) in active markets for identical assets or liabilities (level 1); 

inputs other than quoted prices included within level 1 that are observable for the asset or liability, 

either directly (that is, as prices) or indirectly (that is, derived from prices) (level 2); and 

inputs for the asset or liability that are not based on observable market data (that is, unobservable 

inputs) (level 3). 

The Company has no assets and liabilities that are measured at fair value at December 31, 2016 and 2015. 

The carrying amount of all financial assets and financial liabilities is a reasonable approximation of the fair 

value and therefore information about the fair values of each class has not been disclosed.  

6. Segment Information  

The Company operates in one reportable segment, which comprises the discovery and development of 

innovative, RNA based therapeutics. The management board is identified as the chief operating decision 

maker. The management board reviews the operating results regularly to make decisions about resources 

and to assess overall performance.  

The Company has not generated any sales revenues since inception.  

All non-current assets of the Company are located in the Netherlands. The amounts provided to the 

management board with respect to total assets and liabilities are measured in a manner consistent with that 

of the financial statements. 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
7. Intangible Assets  

dsssds  

Balance at January 1, 2015 

Cost   

Accumulated amortization 

Carrying amount 

Additions 

Amortization 

Movement for the period 

Balance at December 31, 2015 

Cost   

Accumulated amortization 

Carrying amount 

Additions 

Amortization 

Movement for the period 

Balance at December 31, 2016 

Cost   

Accumulated amortization 

Carrying amount 

PAGE 63 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

Licenses 

Software 

€ 1,000 

€ 1,000 

Total 

€ 1,000 

39 

-- 

39 

-- 

-- 

-- 

39 

-- 

39 

-- 

-- 

-- 

39 

-- 

39 

124 

-- 

124 

28 

(50) 

(22) 

152 

(50) 

102 

-- 

(51) 

(51) 

152 

(101) 

51 

163 

-- 

163 

28 

(50) 

(22) 

191 

(50) 

141 

-- 

(51) 

(51) 

191 

(101) 

90 

In 2012, the Company acquired an exclusive license from the Massachusetts General Hospital. The initial 

payment in respect of the license, in the amount of € 39,000, will be amortized over the commercial life of 

products based on the license during the patent-life.  

The amortization charge for 2016 is included in the general and administrative costs for an amount of  

€ 51,000 (2015: € 50,000).  

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
PAGE 64 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

8. Property, Plant and Equipment (‘PP&E’) 

dsssds  

Balance at January 1, 2015 

Cost   

Accumulated depreciation 

Carrying amount 

Additions 

Depreciation 

Disposals 

Movement for the period 

Balance at December 31, 2015 

Cost   

Accumulated depreciation 

Carrying amount 

Additions 

Depreciation 

Transfer 

Disposals 

Movement for the period 

Balance at December 31, 2016 

Cost   

Accumulated depreciation 

Carrying amount 

Leasehold 
 improvements 

Laboratory 
 equipment 

Other 

Total 

€ 1,000 

€ 1,000 

€ 1,000 

€ 1,000 

326 

(17) 

309 

659 

(77) 

-- 

582 

985 

(94) 

891 

1,166 

(499) 

(196) 

(23) 

448 

1,847 

(508) 

1,339 

769 

(104) 

665 

367 

(201) 

-- 

166 

1,136 

(305) 

831 

806 

(340) 

-- 

-- 

466 

1,957 

(660) 

1,297 

242 

(29) 

213 

415 

(145) 

(6) 

264 

651 

(174) 

477 

461 

(332) 

196 

-- 

325 

1,283 

(481) 

802 

1,337 

(150) 

1,187 

1,441 

(423) 

(6) 

1,012 

2,772 

(573) 

2,199 

2,433 

(1,171) 

-- 

(23) 

1,239 

5,087 

(1,649) 

3,438 

The depreciation charge for 2016 is included in the research and development costs for an amount of  

€ 907,000 (2015: € 361,000) and in the general and administrative costs for an amount of € 264,000  

(2015: € 62,000).  

9. Social Security and Other Taxes 

dsssds  

Value added tax 

Wage tax 

December 31, 
2016 

December 31, 
2015 

€ 1,000 

€ 1,000 

395 

-- 

395 

953 

3 

956 

All receivables are considered short-term and due within one year.  

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
PAGE 65 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

10. Prepayments and Other Receivables  

dsssds  

Prepayments 

Other receivables 

All receivables are considered short-term and due within one year.  

11. Cash and Cash Equivalents  

dsssds  

Cash at banks 

Bank deposits 

December 31, 
2016 

December 31, 
2015 

€ 1,000 

€ 1,000 

1,250 

1,170 

2,420 

1,401 

547 

1,948 

December 31, 
2016 

December 31, 
2015 

€ 1,000 

€ 1,000 

56,354 

2,846 

59,200 

94,865 

-- 

94,865 

The cash at banks is at full disposal of the Company. Bank deposits are convertible into cash upon request of 

the Company. 

12. Shareholders’ Equity  

(a) Share capital  

dsssds  

In issue at January 1 

Issued for cash 

Exercise of share options 

In issue at December 31 – fully paid 

Number of ordinary shares 

2016 

2015 

23,345,965 

23,338,154 

-- 

891 

-- 

7,811 

23,346,856 

23,345,965 

The authorized share capital of the Company amounting to € 3,000,000 consists of 37,500,000 ordinary 

shares and 37,500,000 preference shares with a par value of € 0.04 per share. At December 31, 2016, 

24,520,814 ordinary shares were issued and fully paid in cash, of which 1,173,958 were held by the Company 

as treasury shares (2015: 1,174,849). 

On October 2, 2015, the Company filed a shelf registration statement, which permitted: (a) the offering, 

issuance and sale by the Company of up to a maximum aggregate offering price of $ 200,000,000 of its 

ordinary shares, warrants and/or units; and (b) as part of the $ 200,000,000, the offering, issuance and sale by 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
PAGE 66 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

us of up to a maximum aggregate offering price of $ 60,000,000 of its ordinary shares that may be issued and 

sold under a sales agreement with Cantor Fitzgerald & Co in one or more at-the-market offerings. At 

December 31, 2016, no shares had been sold pursuant to its current at-the-market offering program. 

(b) Equity settled employee benefit reserve  
The costs of share options for employees, members of the supervisory board and members of the 

management board are recognized in the income statement, together with a corresponding increase in 

equity during the vesting period, taking into account (deferral of) corporate income taxes. The accumulated 

expense of share options recognized in the income statement is shown separately in the equity category 

‘equity settled employee benefit reserve’ in the ‘statement of changes in equity’.  

(c) Translation reserve 
The translation reserve comprises all foreign currency differences arising from the translation of the financial 

statements of foreign operations.  

