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Voyager Therapeutics, Inc.

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FY2018 Annual Report · Voyager Therapeutics, Inc.
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2018 Annual Report

To Our Shareholders,

I came to Voyager in mid-2018 to join an exceptional group of people working together on a mission to change the lives of people 
suffering from severe neurological diseases.  As the fi eld of gene therapy has blossomed, we see the potential with this modality 
to develop great medicines for some of the most intractable neurological conditions. Our combined expertise in gene therapy and 
neuroscience gives us the focus and ability to lead in this area and make a true difference for patients. 

In  2018,  we  initiated  RESTORE-1,  the  fi rst  of  two  anticipated  pivotal  trials  for VY-AADC  in  patients  with  Parkinson’s 
disease.  This came on the back of clinical results from our Phase 1b trial showing improvements in patients’ motor function 
and quality of life over sustained periods following a one-time administration.  These results were recently published in the peer-
reviewed Annals of Neurology.  To further our efforts in this devastating but all too common disease, we recently entered into a 
collaboration  agreement  with  Neurocrine  Biosciences  to  co-develop  and  co-promote VY-AADC.    Neurocrine’s  experience  and 
capabilities within the movement disorder specialty are already evident in the early days of our collaboration.  Our current joint 
efforts are focused on screening and enrolling patients in the RESTORE-1 trial, and on preparing for the initiation of the RESTORE-2 
trial targeted for next year.

Our preclinical pipeline has also advanced over the past year.  At the European Society of Gene and Cell Therapy conference in 
the fall of 2018, we presented positive preclinical results of VY-HTT01 for Huntington’s disease and VY-SOD102 for ALS.  The results 
refl ected the optimization efforts undertaken around the delivery of these gene therapies aimed at suppressing the expression of 
disease-causing genes.  In the case of our Huntington’s disease program, we achieved meaningful reductions of Huntingtin gene 
expression in both the deeper and outer layers of the brain of non-human primates.  For our ALS-SOD1 program, intended for a 
subset of patients with a particularly fast-progressing form of ALS, our preclinical work showed that direct injection of VY-SOD102 
in the cervical motor neurons also generated meaningful reductions of the toxic SOD1 gene in multiple locations throughout a large 
animal spinal cord, including the regions most critical for maintaining respiratory function.  Preclinical pharmacology and toxicology 
studies are underway to support the potential fi ling of IND applications for both programs later this year. 

As we apply learnings from our more advanced programs and from the broader fi eld, we are intensifying our efforts 
on our research pipeline.  Our use of AAV has initially been directed at gene replacement and gene knock-down, and many 
compelling targets remain in both applications.  Through a pair of new collaborations with AbbVie, we are now also using AAV to 
vectorize antibodies.  By utilizing our novel blood-brain barrier penetrant capsids, we aim to overcome one of the major limitations of 
biologic therapies for neurodegenerative diseases: the suffi cient transport of therapeutic antibodies across the blood-brain barrier.  
Our initial programs in this additional application of AAV are targeted at pathological species of tau for Alzheimer’s disease and 
other tauopathies and alpha-synuclein for Parkinson’s and other synucleinopathies.  In addition to building on this work, we plan to 
further our efforts towards expanding the portfolio and continuing to enhance our gene therapy platform. 

Signifi cant progress has already been made in 2019, and we anticipate that this year will be pivotal for Voyager as we continue 
to build on the exciting momentum established in recent months and advance our lead and preclinical pipeline programs.  As 
our organization continues to grow, we remain doggedly focused on connecting and listening to our science partners, healthcare 
providers, and most importantly patients, each of whom inform our decisions and fuel our drive.

Sincerely,

Andre Turenne

President and Chief Executive Offi cer

UNITED STATES  

SECURITIES AND EXCHANGE COMMISSION 
Washington, DC 20549  

FORM 10-K 

 (Mark One)  
  ANNUAL REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934  

For the fiscal year ended December 31, 2018 

OR 

 

TRANSITION REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934  

For the transition period from                          to                       

Commission File Number: 001-37625  

Voyager Therapeutics, Inc. 

(Exact Name of Registrant as Specified in Its Charter)  

 Delaware 
(State or Other Jurisdiction of 
Incorporation or Organization) 

75 Sidney Street, 
Cambridge, Massachusetts 
(Address of Principal Executive Offices) 

46-3003182 
(IRS Employer 
Identification No.) 

02139 
(Zip Code) 

(857) 259-5340  
(Registrant’s Telephone Number, Including Area Code)  

Indicate by check mark if the registrant is a well-known seasoned issuer, as defined in Rule 405 of the Securities Act. Yes     No   

Indicate by check mark if the registrant is not required to file reports pursuant to Section 13 or Section 15(d) of the Act. Yes      No   

Indicate by check mark whether the registrant: (1) has filed all reports required to be filed by Section 13 or 15(d) of the Securities Exchange Act of 1934 during the 

preceding 12 months (or for such shorter period that the registrant was required to file such reports), and (2) has been subject to such filing requirements for the past 
90 days. Yes      No   

Indicate by check mark whether the registrant has submitted every Interactive Data File required to be submitted pursuant to Rule 405 of Regulation S-T (§ 232.405 

of this chapter) during the preceding 12 months (or for such shorter period that the registrant was required to submit such files). Yes      No   

Indicate by check mark if disclosure of delinquent filers pursuant to Item 405 of Regulation S-K (§ 229.405 of this chapter) is not contained herein, and will not be 
contained, to the best of registrant’s knowledge, in definitive proxy or information statements incorporated by reference in Part III of this Form 10-K or any amendment 
to this Form 10-K.  

Indicate by check mark whether the registrant is a large accelerated filer, an accelerated filer, a non-accelerated filer, a smaller reporting company, or an emerging 
growth company. See the definitions of “large accelerated filer,” “accelerated filer,” “smaller reporting company,” and “emerging growth company” in Rule 12b-2 of 
the Exchange Act.  

Large accelerated filer 
Non-accelerated filer 

  
  

   Accelerated filer 
   Smaller reporting company 
  Emerging growth company 

  
  
  

If an emerging growth company, indicate by check mark if the registrant has elected not to use the extended transition period for complying with any new or 

revised financial accounting standards provided pursuant to Section 13(a) of the Exchange Act.  

Indicate by check mark whether the registrant is a shell company (as defined in Rule 12b-2 of the Act). Yes      No   

The aggregate market value of Common Stock held by non-affiliates of the registrant computed by reference to the price of the registrant’s Common Stock as of 
June 29, 2018, the last business day of the registrant’s most recently completed second fiscal quarter, was approximately $442.0 million (based on the last reported sale 
price on the Nasdaq Global Select Market as of such date).  

As of February 22, 2019, there were 32,616,999 shares of the registrant’s common stock, par value $0.001 per share, outstanding.  

DOCUMENTS INCORPORATED BY REFERENCE  

Portions of the registrant’s definitive Proxy Statement relating to its 2019 Annual Meeting of Stockholders are incorporated by reference into Part III of this Annual 

Report on Form 10-K where indicated. Such Proxy Statement is expected to be filed with the U.S. Securities and Exchange Commission not later than 120 days after 
the end of the fiscal year to which this report relates.  



 
 
 
 
 
 
 
 
 
 
 
 
 
  
  
  
  
  
  
 
 
 
 
 
 
 
 
 
 
 
 
 
Table of Contents  

  Page 

  Business 
  Risk Factors 
  Unresolved Staff Comments 
  Properties 
  Legal Proceedings 
  Mine Safety Disclosures 

Market for Registrant’s Common Equity, Related Stockholder Matters and Issuer 
Purchases of Equity Securities 
  Selected Financial Data 
  Management’s Discussion and Analysis of Financial Condition and Results of Operations 
  Quantitative and Qualitative Disclosures about Market Risk 
  Financial Statements and Supplementary Data 
  Changes in and Disagreements with Accountants on Accounting and Financial Disclosure   
  Controls and Procedures 
  Other Information 

  Directors, Executive Officers and Corporate Governance 
  Executive Compensation 
Security Ownership of Certain Beneficial Owners and Management and Related 
Stockholder Matters 
  Certain Relationships and Related Transactions, and Director Independence 
  Principal Accountant Fees and Services 

  Exhibits, Financial Statement Schedules 
  Form 10-K Summary 

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F-37

PART I. 
Item 1. 
Item 1A. 
Item 1B. 
Item 2. 
Item 3. 
Item 4. 

PART II. 
Item 5. 

Item 6. 
Item 7. 
Item 7A. 
Item 8. 
Item 9. 
Item 9A. 
Item 9B. 

PART III. 
Item 10. 
Item 11. 
Item 12. 

Item 13. 
Item 14. 

PART IV. 
Item 15. 
Item 16. 

Signatures 

2 

 
 
   
   
 
 
 
 
 
 
 
 
 
   
 
   
 
 
 
 
 
 
 
 
 
 
   
 
   
 
 
 
 
 
 
 
 
   
 
   
 
 
 
 
   
 
   
 
 
 
 
[This page intentionally left blank] 

Forward-Looking Statements  

This Annual Report on Form 10-K contains forward-looking statements that involve substantial risks and 

uncertainties. All statements other than statements of historical facts contained in this Annual Report on Form 10-K, 
including statements regarding our strategy, future operations, future financial position, future revenue, projected costs, 
prospects, plans, objectives of management and expected market growth, are forward-looking statements. These 
statements involve known and unknown risks, uncertainties and other important factors that may cause our actual results, 
performance or achievements to be materially different from any future results, performance or achievements expressed 
or implied by the forward-looking statements.  

The words “anticipate,” “believe,” “estimate,” “expect,” “intend,” “may,” “might,” “plan,” “predict,” “project,” 

“target,” “potential,” “contemplate,” “anticipate,” “goals,” “will,” “would,” “could,” “should,” “continue,” and similar 
expressions are intended to identify forward-looking statements, although not all forward-looking statements contain 
these identifying words. These forward-looking statements include, among other things, statements about: 

 

 

 

 

 

 

 

 

 

 

 

 

 

our plans to develop and commercialize our product candidates based on adeno-associated virus, or AAV, gene 
therapy; 

our ability to identify and optimize product candidates and novel AAV gene therapy capsids; 

our ongoing and planned clinical trials and related timelines, including our ability to continue to advance VY-
AADC as a treatment for Parkinson’s disease through the ongoing Phase 1b clinical trial and the RESTORE-1 
Phase 2 clinical trial, and our preclinical development efforts and studies; 

formulation changes to our product candidates may require us to conduct additional clinical studies to bridge 
our modified product candidates to earlier versions; 

the timing of and our ability to submit applications for, and obtain and maintain regulatory approvals for our 
product candidates, including our ability to file Investigational New Drug applications for our programs 
including VY-SOD102 for the treatment of a monogenic form of amyotrophic lateral sclerosis, VY-HTT01 for 
the treatment of Huntington’s disease, and VY-FXN01 for the treatment of Friedreich’s ataxia; 

our estimates regarding expenses, future revenues, capital requirements, and needs for additional financing; 

our ability to continue to develop our gene therapy platform; 

our ability to develop a manufacturing capability compliant with current good manufacturing practices for our 
product candidates;  

our ability to access, develop, and obtain regulatory clearance for devices to deliver our AAV gene therapies to 
critical targets of neurological disease;  

our intellectual property position and our ability to obtain, maintain and enforce intellectual property protection 
for our proprietary assets;  

our estimates regarding the size of the potential markets for our product candidates and our ability to serve those 
markets; 

the rate and degree of market acceptance of our product candidates for any indication once approved;  

our strategic collaborations with Sanofi Genzyme Corporation, or Sanofi Genzyme; AbbVie Biotechnology Ltd 
and AbbVie Ireland Unlimited Company, or collectively AbbVie, and Neurocrine Biosciences, Inc., including 
the possibility and timing of Sanofi Genzyme or AbbVie exercising their respective options to certain of our 
programs as specified in the applicable collaboration agreements; 

3 

 

 

 

 

our plans and ability to raise additional capital, including through equity offerings, debt financings, 
collaborations, strategic alliances, and licensing arrangements;  

our competitive position and the success of competing products that are or become available for the indications 
that we are pursuing; 

the impact of government laws and regulations including in the United States, the European Union, and other 
important geographies such as Japan; 

our ability to maintain consistency with results from our ongoing Phase 1b clinical trial and our separate Phase 
1 clinical trial focused on posterior trajectory in future clinical trials; and 

 

our ability to enter into future collaborations, strategic alliances, or licensing arrangements. 

These forward-looking statements are only predictions and we may not actually achieve the plans, intentions or 

expectations disclosed in our forward-looking statements. You should not place undue reliance on our forward-looking 
statements. Actual results or events could differ materially from the plans, intentions and expectations disclosed in the 
forward-looking statements we make. We have based these forward-looking statements largely on our current 
expectations and projections about future events and trends that we believe may affect our business, financial condition 
and operating results. We have included important factors in the cautionary statements included in this Annual Report on 
Form 10-K, particularly in “Part I, Item 1A - Risk Factors” that could cause actual future results or events to differ 
materially from the forward-looking statements that we make. Our forward-looking statements do not reflect the 
potential impact of any future acquisitions, mergers, dispositions, joint ventures or investments we may make.  

You should read this Annual Report on Form 10-K and the documents that we have filed as exhibits to the 

Annual Report on Form 10-K with the understanding that our actual future results may be materially different from what 
we expect. We do not assume any obligation to update any forward-looking statements whether as a result of new 
information, future events or otherwise, except as required by applicable law.  

ITEM 1.  

   BUSINESS 

PART I 

We are a clinical-stage gene therapy company focused on developing life-changing treatments for patients 

suffering from severe neurological diseases. We focus on neurological diseases where we believe an adeno-associated 
virus, or AAV, gene therapy approach that either increases or decreases the production of a specific protein can slow or 
reduce the symptoms experienced by patients, and therefore have a clinically meaningful impact. We have built a gene 
therapy platform that we believe positions us to be a leading company at the intersection of AAV gene therapy and 
severe neurological disease. Our gene therapy platform enables us to engineer, optimize, manufacture and deliver our 
AAV-based gene therapies that have the potential to provide durable efficacy following a single administration.  

Additionally, we are working to identify novel AAV capsids, which are the outer viral protein shells that 

enclose the genetic material of the virus payload. Our team of experts in the fields of AAV gene therapy and 
neuroscience first identifies and selects severe neurological diseases that are well-suited for treatment using AAV gene 
therapy. We then engineer and optimize AAV vectors for delivery of the virus payload to the targeted tissue or cells. Our 
manufacturing process employs an established system that we believe will enable production of high quality AAV 
vectors at commercial-scale. In addition to our capsid optimization efforts, we leverage novel delivery paradigms, 
established routes of administration, and advances in dosing techniques to optimize delivery of our AAV gene therapies 
to target tissues, regions and cell types that are critical to the disease of interest. We believe we can achieve this directly, 
with targeted infusions to discrete regions of the brain or spinal cord, or systemically, in conjunction with our novel 
capsids. 

Our business strategy focuses on discovering, developing, manufacturing and commercializing our gene therapy 
programs. As part of this strategy, we have developed core competencies specific to AAV gene therapy development and 
manufacturing and are beginning to build our commercial infrastructure. This business strategy also includes business 
development activities that may include in-licensing activities or partnering certain programs in specific geographies 

4 

  
 
with collaborators, as we have demonstrated through our collaborations, including those with Sanofi Genzyme, AbbVie, 
and Neurocrine. Since our inception, our operations have focused on organizing and staffing our company, business 
planning, raising capital, establishing our intellectual property portfolio, determining which neurological diseases to 
pursue, advancing our product candidates including delivery and manufacturing, and conducting preclinical studies and 
clinical trials. We do not have any product candidates approved for sale and have not generated any revenue from 
product sales. We have funded our operations primarily through private placements of redeemable convertible preferred 
stock, public offerings of our common stock, our collaboration with Sanofi Genzyme, or the Sanofi Genzyme 
Collaboration, which commenced in February 2015 and our collaboration with AbbVie focusing on tau-related diseases, 
or the AbbVie Tau Collaboration, which commenced in February 2018. Additionally, we recently entered into 
collaborations with Neurocrine, or the Neurocrine Collaboration, which we expect to commence in the first half of 2019, 
and with AbbVie focusing on pathological species of alpha-synuclein, or the AbbVie Alpha-Synuclein Collaboration.  

Our pipeline of gene therapy programs is summarized in the table below:  

Our pipeline consists of programs for severe neurological indications, including Parkinson’s disease; a 
monogenic form of amyotrophic lateral sclerosis, or ALS; Huntington’s disease; Friedreich’s ataxia; tau-related diseases 
including Alzheimer’s disease, frontotemporal dementia, or FTD, and progressive supranuclear palsy; or PSP, Alpha-
synuclein related diseases for Parkinson’s disease and other synucleinopathies; and severe, chronic pain. We may seek 
orphan drug designation, breakthrough therapy designation, or other expedited review processes for certain of our 
product candidates in the United States, Europe, and Japan.  

Our most advanced clinical candidate, VY-AADC for the treatment of Parkinson’s disease, is being evaluated 

for safety and efficacy of escalating doses in an open-label, Phase 1b clinical trial. We have completed enrolling the 
Phase 1b trial of VY-AADC and continue to monitor results. Preliminary data from Cohorts 1 through 3 from this trial 
were reported beginning in late 2016 and most recently in November 2018.  

5 

 
In December 2018, we announced randomization of the first patient in our RESTORE-1 Phase 2, randomized, 

double-blind, placebo-surgery controlled trial evaluating the safety and efficacy of VY-AADC for the treatment of 
Parkinson’s disease in patients with motor fluctuations that are refractory to medical management. The RESTORE-1 
Phase 2 trial will enroll patients who have been diagnosed with Parkinson’s disease for at least four years, are not 
responding adequately to oral medications, and have at least three or more hours of OFF time during the day as measured 
by a validated self-reported patient diary. Patients who meet the eligibility criteria will be randomized (1:1) to one-time 
administration of VY-AADC (for a total dose of up to 2.5×1012 vector genomes, or vg) or placebo surgery.  

We are pursuing additional product candidates in the preclinical stages of development, including treatment 

programs for ALS, Huntington’s disease, Friedreich’s ataxia, tau-related neurodegenerative diseases and the treatment of 
severe, chronic pain. If preclinical studies prove successful, we plan to file investigational new drug, or IND, 
applications for our ALS and Huntington’s disease programs during 2019.  

In late 2017, we initiated additional preclinical studies to further optimize our ALS program’s therapeutic 

approach, including exploration of additional routes of administration and novel AAV capsids in large animal models. 
VY-SOD102, our clinical candidate for the treatment of a monogenic form of ALS, is composed of an adeno-associated 
virus capsid and a proprietary transgene that selectively knocks down, or reduces, levels of SOD1 mRNA. VY-SOD102 
has the potential to durably reduce the levels of toxic mutant SOD1 protein in the spinal cord to slow the progression of 
disease. In late 2018, we presented data on VY-SOD102 administered with a novel delivery paradigm comprising a one-
time, intraparenchymal infusion after laminectomy to the cervical spinal cord of the mini-pig, which has a spinal cord 
similar in length and diameter to the human spinal cord. This delivery approach yielded safe and significant reduction of 
SOD1 at 4 weeks post-dosing at the site of the infusion and throughout the spinal cord, most notably in the cervical and 
thoracic regions critical for respiratory function. Further preclinical studies are underway with VY-SOD102 which, if 
successful, will support a potential filing of an IND application in 2019. 

In 2017, we selected VY-HTT01 as our clinical candidate for the treatment of Huntington’s disease. Recent 

preclinical delivery studies have further optimized the dosing paradigm to support filing of a potential IND application. 
VY-HTT01 is composed of an adeno-associated virus capsid (AAV1) and proprietary transgene that harnesses the RNA 
interference pathway to selectively knock down, or reduce, levels of HTT mRNA. In late 2018, we presented results 
demonstrating significant reduction of HTT mRNA at five weeks post-dosing in adult non-human primates using a 
magnetic resonance imaging, or MRI, guided surgical delivery of VY-HTT01 and a novel delivery paradigm targeting 
both the putamen and thalamus. Targeting the thalamus in addition to the putamen leverages more extensive and more 
preserved neuronal pathways to the cortex than delivery to the putamen alone. This novel dosing paradigm with VY-
HTT01 resulted in safe and significant suppression of HTT in the striatum and in cortical neurons, which are critical in 
the progression of disease. Further preclinical studies are underway with VY-HTT01 which, if successful, will support a 
potential filing of an IND application in 2019. 

Additional preclinical studies are underway including steps to optimize a lead clinical candidate for the 

treatment of Friedreich’s ataxia. Additionally, we are collaborating with AbbVie on the research and development of 
specified vectorized antibody compounds comprised of an AAV or other viral capsid and a virus vector genome that 
encodes one or more antibodies that target and bind to a tau protein. We are also conducting proof-of-concept studies on 
our VY-NAV01 program for the treatment of severe, chronic pain.  

In January 2019, we announced the Neurocrine Collaboration focused on the development and 
commercialization of our VY-AADC gene therapy program for Parkinson’s disease and VY-FXN01 gene therapy 
program for Friedreich’s ataxia, as well as rights to two programs to be determined. Agreements related to the 
Neurocrine Collaboration including the collaboration and license agreement are subject to certain conditions, including 
the expiration or termination of the applicable waiting period under the Hart-Scott-Rodino Antitrust Improvements Act 
of 1976, as amended, and other customary closing conditions. We expect our work to commence under the Neurocrine 
Collaboration in the first half of 2019. In February 2019, we announced the AbbVie Alpha-Synuclein Collaboration to 
develop and commercialize vectorized antibodies directed at pathological species of alpha-synuclein for the potential 
treatment of Parkinson’s disease and other diseases characterized by the abnormal accumulation of misfolded alpha-
synuclein protein, or synucleinopathies.  

6 

In addition to the programs described above, we continue to evaluate additional severe neurological diseases 
that could be treated using AAV gene therapy through application of either a gene replacement or a gene knockdown 
approach and are also actively exploring additional potential treatment methods that can utilize an AAV vector. 

Finally, we developed our own real-time, intra-operative, MRI compatible device, the Variable Trajectory 

Array Guide, or V-TAG™, that can be used with other neuro-navigational systems for the administration of drug to the 
putamen and other surgical procedures to avoid blood vessels and reduce the risk of potential hemorrhage during surgery 
and to maximize drug coverage of the putamen. In July 2018, the Center for Devices and Radiological Health, or the 
CDRH, of the FDA provided 510(k) clearance for V-TAG. We are currently working with a collaborator on process 
development and manufacturing of the device. Investigators have used an alternative MRI-compatible device called the 
ClearPoint® System in our Phase 1b clinical trial of VY-AADC and Phase 1 posterior trajectory trial. We expect to use 
both our own V-TAG and the ClearPoint System in our RESTORE-1 Phase 2 clinical trial. 

Sanofi Genzyme Collaboration 

In February 2015, we entered into a strategic collaboration with Sanofi Genzyme to leverage our combined 

expertise and assets to develop AAV gene therapies for certain severe neurological diseases. Under the agreement, we 
received $65.0 million in upfront cash, a $30.0 million upfront equity investment, and an in-kind commitment of $5.0 
million, totaling $100.0 million. At the inception of the agreement, we were eligible to receive up to $745.0 million in 
option and milestone payments while retaining U.S. commercial rights to most programs. Under the terms of the 
collaboration, we granted Sanofi Genzyme an exclusive option (i) to license, develop and commercialize ex-U.S. rights 
to the VY-AADC program, VY-FXN01, VY-HTT01, and a future program to be designated by Sanofi Genzyme, which 
we refer to collectively as the Split Territory Programs; (ii) to license, develop and commercialize worldwide rights to 
VY-SMN101; and (iii) to co-commercialize VY-HTT01 in the United States. Each of Sanofi Genzyme’s options to a 
Split Territory Program is triggered following the completion of the first proof-of-principle human clinical study, or POP 
Study, on a program-by-program basis. 

In October 2017, Sanofi Genzyme notified us that it had decided not to exercise its option for the ex-U.S. rights 

to VY-AADC for Parkinson’s disease. As a result, we are no longer entitled to receive $45.0 million and $60.0 million 
of regulatory and commercial milestone payments from Sanofi Genzyme, respectively, related to the Parkinson’s 
program under the Sanofi Genzyme Collaboration. If we use certain Sanofi Genzyme technology in VY-AADC, Sanofi 
Genzyme is entitled to receive low-single-digit royalty payments based on a percentage of net sales by us, and we may 
be obligated to make certain regulatory milestone payments to a third-party licensor.  

AbbVie Tau Collaboration 

In February 2018, we entered into an exclusive collaboration and option agreement with AbbVie for the 
research, development, and commercialization of AAV and other virus-based gene therapy products for the treatment of 
diseases of the central nervous system and other neurodegenerative diseases related to defective or excess aggregation of 
tau protein in the human brain, including Alzheimer’s disease. Under the terms of the agreement, we received an upfront 
payment of $69.0 million and may receive future option fees, development and regulatory milestone payments and 
royalties. Under the terms of the agreement, we will perform specified research, preclinical, and Phase 1 development 
activities regarding vectorized antibodies directed against tau, after which AbbVie may select one or more vectorized 
antibodies to proceed into IND-enabling studies and clinical development. We will be responsible for the research, IND-
enabling studies, and Phase 1 clinical trial activities and costs. Following the completion of Phase 1 clinical 
development, AbbVie has an option to license the vectorized tau antibody program and would then lead further clinical 
development and global commercialization for the product candidates pursuant to the agreement. We may earn up to 
$215.0 million in option exercise fees for such preclinical and Phase 1 development activities. In addition to the upfront 
payment and the potential option exercise payments, we are eligible to receive up to $895.0 million in development and 
regulatory milestones for each vectorized tau antibody compound. We are also eligible to receive tiered, escalating 
royalties in a range, subject to certain specified exceptions, from a high-single digit to a mid-to-high teen percentage of 
the global net sales of the vectorized antibodies for tauopathies, including Alzheimer’s disease and other 
neurodegenerative diseases. We also have an option to share in the costs of clinical development for higher royalty rates. 
Under the terms of the agreement, each party will own the entire right, title and interest in and to all know-how and 

7 

patent rights first made or invented solely by it or its affiliates or its or their sublicensees in the course of the 
collaboration, with certain specified exceptions. We have also agreed to grant AbbVie a worldwide license to certain 
know-how and patent rights developed by us or jointly by the parties arising from the collaboration.  

Neurocrine Collaboration 

In January 2019, we entered into the Neurocrine Collaboration Agreement for the research, development and 
commercialization of four programs including our Parkinson’s disease program, or AADC Program, our Friedreich’s 
ataxia program, or FA Program, and two programs, or the Discovery Programs. The Neurocrine Collaboration 
Agreement is not yet effective and remains subject to customary closing conditions, including certain antitrust approvals, 
that have not been satisfied as of the date of this Annual Report on Form 10-K. Under the terms of the agreement, we 
will receive an upfront payment of $165.0 million, inclusive of $50.0 million for the sale of 4,179,728 shares of our 
common stock, and may receive future development and regulatory milestone payments and royalties. We will use 
commercially reasonable efforts to develop the products in each of these programs. Neurocrine will be responsible for all 
costs incurred by us in conducting these activities for each program, in accordance with an agreed budget.  

Under the terms of the agreement for the AADC Program, Neurocrine will fund the clinical development of the 
RESTORE-1 Phase 2 clinical trials for VY-AADC. After the data readout of the RESTORE-1 Phase 2 trial, we have the 
option to either: (1) co-commercialize VY-AADC with Neurocrine in the U.S. under a 50/50 cost- and profit-sharing 
arrangement and receive milestones and royalties based on ex-U.S. sales, or (2) grant Neurocrine full global commercial 
rights in exchange for milestone payments and royalties based on global sales. Under the terms of the agreement for the 
FA Program, Neurocrine will fund the development through the Phase 1 clinical trial of VY-FXN01. After the data 
readout of the Phase 1 trial, we have the option to either: (1) co-commercialize VY-FXN01 with Neurocrine in the U.S. 
under a 60/40 cost- and profit-sharing arrangement, or (2) grant Neurocrine full worldwide commercial rights in 
exchange for milestone payments and royalties based on global sales, subject to Sanofi Genzyme’s option to 
commercialize the FA Program in countries outside the United States. Under the terms of the agreement for the two 
Discovery Programs, Neurocrine will fund the development of those programs and we have the right to earn milestone 
payments and royalties based on global sales. 

In addition to the upfront payment, we are eligible to receive aggregate development milestone payments under 

(i) the AADC Program of up to $170.0 million, (ii) the FA Program of up to $195.0 million, and (iii) each of the 
Discovery Programs of up to $130.0 million each. We may also be entitled to receive aggregate commercial milestone 
payments for each collaboration product of up to $275.0 million, subject to an aggregate cap on commercial milestone 
payments across all programs of $1.1 billion. We are also eligible to receive royalties, based on future net sales of the 
collaboration products. Such royalty percentages, for net sales in and outside the United States, as applicable, range (i) 
for the AADC Program, from the mid-teens to thirty and the low-teens to twenty, respectively; (ii) for the FA Program, 
from the low-teens to high-teens and high-single digits to mid-teens, respectively; and (iii) for each Discovery Program, 
from the high-single digits to mid-teens and mid-single digits to low-teens, respectively.  

AbbVie Alpha-Synuclein Collaboration 

In February 2019, we entered into an exclusive collaboration and option agreement with AbbVie for the 
development and commercialization of vectorized antibodies directed against pathological species of alpha-synuclein for 
the potential treatment of Parkinson’s disease and other diseases characterized by the abnormal accumulation of 
misfolded alpha-synuclein protein, or synucleinopathies. Under the terms of this agreement, we will receive and upfront 
payment of $65.0 million and may receive future option fees, development, regulatory, and commercial milestone 
payments, and royalties. Under the terms of the agreement, we and AbbVie have agreed to collaborate on the research 
and development of specified vectorized antibody compounds, or research compounds, comprised of an AAV or other 
viral capsid and a virus vector genome that encodes one or more antibodies that target and bind to the alpha-synuclein 
protein. The collaboration is comprised of a research period and a development period. We are obligated to conduct 
research activities directed to constructing one or more virus vectors that encode antibodies designated by AbbVie. We 
are obligated to use diligent efforts to conduct antibody engineering and other research activities to create the Research 
Compounds and to develop product candidates containing or comprised of such Research Compounds, which we refer to 
as Product Candidates. We are solely responsible for the costs and expenses during the research period. During a 

8 

specified portion of the Research Period, AbbVie may exercise one or more of its exclusive development options to 
select up to a total of four Research Compounds and their corresponding Product Candidates to proceed to the 
development period, after which AbbVie may exercise its option to license such Product Candidates following Phase 1 
results, for which we may earn up to $245.0 million in option exercise payments in aggregate. In addition to the upfront 
payment and the potential option exercise payments, we are eligible to receive up to $727.5 million in development and 
regulatory milestones for each Licensed Compound. We are also eligible to receive tiered, escalating royalties, in the 
mid-single-digit percentage range on aggregate net sales of Licensed Products on a Licensed Compound by Licensed 
Compound basis, as well as up to $500.0 million in commercial milestones based on aggregate annual net sales 
thresholds of Licensed Products. The royalties are subject to potential reductions for biosimilar market penetration, 
patent claim expiration, and other provisions, subject to specified limits. Subject to certain exceptions, each of AbbVie 
and the Company has agreed to be financially responsible for all payments owed to a third party with which it has 
contracted for any use of in-licensed intellectual property under the Collaboration Agreement. 

Mission and Strategy 

Our mission is to become the world leader in AAV gene therapy focused on treating severe neurological 

diseases by developing transformative therapies. Our strategy to achieve this mission is to:  

  Optimize and advance VY-AADC for the treatment of Parkinson’s disease. We continue to evaluate the 
dosing and delivery of VY-AADC to determine the optimal and safe dose to achieve meaningful clinical 
benefit for patients with Parkinson’s disease. The November 2018 interim results from the ongoing Phase 
1b clinical trial include data from all 15 patients treated in Cohorts 1, 2 and 3 (five patients in each Cohort) 
including data from patients in Cohort 1 at three years, Cohort 2 at two years and Cohort 3 at 18 months. 
The results continue to demonstrate durable improvements across multiple measures of patients’ motor 
function after a one-time administration of the gene therapy, as evidenced by the patients’ diaries, the 
Unified Parkinson’s Disease Rating Scale, or UPDRS, UPDRS-II and UPDRS-III, and quality of life 
assessments. The average measure of motor function by UPDRS, or UPDRS-III, on medication score was 
13.5 and UPDRS-III off medication score was 37.1, the average measure of activities of daily living by 
UPDRS, or UPDRS-II. The update of results from the ongoing Phase 1b clinical trial of VY-AADC include 
a 2.1-hour mean improvement in good ON time from baseline to three years for patients in Cohort 1, a 2.7-
hour mean improvement from baseline to 2 years in Cohort 2, and a mean improvement of 1.7 hours from 
baseline to eighteen months in Cohort 3. 

For the RESTORE-1 Phase 2 clinical trial, we have selected a dose of up to 2.5 x 1012 vector genomes, 
which is defined as a maximum total bilateral dose. This dose is between the up-to-maximum total vector 
genome doses administered in Cohorts 2 and 3 from the Phase 1b trial when considering the higher volume 
administered with the posterior trajectory and vector produced using the baculovirus system. Having 
selected a dose for the RESTORE-1 Phase 2 trial between the two highest dose Cohorts from the Phase 1b 
trial, we have performed a combined analysis of the outcomes from the ten patients in Cohorts 2 and 3. 
Results from the combined ten patients in Cohorts 2 and 3 demonstrated mean increases from baseline in 
good ON time of 2.4 hours per day at 12 months, the timepoint for the primary endpoint in the RESTORE-
1 Phase 2 trial, and 2.6 hours per day at 18 months, the latest timepoint measured for both Cohorts. Of the 
combined ten patients in Cohorts 2 and 3, seven patients would have met the eligibility criteria for the 
RESTORE-1 Phase 2 trial based on limits in severity of dyskinesia and minimum OFF time at baseline. For 
these seven patients, the RESTORE-1 Phase 2 trial relevant group, the mean improvements in good ON 
time were 2.8 hours at 12 months and 2.5 hours at 18 months. These results were achieved with clinically 
meaningful and sustained reductions in daily oral levodopa and related medications. In December 2018, we 
randomized the first patient in the RESTORE-1 Phase 2 clinical trial.  

  Build and advance a pipeline of gene therapy programs focused on severe neurological diseases. Beyond 
our clinical-stage program for Parkinson’s disease, we have a deep pipeline of AAV gene therapy programs 
in various stages of preclinical development. We plan to file two additional INDs for our preclinical 

9 

programs in 2019. We believe that our leadership position in AAV gene therapy for severe neurological 
diseases and our gene therapy platform provide us with the necessary capabilities to evaluate and capitalize 
on external opportunities. As such, we plan to opportunistically expand our pipeline through acquisition, in-
licensing or other strategic transactions. 

  Continuously invest in our AAV gene therapy platform. We plan to continuously invest in our gene 

therapy platform to maintain our leadership in AAV gene therapy for neurological diseases. Specifically, 
we intend to further develop and enhance our gene therapy platform by focusing on (i) vector engineering 
and optimization; (ii) manufacturing; and (iii) dosing and delivery techniques. We plan to continue 
generating novel AAV vectors by engineering and optimizing vectors best suited to a targeted disease. We 
have built an onsite, state-of-the-art process research and development facility to enable the manufacturing 
of high quality AAV gene therapy vectors at laboratory scale. We expect to utilize established and novel 
techniques for dosing and delivery of our AAV gene therapies to the central nervous system, or CNS.  

  Establish a leadership position in commercial-scale, high quality AAV manufacturing. We believe that 
manufacturing capacity and expertise are critical to successfully treating patients using gene therapy. We 
have established relationships with multiple current good manufacturing practices, or cGMP, contract 
manufacturers. Previously, through one of our collaborations, with MassBiologics, an FDA-licensed 
manufacturer affiliated with the University of Massachusetts Medical School, we initiated cGMP 
production activities. More recently, we announced additional agreements with Brammer Bio and with 
Fujifilm Diosynth Biotechnologies, established contract manufacturers that specialize in gene therapy and 
AAV vectors. We are using the baculovirus/Sf9 AAV production system, a technology for producing AAV 
vectors at scale in insect-derived cells, originally invented and developed by several current and former 
members of our production team while at the National Institutes of Health, or NIH, which we continue to 
improve upon. We believe that having oversight through these key relationships over our own commercial 
manufacturing process is critical to ensuring quality product with commercial yields.  

  Retain commercialization rights to our programs. We hold worldwide rights for our ALS, and severe, 
chronic pain programs. We have retained co-development and co-commercialization rights for our 
Parkinson’s disease and Friedreich’s ataxia programs under our Neurocrine Collaboration and for our 
Huntington’s disease program under our Sanofi Genzyme Collaboration, respectively. As these and other 
programs advance through late-stage clinical development, we intend to build our own sales and marketing 
infrastructure and leverage our partnerships to support our programs where we have retained 
commercialization rights. These collaborations also represent an important advance in our strategy to 
leverage our AAV gene therapy platform and programs through collaborative partnerships with 
biopharmaceutical companies that bring complementary expertise, capabilities, and experience, in addition 
to capital. 

  Expand our intellectual property portfolio. We seek to have an industry leading intellectual property 
portfolio. To that end, we seek patent rights for various aspects of our programs, including vector 
engineering and construct design, our production process, and all features of our clinical products including 
compositions and methods of delivery. We expect to continue to expand our intellectual property portfolio 
by aggressively seeking patent rights for promising aspects of our gene therapy platform and product 
candidates.  

AAV Gene Therapy for Neurological Diseases  

Gene therapy is an approach whereby gene expression is directly altered in patients to address the underlying 

cause or predominant manifestations of disease. We believe that the targeted nature of gene therapy may enable powerful 
treatment options and provide these patients with meaningful and durable benefits.  

10 

 
While AAV gene therapy can potentially be harnessed for multiple treatment methods, we are currently focused 

on gene replacement and gene knockdown approaches. Gene replacement is intended to restore the expression of a 
protein that is not expressed, expressed at abnormally low levels or functionally mutated with loss of function. Gene 
knockdown, or gene silencing, is intended to reduce the expression of a pathologically mutated protein that has 
detrimental effects.  

Our gene therapy approach uses AAV vectors which we believe are ideal vectors for gene therapy for several 

reasons:  

Broad Applicability. AAV is able to transduce, or transfer a therapeutic gene, into numerous cell types 
including target cells in the CNS.  

Safety. AAV is believed to be safe and is not known to cause any disease in humans. No vector-related serious 
adverse effects, or SAEs, have been reported in the more than 1,500 patients, including over 200 patients for 
neurological indications, treated with AAV gene therapy to date.  

Does Not Readily Integrate. AAV does not readily integrate into the genome of the target cell, reducing the 
potential for oncogenesis, or the induction of cancer. 

Scalability. AAV is able to be manufactured at commercial quality and scale.  

We believe that neurological diseases are well-suited for treatment with AAV gene therapy for the following 

reasons:  

Validated Targets. Many neurological diseases are caused by well-defined mutations in genes and these genes 
represent genetically validated drug targets for AAV gene therapy.  

Targeted Delivery. Advances in delivery techniques allow for direct delivery of AAV vectors to discrete 
regions in the brain or broader delivery throughout the spinal cord via the cerebrospinal fluid, or CSF.  

Durable Expression. Long-term gene expression may be achievable in the CNS following one-time dosing and 
transfer of the therapeutic gene with an AAV vector. Neurons in the CNS are terminally differentiated, or no 
longer divide, eliminating the potential for cell division to dilute expression of the therapeutic gene. Repeated or 
continual dosing with direct injection of drugs into the CNS is complex, therefore a one-time AAV gene therapy 
has significant advantages.  

Immune Privileged Site. There is a reduced risk of harmful immune response or reduced efficacy due to 
localized delivery in a self-contained system.  

While we are currently focused on gene replacement and gene knockdown approaches, we are also actively 

exploring additional potential treatment methods that can utilize an AAV vector, including the direct delivery of 
monoclonal antibodies to the CNS (such as in our collaborations with AbbVie), as well as gene editing to correct or 
delete a gene in the cell genome.  

The Voyager Gene Therapy Platform  

We have built a gene therapy platform that we believe positions us to be the leading company at the intersection 

of AAV gene therapy and severe neurological diseases. Our team of experts in the fields of AAV gene therapy and 
neuroscience first identifies and selects severe neurological diseases that are well-suited for treatment using AAV gene 
therapy. We then engineer and optimize AAV vectors for delivery of the virus payload to the targeted tissue or cells. 
Finally, we leverage established routes of administration and advances in dosing techniques to optimize delivery of our 
AAV gene therapies to target cells that are critical to the disease of interest either directly to discrete regions of the brain, 

11 

 
or, more broadly, to the spinal cord region. We believe that optimizing each of these parameters is a key factor for 
overall program success. We expect that our current and future pipeline programs will make use of technological 
advances generated with our gene therapy platform.  

Disease Selection  

We assess potential product programs based upon the following criteria:  

Unmet Need. There is a significant unmet medical need for the indication and substantial commercial potential.  

Target Validation. There is strong evidence that expression of a specific gene, or lack thereof, is causing, or 
critical to, the disease state.  

Delivery Using AAV. There is strong evidence supporting the ability to target the relevant tissue and cells 
using an AAV vector to achieve sufficient target gene expression.  

Clinical Readouts. The clinical impact of an AAV gene therapy can be clearly measured, including through 
well-accepted clinical endpoints and the use of both existing and novel biomarkers.  

Scalability of Manufacturing. Sufficient AAV vector to supply late-stage clinical development and 
commercialization can be manufactured.  

In addition to the criteria above, we also look for groups of diseases where our knowledge can be transferred. 

For instance, we believe that some of the delivery parameters and imaging techniques that are employed in our VY-
AADC program can be applied to AAV gene therapy delivery for Huntington’s disease or other diseases where direct, 
targeted delivery to the brain is warranted.  

Vector Engineering and Optimization  

We have advanced or intend to advance our multiple preclinical programs towards selection of lead clinical 

candidates using AAV vectors that we believe are best suited for each of our programs either through use of our existing 
capsids, through exercising a non-exclusive worldwide commercial license to capsid sequences covered by third parties, 
or by engineering or optimizing novel capsids. The key components of an AAV vector include: (i) the capsid; (ii) the 
therapeutic gene, or transgene; and (iii) the promoter, or the DNA sequence that drives the expression of the transgene.  

Members of our team have co-discovered many of the known naturally occurring AAV capsids and have also 

created promising genetically engineered AAV capsids. Genetically engineered capsids have yielded vectors with 
desirable properties, such as higher biological potency and enhanced tissue specificity. We believe that there is an 
opportunity to further optimize AAV capsids to confer desired characteristics relating to properties such as tissue 
specificity and immunogenicity. We have a significant effort dedicated to designing and screening for novel AAV 
capsids using a number of different scientific approaches. We believe that the information generated by this work will 
enhance our ability to rationally design AAV capsids with specific properties for particular therapeutic applications. In 
September 2016, we announced a co-exclusive worldwide license agreement with the California Institute of Technology, 
or Caltech, related to novel AAV capsids. The license agreement covers all fields of use and includes novel AAV capsids 
that have demonstrated enhanced blood-brain barrier penetration for the potential treatment of neurological diseases 
following systemic administration of an AAV gene therapy vector.  

With respect to the target DNA delivered through AAV gene therapy, we are selecting promoters that we 

believe have the appropriate activity and tissue, selectively for our specific gene therapy programs. We are also 
designing transgenes to provide optimal expression once delivered to the targeted cells.  

Manufacturing at Commercial Quality and Scale  

12 

The ability to produce high quality AAV vectors at commercial-scale is a critical success factor in AAV gene 

therapy. While at the NIH, former members of our production team invented and developed a baculovirus/Sf9 AAV 
production system, which we use and have continued to improve. This system has a number of attributes that we believe 
will enable high quality commercial-scale manufacturing, including:  

High Yield. A single manufacturing run at 500-liter scale can yield many thousands of doses of an AAV gene 
therapy.  

High Purity. A relatively high percentage of AAV vectors contain the therapeutic DNA, reducing the number 
of empty capsids compared to alternative manufacturing approaches. In addition, the baculovirus/Sf9 system 
eliminates the risk of introducing mammalian cell derived impurities.  

Scalability. This process has been reproduced at volumes ranging from 0.02 liters to 250 liters. We believe the 
existing process is scalable to substantially higher volumes.  

We have built a state-of-the-art process research and development production facility for manufacturing 

research-grade AAV vectors onsite at our Cambridge, Massachusetts headquarters. We have also established multiple 
contract manufacturing relationships with companies specializing in the manufacture of gene therapy and AAV vectors. 

Optimized Delivery and Route of Administration  

Identifying the optimal route of administration and delivery parameters for AAV gene therapy, such as infusion 
volume, flow rate, vector concentration and dose and formulation for a specific disease, are critical to achieving safe and 
effective levels of transgene expression in the targeted location in the CNS. We aim to develop clinically feasible 
protocols that yield reproducible results across patients. For our Parkinson’s disease and Huntington’s disease programs, 
we are pursuing direct injection into the brain, called intraparenchymal injection. For our ALS SOD1 and Friedreich’s 
ataxia programs, we are evaluating multiple routes of administration including injection into the CSF within the 
cerebrospinal space, called intrathecal injection, as well as intravenous injection, intraparenchymal injection, and other 
delivery alternatives. 

VTAGTM Intraparenchymal Injection to the Brain  

The surgical approach that we are using for VY-AADC is similar, in some respects, to the stereotactic approach 

used for deep brain stimulation, or DBS, a marketed device-based treatment for Parkinson’s disease. One primary 
difference with our approach is the ability to assist the physician in visualizing the delivery of VY-AADC to the putamen 
using real-time, intra-operative, magnetic resonance imaging, or MRI, to avoid specific blood vessels to reduce the risk 
of potential hemorrhages during the surgical procedure and to maximize the coverage of the putamen.  

Investigators in the Phase 1b clinical trial of VY-AADC and the separate Phase 1 posterior trajectory trial use 

the real-time, intra-operative, MRI system called the ClearPoint System® from MRI Interventions, Inc. However, not all 
neuro-surgical units within the United States utilize this system and may employ other neuro-navigational systems that 
are not compatible with real-time MRI imaging.  

Consequently, we developed the Voyager Trajectory Array Guide, or V-TAG™, as our own device for use as a 

real-time, intra-operative, MRI-compatible device that can be used with other neuro-navigational systems for this and 
other surgical procedures. We received 510(k) clearance from the FDA in July 2018. We are currently working with a 
collaborator on process development and manufacturing of the device. We believe that our experience gained from our 
VY-AADC program, including the use of V-TAG, can be applied to AAV gene therapy delivery for our Huntington’s 
disease program and possibly other projects as well.  

13 

Overview of Intraparenchymal Delivery  

Overview of Our Pipeline  

Courtesy of: Okinawa Institute of Science and Technology. 

We have leveraged our gene therapy platform to assemble a pipeline of novel AAV gene therapies for the 

treatment of severe neurological diseases with high unmet medical need. Depending on the disease, our current AAV 
gene therapies will use either a gene replacement or gene knockdown approach. Our goal is to address the underlying 
cause or the predominant manifestations of a specific disease by significantly increasing or decreasing expression of the 
relevant proteins at targeted sites within the CNS.  

Parkinson’s Disease Program: VY-AADC 

Disease and VY-AADC Overview  

Parkinson’s disease is a chronic, progressive and debilitating neurodegenerative disease that affects 

approximately 1 million people in the United States and 6 million people worldwide. Parkinson’s disease is characterized 
by a loss of dopamine and its function. Dopamine is a chemical “messenger” that is produced in the brain and is involved 
in the control of movement. Some chemicals, like dopamine, are made from other chemicals by proteins called enzymes. 
Dopamine is made in the brain when the enzyme AADC (aromatic l-amino acid decarboxylase) converts the chemical 
levodopa to dopamine. Levodopa, AADC, and dopamine are each present at normal levels in healthy people. 

When dopamine levels decrease in the brain and there is no longer enough to control movement, the motor 

symptoms of Parkinson’s disease including tremors, slow movement or loss of movement, rigidity, and postural 
instability, may occur. When this happens, a doctor may prescribe a levodopa medication, which is converted into 
dopamine by AADC in substantially the same way that naturally occurring levodopa is converted to dopamine. 

As Parkinson’s disease worsens, there is less AADC enzyme in parts of the brain where it is needed to convert 
levodopa to dopamine. Therefore, the amount of dopamine that is produced from each dose of levodopa medicine may 
be reduced. When this happens, patients’ motor function may worsen and a less predictable response to medications may 
occur. 

Our investigational gene therapy VY-AADC is designed to put the AADC enzyme into brain cells where it can 

convert levodopa to dopamine. To do this, the AADC gene is delivered inside a transporter called “adeno-associated 

14 

 
 
viral vector,” which we refer to as AAV, much like a letter that carries the instructions the brain needs to make the 
AADC enzyme with the AAV as the envelope that carries the letter. 

Preclinical Studies  

Preclinical studies conducted by Krystof Bankiewicz, M.D., Ph.D., one of our co-founders, and his colleagues 

at the University of California San Francisco, or UCSF, evaluated the safety, efficacy and pharmacological activity of 
AAV2-AADC gene therapy, a gene therapy substantially similar to VY-AADC, delivered directly to the putamen in a 
non-human primate model of Parkinson’s disease. Overall, the procedure and vector were well-tolerated with no serious 
toxicity issues.  

Positron emission tomography, or PET, imaging with tracers specific for AADC enzyme activity demonstrated 

a significant and sustained increase of activity in the brain region where the vector had been delivered. Increased 
responsiveness to levodopa was also evidenced by significant behavioral improvements observed post-treatment with the 
gene therapy compared to pre-treatment. In five animals, the mean improvement in behavior was determined at various 
doses of levodopa both one month before treatment, as a baseline measure for comparison purposes, and then again six 
months after treatment. A strong PET signal was observed in all five animals following treatment, confirming delivery of 
AADC into the putamen. Animals were significantly more sensitive to levodopa six months following treatment with the 
gene therapy when compared to baseline, as shown below.  

Behavioral Response to Various Doses of levodopa Pre- and Post-Treatment with AAV2-AADC in Non-Human 
Primates(1)  

(1)  Adapted by permission from Macmillan Publishers Ltd; Forsayeth et al, Molecular Therapy (2006), 14 (4); 571-577, copyright 2006. Blue line 

represents base line measurements and yellow line represents six months post-treatment measurements.  

* 

   A result is considered to be statistically significant when the probability of the result occurring by random chance, rather than from the efficacy of the 

treatment, is sufficiently low. The conventional method for measuring the statistical significance of a result is known as the “p-value,” which 
represents the probability that chance caused the result (e.g., a p-value = 0.001 means that there is a 0.1% or less probability that the difference 
between the control group and the treatment group is purely due to random chance).  

The results of these preclinical studies provided support for the initiation of clinical trials.  

15 

 
 
Previous Phase 1 Clinical Trials 

The UPDRS is a standard and widely used four-part clinical rating scale for Parkinson’s disease that evaluates 
cognitive, functional, and motor deficits, as well as medication-related complications. UPDRS Part III measures motor 
function by physician examination. The UPDRS is conducted when patients are taking their Parkinson’s disease 
medications (referred to as “on” medication) and when patients are not taking their Parkinson’s disease medications 
(referred to as “off” medication). In addition, a patient-completed Hauser diary records the patient’s motor response over 
the course of several days as ON time when they have good mobility with or without non-troublesome dyskinesia, or 
uncontrolled, involuntary movement; OFF time when they have poor mobility; and ON time with troublesome 
dyskinesia when they have uncontrolled movements. As shown in the figure below, diary ON time decreases, while OFF 
time and dyskinesias increase as patients progress from the early honeymoon period into later stages of Parkinson’s 
disease.  

Overview of Progression of Parkinson’s Disease (PD)  

In a completed open-label Phase 1 clinical trial conducted at UCSF, VY-AADC was delivered directly to the 
putamen of Parkinson’s disease patients. The primary endpoints of this trial were safety and tolerability of VY-AADC. 
These endpoints were met as VY-AADC was well-tolerated and no treatment related SAEs were reported. Furthermore, 
pharmacologic activity of VY-AADC was observed. This trial was completed prior to our involvement in the program.  

The Phase 1 clinical trial at UCSF was conducted in a total of 10 patients with Parkinson’s disease. Two doses 

of VY-AADC were tested, 9×1010 vector genomes, or vg, and 3×1011 vg, with five patients per dose Cohort. The 
infusion volume was 100µl per putamen, or 200µl per patient. Patients in both Cohorts treated with VY-AADC showed 
modest improvements in motor fluctuations. At six months following treatment, diary OFF time was observed to be 
reduced by an average of approximately three hours and a corresponding increase in diary ON time without dyskinesias 
was also observed. In addition, at six months following treatment, an approximately 30% improvement in UPDRS total 
score both on-medication and off-medication UPDRS Total score, was observed, as shown in the table below. 

16 

 
Summary of UPDRS Results from Phase 1 Trial(1)  

(1)  Christine et al, Neurology (2009), 73: 1662-1669. The row titled “Low-dose Cohort” represents data from the five patients treated with 9 X 1010 vg of 
VY-AADC01. The row titled “High-dose Cohort” represents data from the five patients treated with 3 X 1011 vg of VY-AADC01. The row titled 
“Combined Cohorts” represents data from all ten patients treated with VY-AADC01. The data in the columns under the header “Off medications” 
represents periods during which patients’ medications were not working as measured by a patient’s total UPDRS score at baseline, before treatment 
with VY-AADC01, and at six months following treatment with VY-AADC01, along with percent change from baseline to six months and the 
corresponding p-value. The data in the columns under the header “On medications” represents periods during which patients’ medications were 
working as measured by a patient’s total UPDRS score at baseline, before treatment with VY-AADC01 and at six months following treatment with 
VY-AADC01, along with percent change from baseline to six months and the corresponding p-value. A result is considered to be statistically 
significant when the probability of the result occurring by random chance, rather than from the efficacy of the treatment, is sufficiently low. The 
conventional method for measuring the statistical significance of a result is known as the “p-value,” which represents the probability that chance 
caused the result (e.g., a p-value = 0.001 means that there is a 0.1% or less probability that the difference between the control group and the treatment 
group is purely due to random chance). Because of the small size of this trial, the p-values may not be reliable or repeatable, and may not be 
duplicated in future trials. 

While no gene therapy related SAEs were reported, three patients experienced minor hemorrhages related to the 

surgical procedure. Two of the hemorrhages were asymptomatic, noticed only on imaging, and one was symptomatic 
with the patient making an almost complete recovery. Nevertheless, the stereotactic injection protocol used in the 
surgical procedure was modified to avoid specific blood vessels and no further hemorrhages were reported. In our 
ongoing Phase 1b clinical trial, we implemented the use of real-time, intra-operative MRI guidance. The use of this intra-
operative MRI guidance is a significant advancement in vector delivery.  

The 10 patients in this Phase 1 clinical trial were followed for up to four years after treatment, and a durable, 

dose-dependent expression of AADC enzyme activity was observed. Patients treated with both doses of the gene therapy 
had an increased PET signal, or uptake of the [18F]fluoro-L-m-tyrosine tracer indicative of AADC enzyme activity that 
persisted for up to four years. Patients treated with the high dose gene therapy had a greater PET signal on average when 
compared to the low dose Cohort. 

17 

 
 
Long-Term AADC Expression as Measured by PET Imaging in Patients Treated with High and Low Doses of 
AAV Gene Therapy in a Previous Phase 1 Clinical Trial (1) 

(1)  Mittermeyer et al, Human Gene Therapy (2012), 23: 377-381. The publisher for this copyrighted material is Mary Ann Liebert, Inc. publishers. Blue 

lines represent patients treated with the low dose and yellow lines represent patients treated with the high dose. 

A similar Phase 1 clinical trial was conducted at Jichi Medical University, or JMU, in Japan using the same 

vector that was used in the UCSF trial. The primary endpoint of this trial was safety of the treatment. This endpoint was 
met as the treatment was well-tolerated and no treatment related SAEs were reported. Six patients were treated in this 
trial and an enhanced PET signal was observed in a subset of patients monitored 96 weeks following treatment. A 
second, open-label Phase 1/2 trial is currently being conducted at JMU. The primary endpoint of this trial is also safety. 
This trial is using lower infusion volumes and total doses compared to our ongoing Phase 1b and Phase 2 clinical trials. 
Importantly, the JMU trial is not using real-time, intra-operative MRI guidance.  

While the prior UCSF and JMU clinical results were encouraging and provided evidence of long-term AADC 
enzyme expression, the magnitude of the clinical benefits observed did not exceed placebo effects observed in previous 
surgical therapy trials in Parkinson’s disease patients, and the UCSF and JMU trials were not blinded. Further, based on 
post-operative imaging and our current work using real-time, intra-operative MRI monitoring, we estimate that less than 
10% of the putamen volume was covered by the infusion in these trials, which reflects suboptimal distribution of the 
gene therapy vector in the putamen. We believe that by further optimizing the delivery, dose and infusion volume to 
substantially increase the coverage of the putamen, a more substantial clinical benefit can be achieved.  

Voyager VY-AADC Phase 1b Trial  

In 2014, UCSF initiated an open-label Phase 1b clinical trial to optimize the development of VY-AADC. The 

IND for the Phase 1b trial was filed by UCSF in July 2013 and was transferred to us in October 2015. In November 
2017, we completed enrolling this open-label, dose-escalating Phase 1b trial of VY-AADC. The trial included 15 
patients with Parkinson’s disease and was designed to evaluate the safety and efficacy of escalating doses of VY-AADC. 
In this trial, one-time administration of VY-AADC led to improvements in patients’ motor function, and patients were 
able to reduce their daily levodopa and other Parkinson’s disease medications. To date, administration of VY-AADC has 
been well-tolerated. In patients treated in this trial, there have been no vector-related serious adverse events reported.  

18 

 
 
Patients in three Cohorts of five patients each were treated with a single administration of ascending doses of 

VY-AADC administered under MRI guidance to the putamen, a region of the brain associated with impaired motor 
function in Parkinson’s disease. The primary endpoints of this trial are safety and tolerability of the treatment. This trial 
incorporates three key design features:  

  Use of real-time, intra-operative MRI system during surgery to assist the physician in visualizing the 

delivery of VY-AADC to the putamen and to avoid specific blood vessels during the surgical procedure, 
with the goal of reducing the risk of hemorrhages.  

  Larger infusion volumes designed to increase coverage of the putamen with VY-AADC.  

  Higher concentrations of VY-AADC vector compared to the previously completed UCSF Phase 1 trial.  

Secondary endpoints of this trial, which are being used to assess the potential pharmacologic activity of VY-
AADC, include UPDRS, AADC PET imaging, quality of life, a patient-completed Hauser diary monitoring good ON 
time without troublesome dyskinesia, and a behavioral test using intravenous levodopa treatment to measure changes in a 
patients’ sensitivity to levodopa as well as endpoints to measure motor functions.  

In November 2018, we updated interim results from the ongoing, open-label Phase 1b clinical trial of VY-

AADC for the treatment of Parkinson’s disease. Interim results include data from all 15 patients treated in Cohorts 1, 2 
and 3 (five patients in each Cohort) including data from patients in Cohort 1 at three years, Cohort 2 at two years and 
Cohort 3 at 18 months. Cohort 1 patients received a single administration of VY-AADC at a concentration of 8.3×1011 
vg per milliliter, or vg/ml, using an infusion volume of up to 450 µL per putamen, or up to 900 µL per patient, for a total 
dose of 7.5×1011 vg. Cohort 2 patients received a single administration of VY-AADC at a concentration of 8.3×1011 
vg/ml, using an infusion volume of up to 900 µL per putamen, or up to 1,800 µL per patient, for a total dose of 1.5×1012 
vg. Cohort 3 patients received a three-fold higher vg concentration of 2.6×1012 with the same infusion volumes of VY-
AADC similar to those received by Cohort 2 patients (up to 900 µL per putamen), for a total dose of up to 4.5×1012 vg. 

Administration of VY-AADC has been well-tolerated in all fifteen patients treated in the three Cohorts with no 

reported vector-related SAEs. Fourteen of the 15 patients were discharged from the hospital within two days following 
surgery. As previously reported, one patient experienced two SAEs: a pulmonary embolism or blood clot in the lungs, 
and related heart arrhythmia or irregular heartbeat. Investigators determined that these SAEs were most likely related to 
immobility during the administration of the product; consequently, deep vein thrombosis prophylaxis has been added to 
the clinical trial protocol. 

Key findings from this trial to date include: 

  The use of real-time, MRI-guided delivery and increasing infusion volumes resulted in progressively 

greater coverage of the putamen, 21% mean coverage of the volume of the putamen with VY-AADC in 
Cohort 1, 34% mean coverage in Cohort 2, and 42% mean coverage in Cohort 3. 

  VY-AADC treatment resulted in a 13% increase, a 56% increase, and a 79% increase in mean putaminal 
AADC enzyme activity in Cohort 1, 2, and 3, respectively, at six months relative to baseline as measured 
by 18F-Dopa PET scan. Coverage of the putamen and AADC enzyme activity were highly correlated 
(r=0.84, p=0.0002). 

  One-time administration of VY-AADC resulted in reduced daily doses of oral levodopa and related 

medications. Six months after VY-AADC administration, patients in Cohort 1 had a reduction in levodopa 
equivalent daily dose, or LED, of 15%, Cohort 2 had a LED reduction of 33%, and Cohort 3 had a LED 
reduction of 42%. LED reductions were maintained through last follow up in Cohort 2 (24 months) and 
Cohort 3 (18 months). 

Patients enrolled in Cohorts 1, 2 and 3 were: 

19 

  On average, 58 years of age with a Parkinson’s disease diagnosis for an average of 10 years. 

  Candidates for surgical intervention including deep-brain stimulation due to disabling motor complications 

despite treatment with optimal anti-Parkinsonian medication. 

  At baseline, the average patient diary ON time without troublesome dyskinesia was 10.5 hours and, average 

diary OFF time was 4.6 hours; both diary measures were normalized to a 16-hour waking day. 

  Average UPDRS-III (motor function) on medication score was 13.5 and UPDRS-III off-medication score 
was 37.1; average UPDRS-II (activities of daily living) on medication score was 3.9 and UPDRS-II off 
medication score was 16.5. Patients in Cohort 3 entered the trial with more severe dyskinesia at baseline 
than patients in Cohorts 1 and 2 based on the Unified Dyskinesia Rating Scale, with a mean score of 30.2 
for Cohort 3 compared with a mean score of 19.2 and 17.4 for Cohorts 1 and 2, respectively. 

  At baseline, patients were treated with optimal levels of multiple dopaminergic medications including, in 
many cases, amantadine for the treatment of dyskinesia, or uncontrolled or involuntary movements. 
Patients’ average amount of Parkinson’s disease medications at baseline was 1,526 mg of oral LEDs per 
day.  

The results continue to demonstrate durable improvements across multiple measures of patients’ motor function 

after a one-time administration of the gene therapy, as evidenced by the patients’ diaries and UPDRS-III, and quality of 
life assessments. The update of results from the ongoing Phase 1b trial of VY-AADC include a 2.1-hour mean 
improvement in good ON time from baseline to three years for patients in Cohort 1, a 2.7-hour mean improvement from 
baseline to two years in Cohort 2, and a mean improvement of 1.7 hours from baseline to 18 months in Cohort 3 as 
shown in the table below. 

For the RESTORE-1 Phase 2 clinical trial, we have selected a dose of up to 2.5 x 1012 vg, which is defined as a 

maximum total bilateral dose. This dose is between the up-to-maximum total vector genome doses administered in 
Cohorts 2 and 3 from the Phase 1b trial when considering the higher volume administered with the posterior trajectory 
and vector produced using the baculovirus system. Having selected a dose for the RESTORE-1 Phase 2 trial between the 
two highest dose Cohorts from the Phase 1b trial, we have performed a combined analysis of the outcomes from ten 
patients in Cohorts 2 and 3. Results from the combined ten patients in Cohorts 2 and 3 demonstrated mean increases 
from baseline in good ON time of 2.4 hours per day at 12 months, the timepoint for the primary endpoint in the 
RESTORE-1 Phase 2 trial, and 2.6 hours per day at 18 months, the latest timepoint measured for both Cohorts as shown 
in the table below. Of the combined ten patients in Cohorts 2 and 3, seven patients would have met the eligibility criteria 
for the RESTORE-1 Phase 2 trial based on limits in severity of dyskinesia and minimum OFF time at baseline. For these 
seven patients, the RESTORE-1 Phase 2 trial relevant group, the mean improvements in good ON time were 2.8 hours at 
12 months and 2.5 hours at 18 months. These results were achieved with clinically meaningful and sustained reductions 
in daily oral levodopa and related medications. 

20 

 
Voyager Phase 1 Posterior Trajectory Clinical Trial 

In a separate Phase 1 clinical trial, we have changed the trajectory for administration of VY-AADC to a 
posterior, or back of the head, approach into the putamen, compared to a transfrontal, or top of the head, delivery 
approach used in Cohorts 1 through 3 of the ongoing Phase 1b clinical trial described above. A posterior approach better 
aligns the infusion of VY-AADC with the anatomical structure of the putamen, which reduces the number of trajectories 
needed and potentially reduces the total procedure time and increases the total coverage of the putamen. Administration 
of VY-AADC with this posterior approach has been well-tolerated in the eight patients treated with no reported SAEs. 
Most patients were discharged from the hospital the day after surgery. This trial utilized the same dose concentration as 
Cohort 3 of our Phase 1b clinical trial at a higher volume, yielding a total dose of up to 9.0×1012 vg compared with a 
total dose of up to 4.5×1012 vg in Cohort 3. As we announced in July 2018, interim results suggest that the posterior 
approach is associated with greater average putaminal coverage (approximately 50%), reduced total procedure times 
compared with the transfrontal approach by two to three hours, and improvements in patients’ motor function at six 
months, which were consistent with improvements achieved from patients in Cohorts 2 and 3 at the same time point in 
our Phase 1b trial with VY-AADC. We have determined that the posterior approach is the preferred surgical route of 
administration for the RESTORE-1 Phase 2 clinical trial. 

We continue to follow patients from Cohorts 1 through 3 in the Phase 1b clinical trial of VY-AADC and 

patients in the Phase 1 posterior trajectory trial, and plan to report updated results from these trials from time to time. 

Voyager VY-AADC RESTORE-1 Phase 2 and RESTORE-2 Phase 3 Clinical Trials  

In December 2017, we submitted an IND for VY-AADC which has become effective. As part of this IND, the 

chemistry, manufacturing, and controls section included data demonstrating comparability between VY-AADC using our 
baculovirus/Sf9 manufacturing process and VY-AADC produced using a mammalian cell system consisting of triple-
transfection of HEK293 cells, which was used in our two Phase 1 clinical trials. Both were produced under good 
manufacturing practice or GMP. Our baculovirus/Sf9 manufacturing process is designed for production of AAV vectors 
at clinical and commercial scale, with the potential for increased yields and efficient scalability compared with 
mammalian-based systems. We have demonstrated that this production platform change resulted in comparable vector 
quality and activity. We are using VY-AADC manufactured in our baculovirus/Sf9 system in our global RESTORE-1 
Phase 2 clinical trial and the planned RESTORE-2 Phase 3 clinical trial. In June 2018, the FDA granted RMAT 
designation for our VY-AADC gene therapy treatment, which provides for an enhanced level of interactions between the 
company sponsor and the FDA throughout the development program. The designation was based on our Phase 1b 
clinical data with VY-AADC. Previously, the FDA also granted fast-track designation for VY-AADC.  

In December 2018, we announced randomization of the first patient in RESTORE-1 Phase 2, randomized, 

double-blind, placebo-surgery controlled trial evaluating the safety and our efficacy of VY-AADC for the treatment of 
Parkinson’s disease in patients with motor fluctuations that are refractory to medical management. The RESTORE-1 
Phase 2 trial will enroll patients who have been diagnosed with Parkinson’s disease for at least four years, are not 
responding adequately to oral medications, and have at least three or more hours of OFF time during the day as measured 
by a validated self-reported patient diary. Patients who meet the eligibility criteria will be randomized (1:1) to one-time 

21 

 
administration of VY-AADC (for a total dose of up to 2.5×1012 vg) or placebo surgery. In December 2018, we 
randomized the first patient in the RESTORE-1 Phase 2 clinical trial.  

The primary endpoint of RESTORE-1 is good ON time, as measured by a validated self-reported patient diary 

at 12 months. Secondary endpoints include diary OFF time, other motor function UPDRS-II and UPDRS-III scores, 
assessments from the Parkinson’s Disease Questionnaire (PDQ-39), and patient’s global function as measured by the 
proportion of participants with improvement on the Clinical Global Impression, or CGI, score. The trial will also 
measure non-motor symptoms from the Non-Motor Symptom Scale, or NMSS, as well as safety.  

Biomarker data include measurements of the coverage of the putamen, the specific region of the brain targeted 
with VY-AADC, and measurements of AADC enzyme expression and activity in the putamen measured by PET using 
fluorodopa F-18. Changes in patients’ daily doses of oral levodopa and related medications will also be recorded. 

In December 2018, we held a Type B meeting with the FDA to discuss the overall development program for 

VY-AADC. Based on the meeting discussion and subsequent written feedback from the FDA, we plan to submit a 
revised trial protocol that will include an increase in the target number of patients in the RESTORE-1 Phase 2 trial, 
resulting in 75 to 100 total patients in the trial, and to conduct a staggered-parallel RESTORE-2 Phase 3 trial of similar 
size and design to RESTORE-1. These updates incorporate guidance from the FDA from the Type B meeting to conduct 
two adequate and well-controlled clinical trials for a large patient population such as Parkinson’s disease. 

We expect that the RESTORE-1 Phase 2 trial will enroll 75 to 100 patients. Movement disorder specialists will 

identify and screen potential patients prior to referring to a surgical site for VY-AADC administration and provide 
clinical and safety follow-up after VY-AADC administration.  

We expect patient enrollment to take 15 to 21 months from first patient enrolled, for the RESTORE-1 Phase 2 

clinical trial. We plan to begin enrolling the RESTORE-2 Phase 3 clinical trial in the first half of 2020. We anticipate 
that, if positive, results from the RESTORE-1 Phase 2 clinical trial and the planned RESTORE-2 Phase 3 clinical trial 
could potentially form the basis for submission of a BLA, to the FDA for VY-AADC for the treatment of Parkinson’s 
disease. 

In January 2019, we entered into the Neurocrine Collaboration Agreement for the research, development, and 

commercialization of four programs including our Parkinson’s disease program.  

ALS Program: VY-SOD102 

Disease Overview  

ALS is a fatal neurodegenerative disease that leads to muscle atrophy, spasticity and weakness as well as 

impaired speech, swallowing and breathing, with many patients requiring ventilator support as the disease progresses. 
The average age of onset of ALS is 55 years, and median survival is approximately three years after initial symptoms 
appear. It is estimated that there are approximately 20,000 patients in the United States who are living with the disease. 
Familial, or inherited, ALS accounts for approximately 10% of ALS cases, and an estimated 20% of familial ALS is 
caused by mutations in the superoxide dismutase 1, or SOD1, gene. Therefore, there are an estimated 400-800 patients in 
the United States with ALS caused by mutations in the SOD1 gene.  

The normal function of the SOD1 protein is to catalyze the conversion of superoxide anion (O2

-) to hydrogen 
peroxide (H2O2) and oxygen (O2). Mutations in SOD1 have been shown to lead to the formation of toxic aggregates of 
the SOD1 protein, resulting in the dysfunction and death of motor neurons. Patients with familial ALS caused by certain 
mutations in the SOD1 gene progress more rapidly than patients with other forms of ALS, although the reason for this 
more rapid progression is unknown.  

22 

There are currently only two FDA-approved treatments for ALS, Riluzole by Sanofi, which has been shown to 

have only modest efficacy, prolonging life by a few months, and Edaravone, which has been shown to slow decline of 
daily functioning.  

Our Treatment Approach  

We believe that AAV gene therapy is an attractive approach to treating monogenic ALS caused by SOD1 

mutations. Since the SOD1 gene mutations that cause ALS are toxic gain-of-function mutations, we believe that we can 
employ an AAV gene therapy approach that targets the knockdown of SOD1 gene expression. In addition, the primary 
target cells - motor neurons - reside within the spinal cord, which we believe can be effectively transduced with AAV 
gene therapy through intraparenchymal injection as well as other routes of administration. The mechanism of action of 
VY-SOD102 is knockdown of SOD1 expression in motor neurons, thereby potentially reducing the level of toxicity 
associated with mutated protein, and slowing functional decline and prolonging ventilator-independent survival.  

We believe that there is also the potential to leverage our approach for the treatment of other genetically defined 

forms of ALS.  

Preclinical Studies Targeting SOD1 for Monogenic ALS  

Results from our preclinical studies using intraparenchymal delivery of AAV vector to the spinal cord support 
targeting mutant SOD1 for the treatment of monogenic ALS. In the mini-pig, used as an animal model as it has a spinal 
cord similar in size to the human spinal cord, significant knockdown of SOD1 expression was observed following 
intraparenchymal spinal cord injection of an AAV vector carrying a transgene designed to inhibit SOD1 expression. This 
novel delivery approach with VY-SOD102 reduced SOD1 mRNA in the spinal cord on average by 70% and 50% in the 
cervical and thoracic regions, respectively, both regions critical for respiratory function, and 82% near the site of cervical 
injection. In addition, VY-SOD102 reduced SOD1 mRNA by 22% in the lumbar region.  

The knockdown of SOD1 has also been reported to provide significant survival benefits in animal models of 

ALS. As shown in the example below, mice with a SOD1 mutation treated with an AAV vector to knock down 
expression of the mutant human SOD1 gene extended median survival by 87 days compared to mice treated with a 
control vector.  

Improved Survival Post Knockdown of SOD1(1) 

(1)  Reprinted by permission from Macmillan Publishers Ltd: Foust et al, Molecular Therapy (2013), 21 (12); 2148-2159, copyright (2013). Purple line 

represents mice treated with AAV gene therapy, while gray line represents control mice. 

23 

 
 
These studies provide proof-of-principle for our approach to treating monogenic ALS due to SOD1 mutations 

with VY-SOD102.  

Our Program Status  

In late 2016, we identified VY-SOD101 as a lead clinical candidate after screening a series of capsids, 

microRNA expression cassettes, (a segment of DNA that contains the sequence that targets SOD1 gene expression 
selectively for knockdown), and encoded payloads. We screened more than 100 RNAi sequences, each represented by a 
bar in the graph below, and successfully identified multiple, highly-potent RNAi sequences targeting SOD1, as 
highlighted by the yellow bars in the figure below:  

Overview of miRNA Target Sequences for Knockdown of SOD1 

The most potent RNAi sequences targeting SOD1 gene expression were evaluated in multiple microRNA 

expression cassettes and with a number of vector genome configurations. We have completed the necessary experiments 
to evaluate these potential lead candidates based upon criteria that include safety, selectivity, potency, and efficiency and 
precision of microRNA processing.  

In late 2017, we initiated additional preclinical studies to further optimize our ALS program’s therapeutic 

approach, including exploration of additional routes of administration and novel AAV capsids in large animal models. 
Based on these studies, we selected VY-SOD102 as our lead candidate. VY-SOD102, our clinical candidate for the 
treatment of a monogenic form of ALS, is composed of an adeno-associated virus capsid and a proprietary transgene to 
selectively knock down levels of SOD1 mRNA. VY-SOD102 has the potential to durably reduce the levels of toxic 
mutant SOD1 protein in the spinal cord to slow the progression of disease. In late 2018 we presented data on VY-
SOD102 administered with a novel delivery paradigm comprising a one-time, intraparenchymal infusion after 
laminectomy to the cervical spinal cord of the mini-pig, which has a spinal cord similar in length and diameter to the 
human spinal cord. This delivery approach yielded safe and significant reduction of SOD1 at four weeks post-dosing at 
the site of the infusion in the spinal cord, most notably in the cervical and thoracic regions critical for respiratory 
function. Further preclinical studies are underway with VY-SOD102 which, if successful, will support a potential filing 
of an IND application in 2019.  

Friedreich’s ataxia Program: VY-FXN01 

Disease Overview  

Friedreich’s ataxia is a debilitating neurodegenerative disease resulting in poor coordination of legs and arms, 

progressive loss of the ability to walk, generalized weakness, loss of sensation, scoliosis, diabetes and cardiomyopathy as 
well as impaired vision, hearing and speech. The typical age of onset is 10 to 12 years, and life expectancy is severely 
reduced with patients generally dying of neurological and cardiac complications between the ages of 35 and 45. 

24 

 
According to the Friedreich’s Ataxia Research Alliance, there are approximately 6,400 patients living with the disease in 
the United States. There are currently no FDA-approved treatments for the disease.  

Friedreich’s ataxia patients have mutations of the FXN gene that reduce production of the frataxin protein, 

resulting in the degeneration of sensory pathways and a variety of debilitating symptoms. Friedreich’s ataxia is an 
autosomal recessive disorder, meaning that a person must obtain a defective copy of the FXN gene from both parents in 
order to develop the condition. One healthy copy of the FXN gene, or 50% of normal frataxin protein levels, is sufficient 
to prevent the disease phenotype. We therefore believe that restoring FXN protein levels to at least 50% of normal levels 
by AAV gene therapy might lead to a successful therapy.  

Our Treatment Approach  

We are developing an AAV gene therapy approach that we believe will deliver a functional version of the FXN 
gene to the sensory pathways through intravenous injection. We think this approach has the potential to improve balance, 
ability to walk, sensory capability, coordination, strength and functional capacity of Friedreich’s ataxia patients. Most 
Friedreich’s ataxia patients produce low levels of the frataxin protein, which although insufficient to prevent the disease, 
exposes the patient’s immune system to frataxin. This reduces the likelihood that the FXN protein expressed by AAV 
gene therapy will trigger a harmful immune response.  

Preclinical Studies  

We initially conducted preclinical studies in non-human primates and achieved high FXN expression levels 
within the target sensory ganglia, or clusters of neurons, along the spinal region following intrathecal injection. More 
recently, we conducted preclinical studies in non-human primates with intravenous injection and achieved target FXN 
expression levels within sensory ganglia and the heart. The levels of FXN expression observed using an AAV vector 
were, on average, greater than FXN levels present in control normal human brain tissue. FXN expression was also 
observed in the cerebellar dentate nucleus, another area of the CNS that is often affected in Friedreich’s ataxia, and that 
is often considered difficult to target therapeutically.  

Our Program Status  

VY-FXN01 is currently in preclinical development. We are in the process of identifying a lead candidate which 

will comprise an optimal capsid, promoter, and FXN transgene. We are completing several AAV capsid screening 
experiments to identify capsids that effectively distribute to disease target tissues in a desired manner following 
intravenous injection. Criteria for evaluating these capsids include safety, overall level of transgene expression achieved, 
distribution of transgene expression and the specific cell types transduced. In addition, we are optimizing the promoter 
for VY-FXN01. To evaluate the therapeutic potential of our vectors, we have conducted testing in a new genetic mouse 
model of Friedreich’s ataxia. In this preclinical model of Friedreich’s ataxia, our gene therapy candidates durably 
improved sensory function, and rescued the Friedreich’s ataxia phenotype based on multiple functional tests. In 
physiological and behavioral assays, our gene therapy candidates demonstrated dose-dependent and durable responses 
for more than 10 months after a single administration, preventing central and peripheral disease progression. We also 
have a significant effort focused on better understanding the clinical course of Friedreich’s ataxia and identifying 
potential clinical endpoints for future clinical trials.  

Once we identify a lead candidate for this program, we plan to complete IND enabling studies to evaluate its 

safety and efficacy.  

In January 2019, we entered into the Neurocrine Collaboration Agreement for the research, development, and 

commercialization of four programs including our Friedreich’s ataxia program. 

Huntington’s Disease Program: VY-HTT01 

25 

Disease Overview  

Huntington’s disease is a fatal, inherited neurodegenerative disease that results in the progressive decline of 
motor and cognitive functions and a range of behavioral and psychiatric disturbances. The average age of onset is 39 
years, with patients typically dying approximately 15 to 20 years following diagnosis. According to the Huntington’s 
Disease Society of America, Huntington’s disease affects approximately 30,000 patients in the United States. 
Huntington’s disease is caused by mutations in the huntingtin, or HTT, gene. Huntington’s disease is an autosomal 
dominant disorder, which means that an individual is at risk of inheriting the disease if only one parent is affected. More 
than 200,000 individuals in the United States are at risk for inheriting the mutant gene from an affected parent. While the 
exact function of the HTT gene in healthy individuals is unknown, it is essential for normal development before birth and 
mutations in the HTT gene ultimately lead to the production of abnormal intracellular huntingtin protein aggregates that 
cause neuronal cell death. Currently, there are no approved treatments targeting the underlying cause of the disease and 
only one drug, tetrabenazine, has been approved for the treatment of the specific motor symptoms of Huntington’s 
disease.  

Our Treatment Approach  

We believe that AAV gene therapy is an attractive approach to treating Huntington’s disease. Since HTT 

mutations that cause Huntington’s disease are toxic gain-of-function mutations, we believe that we can employ an AAV 
gene therapy approach designed to knock down expression of the HTT gene. In addition, the targeted cells for treatment 
primarily reside in discrete regions of the brain - the striatum and the cortex - that can be targeted with AAV gene 
therapy delivered directly into the brain. The mechanism of action of VY-HTT01 is knockdown of HTT expression in 
neurons in the striatum and cortex, thereby reducing the level of toxicity associated with mutated protein in these brain 
regions, and slowing the progression of cognitive and motor symptoms. We believe that we can use the same surgical 
approach for this program that has been used for VY-AADC delivery to the brain, allowing us to leverage prior clinical 
experience.  

Preclinical Studies  

Our collaborators at Sanofi Genzyme have completed significant preclinical work focused on AAV gene 
therapy for Huntington’s disease. Sanofi Genzyme’s preclinical studies in a mouse model of Huntington’s disease 
demonstrated the safety and efficacy of AAV gene therapy targeting the knockdown of the HTT gene in the CNS.  

As shown in the figure below, using an AAV vector delivered directly to the CNS, HTT gene expression was 
observed to be reduced by over 50%, on average, in the treatment group as compared to the control group. No signs of 
toxicity were reported.  

Knockdown of HTT Following AAV Delivery(1) 

(1)  Stanek et al, Human Gene Therapy (2014); 25; 461-474. The publisher for this copyrighted material is Mary Ann Liebert, Inc. publishers. 
* 

   p<0.05 

26 

 
 
In addition, significant functional benefit was observed in the treatment group, as measured by the rotarod test 
to assess motor function, and the Porsolt Swim Test to measure depressive behavior in mice. In the figure below, both 
normal or wild type mice, and mice with the HTT mutation, or YAC128, were evaluated following treatment with either 
an AAV vector targeting the knockdown of the HTT gene, labeled as AAV2/1-miRNA-Htt below, or a negative control 
vector, labeled as AAV2/1-Null below. As expected, knocking down HTT in the control mice was observed to have no 
functional impact, whereas knocking down HTT in YAC128 mice was observed to have significant functional benefit.  

Reduction of Behavioral Deficits in an Animal Model of Huntington’s Disease(2)  

(2)  Stanek et al, Human Gene Therapy (2014); 25; 461-474. The publisher for this copyrighted material is Mary Ann Liebert, Inc. publishers. 
* 

   p<0.05 

Our Program Status  

VY-HTT01 is in preclinical development. Sanofi Genzyme’s Huntington’s disease gene therapy program was 

combined with our efforts in connection with entering into the Sanofi Genzyme Collaboration agreement in February 
2015. Through our gene therapy platform, we also constructed and screened a series of microRNA expression cassettes 
and encoded payloads. Multiple rounds of optimization have resulted in potential candidates that are potent and selective 
for knocking down HTT. In addition, many construct configurations were evaluated toward the identification of one 
which would provide excellent yield and genome integrity for manufacturing scale-up in our baculovirus/Sf9 AAV 
manufacturing system in insect-derived cells.  

We also conducted the necessary experiments to evaluate these potential lead candidates based upon criteria 

that include safety, selectivity, potency, and efficiency and precision of microRNA processing, leveraging the learnings 
from the VY-SOD101 program, including the miRNA cassettes and vector genome configurations that we designed for 
the VY-SOD101 program. In June 2017, we reported that we had selected a lead clinical candidate.  

In preclinical studies, a single administration of VY-HTT01 was well-tolerated and resulted in robust and 

widespread knockdown of HTT messenger RNA at five weeks post-dosing in disease-relevant regions of the non-human 
primate central nervous system. The extent of HTT mRNA suppression (greater than 50%) and high precision and 
efficiency of primary microRNA processing in our preclinical studies supported the selection of our lead clinical 
candidate. Additionally, preclinical data in large mammals have demonstrated that a single intraputaminal administration 
results in robust knockdown of HTT in the putamen.  

Recent preclinical delivery studies have further optimized the dosing paradigm. VT-HTT01 is composed of an 

adeno-associated virus capsid (AAV1) and proprietary transgene that harnesses the RNA interference pathway to 
selectively knock down, or reduce, levels of HTT mRNA. In late 2018, we presented results demonstrating significant 

27 

 
 
reduction of HTT mRNA at five weeks post-dosing in adult non-human primates using an MRI-guided surgical delivery 
of VY-HTT01 and a novel delivery paradigm targeting both the putamen and thalamus. Targeting the thalamus in 
addition to the putamen leverages more extensive and more preserved neuronal pathways to the cortex than delivery to 
the putamen alone. This novel dosing paradigm with VY-HTT01 resulted in safe and significant suppression of HTT in 
the striatum and in cortical neurons, which are critical in the progression of disease. A combined infusion of VY-HTT01 
into the putamen and thalamus significantly reduced HTT mRNA by 68% in the caudate, 67% in the putamen, and 73% 
in the thalamus, on average, as measured from tissue punches, and by 32% on average, in laser captured cortical neurons, 
which was also supported by in situ hybridization for HTT mRNA. Further preclinical studies are underway with VY-
HTT01 which, if successful, will support a potential filing of an IND application in 2019. 

Tau Program 

Disease Overview 

In healthy individuals, tau is an abundant soluble cytoplasmic protein that binds to microtubules, which are key 

structural proteins in cells, to promote their stability and function. In Alzheimer’s disease and other tauopathies, tau 
aggregates and forms insoluble tau-containing neurofibrillary tangles. The progressive spread of tau pathology along 
distinct anatomical pathways in the brain closely correlates with disease progression and severity in a number of 
tauopathies, including Alzheimer’s disease, FTD, and PSP. In addition, mutations in the tau gene have been shown to 
cause inherited forms of tauopathies, including FTD and PSP. Because the extent of tau pathology in Alzheimer’s 
disease and other tauopathies closely correlates with the severity of neurodegeneration, synapse loss, and cognitive 
deficits, attempts to prevent, reduce, or slow the development of tau pathology have become important therapeutic 
strategies for these diseases. 

In previous preclinical studies in animal models, despite high weekly or biweekly systemic doses of anti-tau 

monoclonal antibodies administered over three to six months, only very low levels of antibody reached the brain, 
resulting in a modest reduction of tau pathology by ~40–50%. This incomplete and modest reduction in tau pathology 
following treatment with very high and frequent systemic doses of these antibodies may pose therapeutic challenges in 
humans with various tauopathies. To address these limitations, our AbbVie Tau Collaboration attempts to develop AAV 
gene therapies to deliver monoclonal antibodies to the brain directed against tau as potential new treatments for 
Alzheimer’s disease and other tau-related neurodegenerative diseases. 

Our Program Status 

The tau program is currently in the preclinical stage. We have agreed to collaborate with AbbVie on the 
research and development of specified vectorized antibody compounds comprised of an AAV or other viral capsid and a 
virus vector genome that encodes one or more antibodies that target and bind to a tau protein. During the research period 
of our collaboration, we and AbbVie agreed to each identify up to five potential antibodies for evaluation during the 
AbbVie Tau Collaboration. Under the agreement, up to three research antibodies may be selected as candidates for 
creation of research compounds. AbbVie has the right to select two of the three research antibodies. During a specified 
portion of the research period, AbbVie may exercise one or more of its exclusive development options to select up to a 
total of three research compounds and their corresponding product candidates to proceed to the development period.  

Severe, Chronic Pain Program: VY-NAV01 

Disease Overview 

Nav1.7 is a sodium ion channel that is required for transmission of pain signals to the CNS. We believe that an 
AAV gene therapy approach targeting the knockdown of Nav1.7 in sensory neurons could be an effective treatment for 
certain forms of severe, chronic pain. A major challenge for the successful development of small molecules and 
antibodies targeting Nav1.7 has been the selective inhibition of Nav1.7 over closely related sodium channels such as 
Nav1.5 which are important for cardiac function. MicroRNAs, which work by harnessing the RNA interference pathway, 

28 

can achieve a high level of specificity for their messenger RNA targets, and can inhibit Nav1.7 selectively over other 
sodium channel subtypes. Such an approach could avoid the dose-limiting side effects associated with the non-selective 
profile of many current drugs used to treat severe, chronic pain, and also achieve a durable clinical benefit following a 
single administration of the therapy. VY-NAV01 leverages our extensive experience designing novel microRNA 
knockdown cassettes and delivering them using AAV, an approach that we are using for our ALS and Huntington’s 
disease programs. 

Our Program Status 

VY-NAV01 is currently in the research stage. We are in the process of conducting proof-of-concept studies to 
establish the level of Nav1.7 knockdown needed to relieve pain in animal models. We will then identify a lead candidate 
which will comprise an optimal capsid, promoter, and microRNA targeting Nav1.7. We have also initiated proof-of-
concept studies to evaluate knockdown of another sodium channel subtype implicated in chronic pain that may be 
combined with Nav1.7 knockdown. We are completing several AAV capsid screening experiments to identify capsids 
that effectively distribute to pain sensory neurons in a desired manner. We are comparing capsids in non-human primates 
following intrathecal and intravenous injection, and evaluating these capsids based upon multiple criteria including 
safety, overall level of transgene expression achieved, distribution of transgene expression and the specific cell types 
transduced. 

Future Programs  

We are evaluating additional severe neurological diseases that could be treated using AAV gene therapy 

through application of either a gene replacement or a gene knockdown approach.  

Collaborations and License Agreements 

Sanofi Genzyme Collaboration  

In February 2015, we entered into a strategic collaboration with Sanofi Genzyme to leverage our combined 

expertise and assets to develop AAV gene therapies for neurological diseases. Under the agreement, we retained U.S. 
rights to VY-AADC and VY-FXN01, as well as at least co-commercialization rights to VY-HTT01 in the United States. 
In October 2017, Sanofi Genzyme decided not to exercise its option for the ex-U.S. rights to VY-AADC, returning 
global rights to VY-AADC to us. VY-SOD102 is not included as part of the Sanofi Genzyme collaboration and we retain 
unencumbered worldwide rights to this program. Sanofi Genzyme maintains exclusive options to license, develop and 
commercialize (i) the Split Territory Programs outside the United States with an incremental option to co-commercialize 
VY-HTT01 in the United States, and (ii) VY-SMN101 worldwide. Sanofi Genzyme’s option for the Split Territory 
Programs and VY-SMN101 is triggered following the completion of the first POP Study on a program-by-program basis. 
In November 2016, we and Sanofi Genzyme elected to deprioritize the development of VY-SMN101 for spinal muscular 
atrophy due to, among other things, the significant progress we have made in our other preclinical programs and the 
evolving competitive landscape. 

Prior to any option exercise by Sanofi Genzyme, we will collaborate with Sanofi Genzyme in the development 
of products under each Split Territory Program and VY-SMN101 pursuant to a written development plan and under the 
guidance of an alliance joint steering committee, comprised of an equal number of our employees and Sanofi Genzyme 
employees.  

We are required to use commercially reasonable efforts to develop products under each Split Territory Program 

and VY-SMN101 through completion of the applicable POP Study. During the development of our joint programs, our 
and Sanofi Genzyme’s activities are guided by a Development Advisory Committee, which we refer to as the DAC. The 
DAC may elect to utilize certain Sanofi Genzyme technology relating to the VY-AADC program, the VY-HTT01 
Program, or generally with the manufacture of Split Territory Program products. If we use certain Sanofi Genzyme 
technology in VY-AADC, Sanofi Genzyme is entitled to received low single digit royalty payments based on a 

29 

percentage of net sales by us, and we may be obligated to make certain regulatory milestone payments to a third-party 
licensor.  

We will be solely responsible for all costs incurred in connection with the development of Split Territory 
Programs and VY-SMN101 products prior to option exercise, subject to the following: (i) Sanofi Genzyme may agree to 
provide additional funds in return for agreed-upon payback or other agreed economic terms; (ii) we may request, and 
upon mutual agreement, Sanofi Genzyme will provide in-kind services valued at up to $5.0 million; and (iii) expenses of 
certain activities under the VY-HTT01 development plan may be funded to the extent such activities are reimbursed 
through financial support that Sanofi Genzyme may receive from a disease foundation group.  

Excluding the VY-AADC program, if we do not initiate a POP Study for a given Split Territory Program by 

December 31, 2026 (or for the Future Program by the tenth anniversary of the date the Future Program is nominated by 
Sanofi Genzyme), and Sanofi Genzyme has not terminated this agreement with respect to such Collaboration program, 
then Sanofi Genzyme shall be entitled, at its sole and exclusive remedy, to a credit of $10.0 million for each such 
program against other amounts payable by Sanofi Genzyme under the Collaboration. However, if we do not initiate a 
POP Study by such date as a result of a regulatory delay or a force majeure event, such time period shall be extended for 
so long as such regulatory delay or force majeure event continues and we shall not be deemed to have failed to initiate a 
POP Study.  

Post-Option Exercise  

Upon Sanofi Genzyme’s exercise of its option to license a given product in a Split Territory Program, which we 
refer to as a Split Territory Licensed Product, we will have sole responsibility for the development of such Split Territory 
Licensed Product in the United States and Sanofi Genzyme shall have sole responsibility for development of such Split 
Territory Licensed Product in the rest of the world. We and Sanofi Genzyme will have shared responsibility for 
execution of ongoing development of such Split Territory Licensed Product that is not specific to either of our territories, 
including costs associated therewith.  

A steering committee for each program will review and approve a written plan and budget for each relevant 

program. In addition, all development activities to be undertaken with respect to each Split Territory Licensed Product 
by or on behalf of either party will be set forth in a written development plan.  

Sanofi Genzyme shall have the sole right to develop VY-SMN101 worldwide. Sanofi Genzyme shall be 

responsible for all of the development costs that occur after the option exercise date for VY-SMN101.  

Commercialization  

We shall be solely responsible, at our expense, for all commercialization activities relating to Split Territory 

Licensed Products in the United States. Sanofi Genzyme shall be solely responsible, at its expense, for all 
commercialization activities relating to Split Territory Licensed Products in the rest of the world. For VY-HTT01, if 
Sanofi Genzyme has exercised its option to co-commercialize VY-HTT01 in the United States, then Sanofi Genzyme 
will be the lead party responsible for all VY-HTT01 commercialization activities in the United States, and these 
activities will be set forth in reasonable detail in a written commercialization plan.  

Sanofi Genzyme shall be solely responsible, at its expense, for all commercialization activities relating to VY-

SMN101 worldwide. Sanofi Genzyme shall use commercially reasonable efforts to commercialize VY-SMN101 in each 
major market specified in the agreement where Sanofi Genzyme has obtained required governmental approvals. 

Financial Terms  

We received $65.0 million in upfront cash, a $30.0 million upfront equity investment and an in-kind 

commitment of $5.0 million, totaling $100.0 million. If Sanofi Genzyme exercises its option for a collaboration program, 

30 

Sanofi Genzyme is required to make an option exercise payment of $20.0 million or $30.0 million for each program. We 
are no longer entitled to receive the regulatory and commercial milestone payments from Sanofi Genzyme related to VY-
AADC. Sanofi Genzyme shall pay us up to $540.0 million across the remaining product programs upon the achievement 
of specified regulatory and commercial milestones.  

In addition, to the extent any Split Territory Licensed Product or the VY-SMN101 Product is commercialized, 
we are entitled to receive tiered royalty payments ranging from the mid-single digits to mid-teens based on a percentage 
of net sales. Sanofi Genzyme is entitled to receive royalty payments from us related to sales of the Split Territory 
Licensed Products ranging from the low-single digits to mid-single digits, depending on whether we use Sanofi 
Genzyme technology in a Split Territory Licensed Product or the VY-SMN101 Product. If Sanofi Genzyme exercises its 
option to co-commercialize VY-HTT01 in the United States, we will share any profits or losses from VY-HTT01 
product sales.  

Term and Termination; Remedies  

Our collaboration agreement with Sanofi Genzyme will continue in effect until the later of (i) the expiration of 
the last to expire of the option rights and (ii) the expiration of all payment obligations unless sooner terminated by us or 
Sanofi Genzyme.  

We and Sanofi Genzyme have customary termination rights including the right to terminate for an uncured 

material breach of the agreement committed by the other party and Sanofi Genzyme has the right to terminate for 
convenience.  

AbbVie Tau Collaboration 

In February 2018, we entered into an exclusive collaboration and option agreement with AbbVie, or the AbbVie 
Tau Collaboration Agreement, for the research, development, and commercialization of AAV and other virus-based gene 
therapy products for the treatment of neurodegenerative diseases related to defective or excess aggregation of tau protein 
in the human brain, including Alzheimer’s disease.  

Under the AbbVie Tau Collaboration Agreement, we have agreed to collaborate with AbbVie on the research 

and development of specified vectorized antibody compounds comprised of an AAV or other viral capsid and a virus 
vector genome that encodes one or more antibodies that target and bind to a tau protein. The collaboration is comprised 
of a research period, a development period, and an exclusive license option. 

Research Period and AbbVie Development Option 

During the research period, each party has agreed to identify up to five antibodies for inclusion in the 

collaboration. Subject to certain conditions and exceptions, the parties will then select up to three antibodies, or the 
Research Antibodies, as candidates for creation of research compounds, or the Research Compounds, with AbbVie 
having the right to select two of the three Research Antibodies. We are obligated to use diligent efforts to conduct 
antibody engineering and other research activities to create Research Compounds and to develop product candidates 
containing or comprised of such Research Compounds. We will be solely responsible for the costs and expenses during 
the Research Period. During a specified portion of the Research Period, or the Development Option Period, AbbVie may 
exercise one or more of its exclusive development options, each of which we refer to as a Development Option, to select 
up to a total of three Research Compounds, or the Selected Research Compounds, and their corresponding product 
candidates, or the Selected Product Candidates, to proceed to the development period. 

Development Period and AbbVie License Option 

During the development period, we are obligated to use diligent efforts to conduct development activities, 

including IND-enabling and Phase 1 clinical trial activities, for the Selected Research Compounds and corresponding 

31 

Selected Product Candidates. We will be solely responsible for the costs and expenses during the development period. 
During a specified portion of the development period, or the License Option Period, AbbVie may exercise its exclusive 
license option, or the License Option, to further develop and commercialize all of the Research Compounds, or the 
Licensed Compounds, and corresponding product candidates, or the Licensed Products. Upon AbbVie’s exercise of its 
License Option, we have agreed to grant to AbbVie an exclusive, worldwide license, with the right to sublicense, under 
certain of our intellectual property rights to develop and commercialize the Licensed Compounds and the Licensed 
Products for all human diagnostic, prophylactic and therapeutic uses. In addition, after AbbVie’s exercise of the License 
Option, we have certain obligations to complete any remaining research and development activities that have not been 
completed for any Research Compounds and Product Candidates. 

Governance 

Our research and development activities will be conducted pursuant to the plans agreed to by the parties and 
overseen by a joint governance committee, or JGC, comprised of an equal number of representatives from each party. 
Prior to AbbVie’s exercise of its License Option, we will have final decision-making authority within the JGC, subject to 
specified limitations; thereafter, AbbVie will have final decision-making authority, subject to specified limitations. Any 
material amendment to the research or development plans, however, must be mutually agreed to by the parties, which 
may be through the JGC. 

Commercialization 

Under the AbbVie Tau Collaboration Agreement, AbbVie is required to use commercially reasonable efforts to 

develop and commercialize at least one Licensed Product in each of the United States, Japan, the United Kingdom, 
Germany, France, Italy and Spain. After exercise of the License Option, AbbVie is solely responsible for all 
development and commercialization activities relating to Licensed Compounds and Licensed Products at its sole cost and 
expense (subject to our obligation to complete any remaining research and development activities set forth in the agreed-
upon plans), except that we may elect to share in AbbVie’s development costs relating to a Licensed Product on an 
indication-by-indication basis in exchange for a specified increase in royalties. If we exercise this cost-sharing option, we 
may either reimburse AbbVie for AbbVie’s applicable development costs or, in the case of certain budget overruns, 
AbbVie may instead deduct applicable development costs, up to a specified cap, from milestone and royalty payments 
owed by AbbVie to us.  

Manufacturing 

During both the research period and the development period, we will be solely responsible for the manufacture 
and supply of all pre-clinical and clinical requirements for the Research Compounds and Product Candidates. If AbbVie 
were to exercise its License Option, we would be required, at AbbVie’s request, to effect a full transfer of the 
manufacturing process for each Licensed Compound and corresponding Licensed Product to AbbVie. Following such 
transfer, we have agreed to disclose, on a continuing basis, all modifications, enhancements and improvements to 
manufacturing processes for the Licensed Products, and AbbVie has agreed to grant to us a non-exclusive, royalty-free 
license to modifications to the manufacturing process made by AbbVie, in each case subject to specified limitations. 

Financial Terms 

Under the terms of the AbbVie Tau Collaboration Agreement, AbbVie paid us an upfront payment of $69.0 

million in February 2018. AbbVie has also agreed to pay us within 30 days after the applicable exercise date: (1) upon 
AbbVie’s exercise of a Development Option, (a) $80.0 million for the first Selected Research Compound and its 
corresponding Selected Product Candidate and (b) $30.0 million each for up to two additional Selected Research 
Compounds and their corresponding Selected Product Candidates, and (2) upon AbbVie’s exercise of the License 
Option, a one-time payment of $75.0 million. We will be eligible to receive (1) specified development and first-sale 
milestone payments for each Licensed Compound of up to an aggregate of $550.0 million in the case of an Alzheimer’s 
disease indication, up to $230.0 million in the case of the first indication other than Alzheimer’s disease, and $115.0 
million for subsequent non-Alzheimer’s disease indication; and (2) tiered, escalating royalties, in a range from a high-

32 

single digit to a mid-to-high teen (or, if we have exercised our cost-sharing option, low-twenties) percentage of aggregate 
net sales of Licensed Products on a Licensed Compound by Licensed Compound basis. The royalties are subject to 
potential reductions for biosimilar market penetration, patent claim expiration, and other provisions, subject to specified 
limits. For each Licensed Product, AbbVie may make a one-time request either to decrease its royalty payments to a 
specified low-single digit percentage or to terminate them altogether in exchange for a one-time payment by AbbVie at a 
fair market value to be negotiated by the parties. If the parties are not able to agree to the terms of such buy-down, the 
parties may seek a fair market value determination for the buy-down pursuant to dispute resolution procedures specified 
in the agreement.  

Intellectual Property 

Under the terms of the AbbVie Tau Collaboration Agreement, each party will own the entire right, title and 

interest in and to all know-how and patent rights first made or invented solely by it or its affiliates or its or their 
sublicensees in the course of the collaboration, with certain specified exceptions. Also subject to specified exceptions, 
the parties will jointly own all rights, title and interest in and to all know-how and patent rights first made or invented 
jointly by such party or its affiliates or its or their sublicensees in the course of the collaboration. Regardless of whether 
AbbVie has exercised a Development Option or the License Option, we have agreed to grant AbbVie perpetual, 
exclusive or non-exclusive (as the case may be), worldwide licenses to certain know-how and patent rights developed by 
us or jointly by the parties arising from the collaboration. 

Exclusivity 

During the term of the AbbVie Tau Collaboration Agreement, (1) neither party nor any of its respective 

affiliates is permitted to directly or indirectly exploit any vectorized antibody compound targeting a tau protein, which 
we refer to as Vectorized Antibody Exclusivity, and (2) neither we nor any of our affiliates is permitted to directly 
exploit any Research Antibody targeting a tau protein, which we refer to as Research Antibody Exclusivity, in each case 
subject to specified exceptions, including our conduct of basic research.  

Termination 

Unless earlier terminated, the AbbVie Tau Collaboration Agreement will expire on the earliest to occur of the 
expiration of (1) the Development Option Period, without AbbVie’s exercise of a Development Option; (2) the License 
Option Period, without AbbVie’s exercise of its License Option; and (3) the last-to-expire royalty term with respect to all 
Licensed Products in all countries. Subject to a cure period, either we or AbbVie may terminate the AbbVie Tau 
Collaboration Agreement, in whole or, in the case of us, in part, subject to specified conditions, in the event of the other 
party’s uncured material breach. Either we or AbbVie may also terminate, subject to specified conditions, for insolvency 
of the other party, certain failures or delays to obtain certain regulatory clearances of the collaboration, or a joint 
determination of scientific infeasibility by the parties. AbbVie may terminate the AbbVie Tau Collaboration Agreement 
(1) without cause, in its entirety or, after its exercise of the License Option, on a country-by-country basis, with 180 
days’ prior written notice or (2) for our non-compliance with certain anti-bribery or anti-corruption covenants. We may 
terminate the AbbVie Tau Collaboration Agreement, subject to specified conditions, if AbbVie or its affiliates challenge 
the validity or enforceability of certain of our, or jointly-held intellectual property rights.  

Upon termination in certain cases, AbbVie has agreed to grant to us reversionary licenses to certain Licensed 

Compounds. In such case, we may be required to pay royalties to AbbVie in a range from a low to high single digit 
percentage of net sales of Licensed Products containing or comprised of such License Compound, subject to potential 
reduction in some cases. Additionally, upon termination in certain cases, the Vectorized Antibody Exclusivity and 
Research Antibody Exclusivity will survive until the third anniversary of the termination date. If the parties mutually 
agree to terminate for infeasibility or AbbVie terminates for our failure to deliver a final research or development report, 
neither us nor any of its affiliates may directly or indirectly exploit a vectorized antibody compound that targets or binds 
to a tau protein for 18 months after the termination date. 

Neurocrine Collaboration 

33 

In January 2019, we entered into the Neurocrine Collaboration Agreement for the research, development and 

commercialization of certain of our AAV gene therapy products. Under the Neurocrine Collaboration Agreement, upon 
the expiration or termination of applicable waiting periods and the receipt of any required approvals or clearances 
including antitrust clearance, we have agreed to collaborate on the conduct of four collaboration programs, which we 
refer to collectively as the Neurocrine Programs: the AADC Program for the treatment of Parkinson’s disease, our FA 
Program for the treatment of Friedreich’s ataxia including the development of the VY-FXN01 product candidate, which 
together with the AADC Program, we refer to as the Existing Programs, and two programs to be determined by us and 
Neurocrine at a later date, which we refer to as the Discovery Programs. 

Collaboration and Licenses 

Under the terms of the Neurocrine Collaboration Agreement, subject to the rights retained by us thereunder, we 
have agreed to collaborate with Neurocrine on, and to grant, exclusive, royalty-bearing, non-transferable, sublicensable 
licenses to certain of our intellectual property rights, for all human and veterinary diagnostic, prophylactic, and 
therapeutic uses, for the research, development, and commercialization of gene therapy products, which we refer to as 
the Collaboration Products, under (i) the AADC Program, on a worldwide basis; (ii) the FA Program, in the United 
States and, upon expiration of Sanofi Genzyme’s option to the FA Program pursuant to the Sanofi Genzyme 
Collaboration without exercise of such option, all countries in the world in which the Neurocrine Collaboration 
Agreement remains in effect with respect to the FA Program; and (iii) each Discovery Program, on a worldwide basis. 

Pursuant to development plans to be agreed by the parties, which will be overseen by a joint steering committee, 

or JSC, we have operational responsibility, subject to certain exceptions, for the conduct of each Neurocrine Program 
prior to the Transition Event for each Program, as described below, and are required to use commercially reasonable 
efforts to develop the Collaboration Products. Neurocrine has agreed to be responsible for all costs incurred by us in 
conducting these activities for each Neurocrine Program, in accordance with an agreed budget. If we breach our 
development responsibilities or in certain circumstances upon a change in control of us, Neurocrine has the right but not 
the obligation to assume the activities under such Neurocrine Program. 

Upon the occurrence of a specified event for each Neurocrine Program, or a Transition Event, Neurocrine has 
agreed to assume responsibility for development, manufacturing and commercialization activities for such Neurocrine 
Program from us and to pay milestones and royalties on future net sales as described further below. For each Existing 
Program, we have the option, or a Co-Co Option, to co-develop and co-commercialize such Neurocrine Program upon 
the occurrence of a specified event, or a Co-Co Trigger Event. Should we elect to exercise our Co-Co Option, we have 
agreed to enter into a cost- and profit-sharing arrangement with Neurocrine, or a Co-Co Agreement, whereby we have 
agreed to jointly develop and commercialize Collaboration Products for such Neurocrine Program, or Co-Co Products, 
and share in its costs, profits and losses, and we agree to forfeit certain milestones and royalties on net sales in the United 
States during the effective period of the applicable Co-Co Agreement. The Transition Events are (i) with respect to the 
AADC Program, our receipt of topline data for the ongoing RESTORE-1 Phase 2 clinical trial for VY-AADC; (ii) with 
respect to the FA Program, our receipt of topline data for the initial Phase 1 clinical trial for an Friedreich’s ataxia 
program product candidate; and (iii) with respect to each Discovery Program, the preparation by us and the approval by 
Neurocrine of an investigational new drug application to be filed with the FDA by Neurocrine for the first development 
candidate in such Discovery Program. The Co-Co Trigger Events are (i) with respect to the AADC Program, our receipt 
of topline data for the ongoing RESTORE-1 Phase 2 clinical trial for VY-AADC and (ii) with respect to the FA 
Program, the achievement of milestones or metrics specified in the applicable development plan, as determined by the 
JSC. 

Subject to exceptions specified in the Neurocrine Collaboration Agreement, profits and losses under our Co-Co 
Option are agreed to be allocated (i) 50% to Neurocrine and 50% to us for a Collaboration Product from the VY-AADC 
program and (ii) 60% to Neurocrine and 40% to us for a Collaboration Product from the FA Program; provided, 
however, that Neurocrine may elect, within a specified period following the acceptance for filing of a BLA from the 
FDA, to pay a $35.0 million rate-shifting fee to us to change the allocation for the VY-AADC Program to 55% to 
Neurocrine and 45% to us. The parties have agreed that each Co-Co Agreement will provide us the right to terminate for 
any reason upon prior written notice to Neurocrine and Neurocrine the right to terminate in certain circumstances upon 
our change of control. 

34 

 
  
  
  
Governance 

Our research and development activities under the Neurocrine Collaboration Agreement are to be conducted 

pursuant to plans agreed to by the parties, on a program-by-program basis, and overseen by the JSC, which is composed 
of an equal number of representatives from the parties. The JSC may delegate matters within its authority to 
subcommittees of the JSC. In addition, the Neurocrine Collaboration Agreement establishes working groups to handle 
specified matters on a subject matter-by-subject matter basis. If a working group or subcommittee cannot agree on a 
matter within its purview within a specified time, such matter is to be referred sequentially to the JSC and then the 
executive officers of the parties. If the executive officers are not able to resolve the matter, then (i) with respect to each 
Existing Program, subject to specified exceptions, (x) Neurocrine has the right to resolve such matter prior to our 
exercise of our Co-Co Option with regard to such Co-Co Product or if such Co-Co Option expires or goes unexercised 
and (y) following the timely exercise by us of our Co-Co Option, depending on the subject of such matter, either 
Neurocrine, in certain instances, or the parties jointly or the JSC, in other instances, would have the right to resolve such 
matter, and (ii) with respect to Discovery Programs, subject to specified exceptions, Neurocrine has the right to resolve 
such matter. 

Candidate Selection 

The parties have committed, following the effective date of the Neurocrine Collaboration Agreement, to agree 

on a list of up to eight target genes, or Targets, from which Neurocrine has the right to nominate Targets for the two 
Discovery Programs. Each Target for the Discovery Programs must be approved by a consensus of the JSC or the 
executive officers. 

Manufacturing 

Prior to the Transition Event for a Neurocrine Program, we are responsible for the manufacture of any 
Collaboration Products for the Program. Following the Transition Event, the parties shall negotiate the manufacturing 
and supply responsibilities, subject to the terms of any applicable Co-Co Agreement. 

Financial Terms 

Under the terms of the Neurocrine Collaboration Agreement, Neurocrine has agreed to pay us an upfront 

payment of $115.0 million within five business days after the effective date, and $50.0 million as an equity purchase of 
4,179,728 shares to purchase our common stock. The Neurocrine Collaboration Agreement provides for aggregate 
development milestone payments from Neurocrine to us for Collaboration Products under (i) the AADC Program of up 
to $170.0 million; (ii) the FA Program of up to $195.0 million, and (iii) each of the two Discovery Programs of up to 
$130.0 million per Discovery Program. We may be entitled to receive aggregate commercial milestone payments for 
each Collaboration Product of up to $275.0 million, subject to an aggregate cap on commercial milestones across all 
Neurocrine Programs of $1.1 billion. 

Neurocrine has also agreed to pay us royalties, based on future net sales of the Collaboration Products. Such 
royalty percentages, for net sales in and outside the United States, as applicable, range (i) for the VY-AADC program, 
from the mid-teens to thirty and the low-teens to twenty, respectively; (ii) for the FA Program, from the low-teens to 
high-teens and high-single digits to mid-teens, respectively; and (iii) for each Discovery Program, from the high-single 
digits to mid-teens and mid-single digits to low-teens, respectively. On a country-by-country and program-by-program 
basis, royalty payments would commence on the first commercial sale of a Collaboration Product and terminate on the 
later of (a) the expiration of the last patent covering the Collaboration Product or its method of use in such country, 
(b) 10 years from the first commercial sale of the Collaboration Product in such country and (c) the expiration of 
regulatory exclusivity in such country, or the Royalty Term. Royalty payments may be reduced by up to 50% in 
specified circumstances, including expiration of patents rights related to a Collaboration Product, approval of biosimilar 
products in a given country or required payment of licensing fees to third parties related to the development and 
commercialization of any Collaboration Product. Additionally, the licenses granted to Neurocrine shall automatically 

35 

  
convert to fully paid-up, non-royalty bearing, perpetual, irrevocable, exclusive licenses on a country-by-country and 
product-by-product basis upon the expiration of the Royalty Term applicable to such Collaboration Product in such 
country. 

Intellectual Property 

Under the terms of the Neurocrine Collaboration Agreement and subject to specified exceptions therein, each 

party owns the entire right, title and interest in and to all intellectual property rights made solely by its employees or 
agents in the course of the collaboration. The parties jointly own all rights, title and interest in and to all intellectual 
property rights made or invented jointly by employees or agents of both parties. 

Exclusivity 

During the term of the Neurocrine Collaboration Agreement, neither party nor any of its respective affiliates is 

permitted to directly or indirectly exploit any AAV-based gene therapy products directed to a Target to which a 
Collaboration Product is directed, subject to specified exceptions, including the parties’ conduct of basic research and 
our activities under the Sanofi Genzyme Collaboration. 

Termination 

Unless earlier terminated, the Neurocrine Collaboration Agreement expires on the later of (i) the expiration of 

the last to expire Royalty Term with respect to a Collaboration Product in all countries in the relevant territory or (ii) the 
expiration or termination of all Co-Co Agreements. Neurocrine may terminate the Neurocrine Collaboration Agreement 
in its entirety or on a program-by-program or country-by-country basis by providing at least (x) 180-day advance notice 
if such notice is provided prior to the first commercial sale of the Collaboration Product to which the termination applies 
or (y) one-year advance notice if such notice is provided after the first commercial sale of the Collaboration Product to 
which the termination applies. We may terminate the Neurocrine Collaboration Agreement, subject to specified 
conditions, if (i) Neurocrine fails to make the equity purchase of 4,179,728 shares of our common stock, for an aggregate 
purchase price of approximately $50.0 million, or (ii) Neurocrine challenges the validity or enforceability of certain of 
our intellectual property rights. Subject to a cure period, either party may terminate the Neurocrine Collaboration 
Agreement in the event of a material breach by the other party in whole or in part, subject to specified conditions. Either 
party may also terminate the Neurocrine Collaboration Agreement if specified regulatory agencies seek to enjoin the 
transaction or if the parties are unable to obtain antitrust clearance within 180 days of the applicable antitrust filings. 

Upon termination in certain cases, Neurocrine has agreed to grant to us licenses to certain Neurocrine 

intellectual property, subject to a negotiation between the parties to establish royalty rates for use of such intellectual 
property. In the event of a breach by us with respect to a Neurocrine Program, if such termination were to occur after a 
Transition Event, then (i) if a Co-Co Agreement is in effect with respect to such program, Neurocrine can terminate the 
Co-Co Agreement for such program and we would no longer have co-development and co-commercialization rights with 
respect to the Collaboration Product and (ii) subject to any license agreements, Neurocrine would no longer have any 
obligations with respect to any Collaboration Products resulting from such program. 

AbbVie Alpha-Synuclein Collaboration 

In February 2019, we entered into an exclusive collaboration and option agreement, the AbbVie Alpha-

Synuclein Collaboration Agreement, with AbbVie, for the research, development and commercialization of AAV and 
other virus-based gene therapy products directed against pathological species of alpha-synuclein for the potential 
treatment of indications including Parkinson’s disease and other synucleinopathies. 

Collaboration and AbbVie Options  

Under the AbbVie Alpha-Synuclein Collaboration Agreement, we and AbbVie have agreed to collaborate on 
the research and development of specified vectorized antibody compounds comprised of an AAV or other viral capsid 

36 

 
  
 
and a virus vector genome that encodes one or more antibodies that target and bind to the alpha-synuclein protein. The 
collaboration is comprised of a research period and a development period. 

Research Period and AbbVie Development Option 

During the research period, we are obligated to conduct research activities directed to constructing one or more 
virus vectors that encode antibodies designated by AbbVie, or AbbVie Designated Antibodies, which initially are to be 
antibodies provided by AbbVie. We are obligated to use diligent efforts to conduct research activities to create research 
compounds and to develop product candidates containing or comprised of such research compounds. We are solely 
responsible for the costs and expenses during the research period. During a specified portion of the research period, 
AbbVie may exercise one or more of its exclusive development options to select up to a total of four research 
compounds and their corresponding Product candidates to proceed to the development period. 

Development Period and AbbVie License Option 

During the development period, we are obligated to use diligent efforts to conduct development activities, 

including IND application-enabling and Phase 1 clinical trial activities, for the selected research compounds and 
corresponding selected product candidates. We are solely responsible for the costs and expenses during the development 
period. During a specified portion of the development period, AbbVie may exercise its exclusive license option to further 
develop and commercialize all of the research compounds and corresponding product candidates. Upon AbbVie’s 
exercise of its license option, we have agreed to grant to AbbVie an exclusive, worldwide license, with the right to 
sublicense, under certain of our intellectual property rights to develop and commercialize the licensed compounds and 
the licensed products for all human diagnostic, prophylactic and therapeutic uses. In addition, after AbbVie’s exercise of 
the license option, we have certain obligations to complete any remaining research and development activities that have 
not been completed for any research compounds and product candidates. 

Governance 

Our research and development activities are to be conducted pursuant to the plans agreed to by the parties and 

overseen by a joint governance committee, or the ASN JGC, comprised of an equal number of representatives from each 
of us and AbbVie. Prior to AbbVie’s exercise of its License Option, we have final decision-making authority within the 
ASN JGC, subject to specified limitations; thereafter, AbbVie is entitled to final decision-making authority, subject to 
specified limitations. Any material amendment to the research or development plans, however, must be mutually agreed 
to by the parties, which may be through the ASN JGC. 

Commercialization 

Under the AbbVie Alpha-Synuclein Collaboration Agreement, AbbVie is required to use commercially 

reasonable efforts to develop and commercialize at least one licensed product in each of the United States, Japan, the 
United Kingdom, Germany, France, Italy and Spain. After exercise of the license option, AbbVie is solely responsible 
for all development and commercialization activities relating to licensed compounds and licensed products at its sole cost 
and expense, subject to our obligation to complete any remaining research and development activities set forth in the 
agreed-upon plans. 

Manufacturing 

During both the research period and the development period, we are solely responsible for the manufacture and 
supply of all pre-clinical and clinical requirements for the research compounds and product candidates. If AbbVie were 
to exercise its license option, we would be required, at AbbVie’s request, to effect a full transfer of the manufacturing 
process for each licensed compound and corresponding licensed product to AbbVie. Following such transfer, we have 
agreed to disclose, on a continuing basis, all modifications, enhancements and improvements to manufacturing processes 
for the licensed products, and AbbVie has agreed to grant to us a non-exclusive, royalty-free license to modifications to 
the manufacturing process made by AbbVie, in each case, subject to specified limitations. 

37 

Financial 

Under the terms of the AbbVie Alpha-Synuclein Collaboration Agreement, AbbVie has agreed to pay us an 

upfront payment of $65.0 million within 15 business days of entry into the AbbVie Alpha-Synuclein Collaboration 
Agreement. AbbVie has also agreed to pay to us, within 30 days after the applicable exercise date: (1) upon AbbVie’s 
exercise of a development option, (a) $80.0 million for the first selected research compound and its corresponding 
selected product candidate and (b) $30.0 million each for up to three additional selected research compounds and their 
corresponding selected product candidates, and (2) upon AbbVie’s exercise of the license option, a one-time payment of 
$75.0 million. We are eligible to receive (1) specified regulatory milestone payments for each licensed compound of up 
to an aggregate of $450.0 million in the case of a Parkinson’s disease indication and up to $185.0 million in the case of 
the first indication other than Parkinson’s disease and $92.5 million for a subsequent non-Parkinson’s disease indication; 
(2) specified commercial milestone payments for all licensed products for all indications up to an aggregate of $500.0 
million; and (3) tiered, escalating royalties, in the mid-single digit percentage range for aggregate net sales of licensed 
products on a licensed compound by licensed compound basis. The royalties are subject to potential reductions for 
biosimilar market penetration, patent claim expiration, and other provisions, subject to specified limits. Subject to certain 
exceptions, we and AbbVie have agreed to be financially responsible for all payments owed to a third party with which it 
has contracted for any use of in-licensed intellectual property under the AbbVie Alpha-Synuclein Collaboration 
Agreement. 