(d) Share options  

The Company operates an equity-settled share-based compensation plan which was introduced in 2013. 

Options may be granted to employees, members of the supervisory board, members of the management 

board and consultants. The compensation expenses included in operating costs for this plan were  

€ 2,454,000 in 2016 (2015: € 1,212,000), of which € 1,480,000 (2015: € 801,000) was recorded in general and 

administrative costs and € 974,000 (2015: € 411,000) was recorded in research and development costs.  

Options granted under this stock option plan are exercisable once vested. Any vesting schedule may be 

attached to the granted options, however the typical vesting period is four years (25% after every year). The 

options expire ten years after date of grant. Options granted under the stock option plan are granted at 

exercise prices which equal the fair value of the ordinary shares of the Company at the date of the grant.  

The Company accounts for its employee stock options under the fair value method. The fair value of the 

options is estimated at the date of grant using the Black-Scholes option-pricing model, with on average the 

following assumptions:  

dsssds  

Risk-free interest rate 

Expected dividend yield 

Expected volatility 

Expected life in years 

Options  
granted in 2016 

Options  
granted in 2015 

1.467% 

0% 

86.3% 

5 years 

1.497% 

0% 

86.8% 

5 years 

The resulting weighted average grant date fair value of the options amounted to € 3.72 in 2016 (2015:  

€ 10.35). The stock options granted have a 10 year life following the grant date and are assumed to be 

exercised five years from date of grant for all awards. 

 
 
 
 
 
 
 
 
 
 
 
Movements in the number of options outstanding and their related weighted average exercise prices are as 

PAGE 67 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

follows:  

dsssds  

Balance at January 1 

Granted 

Forfeited 

Exercised 

Lapsed 

2016 

2015 

Number of 
 options 

Average 
 exercise price 

Number of 
 options 

Average  
exercise price 

1,108,935 

1,214,126 

(116,181) 

(891) 

-- 

€ 4.19 

€ 5.49 

€ 4.64 

€ 2.38 

-- 

998,765 

125,798 

(7,817) 

(7,811) 

-- 

€ 2.78 

€ 15.27 

€ 4.64 

€ 1.78 

-- 

€ 4.19 

Balance at December 31 

2,205,989 

€ 4.88 

1,108,935 

Exercisable 

615,246 

339,352 

The options outstanding at December 31, 2016 had an exercise price in the range of € 1.11 to € 20.34  

(2015: € 1.11 to € 20.34) and a weighted-average contractual life of 8.3 years (2015: 8.3 years). 

The weighted-average share price at the date of exercise for share options exercised in 2016 was € 4.23 

(2015: € 19.30). 

Please refer to Note 23 for the options granted to key management personnel. 

13. Non-current liabilities  

(a) Borrowings  

dsssds  

Innovation credit 

Accrued interest on innovation credit 

December 31, 
2016 

December 31, 
2015 

€ 1,000 

€ 1,000 

4,598 

1,099 

5,697 

4,228 

596 

4,824 

Innovation credit (“Innovatiekrediet”)  

On June 1, 2012, ProQR was awarded an Innovation credit by the Dutch government, through its agency RVO 

of the Ministry of Economic Affairs, for the Company’s cystic fibrosis program. Amounts were drawn under 

this facility in the course of the years 2013 through 2016. The credit covers 35% of the costs incurred in 

respect of the program up to an initial maximum of € 5.0 million.  

The credit is interest-bearing at a rate of 10% per annum. The credit, including accrued interest, is repayable 

in three installments on November 30, 2018, November 30, 2019 and November 30, 2020, depending on the 

technical success of the program.  

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
PAGE 68 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

The assets which are co-financed with the granted innovation credit are subject to a right of pledge for the 

benefit of RVO. 

(b) Finance lease liabilities  

dsssds  

Balance at January 1 

Initial recognition new finance leases  

Interest expense accrued 

Payment of finance lease liabilities 

Balance at December 31 

Current portion at December 31 

2016 

€ 1,000 

2015 

€ 1,000 

15 

-- 

-- 

(15) 

-- 

-- 

-- 

49 

-- 

-- 

(34) 

15 

15 

-- 

Certain of the Company’s property, plant and equipment items are subject to finance leases. These leases 

relate to laboratory equipment. The net carrying amount of leased assets amounts to nil in 2016 (2015: € 

48,000). 

Future minimum lease payments under finance leases as at December 31 are as follows:  

dsssds  

2016 

2015 

Minimum 
payments 

Present value  
of payments 

Minimum 
payments 

Present value  
of payments 

Less than 1 year 

Between 1 and 5 years 

More than 5 years 

-- 

-- 

-- 

-- 

-- 

-- 

15 

-- 

-- 

15 

-- 

-- 

The interest used for the present value of payments is 2%.  

14. Current Liabilities  

dsssds  

Current portion finance lease liabilities 

Trade payables 

Social securities and other taxes 

Pension premiums 

Deferred income 

Accrued expenses and other liabilities 

December 31, 
2016 

December 31, 
2015 

€ 1,000 

€ 1,000 

-- 

328 

312 

13 

-- 

6,057 

6,710 

15 

885 

235 

16 

144 

4,191 

5,486 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
PAGE 69 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

At December 31, 2015, current liabilities included deferred income resulting from receipt of the first 

installment of the € 6 million grant from the European Commission (EC) under the Horizon 2020 program to 

finance the clinical development of QR-010. 

15. Other income  

dsssds  

Grant income 

Rental income from property subleases 

2016 

€ 1,000 

1,632 

196 

1,828 

2015 

€ 1,000 

3,188 

47 

3,235 

In August 2014, the Company entered into an agreement with Cystic Fibrosis Foundation Therapeutics, Inc., 

or CFFT, a subsidiary of the Cystic Fibrosis Foundation, pursuant to which CFFT agreed to provide the 

Company with up to $ 3 million to support the clinical development of QR-010. The grant is recognized in 

other income in the same period in which the related R&D costs are recognized. 

In 2015, the European Commission (EC) through its Horizon 2020 program awarded ProQR and its academic 

partners a grant of € 6 million (ProQR: € 4.4 million) to support the clinical development of QR-010 through 

December 31, 2017. Horizon 2020 is one of the largest research and innovation programs in the European 

Union with nearly € 80 billion in available funding for qualified projects from 2014 to 2020.  

Both grants are recognized in other income in the same period in which the related R&D costs are 

recognized. 

16. Research and Development Costs  

Research and development costs amounted to € 31,923,000 in 2016 (2015: € 23,401,000) and comprise 

allocated employee costs, the costs of materials and laboratory consumables, the costs of external studies 

including, amongst others, clinical studies and toxicology studies and external research, license- and IP-costs 

and allocated other costs.  