Intellectual Property 

Under the terms of the AbbVie Alpha-Synuclein Collaboration Agreement, each party owns the entire right, 

title and interest in and to all know-how and patent rights first made or invented solely by it or its affiliates or its or their 
sublicensees in the course of the collaboration, with certain specified exceptions. Also subject to specified exceptions, 
the parties jointly own all rights, title and interest in and to all know-how and patent rights first made or invented jointly 
by such party or its affiliates or its or their sublicensees in the course of the collaboration. Regardless of whether AbbVie 
has exercised a development option or the license option, we have agreed to grant AbbVie perpetual, exclusive or non-
exclusive (as the case may be), worldwide licenses to certain know-how and patent rights developed by the Company or 
jointly by the parties arising from the collaboration. 

Exclusivity 

During the term of the AbbVie Alpha-Synuclein Collaboration Agreement, (1) neither party nor any of its 

respective affiliates is permitted to directly or indirectly exploit any vectorized antibody compound targeting the alpha-
synuclein protein, or Vectorized Antibody Exclusivity and (2) neither us nor any of our affiliates is permitted to directly 
or indirectly exploit any AbbVie Designated Antibody, in each case subject to specified exceptions, including AbbVie’s 
conduct of basic research. 

Termination 

Unless earlier terminated, the AbbVie Alpha-Synuclein Collaboration Agreement expires on the earliest to 

occur of the expiration of (1) the development option period, without AbbVie’s exercise of a development option; (2) the 
license option period, without AbbVie’s exercise of its license option; and (3) the last-to-expire royalty term with respect 
to all licensed products in all countries. Subject to a cure period, either party may terminate the AbbVie Alpha-Synuclein 
Collaboration Agreement, in whole or, in the case of us, in part, subject to specified conditions, in the event of the other 
party’s uncured material breach. Either party may also terminate, subject to specified conditions, for insolvency of the 
other party, certain failures or delays to obtain certain regulatory clearances of the collaboration, or a joint determination 
of scientific infeasibility by the parties. AbbVie may terminate the AbbVie Alpha-Synuclein Collaboration Agreement 
(1) without cause, in its entirety or, after its exercise of the license option, on a country-by-country basis, with 180 days’ 
prior written notice or (2) for our non-compliance with certain anti-bribery or anti-corruption covenants. We may 
terminate the AbbVie Alpha-Synuclein Collaboration Agreement, subject to specified conditions, if AbbVie or its 
affiliates challenge the validity or enforceability of certain of our or jointly-held intellectual property rights. 

38 

Upon termination in certain cases, the Vectorized Antibody Exclusivity and AbbVie Designated Exclusivity 

survives until the third anniversary of the termination date. If the parties mutually agree to terminate for infeasibility or 
AbbVie terminates for our failure to deliver a final research or development report, neither we nor any of our affiliates 
may directly or indirectly exploit a vectorized antibody compound that targets or binds to the alpha-synuclein protein for 
18 months after the termination date. 

License Agreement with University of Massachusetts 

On January 30, 2014, we entered into a license agreement with the University of Massachusetts, or UMass, 

pursuant to which UMass granted us an exclusive, worldwide, royalty-bearing license to certain of its licensed patents to 
make, have made, use, offer for sale, sell, have sold and import certain licensed products in the field of human diseases 
that use gene therapy applications. Our license is subject to any rights that may be required to be granted to the 
government of the United States, and UMass reserves the right to use the licensed patents for education and research and, 
with our consent, for non-commercial patient care, without the payment of any compensation to us.  

In consideration for rights granted to us under the agreement, we made an upfront payment of $0.2 million to 

UMass. We are obligated to pay UMass (i) low-single digit royalty payments based on net sales of the licensed products, 
(ii) annual maintenance payments of $30.0 thousand, which are creditable against royalties payable in such period, 
(iii) minimum aggregate annual royalty payments that are creditable against royalties payable in such period, with the 
minimum aggregate amount payable being in the low-six digits for each of the first four years of this agreement and a 
minimum aggregate amount payable being in the mid-six digits for each year, thereafter, (iv) milestone payments of up 
to $1.8 million, per licensed product for the first five licensed products, based on the achievement of development and 
regulatory milestones and (v) a percentage of sublicensing income that decreases over time from low double digit 
percentages to a mid-single digit percentage. We also agreed to reimburse UMass approximately $0.7 million for patent 
related expenses incurred by UMass as of the effective date of the agreement over a two-year period.  

Under the agreement, we agreed to use commercially reasonable efforts to develop licensed products and to 
introduce such licensed products into the commercial market, and further agreed to certain development milestones.  

The agreement will terminate on the date that is the later of (i) seven years after the first commercial sale of the 
last licensed product under the agreement or (ii) such time as there are no valid claims covering a licensed product. We 
have the right to terminate the agreement for any reason upon 90 days prior written notice, and we and UMass have the 
right to terminate the agreement if the other party fails to cure a written breach within 60 days of receiving written notice 
of such breach.  

MassBiologics and UMass Collaboration Agreement 

On October 20, 2014, we entered into a Collaboration Agreement with UMass and MassBiologics, pursuant to 
which we shall (i) fund certain projects that will be conducted by UMass or MassBiologics, (ii) fund certain educational 
programs of UMass, including post-doctoral research at our laboratories beginning in 2015 and an annual lecture series 
beginning in 2015 and (iii) collaborate with MassBiologics to establish scalable processes for manufacturing 
recombinant AAV vector products using cGMP.  

In November 2014, we agreed to the first project under this agreement whereby we funded approximately 

$2.9 million over a 16-month period for certain research and development services performed by MassBiologics. The 
project commenced in January 2015 and completed during 2016. We and UMass and/or MassBiologics may agree to 
conduct other projects in the future, the terms of which will be agreed upon at such time.  

This agreement will remain in effect for a period of five years and automatically renews for additional one-year 

periods. Either party has the right to terminate this agreement, once in each renewal period, for any reason upon 
providing the other party with 90 days written notice or in the event of a material breach of the agreement by the other 
party that is not cured within 60 days of written notice.  

39 

We will own all intellectual property rights generated under this agreement, either by our employees, UMass 
and/or MassBiologics employees, or jointly by our employees and UMass and/or MassBiologics employees, that cover 
AAV materials. We and UMass and/or MassBiologics, as applicable, will jointly own any intellectual property rights 
generated under this agreement jointly by our employees and the employees of UMass and/or MassBiologics, as 
applicable, that do not cover AAV materials.  

License Agreement with REGENX  

In May 2014, we entered into a license agreement with REGENXBIO Inc., formerly known as ReGenX 
Biosciences, LLC, or REGENX, for the development and commercialization of gene therapies to treat ALS, Friedreich’s 
ataxia and Huntington’s disease. Under this license agreement, REGENX granted us a non-exclusive worldwide license 
to make, have made and use its technology solely for internal research and preclinical development for the identification 
of specific vectors that could be commercialized. Following identification, we have an option to obtain a non-exclusive 
worldwide license under the licensed intellectual property to a single specified AAV vector to make, have made, use, 
import, sell and offer for sale licensed products using the selected vector, which can be exercised for each of ALS, 
Friedreich’s ataxia, or Huntington’s disease. Under the terms of this license agreement, we paid REGENX an upfront fee 
of $0.5 million, an extension fee of $0.1 million and are required to pay an annual maintenance fee.  

In November 2016, we exercised commercial options for the use of REGENX’s NAV® Technology Platform, 
or NAV, vectors for the development and commercialization of gene therapies for specific neurological diseases. Upon 
exercise of the options, REGENX has granted us a non-exclusive worldwide commercial license, with rights to 
sublicense, to three specific NAV vector sequences covered by REGENX’s NAV Technology Platform, each for the 
treatment of a specific neurological disease. In return for these rights, we paid upfront payments of $1.05 million and 
have paid to REGENX an annual maintenance fee payment of five digits based on the number of disease indication 
options exercised. In addition, we will be required to pay to REGENX up to $5.0 million in milestone fees per disease 
indication, mid- to high-single digit royalty percentages on net sales of licensed products, and low- to mid-single digit 
percentages of any sublicense fees that we receive from sublicensees for the licensed intellectual property rights.  

Our license agreement with REGENX will expire upon the expiration, lapse, abandonment, or invalidation of 

the last claim of the licensed intellectual property to expire, lapse, or become abandoned or unenforceable in all the 
countries of the world 

We may terminate the license agreement upon a specified number of days prior written notice. REGENX may 

terminate the license agreement if we, our affiliates, or sublicensees experience insolvency, if we are more than a 
specified number of days late in paying money due under the license agreement, or, effective immediately, if we or our 
affiliates commence any action against REGENX or its licensors to declare or render any claim of the licensed patent 
rights invalid or unenforceable. Either party may terminate the license agreement for material breach that is not cured 
within a specified number of days. 

In February 2019, we provided notice to REGENX of our intent to terminate our commercial license. Upon the 

effective date of the termination, we are required to grant to REGENX a non-exclusive, perpetual, irrevocable, 
worldwide, royalty-free, transferable, sublicensable license to certain improvements made by us during the term of the 
license (including any intellectual property rights with respect thereto). This grant back will allow for REGENX’s use of 
the improvements in the research, development or commercialization of products in any therapeutic indication. 

Competition 

The biopharmaceutical industry is characterized by intense and dynamic competition to develop new 
technologies and proprietary therapies. Any product candidates that we successfully develop into products and 
commercialize may compete with existing therapies and new therapies that may become available in the future. While 
we believe that our gene therapy platform, product programs, product candidates and scientific expertise in the fields of 
gene therapy and neuroscience provide us with competitive advantages, we face potential competition from various 

40 

sources, including larger and better-funded pharmaceutical, specialty pharmaceutical and biotechnology companies, as 
well as from academic institutions, governmental agencies and public and private research institutions.  

We are aware of several companies focused on developing AAV gene therapies in various indications, 
including Abeona Therapeutics, Inc., Adverum Biotechnologies, Inc., Aevitas Therapeutics, Inc., Agilis Biotherapeutics, 
LLC, which was acquired by PTC Therapeutics, Inc. in 2018, Applied Genetic Technologies Corporation, Asklepios 
BioPharmaceutical, Inc., Audentes Therapeutics, Inc., AveXis, Inc., or AveXis which was acquired by Novartis in 2018, 
Axovant Sciences Ltd., GenSight Biologics SA, LogicBio Therapeutics, Inc., Lysogene SA, MeiraGTx Ltd., NightstaRx 
Ltd, Prevail Therapeutics, Inc., REGENXBio Inc., Sarepta Therapeutics, Inc., Solid Biosciences, Inc., uniQure NV, or 
uniQure, Pfizer, Inc., or Pfizer, and Spark Therapeutics, Inc., or Spark, as well as several companies addressing other 
methods for modifying genes and regulating gene expression. Any advances in gene therapy technology made by a 
competitor may be used to develop therapies that could compete against any of our product candidates.  

We expect that VY-AADC will compete with a variety of therapies currently marketed and in development for 
Parkinson’s disease, including DBS marketed by Medtronic plc, Abbott Laboratories (acquired from St. Jude Medical in 
2017), and other medical device companies, DUOPA/Duodopa marketed by AbbVie Inc., as well as other novel, non-
oral forms of levodopa in development, including Mitsubishi Tanabe Pharma’s ND0612 (acquired from NeuroDerm in 
2017), Acorda Therapeutics’ inhaled levodopa, INBRIJA (CVT-301), and Sunovion Pharmaceuticals’, or Sunovion’s, 
sublingual apomorphine, APL-130277. Gene therapy competition for Parkinson’s disease previously included AMT-090 
or AAV-GDNF, but this was deprioritized by uniQure in 2016. Axovant Sciences Ltd. is developing a second generation 
LentiVector gene therapy, AXO-Lenti-PD (previously OXB-102, licensed from Oxford Biomedica in 2018). MeiraGTx 
is developing AAV-GAD, acquired from Vector Neurosciences, Inc. in 2018.  

We expect that our preclinical programs will compete with a variety of therapies in development, including: 

  VY-SOD102 for a monogenic form of ALS will potentially compete with IONIS-SOD1Rx being developed 
by Ionis Pharmaceuticals, Inc., or Ionis, in collaboration with Biogen Inc., or Biogen, and a gene therapy 
being developed by AveXis;  

  VY-FXN01 for Friedreich’s ataxia will potentially compete with AAV gene therapies being developed by 
Pfizer and by Agilis Biotherapeutics, LLC (acquired by PTC Therapeutics, Inc. in 2018), and BMN 290 
being developed by BioMarin Pharmaceutical Inc.;  

  VY-HTT01 for Huntington’s disease will potentially compete with IONIS-HTTRx being developed by 

Ionis in collaboration with F. Hoffmann-La Roche Ltd., or Roche, WVE-120101 and WVE-120102 being 
developed by WAVE Life Sciences in collaboration with Takeda Pharmaceuticals, a Zinc Finger Protein 
(ZFP) therapy being developed by Sangamo Therapeutics, Inc. in collaboration with Shire plc, and gene 
therapies being developed by uniQure and Spark;  

  Our Tau program for tauopathies including Alzheimer’s disease, PSP, and FTD will potentially compete 
with tau antibodies being developed by Roche Genentech Inc. in collaboration with AC Immune SA, Eli 
Lilly & Co., AbbVie Inc., Biogen, and several other companies, as well as an antisense oligonucleotide 
program being developed by Ionis in collaboration with Biogen; and 

  VY-NAV01 for severe, chronic pain will potentially compete with Nav1.7 inhibitors being developed by 

Biogen, Sunovion, Amgen, Inc., and Astellas Pharma Inc, and Nav1.8 inhibitors being developed by Vertex 
Pharmaceuticals, or Vertex.  

In addition, companies that are currently engaged in gene therapy for non-neurological diseases could at any 

time decide to develop gene therapies for neurological diseases.  

41 

Many of our competitors, either alone or with their strategic partners, have substantially greater financial, 

technical and human resources than we do and significantly greater experience in the discovery and development of 
product candidates, obtaining FDA and other regulatory approvals of product candidates and commercializing those 
product candidates. Accordingly, our competitors may be more successful than us in obtaining approval for product 
candidates and achieving widespread market acceptance. Our competitors’ product candidates may be more effective, or 
more effectively marketed and sold, than any product candidate we may commercialize and may render our treatments 
obsolete or non-competitive before we can recover the expenses of developing and commercializing any of our product 
candidates.  

Mergers and acquisitions in the biotechnology and pharmaceutical industries may result in even more resources 

being concentrated among a smaller number of our competitors. These competitors also compete with us in recruiting 
and retaining qualified scientific and management personnel and establishing clinical trial sites and subject registration 
for clinical trials, as well as in acquiring technologies complementary to, or necessary for, our programs. Smaller or 
early-stage companies may also prove to be significant competitors, particularly through collaborative arrangements with 
large and established companies.  

We anticipate that we will face intense and increasing competition as new product candidates enter the market 
and advanced technologies become available. We expect any product candidates that we develop and commercialize to 
compete on the basis of, among other things, efficacy, safety, convenience of administration and delivery, price, and the 
availability of reimbursement from government and other third-party payers.  

Our commercial opportunity could be reduced or eliminated if our competitors develop and commercialize 

products that are safer, more effective, have fewer or less severe side effects, are more convenient or are less expensive 
than any products that we may develop. Our competitors also may obtain FDA or other regulatory approval for their 
product candidates more rapidly than we may obtain approval for ours, which could result in our competitors 
establishing a strong market position before we are able to enter the market.  

Manufacturing 

The manufacture of gene therapy products is technically complex, and necessitates substantial expertise and 
capital investment. Production difficulties caused by unforeseen events may delay the availability of material for our 
clinical studies. To meet the requirements of our current and planned future trials we have developed a proprietary 
manufacturing platform that provides a robust and scalable process for AAV production. We are using the 
baculovirus/Sf9 AAV production system, a technology for producing AAV vectors at scale in insect-derived cells. We 
focus on developing internal processes and capabilities to produce high-yield and high-quality gene therapies. The 
process has been successfully transferred to our contract manufacturing organizations where it is used in manufacturing 
of clinical materials in accordance with the FDA’s cGMP. We have entered into agreements with Brammer Bio and 
Fujifilm Diosynth Biotechnologies to further expand our manufacturing capabilities to support the development of our 
gene therapy programs. We have also built an onsite, state-of-the-art process research and development facility to enable 
the manufacturing of high quality AAV gene therapy vectors at laboratory scale. 

We presently contract with third parties for the manufacturing of our program materials. We currently have no 

plans to build our own clinical or commercial scale manufacturing capabilities. The use of contracted manufacturing and 
reliance on collaboration partners is relatively cost efficient and has eliminated the need for our direct investment in 
manufacturing facilities and additional staff early in development. Although we rely on contract manufacturers, we have 
personnel with manufacturing and quality experience to oversee our contract manufacturers. 

42 

Intellectual Property 

Overview  

We strive to protect the proprietary technology, inventions, and know-how to enhance improvements that are 

commercially important to the development of our business, including seeking, maintaining, and defending patent rights, 
whether developed internally or licensed from third parties. We also rely on trade secrets and know-how relating to our 
proprietary technology platform, on continuing technological innovation and on in-licensing opportunities to develop, 
strengthen and maintain the strength of our position in the field of gene therapy that may be important for the 
development of our business. We additionally may rely on regulatory protection afforded through data exclusivity, 
market exclusivity and patent term extensions where available.  

Our commercial success may depend in part on our ability to: obtain and maintain patent and other protections 

for commercially important technology, inventions and know-how related to our business; defend and enforce our 
patents; preserve the confidentiality of our trade secrets; and operate without infringing the valid enforceable patents and 
intellectual property rights of third parties. Our ability to stop third parties from making, having made, using, selling, 
offering to sell or importing our products may depend on the extent to which we have rights under valid and enforceable 
licenses, patents or trade secrets that cover these activities. In some cases, these rights may need to be enforced by third-
party licensors. With respect to both licensed and company-owned intellectual property, we cannot be sure that patents 
will be granted with respect to any of our pending patent applications or with respect to any patent applications filed by 
us in the future, nor can we be sure that any of our existing patents or any patents that may be granted to us in the future 
will be commercially useful in protecting our commercial products and methods of manufacturing the same.  

We have 182 patent applications pending in the United States and foreign jurisdictions. At least 28 patent 

applications have been filed and are pending in the United States and foreign jurisdictions by or on behalf of universities 
which have granted us exclusive license rights to the technology. To date, 72 patents have issued to our licensors which 
have granted us exclusive license rights to the technology. To date, 148 patents have issued to our licensors which have 
granted us non-exclusive license rights to the technology with 64 applications pending. Our policy is to file patent 
applications to protect technology, inventions and improvements to inventions that are commercially important to the 
development of our business. We seek United States and international patent protection for a variety of technologies, 
including: research tools and methods, methods for transferring genetic material into cells, AAV-based biological 
products, methods of designing novel AAV constructs, methods for treating diseases of interest and methods for 
manufacturing our AAV-based products. We also intend to seek patent protection or rely upon trade secret rights to 
protect other technologies that may be used to discover and validate targets and that may be used to identify and develop 
novel biological products. We seek protection, in part, through confidentiality and proprietary information agreements. 
We are a party to various other license agreements that give us rights to use specific technologies in our research and 
development.  

Company-Owned Intellectual Property  

Parkinson’s Disease 

We own four pending patent families with one issued patent and 35 patent applications directed to AAV 

constructs encoding the gene AADC for therapeutic uses. Patents that grant from these patent families are generally 
expected to start to expire in 2035, subject to possible patent term extensions.  

ALS 

We own four pending patent families with 19 patent applications directed to targeting SOD1 for the treatment 
of ALS, and we have an ownership interest in a fifth patent family with 2 patent applications directed to pharmaceutical 
compositions and methods for the treatment of ALS to protect our intellectual property arising from a funded grant from 

43 

The Amyotrophic Lateral Sclerosis Association. Patents that grant from these patent families are generally expected to 
start to expire in 2035, subject to possible patent term extensions.  

Friedreich’s Ataxia  

We own two pending patent families with seven patent applications and we have an ownership interest in one 
pending patent family with two patent applications directed to AAVs encoding frataxin constructs for the treatment of 
Friedreich’s ataxia. Patents that grant from these patent families are generally expected to start to expire in 2036, subject 
to possible patent term extensions.  

Huntington’s Disease 

We own three pending patent families with 19 patent applications directed to pharmaceutical compositions and 

methods for targeting HTT for the treatment of Huntington’s disease, and we have an ownership interest in a fourth 
patent family with one patent application directed to protect our work for additional pharmaceutical compositions and 
methods for targeting HTT for the treatment of Huntington’s disease. Patents from this family are generally expected to 
start to expire in 2037, subject to possible patent term extensions. 

Tauopathies and Antibodies 

We own five pending patent families directed to antibodies with 11 patent applications. The first patent family 
has four patent applications directed to assays for the detection of neutralizing antibodies. The next three patent families 
have six patent applications directed to vectorized antibodies and other therapies. The last patent family has one patent 
application directed to vectored augmentation of proteins. Patents from these families are generally expected to start to 
expire in 2036, subject to possible patent term extensions. 

We also have one pending patent family with one patent application directed to pharmaceutical compositions 

and methods for the treatment of Alzheimer’s Disease. Patents from this family are generally expected to start to expire 
in 2039, subject to possible patent term extensions. 

Neuropathic Pain 

We own one pending patent family with two patent applications directed to pharmaceutical compositions and 

methods for the treatment of neuropathic pain. Patents from this family are generally expected to start to expire in 2039, 
subject to possible patent term extensions. 

Regulatable Expression 

We own two pending patent families with four patent applications directed to regulatable expression control of 

AAV transgenes. Patents that grant from this patent family are generally expected to start to expire in 2036, subject to 
possible patent term extensions.  

Delivery 

We own one pending patent family with five patent applications directed to the delivery of AAV gene therapies 
to the CNS. Patents that grant from this patent family are generally expected to start to expire in 2036, subject to possible 
patent term extensions.  

We own one pending patent family with one patent application directed to cannula delivery system and methods 

of use. Patents that grant from this patent family are generally expected to start to expire in 2039, subject to possible 
patent term extensions. 

44 

We have an ownership interest in two pending patent families directed to trajectory array delivery devices, 

including the V-TAG device and methods of use. Patents that grant from these patent families are generally expected to 
start to expire in 2037, subject to possible patent term extensions. 

Engineering  

We own seven pending patent families with 14 patent applications directed to AAV production and/or 
engineering of the capsid and we have an ownership interest in one patent family with one patent application directed to 
engineering of the capsid. Patents that grant from these patent families are generally expected to start to expire in 2035, 
subject to possible patent term extensions. 

We own three patent families with 33 patent applications directed to engineering of the vector genome. Patents 

that grant from these patent families are generally expected to start to expire in 2035, subject to possible patent term 
extensions. 

Production; Chemistry, Manufacturing, and Controls 

We own 14 pending patent families with 22 patent applications directed to AAV production and CMC. Patents 

that grant from this patent family are generally expected to start to expire in 2035, subject to possible patent term 
extensions. 

Licensed Intellectual Property  

We have obtained exclusive licenses and non-exclusive licenses to patents directed to both compositions of 

matter and methods of use.  

We have licensed six families of patents and patent applications, in the exclusive field of gene therapy for 

human diseases, directed to RNAi constructs as vector payloads, their design and use in the treatment of neurological 
disorders from the University of Massachusetts. These families of patents and applications are pending and/or granted in 
the United States and other territories and comprises 67 granted patents and 15 applications. Patents have been granted in 
the United States, Canada, Europe, Israel, Japan, Korea and Australia. Nationalization for some members has taken place 
in Germany, Spain, France, Great Britain, Italy, and Netherlands. Patents that grant from these patent families are 
generally expected to expire between 2022 and 2025, subject to possible patent term extensions.  

We have exclusively licensed three families of patents and patent applications directed to novel AAV capsids 
from the University of Massachusetts. These families of patents and applications, pending and/or granted in the United 
States and other territories, and comprises 32 granted patents and 18 applications. Patents have been granted in the 
United States, Europe and Japan. Nationalization for some members has taken place in Switzerland, Germany, Denmark, 
Spain, France, Great Britain, Ireland, Italy, Netherlands, and Sweden. Patents that grant from these patent families are 
generally expected to expire between 2030 and 2035, subject to possible patent term extensions.  

We have non-exclusively licensed a patent family directed to production methods for AAV in insect cells from 
the NIH, U.S. Department of Health and Human Services. This family of patents is granted in the United States, Canada, 
Australia and Europe and further nationalized in Germany, France and Great Britain and comprises 8 granted patents. 
Patents that grant from this patent family are generally expected to expire in 2022, subject to possible patent term 
extensions.  

We have non-exclusively licensed two families of patents and patent applications directed to novel AAV 

capsids from the Board of Trustees of the Leland Stanford Junior University. These families of patents and applications, 
pending and/or granted in the United States, comprise 7 granted patents and one application. Patents that grant from 
these patent families are generally expected to expire between 2027 and 2032, subject to possible patent term extensions.  

45 

We have non-exclusively licensed two families of patents and patent applications directed to AAV capsids from 

REGENX. These families of patents and patent applications are pending and/or granted in the United States and other 
territories and comprise 79 granted patents and 16 applications. Patents have been granted in Australia, Brazil, Canada, 
China, Europe, Hong Kong, Israel, India, Japan, Korea, Mexico, New Zealand, Philippines, Singapore, and the United 
States. Patents that grant from these patent families are generally expected to expire between 2022 and 2026, subject to 
possible patent term extensions. Notice of termination of this license was sent to REGENX in February 2019. 

We have non-exclusively licensed two families of patents and patent applications from Ablexis, LLC. These 

families of patents and patent applications are pending and/or granted in the United States and other territories and 
comprise 24 granted patents and 21 applications. Patents have been granted in Australia, Canada, Europe, Korea, New 
Zealand and the United States. Nationalization for some members has taken place in Austria, Belgium, Denmark, France, 
Germany, Ireland, Italy, Netherlands, Poland, Spain, Switzerland, and United Kingdom. Patents that grant from these 
patent families are generally expected to expire between 2029 and 2030, subject to possible patent term extensions. 

We have non-exclusively licensed two families of patents and patent applications directed to AAV capsids from 
the California Institute of Technology. These families of patents and patent applications are pending in the United States 
and internationally and comprise three granted patents and 21 applications. Patents have been granted in the United 
States. Patents that grant from these patent families are generally expected to start to expire in 2034, subject to possible 
patent term extensions. 

Trademark Protection  

We own U.S. Reg. Nos. 4,545,283 for the service mark VOYAGER THERAPEUTICS and 4,621,083 for the 
service mark VOYAGER THERAPEUTICS Logo for “pharmaceutical research and development in the field of gene 
therapy.” These marks were granted registration on the Principal Register of the United States Patent and Trademark 
Office, or USPTO, on June 3, 2014 and October 14, 2014, respectively. 

We also own pending trademark applications in the USPTO for the marks V-TAG and the V-TAG Logo, for 

“medical system comprised of a surgical device for guiding, locating or placing a diagnostic device or therapeutic 
device, namely, stents, probes, needles, leads, grafts, pumps, syringes, catheters, and implants during a medical 
procedure and related software sold as a unit, for use in the field of neurology; MRI-compatible medical system 
comprised of an MRI-compatible surgical device for guiding, locating or placing a diagnostic device or therapeutic 
device, namely, stents, probes, needles, leads, grafts, pumps, syringes, catheters, and implants during a MRI-guided 
procedure and related software sold as a unit, for use in the field of neurology,” as well as European Community 
trademark registrations for VOYAGER TRAJECTORY ARRAY GUIDE (No. 017430042, registered May 8, 2018) and 
V-TAG (No. 017430182, registered May 8, 2018)for these same goods.  

We plan to register trademarks in connection with our biological products.  

Trade Secret Protection  

Finally, we may rely, in some circumstances, on trade secrets to protect our technology. We seek to protect our 

proprietary technology and processes, in part, by entering into confidentiality agreements with our employees, 
consultants, scientific advisors and contractors. We also seek to preserve the integrity and confidentiality of our data and 
trade secrets by maintaining physical security of our premises and physical and electronic security of our information 
technology systems. While we have confidence in these individuals, organizations and systems, agreements or security 
measures may be breached, and we may not have adequate remedies for any breach. In addition, our trade secrets may 
otherwise become known or be independently discovered by competitors. To the extent that our consultants, contractors 
or collaborators use intellectual property owned by others in their work for us, disputes may arise as to the rights in 
related or resulting know-how and inventions.  

46 

Government Regulation and Product Approval 

In the United States, biological products, including gene therapy products, are licensed by FDA for marketing 

under the Public Health Service Act, or PHS Act, and regulated under the Federal Food, Drug, and Cosmetic Act, or 
FDCA. Both the FDCA and the PHS Act and their corresponding regulations govern, among other things, the testing, 
manufacturing, safety, purity, potency, efficacy, labeling, packaging, storage, record keeping, distribution, import, 
export, reporting, advertising and other promotional practices involving biological products. FDA clearance must be 
obtained before clinical testing of biological products, and each clinical study protocol for a gene therapy product is 
reviewed by the FDA and, in some instances, the NIH, through its RAC. FDA licensure also must be obtained before 
marketing of biological products. The process of obtaining regulatory approvals and the subsequent compliance with 
appropriate federal, state, local and foreign statutes and regulations require the expenditure of substantial time and 
financial resources.  

Within the FDA, the Center for Biologics Evaluation and Research, or CBER, regulates gene therapy products. 

Within CBER, the Office of Tissues and Advanced Therapies (OTAT) is responsible for gene therapy review and 
evaluation. CBER works closely with the NIH and its RAC, which makes recommendations to the NIH on gene therapy 
issues and engages in a public discussion of scientific, safety, ethical and societal issues related to proposed and ongoing 
gene therapy protocols. The FDA and the NIH have published guidance documents with respect to the development and 
submission of gene therapy protocols. The FDA also has published guidance documents related to, among other things, 
gene therapy products in general, their preclinical assessment, observing subjects involved in gene therapy studies for 
delayed adverse events, viral shedding, environmental assessments, potency testing, and chemistry, manufacturing and 
control information in gene therapy INDs. FDA guidance documents provide the agency’s current thinking about a 
particular subject but are not legally binding.  

U.S. Biological Products Development Process  

The process required by the FDA before a biological product may be marketed in the United States generally 

involves the following: 

 

completion of nonclinical laboratory tests and animal studies according to good laboratory practices, or 
GLPs, and applicable requirements for the humane use of laboratory animals or other applicable 
regulations;  

  preparation of clinical trial material in accordance with good manufacturing practices, or GMPs; 
 

submission to the FDA of an application for an IND, which must become effective before human clinical 
trials may begin;  
approval by an institutional review board, or IRB, reviewing each clinical site before each clinical trial may 
be initiated; 
approval by an institutional biosafety committee, or IBC, assessing the safety of the clinical research and 
identifying any potential risk to public health or the environment; 

 

 

  performance of adequate and well-controlled human clinical trials according to the FDA’s regulations 

 

 

commonly referred to as good clinical practice, or GCPs, and any additional requirements for the protection 
of human research subjects and their health information, to establish the safety, purity, potency, and 
efficacy, of the proposed biological product for its intended use;  
submission to the FDA of a BLA, for marketing approval that includes substantive evidence of safety, 
purity, potency, and efficacy from results of nonclinical testing and clinical trials;  
satisfactory completion of an FDA inspection prior to BLA approval of the manufacturing facility or 
facilities where the biological product is produced to assess compliance with cGMP, to assure that the 
facilities, methods and controls are adequate to preserve the biological product’s identity, strength, quality 
and purity;  

47 

  potential FDA audit of the nonclinical and clinical study sites that generated the data in support of the 

BLA;  

  potential FDA Advisory Committee meeting to elicit expert input on critical issues and including a vote by 

external Committee members;  

  FDA review and approval, or licensure, of the BLA, and payment of associated user fees; and 
 

compliance with any post approval requirements, including the potential requirement to implement a Risk 
Evaluation and Mitigation Strategy, or REMS, and the potential requirement to conduct post approval 
studies. 

Before testing any biological product candidate, including a gene therapy product, in humans, the product 

candidate enters the preclinical testing stage. Preclinical tests, also referred to as nonclinical tests, include laboratory 
evaluations of product chemistry, toxicity and formulation, as well as animal studies to assess the potential safety and 
activity of the product candidate. The conduct of the preclinical tests must comply with federal regulations and 
requirements including GLPs.  

Previously, when a gene therapy study was conducted at, or sponsored by, institutions receiving NIH funding 

for recombinant DNA research, prior to the submission of an IND to the FDA, a protocol and related documentation was 
to be submitted to and the study was registered with the NIH Office of Biotechnology Activities, or OBA, pursuant to 
the NIH Guidelines for Research Involving Recombinant DNA Molecules, or NIH Guidelines. Compliance with the NIH 
Guidelines was mandatory for investigators at institutions receiving NIH funds for research involving recombinant DNA, 
however many companies and other institutions not otherwise subject to the NIH Guidelines had voluntarily followed 
them. Under an FDA and NIH proposal in 2018, the role of the Recombinant DNA Advisory Committee, or RAC, in 
reviewing gene therapy protocols would be entirely eliminated and sponsors would no longer be required to submit 
reports to NIH on such protocols. Going forward, NIH says the RAC will continue to function as an advisory board to 
NIH on emerging fields such as gene editing, synthetic biology and neurotechnology.  

The clinical study sponsor must submit the results of the preclinical tests, together with manufacturing 
information, analytical data, any available clinical data or literature and a proposed clinical protocol, to the FDA as part 
of the IND. Some preclinical testing typically continues after the IND is submitted. An IND is an exemption from the 
FDCA that allows an unapproved product to be shipped in interstate commerce for use in an investigational clinical trial 
and a request for FDA authorization to administer an investigational product to humans. The IND automatically becomes 
effective 30 days after receipt by the FDA, unless the FDA requests certain changes to a protocol before the study can 
begin, or the FDA places the clinical study on a clinical hold within that 30-day time period. In such a case, the IND 
sponsor and the FDA must resolve any outstanding concerns before the clinical study can begin. With gene therapy 
protocols, if the FDA allows the IND to proceed, but the RAC decides that full public review of the protocol is 
warranted, the FDA will request at the completion of its IND review that sponsors delay initiation of the protocol until 
after completion of the RAC review process. The FDA may also impose clinical holds on a biological product candidate 
at any time before or during clinical trials due to safety concerns or non-compliance. If the FDA imposes a clinical hold, 
studies may not recommence without FDA authorization and then only under terms authorized by the FDA. 
Accordingly, we cannot be sure that submission of an IND will result in the FDA allowing clinical trials to begin, or that, 
once begun, issues will not arise that suspend or terminate such studies.  

Clinical trials involve the administration of the biological product candidate to healthy volunteers or subjects 

under the supervision of qualified investigators, generally physicians not employed by or under the study sponsor’s 
control. Clinical trials are conducted under protocols detailing, among other things, the objectives of the clinical study, 
dosing procedures, subject selection and exclusion criteria, and the parameters to be used to monitor subject safety, 
including stopping rules that assure a clinical study will be stopped if certain adverse events should occur. Each protocol 
and any amendments to the protocol must be submitted to the FDA as part of the IND. Clinical trials must be conducted 
and monitored in accordance with the FDA’s regulations comprising the GCP requirements, including the requirement 
that all research subjects provide informed consent. Further, each clinical study must be reviewed and approved by an 
independent IRB, at or servicing each institution at which the clinical study will be conducted. An IRB is charged with 
protecting the welfare and rights of study participants and considers such items as whether the risks to individuals 

48 

participating in the clinical trials are minimized and are reasonable in relation to anticipated benefits. The IRB also 
approves the form and content of the informed consent that must be signed by each clinical study subject or his or her 
legal representative and must monitor the clinical study until completed. Additionally, some trials are overseen by an 
independent group of qualified experts organized by the trial sponsor, known as a data safety monitoring board or 
committee. Clinical trials involving recombinant or synthetic (or both) nucleic acid molecules performed at or sponsored 
by an institution that receives any NIH funding for such research also must be reviewed by an IBC, a local institutional 
committee that reviews and oversees basic and clinical research conducted at that institution. The IBC assesses the safety 
of the research and identifies any potential risk to public health or the environment.  

Human clinical trials are typically conducted in three sequential phases that may overlap or be combined:  

  Phase 1. The biological product is initially introduced into healthy human subjects and tested for safety. In 
the case of some products for severe or life-threatening diseases, especially when the product may be too 
inherently toxic to ethically administer to healthy volunteers, the initial human testing is often conducted in 
patients. Guidelines on clinical trials with gene therapy products issued by OTAT state that the FDA has 
determined that the benefit-risk ratio of these products does not warrant their evaluation in healthy human 
subjects. 

  Phase 2. The biological product is evaluated in a limited patient population to identify possible adverse 

effects and safety risks, to preliminarily evaluate the efficacy of the product for specific targeted diseases 
and to determine dosage tolerance, optimal dosage and dosing schedule.  

  Phase 3. Clinical trials are undertaken to further evaluate dosage, clinical efficacy, potency and safety in an 
expanded patient population at geographically dispersed clinical trial sites. These clinical trials are intended 
to establish the overall risk/benefit ratio of the product and provide an adequate basis for product labeling.  

Post-approval clinical trials, sometimes referred to as Phase 4 clinical trials, may be conducted after initial 
marketing approval. These clinical trials are used to gain additional experience from the treatment of patients in the 
intended therapeutic indication, particularly for long-term safety follow-up. The FDA recommends that sponsors observe 
subjects for potential gene therapy-related delayed adverse events for a 15-year period, including a minimum of five 
years of annual examinations followed by ten years of annual queries, either in person or by questionnaire, of trial 
subjects.  

During all phases of clinical development, regulatory agencies require extensive monitoring and auditing of all 

clinical activities, clinical data, and clinical trial investigators. Annual progress reports detailing the results of the clinical 
trials must be submitted to the FDA. Written IND safety reports must be promptly submitted to the FDA, the NIH and 
the investigators for serious and unexpected adverse events, any findings from other studies, tests in laboratory animals 
or in vitro testing that suggest a significant risk for human subjects, or any clinically important increase in the rate of a 
serious suspected adverse reaction over that listed in the protocol or investigator brochure. The sponsor must submit an 
IND safety report within 15 calendar days after the sponsor determines that the information qualifies for reporting. The 
sponsor also must notify the FDA of any unexpected fatal or life-threatening suspected adverse reaction within seven 
calendar days after the sponsor’s initial receipt of the information. Phase 1, Phase 2, and Phase 3 clinical trials may not 
be completed successfully within any specified period, if at all. The FDA or the sponsor or its data safety monitoring 
board may suspend a clinical trial at any time on various grounds, including a finding that the research subjects or 
patients are being exposed to an unacceptable health risk. Similarly, an IRB can suspend or terminate approval of a 
clinical trial at its institution if the clinical trial is not being conducted in accordance with the IRB’s requirements or if 
the biological product has been associated with unexpected serious harm to patients.  

Human gene therapy products are a new category of therapeutics. Because this is a relatively new and 

expanding area of novel therapeutic interventions, there can be no assurance as to the length of the study period, the 
number of patients the FDA will require to be enrolled in the studies in order to establish the safety, efficacy, purity and 
potency of human gene therapy products, or that the data generated in these studies will be acceptable to the FDA to 
support marketing approval. The NIH and the FDA have a publicly accessible database, the Genetic Modification 
Clinical Research Information System which includes information on gene transfer studies and serves as an electronic 

49 

tool to facilitate the reporting and analysis of adverse events on these studies. The FDA has issued various guidance 
documents regarding gene therapies, including draft guidance documents released in July 2018 relating to gene therapies 
for human retinal disorders and gene therapies for rare diseases, and on January 15, 2019, the FDA issued a statement 
that it would issue additional guidance to facilitate the development of gene therapy products. 

Concurrent with clinical trials, companies usually complete additional animal studies and must also develop 

additional information about the physical characteristics of the biological product as well as finalize a process for 
manufacturing the product in commercial quantities in accordance with cGMP requirements. To help reduce the risk of 
the introduction of adventitious agents with use of biological products, the PHS Act emphasizes the importance of 
manufacturing control for products whose attributes cannot be precisely defined. The manufacturing process must be 
capable of consistently producing quality batches of the product candidate and, among other things, the sponsor must 
develop methods for testing the identity, strength, quality, potency and purity of the final biological product. 
Additionally, appropriate packaging must be selected and tested and stability studies must be conducted to demonstrate 
that the biological product candidate does not undergo unacceptable deterioration over its shelf life.  

Information about certain clinical trials must be submitted within specific timeframes to the NIH for public 

dissemination on its ClinicalTrials.gov website. 

Expanded Access to an Investigational Drug for Treatment Use 

Expanded access, sometimes called “compassionate use,” is the use of investigational new drug products 

outside of clinical trials to treat patients with serious or immediately life-threatening diseases or conditions when there 
are no comparable or satisfactory alternative treatment options. The rules and regulations related to expanded access are 
intended to improve access to investigational drugs for patients who may benefit from investigational therapies. FDA 
regulations allow access to investigational drugs under an IND by the company or the treating physician for treatment 
purposes on a case-by-case basis for: individual patients (single-patient IND applications for treatment in emergency 
settings and non-emergency settings); intermediate-size patient populations; and larger populations for use of the drug 
under a treatment protocol or Treatment IND Application. 

When considering an IND application for expanded access to an investigational product with the purpose of 
treating a patient or a group of patients, the sponsor and treating physicians or investigators will determine suitability 
when all of the following criteria apply: patient(s) have a serious or immediately life-threatening disease or condition, 
and there is no comparable or satisfactory alternative therapy to diagnose, monitor, or treat the disease or condition; the 
potential patient benefit justifies the potential risks of the treatment and the potential risks are not unreasonable in the 
context or condition to be treated; and the expanded use of the investigational drug for the requested treatment will not 
interfere initiation, conduct, or completion of clinical investigations that could support marketing approval of the product 
or otherwise compromise the potential development of the product. 

On December 13, 2016, the 21st Century Cures Act established (and the 2017 Food and Drug Administration 

Reauthorization Act later amended) a requirement that sponsors of one or more investigational drugs for the treatment of 
a serious disease(s) or condition(s) make publicly available their policy for evaluating and responding to requests for 
expanded access for individual patients. Although these requirements were rolled out over time, they have now come 
into full effect. This provision requires drug and biologic companies to make publicly available their policies for 
expanded access for individual patient access to products intended for serious diseases. 

Sponsors are required to make such policies publicly available upon the earlier of initiation of a Phase 2 or 

Phase 3 study; or 15 days after the drug or biologic receives designation as a breakthrough therapy, fast track product, or 
regenerative medicine advanced therapy. 

In addition, on May 30, 2018, the Right to Try Act was signed into law. The law, among other things, provides 
a federal framework for certain patients to access certain investigational new drug products that have completed a Phase 
I clinical trial and that are undergoing investigation for FDA approval. Under certain circumstances, eligible patients can 
seek treatment without enrolling in clinical trials and without obtaining FDA permission under the FDA expanded access 

50 

program. There is no obligation for a drug manufacturer to make its drug products available to eligible patients as a 
result of the Right to Try Act, but the manufacturer must develop an internal policy and respond to patient requests 
according to that policy. 

U.S. Review and Approval Processes  

After the completion of clinical trials of a biological product, FDA approval of a BLA, must be obtained before 
commercial marketing of the biological product. The BLA must include results of product development, laboratory and 
animal studies, human studies, information on the manufacture and composition of the product, proposed labeling and 
other relevant information. In addition, under the Pediatric Research Equity Act, or PREA, a BLA or supplement to a 
BLA must contain data to assess the safety and effectiveness of the biological product for the claimed indications in all 
relevant pediatric subpopulations and to support dosing and administration for each pediatric subpopulation for which 
the product is safe and effective. The FDA may grant deferrals for submission of data or full or partial waivers. Unless 
otherwise required by regulation, PREA does not apply to any biological product for an indication for which orphan 
designation has been granted. The testing and approval processes require substantial time and effort and there can be no 
assurance that the FDA will accept the BLA for filing and, even if filed, that any approval will be granted on a timely 
basis, if at all.  

Under the Prescription Drug User Fee Act, or PDUFA, as amended, each BLA must be accompanied by a 

significant user fee. Under federal law, the submission of most applications is subject to an application user fee, which 
for federal fiscal year 2019 is $2,588,478 for an application requiring clinical data. The sponsor of an approved 
application is also subject to an annual program fee, which for fiscal year 2019 is $309,915. Fee waivers or reductions 
are available in certain circumstances, including a waiver of the application fee for the first application filed by a small 
business. Additionally, no user fees are assessed on BLAs for product candidates designated as orphan drugs, unless the 
product candidate also includes a non-orphan indication.  

Within 60 days following submission of the application, the FDA reviews a BLA submitted to determine if it is 

substantially complete before the agency accepts it for filing. The FDA may refuse to file any BLA that it deems 
incomplete or not properly reviewable at the time of submission and may request additional information. In this event, 
the BLA must be resubmitted with the additional information. The resubmitted application also is subject to review 
before the FDA accepts it for filing. The application also needs to be published and submitted in an electronic format that 
can be processed through the FDA’s electronic systems. If the electronic submission is not compatible with FDA’s 
systems, the BLA can be refused to file. Once the submission is accepted for filing, the FDA begins an in-depth 
substantive review of the BLA. The FDA reviews the BLA to determine, among other things, whether the proposed 
product is safe, potent, and effective, for its intended use, and has an acceptable purity profile, and whether the product is 
being manufactured in accordance with cGMP to assure and preserve the product’s identity, safety, strength, quality, 
potency and purity. The FDA may refer applications for novel biological products or biological products that present 
difficult questions of safety or efficacy to an advisory committee, typically a panel that includes clinicians and other 
experts, for review, evaluation and a recommendation as to whether the application should be approved and under what 
conditions. The FDA is not bound by the recommendations of an advisory committee, but it considers such 
recommendations carefully when making decisions. During the biological product approval process, the FDA also will 
determine whether a Risk Evaluation and Mitigation Strategy, or REMS, is necessary to assure the safe use of the 
biological product. If the FDA concludes a REMS is needed, the sponsor of the BLA must submit a proposed REMS; the 
FDA will not approve the BLA without a REMS, if required.  

Before approving a BLA, the FDA will inspect the facilities at which the product is manufactured. The FDA 

will not approve the product unless it determines that the manufacturing processes and facilities are in compliance with 
cGMP requirements and adequate to assure consistent production of the product within required specifications. 
Additionally, before approving a BLA, the FDA will typically inspect one or more clinical trial sites to assure that the 
clinical trials were conducted in compliance with IND study requirements and GCP requirements. To assure cGMP and 
GCP compliance, an applicant must incur significant expenditure of time, money and effort in the areas of training, 
record keeping, production, and quality control.  

51 

Notwithstanding the submission of relevant data and information, the FDA may ultimately decide that the BLA 

does not satisfy its regulatory criteria for approval and deny approval. Data obtained from clinical trials are not always 
conclusive and the FDA may interpret data differently than we interpret the same data. If the agency decides not to 
approve the BLA in its present form, the FDA will issue a complete response letter that usually describes all of the 
specific deficiencies in the BLA identified by the FDA. The deficiencies identified may be minor, for example, requiring 
labeling changes, or major, for example, requiring additional clinical trials. Additionally, the complete response letter 
may include recommended actions that the applicant might take to place the application in a condition for approval. If a 
complete response letter is issued, the applicant may either resubmit the BLA, addressing all of the deficiencies 
identified in the letter, or withdraw the application.  

If a product receives regulatory approval, the approval may be significantly limited to specific diseases and 

dosages or the indications for use may otherwise be limited, which could restrict the commercial value of the product. 
Further, the FDA may require that certain contraindications, warnings or precautions be included in the product labeling. 
The FDA may impose restrictions and conditions on product distribution, prescribing, or dispensing in the form of a risk 
management plan, or otherwise limit the scope of any approval. In addition, the FDA may require post marketing clinical 
trials, sometimes referred to as Phase 4 clinical trials, designed to further assess a biological product’s safety and 
effectiveness, and testing and surveillance programs to monitor the safety of approved products that have been 
commercialized. As a condition for approval, the FDA may also require additional non-clinical testing as a Phase 4 
commitment.  

One of the performance goals agreed to by the FDA under the PDUFA is to review standard BLAs in 10 months 

from filing and priority BLAs in six months from filing, whereupon a review decision is to be made. The FDA does not 
always meet its PDUFA goal dates for standard and priority BLAs and its review goals are subject to change from time 
to time. The review process and the PDUFA goal date may be extended by three months if the FDA requests or the BLA 
sponsor otherwise provides additional information or clarification regarding information already provided in the 
submission within the last three months before the PDUFA goal date.  

Post-Approval Requirements  

Maintaining substantial compliance with applicable federal, state, and local statutes and regulations requires the 

expenditure of substantial time and financial resources. Rigorous and extensive FDA regulation of biological products 
continues after approval, particularly with respect to cGMP. We will rely, and expect to continue to rely, on third parties 
for the production of clinical and commercial quantities of any products that we may commercialize. Manufacturers of 
our products are required to comply with applicable requirements in the cGMP regulations, including quality control and 
quality assurance and maintenance of records and documentation. Following approval, the manufacturing facilities are 
subject to biennial inspections by the FDA’s biologics team and such inspections may result in an issuance of FDA Form 
483 deficiency observations or a warning letter, which can lead to plant shutdown and other more serious penalties and 
fines. Prior to the institution of any manufacturing changes, a determination needs to be made whether FDA approval is 
required in advance. If not done in accordance with FDA expectations, the FDA may restrict supply and may take further 
action. Annual product reports are required to be submitted annually. Other post-approval requirements applicable to 
biological products, include reporting of cGMP deviations that may affect the identity, potency, purity and overall safety 
of a distributed product, record-keeping requirements, reporting of adverse effects, reporting updated safety and efficacy 
information, and complying with electronic record and signature requirements. After a BLA is approved, the product 
also may be subject to official lot release. As part of the manufacturing process, the manufacturer is required to perform 
certain tests on each lot of the product before it is released for distribution. If the product is subject to official release by 
the FDA, the manufacturer submits samples of each lot of product to the FDA together with a release protocol showing a 
summary of the history of manufacture of the lot and the results of all of the manufacturer’s tests performed on the lot. 
The FDA also may perform certain confirmatory tests on lots of some products, such as viral vaccines, before releasing 
the lots for distribution by the manufacturer. In addition, the FDA conducts laboratory research related to the regulatory 
standards on the safety, purity, potency, and effectiveness of biological products. Systems need to be put in place to 
record and evaluate adverse events reported by health care providers and patients and to assess product complaints. An 
increase in severity or new adverse events can result in labeling changes or product recall. Defects in manufacturing of 
commercial products can result in product recalls.  

52 

We also must comply with the FDA’s advertising and promotion requirements, such as those related to direct-
to-consumer advertising, the prohibition on promoting products for uses or in patient populations that are not described 
in the product’s approved labeling (known as “off-label use”), industry-sponsored scientific and educational activities, 
and promotional activities involving the internet. Discovery of previously unknown problems or the failure to comply 
with the applicable regulatory requirements may result in restrictions on the marketing of a product or withdrawal of the 
product from the market as well as possible civil or criminal sanctions. Failure to comply with the applicable U.S. 
requirements at any time during the product development process, approval process or after approval, may subject an 
applicant or manufacturer to administrative or judicial civil or criminal sanctions and adverse publicity. FDA sanctions 
could include refusal to approve pending applications, withdrawal of an approval or license revocation, clinical hold, 
warning or untitled letters, product recalls, product seizures, total or partial suspension of production or distribution, 
injunctions, fines, refusals of government contracts, mandated corrective advertising or communications with doctors, 
debarment, restitution, disgorgement of profits, or civil or criminal penalties. Any agency or judicial enforcement action 
could have a material adverse effect on us.  

Biological product manufacturers and other entities involved in the manufacture and distribution of approved 

biological products are required to register their establishments with the FDA and certain state agencies, and are subject 
to periodic unannounced inspections by the FDA and certain state agencies for compliance with cGMPs and other laws. 
Accordingly, manufacturers must continue to expend time, money, and effort in the area of production and quality 
control to maintain cGMP compliance. Discovery of problems with a product after approval may result in restrictions on 
a product, manufacturer, or holder of an approved BLA, including withdrawal of the product from the market. In 
addition, changes to the manufacturing process or facility generally require prior FDA approval before being 
implemented and other types of changes to the approved product, such as adding new indications and additional labeling 
claims, are also subject to further FDA review and approval.  

Orphan Drug Designation  

Under the Orphan Drug Act, the FDA may grant Orphan Drug Designation, or ODD, to a drug or biological 
product intended to treat a rare disease or condition, which is generally a disease or condition that affects fewer than 
200,000 individuals in the United States, or more than 200,000 individuals in the United States and for which there is no 
reasonable expectation that the cost of developing and making a drug or biological product available in the United States 
for this type of disease or condition will be recovered from sales of the product. ODD must be requested before 
submitting a BLA. After the FDA grants ODD, the identity of the therapeutic agent and its potential orphan use are 
disclosed publicly by the FDA. ODD does not convey any advantage in or shorten the duration of the regulatory review 
and approval process.  

If a product that has ODD receives the first FDA approval for the disease or condition for which it has such 
designation, the product is entitled to orphan product exclusivity, which means that the FDA may not approve any other 
applications to market the same biological product for the same indication for seven years, except in limited 
circumstances, such as not being able to supply the product for patients or showing clinical superiority to the product 
with orphan exclusivity. This is the case despite an earlier court opinion holding that the Orphan Drug Act 
unambiguously required the FDA to recognize orphan exclusivity regardless of a showing of clinical superiority. 

Competitors, however, may receive approval of different products for the indication for which the orphan 
product has exclusivity or obtain approval for the same product but for a different indication for which the orphan 
product has exclusivity. Orphan product exclusivity also could block the approval of one of our products for seven years 
if a competitor obtains approval of the same biological product as defined by the FDA or if our product candidate is 
determined to be contained within the competitor’s product for the same indication or disease. If a biological product 
designated as an orphan product receives marketing approval for an indication broader than what is designated, it may 
not be entitled to orphan product exclusivity. 

53 

Expedited Review and Approval Programs  

The FDA has various programs, including fast track designation, priority review, accelerated approval, and 
breakthrough therapy designation, that are intended to expedite or simplify the process for the development and FDA 
review of biological products that are intended for the treatment of serious or life-threatening diseases or conditions and 
demonstrate the potential to address unmet medical needs. The purpose of these programs is to provide important new 
biological products to patients earlier than under standard FDA review procedures. To be eligible for a fast track 
designation, the FDA must determine, based on the request of a sponsor, that a biological product is intended to treat a 
serious or life-threatening disease or condition and demonstrates the potential to address an unmet medical need. The 
FDA will determine that a product will fill an unmet medical need if it will provide a therapy where none exists or 
provide a therapy that may be potentially superior to existing therapy based on efficacy or safety factors. In addition to 
other benefits, such as the ability to have greater interactions with the FDA, the FDA may initiate review of sections of a 
Fast Track BLA before the application is complete, a process known as rolling review.  

The FDA may give a priority review designation to biological products that treats a serious condition and, if 

approved, would provide a significant improvement in safety or effectiveness. A priority review means that the goal for 
the FDA to review an application is six months, rather than the standard review of ten months under current PDUFA 
guidelines. Most products that are eligible for fast track designation may also be considered appropriate to receive a 
priority review.  

In addition, biological products studied for their safety and effectiveness in treating serious or life-threatening 
illnesses and that provide meaningful therapeutic benefit over existing treatments may receive accelerated approval and 
may be approved on the basis of adequate and well-controlled clinical trials establishing that the biological product has 
an effect on a surrogate endpoint that is reasonably likely to predict clinical benefit, or on a clinical endpoint that can be 
measured earlier than irreversible morbidity or mortality, that is reasonably likely to predict an effect on irreversible 
morbidity or mortality or other clinical benefit, taking into account the severity, rarity or prevalence of the condition and 
the availability or lack of alternative treatments. As a condition of approval, the FDA may require a sponsor of a 
biological product receiving accelerated approval to perform post-marketing studies to verify and describe the predicted 
effect on irreversible morbidity or mortality or other clinical endpoint, and the biological product may be subject to 
accelerated withdrawal procedures.  

Moreover, under the Food and Drug Administration Safety and Innovation Act enacted in 2012, a sponsor can 

request designation of a product candidate as a “breakthrough therapy.” A breakthrough therapy is defined as a 
biological product that is intended, alone or in combination with one or more other drugs, to treat a serious or life-
threatening disease or condition, and preliminary clinical evidence indicates that the biological product may demonstrate 
substantial improvement over existing therapies on one or more clinically significant endpoints, such as substantial 
treatment effects observed early in clinical development. Biological products designated as breakthrough therapies are 
also eligible for accelerated approval. The FDA must take certain actions, such as holding timely meetings and providing 
advice, intended to expedite the development and review of an application for approval of a breakthrough therapy.  

Even if a product qualifies for one or more of these programs, the FDA may later decide that the product no 

longer meets the conditions for qualification or decide that the time period for FDA review or approval will not be 
shortened. Furthermore, fast track designation, priority review, accelerated approval and breakthrough therapy 
designation, do not change the standards for approval and may not ultimately expedite the development or approval 
process.  

Expedited programs for regenerative medicine therapies for serious conditions  

As part of the 21st Century Cures Act, Congress amended the FDCA to create an expedited development and 

approval program for regenerative medicine advanced therapies, which include cell therapies, therapeutic tissue 
engineering products, human cell and tissue products, and combination products using any such therapies or products. As 
of November 2017, the FDA has interpreted this definition as follows: gene therapies, including genetically modified 
cells, that lead to a durable modification of cells or tissues may meet the definition of a regenerative medicine advanced 

54 

therapy. The FDA has now determined that ‘in vitro’ gene therapies will qualify as a regenerative medicine advanced 
therapy based on this definition. VY-AADC is an ‘in vitro’ gene therapy for Parkinson’s disease and received this 
designation on June 18, 2018. Regenerative medicine advanced therapies do not include those human cells, tissues, and 
cellular and tissue-based products regulated solely under section 361 of the Public Health Service Act and 21 CFR Part 
1271. The new program is intended to facilitate efficient development and expedite review of regenerative medicine 
advanced therapies, which are intended to treat, modify, reverse, or cure a serious or life-threatening disease or 
condition.  

A drug sponsor may request that the FDA designate a drug as a regenerative medicine advanced therapy 

concurrently with or at any time after submission of an IND. The FDA has 60 calendar days to determine whether the 
drug meets the criteria, including whether there is preliminary clinical evidence indicating that the drug has the potential 
to address unmet medical needs for a serious or life-threatening disease or condition. A new drug application or BLA for 
a regenerative medicine advanced therapy may be eligible for priority review or accelerated approval through (1) 
surrogate or intermediate endpoints reasonably likely to predict long-term clinical benefit or (2) reliance upon data 
obtained from a meaningful number of sites. Benefits of such designation also include early interactions with the FDA to 
discuss any potential surrogate or intermediate endpoint to be used to support accelerated approval. A regenerative 
medicine advanced therapy that is granted accelerated approval and is subject to post-approval requirements may fulfill 
such requirements through the submission of clinical evidence, clinical studies, patient registries, or other sources of real 
world evidence, such as electronic health records; the collection of larger confirmatory data sets; or post-approval 
monitoring of all patients treated with such therapy prior to its approval. 

U.S. Patent Term Restoration  

Depending upon the timing, duration and specifics of the FDA approval of the use of our product candidates, 

some of our U.S. patents may be eligible for limited patent term extension under the Drug Price Competition and Patent 
Term Restoration Act of 1984, commonly referred to as the Hatch-Waxman Amendments. The Hatch-Waxman 
Amendments permit a patent restoration term of up to five years as compensation for patent term lost during product 
development and the FDA regulatory review process. However, patent term restoration cannot extend the remaining term 
of a patent beyond a total of 14 years from the product’s approval date. The patent term restoration period is generally 
one-half the time between the effective date of an IND and the submission date of a BLA plus the time between the 
submission date of a BLA and the approval of that application, less any time the applicant failed to act with due 
diligence. Only one patent applicable to an approved biological product is eligible for the extension and the application 
for the extension must be submitted prior to the expiration of the patent. The U.S. Patent and Trademark Office, in 
consultation with the FDA, reviews and approves the application for any patent term extension or restoration. In the 
future, we may intend to apply for restoration of patent term for one of our currently owned or licensed patents to add 
patent life beyond its current expiration date, depending on the expected length of the clinical trials and other factors 
involved in the filing of the relevant BLA.  

Market and Data Exclusivity 

The 2010 Patient Protection and Affordable Care Act, or the ACA, which was signed into law on March 23, 
2010, included a subtitle called the Biologics Price Competition and Innovation Act of 2009 or BPCIA. The BPCIA 
established a regulatory scheme authorizing the FDA to approve biosimilars and interchangeable biosimilars. As of 
January 1, 2019, the FDA has approved 17 biosimilar products for use in the United States. No interchangeable 
biosimilars have been approved. The FDA has issued several guidance documents outlining an approach to review and 
approval of biosimilars. Additional guidance is expected to be finalized by FDA in the near term. 