17. Employee Benefits  

dsssds  

Wages and salaries 

Social security costs 

Pension costs – defined contribution plans 

Equity-settled share based payments 

2016 

€ 1,000 

10,184 

1,093 

764 

2,454 

14,495 

2015 

€ 1,000 

7,128 

596 

478 

1,212 

9,414 

Average number of employees for the period 

133.4 

86.1 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
PAGE 70 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

Employees per activity at December 31 (converted to FTE): 

dsssds  

Research and Development 

General and Administrative 

December 31, 
2016 

December 31, 
2015 

100.4 

32.9 

133.3 

72.4 

27.1 

99.5 

Of all employees 128.3 FTE are employed in the Netherlands (2015: 94.5 FTE). 

Included in the wages and salaries for 2016 is a credit of € 807,000 (2015: € 372,000) with respect to WBSO 

subsidies.  

18. Financial Income and Expense  

dsssds  

Interest income 

Current accounts and deposits 

Interest costs 

Interest on loans and borrowings 

Foreign exchange result 

Net foreign exchange benefit/(loss) 

19. Income Taxes  

The calculation of the tax charge is as follows:  

dsssds  

2016 

€ 1,000 

2015 

€ 1,000 

270 

501 

(538) 

(395) 

738 

470 

6,065 

6,171 

2015 

€ 1,000 

2015 

€ 1,000 

Income tax provision based on domestic rate (25%) 

9,776 

5,208 

Tax effect of: 

Non-deductible expenses 

Tax incentives 

Current year losses for which no deferred tax asset was recognized 

Change in unrecognized deductible temporary differences 

Income tax charge 

Effective tax rate 

(622) 

(46) 

(9,045) 

(63) 

-- 

0% 

(309) 

136 

(5,035) 

-- 

-- 

0% 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
PAGE 71 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

Due to the operating losses incurred since inception the Company has no tax provisions as of the balance 

sheet date. Furthermore, no significant temporary differences exist between accounting and tax results.  

Realization of deferred tax assets is dependent on future earnings, if any, the timing and amount of which are 

uncertain. Accordingly, the Company has not yet recognized any deferred tax asset related to operating 

losses. As per December 31, 2016, the Company has a total amount of € 82.9 million (2015: € 46.9 million) tax 

loss carry-forwards available for offset against future taxable profits. According to current tax regulations the 

first amount of the tax loss carry-forwards will expire in 2021.  

20. Earnings Per Share  

(a) Basic and diluted earnings per share  

Basic earnings per share are calculated by dividing the result attributable to equity holders of the Company 

by the weighted average number of shares outstanding during the year.  

dsssds  

Result attributable to equity holders of the Company (€ 1,000) 

Weighted average number of shares  

Basic (and diluted) earnings per share (€ per share) 

2016 

2015 

(39,103) 

(20,832) 

23,346,507 

23,343,262 

€ (1.68) 

€ (0.89) 

(b) Diluted earnings per share  
For the periods included in these financial statements, the share options are not included in the diluted 

earnings per share calculation as the Company was loss-making in all periods. Due to the anti-dilutive nature 

of the outstanding options, basic and diluted earnings per share are equal.  

(c) Dividends per share  

The Company did not declare dividends for any of the years presented in these financial statements.  

21. Operational Leases  

Since 2012, the Company is domiciled in Leiden, the Netherlands where it currently has concluded rental 

agreements for laboratory space and offices. In addition, the Company has one office in the US.  

The lease expenditure charged to the income statement in 2016 amounts to € 1,849,000 (2015: € 703,000). 

The future aggregate minimum lease payments under non-cancellable operating leases are as follows: 

dsssds  

Less than 1 year 

Between 1 and 5 years 

More than 5 years 

December 31, 
2016 

December 31, 
2015 

€ 1,000 

€ 1,000 

1,775 

5,508 

-- 

7,283 

1,938 

7,212 

-- 

9,150 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
PAGE 72 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

The Company leases out a part of its office in the US. In 2016, total sublease income amounted to € 196,000 

(2015: € 47,000), which is recorded in other income. At 31 December, the future minimum lease payments 

under non-cancellable leases are receivable as follows: 

dsssds  

Less than 1 year 

Between 1 and 5 years 

More than 5 years 

December 31, 
2016 

December 31, 
2015 

€ 1,000 

463 

-- 

-- 

463 

€ 1,000 

185 

-- 

-- 

185 

22. Commitments and Contingencies  

(a) Claims  
There are no claims known to management related to the activities of the Company.  

(b) Patent license agreement  
The Company and the General Hospital Corporation (MGH) have entered into a Patent License Agreement 

pursuant to which the Company may have certain royalty obligations. The Company is also obligated to pay 

MGH up to $ 700,000 in milestone payments upon the achievement of certain development and regulatory 

milestones and, beginning after its first commercial sale of a product covered by the licensed patent rights, a 

$ 10,000 annual license fee which is creditable against royalties due to MGH in the same calendar year. In 

addition, the Company is obligated to pay MGH 2% of any net sales by the Company, its affiliates or 

sublicensees on licensed products made or sold in the United States, as well as a low double-digit percentage 

of any payments the Company may receive from any sublicensee anywhere in the world.  

The Company has entered into various other Patent License Agreements, including those with Radboud 

University Medical Center, Leiden University Medical Centre and PARI Pharma GmbH, under which the 

Company is granted world-wide exclusive licenses pursuant to which the Company may have certain royalty 

obligations in relation to its product candidates. Pursuant to the terms of these agreements, the Company 

has made upfront payments, is obligated to make milestone payments and has to make sales-based royalty 

payments after market authorization. In specific cases, the Company has the option to make a one-time 

payment to buy of royalty obligations or in case the Company terminates an agreement before or after 

regulatory approval of the product. The Company may terminate an agreement for any reason. 

(c) Clinical support agreement 
In August 2014, the Company entered into an agreement with Cystic Fibrosis Foundation Therapeutics, Inc., 

or CFFT, a subsidiary of the Cystic Fibrosis Foundation, pursuant to which CFFT agreed to provide the 

Company with up to $ 3 million to support the clinical development of QR-010.  

Pursuant to the terms of the agreement, the Company is obligated to make a one-time milestone payment to 

CFFT of up to approximately $ 80 million, payable in three equal annual installments following the first 

commercial sale of QR-010, the first of which is due within 90 days following the first commercial sale. The 

Company is also obligated to make a one-time milestone payment to CFFT of up to $ 3 million if net sales of 

QR-010 exceed $ 500 million in a calendar year. Lastly, the Company is obligated to make a payment to CFFT 

of up to approximately $ 6 million if it transfers, sells or licenses QR-010 other than for certain clinical or 

development purposes, or if the Company enters into a change of control transaction. Either CFFT or the 

 
 
 
 
 
 
 
 
PAGE 73 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

Company may terminate the agreement for cause, which includes the Company’s material failure to achieve 

certain commercialization and development milestones. The Company’s payment obligations survive the 

termination of the agreement.  

(d) Research and development commitments  
The Company has research and development commitments, mainly with CRO's, amounting to € 8,856,000 at 

December 31, 2016 (2015: € 9,481,000). Of these obligations an amount of € 6,258,000 is due in 2017, the 

remainder is due in 1 to 5 years. 

23. Related-Party Transactions  

Details of transactions between the Company and related parties are disclosed below.  

(a) Compensation of the Supervisory Board  
On June 21, 2016, Mr. James Shannon was appointed to our supervisory board. The remuneration of the 

supervisory board members in 2016 is set out in the table below:  

dsssds  

2016 

Mr. Dinko Valerio   

Mr. Henri Termeer 

Mr. Antoine Papiernik 

Ms. Alison Lawton  

Mr. Paul Baart 

Mr. James Shannon 

Short term 
employee 
benefits 

Post 
employment 
benefits 

Share-based 
payment 

Total 

€ 1,000 

€ 1,000 

€ 1,000 

€ 1,000 

36 

31 

78 

31 

82 

29 

287 

-- 

-- 

-- 

-- 

-- 

-- 

-- 

52 

51 

-- 

74 

-- 

27 

204 

88 

82 

78 

105 

82 

56 

491 

The remuneration of the supervisory board members in 2015 is set out in the table below: 

dsssds  

2015 

Mr. Dinko Valerio   

Mr. Henri Termeer 

Mr. Antoine Papiernik 

Ms. Alison Lawton  

Mr. Paul Baart 

Short term 
employee 
benefits 

Post 
employment 
benefits 

Share-based 
payment 

Total 

€ 1,000 

€ 1,000 

€ 1,000 

€ 1,000 

36 

34 

73 

31 

73 

247 

-- 

-- 

-- 

-- 

-- 

-- 

12 

11 

-- 

48 

-- 

71 

48 

45 

73 

79 

73 

318 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
PAGE 74 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

As at December 31, 2016:  

  Mr. Valerio holds 1,043,420 ordinary shares in the Company, as well as 56,261 options. In 2014, Mr. 

Valerio was granted 64,646 options under the Option Plan to acquire depositary receipts issued for 

ordinary shares at an exercise price of € 1.11 per option. Under this option grant, 32,374 options were 

exercisable immediately, while the remaining 32,272 options vest in four annual equal tranches of 25% 

starting for the first time as of the first anniversary of the date of grant. Mr. Valerio exercised 32,374 

options on June 30, 2014, for which he received 32,374 depositary receipts issued for ordinary shares 

after payment of the exercise price. These depositary receipts have been included in his total number of 

ordinary shares held. In 2016, Mr. Valerio was granted 23,989 options at an average exercise price of € 

6.08 per option. 

  Mr. Termeer holds 1,730,714 ordinary shares in the Company as well as 52,698 options. In 2014, 

Mr. Termeer was granted 57,520 options under the Option Plan to acquire depositary receipts issued 

for ordinary shares at an exercise price of € 1.11 per option. Under this option grant 28,811 options 

were exercisable immediately, while the remaining 28,709 options vest in four annual equal tranches of 

25% starting for the first time as of the first anniversary of the date of grant. Mr. Termeer exercised 

28,811 options on June 30, 2014, for which he received 28,811 depositary receipts issued for ordinary 

shares after payment of the total exercise price. These depositary receipts have been included in his 

total number of ordinary shares. In 2016, Mr. Termeer was granted 23,989 options at an average 

exercise price of € 6.08 per option. 

  Mr. Antoine Papiernik does not hold any shares or options in the Company. As a managing partner of 

Sofinnova Partners SAS, the management company of Sofinnova Capital VII FCPR, holder of 2,769,125 

ordinary shares, Mr. Papiernik may be deemed to have share voting and investment power with respect 

to such shares.  

  Ms. Lawton holds 36,809 options. In 2014, Ms. Lawton was granted 7,850 options under the Option Plan 

to acquire depositary receipts issued for ordinary shares at an exercise price of € 10.03 per option. In 

2015, she was granted 4,970 options with an exercise price of € 16.10 per option. In 2016, she was 

granted 23,989 options with an average exercise price of € 6.08 per option. Under these option grants 

options vest in four annual equal tranches of 25% starting for the first time as of the first anniversary of 

the date of grant. 

  Mr. Paul Baart does not hold any shares or options in the Company.  
  Mr. James Shannon holds 33,069 options. In 2016, he was granted 33,069 options at an exercise price of 
€ 4.32 per option. Under these option grants options vest in four annual equal tranches of 25% starting 
for the first time as of the first anniversary of the date of grant. 

 
 
 
 
(b) Compensation of key management personnel  

Our management board is supported by our officers, or senior management. The total remuneration of the 

management board and senior management in 2016 amounted to € 3,038,000 with the details set out in the 

PAGE 75 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

table below:  

dsssds  

Mr. D.A. de Boer 

Mr. R.K. Beukema  

Management Board 

Senior Management 

2016 

Short term 
employee 
benefits 

Post 
employment 
benefits 

Share-based 
payment 

Total 

€ 1,000 

€ 1,000 

€ 1,000 

€ 1,000 

429 

346 

775 

1,020 

1,795 

7 

13 

20 

48 

68 

391 

165 

556 

619 

1,175 

827 

524 

1,351 

1,687 

3,038 

1  
2  

Short term employee benefits includes a bonus for Mr. Daniel de Boer of € 131,000 based on goals realised in 2016. 
Short term employee benefits includes a bonus for Mr. René Beukema of € 76,000 based on goals realised in 2016. 

The total remuneration of the management board and senior management in 2015 amounted to € 2,420,000 

with the details set out in the table below:  

dsssds  

2015 

Mr. D.A. de Boer 

Mr. R.K. Beukema  

Management Board 

Senior Management 

Short term 
employee 
benefits 

Post 
employment 
benefits 

Share-based 
payment 

Total 

€ 1,000 

€ 1,000 

€ 1,000 

€ 1,000 

3971 

3132 

710 

943 

1,653 

7 

13 

20 

27 

47 

164 

88 

252 

468 

720 

568 

414 

982 

1,438 

2,420 

1  
2  

Short term employee benefits includes a bonus for Mr. Daniel de Boer of € 100,000 based on goals realised in 2015. 
Short term employee benefits includes a bonus for Mr. René Beukema of € 46,000 based on goals realised in 2015. 