Under the BPCIA, a manufacturer may submit an application for licensure of a biologic product that is 
“biosimilar to” or “interchangeable with” a previously approved biological product or “reference product.” In order for 
the FDA to approve a biosimilar product, it must find that there are no clinically meaningful differences between the 
reference product and proposed biosimilar product in terms of safety, purity and potency. For the FDA to approve a 
biosimilar product as interchangeable with a reference product, the FDA must find that the biosimilar product can be 
expected to produce the same clinical results as the reference product, and (for products administered multiple times) that 

55 

the biologic and the reference biologic may be switched after one has been previously administered without increasing 
safety risks or risks of diminished efficacy relative to exclusive use of the reference biologic. 

Under the BPCIA, an application for a biosimilar product may not be submitted to the FDA until four years 

following the date of approval of the reference product. The FDA may not approve a biosimilar product until 12 years 
from the date on which the reference product was approved. Even if a product is considered to be a reference product 
eligible for exclusivity, another company could market a competing version of that product if the FDA approves a full 
BLA for such product containing the sponsor’s own preclinical data and data from adequate and well-controlled clinical 
trials to demonstrate the safety, purity and potency of their product. The BPCIA also created certain exclusivity periods 
for biosimilars approved as interchangeable products. At this juncture, it is unclear whether products deemed 
“interchangeable” by the FDA will, in fact, be readily substituted by pharmacies, which are governed by state pharmacy 
law. 

Pediatric Exclusivity 

Pediatric exclusivity is another type of non-patent exclusivity in the United States and, if granted, provides for 
the attachment of an additional six months of marketing protection to the term of any existing regulatory exclusivity or 
patent protection, including the non-patent and orphan exclusivity. This six-month exclusivity may be granted if an 
application sponsor submits pediatric data that fairly respond to a written request from the FDA for such data. The data 
do not need to show the product to be effective in the pediatric population studied; rather, if the clinical trial is deemed to 
fairly respond to the FDA’s request, the additional protection is granted. If reports of requested pediatric studies are 
submitted to and accepted by the FDA within the statutory time limits, whatever statutory or regulatory periods of 
exclusivity or patent protection cover the product are extended by six months. Thus, pediatric exclusivity adds six 
months to existing exclusivity periods applicable to biological products under the BPCIA—namely, the four-year period 
during which the FDA will not consider an application for a biosimilar product, and the 12-year period during which the 
FDA will not approve a biosimilar application. 

Other Healthcare Laws  

Although we currently do not have any products on the market, we may be subject to additional healthcare 

regulation and enforcement by the federal government and by authorities in the states in which we conduct our business. 
Such laws include, without limitation, state and federal anti-kickback, fraud and abuse, false claims, privacy and security 
and physician sunshine laws and regulations, many of which may become more applicable to us if our product 
candidates are approved and we begin commercialization. If our operations are found to be in violation of any of such 
laws or any other governmental regulations that apply to us, we may be subject to penalties, including, without 
limitation, administrative, civil and criminal penalties, damages, fines, disgorgement, the curtailment or restructuring of 
our operations, exclusion from participation in federal and state healthcare programs and imprisonment, any of which 
could adversely affect our ability to operate our business and our financial results. 

In addition, the ACA is intended to broaden access to health insurance, reduce or constrain the growth of 

healthcare spending, enhance remedies against fraud and abuse, add transparency requirements for the healthcare and 
health insurance industries, impose taxes and fees on the health industry and impose additional health policy reforms. 
With regard to biopharmaceutical products, in addition to the Biologics Price Competition and Innovation Act of 2009 
included in the ACA, among other things, the ACA expanded and increased industry rebates for drugs covered under 
Medicaid programs and made changes to the coverage requirements under the Medicare prescription drug benefit. Some 
of the provisions of the ACA have yet to be fully implemented, while certain provisions have been subject to judicial and 
Congressional challenges. In January 2017, Congress voted to adopt a budget resolution for fiscal year 2017, that while 
not a law, is widely viewed as the first step toward the passage of legislation that would repeal certain aspects of the 
ACA.  

The current administration has also taken executive actions to undermine or delay implementation of the ACA. 

Since January 2017, President Trump has signed two Executive Orders which delay the implementation of certain 
provisions of the ACA or otherwise circumvent some of the requirements for health insurance mandated by the ACA. 

56 

One Executive Order directs federal agencies with authorities and responsibilities under the ACA to waive, defer, grant 
exemptions from, or delay the implementation of any provision of the ACA that would impose a fiscal or regulatory 
burden on states, individuals, healthcare providers, health insurers, or manufacturers of pharmaceuticals or medical 
devices. The second Executive Order terminates the cost-sharing subsidies that reimburse insurers under the ACA. 
Several state Attorneys General filed suit to stop the Trump administration from terminating the subsidies, but their 
request for a restraining order was denied by a federal judge in California on October 25, 2017. In addition, the Center 
for Medicare & Medicaid Services has recently proposed regulations that would give states greater flexibility in setting 
benchmarks for insurers in the individual and small group marketplaces, which may have the effect of relaxing the 
essential health benefits required under the ACA for plans sold through such marketplaces. Further, on June 14, 2018, 
U.S. Court of Appeals for the Federal Circuit ruled that the federal government was not required to pay more than $12 
billion in ACA risk corridor payments to third-party payors who argued were owed to them. The effects of this gap in 
reimbursement on third-party payors, the viability of the ACA marketplace, providers, and potentially our business, are 
not yet known. 

With enactment of the Tax Cuts and Jobs Act of 2017, which was signed by the President on December 22, 

2017, Congress repealed the “individual mandate.”  The repeal of this provision, which requires most Americans to carry 
a minimal level of health insurance, became effective January 1, 2019. According to the Congressional Budget Office, 
the repeal of the individual mandate will cause 13 million fewer Americans to be insured in 2027 and premiums in 
insurance markets may rise. Additionally, on January 22, 2018, President Trump signed a continuing resolution on 
appropriations for fiscal year 2018 that delayed the implementation of certain ACA-mandated fees, including the so-
called “Cadillac” tax on certain high cost employer-sponsored insurance plans, the annual fee imposed on certain health 
insurance providers based on market share, and the medical device excise tax on non-exempt medical devices. 

Further, there have been several recent U.S. congressional inquiries and proposed federal and proposed and 

enacted state legislation designed to, among other things, bring more transparency to drug pricing, review the 
relationship between pricing and manufacturer patient programs, reduce the costs of drugs under Medicare and reform 
government program reimbursement methodologies for drug products. For example, there have been several recent U.S. 
congressional inquiries and proposed federal and proposed and enacted state legislation designed to, among other things, 
bring more transparency to drug pricing, review the relationship between pricing and manufacturer patient programs, 
reduce the costs of drugs under Medicare and reform government program reimbursement methodologies for drug 
products. At the federal level, Congress and the Trump administration have each indicated that it will continue to seek 
new legislative and/or administrative measures to control drug costs. For example, on May 11, 2018, the Trump 
administration issued a plan to lower drug prices. Under this blueprint for action, the Trump administration indicated that 
the Department of Health and Human Services will: take steps to end the gaming of regulatory and patent processes by 
drug makers to unfairly protect monopolies; advance biosimilars and generics to boost price competition; evaluate the 
inclusion of prices in drug makers’ ads to enhance price competition; speed access to and lower the cost of new drugs by 
clarifying policies for sharing information between insurers and drug makers; avoid excessive pricing by relying more on 
value-based pricing by expanding outcome-based payments in Medicare and Medicaid; work to give Part D plan 
sponsors more negotiation power with drug makers; examine which Medicare Part B drugs could be negotiated for a 
lower price by Part D plans, and improving the design of the Part B Competitive Acquisition Program; update 
Medicare’s drug-pricing dashboard to increase transparency; prohibit Part D contracts that include “gag rules” that 
prevent pharmacists from informing patients when they could pay less out-of-pocket by not using insurance; and require 
that Part D plan members be provided with an annual statement of plan payments, out-of-pocket spending, and drug 
price increases. 

At the state level, individual states are increasingly aggressive in passing legislation and implementing 
regulations designed to control pharmaceutical and biological product pricing, including price or patient reimbursement 
constraints, discounts, restrictions on certain product access and marketing cost disclosure and transparency measures, 
and, in some cases, designed to encourage importation from other countries and bulk purchasing. In addition, regional 
health care authorities and individual hospitals are increasingly using bidding procedures to determine what 
pharmaceutical products and which suppliers will be included in their prescription product and other health care 
programs. These measures could reduce the ultimate demand for our products, once approved, or put pressure on our 

57 

product pricing. We expect that additional state and federal healthcare reform measures will be adopted in the future, any 
of which could limit the amounts that federal and state governments will pay for healthcare products and services, which 
could result in reduced demand for our product candidates or additional pricing pressures.  

Additional Regulation  

In addition to the foregoing, state and federal laws regarding environmental protection and hazardous 

substances, including the Occupational Safety and Health Act, the Resource Conservancy and Recovery Act and the 
Toxic Substances Control Act, affect our business. These and other laws govern our use, handling and disposal of 
various biological, chemical and radioactive substances used in, and wastes generated by, our operations. If our 
operations result in contamination of the environment or expose individuals to hazardous substances, we could be liable 
for damages and governmental fines. We believe that we are in material compliance with applicable environmental laws 
and that continued compliance therewith will not have a material adverse effect on our business. We cannot predict, 
however, how changes in these laws may affect our future operations.  

U.S. Foreign Corrupt Practices Act  

The U.S. Foreign Corrupt Practices Act, to which we are subject, prohibits corporations and individuals from 
engaging in certain activities to obtain or retain business or to influence a person working in an official capacity. It is 
illegal to pay, offer to pay or authorize the payment of anything of value to any foreign government official, government 
staff member, political party or political candidate in an attempt to obtain or retain business or to otherwise influence a 
person working in an official capacity. 

Review and Clearance of Companion Diagnostics in the United States  

If safe and effective use of a therapeutic depends on an in vitro diagnostic, then the FDA generally will require 
approval or clearance of that diagnostic, known as a companion diagnostic, at the same time that the FDA approves the 
therapeutic product. In August 2014, the FDA issued final guidance clarifying the requirements that will apply to 
approval of therapeutic products and in vitro companion diagnostics. According to the guidance, for novel drugs, a 
companion diagnostic device and its corresponding therapeutic should be approved or cleared contemporaneously by the 
FDA for the use indicated in the therapeutic product’s labeling. In July 2016, the FDA issued a draft guidance intended 
to assist sponsors of the drug therapeutic and in vitro companion diagnostic device on issues related to co-development 
of the products. 

If FDA determines that a companion diagnostic device or delivery device (combination product) is essential to 

the safe and effective use of a novel therapeutic product or indication, FDA generally will not approve the therapeutic 
product or new therapeutic product indication if the companion diagnostic or delivery device is not approved or cleared 
for that indication. Approval or clearance of the companion diagnostic or delivery device will ensure that the device has 
been adequately evaluated and has adequate performance characteristics in the intended population. The review of in 
vitro companion diagnostics in conjunction with the review of our therapeutic treatments for cancer will, therefore, likely 
involve coordination of review by the FDA’s Center for Drug Evaluation and Research and the FDA’s CDRH Office. 

Under the FDCA, in vitro diagnostics, including companion diagnostics, are regulated as medical devices. In 

the United States, the FDCA and its implementing regulations, and other federal and state statutes and regulations 
govern, among other things, medical device design and development, preclinical and clinical testing, premarket clearance 
or approval, registration and listing, manufacturing, labeling, storage, advertising and promotion, sales and distribution, 
export and import, and post-market surveillance. Unless an exemption applies, diagnostic tests require marketing 
clearance or approval from the FDA prior to commercial distribution. The two primary types of FDA marketing 
authorization applicable to a medical device are premarket notification, also called 510(k) clearance, and premarket 
approval, or PMA approval.  

The PMA process, including the gathering of clinical and preclinical data and the submission to and review by 

the FDA, can take several years or longer. It involves a rigorous premarket review during which the applicant must 

58 

prepare and provide the FDA with reasonable assurance of the device’s safety and effectiveness and information about 
the device and its components regarding, among other things, device design, manufacturing and labeling. PMA 
applications are subject to fees for medical device product review. For federal fiscal year 2019, the standard fee for 
review of a PMA was $322,147 and the small business fee was $80,537.  

A 510(k) must demonstrate that the proposed device is substantially equivalent to another legally marketed 
device, or predicate device, that did not require premarket approval. In evaluating a 510(k), the FDA will determine 
whether the device has the same intended use as the predicate device, and (a) has the same technological characteristics 
as the predicate device, or (b) has different technological characteristics, and (i) the data supporting substantial 
equivalence contains information, including appropriate clinical or scientific data, if deemed necessary by the FDA, that 
demonstrates that the device is as safe and as effective as a legally marketed device, and (ii) does not raise different 
questions of safety and effectiveness than the predicate device. Most 510(k)s do not require clinical data for clearance, 
but the FDA may request such data. The FDA seeks to review and act on a 510(k) within 90 days of submission, but it 
may take longer if the agency finds that it requires more information to review the 510(k). If the FDA concludes that a 
new device is not substantially equivalent to a predicate device, the new device will be classified in Class III and the 
manufacturer will be required to submit a PMA to market the product. On July 23, 2018, the CDRH of the FDA cleared 
the 510(k) for our V-TAGTM device that is compatible for use with MRIs.  

Government Regulation Outside of the United States  

In addition to regulations in the United States, we will be subject to a variety of regulations in other jurisdictions 

governing, among other things, clinical trials and any commercial sales and distribution of our products. Because 
biologically sourced raw materials are subject to unique contamination risks, their use may be restricted in some 
countries.  

Whether or not we obtain FDA approval for a product, we must obtain the requisite approvals from regulatory 

authorities in foreign countries prior to the commencement of clinical trials or marketing of the product in those 
countries. Certain countries outside of the United States have a similar process that requires the submission of a clinical 
trial application much like the IND prior to the commencement of human clinical trials, e.g., a clinical trial application 
for each clinical trial for each EU country in which the trial is conducted; a clinical trial notification is required in Japan.  

If we fail to comply with applicable foreign regulatory requirements, we may be subject to, among other things, 

fines, suspension or withdrawal of regulatory approvals, product recalls, seizure of products, operating restrictions and 
criminal prosecution.  

Coverage, Pricing and Reimbursement for Biopharmaceutical Products  

Sales of our products, when and if approved for marketing, will depend, in part, on the extent to which our 
products will be covered by third-party payors, such as federal, state, and foreign government health care programs, 
commercial insurance and managed healthcare organizations. These third-party payors are increasingly reducing 
reimbursements for medical products, drugs and services. In addition, the U.S. government, state legislatures and foreign 
governments have continued implementing cost containment programs, including price controls, restrictions on coverage 
and reimbursement and requirements for substitution of generic products. Adoption of price controls and cost 
containment measures, and adoption of more restrictive policies in jurisdictions with existing controls and measures, 
could further limit our net revenue and results. Decreases in third-party reimbursement for our product candidates or a 
decision by a third-party payor not to cover our product candidates could reduce physician usage of our products once 
approved and have a material adverse effect on our sales, results of operations and financial condition.  

Our Corporate Information 

We were incorporated under the laws of Delaware in June 2013. Our principal executive offices are located at 
75 Sidney Street, Cambridge, MA 02139. Other operations, including laboratory space, are located at 64 Sidney Street, 

59 

Cambridge, MA 02139. We lease our office and laboratory space, which consist of approximately 74,000 square feet 
located in two locations in Cambridge, Massachusetts. Our lease expires in 2026. 

Employees 

As of December 31, 2018, we employed 123 full-time employees in the United States, including 92 in research 
and development and 31 in general and administrative, and one part-time employee. Thirty-nine of our employees have 
either an MD, PhD, or PharmD. We have never had a work stoppage, and none of our employees is represented by a 
labor organization or under any collective-bargaining arrangements. We consider our employee relations to be positive.  

Legal Proceedings 

As of the date of this Annual Report on Form 10-K, we were not party to any material legal matters or claims. 
In the future, we may become party to legal matters and claims arising in the ordinary course of business, the resolution 
of which we do not anticipate would have a material adverse impact on our financial position, results of operations or 
cash flows. 

Available Information 

Our Internet address is http://www.voyagertherapeutics.com. We make available, free of charge, on or through 

our website our annual reports on Form 10-K, quarterly reports on Form 10-Q, current reports on Form 8-K, proxy 
statements and any amendments to those reports filed or furnished pursuant to Section 13(a) or 15(d) of the Securities 
and Exchange Act as soon as reasonably practicable after such material is electronically filed with or furnished to the 
Securities and Exchange Commission. The information on our website is not part of this Annual Report for the year 
ended December 31, 2018. 

ITEM 1A. 

  RISK FACTORS 

The following risk factors and other information in this Annual Report on Form 10-K, including our financial 

statements and related notes thereto, should be carefully considered. The risks and uncertainties described below are not 
the only ones we face. Additional risks and uncertainties not presently known to us or that we presently deem less 
significant may also impair our business operations. Please see page 3 of this Annual Report on Form 10-K for a 
discussion of some of the forward-looking statements that are qualified by these risk factors. If any of the following risks 
occur, our business, financial condition, results of operations and future growth prospects could be materially and 
adversely affected. 

Risks Related to Our Financial Position and Need for Capital 

We have incurred significant losses since inception and anticipate that we will continue to incur losses for the 
foreseeable future and may never achieve or maintain profitability. 

We are a clinical-stage gene therapy company with a limited operating history and have not yet generated 

revenues from the sales of our product candidates. Investment in biotechnology companies is highly speculative because 
it entails substantial upfront capital expenditures and significant risk that the product candidate will fail to obtain 
regulatory approval or become commercially viable. We have not yet demonstrated the ability to complete any clinical 
trials of our product candidates, obtain marketing approvals, manufacture a commercial-scale product or conduct sales 
and marketing activities necessary for successful commercialization. We continue to incur significant expenses related to 
research and development, and other operations in order to commercialize our product candidates. As a result, we are not 
and have never been profitable and have incurred losses since our inception. Our net losses were $88.3 million, 
$70.7 million, and $40.2 million for the years ended December 31, 2018, 2017, and 2016 respectively. As of 
December 31, 2018, we had an accumulated deficit of $269.1 million. 

60 

 
We historically have financed our operations primarily through private placements of our redeemable 

convertible preferred stock, public offerings of our common stock; and strategic collaborations, including those with 
Sanofi Genzyme, AbbVie, and Neurocrine, which we collectively refer to our Strategic Collaborations. On November 
16, 2015 we closed our initial public offering whereby we sold 5,750,000 shares of common stock at a public offering 
price of $14.00 per share, including 750,000 shares of common stock issued upon the full exercise by the underwriters of 
their option to purchase additional shares, resulting in net proceeds to us of $72.9 million after deducting underwriting 
discounts, commissions and offering expenses payable by us. On November 7, 2017, we sold 5,175,000 shares of 
common stock to the public at an offering price of $12.00 per share, including 675,000 shares of common stock issued 
upon the full exercise by the underwriters of their option to purchase additional shares, resulting in net proceeds to us of 
$58.0 million after deducting underwriting discounts, commissions, and offering expenses payable by us. On January 28, 
2019, in connection with our collaboration with Neurocrine, we agreed to sell 4,179,728 shares of common stock to 
Neurocrine at a price of $11.9625 per share, for an aggregate purchase price of approximately $50.0 million. The sale of 
shares to Neurocrine is subject to customary closing conditions, including certain antitrust approvals, that have not been 
satisfied as of the date of this Annual Report on Form 10-K. As a result, we have not yet issued such shares to 
Neurocrine, but, if the applicable closing conditions are met, we expect to do so promptly.  

To date, we have devoted substantially all of our financial resources to building our gene therapy platform, 
selecting product programs, conducting research and development, including preclinical development of our product 
candidates, building our intellectual property portfolio, building our team, and establishing Strategic Collaborations. We 
expect that it could be several years, if ever, before we have a commercialized product candidate. We expect to continue 
to incur significant expenses and increasing operating losses for the foreseeable future. The net losses we incur may 
fluctuate significantly from quarter to quarter. We anticipate that our expenses will increase substantially if, and as, we: 

 

 

 

 

 

 

continue investing in our gene therapy platform to optimize vector engineering, manufacturing and dosing 
and delivery techniques;  

continue to advance our clinical candidate, VY-AADC, as a treatment for Parkinson’s disease through the 
ongoing Phase 1b clinical trial and into our RESTORE-1 Phase 2 clinical trial;  

initiate additional preclinical studies and clinical trials for, and continue research and development of, our 
other programs;  

conduct joint research and development under our Strategic Collaborations for the research, development, 
and commercialization of certain of our pipeline programs; 

continue our process research and development activities, as well as establish our research-grade and 
commercial manufacturing capabilities;  

identify additional neurological diseases for treatment with our AAV gene therapies and develop additional 
programs or product candidates;  

  work to identify and optimize novel AAV capsids; 

 

 

develop, obtain and maintain regulatory clearances for devices to deliver our AAV gene therapies; 

seek marketing and regulatory approvals for VY-AADC or other product candidates or devices that arise 
from our programs that successfully complete clinical development; 

  maintain, expand, protect and enforce our intellectual property portfolio;  

 

identify, acquire or in-license other product candidates and technologies; 

61 

 

 

 

develop a sales, marketing and distribution infrastructure to commercialize any product candidates for 
which we may obtain marketing approval; 

expand our operational, financial and management systems and personnel, including personnel to support 
our clinical development, manufacturing and commercialization efforts and our operations as a public 
company;  

increase our product liability and clinical trial insurance coverage as we expand our clinical trials and 
commercialization efforts; and  

 

continue to operate as a public company. 

Because of the numerous risks and uncertainties associated with pharmaceutical product development, we are 

unable to accurately predict the timing or amount of increased expenses or when, or if, we will be able to achieve 
profitability. Our expenses will increase if: 

  we are required by the U.S. Food and Drug Administration, or FDA, or the European Medicines Agency, or 
EMA, or other regulatory agencies to perform trials or studies in addition to those currently expected; 

 

 

there are any delays in receipt of regulatory clearance to begin our planned clinical programs or to use 
companion devices required in such clinical programs; or  

there are any delays in enrollment of patients in or completing our clinical trials or the development of our 
product candidates. 

To become and remain profitable, we must develop and eventually commercialize, alone or with our 
collaborators, product candidates with significant market potential, which will require us to be successful in a range of 
challenging activities. These activities can include completing preclinical studies and clinical trials of our product 
candidates; obtaining marketing approval for these product candidates; developing and obtaining marketing approval of 
any required companion devices; manufacturing at clinical and commercial scale; marketing and selling those products 
that are approved; satisfying any post-marketing requirements and achieving an adequate level of market acceptance of 
and obtaining and maintaining adequate coverage and reimbursement from third-party payors for such products; and 
protecting our rights to our intellectual property portfolio. We may never succeed in any or all of these activities and, 
even if we do, we may never generate revenues that are significant or large enough to achieve profitability. If we do 
achieve profitability, we may not be able to sustain or increase profitability on a quarterly or annual basis. Our failure to 
become and remain profitable would decrease the value of our company and could impair our ability to raise capital, 
maintain our research and development efforts, expand our business or continue our operations. A decline in the value of 
our company also could cause our stockholders to lose all or part of their investment. 

We may not be able to generate sufficient revenue from the commercialization of our product candidates and may 
never be profitable. 

Our ability to generate revenue and achieve profitability depends on our ability, alone or with our collaborative 
partners, to successfully complete the development of, and obtain the regulatory approvals necessary to commercialize, 
our current and future product candidates. Our lead product candidate VY-AADC is being evaluated in a Phase 1b 
clinical trial, and we are randomizing patients in our RESTORE-1 Phase 2 clinical trial. We do not anticipate generating 
revenues from product sales for the next several years, and we may never succeed in doing so. Our ability to generate 
future revenues from product sales depends heavily on our and our collaborators’ success in: 

 

completing preclinical and clinical development of our product candidates and any required companion 
devices and identifying new product candidates; 

62 

 

 

 

seeking and obtaining regulatory and marketing approvals for product candidates for which we complete 
clinical trials; 

launching and commercializing product candidates for which we obtain regulatory and marketing approval 
by establishing a sales force, marketing and distribution infrastructure or, alternatively, collaborating with a 
commercialization partner; 

obtaining and maintaining adequate coverage and reimbursement by government and third-party payors for 
our product candidates if and when approved; 

  maintaining and enhancing a sustainable, scalable, reproducible and transferable manufacturing process for 

our vectors and product candidates; 

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establishing and maintaining supply and manufacturing relationships with third parties that can provide 
adequate products and services, in both amount and quality, to support clinical development and the market 
demand for our product candidates, if and when approved; 

obtaining an adequate level of market acceptance of our product candidates as a viable treatment option; 

addressing any competing technological and market developments; 

implementing additional internal systems and infrastructure, as needed; 

negotiating favorable terms in any collaboration, licensing or other arrangements into which we may enter 
and performing our obligations in such collaborations; 

obtaining, maintaining, protecting, enforcing and expanding our portfolio of intellectual property rights, 
including patents, trade secrets and know-how; 

avoiding and defending against third-party claims of interference or infringement; and 

attracting, hiring and retaining qualified personnel. 

Even if one or more of the product candidates that we develop is approved for commercial sale, we anticipate 
incurring significant costs associated with commercializing any approved product candidate. Our expenses could 
increase beyond expectations if we are required by the FDA, EMA, or other regulatory authorities to perform preclinical 
studies and clinical trials in addition to those that we currently anticipate. Even if we are able to generate revenues from 
the sale of any approved products, we may not become profitable and may need to obtain additional funding to continue 
operations. 

We will need to raise additional funding, which may not be available on acceptable terms, or at all. Failure to obtain 
this necessary capital when needed may force us to delay, limit or terminate certain of our product development 
efforts or other operations. 

We expect our expenses to increase in connection with our ongoing and planned activities, particularly as we 

continue the research and development of, continue or initiate clinical trials of, and seek marketing approval for, our 
product candidates. In addition, if we obtain marketing approval for any of our product candidates, we expect to incur 
significant expenses related to product sales, medical affairs, marketing, manufacturing and distribution. Since the 
completion of our IPO on November 16, 2015, we have also incurred costs associated with operating as a public 
company. Accordingly, we will need to obtain substantial additional funding in connection with our continuing 
operations. If we are unable to raise capital or enter into business development transactions when needed or on 

63 

 
acceptable terms, we could be forced to delay, reduce or eliminate certain of our research and development programs or 
any future commercialization efforts. 

Our operations have consumed significant amounts of cash since inception. As of December 31, 2018, our cash, 
cash equivalents, and marketable debt securities were $155.8 million. Based upon our current operating plan, we expect 
that our existing cash, cash equivalents, and marketable debt securities, as well as amounts expected from the upfront 
payment and expected reimbursement of development costs from the Neurocrine Collaboration Agreement and the 
upfront payment from the AbbVie Alpha-Synuclein Collaboration Agreement entered into in 2019, will enable us to 
fund our operating expenses and capital expenditure requirements into mid-2022. 

Our future capital requirements will depend on many factors, including: 

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the scope, progress, results, and costs of product discovery, preclinical studies and clinical trials for our 
product candidates and any required companion devices; 

the scope, progress, results, costs, prioritization, and number of our research and development programs; 

the progress and status of our Strategic Collaborations including any research and development costs for 
which we are responsible, the potential exercise by our collaboration partners of options to develop or 
license certain products and product candidates, and our potential receipt of future milestone payments and 
royalties from our collaboration partners; 

the extent to which we are obligated to reimburse, or entitled to reimbursement of, preclinical development 
and clinical trial costs, or the achievement of milestones or occurrence of other developments that trigger 
payments, under any other collaboration agreement to which we might become a party; 

the costs, timing and outcome of regulatory review of our product candidates; 

our ability to establish and maintain collaboration, distribution, or other marketing arrangements for our 
product candidates on favorable terms, if at all; 

the costs and timing of preparing, filing and prosecuting patent applications, maintaining and enforcing our 
intellectual property rights and defending intellectual property-related claims; 

the extent to which we acquire or in-license other product candidates and technologies or acquire or invest 
in other businesses, such as our investment in MRI Interventions, Inc., or MRIC; 

the costs related to evaluating possible alternative devices that may be useful in the delivery of our product 
candidates, including our ongoing development of V-TAG; 

the costs of securing manufacturing arrangements for pre-commercial and commercial production;  

the level of product sales from any product candidates for which we obtain marketing approval in the 
future; and 

the costs of establishing or contracting for sales, manufacturing, marketing, distribution, and other 
commercialization capabilities if we obtain regulatory approvals to market our product candidates. 

Identifying potential product candidates and conducting preclinical studies and clinical trials is a time-
consuming, expensive, and uncertain process that takes years to complete. We may never generate the necessary data or 
results required to obtain marketing approval and achieve product sales. In addition, our product candidates, if approved, 
may not achieve commercial success. Our product revenues, if any, and any commercial milestones or royalty payments 

64 

under our collaboration agreements, will be derived from sales of products that may not be commercially available for 
many years, if at all. Accordingly, we will need to continue to rely on additional financing and business development to 
achieve our business objectives. Adequate additional financing or business development transactions may not be 
available to us on acceptable terms, or at all. In addition, we may seek additional capital due to favorable market 
conditions or strategic considerations, even if we believe we have sufficient funds for our current or future operating 
plans. 

Raising additional capital may cause dilution to our stockholders, restrict our operations or require us to relinquish 
rights to our technologies or product candidates. 

Until such time, if ever, as we can generate product revenues sufficient to achieve profitability, we expect to 

finance our cash needs through a combination of equity offerings, debt financings, collaborations, strategic alliances, and 
licensing arrangements. We do not have any committed external source of funds other than the amounts we are entitled 
to receive from our collaboration partners for potential option exercises, the achievement of specified regulatory and 
commercial milestones, and royalty payments under our collaboration agreements. To the extent that we raise additional 
capital through the sale of equity or equity-linked securities, including convertible debt, our stockholders’ ownership 
interests will be diluted, and the terms of these securities may include liquidation or other preferences that adversely 
affect our existing stockholders’ rights as holders of our common stock. Debt financing and preferred equity financing, if 
available, may involve agreements that include covenants limiting or restricting our ability to take specific actions, such 
as incurring additional debt, obtaining additional capital, acquiring or divesting businesses, making capital expenditures 
or declaring dividends. Our issuance of additional securities, whether equity or debt, or the possibility of such issuance, 
may cause the market price of our common stock to decline. Further, our existing stockholders may not agree with the 
terms of such financings. 

If we raise additional funds through collaborations, strategic alliances, or licensing arrangements with third 
parties, we may have to relinquish valuable rights to our technologies, future revenue streams, research programs or 
product candidates or to grant licenses on terms that may not be favorable to us. If we are unable to raise additional funds 
through equity or debt financings when needed, we may be required to delay, limit, reduce or terminate our product 
development or future commercialization efforts or grant rights to develop and market products or product candidates 
that we would otherwise prefer to develop and market ourselves. Such collaborations, alliances, or licensing 
arrangements could therefore cause the market price of common stock to decline.  

Our limited operating history may make it difficult for our stockholders to evaluate the success of our business to date 
and to assess our future viability. 

We are an early-stage company. Our operations to date have been limited to building our team, business 

planning, raising capital, establishing our intellectual property portfolio, determining which neurological diseases to 
pursue, advancing our product candidates including delivery and manufacturing and conducting preclinical studies and 
clinical trials. Consequently, any predictions about our future success or viability may not be as accurate as they could be 
if we had a longer operating history. 

In addition, as a new business, we may encounter unforeseen expenses, difficulties, complications, delays and 

other known and unknown factors. To achieve our current goals, we will need to transition in the future from a company 
with a research and development focus to a company capable of supporting commercial activities. We may not be 
successful in such a transition. 

We expect our financial condition and operating results to continue to fluctuate significantly from quarter-to-

quarter and year-to-year due to a variety of factors, many of which are beyond our control. Accordingly, our 
stockholders should not rely upon the results of any quarterly or annual periods as indications of future operating 
performance. 

65 

Risks Related to the Development and Regulatory Approval of Our Product Candidates  

Our AAV gene therapy product candidates are based on a relatively novel technology, which makes it difficult to 
predict the time and cost of development and of subsequently obtaining regulatory approval, if at all. Only one AAV 
gene therapy product has been approved in the United States. In Europe, only two AAV gene therapy products have 
been approved. 

We have concentrated our research and development efforts to date on our gene therapy platform, identifying 
our initial targeted disease indications, and our initial product candidates. Our future success depends on our successful 
development of viable AAV gene therapy product candidates. Currently, only one of our product candidates, VY-
AADC, is in clinical development, and the remainder of our product candidates are in preclinical development. We 
cannot accurately predict when or if any of our product candidates will prove effective or safe in humans or whether 
these product candidates will receive marketing approval. There can be no assurance that we will not experience 
problems or delays in developing our product candidates and that such problems or delays will not cause unanticipated 
costs, or that any such development problems can be solved. We also may experience unanticipated problems or delays 
in expanding our manufacturing capacity. 

The clinical trial requirements of the FDA, the EMA and other regulatory authorities and the criteria these 

regulators use to determine the safety and efficacy of a product candidate vary substantially according to the type, 
complexity, novelty and intended use and market of the product candidate. The regulatory approval process for novel 
product candidates such as gene therapies can be more expensive and take longer than for other, better known or more 
extensively studied product candidates. Until August 2017, the FDA had never approved a gene therapy product. Since 
that time, it has approved Luxturna, an AAV gene therapy product by Spark Therapeutics, Inc., or Spark, for patients 
with an inherited form of vision loss. The FDA has also approved two other non-AAV gene therapy products, Kymriah 
by Novartis International AG, for pediatric and young adult patients with a form of acute lymphoblastic leukemia and 
Yescarta by Kite Pharma, Inc., for adult patients with certain forms of non-Hodgkin lymphoma. In Europe, two AAV 
gene therapy products, Glybera by uniQure N.V., or uniQure, and Luxturna by Spark Therapeutics, Inc., or Spark, have 
been granted marketing authorization; however, uniQure decided not to pursue renewal of such authorization in 2017 
and has since withdrawn Glybera from the European market. The European Commission also approved two other non-
AAV gene therapy products, Kymriah by Novartis International AG, and Yescarta by Kite Pharma, Inc. 

It is difficult to determine how long it will take or how much it will cost to obtain regulatory approvals for our 

product candidates in either the United States or the European Union or how long it will take to commercialize our 
product candidates. The few regulatory approvals to date may not be indicative of what the FDA, European Commission, 
or other regulatory authorities may require for approval or whether different or additional preclinical studies or clinical 
trials may be required to support regulatory approval in a particular jurisdiction. Delay or failure to obtain, or unexpected 
costs in obtaining, the regulatory approval necessary to bring a potential product candidate to market could decrease our 
ability to generate sufficient product revenue, and our business, financial condition, results of operations and prospects 
may be harmed. 

Regulatory requirements governing gene and cell therapy products have changed frequently and may continue to 
change in the future. Such requirements may lengthen the regulatory requirements review process, require us to 
perform additional studies, and increase our development costs or may force us to delay, limit, or terminate certain of 
our programs. 

The Center for Biologics Evaluation and Research, or CBER, of the FDA regulates biological products for 

human use. The Office of Tissues and Advanced Therapies, or OTAT, formerly known as the Office of Cellular, Tissue 
and Gene Therapies, within CBER reviews gene therapy and related products and has established the Cellular, Tissue 
and Gene Therapies Advisory Committee to advise CBER in its review.  

NIH-funded institutions need to have their institutional biosafety committee, or IBC, as well as their 
institutional review board, or IRB, review proposed clinical trials to assess the safety of the trial. The Phase 1b clinical 
trial of VY-AADC and the separate Phase 1 trial exploring the delivery of VY-AADC using a posterior trajectory are 
being conducted at the University of California San Francisco, or UCSF, and University of Pittsburgh Medical Center, or 

66 

UPMC, and two other sites and therefore are subject to oversight by these authorities. Such trials will need to be re-
reviewed by the respective institutional IRBs if the protocol for the trial is further amended. For any new clinical trial 
protocols, including those of our RESTORE-1 Phase 2 clinical trial, the same processes and issues apply.  

Adverse developments in clinical trials of gene therapy products conducted by us or others may cause the FDA 
or other oversight bodies to change the requirements for approval of any of our product candidates. Similarly, EMA and 
local health authorities of individual countries within the European Union may issue new guidelines concerning the 
clinical development and marketing authorization for gene therapy medicinal products and require that we comply with 
these new guidelines. The EMA and agencies at both the federal and state level in the United States have expressed an 
interest in further regulating new biotechnologies, including gene therapy. In addition, gene therapy products are 
considered genetically-modified organism, or GMO, products and are regulated as such in each country. Designation of 
the type of GMO product and subsequent handling and disposal requirements can vary across countries and is variable 
throughout the EU. Addressing each specific country requirement and obtaining approval to commence a clinical trial in 
these countries could result in delays in starting, conducting, or completing a clinical trial. Similar issues could be faced 
in other regions of the world including the Asia-Pacific region. 

These regulatory review committees and advisory groups and the new guidelines they promulgate may lengthen 

the regulatory review process, require us to perform additional studies, increase our development costs, lead to changes 
in regulatory positions and interpretations, delay or prevent approval and commercialization of these product candidates 
or lead to significant post-approval limitations or restrictions. As we advance our product candidates, we will be required 
to consult with these regulatory and advisory groups, and comply with applicable guidelines. We have requested 
feedback from the FDA on among other matters, the regulatory pathway for VY-AADC and design of our proposed 
pivotal program, including a randomized, placebo-controlled RESTORE-1 Phase 2 clinical trial and, if required, a 
randomized, placebo-controlled RESTORE-2 Phase 3 clinical trial. We had multiple interactions with the FDA 
throughout 2018 and received certain written feedback requiring additional clarification.  In December 2018, we held a 
Type B meeting with the FDA to discuss the overall development and pivotal program for VY-AADC. Based on the 
meeting discussion and subsequent written feedback from the FDA, we plan to submit a revised trial protocol that will 
include an increase in the target number of patients in the RESTORE-1 Phase 2 trial, resulting in 75 to 100 total patients 
in the trial, and to conduct, in staggered-parallel, the RESTORE-2 Phase 3 trial of similar size and design to RESTORE-
1. These updates incorporate guidance from the FDA from the Type B meeting to conduct two adequate and well-
controlled clinical trials for a large patient population such as Parkinson’s disease. We plan to continue to seek and 
incorporate FDA guidance in our ongoing development plans. If we fail to consult or solicit guidance from regulators, 
we may be required to delay or discontinue development of certain of our product candidates. These additional processes 
may result in a review and approval process that is longer than we otherwise would have expected. Delays as a result of 
increased or lengthier regulatory approval process and further restrictions on development of our product candidates can 
be costly and could negatively impact our or our collaborators’ ability to complete clinical trials and commercialize our 
current and future product candidates in a timely manner, if at all.  

Results from preclinical studies and early-stage clinical trials may not be indicative of efficacy in late-stage clinical 
trials. 

All of our product candidates are in early stages of development. Clinical testing is expensive, is difficult to 

design and implement, can take many years to complete and is uncertain as to outcome. A failure of one or more clinical 
trials can occur at any stage of testing. The outcome of preclinical testing and early clinical trials may not be predictive 
of the success of later stage clinical trials, interim results of a clinical trial do not necessarily predict final results and 
results from one completed clinical trial may not be replicated in a subsequent clinical trial with a similar study design. 
Some of our clinical trials, including our Phase 1 clinical trials and our Phase 1b clinical trial for VY-AADC, were 
conducted with small patient populations and were not blinded or placebo-controlled, making it difficult to predict 
whether the favorable results that we observed in such trials will be repeated in larger and more advanced clinical trials. 
Moreover, preclinical and clinical data are often susceptible to varying interpretations and analyses, and many companies 
that have believed their product candidates performed satisfactorily in preclinical studies and clinical trials have 
nonetheless failed to obtain marketing approval of their products. 

67 

 
A number of companies in the pharmaceutical and biotechnology industries have suffered significant setbacks 

in late-stage clinical trials even after achieving promising results in early-stage clinical trials. If a larger population of 
patients does not experience positive results, if these results are not reproducible, or if our products show diminishing 
activity over time, our products may not receive approval from the EMA or the FDA. Data obtained from preclinical and 
clinical activities are subject to varying interpretations, which may delay, limit or prevent regulatory approval. In 
addition, we may encounter regulatory delays or rejections as a result of many factors, including changes in regulatory 
policy during the period of product development. Failure to confirm favorable results from earlier trials by demonstrating 
the safety and effectiveness of our products in late-stage clinical trials with larger patient populations could harm our 
business and we may never succeed in commercialization or generating product revenue. 

The doses and coverage of the putamen being employed in the ongoing VY-AADC Phase 1b clinical trial and the 
RESTORE-1 Phase 2 clinical trial are higher than those used in prior trials and may need to be further optimized or 
we may not generate sufficient clinical data in a placebo-controlled trial to achieve market authorization. Further, 
any favorable results which we obtain from our Phase 1b clinical trial might not be replicated in subsequent trials. 

The clinical trial results of some of our collaborators have been negatively affected by factors that had not been 

fully anticipated prior to the examination of the trial results. For example, the magnitude of some of the clinical 
responses seen in the Phase 1 clinical trial of AAV2-AADC conducted by UCSF were similar to placebo effects 
observed in previous surgical therapies for Parkinson’s disease. As a result, we are unable to rely on the results of this 
Phase 1 trial for an indication of the efficacy of treatment with VY-AADC. We believe that there is a need to optimize 
the dose and volume of infusion of VY-AADC to substantially increase the coverage of the putamen, the region of the 
brain targeted by VY-AADC, to achieve a clinical benefit. However, we can provide no assurances that we will be able 
to optimize these parameters and thereby achieve sufficient coverage of the putamen to achieve a clinical benefit. 

The ongoing Phase 1b clinical trial of VY-AADC incorporated several design features in an attempt to increase 

the area of the putamen, particularly the posterior putamen, which receives VY-AADC treatment. We employed larger 
infusion volumes and higher doses of VY-AADC, and we used the ClearPoint® System to provide real-time, intra-
operative, magnetic resonance imaging, or MRI, assistance to the physician surgically administering VY-AADC to the 
patient. 

In a separate Phase 1 clinical trial, we are also exploring posterior, or back of the head, delivery of drug into the 

putamen, compared to a transfrontal, or top of the head, delivery approach used in Cohorts 1 through 3 of the ongoing 
Phase 1b clinical trial described above. A posterior approach better aligns the infusion of VY-AADC with the anatomical 
structure of the putamen to potentially reduce the total procedure time and increase the total coverage of the putamen. 
Administration of VY-AADC with this posterior approach has been well-tolerated with no reported serious adverse 
events, or SAEs, with most patients discharged from the hospital the day after surgery. 

Due to the nature of the techniques used in the Phase 1b clinical trial and the numerous variables that can be 

changed, it is possible that the data generated from this trial may not provide evidence of clinical benefit. For example, 
physicians may use cannulas, which are small tubes, of differing lengths in the infusion procedure, or may use differing 
infusion speeds or infusion angles. These differences could affect the dose of VY-AADC that ultimately reaches the 
putamen, leading to highly variable results. Similarly, we have limited experience to date with the posterior delivery 
approach which is the preferred surgical route of administration for the RESTORE-1 Phase 2 clinical trial, and it may not 
generate outcomes that are clinically superior to those of the transfrontal approach.  

For the RESTORE-1 Phase 2 clinical trial, we have selected a dose of up to 2.5 x 1012 vector genomes, which is 

defined as a maximum total bilateral dose. This dose is between the up to maximum total vector genome doses 
administered in Cohorts 2 and 3 from the Phase 1b trial when considering the higher volume administered with the 
posterior trajectory and vector produced using the baculovirus system. We have not previously evaluated this dose in a 
clinical trial. The dose concentration and volume we have selected for the RESTORE-1 Phase 2 clinical trial may not 
achieve the desired safety and efficacy in the RESTORE-1 Phase 2 randomized, controlled clinical trial. 

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We expect that the RESTORE-1 Phase 2 trial will enroll 75-100 patients. Movement disorder specialists will 

identify and screen potential patients prior to referring to a surgical site for VY-AADC administration and provide 
clinical and safety follow-up after VY-AADC administration. 

The RESTORE-1 Phase 2 trial is a randomized, double-blind, placebo-surgery controlled trial with a planned 

enrollment of 75 to 100 patients who have been diagnosed with Parkinson’s disease for at least four years, are not 
responding adequately to oral medications, and have at least three hours of OFF time during the day as measured by a 
validated self-reported patient diary. Patients will be randomized 1:1 to either VY-AADC or placebo surgery.  

The primary efficacy endpoint of the RESTORE-1 Phase 2 trial is the mean improvement from baseline to 12 

months on time without troublesome dyskinesia, or good ON time, as measured by a validated self-reported patient diary 
at 12 months. Secondary endpoints include diary OFF time, other motor function and quality of life measures from the 
United Parkinson’s Disease Rating Scales (UPDRS-II,-III scores), the Parkinson’s Disease Questionnaire (PDQ-39), and 
patient’s global function as measured by the proportion of participants with improvement on the Clinical Global 
Impression (CGI) score. The trial will also measure non-motor symptoms from the Non-Motor Symptom Scale (NMSS), 
as well as safety.  

Biomarker data include measurements of the coverage of the putamen, the specific region of the brain targeted 

with VY-AADC, and measurements of AADC enzyme expression and activity in the putamen measured by positron 
emission tomography (PET) using fluorodopa F-18. Changes in patients’ daily doses of oral levodopa and related 
medications will also be recorded. 

We are using a different manufacturing process for our AAV gene therapy vector in our global RESTORE-1 
Phase 2 clinical trial and our planned RESTORE-2 Phase 3 clinical trial. We have begun to manufacture VY-AADC 
using our baculovirus/Sf9 system as opposed to manufacturing in HEK 293 cells, which were used in our Phase 1 
clinical trials. We have conducted studies to demonstrate comparability between the current version and the new version. 
It is possible, however, that the results of our RESTORE-1 Phase 2 clinical trial and our planned RESTORE-2 Phase 3 
clinical trial in Parkinson’s disease may differ from the results of our Phase 1b or our Phase 1 posterior clinical trials 
based on VY-AADC manufactured using our baculovirus/Sf9 system. 

We intend to conduct, and may in the future conduct, clinical trials for product candidates at sites outside the United 
States, and the FDA may not accept data from trials conducted in such locations. 

To date, we have only conducted clinical trials in the United States. However, we may in the future choose to 

conduct, one or more of our clinical trials or include sites in current or future clinical trials outside the United States. We 
generally plan to conduct our later stage and pivotal clinical trials of our product candidates globally. We currently plan 
to include sites located in Poland in our RESTORE-1 Phase 2 clinical trial.  

Although the FDA may accept data from sites or clinical trials outside the United States, acceptance of these 
data is subject to conditions imposed by the FDA. For example, the clinical trial must be well designed and conducted 
and performed by qualified investigators in accordance with ethical principles. The trial population must also adequately 
represent the U.S. population, and the data must be applicable to the U.S. population and U.S. medical practice in ways 
that the FDA deems clinically meaningful. In addition, while these clinical trials or trial sites are subject to the applicable 
local laws, FDA acceptance of the data will depend on its determination that the trials or trial sites also complied with all 
applicable U.S. laws and regulations. If the FDA does not accept the data from any trial or trial site outside the United 
States, it would likely result in the need for additional trials, which would be costly and time-consuming and would delay 
or permanently halt our development of the applicable product candidates. 

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Other risks inherent in conducting international clinical trials or using international trial sites include: 

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foreign regulatory requirements that could restrict or limit our ability to conduct our clinical trials; 

administrative burden of complying with a variety of foreign laws, medical standards and regulatory 
requirements, including the regulation of pharmaceutical and biotechnology products and treatment;  

failure of enrolled patients to adhere to clinical protocols or inadequate collection and assessment of 
clinical data as a result of differences in healthcare services or cultural customs;  

foreign exchange fluctuations;  

diminished or loss of protection of intellectual property in the relevant jurisdiction; and  

political and economic risks relevant to foreign countries. 

We may encounter substantial delays or difficulties in commencement, enrollment or completion of our clinical trials 
or we may fail to demonstrate safety and efficacy to the satisfaction of applicable regulatory authorities, which could 
prevent us from commercializing our current and future product candidates on a timely basis, if at all. 

Before obtaining marketing approval from regulatory authorities for the sale of our current and future product 

candidates, we must conduct extensive clinical trials to demonstrate the safety and efficacy of the product candidates. 
Clinical trials are expensive, time-consuming and their outcomes are uncertain. 

We have very limited experience with clinical trials. The RESTORE-1 Phase 2 clinical trial of VY-AADC is 
being conducted at several locations including UCSF and UPMC. We cannot guarantee that any clinical trials will be 
conducted as planned or completed on schedule, if at all. A clinical trial failure can occur at any stage of testing. 
Similarly, there may be delays or difficulties in our initiation of future clinical trials. Due to the additional regulatory 
uncertainties associated with gene therapy products, we did not initiate our RESTORE-1 Phase 2 clinical trial for our 
clinical candidate, VY-AADC, as a treatment for Parkinson’s disease until we met with OTAT to discuss our proposed 
trial design and overall development plan. While we have received OTAT’s feedback, and incorporated it as appropriate 
in our plans, the clinical trial as designed may not achieve the prospectively defined primary clinical endpoints or 
provide a favorable benefit to risk ratio to support a BLA filing or approval.  

Identifying and qualifying patients to participate in clinical trials of our product candidates is critical to our 
success. We may not be able to identify, recruit and enroll a sufficient number of patients, or those with required or 
desired characteristics, to complete our clinical trials in a timely manner or at all pursuant to the requirements of the 
FDA, EMA, or other regulatory authorities. Patient enrollment and trial completion are affected by many factors 
including: 

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perceived risks and benefits of AAV gene therapy approaches for the treatment of neurological diseases; 

perceived risks of the delivery procedure, such as intracranial infusion for VY-AADC; 

formulation changes to our product candidates may require us to conduct additional clinical studies to 
bridge our modified product candidates to earlier versions; 

size of the patient population and process for identifying patients; 

design of the trial protocol; 

eligibility and exclusion criteria; 

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patients with preexisting antibodies to the gene therapy vector that preclude their participation in the trial; 

perceived risks and benefits of the product candidate under study; 

availability of competing therapies and clinical trials; 

severity of the disease under investigation; 

availability of genetic testing for potential patients; 

proximity and availability of clinical trial sites for prospective patients; 

lack of adequate compensation of patients; 

ability to obtain and maintain patient consent; 

risk that enrolled patients will drop out before completion of the trial; 

our ability to locate appropriately trained physicians to conduct such clinical trials, which may be 
particularly difficult for the VY-AADC RESTORE-1 Phase 2 and RESTORE-2 Phase 3 clinical trials. We 
have historically used, and expect to use, the ClearPoint System, which is only available at a small number 
of academic medical centers in the United States; 

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our ability to commercially launch V-TAG, our real-time, intra-operative, MRI-compatible device, and to 
train physicians to conduct clinical trials using the device; 

  willingness of patients to participate in a placebo-controlled trial;  

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patient referral practices of physicians; and 

ability to monitor patients adequately during and after treatment. 

Further, we plan to seek marketing approvals in the United States, the European Union and other jurisdictions, 

which may require that we conduct clinical trials in foreign countries. Our ability to successfully initiate, enroll and 
complete a clinical trial in any foreign country is subject to numerous risks unique to conducting business in foreign 
countries, including: 

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difficulty in establishing or managing relationships with clinical research organizations, or CROs, and 
physicians; 

different standards for the conduct of clinical trials; 

absence in some countries of established groups with sufficient regulatory expertise for review of AAV 
gene therapy protocols; 

our inability to locate qualified local partners or collaborators for such clinical trials; and 

the potential burden of complying with a variety of foreign laws, medical standards and regulatory 
requirements, including the regulation of pharmaceutical and biotechnology products and treatment. 

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If we have difficulty enrolling a sufficient number of patients to conduct our clinical trials as planned, we may 

need to delay, limit or terminate ongoing or planned clinical trials, any of which would harm our business, financial 
condition, results of operations and prospects. 

Other events that may prevent successful or timely completion of clinical development include: 

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delays in reaching a consensus with regulatory authorities on trial design; 

delays in reaching agreement on acceptable terms with prospective CROs, and clinical trial sites; 

delays in opening clinical trial sites or obtaining required IRB or independent ethics committee approval at 
each clinical trial site; 

imposition of a clinical hold by regulatory authorities as a result of a serious adverse event or after an 
inspection of our clinical trial operations or trial sites; or our decision or the requirement of regulators or 
institutional review boards to suspend or terminate clinical research for various reasons, including 
noncompliance with regulatory requirements or a finding that the participants are being exposed to 
unacceptable health risks; 

failure by us, any CROs we engage, or any other third parties to adhere to clinical trial protocols or 
regulatory requirements; 

failure by us, any CROs we engage, or any other third parties to perform in accordance with the FDA’s 
good clinical practices, or GCPs, or applicable regulatory guidelines in the European Union; 

failure by physicians to adhere to delivery protocols leading to variable results; 

delays in the testing, validation, manufacturing and delivery of our product candidates to the clinical sites, 
including delays by third parties with whom we have contracted to perform certain of those functions; 

insufficient or inadequate supply or quality of our product candidates or other materials necessary to 
conduct clinical trials of our product candidates, including potential delays in our RESTORE-1 Phase 2 
clinical trial in Parkinson’s disease associated with the commercial availability of V-TAG; 

delays in having patients complete participation in a trial or return for post-treatment follow-up; 

clinical trial sites or patients dropping out of a trial at a rate higher than we anticipate; 

selection of clinical endpoints that require prolonged periods of clinical observation or analysis of the 
resulting data; 

receipt of negative or inconclusive clinical trial results; 

occurrence of serious adverse events associated with the product candidate that are viewed to outweigh its 
potential benefits; 

occurrence of serious adverse events in trials of the same class of agents conducted by other sponsors;  

changes in regulatory requirements and guidance that require amending or submitting new clinical 
protocols; or 

72 

 

the cost of clinical trials of our product candidates may be greater than we anticipate. 

Any inability to successfully complete preclinical studies and clinical trials could result in additional costs to us 
or impair our ability to generate revenues from product sales, regulatory and commercialization milestones and royalties. 
In addition, if we make manufacturing or formulation changes to our product candidates, we may need to conduct 
additional studies to bridge our modified product candidates to earlier versions. Clinical trial delays also could shorten 
any periods during which we may have the exclusive right to commercialize our product candidates or allow our 
competitors to bring products to market before we do, which could impair our ability to successfully commercialize our 
product candidates and may harm our business, financial condition, results of operations and prospects. 

Additionally, if the results of our clinical trials are inconclusive or if there are safety concerns or serious adverse 

events associated with our product candidates, we may: 

 

 

 

 

 

 

 

 

 

be delayed in obtaining marketing approval for our product candidates, if at all; 

obtain approval for indications or patient populations that are not as broad as intended or desired; 

obtain approval with labeling that includes significant use or distribution restrictions or safety warnings; 

be subject to changes in the way the product is administered; 

be required to perform additional clinical trials to support approval or be subject to additional 
post-marketing testing requirements; 

have regulatory authorities withdraw, or suspend, their approval of the product or impose restrictions on its 
distribution in the form of a Risk Evaluation and Mitigation Strategy, or REMS; 

be subject to the addition of labeling statements, such as warnings or contraindications; 

be sued; or 

experience damage to our reputation. 

Our product candidates or the process for administering our product candidates may cause undesirable side effects or 
have other properties that could delay or prevent their regulatory approval, limit their commercial potential or result 
in significant negative consequences following any potential marketing approval. 

In past clinical trials that were conducted by others with non-AAV gene therapy vectors, several significant side 

effects were caused by gene therapy treatments, including reported cases of leukemia and death. Other potential side 
effects could include an immunologic reaction and insertional oncogenesis, which is the process whereby the insertion of 
a functional gene near a gene that is important in cell growth or division results in uncontrolled cell division, which 
could potentially enhance the risk of malignant transformation. If our vectors demonstrate a similar adverse effect, or 
other adverse effects, we may be required to halt or delay further clinical development of our product candidates or 
withdraw the product from the market post-approval. For example, in a recently published review of patients with 
hepatocellular carcinomas, it was shown that a small subset contained an integrated genome sequence of wild-type 
AAV2 and it was suggested that AAV2 may be associated with insertional oncogenesis.  

In addition to side effects caused by the product candidate, the administration process or related procedures also 

can cause side effects. VY-AADC and VY-HTT01 will be administered directly to the targeted areas and cells in the 
brain, requiring the patient to undergo brain surgery. In a previous Phase 1 clinical trial conducted by UCSF, three 
patients experienced hemorrhages caused by the surgical procedure for administering VY-AADC. In the RESTORE-1 
Phase 2 clinical trial of VY-AADC, we are using the ClearPoint System, which has only been used in limited gene 

73 

therapy neurosurgeries to date to provide accurate placement of the cannula in the putamen, to allow for real-time, 
intra-operative MRI to assist the physician in visualizing the delivery of VY-AADC to the putamen and to avoid specific 
blood vessels during the duration of the surgical procedure, with the goal of reducing the risk of hemorrhages. One 
patient in the ongoing Phase 1b trial at UCSF experienced two SAEs, a pulmonary embolism, or blood clot in the lungs, 
and related heart arrhythmia, or irregular heartbeat, which were determined to be related to the surgical procedure and 
prolonged immobility, not VY-AADC. In our Phase 2 and future trials, we may use V-TAG, a proprietary real-time, 
intra-operative, MRI-compatible device that we are currently developing. For VY-SOD102 in the treatment for ALS, the 
product candidate is planned to be injected directly into the spinal cord. Limited clinical data are available for this route 
of administration. If other side effects were to occur in connection with the surgical procedure, or problems were 
encountered with the use of V-TAG, our clinical trials could be suspended or terminated. 

If in the future we are unable to demonstrate that such side effects were caused by the administration process or 

related procedures or are unable to modify the trial protocol adequately to address such side effects, the FDA, the 
European Commission, the EMA or other regulatory authorities could order us to cease further development of, or deny 
approval of, our product candidates for any or all targeted indications. For products that “knock down” or reduce the 
amount of a gene or its encoded protein, their effects on other parts of the body, or “off target” effects, could result in 
unforeseen toxicity. Even if we are able to demonstrate that any future SAEs are not product-related, and regulatory 
authorities do not order us to cease further development of our product candidates, such occurrences could affect patient 
recruitment or the ability of enrolled patients to complete the trial. Moreover, if we elect, or are required, to delay, 
suspend or terminate any clinical trial of any of our product candidates, the commercial prospects of such product 
candidates may be harmed and our ability to generate product revenues from any of these product candidates may be 
delayed or eliminated. Any of these occurrences may harm our ability to develop other product candidates, and may 
harm our business, financial condition and prospects significantly. 

Additionally, if any of our product candidates receives marketing approval, the FDA could require us to adopt a 

REMS to ensure that the benefits outweigh its risks. Such REMS may include, among other things, a medication guide 
outlining the risks of the product for distribution to patients and a communication plan to health care practitioners or the 
limitation of the use of the product to specifically trained neurosurgeons and/or certain centers. Furthermore, adverse 
events which were initially considered unrelated to the study treatment of the clinical trial may later be found to be 
caused by the study treatment. If we or others later identify undesirable side effects caused by our product candidate, 
several potentially significant negative consequences could result, including: 

 

 

regulatory authorities may suspend or withdraw approvals of such product candidate; 

regulatory authorities may require additional warnings on the label; 

  we may be required to change the way a product candidate is administered or conduct additional clinical 

trials; 

  we could be sued and held liable for harm caused to patients; and 

 

our reputation may suffer. 

Any of these events could prevent us from achieving or maintaining market acceptance of our product 

candidates and could significantly harm our business, prospects, financial condition and results of operations. 

74 

We may be unable to obtain orphan drug designation or exclusivity for any of our product candidates for which we 
seek such designation. If our competitors are able to obtain orphan drug exclusivity for products that constitute the 
same drug and treat the same indications as our product candidates, we may not be able to have competing products 
approved by the applicable regulatory authority for a significant period of time. For products for which we may 
obtain orphan drug designation or exclusivity, we may be unable to prevent the approval or marketing authorization 
of other similar products based upon regulator decisions regarding product “sameness”. 