As at December 31, 2016:  

  Mr. de Boer holds 1,171,208 ordinary shares in the Company as well as 209,621 options. In 2014, Mr. de 

Boer was awarded a total number of 55,992 options to acquire ordinary shares at € 3.04 per option. In 

2015, he was awarded 23,902 options at an exercise price of € 16.10 per option. In 2016, he was 

awarded 129,727 options at an exercise price of € 6.64 per option. These options vest over four years in 

equal annual installments and had a remaining weighted-average contractual life of 8.5 years at 

December 31, 2016.  

  Mr. Beukema holds 300,000 ordinary shares in the Company as well as 197,673 options. In 2014, 

Mr. Beukema was awarded 30,541 options to acquire ordinary shares at € 3.04 per option. In 2015, he 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
PAGE 76 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

was awarded 8,713 options at an exercise price of € 16.10 per option. In 2016, he was awarded 50,608 

options at an exercise price of € 6.64 per option. These options vest over four years in equal annual 

installments and had a remaining weighted-average contractual life of 7.5 years at December 31, 2016.  

ProQR does not grant any loans, advanced payments and guarantees to members of the Management and 

Supervisory Board. 

24. Subsequent events  

On March 27, 2017, the Company announced that it appointed David M. Rodman, MD as Chief Development 

Strategy Officer. David will join ProQR in April 2017 having previously served in leadership roles with Novartis 

Institutes for Biomedical Research (NIBR), Vertex Pharmaceuticals, miRagen Therapeutics and Nivalis 

Therapeutics. Prior to moving to industry in 2005, David had a distinguished academic career, leading the 

Center for Genetic Lung Diseases at the University of Colorado and directing the Cystic Fibrosis Care, 

Teaching and Research efforts at the National Jewish Medical and Research Center in Denver, Colorado. 

During 12 years in industry, David has had global responsibility for driving innovation in the translation of 

cutting-edge science into transformational new therapies for rare diseases including CF, pulmonary fibrosis, 

pulmonary artery hypertension and severe immunologic and inflammatory diseases. At Vertex 

Pharmaceuticals he directed early- and late-stage CF clinical development programs including Kalydeco®, 

Orkambi® and VX-661. David received a BA in Economics from Haverford College in 1976, an MD from the 

University of Pennsylvania in 1980 and completed training in Internal Medicine, Pulmonary and Critical Care 

Medicine at the University of Colorado. He has served as an advisor to the National Institutes of Health, was 

elected to the American Society for Clinical Investigation and is a Fellow of the American Heart Association. 

 
 
 
 
Company balance sheet at December 31, 2016 

PAGE 77 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

(Before appropriation of result) 

dsssds  

ASSETS 

Non-current assets 

Intangible assets 

Property, plant and equipment 

Financial fixed assets 

Current assets 

Social securities and other taxes 

Prepayments and other receivables 

Cash and cash equivalents 

TOTAL ASSETS 

EQUITY 

Shareholders' equity 

Share capital 

Share premium reserve 

Equity settled employee benefits reserve 

Translation reserve 

Accumulated deficit 

Unappropriated result 

LIABILITIES 

Provisions 

Non-current liabilities 

Borrowings 

Current liabilities 

Trade payables 

Social securities and other taxes 

Pension premiums 

Deferred income 

Other current liabilities 

Note 

December 31, 
2016 

December 31, 
2015 

€ 1,000 

€ 1,000 

-- 

-- 

0 

0 

395 

12,217 

59,042 

71,654 

-- 

-- 

0 

0 

774 

1,638 

94,862 

97,274 

71,654 

97,274 

934 

123,597 

4,343 

(15) 

(36,630) 

(39,103) 

53,136 

934 

123,595 

1,899 

1 

(15,798) 

(20,832) 

89,799 

12,175 

1,922 

5,697 

5,697 

-- 

106 

-- 

-- 

540 

646 

4,824 

4,824 

-- 

38 

-- 

144 

547 

729 

27 

28 

29 

30 

31 

32 

13 

33 

TOTAL EQUITY AND LIABILITIES 

71,654 

97,274 

The accompanying notes are an integral part of these financial statements. 

 
 
 
 
 
 
 
 
 
 
 
  
 
 
 
 
 
 
 
 
 
 
 
 
  
 
 
 
 
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
 
 
 
 
 
PAGE 78 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

Company income statement for the year ended December 31, 2016 

dsssds  

Note 

Share in results of participating interests, after taxation 

27 

Other result after taxation 

Net result for the year 

The accompanying notes are an integral part of these financial statements. 

2016 

€ 1,000 

(37,537) 

(1,566) 

2015 

€ 1,000 

(14,104) 

(6,728) 

(39,103) 

(20,832) 

 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
 
 
 
 
 
 
PAGE 79 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

Notes to the Company financial statements for the year ended December 31, 2016 

25. General 

The company financial statements are part of the 2016 financial statements of ProQR Therapeutics N.V. (the 

‘Company’) and have been prepared in accordance with the legal requirements of Part 9, Book 2 of the 

Netherlands Civil Code. 

With reference to the income statement of the company, use has been made of the exemption pursuant to 

Section 402 of Book 2 of the Netherlands Civil Code. 

26. Principles for the measurement of assets and liabilities and the determination of the 
result 

For setting the principles for the recognition and measurement of assets and liabilities and determination of 

the result for its company financial statements, the Company makes use of the option provided in section 

2:362(8) of the Netherlands Civil Code. This means that the principles for the recognition and measurement 

of assets and liabilities and determination of the result (hereinafter referred to as principles for recognition 

and measurement) of the company financial statements of the Company are the same as those applied for 

the consolidated IFRS financial statements. See page 51 for a description of these principles. 

Participating interests in group companies 

Participating interests in group companies are accounted for in the company financial statements according 

to the equity method. If the net asset value is negative, the participating interest is valued at nil. This likewise 

takes into account other long-term interests that should effectively be considered part of the net investment 

in the participating interest. If the company fully or partly guarantees the liabilities of the associated company 

concerned, or has the effective obligation respectively to enable the associated company to pay its (share of 

the) liabilities, a provision is formed. Upon determining this provision, provisions for doubtful debts already 

deducted from the receivables from the associated company are taken into account. Refer to the basis of 

consolidation accounting policy in the consolidated financial statements. 

Result of participating interests 
The share in the result of participating interests consists of the share of the Company in the result of these 

participating interests. In so far as gains or losses on transactions involving the transfer of assets and 

liabilities between the Company and its participating interests or between participating interests themselves 

can be considered unrealised, they have not been recognised. 