Regulatory authorities in some jurisdictions, including the United States and the European Union, may 

designate drugs for relatively small patient populations as orphan drugs. Under the Orphan Drug Act of 1983, or the 
Orphan Drug Act, the FDA may designate a product candidate as an orphan drug or biological product if it is intended to 
treat a rare disease or condition, which is generally defined as having a patient population of fewer than 200,000 
individuals in the United States, or a patient population greater than 200,000 in the United States where there is no 
reasonable expectation that the cost of developing the drug or biological product will be recovered from sales in the 
United States. We have received feedback from the FDA that VY-AADC for the treatment of Parkinson’s disease does 
not qualify for orphan disease designation because the potential for its use in earlier stages of Parkinson’s disease 
exceeds the 200,000 patient population criteria in the United States. In the European Union, EMA’s Committee for 
Orphan Medicinal Products grants orphan drug designation to promote the development of products that are intended for 
the diagnosis, prevention or treatment of a life-threatening or chronically debilitating condition affecting not more than 
five in 10,000 persons in the European Union. Additionally, orphan designation is granted for products intended for the 
diagnosis, prevention or treatment of a life-threatening, seriously debilitating or serious and chronic condition and when, 
without incentives, it is unlikely that sales of the drug in the European Union would be sufficient to justify the necessary 
investment in developing the drug or biologic product. We have received feedback from the Committee for Orphan 
Medicinal Products that orphan designation likely would not be granted for VY-AADC in Parkinson’s disease since the 
Committee does not grant such status for products targeting more severe stages of a disease. 

Generally, if a product candidate with an orphan drug designation receives the first marketing approval for the 
indication for which it has such designation, the product is entitled to a period of marketing exclusivity, which precludes 
the FDA or the European Commission from approving another marketing application for a product that constitutes the 
same drug treating the same indication for that marketing exclusivity period, except in limited circumstances. If another 
sponsor receives such approval before we do (regardless of our orphan drug designation), we may be precluded from 
receiving marketing approval for our product for the applicable exclusivity period. The applicable period is seven years 
in the United States and 10 years in the European Union. The exclusivity period in the United States can be extended by 
nine months if the BLA sponsor submits pediatric data that adequately respond to a written request from the FDA for 
such data. The exclusivity period in the European Union can be reduced to six years if a product no longer meets the 
criteria for orphan drug designation or if the product is sufficiently profitable so that market exclusivity is no longer 
justified. Orphan drug exclusivity may be revoked if any regulatory agency determines that the request for designation 
was materially defective or if the manufacturer is unable to assure sufficient quantity of the product to meet the needs of 
patients with the rare disease or condition. 

We believe that all of our current programs may qualify for orphan drug designation except for VY-AADC for 

Parkinson’s disease. Even if we obtain orphan drug exclusivity for a product candidate, that exclusivity may not 
effectively protect the product candidate from competition because different drugs or biological products can be 
approved for the same condition. In the United States, even after an orphan drug is approved, the FDA may subsequently 
approve another drug or biological product for the same condition if the FDA concludes that the latter drug or biological 
product is not the same drug or biological product or is clinically superior in that it is shown to be safer, more effective 
or makes a major contribution to patient care. In the European Union, marketing authorization may be granted to a 
similar medicinal product for the same orphan indication if: 

 

 

the second applicant can establish in its application that its medicinal product, although similar to the 
orphan medicinal product already authorized, is safer, more effective or otherwise clinically superior; 

the holder of the marketing authorization for the original orphan medicinal product consents to a second 
orphan medicinal product application; or 

75 

 

the holder of the marketing authorization for the original orphan medicinal product cannot supply sufficient 
quantities of orphan medicinal product. 

Even if we seek orphan drug designation from the FDA, the European Commission or other regulatory agencies 

for a product candidate, there can be no assurances that the regulatory agency or agencies will grant such designation. 
Additionally, the designation of any of our product candidates as an orphan drug does not guarantee that any regulatory 
agency will ultimately approve that product candidate or prevent other products from receiving marketing authorization 
due to decisions of the applicable regulatory agency regarding “sameness” of the products.  

On August 3, 2017, the Congress passed the FDA Reauthorization Act of 2017, or FDARA. FDARA, among 
other things, codified the FDA’s pre-existing regulatory interpretation, to require that a drug sponsor demonstrate the 
clinical superiority of an orphan drug that is otherwise the same as a previously approved drug for the same rare disease 
in order to receive orphan drug exclusivity. The new legislation reverses prior precedent holding that the Orphan Drug 
Act unambiguously requires that the FDA recognize the orphan exclusivity period regardless of a showing of clinical 
superiority. The FDA may further reevaluate the Orphan Drug Act and its regulations and policies. We do not know if, 
when, or how the FDA may change the orphan drug regulations and policies in the future, and it is uncertain how any 
changes might affect our business. Depending on what changes the FDA may make to its orphan drug regulations and 
policies, our business could be adversely impacted. 

A potential breakthrough therapy designation by the FDA for our product candidates may not lead to a faster 
development or regulatory review or approval process, and it does not increase the likelihood that our product 
candidates will receive marketing approval. 

We have sought and may in the future seek a breakthrough therapy designation for some of our product 

candidates. A breakthrough therapy is defined as a drug or biological product that is intended, alone or in combination 
with one or more other drugs, to treat a serious or life-threatening disease or condition, and preliminary clinical evidence 
indicates that the drug or biological product may demonstrate substantial improvement over existing therapies on one or 
more clinically significant endpoints, such as substantial treatment effects observed early in clinical development. For 
drugs or biological products that have been designated as breakthrough therapies, interaction and communication 
between the FDA and the sponsor of the trial can help to identify the most efficient path for clinical development while 
minimizing the number of patients placed in ineffective control regimens. Drugs designated as breakthrough therapies by 
the FDA may also be eligible for accelerated approval. 

Designation as a breakthrough therapy is within the discretion of the FDA. Accordingly, even if we believe one 
of our product candidates meets the criteria for designation as a breakthrough therapy, the FDA may disagree and instead 
determine not to make such designation. In any event, the receipt of a breakthrough therapy designation for a product 
candidate may not result in a faster development process, review or approval compared to drugs considered for approval 
under conventional FDA procedures and does not assure ultimate approval by the FDA. In addition, even if one or more 
of our product candidates qualify as breakthrough therapies, the FDA may later decide that the drugs or biological 
products no longer meet the conditions for qualification. 

A potential regenerative medicine advanced therapy designation by the FDA for our product candidates may not lead 
to a faster development or regulatory review or approval process, and it does not increase the likelihood that our 
product candidates will receive marketing approval. 

We have sought and may in the future seek a regenerative medicine advanced therapy, or RMAT, designation 

for some of our product candidates. Under the 21st Century Cures Act, or the Cures Act, to be eligible to receive RMAT 
designation from the FDA, a product candidate must be (i) considered a “regenerative medicine therapy” as defined in 
the Cures Act; (ii) intended to treat, modify, reverse, or cure one or more serious or life-threatening diseases or 
conditions; and (iii) indicated, in preliminary clinical evidence, to have the potential to address unmet medical needs for 
such diseases or conditions. Gene therapies, including genetically modified cells, that lead to a durable modification of 
cells or tissues may meet the definition in the Cures Act of a regenerative medicine therapy.  

76 

 
The RMAT program is intended to facilitate efficient development and expedite review of such therapies. A 

new drug application or a BLA for a product candidate that has received an RMAT designation may be eligible for 
priority review or accelerated approval through (1) surrogate or intermediate endpoints reasonably likely to predict long-
term clinical benefit or (2) reliance upon data obtained from a meaningful number of sites. Benefits of such designation 
also include early interactions with FDA to discuss any potential surrogate or intermediate endpoint to be used to support 
accelerated approval. A product candidate that has received an RMAT designation that is granted accelerated approval 
and is subject to post-approval requirements may fulfill such requirements through the submission of clinical evidence, 
clinical studies, patient registries, or other sources of real world evidence, such as electronic health records; the 
collection of larger confirmatory data sets; or post-approval monitoring of all patients treated with such therapy prior to 
its approval.  

In June 2018, the FDA granted RMAT designation for our VY-AADC gene therapy treatment for Parkinson’s 

disease in patients with motor fluctuations that are refractory to medical management. The designation was based on data 
from our Phase 1b clinical trial. 

RMAT designation is within the discretion of the FDA. Accordingly, even if we believe one of our other 

product candidates meets the criteria for RMAT designation, the FDA may disagree and instead determine not to make 
such designation. In any event, the receipt of RMAT designation for a product candidate may not result in a faster 
development process, review or approval compared to drugs considered for approval under conventional FDA 
procedures and does not assure ultimate approval by the FDA. In addition, the FDA may later decide that product 
candidate that received RMAT designation no longer meets the conditions for designation. 

Fast track designation by the FDA may not actually lead to a faster development or regulatory review or approval 
process and does not assure FDA approval of our product candidate. 

If a drug is intended for the treatment of a serious or life-threatening condition and the drug demonstrates the 
potential to address unmet medical need for this condition, the drug sponsor may apply for FDA fast track designation. 
VY-AADC has been granted fast track designation by the FDA. We may seek such a designation for our other product 
candidates. A fast track designation does not ensure that the product candidate will receive marketing approval or that 
approval will be granted within any particular timeframe. Thus, fast track products may not experience a faster 
development process, review or approval compared to conventional FDA procedures. In addition, the FDA may 
withdraw fast track designation if it believes that the designation is no longer supported by data from our clinical 
development program. Fast track designation alone does not guarantee qualification for the FDA’s priority review 
procedures. 

Priority review designation by the FDA may not lead to a faster regulatory review or approval process and, in 

any event, does not assure FDA approval of our product candidate. 

If the FDA determines that a product candidate offers major advances in treatment or provides a treatment 

where no adequate therapy exists, the FDA may designate the product candidate for priority review. A priority review 
designation means that the FDA’s goal to review an application is six months, rather than the standard review period of 
ten months. We may request priority review for our product candidates. The FDA has broad discretion with respect to 
whether or not to grant priority review status to a product candidate, so even if we believe a particular product candidate 
is eligible for such designation or status, the FDA may decide not to grant it. Moreover, a priority review designation 
does not necessarily mean a faster regulatory review process or necessarily confer any advantage with respect to 
approval compared to conventional FDA procedures. Receiving priority review from the FDA does not guarantee 
approval within the six-month review cycle or thereafter. 

77 

 
 
Even if we complete the necessary preclinical studies and clinical trials, the marketing approval process is expensive, 
time-consuming and uncertain and may prevent us from obtaining approvals for the commercialization of some or all 
of our product candidates. If we or any current or future collaborators are not able to obtain, or if there are delays in 
obtaining, required regulatory approvals, we or they may not be able to commercialize our products, and our ability to 
generate revenue may be materially impaired.  

Our product candidates and the activities associated with their development and commercialization, including 
their design, testing, manufacture, safety, efficacy, recordkeeping, labeling, storage, approval, advertising, promotion, 
sale and distribution, export and import, are subject to comprehensive regulation by the FDA and other regulatory 
agencies in the United States and by the EMA and comparable regulatory authorities in other countries. Failure to obtain 
marketing approval for a product candidate will prevent us from commercializing the product candidate. We have not 
received approval to market any of our product candidates from regulatory authorities in any jurisdiction. We have only 
limited experience in filing and supporting the applications necessary to gain marketing approvals and expect to rely on 
third-party contract research organizations to assist us in this process.  

Securing marketing approval requires the submission of extensive preclinical and clinical data and supporting 

information to the various regulatory authorities for each therapeutic indication to establish the product candidate’s 
safety and efficacy. Securing regulatory approval also requires the submission of information about the product 
manufacturing process to, and inspection of manufacturing facilities by, the relevant regulatory authority. Our product 
candidates may not be effective, may be only moderately effective or may prove to have undesirable or unintended side 
effects, toxicities or other characteristics that may preclude our obtaining marketing approval or prevent or limit 
commercial use.  

The process of obtaining marketing approvals, both in the United States and abroad, is expensive; may take 

many years if additional clinical trials are required, if approval is obtained at all, and can vary substantially based upon a 
variety of factors, including the type, complexity and novelty of the product candidates involved. In the United States, 
for example, the submission fee to obtain U.S. marketing approval is more than $2.0 million, and may be higher in the 
future. Changes in marketing approval policies during the development period, in or the enactment of additional statutes 
or regulations, or in regulatory review for each submitted product application, may cause delays in the approval or 
rejection of an application. The FDA and comparable authorities in other countries have substantial discretion in the 
approval process and may refuse to accept any application or may decide that our data are insufficient for approval and 
require additional preclinical, clinical or other studies. In addition, varying interpretations of the data obtained from 
preclinical and clinical testing could delay, limit or prevent marketing approval of a product candidate. Any marketing 
approval we, or any future collaborators, ultimately obtain may be limited or subject to restrictions or post-approval 
commitments that render the approved product not commercially viable.  

Accordingly, if we or any future collaborators experience delays in obtaining approval or if we or they fail to 

obtain approval of our product candidates, the commercial prospects for our product candidates may be harmed and our 
ability to generate revenues will be materially impaired. 

Even if we obtain regulatory approval for a product candidate, our products will remain subject to regulatory 
oversight. 

Even if we obtain any regulatory approval for our product candidates, they will be subject to ongoing regulatory 

requirements for manufacturing, labeling, packaging, storing, advertising, promoting, sampling, record-keeping and 
submitting safety and other post-market information. Any regulatory approvals that we receive for our product 
candidates also may be subject to a REMS, limitations on the approved indicated uses for which the product may be 
marketed or to the conditions of approval or contain requirements for potentially costly post-marketing testing, including 
Phase 4 clinical trials, and surveillance to monitor the quality, safety and efficacy of the product. For example, the holder 
of an approved BLA is obligated to monitor and report adverse events and any failure of a product to meet the 
specifications in the BLA. FDA guidance advises that patients treated with some types of gene therapy undergo 
follow-up observations for potential adverse events for as long as 15 years. The holder of an approved BLA also must 
submit new or supplemental applications and obtain FDA approval for certain changes to the approved product, product 

78 

labeling or manufacturing process. Advertising and promotional materials must comply with FDA rules and are subject 
to FDA review, in addition to other potentially applicable federal and state laws. 

In addition, product manufacturers and their facilities are subject to payment of user fees and continual review 

and periodic inspections by the FDA and other regulatory authorities for compliance with cGMP, requirements and 
adherence to commitments made in the BLA or foreign marketing application. If we, or a regulatory authority, discover 
previously unknown problems with a product, such as adverse events of unanticipated severity or frequency, or problems 
with the facility where the product is manufactured or such regulatory authority disagrees with the promotion, marketing 
or labeling of that product, the regulatory authority may impose restrictions relative to that product, the manufacturing 
facility or us, including requiring recall or withdrawal of the product from the market or suspension of manufacturing. 

If we fail to comply with applicable regulatory requirements following approval of any of our product 

candidates, a regulatory authority may: 

 

 

 

 

 

 

 

 

 

issue a warning letter asserting that we are in violation of the law; 

seek an injunction or impose administrative, civil or criminal penalties or monetary fines; 

suspend or withdraw regulatory approval; 

suspend any ongoing clinical trials; 

refuse to approve a pending BLA or comparable foreign marketing application, or any supplements thereto, 
submitted by us or our strategic partners; 

restrict the marketing or manufacturing of the product; 

seize or detain the product or otherwise require the withdrawal of the product from the market; 

refuse to permit the import or export of products; or 

refuse to allow us to enter into supply contracts, including government contracts. 

Any government investigation of alleged violations of law could require us to expend significant time and 

resources in response and could generate negative publicity. The occurrence of any event or penalty described above 
may inhibit our ability to commercialize our product candidates and adversely affect our business, financial condition, 
results of operations and prospects. 

In addition, FDA policies, and those of equivalent foreign regulatory agencies, may change and additional 

government regulations may be enacted that could prevent, limit or delay regulatory approval of our product candidates. 
We cannot predict the likelihood, nature or extent of government regulation that may arise from future legislation or 
administrative action, either in the United States or abroad. If we are slow or unable to adapt to changes in existing 
requirements or the adoption of new requirements or policies, or if we are not able to maintain regulatory compliance, 
we may lose any marketing approval that we may have obtained and we may not achieve or sustain profitability, which 
would harm our business, financial condition, results of operations and prospects. 

79 

We face significant competition in an environment of rapid technological change and the possibility that our 
competitors may achieve regulatory approval before us or develop therapies that are more advanced or effective than 
ours, which may harm our business and financial condition, and our ability to successfully market or commercialize 
our product candidates. 

The biopharmaceutical industry is characterized by intense and dynamic competition to develop new 
technologies and proprietary therapies. Any product candidates that we successfully develop into products and 
commercialize may compete with existing therapies and new therapies that may become available in the future. While 
we believe that our gene therapy platform, product programs, product candidates and scientific expertise in the fields of 
gene therapy and neuroscience provide us with competitive advantages, we face potential competition from various 
sources, including larger and better-funded pharmaceutical, specialty pharmaceutical and biotechnology companies, as 
well as from academic institutions, governmental agencies and public and private research institutions.  

We are aware of several companies focused on developing AAV gene therapies in various indications, 
including Abeona Therapeutics, Inc., Adverum Biotechnologies, Inc., Aevitas Therapeutics, Inc., Agilis Biotherapeutics, 
LLC (acquired by PTC Therapeutics, Inc. in 2018), Applied Genetic Technologies Corporation, Asklepios 
BioPharmaceutical, Inc., Audentes Therapeutics, Inc., AveXis, Inc. (acquired by Novartis in 2018), Axovant Sciences 
Ltd., GenSight Biologics SA, LogicBio Therapeutics, Inc., Lysogene SA, MeiraGTx Ltd., NightstaRx Ltd, Prevail 
Therapeutics, Inc., REGENXBio Inc., Sarepta Therapeutics, Inc., Solid Biosciences, Inc., uniQure, Pfizer, and Spark 
Therapeutics, Inc., as well as several companies addressing other methods for modifying genes and regulating gene 
expression. Any advances in gene therapy technology made by a competitor may be used to develop therapies that could 
compete against any of our product candidates.  

We expect that VY-AADC will compete with a variety of therapies currently marketed and in development for 
Parkinson’s disease, including DBS marketed by Medtronic plc, Abbott Laboratories (acquired from St. Jude Medical in 
2017), and other medical device companies, DUOPA/Duodopa marketed by AbbVie Inc., as well as other novel, non-
oral forms of levodopa in development, including Mitsubishi Tanabe Pharma’s ND0612 (acquired from NeuroDerm in 
2017), Acorda Therapeutics’ inhaled levodopa, INBRIJA (CVT-301), and Sunovion Pharmaceuticals’ sublingual 
apomorphine, APL-130277. Gene therapy competition for Parkinson’s disease previously included AMT-090 or AAV-
GDNF, but this was deprioritized by uniQure in 2016. Axovant is developing a second generation LentiVector gene 
therapy, AXO-Lenti-PD (previously OXB-102, licensed from Oxford Biomedica in 2018). MeiraGTx is developing 
AAV-GAD, acquired from Vector Neurosciences, Inc. in 2018.  

We expect that our preclinical programs will compete with a variety of therapies in development, including: 

  VY-SOD102 for a monogenic form of ALS will potentially compete with IONIS-SOD1Rx being developed 
by Ionis Pharmaceuticals, Inc., or Ionis, in collaboration with Biogen Inc., or Biogen, and a gene therapy 
being developed by AveXis (acquired by Novartis in 2018);  

  VY-FXN01 for Friedreich’s ataxia will potentially compete with AAV gene therapies being developed by 
Pfizer and by Agilis Biotherapeutics, LLC (acquired by PTC Therapeutics, Inc. in 2018), and BMN 290 
being developed by BioMarin Pharmaceutical Inc.;  

  VY-HTT01 for Huntington’s disease will potentially compete with IONIS-HTTRx being developed by 

Ionis in collaboration with F. Hoffmann-La Roche Ltd., or Roche, WVE-120101 and WVE-120102 being 
developed by WAVE Life Sciences in collaboration with Takeda Pharmaceuticals, a Zinc Finger Protein 
(ZFP) therapy being developed by Sangamo Therapeutics, Inc. in collaboration with Shire plc, and gene 
therapies being developed by uniQure and Spark;  

  Our Tau program for tauopathies including Alzheimer’s disease, PSP, and FTD will potentially compete 
with tau antibodies being developed by Roche Genentech Inc. in collaboration with AC Immune SA, Eli 
Lilly & Co., AbbVie Inc., Biogen, and several other companies, as well as an antisense oligonucleotide 
program being developed by Ionis in collaboration with Biogen; and 

80 

VY-NAV01 for severe, chronic pain will potentially compete with Nav1.7 inhibitors being developed by 
Biogen, Sunovion, Amgen, Inc., and Astellas Pharma Inc, and Nav1.8 inhibitors being developed by Vertex.  

We are also aware of several companies and institutions who have developed or are developing real-time, intra-
operative, MRI-compatible devices that would compete with V-TAG. Investigators in the Phase 1b, the separate Phase 1 
posterior trajectory trial, and the RESTORE-1 Phase 2 clinical trial of VY-AADC, have used and are using the 
ClearPoint System from MRIC.  

Many of our potential competitors, alone or with their strategic partners, have substantially greater financial, 

technical and other resources, such as larger research and development, clinical, marketing and manufacturing 
organizations. Mergers and acquisitions in the biotechnology and pharmaceutical industries may result in even more 
resources being concentrated among a smaller number of competitors. Smaller and other early stage companies may also 
prove to be significant competitors, particularly through collaborative agreements with large and established companies. 
Our commercial opportunity could be reduced or eliminated if competitors develop and commercialize products that are 
safer, more effective, have fewer or less severe side effects, are more convenient or are less expensive than any products 
that we may develop. Competitors also may obtain FDA or other regulatory approval for their products more rapidly or 
earlier than us or may obtain orphan drug or other marketing exclusivity, which could result in our competitors 
establishing a strong market position before we are able to enter the market or reducing the number of available subjects 
for enrollment in our clinical trials to support regulatory submissions and approvals of our product. Additionally, 
technologies developed or acquired by our competitors may render our potential product candidates uneconomical or 
obsolete, and we may not be successful in marketing our product candidates against competitors. These third parties also 
compete with us in recruiting and retaining qualified scientific and management personnel, establishing clinical trial 
sites, and registering patients for clinical trials. 

In addition, as a result of the expiration or successful challenge of our patent rights, we could face more 

litigation with respect to the validity and scope of patents relating to our competitors’ products. The availability of our 
competitors’ products could limit the demand, and the price we are able to charge, for any products that we may develop 
and commercialize. If we are not able to compete effectively against potential competitors, our business will not grow 
and our financial condition and operations will be harmed. 

Even if we obtain and maintain approval for our product candidates from the FDA, we may never obtain approval for 
our product candidates outside of the United States, which would limit our market opportunities and adversely affect 
our business. 

Approval of a product candidate in the United States by the FDA does not ensure approval of such product 

candidate by regulatory authorities in other countries or jurisdictions, and approval by one foreign regulatory authority 
does not ensure approval by regulatory authorities in other foreign countries or by the FDA. Sales of our product 
candidates outside of the United States will be subject to foreign regulatory requirements governing clinical trials and 
marketing approval. Even if the FDA grants marketing approval for a product candidate, comparable regulatory 
authorities of foreign countries also must approve the manufacturing and marketing of the product candidates in those 
countries. Approval procedures vary among jurisdictions and can involve requirements and administrative review 
periods different from, and more onerous than, those in the United States, including additional preclinical studies or 
clinical trials or manufacturing control requirements. In many countries outside the United States, a product candidate 
must be separately approved for reimbursement before it can be approved for sale in that country. In some cases, the 
price that we intend to charge for our products, if approved, is also subject to approval. We intend to submit a marketing 
authorization application to EMA for approval of our product candidates in the European Union but obtaining such 
approval from the European Commission following the opinion of EMA is a lengthy and expensive process. Even if a 
product candidate is approved, the FDA or the European Commission may limit the indications for which the product 
may be marketed, require extensive warnings on the product labeling or require expensive and time-consuming 
additional clinical trials or reporting as conditions of approval. Regulatory authorities in countries outside of the United 
States and the European Union also have requirements for approval of product candidates with which we must comply 
prior to marketing in those countries. Obtaining foreign regulatory approvals and compliance with foreign regulatory 
requirements could result in significant delays, difficulties and costs for us and could delay or prevent the introduction of 
our product candidates in certain countries. 

81 

Further, clinical trials conducted in one country may not be accepted by regulatory authorities in other 

countries. Regulatory approval for any of our product candidates may be withdrawn. If we fail to comply with the 
regulatory requirements, our target market will be reduced and our ability to realize the full market potential of our 
product candidates will be harmed and our business, financial condition, results of operations and prospects will be 
harmed. 

Additionally, on June 23, 2016, the electorate in the United Kingdom voted in favor of leaving the European 
Union, commonly referred to as Brexit. On March 29, 2017, the country formally notified the European Union of its 
intention to withdraw pursuant to Article 50 of the Lisbon Treaty. Since a significant proportion of the regulatory 
framework in the United Kingdom is derived from European Union directives and regulations, the referendum could 
materially impact the regulatory regime with respect to the approval of our product candidates in the United Kingdom or 
the European Union. Any delay in obtaining, or an inability to obtain, any marketing approvals, as a result of Brexit or 
otherwise, would prevent us from commercializing our product candidates in the United Kingdom and/or the European 
Union and restrict our ability to generate revenue and achieve and sustain profitability. If any of these outcomes occur, 
we may be forced to restrict or delay efforts to seek regulatory approval in the United Kingdom and/or European Union 
for our product candidates, which could significantly and materially harm our business. 

The United Kingdom has a period of a maximum of two years from the date of its formal notification to 

negotiate the terms of its withdrawal from, and future relationship with, the European Union. If no formal withdrawal 
agreement is reached between the United Kingdom and the European Union, then it is expected the United Kingdom’s 
membership of the European Union will automatically terminate two years after the submission of the notification of the 
United Kingdom’s intention to withdraw from the European Union. Discussions between the United Kingdom and the 
European Union focused on finalizing withdrawal issues and transition agreements are ongoing. However, limited 
progress to date in these negotiations and ongoing uncertainty within the UK Government and Parliament sustains the 
possibility of the United Kingdom leaving the European Union on March 29, 2019 without a withdrawal agreement and 
associated transition period in place, which is likely to cause significant market and economic disruption. 

Risks Related to Third Parties 

To date, all of our revenue has been derived from our collaborations with Sanofi Genzyme and AbbVie, and if any of 
these, our other Strategic Collaborations, or future collaboration agreements were to be terminated, our business 
financial condition, results of operations and prospects would be harmed. 

In February 2015, we entered into the Sanofi Genzyme Collaboration Agreement to leverage our combined 

expertise and assets in gene therapy for neurological diseases. Under the Sanofi Genzyme Collaboration Agreement, we 
received an upfront commitment of approximately $100.0 million. Pursuant to the agreement, we granted Sanofi 
Genzyme an exclusive option to license, develop and commercialize (i) ex-U.S. rights to our Parkinson’s disease, 
Friedreich’s ataxia and Huntington’s disease programs and a future program, or the Split Territory Programs, with an 
incremental option to co-commercialize the product candidate from our Huntington’s disease program in the United 
States and (ii) worldwide rights to our spinal muscular atrophy program. If Sanofi Genzyme exercises an option for a 
Split Territory Program, except for our Parkinson’s disease program, it is required to make an option exercise payment to 
us. Furthermore, at the inception of the agreement, we were eligible to receive up to $745.0 million in the aggregate upon 
the achievement of specified regulatory and commercial milestones, as well as tiered royalty payments based on a 
percentage of net sales of product candidates from the programs for which Sanofi Genzyme exercised its option, or the 
Optioned Programs. Our research and development activities in connection with the collaboration might not be 
successful. We cannot accurately predict when or if any of our product candidates will prove effective or safe in humans 
or whether these product candidates will receive marketing approval. There can be no assurance that we will not 
experience problems or delays in developing our product candidates and that such problems or delays will not cause 
unanticipated costs, or that any such development problems can be solved. If Sanofi Genzyme were to elect not to 
exercise an option, we would have incurred significant development expenses but not receive the option exercise 
payment or be eligible to receive future milestone or royalty payments related to such program. For example, in October 
2017, Sanofi Genzyme notified us that it had decided not to exercise its option for the ex-U.S. rights to VY-AADC and 
terminated the portion of its collaboration with us concerning Parkinson’s disease. As a result of Sanofi Genzyme’s 
decision, the Company is no longer entitled to receive up to $105.0 million of milestone payments related to the 
Parkinson’s program. 

82 

If Sanofi Genzyme exercises one or more options, following such exercise, Sanofi Genzyme will have sole 

responsibility for the development and commercialization of the product candidates from such program in the applicable 
territory. Sanofi Genzyme will have the sole discretion to determine and direct its efforts and resources, including the 
ability to discontinue all efforts and resources it applies to the development and, if approval is obtained, 
commercialization and marketing of the product candidates covered by the Optioned Programs in the applicable 
territories. Sanofi Genzyme may not be effective in obtaining approvals for the product candidates developed from the 
Optioned Programs or in marketing, or arranging for necessary supply, manufacturing or distribution relationships for, 
any approved products. Furthermore, Sanofi Genzyme may change its strategic focus or pursue alternative technologies 
in a manner that results in reduced, delayed or no revenue to us. Sanofi Genzyme has a variety of marketed products and 
product candidates under collaboration with other companies, including some of our competitors, and its own corporate 
objectives may not be consistent with our best interests. If Sanofi Genzyme fails to develop, obtain regulatory approval 
for or ultimately commercialize any product candidate from the Optioned Programs in the applicable territories, or if 
Sanofi Genzyme terminates our collaboration, our business, financial condition, results of operations and prospects 
would be harmed.  

In addition, any dispute or litigation proceedings we may have with Sanofi Genzyme in the future could delay 

development programs, create uncertainty as to ownership of or access to intellectual property rights, distract 
management from other business activities and generate substantial expense. Finally, we also may not be able to seek 
and obtain a viable, alternative collaborator to partner with for the development and commercialization of the Split 
Territory Programs or the Optioned Programs. 

In February 2018, we entered into an exclusive collaboration and option agreement with AbbVie, which we 
refer to as the AbbVie Tau Collaboration Agreement, for the research, development, and commercialization of AAV 
gene therapy products for the treatment of diseases of the central nervous system and other neurodegenerative diseases 
related to defective or excess aggregation of tau protein in the human brain, including Alzheimer’s disease. Under the 
terms of the AbbVie Tau Collaboration Agreement, we received an upfront payment of $69.0 million and may receive 
option exercise payments, future development and regulatory milestone payments and royalties.  

Under the AbbVie Tau Collaboration Agreement, we are obligated to use diligent efforts to conduct research 

and development activities, including IND-enabling and Phase 1 clinical trial activities, for which we are solely 
financially responsible. As described above, our research and development activities in connection with a collaboration 
might not be successful. The AbbVie Tau Collaboration Agreement shall automatically terminate if AbbVie decides not 
to exercise one or more of its options prior to the expiration of the specified periods of the collaboration. If AbbVie did 
not exercise one or more of its options, we would have incurred significant research and development expenses but 
would not receive any future option exercise, milestone payments, or royalty payments under the collaboration. 

Such research and development activities shall be pursuant to plans agreed to by the parties and overseen by a 
joint governance committee, or JGC. Initially, we will have final decision-making authority within the JGC, subject to 
specified limitations. If AbbVie exercises its license option, however, AbbVie will assume final decision-making 
authority within the JGC. If AbbVie exercises its license option, it will also be solely responsible for all development and 
commercialization activities relating to compounds and products licensed pursuant to the agreement, and, at AbbVie’s 
request, we could be required to effect a full transfer of the manufacturing process for such compound and products. 
Even if AbbVie does not exercise any of its options under the AbbVie Tau Collaboration Agreement, we have agreed to 
grant AbbVie perpetual, exclusive or non-exclusive (as the case may be), worldwide licenses to certain know-how and 
patent rights developed by us or jointly by us and AbbVie arising from the collaboration. 

AbbVie might not be effective in administering the JGC, obtaining approvals for the product candidates arising 

from our collaboration, or commercializing or manufacturing the resulting products. Further, AbbVie’s objectives in 
connection with the collaboration may not be consistent with our best interests. AbbVie could use its leadership of the 
JGC or intellectual property it has licensed under the AbbVie Tau Collaboration Agreement in a manner adverse to us, or 
it could halt, slow, or deprioritize its development and commercialization efforts under the collaboration. In any such 
instances, our business, financial condition, results of operations and prospects could be materially harmed. 

In January 2019, we entered into the Neurocrine Collaboration Agreement for the research, development and 
commercialization of four programs including our Parkinson’s disease program, or AADC Program, our Friedreich’s 
ataxia program, or FA Program, and two programs to be determined by us and Neurocrine at a later date, or the 

83 

Discovery Programs. Under the terms of the agreement, we will receive an upfront payment of $165.0 million, inclusive 
of $50.0 million for 4,179,728 shares of our common stock and may receive future development and regulatory 
milestones and royalties. The Neurocrine Collaboration Agreement is subject to certain conditions, including the 
expiration or termination of the applicable waiting period under the Hart-Scott-Rodino Antitrust Improvements Act of 
1976, as amended, and other customary closing conditions.  

Under the Neurocrine Collaboration Agreement, upon the expiration or termination of applicable waiting 

periods and the receipt of any required approvals or clearances including antitrust clearance, we have agreed to 
collaborate on the conduct of four collaboration programs: the AADC Program for the treatment of Parkinson’s disease, 
our FA Program for the treatment of Friedreich’s ataxia including the development of the VY-FXN01 product candidate, 
and two programs Discovery Programs. 

Under the terms of the Neurocrine Collaboration Agreement, subject to the rights retained by us thereunder, we 
have agreed to collaborate with Neurocrine on, and to grant, exclusive, royalty-bearing, non-transferable, sublicensable 
licenses to certain of our intellectual property rights for all human and veterinary diagnostic, prophylactic, and 
therapeutic uses for the research, development, and commercialization of gene therapy Collaboration Products, under (i) 
the AADC Program, on a worldwide basis; (ii) the FA Program, in the United States and, upon expiration of Sanofi 
Genzyme’s option to the Friedreich’s ataxia program pursuant to the Sanofi Genzyme Collaboration without exercise of 
such option, all countries in the world in which the Neurocrine Collaboration Agreement remained in effect with respect 
to the FA Program; and (iii) each Discovery Program, on a worldwide basis. 

Pursuant to development plans to be agreed by the parties, which will be overseen by a joint steering committee, 

we have operational responsibility, subject to certain exceptions, for the conduct of each Neurocrine Program (prior to 
specified transition events for each program), and are required to use commercially reasonable efforts to develop the 
Collaboration Products. Neurocrine has agreed to be responsible for all costs incurred by us in conducting these activities 
for each Neurocrine Program in accordance with an agreed budget. If we breach our development responsibilities or in 
certain circumstances upon a change in control of us, Neurocrine has the right but not the obligation to assume the 
activities under such Neurocrine Program. 

Upon the occurrence of specified events for each program, Neurocrine has agreed to assume responsibility for 

development, manufacturing and commercialization activities for such program and to pay us milestones and royalties on 
future net sales. For each Existing Program, we have the option to co-develop and co-commercialize such Program upon 
the occurrence of a specified event. Should we elect to exercise our co-development and co-commercialization option, 
we and Neurocrine have agreed to enter into a cost- and profit-sharing arrangement whereby we and Neurocrine agree to 
jointly develop and commercialize Collaboration Products for such program and share in its costs, profits and losses, and 
we have agreed to forfeit certain milestones and royalties on net sales in the United States during the effective period of 
the applicable co-development and co-commercialization agreement. As described above, our research and development 
activities in connection with a collaboration might not be successful. Neurocrine may terminate the collaboration 
agreement in its entirety or on a program-by-program or country-by-country basis by providing at least 180-day advance 
notice if such notice is provided prior to the first commercial sale of the Collaboration Product to which the termination 
applies or one-year advance notice if such notice is provided after the first commercial sale of the Collaboration Product 
to which the termination applies. If Neurocrine were to terminate the agreement, we would become responsible for all 
research and development expenses relating to the Neurocrine Programs, and would not receive any future milestone 
payments or royalty payments under the Neurocrine Collaboration Agreement. The Neurocrine Collaboration Agreement 
may also be terminated if specified regulatory agencies seek to enjoin the transaction or if we are unable to obtain 
antitrust clearance within 180 days of the applicable antitrust filings, which would prevent us from receiving any 
payments from Neurocrine under the agreement. 

Neurocrine might not be successful in obtaining approvals for the product candidates arising from our 
collaboration, or commercializing or manufacturing the resulting products. Further, Neurocrine’s objectives in 
connection with the collaboration may not be consistent with our best interests. With respect to the rights granted to 
Neurocrine by us, Neurocrine could take actions that may be adverse to us, or it could halt, slow, or deprioritize its 
development and commercialization efforts under the collaboration. In any such instances, our business, financial 
condition, results of operations and prospects could be materially harmed. 

84 

 
  
 
In February 2019, we entered into the AbbVie Alpha-Synuclein Collaboration Agreement for the research, 
development, and commercialization of AAV gene therapy products directed against alpha-synuclein for indications 
including Parkinson’s disease and other synucleinopathies. Under the terms of the AbbVie Alpha-Synuclein 
Collaboration Agreement, we received an upfront payment of $65.0 million and may receive option exercise payments, 
future development and regulatory milestone payments and royalties.  

Under the terms of the agreement, we and AbbVie have agreed to collaborate on the research and development 

of the specified vectorized antibody compounds, or the Research Compounds. We are obligated to conduct research 
activities directed to constructing one or more virus vectors that encode antibodies designated by AbbVie. We are 
obligated to use diligent efforts to conduct antibody engineering and other research activities to create Research 
Compounds and to develop product candidates containing or comprised of such Research Compounds, which we refer to 
as Product Candidates. We are solely responsible for the costs and expenses during the research period. During a 
specified portion of the Research Period, AbbVie may exercise one or more of its exclusive development options to 
select up to a total of four Research Compounds and their corresponding product candidates to proceed to the 
Development Period. As described above, our research and development activities in connection with a collaboration 
might not be successful. The AbbVie Alpha-Synuclein Collaboration Agreement shall automatically terminate if AbbVie 
decides not to exercise one or more of its options prior to the expiration of the specified periods of the collaboration. If 
AbbVie did not exercise one or more of its options, we would have incurred significant research and development 
expenses but would not receive any future option exercise, milestone payments, or royalty payments under the 
collaboration. 

Such research and development activities shall be pursuant to plans agreed to by the parties and overseen by a 
JGC. Initially, we will have final decision-making authority within the JGC, subject to specified limitations. If AbbVie 
exercises its license option, however, AbbVie will assume final decision-making authority within the JGC. If AbbVie 
exercises its license option, it will also be solely responsible for all development and commercialization activities 
relating to compounds and products licensed pursuant to the agreement, and, at AbbVie’s request, we could be required 
to effect a full transfer of the manufacturing process for such compound and products. Even if AbbVie does not exercise 
any of its options under the AbbVie Alpha-Synuclein Collaboration Agreement, we have agreed to grant AbbVie 
perpetual, exclusive or non-exclusive (as the case may be), worldwide licenses to certain know-how and patent rights 
developed by us or jointly by us and AbbVie arising from the collaboration. 

AbbVie might not be effective in administering the JGC, obtaining approvals for the product candidates arising 

from our collaboration, or commercializing or manufacturing the resulting products. Further, AbbVie’s objectives in 
connection with the collaboration may not be consistent with our best interests. AbbVie could use its leadership of the 
JGC or intellectual property it has licensed under the AbbVie Alpha-Synuclein Collaboration Agreement in a manner 
adverse to us, or it could halt, slow, or deprioritize its development and commercialization efforts under the 
collaboration. In any such instances, our business, financial condition, results of operations and prospects could be 
materially harmed. 

We have only used the ClearPoint System to deliver our product candidates. While other devices for delivery may be 
used in the future, any issues with the ClearPoint System or the manufacturer of the ClearPoint System, may result in 
delays in the development and commercialization of certain of our product candidates, which could have an adverse 
impact on our business. 

The surgical approach that we are using for VY-AADC is similar, in some respects, to the stereotactic approach 

used for DBS. One primary difference with our approach is the ability to assist the physician in visualizing the delivery 
of VY-AADC to the putamen using real-time, intra-operative, MRI to avoid specific blood vessels to potentially reduce 
the risk of hemorrhages during the surgical procedure and to maximize the coverage of the putamen.  

Investigators in the Phase 1b clinical trial, the separate Phase 1 posterior trajectory trial, and the RESTORE-1 

Phase 2 clinical trial for VY-AADC, have used and are using the real-time, intra-operative, MRI system called the 
ClearPoint System from MRIC. However, not all neurosurgical units within the United States utilize this system and may 
employ other neuro-navigational systems that are not compatible with real-time MRI imaging. We intend to use the 
ClearPoint System at certain sites in our RESTORE-1 Phase 2 clinical trial and may choose to use it in future clinical 
trials of VY-AADC and any other of our product candidates that are injected directly into the brain. Therefore, any 

85 

issues with the ClearPoint System, such as a finding that use of the ClearPoint System causes adverse events or a product 
recall, or issues with the manufacturer of the ClearPoint System, such as bankruptcy or a decision to stop production of 
the system due to lack of profitability, could delay the development or commercialization of certain of our product 
candidates, including VY-AADC, as there currently is no other manufacturer of the ClearPoint System. Outside the 
United States, the ClearPoint System is not widely available or utilized in neurosurgical units.  

Management of MRIC, the manufacturer of the ClearPoint System, has expressed substantial doubt as to the 
ability of MRIC to continue as a going concern on several occasions. As of September 30, 2018, MRIC reported cash 
and cash equivalents of $3.7 million and secured debt (senior and junior) totaling approximately $3.3 million on its 
balance sheet. MRIC also reported a net loss of $1.4 million and $4.9 million for the three and nine months ended 
September 30, 2018. 

We developed V-TAG as our own real-time, intra-operative, MRI-compatible device that can be used with 

other neuro-navigational systems to dose VY-AADC and for other surgical procedures. We believe that the experience 
we have gained from delivering VY-AADC in our clinical trials to date and our work to develop V-TAG may inform 
AAV gene therapy delivery for our Huntington’s disease program and other projects. In July 2018, we received 510(k) 
regulatory clearance of V-TAG from the Center for Devices and Radiological Health of the FDA. There are additional 
steps needed in making this device available for use including the manufacture of the product and compliance with state 
and federal laws and regulations for medical devices. 

We are currently exploring collaborations for V-TAG and expect to rely on third parties in the development and 

manufacture of the device. In May 2018, for example, we entered into a master services and supply agreement with 
MRIC which provides for MRIC to perform certain manufacturing, supply, development, and services as requested by 
us, including the supply of the ClearPoint System and cannula devices as well as to collaborate on V-TAG. 
Collaborations, including our collaboration with MRIC, are subject to risks similar to those described elsewhere in this 
“Risk Factors” section, and the corporate objectives of such current and potential future collaborators may not be 
consistent with our best interests. If we are unable to enter into additional collaborations or any collaborations are not 
successful, use of V-TAG in our clinical trials could be adversely affected, and our clinical trials, including our 
RESTORE-1 Phase 2 clinical trial, could be delayed.  

We may seek to enter into collaborations in the future with other third parties. If we are unable to enter into such 
collaborations, or if these collaborations are not successful, our business could be adversely affected. 

We may seek to enter into additional collaborations in the future, including sales, marketing, distribution, 

development, licensing, and/or broader collaboration agreements. Our likely collaborators include large and mid-size 
pharmaceutical companies, regional and national pharmaceutical companies, biotechnology companies, and medical 
device manufacturers. However, we may not be able to enter into additional collaborations on favorable terms or at all. 
Our ability to generate revenues from our collaborations will depend on our and our collaborators’ abilities to 
successfully perform the functions assigned to each of us in these arrangements. In addition, our collaborators might 
have the ability to abandon research or development projects and terminate applicable agreements. Moreover, an 
unsuccessful outcome in any clinical trial for which our collaborator is responsible could be harmful to the public 
perception and prospects of our gene therapy platform. 

Our relationship with any future collaborations may pose several risks, including the following: 

 

 

 

collaborators have significant discretion in determining the amount and timing of the efforts and resources 
that they will apply to these collaborations; 

collaborators may not perform their obligations as expected; 

the preclinical studies and clinical trials conducted as part of these collaborations may not be successful; 

86 

 

 

collaborators may not pursue development and commercialization of any product candidates that achieve 
regulatory approval or may elect not to continue or renew development or commercialization programs 
based on preclinical study or clinical trial results, changes in the collaborators’ strategic focus or available 
funding or external factors, such as an acquisition, that divert resources or create competing priorities; 

collaborators may delay preclinical studies and clinical trials, provide insufficient funding for preclinical 
studies and clinical trials, stop a preclinical study or clinical trial or abandon a product candidate, repeat or 
conduct new preclinical studies or clinical trials or require a new formulation of a product candidate for 
preclinical studies or clinical trials; 

  we may not have access to, or may be restricted from disclosing, certain information regarding product 

candidates being developed or commercialized under a collaboration and, consequently, may have limited 
ability to inform our stockholders about the status of such product candidates; 

 

 

 

 

 

 

 

 

 

collaborators could independently develop, or develop with third parties, products that compete directly or 
indirectly with our product candidates if the collaborators believe that competitive products are more likely 
to be successfully developed or can be commercialized under terms that are more economically attractive 
than ours; 

product candidates developed in collaboration with us may be viewed by our collaborators as competitive 
with their own product candidates or products, which may cause collaborators to cease to devote resources 
to the commercialization of our product candidates; 

a collaborator with marketing and distribution rights to one or more of our product candidates that achieve 
regulatory approval may not commit sufficient resources to the marketing and distribution of any such 
product candidate; 

disagreements with collaborators, including disagreements over proprietary rights, contract interpretation or 
the preferred course of development of any product candidates, may cause delays or termination of the 
research, development or commercialization of such product candidates, may lead to additional 
responsibilities for us with respect to such product candidates or may result in litigation or arbitration, any 
of which would be time-consuming and expensive; 

collaborators may not properly maintain or defend our intellectual property rights or may use our 
proprietary information in such a way as to invite litigation that could jeopardize or invalidate our 
intellectual property or proprietary information or expose us to potential litigation; 

disputes may arise with respect to the ownership or inventorship of intellectual property developed 
pursuant to our collaborations; 

collaborators may infringe the intellectual property rights of third parties, which may expose us to litigation 
and potential liability; 

the terms of our collaboration agreement may restrict us from entering into certain relationships with other 
third parties, thereby limiting our options; and 

collaborations may be terminated for the convenience of the collaborator and, if terminated, we could be 
required to raise additional capital to pursue further development or commercialization of the applicable 
product candidates. 

Collaboration agreements may not lead to the development or commercialization of product candidates in the 

most efficient manner, or at all. If our collaborations do not result in the successful development and commercialization 
of products, or if one of our collaborators terminates its agreement with us, we may not receive any future research 

87 

funding or milestone or royalty payments under the collaboration. If we do not receive the funding we expect under these 
agreements, our development of our product candidates could be delayed, and we may need additional resources to 
develop our product candidates. Additionally, subject to its contractual obligations to us, if a collaborator of ours were to 
be involved in a business combination, it might deemphasize or terminate the development or commercialization of any 
product candidate licensed to it by us. If one of our collaborators terminates its agreement with us, we may find it more 
difficult to attract new collaborators, and the perception of us in the business and financial communities could be 
adversely affected. All of the risks relating to product development, regulatory approval and commercialization 
described in this periodic report also apply to the activities of our collaborators. 

We will face significant competition in seeking appropriate collaborators, and the negotiation process is time-

consuming and complex. Our ability to reach a definitive collaboration agreement with any future collaborators will 
depend, among other things, upon our assessment of the collaborator’s resources and expertise, the terms and conditions 
of the proposed collaboration and the proposed collaborator’s evaluation of several factors. Those factors may include 
the design or results of clinical trials, the likelihood of approval by the FDA or similar regulatory authorities outside the 
United States, the potential market for the subject product candidate, the costs and complexities of manufacturing and 
delivering such product candidate to patients, the potential of competing products, the existence of uncertainty with 
respect to our ownership of technology, which can exist if there is a challenge to such ownership without regard to the 
merits of the challenge, and industry and market conditions generally. The collaborator may also consider alternative 
product candidates or technologies for similar indications that may be available to collaborate on and whether such a 
collaboration could be more attractive than the one with us for our product candidate. We may also be restricted under 
future license agreements from entering into agreements on certain terms with potential collaborators. In addition, there 
have been a significant number of recent business combinations among large pharmaceutical companies that have 
resulted in a reduced number of potential future collaborators.  

If we are unable to reach agreements with suitable collaborators on a timely basis, on acceptable terms, or at all, 
we may have to curtail the development of a product candidate, reduce or delay its development program or one or more 
of our other development programs, delay its potential commercialization or reduce the scope of any sales or marketing 
activities, or increase our expenditures and undertake development or commercialization activities at our own expense. If 
we elect to fund and undertake development or commercialization activities on our own, we may need to obtain 
additional expertise and additional capital, which may not be available to us on acceptable terms or at all. If we fail to 
enter into collaborations and do not have sufficient funds or expertise to undertake the necessary development and 
commercialization activities, we may not be able to further develop our product candidates or bring them to market or 
continue to develop our gene therapy platform. If we license rights to product candidates, we may not be able to realize 
the benefit of such transactions if we are unable to successfully integrate them with our existing operations and company 
culture. 

We have relied, and we expect to continue to rely on third parties to conduct, supervise and monitor our clinical trials, 
and if these third parties perform in an unsatisfactory manner, our business could be harmed. 

We expect to rely on CROs and clinical trial sites to ensure our clinical trials are conducted properly and on 

time. We may also engage third parties such as clinical data management organizations, medical institutions and clinical 
investigators to conduct or assist in our clinical trials or other clinical development work. While we will have agreements 
governing their activities, we will have limited influence over their actual performance. We will control only certain 
aspects of our third-party service providers’ activities. Nevertheless, we will be responsible for ensuring that each of our 
clinical studies is conducted in accordance with the applicable protocol, legal, regulatory and scientific standards. Our 
reliance on these third-parties does not relieve us of our regulatory responsibilities. For example, our Phase 1b clinical 
trial of VY-AADC and our separate Phase 1 trial exploring the delivery of VY-AADC using a posterior trajectory were 
conducted at several locations, including UCSF and UPMC. We expect to conduct our RESTORE-1 Phase 2 clinical trial 
at over twenty clinical trial sites, including neurosurgical and neurology patient referral sites in the United States and 
Europe. If any locations terminate the clinical trial, we would be required to find another party to conduct any new trials. 
We may be unable to find a new party to conduct new trials of our product candidates or obtain clinical supply of our 
product candidates or AAV vectors for such trials. 

We and our third-party service providers are required to comply with the FDA’s GCPs for conducting, 

recording and reporting the results of IND-enabling studies and clinical studies to assure that the data and reported 
results are credible and accurate and that the rights, integrity and confidentiality of clinical trial participants are 

88 

protected. We are also required to register ongoing clinical trials and post the results of completed clinical trials on a 
government-sponsored database, ClinicalTrials.gov, within specified timeframes. The FDA enforces these GCPs through 
periodic inspections of trial sponsors, principal investigators and clinical trial sites at which the FDA may determine that 
our clinical trials did not comply with GCPs. If we or our third-party service providers fail to comply with applicable 
GCPs, the clinical data generated in our future clinical trials may be deemed unreliable and the FDA may require us to 
perform additional clinical trials before approving any marketing applications. In addition, our future clinical trials will 
require a sufficient number of patients to evaluate the safety and effectiveness of our product candidates. Accordingly, if 
we or our third-party service providers fail to comply with these regulations or fail to recruit a sufficient number of 
patients, we may be required to repeat such clinical trials, which would delay the regulatory approval process. Failure to 
comply can also result in fines, adverse publicity, and civil and criminal sanctions.  

Our third-party service providers are not our employees, and we are therefore unable to directly monitor 

whether or not they devote sufficient time and resources to our clinical and nonclinical programs. These third-party 
service providers may also have relationships with other commercial entities, including our competitors, for whom they 
may also be conducting clinical trials or other drug development activities that could harm our competitive position. If 
our third-party service providers do not successfully carry out their contractual duties or obligations, fail to meet 
expected deadlines, or if the quality or accuracy of the clinical data they obtain is compromised due to the failure to 
adhere to our clinical protocols or regulatory requirements, or for any other reasons, our clinical trials may be extended, 
delayed or terminated, and we may not be able to obtain regulatory approval for, or successfully commercialize our 
product candidates. As a result, our financial results and the commercial prospects for our product candidates would be 
harmed, our costs could increase, and our ability to generate revenues could be delayed. 

Risks Related to Manufacturing 

Gene therapies and their companion diagnostics are novel, complex and difficult to manufacture. We could 
experience manufacturing problems that result in delays in the development or commercialization of our product 
candidates or otherwise harm our business. 

The manufacture of gene therapy products is technically complex and necessitates substantial expertise and 
capital investment. Production difficulties caused by unforeseen events may delay the availability of material for our 
clinical studies. To meet the requirements of our current and planned future trials we have developed a proprietary 
manufacturing platform that provides a robust and scalable process for AAV production. We are using the 
baculovirus/Sf9 AAV production system, a technology for producing AAV gene therapy vectors at scale in insect-
derived cells. The process has been successfully transferred to our contract manufacturing organizations where it is used 
in manufacturing clinical materials in accordance with the FDA’s current good manufacturing practices, or cGMPs. We 
have also built an onsite, state-of-the-art process research and development facility to enable the manufacturing of high 
quality AAV gene therapy vectors at laboratory scale. 

We presently contract with third parties for the manufacturing of our program materials. We currently have no 

plans to build our own clinical or commercial scale manufacturing capabilities. The use of contracted manufacturing and 
reliance on collaboration partners is relatively cost efficient and has eliminated the need for our direct investment in 
manufacturing facilities and additional staff early in development. Although we rely on contract manufacturers, we have 
personnel with manufacturing and quality experience to oversee our contract manufacturers. 

To date, our third-party manufacturers have met our manufacturing requirements for our program materials. We 

expect third-party manufacturers to be capable of providing sufficient quantities of our program materials to meet 
anticipated clinical trial scale demands. To meet our projected needs for commercial manufacturing, third parties with 
whom we currently work might need to increase their scale of production or we will need to secure alternate suppliers. 
We believe that there are alternate sources of supply for our program materials that can satisfy our clinical and 
commercial requirements, although we cannot be certain that identifying and establishing relationships with such 
sources, if necessary, would not result in significant delay or material additional costs. 

To date, our third-party manufacturers have met our quality standards for our program materials. The 

manufacturers of pharmaceutical products must comply with strictly enforced cGMP requirements, state and federal 

89 

regulations, as well as foreign requirements when applicable. Any failure of us or our contract manufacturing 
organizations to adhere to or document compliance to such regulatory requirements could lead to a delay or interruption 
in the availability of our program materials for clinical study. If we or our manufacturers were to fail to comply with the 
FDA, EMA, or other regulatory authority, it could result in sanctions being imposed on us, including clinical holds, 
fines, injunctions, civil penalties, delays, suspension or withdrawal of approvals, license revocation, seizures or recalls of 
product candidates or products, operating restrictions and criminal prosecutions, any of which could significantly and 
adversely affect supplies of our product candidates. Our potential future dependence upon others for the manufacture of 
our product candidates may also adversely affect our future profit margins and our ability to commercialize any product 
candidates that receive regulatory approval on a timely and competitive basis. 

Biological products are inherently difficult to manufacture. Our program materials are manufactured using 

technically complex processes requiring specialized equipment and facilities, highly specific raw materials, cells, and 
reagents, and other production constraints. Our production process requires a number of highly specific raw materials, 
cells, and reagents with limited suppliers. Even though we aim to have backup supplies of raw materials, cells, and 
reagents whenever possible, we cannot be certain they will be sufficient if our primary sources are unavailable. A 
shortage of a critical raw material, cell line, or reagent, or a technical issue during manufacturing may lead to delays in 
clinical development or commercialization plans. Any changes in the manufacturing of components of the raw materials 
we use could result in unanticipated or unfavorable effects in our manufacturing processes, resulting in delays.  

Companion diagnostic devices may be required to diagnose a genetic disease or to determine patient antibody 

levels to certain components in a product, and could also require a sophisticated, technically complex manufacturing 
processes. If we or our contract manufacturing organizations fail to manufacture such diagnostics or comply with 
relevant regulatory requirements or approvals, we might seek to transition such manufacturing processes to another 
contract manufacturing organization. We might not be able to transition such processes in a timely manner or at all, and 
our commercialization and development efforts could be delayed. 

Delays in obtaining regulatory approval of our or our collaborators’ manufacturing process and facility or 
disruptions in our manufacturing process may delay or disrupt our commercialization efforts. Until recently, no 
cGMP gene therapy manufacturing facility in the United States had received approval from the FDA for the 
manufacture of an approved gene therapy product. 

Before we can begin to commercially manufacture our product candidates in our own facility, or the facility of a 

collaborator, we must obtain regulatory approval from the FDA for our manufacturing process and our collaborator’s 
facility. A manufacturing authorization must also be obtained from the appropriate European Union regulatory 
authorities. Until recently, no cGMP gene therapy manufacturing facility in the United States had received approval from 
the FDA for the manufacture of an approved gene therapy product and, therefore, the timeframe required for us to obtain 
such approval is uncertain. In addition, we must pass a pre-approval inspection of our or our collaborator’s 
manufacturing facility by the FDA and other relevant regulatory authorities before any of our product candidates can 
obtain marketing approval. In order to obtain approval, we will need to ensure that all of our processes, methods and 
equipment are compliant with cGMP, and perform extensive audits of vendors, contract laboratories and suppliers. If any 
of our vendors, contract laboratories or suppliers is found to be out of compliance with cGMP, we may experience delays 
or disruptions in manufacturing while we work with these third parties to remedy the violation or while we work to 
identify suitable replacement vendors. The cGMP requirements govern quality control of the manufacturing process and 
documentation policies and procedures. In complying with cGMP, we will be obligated to expend time, money and effort 
in production, record keeping and quality control to assure that the product meets applicable specifications and other 
requirements. If we fail to comply with these requirements, we would be subject to possible regulatory action and may 
not be permitted to sell any products that we may develop. 

Failure to comply with ongoing regulatory requirements could cause us to suspend production or put in place costly 
or time-consuming remedial measures. 

The regulatory authorities may, at any time, following approval of a product for sale, audit the manufacturing 
facilities for such product, or institute biennial inspections. If any such inspection or audit identifies a failure to comply 
with applicable regulations, or if a violation of product specifications or applicable regulations occurs independent of 
such an inspection or audit, the relevant regulatory authority may require remedial measures that may be costly or 

90 

time-consuming to implement and that may include the temporary or permanent suspension of a clinical trial or 
commercial sales or the temporary or permanent closure of a manufacturing facility. Any such remedial measures 
imposed upon our third-party manufacturers or us could harm our business, financial condition, results of operations and 
prospects. 

If our third-party manufacturers or we fail to comply with applicable cGMP regulations, FDA and foreign 
regulatory authorities can impose regulatory sanctions including, among other things, refusal to approve a pending 
application for a new product candidate or suspension or revocation of a pre-existing approval. Such an occurrence may 
cause our business, financial condition, results of operations and prospects to be harmed. 

Additionally, if supply from any third-party manufacturers is delayed or interrupted, there could be a significant 

disruption in the supply of our clinical or commercial material. We have agreements in place with our contract 
manufacturers pursuant to which we are collaborating on cGMP manufacturing processes and analytical methods for the 
manufacture of our AAV product candidates. Therefore, if we are unable to enter into an agreement with our contract 
manufacturers to manufacture clinical or commercial material for our product programs, or if our agreement with our 
contract manufacturers were terminated, we would have to find suitable alternative manufacturers. This could delay our 
or our collaborators’ ability to conduct clinical trials or commercialize our current and future product candidates. The 
regulatory authorities also may require additional trials if a new manufacturer is relied upon for commercial production. 
Switching manufacturers may involve substantial costs and could result in a delay in our desired clinical and commercial 
timelines. 

Any contamination in the manufacturing process for our products or product candidates, shortages of raw materials, 
cells or reagents, or failure of any of our key suppliers to deliver necessary components could result in delays in our 
clinical development or marketing schedules. 

Given the nature of biologics manufacturing, there is a risk of contamination. Any contamination could 

adversely affect our ability to produce product candidates on schedule and could, therefore, harm our results of 
operations and cause reputational damage. 

Some of the raw materials required in our manufacturing process are derived from biologic sources. Such raw 

materials are difficult to procure and may be subject to contamination or recall. A material shortage, contamination, 
recall or restriction on the use of biologically derived substances in the manufacture of our product candidates could 
adversely impact or disrupt the commercial manufacturing or the production of clinical material, which could adversely 
affect our development timelines and our business, financial condition, results of operations and prospects. 

Interruptions in the supply of product candidates or inventory loss may harm our operating results and financial 
condition. 

Our product candidates are manufactured using technically complex processes requiring specialized facilities, 

highly specific raw materials and other production constraints. The complexity of these processes, as well as strict 
government standards for the manufacture and storage of our product candidates, subjects us to manufacturing risks. 
While product candidate batches released for use in clinical trials or for commercialization undergo sample testing, some 
defects may only be identified following product release. In addition, process deviations or unanticipated effects of 
approved process changes may result in these intermediate products not complying with stability requirements or 
specifications. Our product candidates must be stored and transported at temperatures within a certain range. If these 
environmental conditions deviate, our product candidates’ remaining shelf-lives could be impaired or their efficacy and 
safety could be negatively impacted, making them no longer suitable for use. 

The occurrence, or suspected occurrence, of manufacturing and distribution difficulties can lead to lost 
inventories and, in some cases, product recalls, with consequential reputational damage and the risk of product liability. 
The investigation and remediation of any identified problems can cause production delays, substantial expense, lost sales 
and delays of new product launches. Any interruption in the supply of finished products or the loss thereof could hinder 
our ability to timely distribute our products and satisfy customer demand. Any unforeseen failure in the storage of the 
product or loss in supply could delay our clinical trials and, if our product candidates are approved, result in a loss of our 
market share and negatively affect our business, financial condition, results of operations and prospects. 

91 

Risks Related to Our Business Operations 

We may not be successful in our efforts to identify or discover additional product candidates and may fail to capitalize 
on programs or product candidates that may be a greater commercial opportunity, or for which there is a greater 
likelihood of success. 

The success of our business depends upon our ability to identify, develop and commercialize product candidates 

generated through our gene therapy platform. Research programs to identify new product candidates require substantial 
technical, financial and human resources. Although VY-AADC is currently in clinical development and our other 
product candidates are in preclinical development, we may fail to identify other potential product candidates for clinical 
development for several reasons. For example, our research may be unsuccessful in identifying potential product 
candidates or our potential product candidates may be shown to have harmful side effects, may be commercially 
impracticable to manufacture or may have other characteristics that may make the products unmarketable or unlikely to 
receive marketing approval. 

Additionally, because we have limited resources, we may forego or delay pursuit of opportunities with certain 

programs or product candidates or for indications that later prove to have greater commercial potential. Our spending on 
current and future research and development programs may not yield any commercially viable products. If we do not 
accurately evaluate the commercial potential for a particular product candidate, we may relinquish valuable rights to that 
product candidate through strategic collaboration, licensing or other arrangements in cases in which it would have been 
more advantageous for us to retain sole development and commercialization rights to such product candidate. 
Alternatively, we may allocate internal resources to a product candidate in a therapeutic area in which it would have been 
more advantageous to enter into a partnering arrangement. 

If any of these events occur, we may be forced to abandon our development efforts with respect to a particular 
product candidate or fail to develop a potentially successful product candidate, which could harm our business, financial 
condition, results of operations and prospects. 

Our future success depends on our ability to retain key members of our management team, and to attract, retain and 
motivate qualified personnel. 

We are highly dependent on the management, technical, and scientific expertise of principal members of our 

management, scientific, and clinical teams including our former President and Chief Executive Officer, Steven M. Paul, 
M.D., who now serves as a senior advisor, director, and member of our Science and Technology Committee, and G. 
Andre Turenne, who joined us as President and Chief Executive Officer in July 2018. While we have entered into 
employment agreements or offer letters with each of our executive officers, any of them could leave our employment at 
any time, as all of our employees are “at will” employees. We currently do not have “key person” insurance on any of 
our employees. The loss of the services of one or more of our current employees might impede the achievement of our 
research, development and commercialization objectives. 

Recruiting and retaining other qualified employees, consultants and advisors for our business, including 

scientific and technical personnel is also critical to our success. There currently is a shortage of skilled individuals with 
substantial gene therapy experience, which is likely to continue. As a result, competition for skilled personnel, including 
in gene therapy research and vector manufacturing, is intense and the turnover rate can be high. We may not be able to 
attract and retain personnel on acceptable terms, if at all, given the competition among numerous pharmaceutical and 
biotechnology companies and academic institutions for individuals with similar skill sets. Our consultants and advisors 
may be employed by employers other than us and may have commitments under consulting or advisory contracts with 
other entities that may limit their availability to us. In addition, failure to succeed in preclinical or clinical trials or 
applications for marketing approval may make it more challenging to recruit and retain qualified personnel. The inability 
to recruit, or loss of services of certain executives, key employees, consultants or advisors, may impede the progress of 
our research, development and commercialization objectives and could harm our business, financial condition, results of 
operations and prospects. 

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If we are unable to manage expected growth in the scale and complexity of our operations, our performance may 
suffer. 

If we are successful in executing our business strategy, we will need to expand our managerial, operational, 
financial and other systems and resources to manage our operations, continue our research and development activities 
and, in the longer term, build a commercial infrastructure to support commercialization of any of our product candidates 
that are approved for sale. We can provide no assurances that we will have sufficient resources in the future to manage 
all of our planned programs. Future growth would impose significant added responsibilities on members of management, 
may lead to significant added costs, and may divert our management and business development resources. It is likely that 
our management, finance, development personnel, systems and facilities currently in place may not be adequate to 
support this future growth. Our need to effectively manage our operations, growth and product candidates requires that 
we continue to develop more robust business processes and improve our systems and procedures in each of these areas 
and to attract and retain sufficient numbers of talented employees. We may be unable to successfully implement these 
tasks on a larger scale and, accordingly, may not achieve our research, development and growth goals. 

Our employees, principal investigators, consultants and commercial partners may engage in misconduct or other 
improper activities, including non-compliance with regulatory standards and requirements and insider trading. 

We are exposed to the risk of fraud or other misconduct by our employees, principal investigators, consultants 

and commercial partners. Misconduct by these parties could include intentional failures to comply with FDA regulations 
or the regulations applicable in the European Union and other jurisdictions, provide accurate information to the FDA, the 
European Commission and other regulatory authorities, comply with healthcare fraud and abuse laws and regulations in 
the United States and abroad, report financial information or data accurately or disclose unauthorized activities to us. In 
particular, sales, marketing and business arrangements in the healthcare industry are subject to extensive laws and 
regulations intended to prevent fraud, misconduct, kickbacks, self-dealing and other abusive practices. These laws and 
regulations restrict or prohibit a wide range of pricing, discounting, marketing and promotion, sales commission, 
customer incentive programs and other business arrangements. Such misconduct also could involve the improper use of 
information obtained in the course of clinical trials or interactions with the FDA or other regulatory authorities, which 
could result in regulatory sanctions and cause serious harm to our reputation. We have adopted a code of conduct 
applicable to all of our employees, but it is not always possible to identify and deter employee misconduct, and the 
precautions we take to detect and prevent this activity may not be effective in controlling unknown or unmanaged risks 
or losses or in protecting us from government investigations or other actions or lawsuits stemming from a failure to 
comply with these laws or regulations. If any such actions are instituted against us, and we are not successful in 
defending ourselves or asserting our rights, those actions could have a significant impact on our business, financial 
condition, results of operations and prospects, including the imposition of significant fines or other sanctions. 

Current and future legislation may increase the difficulty and cost for us and any future collaborators to obtain 
marketing approval of and commercialize our product candidates and affect the prices we, or they, may obtain. 

In the United States and some foreign jurisdictions, there have been and continue to be a number of legislative 

and regulatory changes and proposed changes regarding the healthcare system that could, among other things, prevent or 
delay marketing approval of our product candidates, restrict or regulate post-approval activities and affect our ability, or 
the ability of any future collaborators, to profitably sell any products for which we, or they, obtain marketing approval. 
We expect that current laws, as well as other healthcare reform measures that may be adopted in the future, may result in 
more rigorous coverage criteria and in additional downward pressure on the price that we, or any future collaborators, 
may receive for any approved products.  

In March 2010, President Obama signed into law the Patient Protection and Affordable Care Act, as amended 

by the Health Care and Education Reconciliation Act, which we refer to, collectively, as the ACA. Among the provisions 
of the ACA of potential importance to our business and our product candidates are the following:   

 

an annual, non-deductible fee on any entity that manufactures or imports specified branded 
prescription drugs and biologic agents;   

93 

 

 

 

 

 

 

 

 

 

an increase in the statutory minimum rebates a manufacturer must pay under the Medicaid Drug 
Rebate Program;   

expansion of healthcare fraud and abuse laws, including the civil False Claims Act and the federal 
Anti-Kickback Statute, new government investigative powers and enhanced penalties for 
noncompliance;   

a new Medicare Part D coverage gap discount program, in which manufacturers must agree to offer 
50% point-of-sale discounts off negotiated prices;   

extension of manufacturers’ Medicaid rebate liability;   

expansion of eligibility criteria for Medicaid programs;   

expansion of the entities eligible for discounts under the Public Health Service pharmaceutical pricing 
program;   

new requirements to report certain financial arrangements with physicians and teaching hospitals;   

a new requirement to annually report drug samples that manufacturers and distributors provide to 
physicians; and   

a new Patient-Centered Outcomes Research Institute to oversee, identify priorities in, and conduct 
comparative clinical effectiveness research, along with funding for such research.  

In addition, other legislative changes have been proposed and adopted since the ACA was enacted. These 

changes include the Budget Control Act of 2011, which, among other things, led to aggregate reductions to Medicare 
payments to providers of up to 2% per fiscal year that started in 2013 and, due to subsequent legislative amendments to 
the statute, will stay in effect through 2025 unless additional congressional action is taken, and the American Taxpayer 
Relief Act of 2012, which, among other things, reduced Medicare payments to several types of providers and increased 
the statute of limitations period for the government to recover overpayments to providers from three to five years. These 
new laws may result in additional reductions in Medicare and other healthcare funding and otherwise affect the prices we 
may obtain for any of our product candidates for which we may obtain regulatory approval or the frequency with which 
any such product candidate is prescribed or used. Further, there have been several recent U.S. congressional inquiries 
and proposed state and federal legislation designed to, among other things, bring more transparency to drug pricing, 
review the relationship between pricing and manufacturer patient programs, reduce the costs of drugs under Medicare 
and reform government program reimbursement methodologies for drug products.  

We expect that these healthcare reforms, as well as other healthcare reform measures that may be adopted in the 

future, may result in additional reductions in Medicare and other healthcare funding, more rigorous coverage criteria, 
new payment methodologies and additional downward pressure on the price that we receive for any approved product 
and/or the level of reimbursement physicians receive for administering any approved product we might bring to market. 
Reductions in reimbursement levels may negatively impact the prices we receive or the frequency with which our 
products are prescribed or administered. Any reduction in reimbursement from Medicare or other government programs 
may result in a similar reduction in payments from private payors. 

With enactment of the Tax Cuts and Jobs Act of 2017, which was signed by the President on December 22, 

2017, Congress repealed the “individual mandate.” The repeal of this provision, which requires most Americans to carry 
a minimal level of health insurance, became effective January 1, 2019. According to the Congressional Budget Office, 
the repeal of the individual mandate will cause 13 million fewer Americans to be insured in 2027 and premiums in 
insurance markets may rise. Further, each chamber of the Congress has put forth multiple bills designed to repeal or 
repeal and replace portions of the ACA. Although none of these measures has been enacted by Congress to date, 
Congress may consider other legislation to repeal and replace elements of the ACA in the future.  

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The Trump administration has also taken executive actions to undermine or delay implementation of the ACA. 
Since January 2017, President Trump has signed two Executive Orders designed to delay the implementation of certain 
provisions of the ACA or otherwise circumvent some of the requirements for health insurance mandated by the ACA. 
One Executive Order directs federal agencies with authorities and responsibilities under the ACA to waive, defer, grant 
exemptions from, or delay the implementation of any provision of the ACA that would impose a fiscal or regulatory 
burden on states, individuals, healthcare providers, health insurers, or manufacturers of pharmaceuticals or medical 
devices. The second Executive Order terminates the cost-sharing subsidies that reimburse insurers under the ACA. 
Several state Attorneys General filed suit to stop the Trump administration from terminating the subsidies, but their 
request for a restraining order was denied by a federal judge in California on October 25, 2017. In addition, CMS has 
recently proposed regulations that would give states greater flexibility in setting benchmarks for insurers in the 
individual and small group marketplaces, which may have the effect of relaxing the essential health benefits required 
under the ACA for plans sold through such marketplaces. Further, on June 14, 2018, U.S. Court of Appeals for the 
Federal Circuit ruled that the federal government was not required to pay more than $12 billion in ACA risk corridor 
payments to third-party payors who argued were owed to them. The effects of this gap in reimbursement on third-party 
payors, the viability of the ACA marketplace, providers, and potentially our business, are not yet known. 

Further, there have been several recent U.S. congressional inquiries and proposed federal and proposed and 

enacted state legislation designed to, among other things, bring more transparency to drug pricing, review the 
relationship between pricing and manufacturer patient programs, reduce the costs of drugs under Medicare and reform 
government program reimbursement methodologies for drug products. For example, there have been several recent U.S. 
congressional inquiries and proposed federal and proposed and enacted state legislation designed to, among other things, 
bring more transparency to drug pricing, review the relationship between pricing and manufacturer patient programs, 
reduce the costs of drugs under Medicare and reform government program reimbursement methodologies for drug 
products. At the federal level, Congress and the Trump administration have each indicated that it will continue to seek 
new legislative and/or administrative measures to control drug costs. For example, on May 11, 2018, the Trump 
administration issued a plan to lower drug prices. Under this blueprint for action, the Trump administration indicated that 
the Department of Health and Human Services will: take steps to end the gaming of regulatory and patent processes by 
drug makers to unfairly protect monopolies; advance biosimilars and generics to boost price competition; evaluate the 
inclusion of prices in drug makers’ ads to enhance price competition; speed access to and lower the cost of new drugs by 
clarifying policies for sharing information between insurers and drug makers; avoid excessive pricing by relying more on 
value-based pricing by expanding outcome-based payments in Medicare and Medicaid; work to give Part D plan 
sponsors more negotiation power with drug makers; examine which Medicare Part B drugs could be negotiated for a 
lower price by Part D plans, and improving the design of the Part B Competitive Acquisition Program; update 
Medicare’s drug-pricing dashboard to increase transparency; prohibit Part D contracts that include “gag rules” that 
prevent pharmacists from informing patients when they could pay less out-of-pocket by not using insurance; and require 
that Part D plan members be provided with an annual statement of plan payments, out-of-pocket spending, and drug 
price increases. 

At the state level, individual states are increasingly aggressive in passing legislation and implementing 
regulations designed to control pharmaceutical and biological product pricing, including price or patient reimbursement 
constraints, discounts, restrictions on certain product access and marketing cost disclosure and transparency measures, 
and, in some cases, designed to encourage importation from other countries and bulk purchasing. In addition, regional 
health care authorities and individual hospitals are increasingly using bidding procedures to determine what 
pharmaceutical products and which suppliers will be included in their prescription drug and other health care programs. 
These measures could reduce the ultimate demand for our products, once approved, or put pressure on our product 
pricing. We expect that additional state and federal healthcare reform measures will be adopted in the future, any of 
which could limit the amounts that federal and state governments will pay for healthcare products and services, which 
could result in reduced demand for our product candidates or additional pricing pressures. 

Finally, legislative and regulatory proposals have also been made to expand post-approval requirements and 
restrict sales and promotional activities for pharmaceutical products. We cannot be sure whether additional legislative 
changes will be enacted, or whether the FDA regulations, guidance or interpretations will be changed, or what the impact 
of such changes on the marketing approvals of our product candidates, if any, may be. In addition, increased scrutiny by 
the U.S. Congress of the FDA’s approval process may significantly delay or prevent marketing approval, as well as 

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subject us and any future collaborators to more stringent product labeling and post-marketing testing and other 
requirements. 

Under the Cures Act and the Trump Administration’s regulatory reform initiatives, the FDA’s policies, regulations 
and guidance may be revised or revoked and that could prevent, limit or delay regulatory approval of our product 
candidates, which would impact our ability to generate revenue.  

In December 2016, the Cures Act was signed into law. The Cures Act, among other things, is intended to 

modernize the regulation of drugs and spur innovation, but its ultimate implementation is unclear. If we are slow or 
unable to adapt to changes in existing requirements or the adoption of new requirements or policies, or if we are not able 
to maintain regulatory compliance, we may lose any marketing approval that we may have obtained and we may not 
achieve or sustain profitability, which would adversely affect our business, prospects, financial condition and results of 
operations.  

We also cannot predict the likelihood, nature or extent of government regulation that may arise from future 

legislation or administrative or executive action, either in the United States or abroad. For example, certain policies of 
the Trump administration may impact our business and industry. Namely, the Trump administration has taken several 
executive actions, including the issuance of a number of Executive Orders, that could impose significant burdens on, or 
otherwise materially delay, the FDA’s ability to engage in routine regulatory and oversight activities such as the 
implementation of statutes through rulemaking, issuance of guidance, and review and approval of marketing 
applications. An under-staffed FDA could result in delays in the FDA’s responsiveness or in its ability to review 
submissions or applications, issue regulations or guidance, or implement or enforce regulatory requirements in a timely 
fashion or at all. Moreover, on January 30, 2017, President Trump issued an Executive Order, applicable to all executive 
agencies, including the FDA, which requires that for each notice of proposed rulemaking or final regulation to be issued 
in fiscal year 2017, the agency shall identify at least two existing regulations to be repealed, unless prohibited by law. 
These requirements are referred to as the “two-for-one” provisions. This Executive Order includes a budget neutrality 
provision that requires the total incremental cost of all new regulations in the 2017 fiscal year, including repealed 
regulations, to be no greater than zero, except in limited circumstances. For fiscal years 2018 and beyond, the Executive 
Order requires agencies to identify regulations to offset any incremental cost of a new regulation and approximate the 
total costs or savings associated with each new regulation or repealed regulation. In interim guidance issued by the 
Office of Information and Regulatory Affairs within OMB on February 2, 2017, the Trump administration indicates that 
the “two-for-one” provisions may apply not only to agency regulations, but also to significant agency guidance 
documents. In addition, on February 24, 2017, President Trump issued an executive order directing each affected agency 
to designate an agency official as a “Regulatory Reform Officer” and establish a “Regulatory Reform Task Force” to 
implement the two-for-one provisions and other previously issued executive orders relating to the review of federal 
regulations, however it is difficult to predict how these requirements will be implemented, and the extent to which they 
will impact the FDA’s ability to exercise its regulatory authority. If these executive actions impose constraints on the 
FDA’s ability to engage in oversight and implementation activities in the normal course, our business may be negatively 
impacted. 

We may be subject, directly or indirectly, to federal, state, and foreign healthcare laws and regulations, including 
fraud and abuse laws, false claims laws and health information privacy and security laws. If we are unable to comply, 
or have not fully complied, with such laws, we could face substantial penalties. 

If we obtain FDA approval for any of our product candidates and begin commercializing those products in the 
United States, our operations will be directly, or indirectly through our prescribers, customers and purchasers, subject to 
various federal and state laws and regulations, including, without limitation, the federal Anti-Kickback Statute, the 
federal civil and criminal false claims act, and the Physician Payments Sunshine Act and regulations. These laws will 
impact, among other things, our proposed sales, marketing and educational programs. In addition, we may be subject to 
data privacy laws by both the federal government and the states in which we conduct our business. Such laws that may 
constrain the business or financial arrangements and relationships through which we conduct our operations include, but 
are not limited to: 

 

the federal Anti-Kickback Statute, which prohibits, among other things, persons and entities from 
knowingly and willfully soliciting, receiving, offering or paying any remuneration (including any kickback, 
bribe or rebate), directly or indirectly, overtly or covertly, in cash or in kind, to induce or in return for either 

96 

the referral of an individual for, or the purchase, recommendation, leasing or furnishing of, an item or 
service reimbursable under a federal healthcare program, such as the Medicare and Medicaid programs. 
This statute has been interpreted to apply to arrangements between pharmaceutical manufacturers on the 
one hand, and prescribers, purchasers and formulary managers on the other. Further, the ACA amended the 
intent requirement of the federal Anti-Kickback Statute. A person or entity no longer needs to have actual 
knowledge of this statute or specific intent to violate it; 

the federal civil and criminal false claims laws and civil monetary penalty laws, including the civil False 
Claims Act, which prohibit, among other things, individuals or entities from knowingly presenting, or 
causing to be presented, claims for payment or approval from Medicare, Medicaid or other government 
payors that are false or fraudulent, or making a false statement to avoid, decrease, or conceal an obligation 
to pay money to the federal government. The ACA provided and recent government cases against 
pharmaceutical and medical device manufacturers support the view that federal Anti-Kickback Statute 
violations and certain marketing practices, including off-label promotion, may implicate the civil False 
Claims Act; 

the federal Health Insurance Portability and Accountability Act of 1996, or HIPAA, which created 
additional federal criminal statutes that prohibit a person from knowingly and willfully executing or 
attempting to execute a scheme or from making false or fraudulent statements to defraud any healthcare 
benefit program, regardless of the payor (e.g., public or private); 

 

 

  HIPAA, as amended by the Health Information Technology for Economic and Clinical Health Act, or 
HITECH, and its implementing regulations, and as amended again by the final HIPAA omnibus rule, 
Modifications to the HIPAA Privacy, Security, Enforcement, and Breach Notification Rules Under 
HITECH and the Genetic Information Nondiscrimination Act; Other Modifications to HIPAA, published in 
January 2013, which imposes certain requirements relating to the privacy, security and transmission of 
individually identifiable health information without appropriate authorization by entities subject to the rule, 
such as health plans, health care clearinghouses and health care providers; 

 

 

federal transparency laws, including the federal Physician Payments Sunshine Act, which is part of the 
ACA, that requires certain manufacturers of drugs, devices, biologics and medical supplies for which 
payment is available under Medicare, Medicaid, or the Children’s Health Insurance Program, with specific 
exceptions, to report annually to Center for Medicare & Medicaid Services, or CMS, information related to 
payments and other transfers of value provided to physicians and teaching hospitals, and ownership and 
investment interests held by physicians and their immediate family members, by the 90th day of each 
subsequent calendar year, and disclosure of such information is made by CMS on a publicly available 
website; and 

state and/or foreign law equivalents of each of the above federal laws, such as state anti-kickback and false 
claims laws that may apply to arrangements and claims involving health care items or services reimbursed 
by non-governmental third-party payors; state laws that require drug manufacturers to report information 
related to payments and other transfers of value to physicians and other healthcare providers or marketing 
expenditures; state laws that require pharmaceutical companies to comply with the pharmaceutical 
industry’s voluntary compliance guidelines and the relevant compliance guidance promulgated by the 
federal government; and state and foreign laws governing the privacy and security of health information in 
certain circumstances, many of which differ from each other in significant ways and may not have the same 
effect, thus complicating compliance efforts in certain circumstances, such as specific disease states. 

Because of the breadth of these laws and the narrowness of the statutory exceptions and safe harbors available, 

it is possible that some of our business activities could be subject to challenge under one or more of such laws. If our 
operations are found to be in violation of any of the laws described above or any other government regulations that apply 
to us, we may be subject to penalties, including civil and criminal penalties, damages, fines, exclusion from participation 
in government health care programs, such as Medicare and Medicaid, disgorgement, contractual damages, reputational 

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harm, diminished profits and future earnings, imprisonment and the curtailment or restructuring of our operations, any of 
which could adversely affect our ability to operate our business and our results of operations. 

The provision of benefits or advantages to physicians to induce or encourage the prescription, recommendation, 
endorsement, purchase, supply, order or use of medicinal products is prohibited in the European Union. The provision of 
benefits or advantages to physicians is also governed by the national anti-bribery laws of European Union Member 
States, such as the UK Bribery Act 2010. Infringement of these laws could result in substantial fines and imprisonment. 

Payments made to physicians in certain European Union Member States must be publicly disclosed. Moreover, 
agreements with physicians often must be the subject of prior notification and approval by the physician’s employer, his 
or her competent professional organization and/or the regulatory authorities of the individual European Union Member 
States. These requirements are provided in the national laws, industry codes or professional codes of conduct, applicable 
in the European Union Member States. Failure to comply with these requirements could result in reputational risk, public 
reprimands, administrative penalties, fines or imprisonment. 

The collection, use, disclosure, transfer, or other processing of personal data regarding individuals in the EU, 
including personal health data, is subject to the EU General Data Protection Regulation, or the GDPR, which became 
effective on May 25, 2018. The GDPR is wide-ranging in scope and imposes numerous requirements on companies that 
process personal data, including requirements relating to processing health and other sensitive data, obtaining consent of 
the individuals to whom the personal data relates, providing information to individuals regarding data processing 
activities, implementing safeguards to protect the security and confidentiality of personal data, providing notification of 
data breaches, and taking certain measures when engaging third-party processors. The GDPR also imposes strict rules on 
the transfer of personal data to countries outside the EU, including the U.S., and permits data protection authorities to 
impose large penalties for violations of the GDPR, including potential fines of up to €20 million or 4% of annual global 
revenues, whichever is greater. The GDPR also confers a private right of action on data subjects and consumer 
associations to lodge complaints with supervisory authorities, seek judicial remedies, and obtain compensation for 
damages resulting from violations of the GDPR. Compliance with the GDPR will be a rigorous and time-intensive 
process that may increase the cost of doing business or require companies to change their business practices to ensure 
full compliance. Compliance with the GDPR has been and will continue to be a rigorous and time-intensive process that 
has increased and will continue to increase our cost of doing business or require us to change our business practices, and 
despite those efforts, there is a risk that we or our collaborators may be subject to fines and penalties, litigation, and 
reputational harm in connection with any European activities, which could adversely affect our business, prospects, 
financial condition and results of operations. 

Product liability lawsuits against us could cause us to incur substantial liabilities and could limit commercialization 
of any product candidates that we may develop. 

We face an inherent risk of product liability exposure related to the testing of our product candidates in clinical 

trials and may face an even greater risk if we commercialize any products that we may develop. If we cannot 
successfully defend ourselves against claims that our product candidates caused injuries, we could incur substantial 
liabilities. Regardless of merit or eventual outcome, liability claims may result in: 

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decreased demand for any product candidates that we may develop; 

loss of revenue; 

substantial monetary awards to trial participants or patients; 

significant time and costs to defend the related litigation; 

  withdrawal of clinical trial participants; 

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the inability to commercialize any product candidates that we may develop; and 

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 

injury to our reputation and significant negative media attention. 

Although we maintain product liability insurance coverage in the amount of $5.0 million per occurrence and 
$5.0 million in the aggregate, and clinical testing liability insurance in the amount of $10.0 million per occurrence and 
$10.0 million in the aggregate, this insurance may not be adequate to cover all liabilities that we may incur. We 
anticipate that we will need to increase our insurance coverage each time we commence a clinical trial and if we 
successfully commercialize any product candidate. Insurance coverage is increasingly expensive. We may not be able to 
maintain insurance coverage at a reasonable cost or in an amount adequate to satisfy any liability that may arise. 

If we, our collaborators, or any third-party manufacturers engaged by us or our collaborators fail to comply with 
environmental, health and safety laws and regulations, we could become subject to fines or penalties or incur costs 
that could harm our business. 

We, our collaborators, and any third-party manufacturers we engage are subject to numerous environmental, 
health and safety laws and regulations, including those governing laboratory procedures and the generation, handling, 
use, storage, treatment, manufacture, transportation and disposal of, and exposure to, hazardous materials and wastes, as 
well as laws and regulations relating to occupational health and safety. Our operations involve the use of hazardous and 
flammable materials, including chemicals and biologic and radioactive materials. Our operations also produce hazardous 
waste products. We generally contract with third parties for the disposal of these materials and wastes. We cannot 
eliminate the risk of contamination or injury from these materials. In the event of contamination or injury resulting from 
our use of hazardous materials, we could be held liable for any resulting damages, and any liability could exceed our 
resources. We also could incur significant costs associated with civil or criminal fines and penalties. 

Although we maintain general liability insurance and workers’ compensation insurance for certain costs and 
expenses we may incur due to injuries to our employees resulting from the use of hazardous materials or other work-
related injuries, this insurance may not provide adequate coverage against potential liabilities. We do not maintain 
insurance for environmental liability or toxic tort claims that may be asserted against us in connection with our storage or 
disposal of biologic, hazardous or radioactive materials. 

In addition, we may incur substantial costs in order to comply with current or future environmental, health and 

safety laws and regulations, which have tended to become more stringent over time. These current or future laws and 
regulations may impair our research, development or production efforts. Failure to comply with these laws and 
regulations also may result in substantial fines, penalties or other sanctions or liabilities, which could harm our business, 
financial condition, results of operations and prospects. 

Further, with respect to the operations of any current or future collaborators or third-party contract 
manufacturers, it is possible that if they fail to operate in compliance with applicable environmental, health and safety 
laws and regulations or properly dispose of wastes associated with our products, we could be held liable for any resulting 
damages, suffer reputational harm or experience a disruption in the manufacture and supply of our product candidates or 
products. 

Unfavorable global economic conditions could adversely affect our business, financial condition or results of 
operations. 

Our results of operations could be adversely affected by general conditions in the global economy and in the 

global financial markets. The most recent global financial crisis caused extreme volatility and disruptions in the capital 
and credit markets. A severe or prolonged economic downturn, such as the most recent global financial crisis, could 
result in a variety of risks to our business, including weakened demand for our product candidates and our ability to raise 
additional capital when needed on acceptable terms, if at all. This is particularly true in the European Union, which is 
recovering from a severe economic crisis. A weak or declining economy could strain our suppliers, possibly resulting in 
supply disruption, or cause delays in payments for our services by third-party payors or our collaborators. Any of the 
foregoing could harm our business and we cannot anticipate all of the ways in which the current economic climate and 
financial market conditions could adversely impact our business. 

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Our internal computer systems, or those of our collaborators or other contractors or consultants, may fail or suffer 
security breaches, which could result in a material disruption of our product development programs. 

Our internal computer systems and those of our current and any future collaborators and other contractors or 

consultants are vulnerable to damage from cyber-attacks, computer viruses, unauthorized access, sabotage, natural 
disasters, terrorism, war and telecommunication and electrical failures. While we have not experienced any such material 
system failure, accident or security breach to date, if such an event were to occur and cause interruptions in our 
operations or the operations of those third parties with which we contract, it could result in a material disruption of our 
development programs and our business operations, whether due to a loss of our trade secrets or other proprietary 
information or other similar disruptions, and could require a substantial expenditure of resources to remedy. For 
example, the loss of clinical trial data from completed or ongoing clinical trials could result in delays in our regulatory 
approval efforts and significantly increase our costs to recover or reproduce the data. We could also be subject to risks 
caused by misappropriation, misuse, leakage, falsification or intentional or accidental release or loss of information 
maintained in our information systems and networks, including personal information of our employees. Outside parties 
may attempt to penetrate our systems or those of the third parties with which we contract or to fraudulently induce our 
employees or employees of such third parties to disclose sensitive information to gain access to our data. 

The number and complexity of these threats continue to increase over time. Although we develop and maintain 

systems and controls designed to prevent these events from occurring, and we have a process to identify and mitigate 
threats, the development and maintenance of these systems, controls and processes is costly and requires ongoing 
monitoring and updating as technologies change and efforts to overcome security measures become more sophisticated. 
Despite our efforts, the possibility of these events occurring cannot be eliminated entirely. Although we maintain cyber 
risk insurance for certain costs we may incur due to a cyber-related event, this insurance may not provide adequate 
coverage against potential liabilities. To the extent that any disruption or security breach were to result in a loss of, or 
damage to, our data or applications, or inappropriate disclosure of confidential or proprietary information, we could incur 
liability, our competitive position and the market perception of the effectiveness of our security measures could be 
harmed, our credibility could be damaged, and the further development and commercialization of our product candidates 
could be delayed.  

The Tax Cuts and Jobs Act of 2017 could adversely affect our business and financial condition.  

On December 22, 2017, President Trump signed into law the Tax Cuts and Jobs Act of 2017, or TCJA, which 

significantly revised the Internal Revenue Code of 1986, as amended. The TCJA, among other things, contains 
significant changes to corporate taxation, including reduction of the corporate tax rate from a top marginal rate of 35% to 
a flat rate of 21%, limitation of the tax deduction for net interest expense to 30% of adjusted earnings (except for certain 
small businesses), limitation of the deduction for net operating losses to 80% of current year taxable income and 
elimination of net operating loss carrybacks, in each case, for losses arising in taxable years beginning after December 
31, 2017 (though any such net operating losses may be carried forward indefinitely), one-time taxation of offshore 
earnings at reduced rates regardless of whether they are repatriated, elimination of U.S. tax on foreign earnings (subject 
to certain important exceptions), immediate deductions for certain new investments instead of deductions for 
depreciation expense over time, and modifying or repealing many business deductions and credits. Notwithstanding the 
reduction in the corporate income tax rate, the overall impact of the TCJA is uncertain and our business and financial 
condition could be adversely affected. In addition, it is uncertain how various states will respond to the TCJA. The 
impact of this tax reform on holders of our common stock is also uncertain and could be adverse. We urge our 
stockholders to consult with their legal and tax advisors with respect to this legislation and the potential tax 
consequences of investing in or holding our common stock.  

We might not be able to utilize a significant portion of our net operating loss carryforwards.  

As of December 31, 2018, we had both federal and state net operating loss carryforwards of $162.9 million and 

$163.8 million, respectively, which expire beginning in 2033. These net operating loss carryforwards could expire 
unused and be unavailable to offset our future income tax liabilities. Under the newly enacted federal income tax law, 
federal net operating losses incurred in 2018 and in future years may be carried forward indefinitely, but the deductibility 
of such federal net operating losses is limited. It is uncertain how various states will respond to the newly enacted federal 

100 

tax law. If our ability to use our historical net operating loss carryforwards is materially limited, it would harm our future 
operating results by effectively increasing our future tax obligations. 

Risks Related to the Commercialization of Our Product Candidates 

The affected populations for our product candidates may be smaller than we or third parties currently project, which 
may affect the addressable markets for our product candidates. 

Our projections of the number of people who have the diseases we are seeking to treat, as well as the subset of 

people with these diseases who have the potential to benefit from treatment with our product candidates, are estimates 
based on our knowledge and understanding of these diseases. The total addressable market opportunity for our product 
candidates will ultimately depend upon a number of factors including the diagnosis and treatment criteria included in the 
final label, if approved for sale in specified indications, acceptance by the medical community, patient access and 
product pricing and reimbursement. Prevalence estimates are frequently based on information and assumptions that are 
not exact and may not be appropriate, and the methodology is forward-looking and speculative. The process we have 
used in developing an estimated prevalence range for the indications we are targeting has involved collating limited data 
from multiple sources. While we believe these sources are reliable, we have not independently verified the data. 
Accordingly, the prevalence estimates included in our periodic reports and other reports filed with or furnished to the 
SEC should be viewed with caution. Further, the data and statistical information used in such reports, including estimates 
derived from them, may differ from information and estimates made by our competitors or from current or future studies 
conducted by independent sources. 

The use of such data involves risks and uncertainties, and such data is subject to change based on various 

factors. Our estimates may prove to be incorrect and new studies may change the estimated incidence or prevalence of 
the diseases we seek to address. The number of patients with the diseases we are targeting in the United States, the 
European Union and elsewhere may turn out to be lower than expected or may not be otherwise amenable to treatment 
with our products, or new patients may become increasingly difficult to identify or access, all of which would harm our 
results of operations and our business. 

If we are unable to establish sales, medical affairs and marketing capabilities or enter into agreements with third 
parties to market and sell our product candidates, we may be unable to generate any product revenue. 

To successfully commercialize any products that may result from our clinical development programs, we will 
need to further develop these capabilities, either on our own or with others. The establishment and development of our 
own commercial team or the establishment of a contract sales force to market any products we may develop will be 
expensive and time-consuming and could delay any product launch. Moreover, we cannot be certain that we will be able 
to successfully develop this capability.  

Under our collaboration agreement with Sanofi Genzyme, Sanofi Genzyme has an exclusive option to license, 

develop and commercialize ex-U.S. rights to our Friedreich’s ataxia program, a future program to be designated by 
Sanofi Genzyme and our Huntington’s disease program. Additionally, we have granted Sanofi Genzyme an incremental 
option to co-commercialize our Huntington’s disease program in the United States and to acquire worldwide rights to our 
spinal muscular atrophy program. If Sanofi Genzyme exercises any of these options, we would be eligible to receive 
specified option fees. In addition, we would be eligible to receive specified milestone payments and royalties for any 
product developed in such programs.  

Under our collaboration and option agreement with AbbVie related to defective or excess aggregation of tau 

protein in the human brain, AbbVie has options to advance certain compounds into further development and to obtain an 
exclusive, worldwide license, with the right to sublicense, under certain of our intellectual property rights to develop and 
commercialize such compounds and corresponding products for all human diagnostic, prophylactic and therapeutic uses. 
If AbbVie exercises any of these options, we would be eligible to receive specified option fees. In addition, we would be 
eligible to receive specified development and regulatory milestone payments and tiered royalties on the global 
commercial net sales products developed under our tau collaboration.  

101 

Under our collaboration agreement with Neurocrine, Neurocrine will fund the clinical development through the 
readout of the RESTORE-1 Phase 2 clinical trial for VY-AADC. After the data readout of the RESTORE-1 Phase 2 trial, 
we have the option to either: (1) co-commercialize VY-AADC with Neurocrine in the U.S. under a 50/50 cost- and 
profit-sharing arrangement and receive milestones and royalties based on ex-U.S. sales, or (2) grant Neurocrine full 
global commercial rights in exchange for milestone payments and royalties based on global sales. Under the terms of the 
agreement for the FA Program, Neurocrine will fund the development through the Phase 1 clinical trial of VY-FXN01. 
After the data readout of the Phase 1 trial, we have the option to either: (1) co-commercialize VY-FXN01 with 
Neurocrine in the U.S. under a 60/40 cost- and profit-sharing arrangement, or (2) grant Neurocrine full worldwide 
commercial rights in exchange for milestone payments and royalties based on global sales, subject to Sanofi Genzyme’s 
option to commercialize the FA Program in countries outside the United States. Under the terms of the agreement for the 
two Discovery Programs, Neurocrine will fund the development of those programs and we have the right to earn 
milestone payments and royalties based on global sales. Under the terms of the agreement for the two Discovery 
Programs, Neurocrine will fund the development of the programs and we have the right to earn milestone payments and 
royalties based on global sales. 

Under our collaboration and option agreement with AbbVie directed against pathological species of alpha-

synuclein for the potential treatment of Parkinson’s disease and other synucleinopathies, AbbVie has options to advance 
certain compounds into further development and to obtain an exclusive, worldwide license, with the right to sublicense, 
under certain of our intellectual property rights to develop and commercialize such compounds and corresponding 
products for all human diagnostic, prophylactic and therapeutic uses. If AbbVie exercises any of these options, we would 
be eligible to receive specified option fees. In addition, we would be eligible to receive specified regulatory and 
commercial milestone payments and tiered royalties on the global commercial net sales products developed under our 
alpha-synuclein collaboration. 

In the future, we may seek to enter into collaborations regarding other of our product candidates with other 

entities to utilize their established marketing and distribution capabilities, but we may be unable to enter into such 
agreements on favorable terms, if at all. If any current or future collaborators do not commit sufficient resources to 
commercialize our products, or we are unable to develop the necessary capabilities on our own, we will be unable to 
generate sufficient product revenue to sustain our business. We compete with many companies that currently have 
extensive, experienced and well-funded medical affairs, marketing and sales operations to recruit, hire, train and retain 
marketing and sales personnel. We also face competition in our search for third parties to assist us with the sales and 
marketing efforts of our product candidates. We might face unforeseen costs and expenses associated with creating an 
independent sales and marketing organization. Our sales personnel might also face difficulties obtaining access to 
physicians or being able to persuade adequate numbers of physicians to use or prescribe our products or selling our 
products if we lack complementary products, which could disadvantage us compared to companies with more extensive 
product lines. Without an internal team or the support of a third party to perform marketing and sales functions, we may 
be unable to compete successfully against these more established companies. 

Our efforts to educate the medical community and third-party payors on the benefits of our product candidates 

may require significant resources and may never be successful. Such efforts may require more resources than are 
typically required due to the complexity and uniqueness of our potential products. If any of our product candidates is 
approved but fails to achieve market acceptance among physicians, patients, or third-party payors, we will not be able to 
generate significant revenues from such product, which could harm our business, financial condition, results of 
operations and prospects.  

The insurance coverage and reimbursement status of newly-approved products is uncertain. Failure to obtain or 
maintain adequate coverage and reimbursement for our product candidates, if approved, could limit our ability to 
market those products and decrease our ability to generate product revenue. 

We expect the cost of a single administration of gene therapy products, such as those we are developing, to be 
substantial, when and if they receive regulatory approval. We expect that coverage and reimbursement by government 
and private payors will be essential for most patients to be able to afford these treatments. Accordingly, sales of our 
product candidates will depend substantially, both domestically and abroad, on the extent to which the costs of our 
product candidates will be paid by health maintenance, managed care, pharmacy benefit and similar healthcare 
management organizations, or will be reimbursed by government authorities, private health coverage insurers and other 

102 

third-party payors. Coverage and reimbursement by a third-party payor may depend upon several factors, including the 
third-party payor’s determination that use of a product is: 

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 

 

 

a covered benefit under its health plan; 

safe, effective and medically necessary; 

appropriate for the specific patient and the indication; 

convenient and easy-to-administer compared to alternative treatments; 

cost-effective compared to alternative treatments; and 

neither experimental nor investigational. 

No uniform policy requirement for coverage and reimbursement for biopharmaceutical products exists among 

third-party payors. Therefore, coverage and reimbursement for such products can differ significantly from payor to 
payor. As a result, obtaining coverage and reimbursement for a product from third-party payors is a time-consuming and 
costly process that could require us to provide to each different payor supporting scientific, clinical and 
cost-effectiveness data. We may not be able to provide data sufficient to gain acceptance with respect to coverage and 
reimbursement. If coverage and reimbursement are not available, or are available only at limited levels, we may not be 
able to successfully commercialize our product candidates. Even if coverage is provided, the approved reimbursement 
amount may not be adequate to realize a sufficient return on our investment including our research, development, 
manufacture, sales, and distribution expenses. Interim reimbursement levels for new drugs, if applicable, may also not be 
sufficient to cover our costs and may not be made permanent. Reimbursement rates may vary according to the use of the 
drug and the clinical setting in which it is used, may be based on reimbursement levels already set for lower cost drugs 
and may be incorporated into existing payments for other services. Assuming we obtain coverage for a given product by 
a third-party payor, the resulting reimbursement payment rates may not be adequate or may require co-payments that 
patients find unacceptably high. Patients who are prescribed medications for the treatment of their conditions, and their 
prescribing physicians, generally rely on third-party payors to reimburse all or part of the costs associated with their 
prescription drugs. Patients are unlikely to use our products unless coverage is provided and reimbursement is adequate 
to cover all or a significant portion of the cost of our products. Therefore, coverage and adequate reimbursement are 
critical to new product acceptance. Additionally, there may be significant delays in obtaining coverage and 
reimbursement for newly approved drugs and biologics, and coverage may be more limited than the purposes for which 
the drug is approved by the FDA or comparable foreign regulatory authorities. 

There is significant uncertainty related to third-party coverage and reimbursement of newly approved products. 
In the United States, third-party payors, including government payors such as the Medicare and Medicaid programs, play 
an important role in determining the extent to which new drugs and biologics will be covered and reimbursed. The 
Medicare and Medicaid programs increasingly are used as models for how private payors and government payors 
develop their coverage and reimbursement policies. A primary trend in the U.S. healthcare industry and elsewhere is cost 
containment. Government authorities and third-party payors have attempted to control costs by limiting coverage and the 
amount of reimbursement for particular medications. Increasingly, third-party payors are requiring that drug companies 
provide them with predetermined discounts from list prices and are challenging the prices charged for medical products. 

The CMS is responsible for determining whether a product should be approved for coverage and reimbursement 
under the Medicare program. It is difficult to predict what CMS will decide with respect to coverage and reimbursement 
for novel products such as ours, as there is no body of established practices and precedents for these types of products. 
Currently, no gene therapy product has been approved for coverage and reimbursement by the CMS. Moreover, 
reimbursement agencies in the European Union may be more conservative than CMS. For example, several cancer drugs 
have been approved for reimbursement in the United States and have not been approved for reimbursement in certain 
European Union Member States. It is difficult to predict what third-party payors will decide with respect to the coverage 
and reimbursement for our product candidates, especially given that the cost of our product candidates is likely to be 
very high and pricing of such products is highly uncertain. 

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Outside the United States, international operations generally are subject to extensive government price controls 

and other market regulations, and increasing emphasis on cost-containment initiatives in the European Union, Canada 
and other countries may put pricing pressure on us. In many countries, the prices of medical products are subject to 
varying price control mechanisms as part of national health systems. In general, the prices of medicines under such 
systems are substantially lower than in the United States. Other countries allow companies to fix their own prices for 
medical products, but monitor and control company profits. Additional foreign price controls or other changes in pricing 
regulation could restrict the amount that we are able to charge for our product candidates. Accordingly, in markets 
outside the United States, the reimbursement for our products may be reduced compared with the United States and may 
be insufficient to generate commercially reasonable product revenues. 

Moreover, increasing efforts by government and third-party payors in the United States and abroad to cap or 
reduce healthcare costs may cause such organizations to limit both coverage and the level of reimbursement for new 
products approved and, as a result, they may not cover or provide adequate payment for our product candidates. Payors 
increasingly are considering new metrics as the basis for reimbursement rates, such as average sales price, or ASP, 
average manufacturer price, or AMP, and Actual Acquisition Cost. The existing data for reimbursement based on some 
of these metrics is relatively limited, although certain states have begun to survey acquisition cost data for the purpose of 
setting Medicaid reimbursement rates, and CMS has begun making pharmacy National Average Drug Acquisition Cost 
and National Average Retail Price data publicly available on at least a monthly basis. The regulations that govern 
marketing approvals, pricing, coverage and reimbursement for new drug and device products vary widely from country 
to country. Current and future legislation may significantly change the approval requirements in ways that could involve 
additional costs and cause delays in obtaining approvals. Some countries require approval of the sale price of a drug 
before it can be marketed. In many countries, the pricing review period begins after marketing or product licensing 
approval is granted. In some foreign markets, prescription pharmaceutical pricing remains subject to continuing 
governmental control even after initial approval is granted. As a result, we might obtain marketing approval for a product 
in a particular country, but then be subject to price regulations that delay our commercial launch of the product, possibly 
for lengthy time periods, and negatively impact the revenues we are able to generate from the sale of the product in that 
country. To obtain reimbursement or pricing approval in some countries, we may be required to conduct a clinical trial 
that compares the cost-effectiveness of our product candidate to other available therapies. Adverse pricing limitations 
may hinder our ability to recoup our investment in one or more product candidates, even if our product candidates obtain 
marketing approval. 

Therefore, it is difficult to project the impact of these evolving reimbursement metrics on the willingness of 

payors to cover candidate products that we or our partners are able to commercialize. We expect to experience pricing 
pressures in connection with the sale of any of our product candidates due to the trend toward managed healthcare, the 
increasing influence of health maintenance organizations and additional legislative changes. The downward pressure on 
healthcare costs in general, particularly prescription drugs and surgical procedures and other treatments, has become 
intense. As a result, increasingly high barriers are being erected to the entry of new products such as ours. 

The commercial success of any of our product candidates will depend upon its degree of market acceptance by 
physicians, patients, third-party payors and others in the medical community. 

Ethical, social and legal concerns about gene therapy could result in additional regulations restricting or 

prohibiting our products. Even with the requisite approvals from the FDA in the United States, EMA in the European 
Union and other regulatory authorities internationally, the commercial success of our product candidates will depend, in 
part, on the acceptance of physicians, patients and health care payors of gene therapy products in general, and our 
product candidates in particular, as medically necessary, cost-effective and safe. Any product that we commercialize may 
not gain acceptance by physicians, patients, health care payors and others in the medical community. If these products do 
not achieve an adequate level of acceptance, we may not generate significant product revenue and may not become 
profitable. The degree of market acceptance of gene therapy products and, in particular, our product candidates, if 
approved for commercial sale, will depend on several factors, including: 

 

 

the efficacy and safety of such product candidates as demonstrated in clinical trials; 

the potential and perceived advantages of product candidates over alternative treatments; 

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 

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 

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 

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the cost of treatment relative to alternative treatments; 

the clinical indications for which the product candidate is approved by the FDA or the European 
Commission, or other regulatory authorities; 

patient awareness of, and willingness to seek, genotyping; 

the willingness of physicians to prescribe new therapies; 

the willingness of physicians to undergo specialized training with respect to administration of our product 
candidates; 

the willingness of the target patient population to try new therapies; 

the prevalence and severity of any side effects; 

product labeling or product insert requirements of the FDA, EMA or other regulatory authorities, including 
any limitations or warnings contained in a product’s approved labeling or restrictions on the use of our 
products together with other medications; 

relative convenience and ease of administration; 

the strength of marketing and distribution support; 

the timing of market introduction of competitive products; 

publicity concerning our products or competing products and treatments; and 

sufficient third-party payor coverage and reimbursement. 

Even if a potential product displays a favorable efficacy and safety profile in preclinical studies and clinical 

trials, market acceptance of the product will not be fully known until after it is launched. 

Our gene therapy approach utilizes vectors derived from viruses, which may be perceived as unsafe or may result in 
unforeseen adverse events. Negative public opinion and increased regulatory scrutiny of gene therapy may damage 
public perception of the safety of our product candidates and adversely affect our ability to conduct our business or 
obtain regulatory approvals for our product candidates. 

Gene therapy remains a novel technology, with few gene therapy products approved to date in the United States 

and the European Union. Public perception may be influenced by claims that gene therapy is unsafe, and gene therapy 
may not gain the acceptance of the public or the medical community. In particular, our success will depend upon 
physicians who specialize in the treatment of genetic diseases targeted by our product candidates, prescribing treatments 
that involve the use of our product candidates in lieu of, or in addition to, existing treatments with which they are 
familiar and for which greater clinical data may be available. More restrictive government regulations or negative public 
opinion would have an adverse effect on our business, financial condition, results of operations and prospects and may 
delay or impair the development and commercialization of our product candidates or demand for any products we may 
develop. For example, earlier gene therapy trials led to several well-publicized adverse events, including cases of 
leukemia and death seen in other trials using non-AAV gene therapy vectors. Serious adverse events in our clinical trials, 
or other clinical trials involving gene therapy products or our competitors’ products, even if not ultimately attributable to 
the relevant product candidates, and the resulting publicity, could result in increased government regulation, unfavorable 
public perception, potential regulatory delays in the testing or approval of our product candidates, stricter labeling 
requirements for those product candidates that are approved and a decrease in demand for any such product candidates. 

105 

If we obtain approval to commercialize our product candidates outside of the United States, in particular in the 
European Union, a variety of risks associated with international operations could harm our business. 

We expect that we will be subject to additional risks in commercializing our product candidates outside the 

United States, including: 

 

 

 

 

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different regulatory requirements for approval of drugs and biologics in foreign countries; 

reduced or loss of protection under our intellectual property rights; 

unexpected changes in tariffs, trade barriers and regulatory requirements; 

economic weakness, including inflation, or political instability in particular foreign economies and markets; 

compliance with tax, employment, immigration and labor laws for employees living or traveling abroad; 

foreign currency fluctuations, which could result in increased operating expenses and reduced revenues, 
and other obligations incident to doing business in another country; 

  workforce uncertainty in countries where labor unrest is more common than in the United States; 

 

 

shortages resulting from any events affecting raw material supply or manufacturing capabilities abroad; 

business interruptions resulting from geopolitical actions, including war and terrorism or natural disasters 
including earthquakes, typhoons, floods and fires, or from economic or political instability; and 

 

greater difficulty with enforcing our contracts in jurisdictions outside of the United States. 

We must dedicate additional resources to comply with numerous laws and regulations in each jurisdiction in 

which we plan to operate. The Foreign Corrupt Practices Act, or FCPA, prohibits any U.S. individual or business from 
paying, offering, authorizing payment or offering of anything of value, directly or indirectly, to any foreign official, 
political party or candidate for the purpose of influencing any act or decision of the foreign entity in order to assist the 
individual or business in obtaining or retaining business. The FCPA also obligates companies whose securities are listed 
in the United States to comply with certain accounting provisions requiring the company to maintain books and records 
that accurately and fairly reflect all transactions of the corporation, including international subsidiaries, and to devise and 
maintain an adequate system of internal accounting controls for international operations. 

Compliance with the FCPA is expensive and difficult, particularly in countries in which corruption is a 
recognized problem. In many foreign countries, it is common for others to engage in business practices that are 
prohibited by U.S. laws and regulations applicable to us, including the FCPA. In addition, the FCPA presents particular 
challenges in the pharmaceutical industry because, in many countries, hospitals are operated by the government, and 
doctors and other hospital employees are considered foreign officials. Certain payments to hospitals in connection with 
clinical trials and other work have been deemed to be improper payments to government officials and have led to FCPA 
enforcement actions.  

Various laws, regulations and executive orders also restrict the use and dissemination outside of the 
United States, or the sharing with certain non-U.S. nationals, of information classified for national security purposes, as 
well as certain products and technical data relating to those products. If we expand our presence outside of the United 
States, it will require us to dedicate additional resources to comply with these laws, and these laws may preclude us from 
developing, manufacturing, or selling certain products and product candidates outside of the United States, which could 
limit our growth potential and increase our development costs.  

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The failure to comply with laws governing international business practices may result in substantial civil and 

criminal penalties and suspension or debarment from government contracting. The Securities and Exchange Commission 
also may suspend or bar issuers from trading securities on U.S. exchanges for violations of the FCPA’s accounting 
provisions. Although we expect to implement policies and procedures designed to comply with these laws and policies, 
there can be no assurance that our employees, contractors and agents will comply with these laws and policies. If we are 
unable to successfully manage the challenges of international expansion and operations, our business and operating 
results could be harmed. 

Risks Related to Our Intellectual Property 

Our rights to develop and commercialize our product candidates are subject to, in part, the terms and conditions of 
licenses granted to us by others. 

We are reliant upon licenses to certain patent rights and proprietary technology from third parties that are 

important or necessary to the development of our technology and products, including technology related to our 
manufacturing process and our product candidates. These and other licenses may not provide exclusive rights to use such 
intellectual property and technology in all relevant fields of use and in all territories in which we may wish to develop or 
commercialize our technology and products in the future. As a result, we may not be able to prevent competitors from 
developing and commercializing competitive products in territories included in all of our licenses. These licenses may 
also require us to grant back certain rights to licensors and to pay certain amounts relating to sublicensing patent and 
other rights under the agreement. 

In some circumstances, particularly in-licenses with academic institutions, we may not have the right to control 

the preparation, filing and prosecution of patent applications, or to maintain, enforce or defend the patents, covering 
technology that we license from third parties. Therefore, we cannot be certain that these patents and applications will be 
prosecuted, maintained and enforced in a manner consistent with the best interests of our business. If our licensors fail to 
maintain such patents, or lose rights to those patents or patent applications, the rights we have licensed may be reduced 
or eliminated and our right to develop and commercialize any of our products that are the subject of such licensed rights 
could be adversely affected. In certain circumstances, we have or may license technology from third parties on a non-
exclusive basis. In such instances, other licensees may have the right to enforce our licensed patents in their respective 
fields, without our oversight or control. Those other licensees may choose to enforce our licensed patents in a way that 
harms our interest, for example, by advocating for claim interpretations or agreeing on invalidity positions that conflict 
with our positions or our interest. In addition to the foregoing, the risks associated with patent rights that we license from 
third parties will also apply to patent rights we own or may own in the future. 

Further, in many of our license agreements we are responsible for bringing any actions against any third party 

for infringing on the patents we have licensed. Certain of our license agreements also require us to meet development 
thresholds to maintain the license, including establishing a set timeline for developing and commercializing products and 
minimum yearly diligence obligations in developing and commercializing the product. Certain of our license agreements 
contain “no challenge” clauses which preclude and prevent us from taking any action to limit or narrow the intellectual 
property of a licensor. In some cases these limitations extend to any intellectual property of our licensor and not just that 
which is licensed to us. Such constraints may limit our ability to develop or commercialize products or to expand such 
efforts beyond the scope of any license. Disputes may arise regarding intellectual property subject to a licensing 
agreement, including: 

 

 

the scope of rights granted under the license agreement and other interpretation-related issues; 

the extent to which our technology and processes infringe on intellectual property of the licensor that is not 
subject to the licensing agreement; 

 

the sublicensing of patent and other rights under our collaborative development relationships; 

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 

 

our diligence obligations under the license agreement and what activities satisfy those diligence 
obligations; 

the inventorship or ownership of inventions and know-how resulting from the creation or use of intellectual 
property by our licensors and us and our partners; and 

 

the priority of invention of patented technology. 

If disputes over intellectual property that we have licensed prevent or impair our ability to maintain our current 
licensing arrangements on acceptable terms, we may be unable to successfully develop and commercialize the affected 
product candidates. 

If we fail to comply with our obligations under these license agreements, or we are subject to a bankruptcy, the 

licensor may have the right to terminate the license, in which event we would not be able to develop, manufacture, or 
market products covered by the license or may face other penalties under the agreements. Termination of these 
agreements or reduction or elimination of our rights under these agreements may result in our having to negotiate new or 
reinstated agreements with less favorable terms or cause us to lose our rights under these agreements, including our 
rights to important intellectual property or technology. 

Furthermore, the research resulting in certain of our licensed patent rights and technology was funded by the 

U.S. government. As a result, the government may have certain rights, or march-in rights, to such patent rights and 
technology. When new technologies are developed with U.S. government funding, the U.S. government generally 
obtains certain rights in any resulting patents, including a non-exclusive, royalty-free license authorizing the U.S. 
government, or a third party on its behalf, to use the invention for non-commercial purposes. These rights may permit the 
government to disclose our confidential information to third parties and to exercise march-in rights to use or allow third 
parties to use our licensed technology. The U.S. government can exercise its march-in rights if it determines that action is 
necessary because we fail to achieve practical application of the government-funded technology, because action is 
necessary to alleviate health or safety needs, to meet requirements of federal regulations or to give preference to U.S. 
industry. In addition, our rights in such inventions may be subject to certain requirements to manufacture products 
embodying such inventions in the United States. Any exercise by the government, or a third party on its behalf, of such 
rights could harm our competitive position, business, financial condition, results of operations and prospects. 

If we are unable to obtain and maintain patent protection for our products and technology, or if the scope of the 
patent protection obtained is not of sufficient breadth, our competitors could develop and commercialize products and 
technology similar or identical to ours, and our ability to successfully commercialize our products and technology 
may be adversely affected. 

Our success depends, in large part, on our and our licensors’ ability to obtain and maintain patent protection in 
the United States and other countries with respect to our product candidates and manufacturing technology. We and our 
licensors have sought, and we intend to seek in the future, to protect our proprietary position by filing patent applications 
in the United States and abroad related to many of our technologies and product candidates that are important to our 
business. 

The patent prosecution process is expensive, time-consuming and complex, and we may not have and may not 
in the future be able to file, prosecute, maintain, enforce, defend or license all necessary or desirable patent applications 
at a reasonable cost or in a timely manner. For example, in some cases, the work of certain academic researchers in the 
gene therapy field has entered the public domain, which may compromise our ability to obtain patent protection for 
certain inventions related to or building upon such prior work. Consequently, we may not be able to obtain any such 
patents to prevent others from using our technology for, and developing and marketing competing products to treat, these 
indications. It is also possible that we will fail to identify patentable aspects of our research and development output 
before it is too late to obtain patent protection. In some cases, we may be able to obtain patent protection, but such 
protections may expire before we commercialize the product protected by those rights, leaving us no meaningful 
protection for our products. In other cases, where our intellectual property is being managed by a third-party 
collaborator, licensee or partner, that third party may fail to act diligently in prosecuting, maintaining, defending or 

108 

enforcing our patents. Such conduct may result in the failure to maintain or obtain protections, loss of rights, loss of 
patent term or, in cases where a third party has acted negligently or inequitably, patents being found unenforceable. 

The patent position of biotechnology and pharmaceutical companies generally is highly uncertain, involves 

complex legal and factual questions and has, in recent years, been the subject of much litigation. As a result, the 
issuance, scope, validity, enforceability and commercial value of our and our licensors’ patent rights are highly 
uncertain. Our pending and future patent applications may not result in patents being issued which protect our 
technology or product candidates or which effectively prevent others from commercializing competitive technologies 
and product candidates. In particular, during prosecution of any patent application, the issuance of any patents based on 
the application may depend upon our ability to generate additional preclinical or clinical data that support the 
patentability of our proposed claims. We may not be able to generate sufficient additional data on a timely basis, or at all. 
Changes in either the patent laws or interpretation of the patent laws in the United States and other countries may 
diminish the value, narrow the scope, or eliminate the enforceability of our and our licensors’ patent protection. 

We may not be aware of all third-party intellectual property rights potentially relating to our product candidates. 

Publications of discoveries in the scientific literature often lag behind the actual discoveries, and patent applications in 
the United States and other jurisdictions are typically not published until 18 months after filing or, in some cases, only 
upon issuance or not at all. Therefore, we cannot be certain that we, or a licensor, were the first to make the inventions 
claimed in any owned or any licensed patents or pending patent applications, respectively, or which entity was the first to 
file for patent protection until such patent application publishes or issues as a patent. Databases for patents and 
publications, and methods for searching them, are inherently limited, so it is not practical to review and know the full 
scope of all issued and pending patent applications. As a result, the issuance, scope, validity, enforceability, and 
commercial value of our and our licensed patent rights are uncertain. 

Even if the patent applications we license or may own in the future do issue as patents, they may not issue in a 

form that will provide us with any meaningful protection, prevent competitors or other third parties from competing with 
us or otherwise provide us with any competitive advantage. Our competitors or other third parties may be able to 
circumvent our patents by developing similar or alternative technologies or products in a non-infringing manner. 