27. Financial fixed assets 

dsssds  

Participating interests in group companies 

December 31, 
2016 

December 31, 
2015 

€ 1,000 

€ 1,000 

0 

0 

0 

0 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
PAGE 80 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

Movements in financial fixed assets were as follows: 

dsssds  

Net asset value as of January 1 

Share in results of participating interests, after taxation 

Exchange differences 

Change in provisions for negative net asset value 

Net asset value as of December 31 

Participating 
interests  
in group 
 companies  

Total 

€ 1,000 

€ 1,000 

0 

37,537 

(16) 

(37,553) 

0 

0 

37,537 

(16) 

(37,553) 

0 

At December 31, 2016, the Company, having its statutory seat in Leiden, the Netherlands, is the ultimate 

parent company of the following consolidated participating interests: 

Name 

ProQR Therapeutics Holding B.V. 

ProQR Therapeutics I B.V. 

ProQR Therapeutics II B.V. 

ProQR Therapeutics III B.V. 

ProQR Therapeutics IV B.V. 

ProQR Therapeutics VI B.V. 

ProQR Therapeutics VII B.V. 

ProQR Therapeutics VIII B.V. 

ProQR Therapeutics IX B.V. 

ProQR Therapeutics I Inc. 

Location 

Share in issued capital 

Leiden, the Netherlands 

Leiden, the Netherlands 

Leiden, the Netherlands 

Leiden, the Netherlands 

Leiden, the Netherlands 

Leiden, the Netherlands 

Leiden, the Netherlands 

Leiden, the Netherlands 

Leiden, the Netherlands 

Delaware, United States 

100% 

100% 

100% 

100% 

100% 

100% 

100% 

100% 

100% 

100% 

For details on the accounts receivable from participating interests and the other receivables, reference is 

made to note 29. 

28. Social Security and Other Taxes 

dsssds  

Value added tax 

All receivables are considered short-term and due within one year.  

December 31, 
2016 

December 31, 
2015 

€ 1,000 

€ 1,000 

395 

395 

774 

774 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
PAGE 81 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

29. Prepayments and Other Receivables  

dsssds  

Accounts receivable from group companies 

Prepayments 

Other receivables 

All receivables are considered short-term and due within one year.  

30. Cash and Cash Equivalents  

dsssds  

Cash at banks 

Bank deposits 

December 31, 
2016 

December 31, 
2015 

€ 1,000 

€ 1,000 

10,854 

235 

1,128 

12,217 

855 

270 

513 

1,638 

December 31, 
2016 

December 31, 
2015 

€ 1,000 

€ 1,000 

56,196 

2,846 

59,042 

94,862 

-- 

94,862 

The cash at banks is at full disposal of the Company. Bank deposits are convertible into cash upon request of 

the Company. 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
PAGE 82 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

31. Shareholders’ equity 

Share 
Capital 

Share 
Premium 

Equity 
Settled 
Employee 
Benefit 
Reserve 

Trans-
lation 
Reserve 

Accumu-
lated 
Deficit 

Unappro-
priated 
result 

Total 
Equity 

€ 1,000 

€ 1,000 

€ 1,000 

€ 1,000 

€ 1,000 

€ 1,000 

€ 1,000 

Balance at January 1, 2015 

934 

123,581 

687 

Retained result 

Foreign exchange differences 

Recognition of share-based 
payments 

Share options exercised 

Result for the year 

-- 

-- 

-- 

0 

-- 

-- 

-- 

-- 

14 

-- 

-- 

-- 

1,212 

-- 

-- 

Balance at December 31, 
2015 

934 

123,595 

1,899 

Retained result 

Foreign exchange differences 

Recognition of share-based 
payments 

Share options exercised 

Result for the year 

Balance at December 31, 
2016 

-- 

-- 

-- 

0 

-- 

-- 

-- 

-- 

2 

-- 

-- 

-- 

2,454 

-- 

-- 

-- 

-- 

1 

-- 

-- 

-- 

1 

-- 

(16) 

-- 

-- 

-- 

(3,671) 

(12,127) 

109,404 

(12,127) 

12,127  

(20,832) 

(20,832) 

(15,798) 

(20,832) 

89,799 

(20,832) 

20,832 

-- 

1 

1,212 

14 

-- 

(16) 

2,454 

2 

-- 

-- 

-- 

-- 

-- 

-- 

-- 

-- 

-- 

-- 

-- 

-- 

-- 

-- 

(39,103) 

(39,103) 

934 

123,597 

4,353 

(15) 

(36,630) 

(39,103) 

53,136 

The 2015 result was added to the accumulated deficit in accordance with the resolution of the Annual 

General Meeting of shareholders. At the upcoming Annual General Meeting of shareholders, it will be 

proposed to add the 2016 result to the accumulated deficit. For more details we refer to note 12 to the 

consolidated financial statements. 

32. Provisions 

dsssds  

December 31, 
2016 

December 31, 
2015 

Provision for negative equity group companies 

€ 1,000 

€ 1,000 

Balance at January 1 

Provisions made during the year 

Balance at December 31 

1,922 

10,253 

-- 

1,922 

12,175 

1,922 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
33. Current Liabilities  

dsssds  

Trade payables 

Social securities and other taxes 

Pension premiums 

Deferred income 

Accrued expenses and other liabilities 

PAGE 83 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

December 31, 
2016 

December 31, 
2015 

€ 1,000 

€ 1,000 

-- 

106 

-- 

-- 

540 

646 

-- 

38 

-- 

144 

547 

729 

34. Commitments and Contingencies  

(a) Claims  
There are no claims known to management related to the activities of the Company.  

(b) Clinical support agreement 
In August 2014, the Company entered into an agreement with Cystic Fibrosis Foundation Therapeutics, Inc., 

or CFFT, a subsidiary of the Cystic Fibrosis Foundation, pursuant to which CFFT agreed to provide the 

Company with up to $ 3 million to support the clinical development of QR-010.  

Pursuant to the terms of the agreement, the Company is obligated to make a one-time milestone payment to 

CFFT of up to approximately $ 80 million, payable in three equal annual installments following the first 

commercial sale of QR-010, the first of which is due within 90 days following the first commercial sale. The 

Company is also obligated to make a one-time milestone payment to CFFT of up to $ 3 million if net sales of 

QR-010 exceed $ 500 million in a calendar year. Lastly, the Company is obligated to make a payment to CFFT 

of up to approximately $ 6 million if it transfers, sells or licenses QR-010 other than for certain clinical or 

development purposes, or if the Company enters into a change of control transaction. Either CFFT or the 

Company may terminate the agreement for cause, which includes the Company’s material failure to achieve 

certain commercialization and development milestones. The Company’s payment obligations survive the 

termination of the agreement.  

(c) Several liability and guarantees 
The Company has issued declarations of joint and several liabilities for debts arising from the actions of 

Dutch consolidated participating interests, as meant in article 2:403 of the Netherlands Civil Code. 