In spite of a legal presumption of validity, the issuance of a patent is not conclusive as to its inventorship, scope, 
validity, or enforceability which may be challenged in the courts and patent offices in the United States and abroad. Such 
challenges may result in loss of exclusivity or in patent claims being narrowed, invalidated or held unenforceable, which 
could limit our ability to stop others from using or commercializing similar or identical technology and products, or limit 
the duration of the patent protection of our technology and product candidates. Given the amount of time required for the 
development, testing and regulatory review of new product candidates, patents protecting such candidates might expire 
before or shortly after such candidates are commercialized. As a result, our intellectual property may not provide us with 
sufficient rights to exclude others from commercializing products similar or identical to ours. 

Our intellectual property licenses with third parties may be subject to disagreements over contract interpretation, 
which could narrow the scope of our rights to the relevant intellectual property or technology, resulting in 
termination of our access to such intellectual property, or increase our financial or other obligations to our licensors. 

The agreements under which we currently license intellectual property or technology from third parties are 

complex, and certain provisions in such agreements may be susceptible to multiple interpretations. The resolution of any 
contract interpretation disagreement that may arise could narrow what we believe to be the scope of our rights to the 
relevant intellectual property or technology, result in loss of access, or increase what we believe to be our financial or 
other obligations under the relevant agreement, any of which could harm our business, financial condition, results of 
operations and prospects. 

We may not be successful in obtaining necessary rights to our product candidates through acquisitions and 
in-licenses. 

We currently have rights to certain intellectual property, through licenses from third parties, to develop our 

product candidates. Because our programs may require the use of proprietary rights held by third parties, the growth of 

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our business likely will depend, in part, on our ability to acquire, in-license or use these proprietary rights. We may be 
unable to acquire or in-license any compositions, methods of use, processes or other intellectual property rights from 
third parties that we identify as necessary for our product candidates. The licensing or acquisition of third-party 
intellectual property rights is a competitive area, and several more established companies may pursue strategies to 
license or acquire third-party intellectual property rights that we may consider attractive. These established companies 
may have a competitive advantage over us due to their size, capital resources and greater clinical or technical 
development and commercialization capabilities. In addition, companies that perceive us to be a competitor may be 
unwilling to assign or license rights to us. We also may be unable to license or acquire third-party intellectual property 
rights on terms that would allow us to make an appropriate return on our investment. 

We sometimes collaborate with non-profit and academic institutions to accelerate our preclinical research or 

development under written agreements with these institutions. Typically, these institutions provide us with an option to 
negotiate a license to any of the institution’s rights in technology resulting from the collaboration. Regardless of such 
option, we may be unable to negotiate a license within the specified timeframe or under terms that are acceptable to us. If 
we are unable to do so, the institution may offer the intellectual property rights to other parties, potentially blocking our 
ability to develop our program. 

If we are unable to successfully obtain rights to required third-party intellectual property rights or maintain the 

existing intellectual property rights we have, we may be required to expend significant time and resources to redesign 
our product candidates or the methods for manufacturing them or to develop or license replacement technology, all of 
which may not be feasible on a technical or commercial basis. If we are unable to do so, we may be unable to develop or 
commercialize the affected product candidates, which could harm our business significantly. 

Obtaining and maintaining our patent protection depends on compliance with various procedural, document 
submission, fee payment and other requirements imposed by government patent agencies, and our patent protection 
could be reduced or eliminated for non-compliance with these requirements. 

Periodic maintenance fees, renewal fees, annuity fees and various other government fees on patents and/or 

applications will be due to be paid to the United States Patent and Trademark Office, or USPTO, and various 
government patent agencies outside of the United States over the lifetime of our licensed patents and/or applications and 
any patent rights we may own in the future. We rely on our outside counsel or our licensing partners to pay these fees 
due to patent agencies. The USPTO and various non-U.S. government patent agencies require compliance with several 
procedural, documentary, fee payment and other similar provisions during the patent application process. We employ 
reputable law firms and other professionals to help us comply and we are also dependent on our licensors to take the 
necessary action to comply with these requirements with respect to our licensed intellectual property. In many cases, an 
inadvertent lapse can be cured by payment of a late fee or by other means in accordance with the applicable rules. There 
are situations, however, in which non-compliance can result in abandonment or lapse of the patent or patent application, 
resulting in partial or complete loss of patent rights in the relevant jurisdiction, and may compromise the strength of 
other intellectual property in our portfolio. In such an event, potential competitors might be able to enter the market and 
this circumstance could harm our business.  

On February 1, 2019 the government of Venezuela, in response to certain US sanctions, began to require that  

foreign entities pay all official fees, including patent fees (either for pending matters or new petitions), in PETRO, a 
“cryptocurrency” created by the Nicolás Maduro administration in February 2018 as a way to collect U.S. dollars while 
avoiding American financial sanctions issued under an Executive Order of President Trump on March 19, 2018. The 
Executive Order banned transactions involving “any digital currency, digital coin, or digital token, that was issued by, 
for, or on behalf of the Government of Venezuela on or after January 9, 2018.” The prohibition is applicable to any U.S. 
entity unless exempted by license. We do not hold such a license and therefore may not be able to secure patents in 
Venezuela.  

We may not be able to protect our intellectual property rights throughout the world. 

Filing, prosecuting and defending patents on product candidates in all countries throughout the world would be 
prohibitively expensive. Our intellectual property rights may vary from country to country and foreign protections could 

110 

be less extensive than those in the United States. In addition, the laws of some foreign countries do not protect 
intellectual property rights to the same extent as federal and state laws in the United States. Consequently, we may not be 
able to prevent third parties from practicing our inventions in all countries outside the United States, or from selling or 
importing products made using our inventions in and into the United States or other jurisdictions. Competitors may use 
our technologies in jurisdictions where we have not obtained patent protection to develop their own products and, 
further, may export otherwise infringing products to territories where we have patent protection, but enforcement is not 
as strong as that in the United States. These products may compete with our products and our patents or other intellectual 
property rights may not be effective or sufficient to prevent them from competing. 

Many companies have encountered significant problems in protecting and defending intellectual property rights 
in foreign jurisdictions. The legal systems of certain countries, particularly certain developing countries, do not favor the 
enforcement of patents, trade secrets and other intellectual property protection, particularly those relating to 
biotechnology products or methods of treatment, which could make it difficult for us to stop the infringement of our 
patents or marketing of competing products in violation of our proprietary rights generally. For example, an April 2014 
report from the Office of the United States Trade Representative identified a number of countries, including India and 
China, where challenges to the procurement and enforcement of patent rights have been reported. Several countries, 
including India and China, have been listed in the report every year since 1989. With Brexit, there is uncertainty 
associated with obtaining, defending, and enforcing intellectual property rights in the United Kingdom. International 
treaties and regulations promulgated as a result of this transition could impede or eliminate our ability to obtain or 
maintain meaningful intellectual property rights in the United Kingdom. Proceedings to enforce our patent rights in 
foreign jurisdictions could result in substantial costs and divert our efforts and attention from other aspects of our 
business, could put our patents at risk of being invalidated or interpreted narrowly and our patent applications at risk of 
not issuing and could provoke third parties to assert claims against us. We may not prevail in any lawsuits that we 
initiate, and the damages or other remedies awarded, if any, may not be commercially meaningful. Accordingly, our 
efforts to enforce our intellectual property rights around the world may be inadequate to obtain a significant commercial 
advantage from the intellectual property that we develop or license. 

Issued patents covering our technology or product candidates could be found invalid or unenforceable if challenged 
in court. We may not be able to protect our trade secrets in court. 

If one of our licensing partners or we initiate legal proceedings against a third party to enforce a patent covering 

our technology or one of our product candidates, the defendant could counterclaim that the patent covering such 
technology or product candidate is invalid or unenforceable. In patent litigation in the United States, defendant 
counterclaims alleging invalidity or unenforceability are commonplace. Grounds for a validity challenge could be an 
alleged failure to meet any of several statutory requirements, including lack of novelty, obviousness, lack of written 
description or non-enablement. Grounds for an unenforceability assertion could be an allegation that an individual 
connected with prosecution of the patent, including an inventor, an employee of the company, a collaborator or advisor, 
withheld information material to patentability from the USPTO, or made a misleading statement, during prosecution. 
Third parties also may raise similar claims before administrative bodies in the United States or abroad, even outside the 
context of litigation. Such mechanisms include pre-issuance submissions, ex parte re-examination, post-grant review, 
inter partes review and equivalent proceedings in foreign jurisdictions. Some of these mechanisms may even be 
exploited anonymously by third parties. Such proceedings could result in the revocation or cancellation of or amendment 
to our patents in such a way that they no longer cover our technology or product candidates. The outcome following 
legal assertions of invalidity and unenforceability is unpredictable. With respect to the validity question, for example, we 
cannot be certain that there is no invalidating prior art, of which the patent examiner and we or our licensing partners 
were unaware during prosecution. If a defendant were to prevail on a legal assertion of invalidity or unenforceability, we 
could lose part or, all of the patent protection on one or more of our product candidates or our supporting technology. 
Such a loss of patent protection could harm our business. 

In addition to the protection afforded by patents, we rely on trade secret protection, nondisclosure, and 
confidentiality agreements to protect proprietary know-how that is not patentable or that we elect not to patent, processes 
for which patents are difficult to enforce and any other elements of our product candidate discovery and development 
processes that involve proprietary know-how, information or technology that is not covered by patents. However, trade 
secrets can be difficult to protect. Some courts inside and outside the United States are less willing or unwilling to 

111 

protect trade secrets. We seek to protect our proprietary technology and processes, in part, by entering into 
confidentiality agreements with our employees, consultants, scientific advisors, collaborators, contractors, and other third 
parties. We cannot guarantee that we have entered into such agreements with each party that may have or have had 
access to our trade secrets or proprietary technology and processes. We also seek to preserve the integrity and 
confidentiality of our data and trade secrets by maintaining physical security of our premises and physical and electronic 
security of our information technology systems. While we have confidence in these individuals, organizations and 
systems, agreements or security measures may be breached, and we may not have adequate remedies for any breach. In 
addition, our trade secrets may otherwise become known or be independently discovered by competitors. 

Third parties may initiate legal proceedings alleging that we are infringing their intellectual property rights, the 
outcome of which would be uncertain and could harm our business. 

Our commercial success depends upon our ability and the ability of our collaborators to develop, manufacture, 
market and sell our product candidates and use our proprietary technologies without infringing the proprietary rights and 
intellectual property of third parties. The biotechnology and pharmaceutical industries are characterized by extensive and 
complex litigation regarding patents and other intellectual property rights. We may become party to, or threatened with, 
infringement litigation claims regarding our products and technology, including claims from competitors or from non-
practicing entities that have no relevant product revenue and against whom our own patent portfolio may have no 
deterrent effect. Moreover, we may become party to, or be threatened with, adversarial proceedings or litigation 
regarding intellectual property rights with respect to our product candidates and technology, including ex parte re-
examination, post grant review and inter partes review before the USPTO or foreign patent offices. Third parties may 
assert infringement claims against us based on existing patents or patents that may be granted in the future, regardless of 
the merit of the claim. There is a risk that third parties may choose to engage in litigation with us to enforce or to 
otherwise assert their patent rights against us. Even if we believe such claims are without merit, a court of competent 
jurisdiction could hold that these third-party patents are valid, enforceable and infringed, which could adversely affect 
our ability to commercialize our product candidates or any other of our product candidates or technologies covered by 
the asserted third-party patents. In order to successfully challenge the validity of any such asserted third-party U.S. patent 
in federal court, we would need to overcome a presumption of validity. As this burden is a high one requiring us to 
present clear and convincing evidence as to the invalidity of any such U.S. patent claim, there is no assurance that a court 
of competent jurisdiction would invalidate the claims of any such U.S. patent. Similar challenges exist in other 
jurisdictions. If we are found to infringe a third-party’s valid and enforceable intellectual property rights, we could be 
required to obtain a license from such third-party to continue developing, manufacturing and marketing our product 
candidates and technology. However, we may not be able to obtain any required license on commercially reasonable 
terms or at all. Even if we were able to obtain a license, it could be non-exclusive, thereby giving our competitors and 
other third parties access to the same technologies licensed to us, and it could require us to make substantial licensing 
and royalty payments. We could be forced, including by court order, to cease developing, manufacturing and 
commercializing the infringing technology or product candidates. In addition, we could be found liable for monetary 
damages, including treble damages and attorneys’ fees, if we are found to have willfully infringed a patent or other 
intellectual property right. A finding of infringement could prevent us from manufacturing and commercializing our 
product candidates or force us to cease some of our business operations, which could harm our business. In addition, we 
may be forced to redesign our product candidates, seek new regulatory approvals, and indemnify third parties pursuant to 
contractual agreements. Claims that we have misappropriated the confidential information or trade secrets of third parties 
could have a similar negative impact on our business, reputation, financial condition, results of operations and prospects. 

Intellectual property litigation could cause us to spend substantial resources and distract our personnel from their 
normal responsibilities. 

Competitors may infringe our intellectual property rights or the intellectual property rights of our licensing 

partners, or we may be required to defend against claims of infringement. To counter infringement or unauthorized use 
claims or to defend against claims of infringement can be expensive and time consuming. Even if resolved in our favor, 
litigation or other legal proceedings relating to intellectual property claims may cause us to incur significant expenses 
and could distract our technical and management personnel from their normal responsibilities. In addition, there could be 
public announcements of the results of hearings, motions or other interim proceedings or developments. If securities 
analysts or investors perceive these results to be negative, it could have a substantial adverse effect on the price of our 

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common stock. Such litigation or proceedings could substantially increase our operating losses and reduce the resources 
available for development activities or any future sales, marketing or distribution activities. We may not have sufficient 
financial or other resources to conduct such litigation or proceedings adequately. Some of our competitors may be able to 
sustain the costs of such litigation or proceedings more effectively than we can because of their greater financial 
resources and more mature and developed intellectual property portfolios. Uncertainties resulting from the initiation and 
continuation of patent litigation or other proceedings could adversely affect our ability to compete in the marketplace. 

We may be subject to claims asserting that our employees, consultants or advisors have wrongfully used or disclosed 
alleged trade secrets of their current or former employers or claims asserting ownership of what we regard as our own 
intellectual property. 

Many of our directors, employees, consultants, and advisors are currently, or were previously, employed at 
universities or other biotechnology or pharmaceutical companies, including our competitors or potential competitors. 
Although we try to ensure that these individuals do not use the proprietary information or know-how of others in their 
work for us, we may be subject to claims that these individuals or we have used or disclosed intellectual property, 
including trade secrets or other proprietary information, of any such individual’s current or former employer. Litigation 
may be necessary to defend against these claims. If we fail in defending any such claims, in addition to paying monetary 
damages, we may lose valuable intellectual property rights or personnel. Even if we are successful in defending against 
such claims, litigation could result in substantial costs and be a distraction to management. 

In addition, while it is our policy to require our employees, consultants, advisors and contractors who may be 

involved in the conception or development of intellectual property to execute agreements assigning such intellectual 
property to us, we may be unsuccessful in executing such an agreement with each party who, in fact, conceives or 
develops intellectual property that we regard as our own. The assignment of intellectual property rights may not be 
self-executing or the assignment agreements may be breached, and we may be forced to bring claims against third 
parties, or defend claims that they may bring against us, to determine the ownership of what we regard as our intellectual 
property. 

If we fail in prosecuting or defending any such claims, in addition to paying monetary damages, we may lose 

valuable intellectual property rights or personnel. Even if we are successful in prosecuting or defending against such 
claims, litigation could result in substantial costs and be a distraction to management.  

Changes in U.S. patent law could diminish the value of patents in general, thereby impairing our ability to protect our 
products. 

Patent reform legislation could increase the uncertainties and costs surrounding the prosecution of patent 
applications and the enforcement or defense of issued patents. On September 16, 2011, the Leahy-Smith America 
Invents Act, or the Leahy-Smith Act, was signed into law. The Leahy-Smith Act includes several significant changes to 
U.S. patent law. These include provisions that affect the way patent applications are prosecuted and also may affect 
patent litigation. These also include provisions that switched the United States from a “first-to-invent” system to a 
“first-inventor-to-file” system, allow third-party submission of prior art to the USPTO during patent prosecution and set 
forth additional procedures to attack the validity of a patent by the USPTO administered post grant proceedings. Under a 
first-inventor-to-file system, assuming the other requirements for patentability are met, the first inventor to file a patent 
application generally will be entitled to the patent on an invention regardless of whether another inventor had made the 
invention earlier. The USPTO has promulgated regulations and procedures to govern administration of the Leahy-Smith 
Act, and many of the substantive changes to patent law associated with the Leahy-Smith Act, and in particular, the 
first-inventor-to-file provisions, became effective on March 16, 2013. The Leahy-Smith Act has resulted in an increased 
investment in filing applications earlier, and consequently has increased the uncertainties and costs surrounding the 
prosecution of our patent applications, and may increase the enforcement or defense of our issued patents, all of which 
could harm our business, financial condition, results of operations and prospects. 

The  administrative tribunal created by the Leahy-Smith Act, known as the Patent Trial and Appeals Board, or 

PTAB, may have an impact on the operation of our business in the future. For example, the initial results of patent 
challenge proceedings before the PTAB since its inception in 2013 have resulted in the invalidation of many U.S. patent 

113 

claims. The availability of the PTAB as a lower-cost, faster and potentially more potent tribunal for challenging patents 
could therefore increase the likelihood that our own licensed patents will be challenged, thereby increasing the 
uncertainties and costs of maintaining and enforcing them. Moreover, if such challenges occur, we may not have the 
right to control the defense. In certain situations, we may be required to rely on our licensor to consider our suggestions 
and to defend such challenges, with the possibility that it may not do so in a way that best protects our interests.  

We also may be subject to a third-party pre-issuance submission of prior art to the USPTO or become involved 

in other contested proceedings such as opposition, derivation, reexamination, inter partes review, or post-grant review 
proceedings challenging our patent rights or the patent rights of others. An adverse determination in any such 
submission, proceeding or litigation could reduce the scope of, or invalidate, our patent rights, allow third parties to 
commercialize our technology or products and compete directly with us, without payment to us, or result in our inability 
to manufacture or commercialize products without infringing third-party patent rights. In addition, if the breadth or 
strength of protection provided by our patents and patent applications is threatened, it could dissuade companies from 
collaborating with us to license, develop or commercialize current or future products. 

The patent positions of companies engaged in the development and commercialization of biologics and 

pharmaceuticals are particularly uncertain as the courts address issues such as patenting genes or gene products. The 
most recent guidance provided under  Berkheimer v HP, Inc. (April 19, 2018) and Vanda Pharmaceuticals, Inc. v West-
Ward Pharmaceuticals (June 7, 2018) instruct USPTO examiners on the ramifications of the court rulings as applied to 
natural products and principles including all naturally occurring nucleic acids. Patents for certain of our product 
candidates contain claims related to specific DNA sequences that are naturally occurring and, therefore, could be the 
subject of future challenges made by third parties. In addition, the recent USPTO guidance could make it impossible for 
us to pursue similar patent claims in patent applications we may prosecute in the future. 

We cannot assure you that our efforts to seek patent protection for our technology and products will not be 

negatively impacted by the court decisions referenced above, rulings in other cases or changes in guidance or procedures 
issued by the USPTO. We cannot fully predict what impact decisions from the Supreme Court’s decisions in Mayo 
Collaborative Services v. Prometheus Laboratories and Molecular Pathology v. Myriad Genetics, Inc. or other 
applicable court decisions may have on the ability of life science companies to obtain or enforce patents relating to their 
products and technologies in the future. These decisions, the guidance issued by the USPTO and rulings in other cases or 
changes in USPTO guidance or procedures could have an adverse effect on our existing patent portfolio and our ability 
to protect and enforce our intellectual property in the future. 

Moreover, although the Supreme Court has held that isolated segments of naturally occurring DNA are not 

patent-eligible subject matter, certain third parties could allege that activities that we may undertake infringe other 
gene-related patent claims, and we may deem it necessary to defend ourselves against these claims by asserting 
non-infringement and/or invalidity positions, or paying to obtain a license to these claims. In any of the foregoing or in 
other situations involving third-party intellectual property rights, if we are unsuccessful in defending against claims of 
patent infringement, we could be forced to pay damages or be subjected to an injunction that would prevent us from 
utilizing the patented subject matter. Such outcomes could harm our business, financial condition, results of operations 
or prospects. 

Outside the United States, other courts have also begun to address the patenting of genetic material. In August 
2015, the Australian High Court ruled that isolated genes cannot be patented in Australia. The decision did not address 
methods of using genetic material. Any ruling of a similar scope in other countries could affect the scope of our 
intellectual property rights. The ambiguities and changing law in all countries as to patenting genetic material may 
directly affect our ability to secure and/or maintain patent protection for our products. 

If we do not obtain patent term extension and data exclusivity for our product candidates, our business may be 
harmed. 

Depending upon the timing, duration and specifics of any FDA marketing approval of our product candidates, 
one or more of our U.S. patents, which may cover non-gene therapy compounds, may be eligible for limited patent term 
extension under the Drug Price Competition and Patent Term Restoration Act of 1984, or the Hatch-Waxman Act. The 

114 

Hatch-Waxman Act permits a patent extension term of up to five years as compensation for patent term lost during the 
FDA regulatory review process. A patent term extension cannot extend the remaining term of a patent beyond a total of 
14 years from the date of product approval, only one patent may be extended per FDA-approved product, and only those 
claims covering the approved drug, a method for using it or a method for manufacturing it may be extended. Further, 
certain of our licenses currently or in the future may not provide us with the right to control decisions the licensor or its 
other licensees on Orange Book listings or patent term extension decisions under the Hatch-Waxman Act. Thus, if one of 
our important licensed patents is eligible for a patent term extension under the Hatch-Waxman Act, and it covers a 
product of another licensee in addition to our own product candidate, we may not be able to obtain that extension if the 
other licensee seeks and obtains that extension first. However, we may not be granted an extension because of, for 
example, failing to exercise due diligence during the testing phase or regulatory review process, failing to apply within 
applicable deadlines, failing to apply prior to expiration of relevant patents or otherwise failing to satisfy applicable 
requirements.  

The Biologics Price Competition and Innovation Act of 2009 provides 12 years of market exclusivity for a 

reference biological product. We may not be able to obtain such exclusivity for our products. Moreover, the applicable 
time-period or the scope of patent protection afforded during any such extension could be less than we request. If we are 
unable to obtain patent term extension or the scope of term of any such extension is less than we request, the period 
during which we will have the right to exclusively market our product may be shortened and our competitors may obtain 
approval of competing products following our patent expiration, and our revenue could be materially reduced. 

If our trademarks and trade name are not adequately protected, then we may not be able to build name recognition in 
our markets of interest and our business may be adversely affected. 

We own service mark registrations in the USPTO for the marks “VOYAGER THERAPEUTICS” and 
“VOYAGER THERAPEUTICS Logo” and European Community trademark registrations for the marks “V-TAG” and 
“VOYAGER TRAJECTORY ARRAY GUIDE.” We also own pending trademark applications in the USPTO for the 
marks “V-TAG” and the V-TAG Logo. Our trademarks or our trade name may be challenged, infringed, circumvented or 
declared generic or found to infringe prior third-party marks. We may not be able to protect our rights in our trademarks 
or in our trade name, which we need in order to build name recognition among potential partners or customers in our 
markets of interest. It is possible that competitors may adopt trade names or trademarks similar to ours, thereby impeding 
our ability to build brand identity and possibly leading to market confusion. In addition, there could be potential trade 
name or trademark infringement claims brought by owners of prior registered trademarks or trademarks that incorporate 
variations of our registered or unregistered trademarks or trade name. Over the long term, if we are unable to establish 
name recognition based on our trademarks and trade name, then we may not be able to compete effectively, and our 
business may be adversely affected. Our efforts to enforce and protect our proprietary rights related to trademarks, trade 
secrets, domain names, copyrights and other intellectual property may be ineffective and could result in substantial costs 
and diversion of resources and could adversely impact our financial condition or results of operations. 

Intellectual property rights do not necessarily address all potential threats. 

The degree of future protection afforded by our intellectual property rights is uncertain because intellectual 
property rights have limitations, and such rights may not adequately protect our business or permit us to maintain our 
competitive advantage. For example: 

 

others may be able to make gene therapy products that are similar to our product candidates but that are not 
covered by the claims of the patents that we own, license or may access in the future; 

  we, or our license partners or current or future collaborators, might not have been the first to make the 

inventions covered by the issued patent or pending patent application that we license or may own in the 
future; 

  we, or our license partners or current or future collaborators, might not have been the first to file patent 

applications covering certain of our or their inventions; 

115 

 

 

 

 

others may independently develop similar or alternative technologies or duplicate any of our technologies 
without infringing our owned or licensed intellectual property rights; 

it is possible that our pending patent applications or those that we may own in the future will not lead to 
issued patents; 

issued patents that we hold rights to may be held invalid or unenforceable, including as a result of legal 
challenges by our competitors; 

our competitors might conduct research and development activities in countries where we do not have 
patent rights and then use the information learned from such activities to develop competitive products for 
sale in our major commercial markets; 

  we may not develop additional proprietary technologies that are patentable; 

 

the patents of others may have an adverse effect on our business; and 

  we may choose not to file a patent for certain inventions, trade secrets or know-how, and a third party may 

subsequently file a patent covering such intellectual property. 

Should any of these events occur, they could significantly harm our business, financial condition, results of 

operations and prospects. 

Our reliance on third parties requires us to share our trade secrets, which increases the possibility that a competitor 
will discover them or that our trade secrets will be misappropriated or disclosed. 

Because we currently rely on certain third parties to manufacture all or part of our product candidates and to 

perform quality testing, and because we collaborate with various organizations and academic institutions for the 
advancement of our gene therapy platform and pipeline, we must, at times, share our proprietary technology and 
confidential information, including trade secrets, with them. We seek to protect our proprietary technology, in part, by 
entering into confidentiality agreements and, if applicable, material transfer agreements, collaborative research 
agreements, consulting agreements or other similar agreements with our collaborators, advisors, employees and 
consultants prior to beginning research or disclosing proprietary information. These agreements typically limit the rights 
of the third parties to use or disclose our confidential information. Despite the contractual provisions employed when 
working with third parties, the need to share trade secrets and other confidential information increases the risk that such 
trade secrets become known by our competitors, are inadvertently incorporated into the technology of others or are 
disclosed or used in violation of these agreements. Given that our proprietary position is based, in part, on our know-how 
and trade secrets, a competitor’s discovery of our proprietary technology and confidential information or other 
unauthorized use or disclosure would impair our competitive position and may harm our business, financial condition, 
results of operations and prospects. 

Despite our efforts to protect our trade secrets, our competitors may discover our trade secrets, either through 
breach of these agreements, independent development or publication of information including our trade secrets by third 
parties. A competitor’s discovery of our trade secrets would impair our competitive position and have an adverse impact 
on our business, financial condition, results of operations and prospects. 

Risks Related to Ownership of Our Common Stock  

Our executive officers, directors, principal stockholders and their affiliates exercise significant influence over our 
company. 

The holdings of our executive officers, directors, principal stockholders and their affiliates, including 

investment funds affiliated with Third Rock Ventures, Bellevue Asset Management, Armistice Capital, UBS 
Switzerland, and BlackRock Institutional Trust Company, N.A., represent beneficial ownership, in the aggregate, of 
approximately 56% of our outstanding common stock as of December 31, 2018. As a result, these stockholders, if they 

116 

act together, will be able to influence our management and affairs and the outcome of matters submitted to our 
stockholders for approval, including the election of directors and any sale, merger, consolidation, or sale of all or 
substantially all of our assets. In addition, this concentration of ownership might adversely affect the market price of our 
common stock by: 

 

 

 

delaying, deferring or preventing a change of control of us; 

impeding a merger, consolidation, takeover or other business combination involving us; or 

discouraging a potential acquirer from making a tender offer or otherwise attempting to obtain control of 
us. 

An active trading market for our common stock may not be sustained. 

Our shares of common stock began trading on the Nasdaq Global Select Market on November 11, 2015. Given 

the limited trading history of our common stock, there is a risk that an active trading market for our shares will not be 
sustained, which could put downward pressure on the market price of our common stock and thereby affect the ability of 
our stockholders to sell their shares. 

If securities analysts do not publish research or reports about our business or if they publish negative evaluations of 
our stock, the price of our stock could decline.  

The trading market for our common stock will rely in part on the research and reports that industry or financial 

analysts publish about us or our business. If no or few analysts maintain coverage of us, the trading price of our stock 
would likely decrease. If one or more of the analysts covering our business downgrade their evaluations of our stock, the 
price of our stock could decline. If one or more of these analysts cease to cover our stock, we could lose visibility in the 
market for our stock, which in turn could cause our stock price to decline.  

Sales of a substantial number of shares of our common stock in the public market could cause our stock price to fall. 

Persons who were our stockholders prior to our initial public offering continue to hold a substantial number of 

shares of our common stock. If such persons sell, or indicate an intention to sell, substantial amounts of our common 
stock in the public market, the trading price of our common stock could decline. 

In addition, shares of common stock that are either subject to outstanding options or restricted stock units, or 

RSUs, or reserved for future issuance under our stock incentive plans will become eligible for sale in the public market 
to the extent permitted by the provisions of various vesting schedules and Rule 144 and Rule 701 under the Securities 
Act of 1933, as amended, and, in any event, we have filed a registration statement permitting shares of common stock 
issued on exercise of options or the settlement of RSUs to be freely sold in the public market. If these additional shares 
of common stock are sold, or if it is perceived that they will be sold, in the public market, the trading price of our 
common stock could decline. In addition, we have registered on a registration statement on Form S-3 that has been 
declared effective, the sale of up to $250.0 million in aggregate of an indeterminate number of shares of common stock 
and preferred stock, an indeterminate principal amount of debt securities, and an indeterminate number of warrants. In 
November 2017, under such shelf registration statement, we issued and sold 5,175,000 shares of common stock to the 
public at an offering price of $12.00 per share, including 675,000 shares of common stock issued upon the full exercise 
by the underwriter of their option to purchase additional shares, resulting in net proceeds of $58.0 million after deducting 
underwriting discounts, commissions, and offering expenses payable by us. The registration statement also registers the 
offering, issuance, and sale of common stock having up to a maximum aggregate offering price of $75.0 million that we 
may issue and sell in at-the-market offerings or negotiated transactions under a sales agreement we entered into with 
Cowen and Company, LLC on December 1, 2016 pursuant to a sales agreement prospectus that forms a part of the 
registration statement. As of February 26, 2019, $75.0 million in shares of common stock remained eligible for sale 
under the sales agreement. In January 2019, we executed a stock purchase agreement to sell common stock to Neurocrine 
for an aggregate purchase price of approximately $50.0 million. The sale of shares to Neurocrine is subject to customary 
closing conditions, including certain antitrust approvals, that have not been satisfied as of the date of this Annual Report 
on Form 10-K. As a result, we have not yet issued such shares to Neurocrine, but, if the applicable closing conditions are 
met, we expect to do so promptly. 

117 

Certain holders of our common stock have rights, subject to specified conditions, to require us to file 

registration statements covering their shares or to include their shares in registration statements that we may file for 
ourselves or other stockholders. Any sales of securities by these stockholders could have a material adverse effect on the 
trading price of our common stock. 

The price of our common stock may be volatile and fluctuate substantially, which could result in substantial losses for 
purchasers of our common stock. 

The price of our common stock is likely to be volatile and may fluctuate substantially. From January 1, 2018 

through December 31, 2018, the sales price of our common stock ranged from a high of $31.91 to a low of $8.30 on the 
Nasdaq Global Select Market. As a result of this volatility, our stockholders may not be able to sell their common stock 
at or above the price at which they purchased it. The market price for our common stock may be influenced by many 
factors, including: 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

our success in commercializing any product candidates for which we obtain marketing approval; 

regulatory action and results of clinical trials of our product candidates or those of our competitors; 

the success of competitive products or technologies; 

the results of clinical trials of our product candidates; 

the results of clinical trials of product candidates of our competitors; 

the commencement, termination, and success of our collaborations; 

regulatory or legal developments in the United States and other countries; 

developments or disputes concerning patent applications, issued patents or other proprietary rights; 

the recruitment or departure of key personnel; 

the level of expenses related to any of our product candidates or clinical development programs; 

the results of our efforts to discover, develop, acquire or in-license additional product candidates or 
technologies, the cost of commercializing such product candidates, and the cost of development of any such 
product candidates or technologies; 

actual or anticipated changes in estimates as to financial results, development timelines or 
recommendations by securities analysts; 

variations in our financial results or those of companies that are perceived to be similar to us; 

the ability to secure third-party reimbursement for our product candidates; 

changes in the structure of healthcare payment systems; 

  market conditions in the pharmaceutical and biotechnology sectors; 

 

 

general economic, industry and market conditions; and 

the other factors described in this “Risk Factors” section and elsewhere in this Annual Report on Form 10-
K.  

If our operating results fall below the expectations of investors or securities analysts for a given period, the 

price of our common stock could decline substantially. Furthermore, any fluctuations in our operating results from period 

118 

to period may, in turn, cause the price of our stock to fluctuate substantially. We believe that such comparisons of our 
financial results are not necessarily meaningful and should not be relied upon as an indication of our future performance. 

In the past, following periods of volatility in the market price of a company’s securities, securities class-action 
litigation often has been instituted against that company. We also may face securities class-action litigation if we cannot 
obtain regulatory approvals for or if we otherwise fail to commercialize our product candidates. Such litigation, if 
instituted against us, could cause us to incur substantial costs to defend such claims and divert management’s attention 
and resources, which could seriously harm our business, financial condition, results of operations and prospects. 

We have broad discretion in how we apply our available funds, and we may not use these funds effectively, which 
could affect our results of operations and cause our stock price to decline.  

Our management will have broad discretion in the application of our existing cash, cash equivalents and 

marketable securities and could spend these funds in ways that do not improve our results of operations or enhance the 
value of our common stock. The failure by our management to apply our available funds effectively could result in 
financial losses that could cause the price of our common stock to decline and delay the development of our product 
candidates and preclinical programs. Pending their use, we may invest our available funds in a manner that does not 
produce income or that loses value.  

We are an “emerging growth company” and a “smaller reporting company” and the reduced disclosure requirements 
applicable to such companies may make our common stock less attractive to investors. 

For so long as we remain an “emerging growth company,” or EGC, as defined in the JOBS Act, we are 

permitted and intend to rely on exemptions from certain disclosure requirements that are applicable to other public 
companies that are not EGCs. These exemptions include: 

 

 

 

 

not being required to comply with the auditor attestation requirements in the assessment of our internal 
control over financial reporting pursuant to Section 404 of the Sarbanes-Oxley Act of 2002; 

not being required to comply with any requirement that may be adopted by the Public Company 
Accounting Oversight Board regarding mandatory audit firm rotation or a supplement to the auditor’s 
report providing additional information about the audit and the financial statements; 

reduced disclosure obligations regarding executive compensation; and 

exemptions from the requirements of holding a nonbinding advisory vote on executive compensation and 
stockholder approval of any golden parachute payments not previously approved. 

We may take advantage of these exemptions until we are no longer an EGC. We would cease to be an EGC 

upon the earlier of: (i) the last day of the fiscal year in which we have total annual gross revenue of $1.07 billion or 
more; (ii) the last day of the fiscal year following the fifth anniversary of the date of the completion of our IPO, which is 
December 31, 2020; (iii) the date on which we have issued more than $1.0 billion in nonconvertible debt during the 
previous three years; or (iv) the date on which we are deemed to be a large accelerated filer under the rules of the 
Securities and Exchange Commission, or SEC, which means the first day of the year following the first year in which the 
market value of our common stock that is held by non-affiliates exceeds $700.0 million as of June 30th. 

We expect to continue to take advantage of some, but not all, of the available exemptions. Even after we no 

longer qualify as an emerging growth company, we might still qualify as a smaller reporting company, or SRC, which 
would allow us to take advantage of many of the same exemptions from disclosure requirements, including reduced 
disclosure obligations regarding executive compensation. We cannot predict whether investors will find our common 
stock less attractive if we rely on certain or all of these exemptions. If some investors find our common stock less 
attractive as a result, there may be a less active trading market for our common stock and our stock price may be more 
volatile and may decline. 

119 

In addition, the JOBS Act provides that an EGC may take advantage of an extended transition period for 

complying with new or revised accounting standards. This allows an EGC to delay the adoption of certain accounting 
standards until those standards would otherwise apply to private companies. We have irrevocably elected not to avail 
ourselves of this exemption from new or revised accounting standards and, therefore, we will be subject to the same new 
or revised accounting standards as other public companies that are not emerging growth companies. 

We incur increased costs as a result of operating as a public company, and our management is required to devote 
substantial time to new compliance initiatives. 

As a public company, and particularly after we are no longer an EGC or SRC, we incur significant legal, 

accounting and other expenses that we did not incur as a private company. In addition, the Sarbanes-Oxley Act of 2002, 
the Dodd-Frank Wall Street Reform and Consumer Protection Act, and rules subsequently implemented by the SEC and 
The Nasdaq Stock Market have imposed various requirements on public companies, including establishment and 
maintenance of effective disclosure and financial controls and corporate governance practices. Our management and 
other personnel devote a substantial amount of time to these compliance initiatives. Moreover, these rules and 
regulations have increased our legal and financial compliance costs and have made some activities more time-consuming 
and costly. For example, these rules and regulations have made it more difficult and more expensive for us to obtain 
director and officer liability insurance, and we have been required to accept reduced policy limits and coverage or incur 
substantially higher costs to obtain the same or similar coverage. As a result, it may be more difficult for us to attract and 
retain qualified people to serve on our board of directors, our board committees or as executive officers. 

Pursuant to Section 404 of the Sarbanes-Oxley Act of 2002, or Section 404, we will be required to furnish a 

report by our management on our internal control over financial reporting. However, while we remain an EGC, we will 
not be required to include an attestation report on internal control over financial reporting issued by our independent 
registered public accounting firm. To achieve compliance with Section 404 within the prescribed period, we will be 
engaged in a process to document and evaluate our internal control over financial reporting, which is both costly and 
challenging. In this regard, we will need to continue to dedicate internal resources, potentially engage outside consultants 
and adopt a detailed work plan to assess and document the adequacy of internal control over financial reporting, continue 
steps to improve control processes as appropriate, validate through testing that controls are functioning as documented 
and implement a continuous reporting and improvement process for internal control over financial reporting. Despite our 
efforts, there is a risk that neither we nor our independent registered public accounting firm will be able to conclude 
within the prescribed timeframe, or at all, that our internal control over financial reporting is effective as required by 
Section 404. If we identify one or more material weaknesses in our internal control over financial reporting, it could 
result in an adverse reaction in the financial markets due to a loss of confidence in the reliability of our financial 
statements. 

Provisions in our amended and restated certificate of incorporation and bylaws and Delaware law could make an 
acquisition of us, which may be beneficial to our stockholders, more difficult and may prevent attempts by our 
stockholders to replace or remove our current management. 

Provisions in our amended and restated certificate of incorporation and bylaws may discourage, delay or 
prevent a merger, acquisition or other change in control of us that stockholders may consider favorable, including 
transactions in which our stockholders might otherwise receive a premium for their shares. These provisions also could 
limit the price that investors might be willing to pay in the future for shares of our common stock, thereby depressing the 
market price of our common stock. In addition, because our board of directors is responsible for appointing the members 
of our management team, these provisions may frustrate or prevent any attempts by our stockholders to replace or 
remove our current management by making it more difficult for stockholders to replace members of our board of 
directors. Among other things, these provisions: 

 

establish a classified board of directors such that only one of three classes of members of the board is 
elected each year; 

 

allow the authorized number of our directors to be changed only by resolution of our board of directors; 

120 

 

 

 

 

 

 

limit the manner in which stockholders can remove directors from the board; 

establish advance notice requirements for stockholder proposals that can be acted on at stockholder 
meetings and nominations to our board of directors; 

require that stockholder actions must be effected at a duly called stockholder meeting and prohibit actions 
by our stockholders by written consent; 

limit who may call stockholder meetings; 

authorize our board of directors to issue preferred stock without stockholder approval, which could be used 
to institute a stockholder rights plan, or so-called “poison pill,” that would work to dilute the stock 
ownership of a potential hostile acquirer, effectively preventing acquisitions that have not been approved 
by our board of directors; and 

require the approval of the holders of at least 75% of the votes that all our stockholders would be entitled to 
cast to amend or repeal certain provisions of our amended and restated certificate of incorporation or 
bylaws. 

Moreover, because we are incorporated in Delaware, we are governed by the provisions of Section 203 of the 

Delaware General Corporation Law, which prohibits a person who owns in excess of 15% of our outstanding voting 
stock from merging or combining with us for a period of three years after the date of the transaction in which the person 
acquired in excess of 15% of our outstanding voting stock, unless the merger or combination is approved in a prescribed 
manner. 

Our amended and restated certificate of incorporation designates the Court of Chancery of the State of Delaware as 
the sole and exclusive forum for certain types of actions and proceedings that may be initiated by our stockholders, 
which could limit our stockholders’ ability to obtain a favorable judicial forum for disputes with us or our directors, 
officers or employees. 

Our amended and restated certificate of incorporation, provides that, unless we consent in writing to an 

alternative forum, the Court of Chancery of the State of Delaware will be the sole and exclusive forum for (i) any 
derivative action or proceeding brought on our behalf, (ii) any action asserting a claim of breach of a fiduciary duty owed 
by any of our directors, officers and employees to us or our stockholders, (iii) any action asserting a claim arising 
pursuant to any provision of the Delaware General Corporation Law, our certificate of incorporation or our bylaws or 
(iv) any action asserting a claim that is governed by the internal affairs doctrine, in each case subject to the Court of 
Chancery having personal jurisdiction over the indispensable parties named as defendants therein. Any person 
purchasing or otherwise acquiring any interest in any shares of our capital stock shall be deemed to have notice of and to 
have consented to this provision of our amended and restated certificate of incorporation. This choice of forum provision 
may limit a stockholder’s ability to bring a claim in a judicial forum that he, she or it finds favorable for disputes with us 
or our directors, officers or employees, which may discourage such lawsuits against us and our directors, officers and 
employees even though an action, if successful, might benefit our stockholders. Stockholders who do bring a claim in the 
Court of Chancery could face additional litigation costs in pursuing any such claim, particularly if they do not reside in 
or near the State of Delaware. The Court of Chancery may also reach different judgments or results than would other 
courts, including courts where a stockholder considering an action may be located or would otherwise choose to bring 
the action, and such judgments or results may be more favorable to us than to our stockholders. Alternatively, if a court 
were to find this provision of our amended and restated certificate of incorporation inapplicable to, or unenforceable in 
respect of, one or more of the specified types of actions or proceedings, we may incur additional costs which could have 
a material adverse effect on our business, financial condition or results of operations. 

121 

Because we do not anticipate paying any cash dividends on our capital stock in the foreseeable future, capital 
appreciation, if any, will be our stockholders’ sole source of gain. 

We have never declared or paid cash dividends on our capital stock. We currently intend to retain all of our future 
earnings, if any, to finance the growth and development of our business. In addition, the terms of any future debt 
agreements may preclude us from paying dividends. As a result, capital appreciation, if any, of our common stock will 
be the sole source of gain for our stockholders for the foreseeable future. 

ITEM 1B.  

    UNRESOLVED STAFF COMMENTS 

Not applicable. 

ITEM 2.  

  PROPERTIES 

Our corporate headquarters are located in Cambridge, Massachusetts. Our current leased facilities encompass 

approximately 74,000 square feet of office and laboratory space, located at 75 Sidney Street and 64 Sidney Street, 
Cambridge, Massachusetts.  

ITEM 3.  

  LEGAL PROCEEDINGS 

In the ordinary course of business, we are from time to time involved in lawsuits, claims, investigations, 
proceedings, and threats of litigation relating to intellectual property, commercial arrangements and other matters. While 
the outcome of these proceedings and claims cannot be predicted with certainty, as of December 31, 2018, we were not 
party to any legal matters, claims, or arbitration proceedings that may have, or have had in the recent past, significant 
effects on our financial position or profitability. No governmental proceedings are pending or, to our knowledge, 
contemplated against us. We are not a party to any material proceedings in which any director, member of senior 
management or affiliate of ours is either a party adverse to us or our subsidiaries or has a material interest adverse to us 
or our subsidiaries. 

ITEM 4.  

  MINE SAFETY DISCLOSURES 

Not applicable. 

PART II 

ITEM 5.  

  MARKET FOR REGISTRANT’S COMMON EQUITY, RELATED STOCKHOLDER 

MATTERS AND ISSUER PURCHASES OF EQUITY SECURITIES. 

Our common stock has been traded on the Nasdaq Global Select Market under the symbol “VYGR” since 

November 11, 2015. Prior to this time, there was no public market for our common stock. On February 22, 2019, the last 
reported sale price for our common stock on the Nasdaq Global Select Market was $13.12 per share.  

Stockholders  

As of February 22, 2019, there were approximately 15 holders of record of our common stock. The actual 
number of stockholders is greater than this number of record holders, and includes stockholders who are beneficial 
owners, but whose shares are held in street name by brokers and other nominees. This number of holders of record also 
does not include stockholders whose shares may be held in trust by other entities.  

Dividends  

We have not paid any cash dividends on our common stock since inception and do not anticipate paying cash 

dividends in the foreseeable future.  

122 

  
 
  
  
 
  
  
  
  
 
 
  
  
 
 
Securities Authorized for Issuance Under Equity Compensation Plans  

Information about our equity compensation plans is incorporated herein by reference to Item 12 of Part III of 

this Annual Report on Form 10-K.  

Unregistered Sales of Equity Securities 

On July 16, 2018, we issued to our new President and Chief Executive Officer, G. Andre Turenne, a non-
statutory stock option to purchase an aggregate of 650,000 shares of our common stock at an exercise price of $18.03 per 
share. This option was granted as an inducement material to Mr. Turenne’s entry into employment with us and was 
issued outside our existing equity compensation plans in accordance with Nasdaq Stock Market Listing Rule 5635(c)(4). 
We also intend to issue non-statutory stock options to purchase an aggregate of 338,750 shares of our common stock and 
an aggregate of 58,125 restricted stock units, or RSUs, to certain of our future executive officers who have executed 
employment agreements but not yet commenced employment with us. These equity awards were granted as inducements 
material to the respective officers’ entry into their employment agreements with us and will be issued outside of our 
existing equity compensation plans in accordance with Nasdaq Stock Market Listing Rule 5635(c)(4) on such officers’ 
first day of employment. 

We intend to file a registration statement on a Form S-8 to register the shares of common stock underlying the 

stock options granted to Mr. Turenne and the stock options and RSUs granted to our future executive officers prior to the 
time at which the shares underlying such options become exercisable or such RSUs become settleable. 

Additionally, on January 28, 2019, in connection with the Neurocrine Collaboration, we agreed to sell 
4,179,728 shares of Common Stock to Neurocrine at a price of $11.9625 per share, for an aggregate purchase price of 
approximately $50.0 million. The sale of shares to Neurocrine is subject to customary closing conditions, including 
certain antitrust approvals, that have not been satisfied as of the date of this Annual Report on Form 10-K. As a result, 
we have not yet issued such shares to Neurocrine, but, if the applicable closing conditions are met, we expect to do so 
promptly thereafter. We expect the shares to be issued in reliance on the exemption from registration under Section 
4(a)(2) of the Securities Act of 1933, as amended, for a transaction by an issuer not involving any public offering within 
the meaning of Section 4(a)(2) thereunder.  

Purchases of Equity Securities by the Issuer or Affiliated Purchasers  

There were no repurchases of shares of common stock made during the year ended December 31, 2018.  

ITEM 6.  

    SELECTED FINANCIAL DATA 

The  following  financial  data  should  be  read  in  conjunction  with  “Management’s  Discussion  and  Analysis  of 
Financial  Condition  and  Results  of  Operations”,  the  financial  statements  and  related  notes,  and  other  financial 
information included in this Annual Report on Form 10-K. 

123 

 
  
  
 
 
We have derived the statements of operations data for the years ended December 31, 2018, 2017, and 2016, and 

the balance sheet data as of December 31, 2018 and 2017, from our audited financial statements included elsewhere in 
this Annual Report on Form 10-K. We have derived the statements of operations data for the years ended December 31, 
2015 and 2014, and the balance sheet data as of December 31, 2016, 2015, and 2014, from our audited consolidated 
financial statements not included in this Annual Report on Form 10-K. Historical results are not necessarily indicative 
of the results to be expected in future periods.  

Consolidated statements of operations 
data: 
Collaboration revenue 
Operating expenses: 

Research and development 
General and administrative 

Total operating expenses 

Loss from operations 

Interest income (expense), net 
Other (expense) income, net 

Loss before income taxes 
Income tax (benefit) provision 
Net loss 
Other comprehensive loss 

Net unrealized gain (loss) on available-
for-sale-securities, net 

Comprehensive loss 
Reconciliation of net loss to net loss 
attributable to common stockholders: 
Net loss 

Accretion of preferred stock to 
redemption value 
Accrued dividends on series A preferred 
stock 

Net loss attributable to common 
stockholders 
Net loss per share attributable to common 
stockholders—basic and diluted(1) 
Weighted average number of common 
shares used in net loss per share attributable 
to common stockholders—basic and 
diluted(1) 

Year ended December 31,  

2018 

2017 

2016 

2015 

2014 

(amounts in thousands, except share and per share data) 

 $ 

 7,619   $ 

 10,135   $ 

 14,220   $ 

 17,334   $ 

 —  

 64,905    
 33,809  
 98,714  
 (91,095) 
 3,310  
 (683) 
 (88,468) 
 (180) 
 (88,288)  $ 

 62,260  
 19,738  
 81,998  
 (71,863) 
 1,227  
 (62) 
 (70,698) 
 —  
 (70,698)  $ 

 42,249  
 13,270  
 55,519  
 (41,299) 
 976  
 182  
 (40,141) 
 52  
 (40,193)  $ 

 27,679  
 9,909  
 37,588  
 (20,254) 
 332  
 (9,750) 
 (29,672) 
 —  

 8,898  
 5,469  
 14,367  
 (14,367) 
 (1) 
 (1,949) 
    (16,317) 
 —  
 (29,672)  $   (16,317) 

 34  
 (88,254)  $ 

 (235) 
 (70,933)  $ 

 199  
 (39,994)  $ 

 (251) 

 —  
 (29,923)  $   (16,317) 

  $ 

  $ 

  $ 

 (88,288)  $ 

 (70,698)  $ 

 (40,193)  $ 

 (29,672)  $   (16,317) 

 —  

 —  

 —  

 —  

 —  

 —  

 (7,373) 

 (1,366) 

 (1,245) 

 —  

 $ 

 $ 

 (88,288)  $ 

 (70,698)  $ 

 (40,193)  $ 

 (38,290)  $   (17,683) 

 (2.75)  $ 

 (2.64)  $ 

 (1.59)  $ 

 (9.14)  $ 

 (27.83) 

     32,065,781  

   26,803,711  

   25,302,414  

   4,191,210  

   635,448  

124 

 
 
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
    
     
     
 
 
 
 
  
 
   
 
 
 
 
 
 
 
 
 
 
 
  
 
   
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
  
  
  
  
  
  
  
  
  
    
 
  
 
  
 
    
  
  
  
 
 
 
 
 
 
 
 
 
  
 
 
 
 
 
 
 
 
 
 
 
2018 

2017 

As of December 31,  

2016 
(in thousands) 

2015 

2014 

Consolidated balance sheet data: 
Cash, cash equivalents, and marketable debt 
securities 
Working capital(2) 
Total assets 
Redeemable convertible preferred stock 
Common stock and additional paid-in capital      
Total stockholders’ equity (deficit) 

  $ 

 155,806   $ 
 130,808  
 177,029  
 —  
 315,630  
 46,446  

 169,052   $ 
 155,893  
 184,477  
 —  
 295,051  
 134,051  

 174,418   $ 
 164,984  
 189,566  
 —  
 225,989  
 135,922  

 224,345   $ 
 171,963     
 229,457     
 —     
 219,147     
 169,074     

 7,035  
 5,884  
 11,497  
 21,979  
 1  
 (20,830) 

(1) See Statements of Operations Data and Note 2 to our financial statements for further details on the calculation of net loss per share, basic and 
diluted, attributable to common stockholders and the weighted-average number of shares used in the computation of the per share amounts.  

(2) We define working capital as current assets less current liabilities. See our financial statements for further details regarding our current assets and 
current liabilities. 

ITEM 7.  

    MANAGEMENT’S DISCUSSION AND ANALYSIS OF FINANCIAL CONDITION AND 

RESULTS OF OPERATIONS 

You should read the following discussion and analysis of our financial condition and results of operations 
together with our consolidated financial statements and related notes appearing elsewhere in this Annual Report on 
Form 10-K. In addition to historical information, this discussion and analysis contains forward-looking statements that 
involve risks, uncertainties and assumptions. Our actual results may differ materially from those anticipated in these 
forward-looking statements as a result of certain factors. We discuss factors that we believe could cause or contribute to 
these differences below and elsewhere in this report, including those set forth under Item 1A. “Risk Factors” and under 
“Forward-Looking Statements” in this Annual Report on Form 10-K.  

We are a clinical-stage gene therapy company focused on developing life-changing treatments for patients 

suffering from severe neurological diseases. We focus on neurological diseases where we believe an adeno-associated 
virus, or AAV, gene therapy approach that either increases or decreases the production of a specific protein can slow or 
reduce the symptoms experienced by patients, and therefore have a clinically meaningful impact. We have built a gene 
therapy platform that we believe positions us to be a leading company at the intersection of AAV gene therapy and 
severe neurological disease. Our gene therapy platform enables us to engineer, optimize, manufacture and deliver our 
AAV-based gene therapies that have the potential to provide durable efficacy following a single administration.  

Additionally, we are working to identify novel AAV capsids, which are the outer viral protein shells that 

enclose the genetic material of the virus payload. Our team of experts in the fields of AAV gene therapy and 
neuroscience first identifies and selects severe neurological diseases that are well-suited for treatment using AAV gene 
therapy. We then engineer and optimize AAV vectors for delivery of the virus payload to the targeted tissue or cells. Our 
manufacturing process employs an established system that we believe will enable production of high quality AAV 
vectors at commercial-scale. In addition to our capsid optimization efforts, we leverage novel delivery paradigms, 
established routes of administration, and advances in dosing techniques to optimize delivery of our AAV gene therapies 
to target tissues, regions and cell types that are critical to the disease of interest. We believe we can achieve this directly, 
with targeted infusions to discrete regions of the brain or spinal cord, or systemically, in conjunction with our novel 
capsids. 

Our business strategy focuses on discovering, developing, manufacturing and commercializing our gene therapy 
programs. As part of this strategy, we have developed core competencies specific to AAV gene therapy development and 
manufacturing and are beginning to build our commercial infrastructure. This business strategy also includes business 
development activities that may include in-licensing activities or partnering certain programs in specific geographies 
with collaborators, as we have demonstrated through our collaborations, including those with Sanofi Genzyme 

125 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
 
    
    
    
  
  
 
 
  
     
 
 
 
 
 
 
 
 
 
   
 
 
    
  
    
  
    
  
  
    
  
 
 
 
 
  
  
Corporation, which we refer to as Sanofi Genzyme, AbbVie Biotechnology Ltd and its affiliates, which we collectively 
refer to as AbbVie, and Neurocrine Biosciences, Inc., which we refer to as Neurocrine. Since our inception, our 
operations have focused on organizing and staffing our company, business planning, raising capital, establishing our 
intellectual property portfolio, determining which neurological diseases to pursue, advancing our product candidates 
including delivery and manufacturing, and conducting preclinical studies and clinical trials. We do not have any product 
candidates approved for sale and have not generated any revenue from product sales. We have funded our operations 
primarily through private placements of redeemable convertible preferred stock, public offerings of our common stock, 
our collaboration with Sanofi Genzyme, or the Sanofi Genzyme Collaboration, which commenced in February 2015 and 
our collaboration with AbbVie focusing on tau-related disease, or the AbbVie Tau Collaboration, which commenced in 
February 2018. Additionally, we recently entered into collaborations with Neurocrine, or the Neurocrine Collaboration, 
which we expect to commence in the first half of 2019, and with AbbVie focusing on pathological species of alpha-
synuclein, or the AbbVie Alpha-Synuclein Collaboration. 

On November 7, 2017, we completed the sale of 5,175,000 shares of common stock to the public at an offering 
price of $12.00 per share, including 675,000 shares of common stock issued upon the full exercise by the underwriters of 
their option to purchase additional shares, resulting in net proceeds of $58.0 million after deducting underwriting 
discounts, commissions, and offering expenses payable by us. 

Since inception, we have incurred significant operating losses. Our net losses were $88.3 million, $70.7 million, 

and $40.2 million for the years ended December 31, 2018, 2017, and 2016, respectively. As of December 31, 2018, we 
had an accumulated deficit of $269.1 million. We expect to continue to incur significant expenses and operating losses 
for the foreseeable future. We anticipate that our expenses will increase significantly in connection with our ongoing 
activities, as we: 

 

 

 

 

 

 

continue investing in our gene therapy platform to optimize vector engineering, manufacturing and dosing 
and delivery techniques;  

continue to advance our clinical candidate, VY-AADC, as a treatment for Parkinson’s disease through the 
ongoing Phase 1b clinical trial and our RESTORE-1 Phase 2 clinical trial;  

initiate additional preclinical studies and clinical trials for, and continue research and development of, our 
other programs;  

conduct joint research and development under our strategic collaborations for the research, development, 
and commercialization of certain of our pipeline programs; 

continue our process research and development activities, as well as establish our research-grade and 
commercial manufacturing capabilities;  

identify additional neurological diseases for treatment with our AAV gene therapies and develop additional 
programs or product candidates;  

  work to identify and optimize novel AAV capsids; 

 

 

develop, obtain and maintain regulatory clearances for devices to deliver our AAV gene therapies; 

seek marketing and regulatory approvals for VY-AADC or other product candidates or devices that arise 
from our programs that successfully complete clinical development; 

  maintain, expand, protect and enforce our intellectual property portfolio;  

 

identify, acquire or in-license other product candidates and technologies; 

126 

 

 

 

develop a sales, marketing and distribution infrastructure to commercialize any product candidates for 
which we may obtain marketing approval; 

expand our operational, financial and management systems and personnel, including personnel to support 
our clinical development, manufacturing and commercialization efforts and our operations as a public 
company;  

increase our product liability and clinical trial insurance coverage as we expand our clinical trials and 
commercialization efforts; and  

 

continue to operate as a public company. 

Financial Operations Overview 

Revenue 

To date, we have not generated any revenue from product sales and do not expect to generate any revenue from 

product sales for the foreseeable future. For the year ended December 31, 2018, we recognized $0.7 million of 
collaboration revenue from the Sanofi Genzyme Collaboration and $6.9 million of collaboration revenue from the 
AbbVie Tau Collaboration. For additional information about our revenue recognition policy related to the collaborations, 
see the section titled “—Critical Accounting Policies and Estimates—Revenue.” 

For the foreseeable future, we expect substantially all of our revenue will be generated from our collaboration 
agreements with Sanofi Genzyme, AbbVie, Neurocrine, and any other strategic relationships we may enter into. If our 
development efforts are successful, we may also generate revenue from product sales. 

Expenses 

Research and Development Expenses 

Research and development expenses consist primarily of costs incurred for our research activities, including our 

program discovery efforts, and the development of our programs and gene therapy platform, which include: 

 

 

 

 

 

 

employee-related expenses including salaries, benefits, and stock-based compensation expense; 

costs of funding research performed by third parties that conduct research and development, preclinical 
activities, manufacturing and production design on our behalf; 

the cost of purchasing lab supplies and non-capital equipment used in designing, developing and 
manufacturing preclinical study materials; 

consultant fees; 

facility costs including rent, depreciation and maintenance expenses; and 

fees for maintaining licenses under our third-party licensing agreements. 

Research and development costs are expensed as incurred. Costs for certain activities, such as manufacturing, 
preclinical studies, and clinical trials, are generally recognized based on an evaluation of the progress to completion of 
specific tasks using information and data provided to us by our vendors and collaborators. 

127 

At this time, we cannot reasonably estimate or know the nature, timing and estimated costs of the efforts that 
will be necessary to complete the development of our product candidates. We are also unable to predict when, if ever, 
material net cash inflows will commence from sales of our product candidates. This is due to the numerous risks and 
uncertainties associated with developing such product candidates, including the uncertainty of: 

 

 

 

 

 

 

 

 

successful enrollment in and completion of clinical trials; 

establishing an appropriate safety profile; 

establishing commercial manufacturing capabilities or making arrangements with third-party 
manufacturers; 

receipt of marketing approvals from applicable regulatory authorities; 

commercializing the product candidates, if and when approved, whether alone or in collaboration with 
others; 

obtaining and maintaining patent and trade secret protection and regulatory exclusivity for our product 
candidates; 

continued acceptable safety profiles of the products following approval; and 

retention of key research and development personnel. 

A change in the outcome of any of these variables with respect to the development of any of our product 

candidates would significantly change the costs, timing and viability associated with the development of that product 
candidate. 