The Company constitutes a tax entity with its Dutch subsidiaries for corporate income tax purposes; the 

standard conditions prescribe that all companies of the tax entity are jointly and severally liable for the 

corporate income tax payable. 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
PAGE 84 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

35. Auditor fees 

The fees for services provided by our external auditor, Deloitte Accountants B.V., are specified below for each 

of the financial years indicated: 

dsssds  

Audit fees 

Audit-related fees 

Tax fees 

All other fees 

Audit fees 

2016 

€ 1,000 

2015 

€ 1,000 

165 

39 

-- 

-- 

204 

193 

-- 

-- 

-- 

193 

Consist of aggregate fees for professional services provided in connection with the annual audit of our 

financial statements, the review of our quarterly financial statements, consultations on accounting matters 

directly related to the audit, and comfort letters, consents and assistance with and review of documents filed 

with the SEC.  

 
 
 
 
 
 
 
 
 
 
 
 
 
PAGE 85 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

Signing of the Annual Report 

Leiden, March 31, 2017, 

D.A. de Boer 

D. Valerio 

R.K. Beukema 

H.A. Termeer 

A.B. Papiernik 

A. Lawton 

P.R. Baart 

J.S.S. Shannon (as of June 21, 2016) 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
PAGE 86 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

Other information 

Independent auditor’s report  

Reference is made to the independent auditor’s report as included hereinafter. 

Statutory arrangement concerning the appropriation of the result 

In Article 21 of the Company statutory regulations the following has been presented concerning the 

appropriation of result: 

1. The profit is at the free disposal of the General Meeting of Shareholders. 

2. The Company may only distribute profits to shareholders and other recipients to distributable profits to 

the extent that the equity exceeds the paid-up capital plus the reserves required by law. 

3. Distribution of profits shall take place after adoption of the annual accounts from which it becomes clear 

that distribution is permissible. 

4. When calculating the distribution of profits shares held by the Company shall be disregarded, unless this 

shares has been encumbered with usufruct or right of pledge or certificates thereof are issued as a result of 

which the entitlement to profits accrue to the usufructuary, pledgee or holder of the certificates. 

5. Certificates held by the Company or whereon the Company holds limited rights as a result of which the 

Company is entitled to distribution of profits shall also be disregarded when calculating the distribution of 

profits. 

6. The Company may make interim distributions, only if the requirements in paragraph 2 are met.  

 
 
 
 
 
 
 
 
 
 
PAGE 87 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

Independent auditor’s report 

To the Shareholders and the Supervisory Board of ProQR Therapeutics N.V.  

Report on the financial statements 2016  

Our Opinion 

We have audited the financial statements 2016 of ProQR Therapeutics N.V., based in Leiden, the Netherlands. 

The financial statements include the consolidated financial statements and the company financial 

statements. 

In our opinion: 

 

The consolidated financial statements give a true and fair view of the financial position of ProQR 

Therapeutics N.V. as at December 31, 2016, and of its result and its cash flows for 2016 in accordance 

with International Financial Reporting Standards as adopted by the European Union (EU-IFRS) and with 

Part 9 of Book 2 of the Dutch Civil Code. 

 

The company financial statements give a true and fair view of the financial position of ProQR 

Therapeutics N.V. as at December 31, 2016, and of its result for 2016 in accordance with Part 9 of Book 

2 of the Dutch Civil Code. 

The consolidated financial statements comprise: 

 

 

 

The consolidated statement of financial position as at December 31, 2016. 

The following statements for 2016: the consolidated statement of profit or loss and comprehensive 

income, changes in equity and cash flows. 

The notes comprising a summary of the significant accounting policies and other explanatory 

information.  

The company financial statements comprise: 

 

 

 

The company balance sheet as at December 31, 2016. 

The company income statement for 2016. 

The notes comprising a summary of the accounting policies and other explanatory information. 

Basis for our opinion 

We conducted our audit in accordance with Dutch law, including the Dutch Standards on Auditing. Our 

responsibilities under those standards are further described in the “Our responsibilities for the audit of the 

financial statements” section of our report. 

We are independent of ProQR Therapeutics N.V. in accordance with the Verordening inzake de 

onafhankelijkheid van accountants bij assurance-opdrachten (ViO) and other relevant independence 

regulations in the Netherlands. Furthermore we have complied with the Verordening gedrags- en 

beroepsregels accountants (VGBA). 

We believe the audit evidence we have obtained is sufficient and appropriate to provide a basis for our 

opinion. 

 
 
 
 
PAGE 88 / 91 
Financial Statements 2016 
ANNUAL REPORT 2016 

Materiality 

Based on our professional judgement we determined the materiality for the financial statements as a whole 

at EUR 2.5 million. The materiality is based on 7.5% of normalized loss before tax. We have also taken into 

account misstatements and/or possible misstatements that in our opinion are material for the users of the 

financial statements for qualitative reasons. 

We agreed with the Supervisory Board that misstatements in excess of EUR 125,000, which are identified 

during the audit, would be reported to them, as well as smaller misstatements that in our view must be 

reported on qualitative grounds.  

Scope of the group audit 

ProQR Therapeutics N.V. is at the head of a group of entities. The financial information of this group is 

included in the consolidated financial statements of ProQR Therapeutics N.V.. 

The financial administration for all group entities is centralized in the Netherlands. Consequently, we have 

centralized our audit approach and we have been able to obtain sufficient and appropriate audit evidence 

about the group’s financial information to provide an opinion about the financial statements. 

Our key audit matters 

Key audit matters are those matters that, in our professional judgement, were of most significance in our 

audit of the financial statements. We have communicated the key audit matters to the Supervisory Board. 

The key audit matters are not a comprehensive reflection of all matters discussed. 

These matters were addressed in the context of our audit of the financial statements as a whole and in 

forming our opinion thereon, and we do not provide a separate opinion on these matters.  

Research and development expenses 

The total research and development expenses for the year 2016 amounts to EUR 31.9 million. These research 

and development expenses consists of payroll costs of employees as well as outsourced research and 

development activities with third party suppliers. The research and development activities with these 

suppliers are concluded in master service agreements and statements of work. These outsourced research 

and development activities are typically performed over a period of time and allocation of expenses in each 

reporting period based on the progress of the work involves judgement. Our audit procedures included, 

amongst others, the review of the agreements with suppliers and the related accounting evaluation as well as 

the timing of expenses recognized. 

Significant contracts 

ProQR Therapeutics N.V. concluded several significant contracts, amongst others, the agreements with 

European Commission in relation to H2020 grant and the above mentioned research and development 

agreements. These contracts contain terms and conditions that may require complex accounting and/or 

significant long-term commitments that require disclosure in the financial statements. Our audit procedures 

included, amongst others, the review of the contract register, review of the contract terms and related 

accounting evaluation of the impact on the financial statements including disclosures of the commitments.   

Cash and cash equivalents 

The total cash and cash equivalents as per December 31, 2016 amounts to EUR 59.2 million. We focused on 

this area as the cash and cash equivalents are material to the financial statements. We reconciled the bank 

balances to bank confirmations, recalculated the foreign exchange result on these balances and reviewed the 

bank confirmations and underlying agreements for deposit balances to assess the presentation and 

disclosure in the financial statements. 

 
 
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ANNUAL REPORT 2016 

Report on the other information included in the annual accounts 

In addition to the financial statements and our auditor’s report, the annual accounts contain other 

information that consists of: 

  Management Board's Report. 

 

Other Information as required by Part 9 of Book 2 of the Dutch Civil Code. 

Based on the following procedures performed, we conclude that the other information: 

 

 

Is consistent with the financial statements and does not contain material misstatements. 

Contains the information as required by Part 9 of Book 2 of the Dutch Civil Code.  

We have read the other information. Based on our knowledge and understanding obtained through our audit 

of the financial statements or otherwise, we have considered whether the other information contains 

material misstatements. 

By performing these procedures, we comply with the requirements of Part 9 of Book 2 of the Dutch Civil Code 

and the Dutch Standard 720. The scope of the procedures performed is substantially less than the scope of 

those performed in our audit of the financial statements. 

Management is responsible for the preparation of other information, including the Management Board's 

Report in accordance with Part 9 of Book 2 of the Dutch Civil Code, and the other information as required by 

Part 9 of Book 2 of the Dutch Civil Code.  

Report on other legal and regulatory requirements 

Engagement 

We were engaged by the Supervisory Board as auditor of ProQR Therapeutics N.V. as of the audit for year 

2012 and have operated as statutory auditor ever since that date.  

Description of responsibilities for the financial statements 

Responsibilities of management and the Supervisory Board for the financial statements 

Management is responsible for the preparation and fair presentation of the financial statements in 

accordance with EU-IFRS and Part 9 of Book 2 of the Dutch Civil Code. Furthermore, management is 

responsible for such internal control as management determines is necessary to enable the preparation of 

the financial statements that are free from material misstatement, whether due to fraud or error. 

As part of the preparation of the financial statements, management is responsible for assessing the 

company’s ability to continue as a going concern. Based on the financial reporting framework mentioned, 

management should prepare the financial statements using the going concern basis of accounting unless 

management either intends to liquidate the company or to cease operations, or has no realistic alternative 

but to do so.  

Management should disclose events and circumstances that may cast significant doubt on the company’s 

ability to continue as a going concern in the financial statements. 

 
 
 
 
 
 
 
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The Supervisory Board is responsible for overseeing the company’s financial reporting process. 

Our responsibilities for the audit of the financial statements 

Our objective is to plan and perform the audit assignment in a manner that allows us to obtain sufficient and 

appropriate audit evidence for our opinion. 

Our audit has been performed with a high, but not absolute, level of assurance, which means we may not 

have detected all material errors and fraud. 

Misstatements can arise from fraud or error and are considered material if, individually or in the aggregate, 

they could reasonably be expected to influence the economic decisions of users taken on the basis of these 

financial statements. The materiality affects the nature, timing and extent of our audit procedures and the 

evaluation of the effect of identified misstatements on our opinion. 

We have exercised professional judgment and have maintained professional skepticism throughout the audit, 

in accordance with Dutch Standards on Auditing, ethical requirements and independence requirements. Our 

audit included e.g.: 

 

Identifying and assessing the risks of material misstatement of the financial statements, whether due to 

fraud or error, designing and performing audit procedures responsive to those risks, and obtaining 

audit evidence that is sufficient and appropriate to provide a basis for our opinion. The risk of not 

detecting a material misstatement resulting from fraud is higher than for one resulting from error, as 

fraud may involve collusion, forgery, intentional omissions, misrepresentations, or the override of 

 

 

 

 

 

internal control. 

Obtaining an understanding of internal control relevant to the audit in order to design audit procedures 

that are appropriate in the circumstances, but not for the purpose of expressing an opinion on the 

effectiveness of the company’s internal control. 

Evaluating the appropriateness of accounting policies used and the reasonableness of accounting 

estimates and related disclosures made by management. 

Concluding on the appropriateness of management’s use of the going concern basis of accounting, and 

based on the audit evidence obtained, whether a material uncertainty exists related to events or 

conditions that may cast significant doubt on the company’s ability to continue as a going concern. If we 

conclude that a material uncertainty exists, we are required to draw attention in our auditor’s report to 

the related disclosures in the financial statements or, if such disclosures are inadequate, to modify our 

opinion. Our conclusions are based on the audit evidence obtained up to the date of our auditor’s 

report. However, future events or conditions may cause the company to cease to continue as a going 

concern. 

Evaluating the overall presentation, structure and content of the financial statements, including the 

disclosures.  

Evaluating whether the financial statements represent the underlying transactions and events in a 

manner that achieves fair presentation. 

Because we are ultimately responsible for the opinion, we are also responsible for directing, supervising and 

performing the group audit. In this respect we have determined the nature and extent of the audit 

procedures to be carried out for group entities. Decisive were the size and/or the risk profile of the group 

entities or operations. On this basis, we selected group entities for which an audit or review had to be carried 

out on the complete set of financial information or specific items. 

 
 
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We communicate with the Supervisory Board regarding, among other matters, the planned scope and timing 

of the audit and significant audit findings, including any significant findings in internal control that we identify 

during our audit. 

We provide the Supervisory Board with a statement that we have complied with relevant ethical 

requirements regarding independence, and to communicate with them all relationships and other matters 

that may reasonably be thought to bear on our independence, and where applicable, related safeguards. 

From the matters communicated with the Supervisory Board, we determine those matters that were of most 

significance in the audit of the financial statements of the current period and are therefore the key audit 

matters. We describe these matters in our auditor’s report unless law or regulation precludes public 

disclosure about the matter or when, in extremely rare circumstances, not communicating the matter is in 

the public interest. 

Amsterdam, March 31, 2017 

Deloitte Accountants B.V. 

P.J.M.A. van de Goor 

 
 
 
ANNUAL

REPORT 

2016

Acting in the interest of patients

ProQR Therapeutics N.V. 

T  : +31 88 166 7000 
W : www.proqr.com 
E  : info@proqr.com

Headquarters Leiden:
Zernikedreef 9, 2333 CK Leiden,
the Netherlands

Office Palo Alto:
543 Bryant Street, Palo Alto,
CA 94301, USA