Research and development activities are central to our business model. We expect research and development 

costs to increase significantly for the foreseeable future as our development programs progress, including as we continue 
to support the ongoing Phase 1b clinical trial and the separate Phase 1 trial exploring a posterior delivery approach and 
continue to enroll the RESTORE-1 Phase 2 clinical trial of VY-AADC as a treatment for Parkinson’s disease, and move 
our other product candidates into clinical trials. Additionally, we expect research and development costs associated with 
activities under our Strategic Collaborations to increase. There are numerous factors associated with the successful 
commercialization of any of our product candidates, including future trial design and various regulatory requirements, 
many of which cannot be determined with accuracy at this time based on our stage of development. Additionally, future 
commercial and regulatory factors beyond our control will impact our clinical development programs and plans. 

General and Administrative Expenses 

General and administrative expenses consist primarily of salaries and other related costs, including stock-based 

compensation, for personnel in executive, finance, accounting, business development, legal and human resource 
functions. Other significant costs include corporate facility costs not otherwise included in research and development 
expenses, legal fees related to patent and corporate matters and fees for accounting and consulting services. 

We anticipate that our general and administrative expenses will increase in the future to support continued 

research and development activities, including the RESTORE-1 Phase 2 clinical trial of VY-AADC, the expanded efforts 
in connection with our Strategic Collaborations, and the ongoing research and development activities and initiation of 
clinical trials for our other product candidates. These increases will likely include increased costs related to the hiring of 
additional personnel and fees to outside consultants. We also anticipate increased expenses associated with being a 
public company, including costs for audit, legal, regulatory, and tax-related services, director and officer insurance 
premiums, business development activities, and investor relations costs. 

128 

Other Income (Expense) 

Other income (expense) consists primarily of the gain (loss) on the equity securities investment in MRI 

Interventions.  

Critical Accounting Policies and Estimates 

Our management’s discussion and analysis of our consolidated financial condition and results of operations are 

based on our consolidated financial statements, which have been prepared in accordance with U.S. generally accepted 
accounting principles. The preparation of these consolidated financial statements requires us to make judgments and 
estimates that affect the reported amounts of assets, liabilities, revenues and expenses and the disclosure of contingent 
assets and liabilities in our consolidated financial statements. We base our estimates on historical experience, known 
trends and events, and various other factors that are believed to be reasonable under the circumstances. Actual results 
may differ from these estimates under different assumptions or conditions. On an ongoing basis, we evaluate our 
judgments and estimates in light of changes in circumstances, facts and experience. The effects of material revisions in 
estimates, if any, will be reflected in the financial statements prospectively from the date of change in estimates.  

While our significant accounting policies are described in more detail in the notes to our consolidated financial 
statements appearing elsewhere in this Annual Report on Form 10-K, we believe the following accounting policies used 
in the preparation of our financial statements require the most significant judgments and estimates.  

Revenue Recognition – ASC 606 

As of December 31, 2018, our revenue was generated from the Sanofi Genzyme Collaboration and the AbbVie 

Tau Collaboration. We recognize revenue in accordance with Financial Accounting Standards Board, or FASB, 
Accounting Standards Codification, or ASC, Topic 606 Revenue from Contracts with Customers, or ASC 606. Effective 
January 1, 2018, we adopted the provisions of ASC 606 using the modified retrospective transition method. Under this 
method, we recorded the cumulative effect of initially applying the new standard to all contracts as of the date of 
adoption.  

We enter into collaboration agreements which are within the scope of ASC 606, under which we license rights 

to certain of our product candidates and perform research and development services. The terms of these arrangements 
typically include payment of one or more of the following: non-refundable, upfront fees; reimbursement of research and 
development costs; development, regulatory and commercial milestone payments; and royalties on net sales of licensed 
products. 

Under ASC 606, an entity recognizes revenue when its customer obtains control of promised goods or services, 

in an amount that reflects the consideration which the entity expects to receive in exchange for those goods or services. 
To determine the appropriate amount of revenue to be recognized for arrangements determined to be within the scope of 
ASC 606, we perform the following five steps: (i) identification of the promised goods or services in the contract; (ii) 
determination of whether the promised goods or services are performance obligations including whether they are distinct 
in the context of the contract; (iii) measurement of the transaction price, including the constraint on variable 
consideration; (iv) allocation of the transaction price to the performance obligations; and (v) recognition of revenue when 
(or as) we satisfy each performance obligation. We only apply the five-step model to contracts when it is probable that 
we will collect consideration we are entitled to in exchange for the goods or services we transfer to the customer.  

The promised goods or services in our arrangement typically consist of license rights to our intellectual property 

or research and development services. We provide options to additional items in the contracts, which are accounted for 
as separate contracts when the customer elects to exercise such options, unless the option provides a material right to the 
customer. We evaluate the customer options for material rights, or options to acquire additional goods or services for free 
or at a discount. If the customer options are determined to represent a material right, the material right is recognized as a 
separate performance obligation at the outset of the arrangement. Performance obligations are promised goods or 
services in a contract to transfer a distinct good or service to the customer and are considered distinct when (i) the 

129 

customer can benefit from the good or service on its own or together with other readily available resources and (ii) the 
promised good or service is separately identifiable from other promises in the contract. In assessing whether promised 
goods or services are distinct, we consider factors such as the stage of development of the underlying intellectual 
property, the capabilities of the customer to develop the intellectual property on their own or whether the required 
expertise is readily available and whether the goods or services are integral or dependent to other goods or services in the 
contract. 

We estimate the transaction price based on the amount expected to be received for transferring the promised 
goods or services in the contract. The consideration may include fixed consideration or variable consideration. At the 
inception of each arrangement that includes variable consideration, we evaluate the amount of potential payment and the 
likelihood that the payments will be received. We utilize either the most likely amount method or expected amount 
method to estimate the amount expected to be received based on which method best predicts the amount expected to be 
received. The amount of variable consideration which is included in the transaction price may be constrained, and is 
included in the transaction price only to the extent that it is probable that a significant reversal in the amount of the 
cumulative revenue recognized will not occur in a future period.  

Our contracts often include development and regulatory milestone payments which are assessed under the most 

likely amount method and constrained if it is probable that a significant revenue reversal would occur. Milestone 
payments that are not within our control or the licensee’s control, such as regulatory approvals, are not considered 
probable of being achieved until those approvals are received. At the end of each reporting period, we re-evaluate the 
probability of achievement of such development milestones and any related constraint, and if necessary, adjust our 
estimate of the overall transaction price. Any such adjustments are recorded on a cumulative catch-up basis, which 
would affect collaboration revenues in the period of adjustment. To date, we have not recognized any consideration 
related to the achievement of development, regulatory, or commercial milestone revenue resulting from any of our 
collaboration arrangements. 

For arrangements that include sales-based royalties, including milestone payments based on the level of sales, 

and the license is deemed to be the predominant item to which the royalties relate, we recognize revenue at the later of (i) 
when the related sales occur, or (ii) when the performance obligation to which some or all of the royalty has been 
allocated has been satisfied (or partially satisfied). To date, we have not recognized any consideration related to sales-
based royalty revenue resulting from any of our collaboration arrangements. 

We allocate the transaction price based on the estimated stand-alone selling price of each of the performance 

obligations. We must develop assumptions that require judgment to determine the stand-alone selling price for each 
performance obligation identified in the contract. We utilize key assumptions to determine the stand-alone selling price 
for service obligations, which may include other comparable transactions, pricing considered in negotiating the 
transaction and the estimated costs. Additionally, in determining the standalone selling price for material rights, we 
utilize comparable transactions, industry standards for product development and clinical trial success probabilities and 
estimates of option exercise likelihood. Variable consideration is allocated specifically to one or more performance 
obligations in a contract when the terms of the variable consideration relate to the satisfaction of the performance 
obligation and the resulting amounts allocated are consistent with the amounts we would expect to receive for the 
satisfaction of each performance obligation.  

The consideration allocated to each performance obligation is recognized as revenue when control is transferred 

for the related goods or services. For performance obligations which consist of licenses and other promises, we utilize 
judgment to assess the nature of the combined performance obligation to determine whether the combined performance 
obligation is satisfied over time or at a point in time and, if over time, the appropriate method of measuring progress. We 
evaluate the measure of progress each reporting period and, if necessary, adjust the measure of performance and related 
revenue recognition.  

Upfront payments and fees are recorded as deferred revenue upon receipt or when due until we perform our 

obligations under these arrangements. Amounts are recorded as accounts receivable when our rights to consideration are 
unconditional.  

130 

Accrued Research and Development Expenses  

As part of the process of preparing our financial statements, we are required to estimate our accrued expenses as 
of each balance sheet date. This process involves reviewing open contracts and purchase orders, communicating with our 
personnel to identify services that have been performed on our behalf and estimating the level of service performed and 
the associated cost incurred for the service when we have not yet been invoiced or otherwise notified of the actual cost. 
The majority of our service providers invoice us monthly in arrears for services performed or when contractual 
milestones are met. We make estimates of our accrued expenses as of each balance sheet date based on facts and 
circumstances known to us at that time. We periodically confirm the accuracy of our estimates with the service providers 
and make adjustments if necessary. The significant estimates in our accrued research and development expenses include 
the costs incurred for services performed by our vendors in connection with research and development activities for 
which we have not yet been invoiced.  

We record our expenses related to research and development activities on our estimates of the services received 

and efforts expended pursuant to quotes and contracts with vendors that conduct research and development on our 
behalf. The financial terms of these agreements are subject to negotiation, vary from contract to contract and may result 
in uneven payment flows. There may be instances in which payments made to our vendors will exceed the level of 
services provided and result in a prepayment of the research and development expense. In accruing service fees, we 
estimate the time period over which services will be performed and the level of effort to be expended in each period. If 
the actual timing of the performance of services or the level of effort varies from our estimate, we adjust the accrual or 
prepaid accordingly. Non-refundable advance payments for goods and services that will be used in future research and 
development activities are expensed when the activity has been performed or when the goods have been received rather 
than when the payment is made.  

Although we do not expect our estimates to be materially different from amounts actually incurred, if our 

estimates of the status and timing of services performed differ from the actual status and timing of services performed, it 
could result in us reporting amounts that are too high or too low in any particular period. To date, there have been no 
material differences between our estimates of such expenses and the amounts actually incurred.  

Stock-based Compensation  

We account for our stock-based compensation awards in accordance with ASC Topic 718, Compensation—
Stock Compensation, or ASC 718. ASC 718 requires all stock-based payments to employees and directors, including 
grants of restricted stock and stock options, to be recognized as expense in the statements of operations based on their 
grant date fair values. Grants of restricted stock and stock options to other service providers, referred to as non-
employees, are required to be recognized as expense in the statements of operations based on their vesting date fair 
values. We estimate the fair value of options granted using the Black-Scholes option pricing model. We use the fair 
value of our common stock to determine the fair value of restricted stock awards.  

The Black-Scholes option pricing model requires inputs based on certain subjective assumptions, including 

(a) the expected stock price volatility, (b) the calculation of expected term of the award, (c) the risk-free interest rate and 
(d) expected dividends. Due to a lack of company-specific historical and implied volatility data, we have based the 
estimate of expected volatility on the historical volatility of a group of similar companies that are publicly traded, 
blended with the most recent period of historic volatility of our common stock. The historical volatility is calculated 
based on a period of time commensurate with the expected term assumption. The computation of expected volatility is 
based on the historical volatility of a representative group of companies with similar characteristics to us, including stage 
of product development and life science industry focus. We use the simplified method as prescribed by the SEC Staff 
Accounting Bulletin No. 107, Share-Based Payment, to calculate the expected term for options granted to employees and 
directors as we do not have sufficient historical exercise data to provide a reasonable basis upon which to estimate the 
expected term. For options granted to non-employees, we utilize the contractual term of the arrangement as the basis for 
the expected term assumption. The risk-free interest rate is based on a treasury instrument whose term is consistent with 

131 

the expected term of the stock options. The expected dividend yield is assumed to be zero as we have never paid 
dividends and do not have current plans to pay any dividends on common stock.  

The fair value of each option issued to employees and directors was estimated at the date of grant using the 

Black-Scholes option pricing model with the following weighted-average assumptions:  

Risk-free interest rate 
Expected dividend yield 
Expected term (in years) 
Expected volatility 

Year ended December 31,  

2018 

2017 

2016 

 2.8 %  
 — %  
 6.0  
 74.4 %  

 2.0 %  
 — %  
 6.0  
 73.7 %  

 1.5 %  
 — %  
 6.0  
 73.1 %  

The fair value of each option issued to non-employees was estimated at each vesting and reporting date using 

the Black-Scholes option pricing model. The reporting date fair value was determined using the following 
weighted-average assumptions:  

Risk-free interest rate 
Expected dividend yield 
Expected term (in years) 
Expected volatility 

2018 

As of December 31,  
2017 

 2.6 %   
 — %   
 7.6  
 73.2 %   

 2.4 %   
 — %   
 8.5  
 76.2 %   

2016 

 2.1 %   
 — %   
 9.1  
 83.3 %   

We expense the fair value of our stock-based compensation awards to employees and directors on a straight-line 

basis over the associated service period, which is generally the period in which the related services are received. Stock-
based compensation awards to non-employees are adjusted through stock-based compensation expense at each reporting 
period end to reflect the current fair value of such awards and are expensed on a straight-line basis.  

We record the expense for stock-based compensation awards subject to performance-based milestone vesting 

over the remaining service period when management determines that achievement of the milestone is probable. 
Management evaluates when the achievement of a performance-based milestone is probable based on the expected 
satisfaction of the performance conditions as of the reporting date. Management concluded that the achievement of the 
performance milestone for one of the three performance-based awards had been met during 2016. Accordingly, stock-
based compensation expense in the amount of $0.3 million, $1.4 million, and $1.1 million was recorded in the years 
ended December 31, 2018, 2017, and 2016, respectively.  

Stock-based compensation totaled approximately $15.7 million, $9.1 million, and $6.3 million the years ended 

December 31, 2018, 2017, and 2016 respectively. As of December 31, 2018, we had $0.1 million and $26.1 million of 
unrecognized compensation expense related to restricted stock awards and stock option awards, respectively, which are 
expected to be recognized over weighted-average remaining vesting periods at the time the milestones have been met 
and approximately 2.78 years, respectively. We expect the impact of our stock-based compensation expense for 
restricted stock and stock options granted to employees, directors and other service providers to grow in future periods 
due to the potential increases in the value of our common stock and headcount. 

132 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
    
 
 
 
  
  
  
 
 
 
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
     
 
 
 
  
  
  
 
 
 
  
 
 
Results of Operations 

Comparison of the years ended December 31, 2018 and 2017: 

The following table summarizes our results of operations for the years ended December 31, 2018 and 2017, 

respectively, together with the changes in those items in dollars: 

Collaboration revenue 
Operating expenses: 

Research and development 
General and administrative 
Total operating expenses 

Other income: 

Interest income 
Other expense 

Total other income 

Loss before income taxes 
Income tax provision 
Net loss 

Collaboration Revenue 

Year ended  
December 31,  

2018 

2017 
(in thousands) 

Change 

  $ 

 7,619     $ 

 10,135     $ 

 (2,516) 

 64,905  
 33,809  
 98,714  

 62,260  
 19,738  
 81,998  

 3,310  
 (683) 
 2,627  
 (88,468) 
 (180) 
 (88,288)  $ 

 1,227  
 (62) 
 1,165  
 (70,698)
 —  
 (70,698)  $ 

  $ 

 2,645  
 14,071  
 16,716  

 2,083  
 (621) 
 1,462  

 (180) 
 (180) 

Collaboration revenue was $7.6 million for the year ended December 31, 2018, and $10.1 million for the year 

ended December 31, 2017. Collaboration revenue included amounts related to the Sanofi Genzyme Collaboration in 
addition to research services related to the AbbVie Tau Collaboration, which commenced in February 2018. The 
decrease in revenue is primarily related the recognition of amounts related to our Parkinson’s program under the Sanofi 
Genzyme Collaboration, or the PD Option, in the year ended December 31, 2017, the adoption of ASC 606 as of January 
1, 2018, which resulted in the use of a proportional performance method in 2018 as compared to the use of straight-line 
method in 2017, as well as changes in estimates of costs to reach proof of principle on our Huntington’s disease and 
Friedreich’s ataxia programs. These reductions were offset by revenue recognized on research services performed under 
the AbbVie Tau Collaboration.  

We recognized $5.5 million of revenue in the year ended December 31, 2017 related to the portion of the 

upfront consideration under the Sanofi Genzyme Collaboration which had been allocated to Sanofi Genzyme’s rights 
related to the PD Option. In the year ended December 31, 2017, Sanofi Genzyme decided not to exercise the PD Option, 
and we recognized all amounts allocated to their option at that time. Additionally, effective January 1, 2018, we adopted 
the provisions of ASC 606 for revenue recognition.  

Research and Development Expense 

Research and development expense increased by $2.6 million from $62.3 million for the year ended 

December 31, 2017 to $64.9 million for the year ended December 31, 2018. The following table summarizes our 
research and development expenses, for the years ended December 31, 2018 and 2017: 

External research and development expenses 
Employee and contractor related expenses 

  $ 

 28,890     $ 
 26,075  

 33,816   $ 
 20,919  

 (4,926) 
 5,156  

Year ended  
December 31,  

2018 

2017 
(in thousands) 

      Change 

133 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
     
     
 
 
 
 
   
 
 
 
 
 
 
 
 
  
  
  
  
  
  
  
  
  
   
 
 
 
 
 
 
 
 
  
  
  
  
  
  
  
  
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
     
 
 
 
 
    
  
  
Facility, technology, and other expenses 
License fees 

Total research and development expenses 

$ 

 9,305  
 635  
 64,905   $ 

 6,705  
 820  
 62,260   $ 

 2,600  
 (185) 
 2,645  

The change in research and development expense for the year ended December 31, 2018 was primarily 

attributable to the following: 

 

 

 

approximately $5.2 million for increased research and development employee compensation costs as we 
continue to increase research and development headcount to support our ongoing development of our 
clinical and pre-clinical programs and platform;  

approximately $2.6 million for increased facility and other costs including rent, depreciation, maintenance 
and other expenses due to the additional space leased;  

offset by approximately $4.8 million for decreased costs of funding research performed by third parties that 
conduct research and development. This reduction includes a decrease in preclinical activities, offset by an 
increase in clinical and manufacturing activities; further decreased by a reduction of approximately 
$0.1 million attributable to in-kind research and development services incurred by Sanofi Genzyme and 
provided to us under the Sanofi Genzyme Collaboration; and approximately $0.2 million related to 
decreased licensing costs. 

General and Administrative Expense 

General and administrative expense increased by $14.1 million from $19.7 million for the year ended 
December 31, 2017 to $33.8 million for the year ended December 31, 2018. The change in general and administrative 
expense was primarily attributable to the following: 

 

 

 

approximately $9.6 million for increased employee compensation costs as we increase our administrative 
headcount to support our growing business. The increase included the recognition of $5.4 million of stock-
based compensation related to the retirement agreement with our former Chief Executive Officer, Dr. 
Steven Paul;  

approximately $2.7 million for increased legal costs for general, business development, and intellectual 
property support; and 

approximately $1.7 million for increased facility and other costs including rent, depreciation, maintenance 
and other expenses. 

Other Income, Net 

Other income of approximately $2.6 million and $1.2 million was recognized in the years ended 
December 31, 2018 and 2017 related to interest income on marketable securities balances offset by losses on our 
warrants to purchase shares of common stock and our common stock investment in MRI Interventions, Inc., or MRIC. 
The increase in other income is largely a result of higher cash balances year over year.  

Income Tax 

We recorded an income tax provision of $0.2 million related to our alternative minimum tax, or AMT, liability 

resulting in an income tax payable of $0.1 million for the year ended December 31, 2017. The payable was due to the 
recognition of deferred revenue related to the Sanofi Genzyme Collaboration for income tax purposes. There was no 
income tax payable for the year ended December 31, 2018. 

134 

    
  
  
    
  
  
 
 
Comparison of year ended December 31, 2017 and 2016:  

The following table summarizes our results of operations for the year ended December 31, 2017 and 2016, 

respectively, together with the changes in those items in dollars:  

Collaboration revenue 
Operating expenses: 

Research and development 
General and administrative 
Total operating expenses 

Other income: 

Interest income 
Other (expense) income, net 

Total other income 

Loss before income taxes 
Income tax provision 
Net loss 

Collaboration Revenue  

Year ended  
December 31,  

2017 

2016 
(in thousands) 

Change 

  $ 

 10,135     $ 

 14,220     $ 

 (4,085)  

 62,260  
 19,738  
 81,998  

 42,249  
 13,270  
 55,519  

 1,227  
 (62) 
 1,165  
 (70,698) 
 —  
 (70,698)  $ 

 976  
 182  
 1,158  
 (40,141)  
 52  
 (40,193)   $ 

  $ 

 20,011  
 6,468  
 26,479  

 251  
 (244)  
 7  
 (30,557)  
 (52)  
 (52)  

Collaboration revenue was $10.1 million for the year ended December 31, 2017, and $14.2 million for the year 
ended December 31, 2016, all of which related to the Sanofi Genzyme Collaboration. In October 2017, Sanofi Genzyme 
decided not to exercise its option for the ex-U.S. rights to the PD Option. Therefore, in the year ended December 31, 
2017, we recognized $5.5 million of revenue related to the portion of the upfront consideration which had been allocated 
to the PD Option. In addition, revenue recognized during the year ended December 31, 2017 and 2016 includes amounts 
recognized related to consideration allocated to research and development services for various programs under the Sanofi 
Genzyme Collaboration. During 2017 we reassessed the estimated period of performance for each of the units of 
accounting and determined that the estimated period would be extended for two units of accounting. During 2016 we 
deprioritized the development of VY-SMN101. These adjustments were made on a prospective basis and resulted in 
decreases in revenue recognized by $2.1 million and $9.5 million, respectively, for the year ended December 31, 2017. 

Research and Development Expense  

Research and development expense increased by $20.0 million from $42.2 million for the year ended 

December 31, 2016 to $62.2 million for the year ended December 31, 2017. The following table summarizes our 
research and development expenses, for the year ended December 31, 2017 and 2016, respectively:  

External research and development expenses 
Employee and contractor related expenses 
Facility, technology, and other expenses 
License fees 

  $ 

Total research and development expenses 

$ 

Year ended  
December 31,  

2017 

2016 
(in thousands) 

      Change 

 33,816     $ 
 20,919  
 6,705  
 820  
 62,260   $ 

 20,413   $ 
 15,530  
 4,553  
 1,753  
 42,249   $ 

 13,403 
 5,389 
 2,152 
 (933)
 20,011 

135 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
     
     
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
  
  
  
  
  
  
  
  
 
 
 
 
 
 
 
 
 
 
  
  
  
  
  
  
  
  
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
     
 
 
    
  
 
    
  
  
    
  
  
The change in research and development expense was primarily attributable to research and development, and 

included the following: 

 

 

 

approximately $14.5 million for increased costs of funding research performed by third parties that conduct 
research and development, preclinical and clinical activities and manufacturing and production design on 
our behalf and increased purchases of lab supplies and non-capital equipment used in designing, developing 
and manufacturing preclinical study materials, and an additional expense of approximately $1.1 million 
attributable to in-kind research and development services incurred by Sanofi Genzyme and provided to us 
under the Sanofi Genzyme Collaboration; 

approximately $5.4 million for increased research and development employee-related and consultant 
compensation costs;  

approximately $2.2 million for increased facility and other costs including rent, depreciation, maintenance 
and other expenses due to the additional leased space dedicated to research and development efforts at 64 
Sidney Street and 75 Sidney Street; and  

 

offset by approximately $0.9 million for decreased intellectual property and license fees.  

General and Administrative Expense  

General and administrative expense increased by $6.4 million from $13.3 million for the year ended 
December 31, 2016 to $19.7 million for the year ended December 31, 2017. The change in general and administrative 
expense was primarily attributable to the following: 

 

 

approximately $3.5 million for increased employee compensation cost due to increases in headcount and 
stock-based compensation; 

approximately $0.8 million for increased facility and other costs including rent, depreciation, maintenance 
and other expenses; and 

 

approximately $2.1 million for increased legal and intellectual property expenses. 

Other Income, Net  

Other income of approximately $1.2 million was recognized in the years ended December 31, 2017 and 2016 

related to interest income on marketable securities balances offset by losses on our warrants to purchase shares of 
common stock of MRIC. 

Income Tax 

We recorded an income tax provision of $0.2 million related to our alternative minimum tax, or AMT, liability 

resulting in an income tax payable of $0.1 million for the year ended December 31, 2016. The payable was due to the 
recognition of deferred revenue related to the Sanofi Genzyme Collaboration for income tax purposes. There was no 
income tax payable for the year ended December 31, 2017. Our overall income tax provision was offset by an income tax 
benefit recorded to continuing operations of $0.1 million associated with the recognition of the corresponding income tax 
associated with unrealized gains included in other comprehensive income. The net tax effect resulted in an overall 
income tax provision recorded to continuing operations of $0.1 million. We recorded no income tax provision (benefit) 
for the year ended December 31, 2017. 

136 

Liquidity and Capital Resources 

Sources of Liquidity 

We have funded our operations primarily through private placements of redeemable convertible preferred stock, 

public offerings of our common stock, the Sanofi Genzyme Collaboration which commenced in February 2015, and the 
AbbVie Tau Collaboration which commenced in February 2018. Additionally, in January 2019, we entered into a 
Collaboration and License Agreement with Neurocrine Biosciences, Inc., or the Neurocrine Collaboration, which we 
expect to commence in the first half of 2019, and in February 2019, we entered into a Collaboration Agreement with 
AbbVie Ireland Unlimited Company, or the AbbVie Alpha-Synuclein Collaboration. Under the Neurocrine 
Collaboration, we will receive an upfront payment of $165.0 million, including a $50.0 million investment through the 
purchase of 4,179,728 shares of our common stock in a private placement, and under the AbbVie Alpha-Synuclein 
Collaboration, we will receive an upfront payment of $65.0 million.  

On November 16, 2015, we closed our IPO whereby we sold 5,750,000 shares of common stock, at a public 

offering price of $14.00 per share, including 750,000 shares of common stock issued upon the full exercise by the 
underwriters of their option to purchase additional shares, resulting in net proceeds to us of $72.9 million after deducting 
underwriting discounts and commissions and offering expenses payable by us. On November 7, 2017, we sold 5,175,000 
shares of common stock to the public at an offering price of $12.00 per share, including 675,000 shares of common stock 
issued upon the full exercise by the underwriters of their option to purchase additional shares, resulting in net proceeds to 
us of $58.0 million after deducting underwriting discounts, commissions, and offering expenses payable by us.  

As of December 31, 2018, we had cash, cash equivalents, and marketable debt securities of $155.8 million. 

Cash Flows 

The following table provides information regarding our cash flows for the years ended December 31, 2018, 

2017, and 2016: 

Net cash (used in) provided by: 

Operating activities 
Investing activities 
Financing activities 

  $ 

Net increase (decrease) in cash and cash equivalents   $ 

Cash Flows from Operating Activities 

2018 

Year ended  
December 31,  
2017 
(in thousands) 

2016 

 (15,887)  $ 
 26,467  
 4,749  
 15,329   $ 

 (61,350)  $ 
 (3,681) 
 59,920  
 (5,111)  $ 

 (42,482)
 47,721 
 514 
 5,753 

Net cash used in operating activities was $15.9 million during the year ended December 31, 2018 compared to 

$61.4 million during the year ended December 31, 2017. The decrease in cash used for operating activities was primarily 
due to the receipt of $69.0 million related to the AbbVie Tau Collaboration in 2018, offset by increases in operating 
expenses due to increased research and development activities, as well as higher general and administrative expenses as a 
result of a higher headcount, legal fees, and other costs year over year. The decrease in cash used in operating activities 
was also offset by an increase in prepaid expenses and other current assets as well as a decrease in accrued expenses.  

 Net cash used in operating activities was $61.4 million during the year ended December 31, 2017 compared to 

$42.5 million of cash used in operating activities during the year ended December 31, 2016. The increase in cash used 
for operating activities was primarily due to an increase in operating expenses. The increase in operating expenses is 
primarily due to increased research and development activities as we advance our clinical and preclinical programs, as 
well as higher general and administrative expenses to support the increased research and development operations. 

137 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
    
     
 
 
    
 
 
  
 
 
 
 
 
 
 
 
 
  
  
  
 
  
  
  
Cash Flows from Investing Activities 

Net cash provided by investing activities was $26.5 million during the year ended December 31, 2018. The cash 
provided by investing activities for the year ended December 31, 2018 was primarily due to proceeds from maturities of 
marketable securities of $364.0 million offset by purchase of marketable securities of $333.2 million and purchases of 
property and equipment of $4.9 million.  

Net cash used in investing activities was $3.7 million during the year ended December 31, 2017. The cash used 

in investing activities for the year ended December 31, 2017 was due to purchases of property and equipment of $4.0 
million, offset by net proceeds from maturities and purchases of marketable securities of $0.3 million. 

Net cash provided by investing activities was $47.7 million during the year ended December 31, 2016. The cash 

provided by investing activities for the year ended December 31, 2016 was due to proceeds from maturities of 
marketable securities of $165.1 million, partially offset by purchases of marketable securities of $112.4 million and 
purchases of property and equipment of $5.0 million.  

Cash Flows from Financing Activities 

Net cash provided by financing activities was $4.7 million during the year ended December 31, 2018 related to 

proceeds from exercises of stock options, and purchases by our employees of our common stock under our Employee 
Stock Purchase Plan.  

Net cash provided by financing activities was $59.9 million during the year ended December 31, 2017 and was 

largely driven by the $58.0 million of net proceeds from our sale of common stock in November 2017 and proceeds from 
exercises of stock options. 

Net cash provided by financing activities was $0.5 million during the year ended December 31, 2016 and 

consisted of proceeds from exercises of stock options. 

Funding Requirements 

We expect our expenses to increase in connection with our ongoing activities, particularly as we continue the 

research and development of, continue or initiate clinical trials of, and seek marketing approval for, our product 
candidates. In addition, if we obtain marketing approval for any of our product candidates, we expect to incur significant 
expenses related to program sales, marketing, manufacturing and distribution to the extent that such sales, marketing and 
distribution are not the responsibility of potential collaborators. Furthermore, we expect to incur additional costs 
associated with operating as a public company. Accordingly, we will need to obtain substantial additional funding in 
connection with our continuing operations. If we are unable to raise capital or enter into business development 
transactions when needed or on acceptable terms, we could be forced to delay, reduce or eliminate our research and 
development programs or any future commercialization efforts. 

Based upon our current operating plan, we expect that our existing cash, cash equivalents, and marketable debt 

securities, as well as amounts expected from the upfront payment and expected reimbursement of development costs 
from the Neurocrine Collaboration Agreement and the upfront payment from the AbbVie Alpha-Synuclein Collaboration 
Agreement entered into in 2019, will enable us to fund our operating expenses and capital expenditure requirements into 
mid-2022. Our future capital requirements will depend on many factors, including: 

 

the scope, progress, results, and costs of product discovery, preclinical studies and clinical trials for our 
product candidates and any required companion devices; 

 

the scope, progress, results, costs, prioritization, and number of our research and development programs; 

138 

 

 

 

 

 

 

 

 

 

 

the progress and status of our strategic collaborations, including any research and development costs for 
which we are responsible, the potential exercise by our collaboration partners of options to develop or 
license certain products and product candidates, and our potential receipt of future milestone payments and 
royalties from our collaboration partners; 

the extent to which we are obligated to reimburse, or entitled to reimbursement of, preclinical development 
and clinical trial costs, or the achievement of milestones or occurrence of other developments that trigger 
payments, under any other collaboration agreement to which we might become a party; 

the costs, timing and outcome of regulatory review of our product candidates; 

our ability to establish and maintain collaboration, distribution, or other marketing arrangements for our 
product candidates on favorable terms, if at all; 

the costs and timing of preparing, filing and prosecuting patent applications, maintaining and enforcing our 
intellectual property rights and defending intellectual property-related claims; 

the extent to which we acquire or in-license other product candidates and technologies or acquire or invest 
in other businesses, such as our investment in MRIC; 

the costs related to evaluating possible alternative devices that may be useful in the delivery of our product 
candidates, including our ongoing development of V-TAG; 

the costs of securing manufacturing arrangements for pre-commercial and commercial production;  

the level of product sales from any product candidates for which we obtain marketing approval in the 
future; and 

the costs of establishing or contracting for sales, manufacturing, marketing, distribution, and other 
commercialization capabilities if we obtain regulatory approvals to market our product candidates. 

Identifying potential product candidates and conducting preclinical studies and clinical trials is a 

time-consuming, expensive and uncertain process that takes years to complete. We may never generate the necessary 
data or results required to obtain marketing approval and achieve product sales. In addition, our product candidates, if 
approved, may not achieve commercial success. Our product revenues, if any, and any commercial milestones or royalty 
payments under our collaboration agreements, will be derived from sales of products that may not be commercially 
available for many years, if at all. Accordingly, we will need to continue to rely on additional financing and business 
development transactions to achieve our business objectives. Adequate additional financing may not be available to us 
on acceptable terms, or at all. 

Until such time, if ever, as we can generate product revenues sufficient to achieve profitability, we expect to 

finance our cash needs through a combination of equity offerings, debt financings, collaborations, strategic alliances and 
licensing arrangements. To the extent that we raise additional capital through the sale of equity or equity-linked 
securities, including convertible debt, our stockholders’ ownership interests will be diluted, and the terms of these 
securities may include liquidation or other preferences that adversely affect our existing stockholders’ rights as holders 
of our common stock. Debt financing and preferred equity financing, if available, may involve agreements that include 
covenants limiting or restricting our ability to take specific actions, such as incurring additional debt, obtaining 
additional capital, acquiring or divesting businesses, making capital expenditures or declaring dividends. 

If we raise additional funds through collaborations, strategic alliances or licensing arrangements with third 
parties, we may have to relinquish valuable rights to our technologies, future revenue streams, research programs or 
product candidates or to grant licenses on terms that may not be favorable to us. If we are unable to raise additional funds 

139 

through equity or debt financings when needed, we may be required to delay, limit, reduce or terminate our product 
development or future commercialization efforts or grant rights to develop and market product candidates that we would 
otherwise prefer to develop and market ourselves. 

Contractual Obligations 

The following table summarizes our significant contractual obligations as of payment due date by period at 

December 31, 2018: 

      Total 

    Less Than 
     1 Year 

     More than  
     1 to 3 Years     3 to 5 Years       5 Years   
(in thousands) 

Operating lease commitments(1) 

  $ 40,844   $  4,325   $ 

 9,594   $  10,142   $ 16,783  

(1)  We lease office space at 75 Sidney Street and 64 Sidney Street in Cambridge, Massachusetts under non-cancelable 

operating leases that expire in November 2026. 

In February 2018, we executed a second amendment for additional space located at 75 Sidney Street in 
Cambridge, Massachusetts, concurrent to the existing leases with terms going through December 2024. In June 2018, we 
executed a third amendment for additional space located at 75 Sidney Street, including an extension to the term through 
November 2026. Additionally, we executed an amendment to the lease at 64 Sidney Street to extend the term through 
November 2026.  

We enter into agreements in the normal course of business with clinical research organizations, contract 
manufacturing organizations, and institutions to license intellectual property. We have not included these future 
payments in the table of contractual obligations above since the contracts are cancelable at any time by us, generally 
upon 30 to 90 days prior written notice. 

Our agreements to license intellectual property include potential milestone payments that are dependent upon 

the development of products using the intellectual property licensed under the agreements and contingent upon the 
achievement of clinical trial or regulatory approval milestones. We may also be required to pay annual maintenance fees 
or minimum amounts payable ranging from low-four digits to low five-digits depending upon the terms of the applicable 
agreement. 

In February 2015, we entered into an agreement in connection with the Sanofi Genzyme Collaboration, or the 

Sanofi Genzyme Collaboration Agreement, pursuant to which we granted Sanofi Genzyme an exclusive option to 
license, develop, and commercialize (i) ex-U.S. rights to several programs including our programs for Parkinson’s 
disease and Friedreich’s ataxia, (ii) ex-U.S. rights and U.S. co-commercialization rights to our program for Huntington’s 
disease, and (iii) worldwide rights to our program for Spinal Muscular Atrophy. Sanofi Genzyme’s option for each 
program is triggered following the completion of the first proof of principle human clinical study on a program-by-
program basis. 

Subject to specified exceptions, we are solely responsible for all costs incurred in connection with the 
development of each of the covered programs prior to Sanofi Genzyme’s exercise of its option for such program. Upon 
Sanofi Genzyme’s exercise of its option to license most of the programs pursuant to our collaboration, we will have sole 
responsibility for the development and commercialization of such product in the United States, and Sanofi Genzyme 
shall have sole responsibility for development and commercialization of such product in the rest of the world. In the case 
of our Huntington’s disease program, if Sanofi Genzyme exercises its co-commercialization rights, it will also be 
responsible for commercialization activities related to the Huntington’s disease product in the United States. In the case 
of our Spinal Muscular Atrophy product, if Sanofi Genzyme exercises its option, it will be responsible for all 
development costs occurring following the option exercise date and all worldwide commercialization activities. 

In February 2018, we entered into an agreement in connection with the AbbVie Tau Collaboration, or the 

AbbVie Tau Collaboration Agreement, pursuant to which we granted AbbVie an exclusive option to license, develop, 

140 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
and commercialize AAV and other virus-based gene therapy products for the treatment of neurodegenerative diseases 
related to defective or excess aggregation of tau protein in the human brain, including Alzheimer’s disease. The 
collaboration is comprised of a research period, a development period, and an exclusive license option. During the 
research period, we are obligated to use diligent efforts to conduct antibody engineering and other research activities to 
create research compounds and to develop product candidates. We will be solely responsible for the costs and expenses 
during the research period. During a specified portion of the research period, AbbVie may exercise one or more of its 
exclusive development options to select up to a total of three research compounds and their corresponding product 
candidates to proceed to the development period. 

During the development period, we are obligated to use diligent efforts to conduct development activities, 

including IND-enabling and Phase 1 clinical trial activities, for such selected research compounds and their 
corresponding product candidates. We will be solely responsible for the costs and expenses during the development 
period. During a specified portion of the development period, AbbVie may exercise its exclusive license option to further 
develop and commercialize all of the research compounds and corresponding product candidates. Upon the exercise of 
its license option, AbbVie would be solely responsible for all development and commercialization activities relating to 
the licensed compounds and corresponding licensed products, subject to certain exceptions. We also may elect to share 
in AbbVie’s development costs relating to a licensed product on an indication-by-indication basis in exchange for a 
specified increase in royalties. 

In January 2019, we entered into an agreement in connection with the Neurocrine Collaboration, or the 
Neurocrine Collaboration Agreement, for the development and commercialization of our VY-AADC program, our gene 
therapy program for the treatment of Friedreich’s ataxia including VY-FXN01, and two programs to be determined by us 
and Neurocrine at a later date, which we refer to herein as the Discovery Programs. Under the terms of the Neurocrine 
Collaboration Agreement, Neurocrine has agreed to pay us an upfront payment of $165.0 million, along with funding for 
ongoing development of each program in accordance with an agreed budget, and up to $1.7 billion in potential 
development, regulatory, and commercial milestone payments. Under the terms of the Neurocrine Collaboration 
Agreement, Neurocrine will fund the clinical development of the Phase 2 clinical program for VY-AADC for 
Parkinson’s disease. After our receipt of topline data of the RESTORE-1 Phase 2 trial, we have the option to either (1) 
co-commercialize VY-AADC in the United States under a 50/50 cost- and profit-sharing arrangement and receive 
milestones and royalties based on ex-U.S. sales, or (2) grant Neurocrine full global commercial rights in exchange for 
milestone payments and royalties based on global sales. Under the terms of the Neurocrine Collaboration Agreement, 
Neurocrine will fund the development for VY-FXN01 for Friedreich’s ataxia through the Phase 1 clinical trial. After the 
data readout of the Phase 1 trial, we have the option to either (1) co-commercialize VY-FXN01 with Neurocrine in the 
U.S. under a 60/40 cost- and profit-sharing arrangement, or (2) grant Neurocrine full worldwide commercial rights in 
exchange for milestone payments and royalties based on global sales, subject to Sanofi Genzyme’s option to 
commercialize the FA Program in countries outside the United States. Under the terms of the agreement for the two 
Discovery Programs, Neurocrine will fund the development of those programs and we have the right to earn milestone 
payments and royalties based on global sales. Sanofi Genzyme retains an option for ex-U.S. rights to VY-FXN01 
following the data readout of the Phase 1 trial; however, if Sanofi Genzyme declines its option, the ex-U.S. rights to VY-
FXN01 would revert to us and be included under the territories licensed to Neurocrine. Under the terms of the 
Neurocrine Collaboration Agreement, Neurocrine will fund the development of the two Development Programs, and we 
will have the right to earn milestone payments and royalties on global sales. 

In February 2019, we entered into an exclusive collaboration and option agreement with AbbVie to develop and 

commercialize vectorized antibodies directed against pathological species of alpha-synuclein for the potential treatment 
of Parkinson’s disease and other diseases characterized by the abnormal accumulation of misfolded alpha-synuclein 
protein, or synucleinopathies. Under the terms of this agreement, we will receive an upfront payment of $65.0 million 
and may receive future option fees, development, regulatory, and commercial milestone payments, and royalties. Under 
the terms of the agreement, we and AbbVie have agreed to collaborate on the research and development of specified 
vectorized antibody compounds, or Research Compound, comprised of an AAV or other viral capsid and a virus vector 
genome that encodes one or more antibodies that target and bind to the alpha-synuclein protein. The collaboration is 
comprised of a research period and a development period. We are obligated to conduct research activities directed to 
constructing one or more virus vectors that encode antibodies designated by AbbVie. We are obligated to use diligent 
efforts to conduct antibody engineering and other research activities to create Research Compounds and to develop 
product candidates containing or comprised of such Research Compounds, Product Candidates. We are solely 

141 

responsible for the costs and expenses during the Research Period. During a specified portion of the Research Period, 
AbbVie may exercise one or more of its exclusive development options to select up to a total of four Research 
Compounds and their corresponding Product Candidates to proceed to the Development Period, after which AbbVie may 
exercise its option to license such Product Candidates following Phase 1 results, for which we may earn up to $245.0 
million in option exercise payments in aggregate. In addition to the upfront payment and the potential option exercise 
payments, we are eligible to receive up to $727.5 million in development and regulatory milestones for each Licensed 
Compound. We are also eligible to receive tiered, escalating royalties, in the mid-single-digit percentage range on 
aggregate net sales of Licensed Products on a Licensed Compound by Licensed Compound basis, as well as up to $500.0 
million in commercial milestones based on aggregate annual net sales thresholds of Licensed Products. The royalties are 
subject to potential reductions for biosimilar market penetration, patent claim expiration, and other provisions, subject to 
specified limits. Subject to certain exceptions, each of AbbVie and the Company has agreed to be financially responsible 
for all payments owed to a third party with which it has contracted for any use of in-licensed intellectual property under 
the Collaboration Agreement.  

Off-Balance Sheet Arrangements 

We did not have, during the periods presented, and we do not currently have, any off-balance sheet 

arrangements, as defined under applicable Securities and Exchange Commission rules. 

JOBS Act 

In April 2012, the JOBS Act, was enacted. Section 107 of the JOBS Act provides that an emerging growth company, 

or EGC, can take advantage of the extended transition period provided in Section 7(a)(2)(B) of the Securities Act of 1933, as 
amended, or the Securities Act, for complying with new or revised accounting standards. Thus, an EGC can delay the adoption 
of certain accounting standards until those standards would otherwise apply to private companies. 

Subject to certain conditions, as an EGC, we intend to rely on certain of these exemptions, including without 

limitation, (i) providing an auditor’s attestation report on our system of internal controls over financial reporting pursuant 
to Section 404(b) of the Sarbanes‑Oxley Act and (ii) complying with any requirement that may be adopted by the Public 
Company Accounting Oversight Board, or PCAOB, regarding mandatory audit firm rotation or a supplement to the 
auditor’s report providing additional information about the audit and the financial statements, known as the auditor 
discussion and analysis. We will remain an EGC until the earlier of (i) the last day of the fiscal year in which we have 
total annual gross revenues of $1.07 billion or more; (ii) December 31, 2020; (iii) the date on which we have issued more 
than $1.0 billion in nonconvertible debt during the previous three years; or (iv) the date on which we are deemed to be a 
large accelerated filer under the rules of the SEC. 

ITEM 7A.  

   QUANTITATIVE AND QUALITATIVE DISCLOSURES ABOUT MARKET RISK 

We are exposed to market risk related to changes in interest rates. We have policies requiring us to invest in 

high-quality issuers, limit our exposure to any individual issuer, and ensure adequate liquidity. Our primary exposure to 
market risk is interest rate sensitivity, which is affected by changes in the general level of U.S. interest rates, particularly 
because our investments, including cash equivalents, are in the form of money market fund and marketable securities and 
are invested in U.S. Treasury and U.S. government agency obligations. Due to the short-term duration of our investment 
portfolio and the low risk profile of our investments, an immediate 100 basis point change in interest rates would not 
have had a material effect on the fair market value of our portfolio. 

We are not currently exposed to market risk related to changes in foreign currency exchange rates; however, we 
may contract with vendors that are located in Asia and Europe in the future and may be subject to fluctuations in foreign 
currency rates at that time. 

Inflation generally affects us by increasing our cost of labor and clinical trial costs. We do not believe that 

inflation had a material effect on our business, financial condition, or results of operations during the year ended 
December 31, 2018. 

142 

 
  
ITEM 8.  

    FINANCIAL STATEMENTS AND SUPPLEMENTARY DATA 

The financial statements required to be filed pursuant to this Item 8 are appended to this report. An index of 

those financial statements is found in Item 15. 

ITEM 9.  

    CHANGES IN AND DISAGREEMENTS WITH ACCOUNTANTS ON ACCOUNTING AND 

FINANCIAL DISCLOSURE 

None. 

ITEM 9A.  

 CONTROLS AND PROCEDURES 

Management’s Evaluation of Disclosure Controls and Procedures 

We maintain “disclosure controls and procedures,” as defined in Rules 13a-15(e) or 15d-15(e) under the 
Exchange Act to mean controls and other procedures of a company that are designed to ensure that information required 
to be disclosed by a company in the reports that it files or submits under the Exchange Act is recorded, processed, 
summarized, and reported within the time periods specified in the SEC’s rules and forms. Our disclosure controls and 
procedures include, without limitation, controls and other procedures designed to ensure that information required to be 
disclosed by us in the reports we file or submit under the Exchange Act is accumulated and communicated to our 
management, including our principal executive officer and principal financial officer, as appropriate, to allow timely 
decisions regarding required disclosure.  

Our management, with the participation of our principal executive officer and principal financial officer, 
evaluated the effectiveness of our disclosure controls and procedures as of December 31, 2018. Our management 
recognizes that any controls and procedures, no matter how well designed and operated, can provide only reasonable 
assurance of achieving their objectives, and management necessarily applies its judgment in evaluating the cost-benefit 
relationship of possible controls and procedures. Our principal executive officer and principal financial officer have 
concluded based upon the evaluation described above that, as of December 31, 2018, our disclosure controls and 
procedures were effective at the reasonable assurance level. 

We continue to review and document our disclosure controls and procedures and may from time to time make 

changes aimed at enhancing their effectiveness and to ensure that our systems evolve with our business.  

Management’s Annual Report on Internal Control over Financial Reporting 

Our management is responsible for establishing and maintaining adequate internal control over financial 
reporting. Internal control over financial reporting is defined in Rules 13a-15(f) and 15d-15(f) under the Exchange Act as 
a process designed by, or under the supervision of, a company’s principal executive officer and principal financial 
officer, or persons performing similar functions, and effected by a company’s board of directors, management, and other 
personnel, to provide reasonable assurance regarding the reliability of financial reporting and the preparation of financial 
statements for external purposes in accordance with generally accepted accounting principles and includes those policies 
and procedures that: 

 

 

pertain to the maintenance of records that in reasonable detail accurately and fairly reflect the transactions 
and dispositions of a company’s assets; 

provide reasonable assurance that transactions are recorded as necessary to permit preparation of financial 
statements in accordance with generally accepted accounting principles, and that a company’s receipts and 
expenditures are being made only in accordance with authorizations of the company’s management and 
directors; and 

 

provide reasonable assurance regarding prevention or timely detection of unauthorized acquisition, use or 
disposition of our assets that could have a material effect on the financial statements. 

143 

  
  
 
 
  
  
 
  
  
Because of its inherent limitations, internal control over financial reporting may not prevent or detect 
misstatements. Projections of any evaluation of effectiveness to future periods are subject to the risk that controls may 
become inadequate because of changes in conditions, or that the degree of compliance with the policies or procedures 
may deteriorate. 

Under the supervision of and with the participation of our principal executive officer and principal financial and 

accounting officer, our management assessed the effectiveness of our internal control over financial reporting as of 
December 31, 2018 based on the criteria set forth by the Committee of Sponsoring Organizations of the Treadway 
Commission (COSO) in Internal Control—Integrated Framework (2013 framework). Based on this assessment, 
management concluded that our internal control over financial reporting was effective as of December 31, 2018. 

ITEM 9B.  

  OTHER INFORMATION 

As previously discussed, we are a party to a license agreement with REGENX for the development and 
commercialization of gene therapies to treat ALS, Friedreich’s ataxia and Huntington’s disease. Pursuant to our license 
agreement, REGENX granted us a non-exclusive worldwide license to certain technology and an option to obtain a non-
exclusive worldwide license for intellectual property pertaining to a specified AAV vector. We exercised an option to 
certain technology of REGENX in November 2016. We do not currently utilize any rights licensed under the REGENX 
license agreement. 

On February 21, 2019, we provided REGENX notice that we were exercising our right to terminate our license 
agreement with REGENX in its entirety, effective on May 22, 2019, at the expiration of the required notice period. Upon 
the effective date of termination, we are required to grant to REGENX a non-exclusive, perpetual, irrevocable, 
worldwide, royalty-free, transferable, sublicensable license to certain improvements made by us during the term of the 
license (including any intellectual property rights with respect thereto). This grant back will allow for REGENX’s use of 
the improvements in the research, development or commercialization of products in any therapeutic indication. 
Additionally, all licenses granted to us under the license agreement will terminate upon the effective date of termination. 

PART III 

ITEM 10.  

  DIRECTORS, EXECUTIVE OFFICERS AND CORPORATE GOVERNANCE 

Incorporated by reference from the information in our Proxy Statement for our 2019 Annual Meeting of 

Stockholders, which we expect to file with the SEC within 120 days of the end of the fiscal year to which this Annual 
Report on Form 10-K relates. 

ITEM 11.  

 EXECUTIVE COMPENSATION 

Incorporated by reference from the information in our Proxy Statement for our 2019 Annual Meeting of 

Stockholders, which we expect to file with the SEC within 120 days of the end of the fiscal year to which this Annual 
Report on Form 10-K relates. 

ITEM 12. 

   SECURITY OWNERSHIP OF CERTAIN BENEFICIAL OWNERS AND MANAGEMENT 

AND RELATED STOCKHOLDER MATTERS 

Incorporated by reference from the information in our Proxy Statement for our 2019 Annual Meeting of 

Stockholders, which we expect to file with the SEC within 120 days of the end of the fiscal year to which this Annual 
Report on Form 10-K relates. 

144 

 
  
  
 
 
 
 
  
  
 
 
  
  
 
  
  
  
 
 
ITEM 13. 

   CERTAIN RELATIONSHIPS AND RELATED TRANSACTIONS, AND DIRECTOR 

INDEPENDENCE 

Incorporated by reference from the information in our Proxy Statement for our 2019 Annual Meeting of 

Stockholders, which we expect to file with the SEC within 120 days of the end of the fiscal year to which this Annual 
Report on Form 10-K relates. 

ITEM 14.  

   PRINCIPAL ACCOUNTANT FEES AND SERVICES 

Incorporated by reference from the information in our Proxy Statement for our 2019 Annual Meeting of 

Stockholders, which we expect to file with the SEC within 120 days of the end of the fiscal year to which this Annual 
Report on Form 10-K relates. 

ITEM 15.  

   EXHIBITS, FINANCIAL STATEMENT SCHEDULES 

PART IV 

(a)(1) Financial Statements.  

Report of independent registered public accounting firm 

Consolidated Balance Sheets 

Consolidated Statements of Operations and Comprehensive Loss 

Consolidated Statements of Stockholders’ Equity 

Consolidated Statements of Cash Flows 

Notes to consolidated financial statements 

(a)(2) Financial Statement Schedules.  

Pages 
F-1 

F-2 

F-3 

F-4 

F-5 

F-6 

All schedules have been omitted because they are not required or because the required information is given in the 

Consolidated Financial Statements or Notes thereto set forth under Item 8 above.  

(a)(3) Exhibits.  

See the Exhibit Index immediately preceding the signature page of this Annual Report on Form 10-K. The 
exhibits listed in the Exhibit Index below are filed or incorporated by reference as part of this Annual Report on Form 
10-K.  

ITEM 16.  

   FORM 10-K SUMMARY 

This Annual Report on Form 10-K does not include a summary. 

145 

  
  
 
 
  
  
 
 
 
  
  
 
 
 
 
  
   
   
  
  
   
  
  
   
  
  
   
  
  
   
  
  
   
 
 
 
  
  
 
 
 
Report of Independent Registered Public Accounting Firm  

To the Stockholders and the Board of Directors of Voyager Therapeutics, Inc.  

Opinion on the Financial Statements 

We have audited the accompanying consolidated balance sheets of Voyager Therapeutics, Inc. (the “Company”) as of 
December 31, 2018 and 2017, the related consolidated statements of operations and comprehensive loss, redeemable 
convertible preferred stock and stockholders’ equity (deficit), and cash flows for each of the three years in the period 
ended December 31, 2018, and the related notes (collectively referred to as the “consolidated financial statements”). In 
our opinion, the consolidated financial statements present fairly, in all material respects, the financial position of the 
Company at December 31, 2018 and 2017, and the results of its operations and its cash flows for each of the three years 
in the period ended December 31, 2018, in conformity with U.S. generally accepted accounting principles. 

Adoption of ASU No. 2014-09 Revenue from Contracts with Customers (Topic 606) 

As discussed in Note 2 to the consolidated financial statements, the Company changed its method of accounting for 
revenue in 2018 due to the adoption of Accounting Standard Update (ASU) No. 2014-09, Revenue from Contracts with 
Customers (Topic 606), and the related amendments. 

Basis for Opinion 

These financial statements are the responsibility of the Company’s management. Our responsibility is to express an 
opinion on the Company’s financial statements based on our audits. We are a public accounting firm registered with the 
Public Company Accounting Oversight Board (United States) (PCAOB) and are required to be independent with respect 
to the Company in accordance with the U.S. federal securities laws and the applicable rules and regulations of the 
Securities and Exchange Commission and the PCAOB. 

We conducted our audits in accordance with the standards of the PCAOB. Those standards require that we plan and 
perform the audit to obtain reasonable assurance about whether the financial statements are free of material 
misstatement, whether due to error or fraud. The Company is not required to have, nor were we engaged to perform, an 
audit of its internal control over financial reporting. As part of our audits we are required to obtain an understanding of 
internal control over financial reporting but not for the purpose of expressing an opinion on the effectiveness of the 
Company’s internal control over financial reporting. Accordingly, we express no such opinion.  

Our audits included performing procedures to assess the risks of material misstatement of the financial statements, 
whether due to error or fraud, and performing procedures that respond to those risks. Such procedures included 
examining, on a test basis, evidence regarding the amounts and disclosures in the financial statements. Our audits also 
included evaluating the accounting principles used and significant estimates made by management, as well as evaluating 
the overall presentation of the financial statements. We believe that our audits provide a reasonable basis for our opinion. 

/s/ Ernst & Young LLP 

We have served as the Company’s auditor since 2015. 
Boston, Massachusetts  
February 26, 2019 

F-1 

 
 
 
 
 
 
 
 
  
 
  
 
 
Voyager Therapeutics, Inc. 
Consolidated Balance Sheets 
(amounts in thousands, except share and per share data) 

Assets 
Current assets: 

Cash and cash equivalents 
Marketable securities, current 
Prepaid expenses and other current assets 

Total current assets 

Property and equipment, net 
Deposits and other non-current assets 
Marketable securities, non-current 

Total assets 

Liabilities and stockholders’ equity 
Current liabilities: 

Accounts payable 
Accrued expenses 
Deferred revenue, current portion 

Total current liabilities 

Deferred rent 
Deferred revenue, net of current portion 
Other non-current liabilities 
Total liabilities 

Commitments and contingencies (see note 7) 
Stockholders’ equity: 

Preferred stock, $0.001 par value: 5,000,000 shares authorized; no shares issued and outstanding at 
December 31, 2018 and 2017 
Common stock, $0.001 par value: 120,000,000 shares authorized; 32,364,895 and 31,572,044 
shares issued and outstanding at December 31, 2018 and 2017, respectively 
Additional paid-in capital 
Accumulated other comprehensive loss 
Accumulated deficit 

Total stockholders’ equity 
Total liabilities and stockholders’ equity 

December 31,  

2018 

2017 

$ 

$ 

$ 

 46,859   
 108,947   
 6,675   
 162,481   
 12,771   
 1,149   
 628   
 177,029   

 1,038   
 9,788   
 20,847   
 31,673   
 5,710   
 92,199   
 1,001   
 130,583   

 31,530   
 137,522   
 2,738   
 171,790   
 10,283   
 1,304   
 1,100   
 184,477   

 1,020   
 11,497   
 3,380   
 15,897   
 5,337   
 28,180   
 1,012   
 50,426   

 —   

 —   

 32   
 315,598   
 (133) 
 (269,051) 
 46,446   
 177,029   

$ 

 32   
 295,019   
 (287) 
 (160,713) 
 134,051   
 184,477   

$ 

$ 

$ 

$ 

The accompanying notes are an integral part of these consolidated financial statements. 

F-2 

 
 
 
 
 
 
 
 
 
 
  
 
     
     
  
 
 
     
 
 
     
 
 
 
 
 
 
 
 
 
 
  
  
 
  
  
 
  
  
 
  
  
 
  
  
 
  
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
  
 
  
  
 
  
 
 
  
  
 
  
  
 
  
  
 
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
  
 
  
  
 
  
  
 
  
  
 
  
  
 
 
 
 
 
Voyager Therapeutics, Inc. 
Consolidated Statements of Operations and Comprehensive Loss 
(amounts in thousands, except share and per share data)  

Collaboration revenue 
Operating expenses: 

Research and development 
General and administrative 
Total operating expenses 

Operating loss 
Other income (expense), net: 

Interest income, net 
Other (expense) income, net 
Total other income (expense), net 

Loss before income taxes 
Income tax provision 
Net loss 
Other comprehensive loss 

Net unrealized gain (loss) on available-for-sale-securities, net 
of tax expense of $128 for the year ended December 31, 2016 

Total other comprehensive income (loss) 

Comprehensive loss 
Net loss per share, basic and diluted 
Weighted-average common shares outstanding, basic and diluted 

  $ 

  $ 
  $ 

Year ended  
December 31,  

2018 

  $ 

 7,619     $ 

2017 
 10,135     $ 

2016 
 14,220  

 64,905  
 33,809  
 98,714  
 (91,095) 

 62,260  
 19,738  
 81,998  
 (71,863) 

 3,310  
 (683) 
 2,627  
 (88,468) 
 (180) 
 (88,288)  $ 

 1,227  
 (62) 
 1,165  
 (70,698) 
 —  
 (70,698)  $ 

 42,249  
 13,270  
 55,519  
 (41,299) 

 976  
 182  
 1,158  
 (40,141) 
 52  
 (40,193) 

 34  
 34  
 (88,254)  $ 
 (2.75)  $ 

 (235) 
 (235) 
 (70,933)  $ 
 (2.64)  $ 

 199  
 199  
 (39,994) 
 (1.59) 
    25,302,414  

    32,065,781  

    26,803,711  

The accompanying notes are an integral part of these consolidated financial statements. 

F-3 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
     
     
     
  
 
   
 
   
 
   
 
  
  
  
  
  
  
  
 
  
 
 
 
 
 
   
 
   
 
   
 
  
  
  
  
  
  
  
  
  
 
 
 
 
 
 
 
   
 
   
 
   
 
  
  
  
  
  
  
 
 
 
 
Voyager Therapeutics, Inc.  
Consolidated Statements of Stockholders’ Equity  
 (amounts in thousands, except share data) 

  Accumulated   

  Additional  

Other 

Common Stock 

Paid-In 
      Amount        Capital 

Shares 

  Comprehensive   Accumulated  

Loss 

      Deficit 

Stockholders’    
Equity 

Balance at December 31, 2015 
Vesting of restricted stock 
Exercises of vested stock options 
Stock-based compensation expense 
Unrealized gain on available-for-sale securities, net of tax 
Net loss 
Balance at December 31, 2016 
Vesting of restricted stock 
Exercises of vested stock options 
Issuance of common stock under ESPP 
Issuance of common stock from public offering (net of underwriters discounts and 
issuance costs of $4,100) 
Stock-based compensation expense 
Unrealized loss on available-for-sale securities, net of tax 
Net loss 
Balance at December 31, 2017 
Vesting of restricted stock 
Exercises of vested stock options 
Issuance of common stock under ESPP 
Stock-based compensation expense 
Unrealized gain on available-for-sale securities, net of tax 
Cumulative-effect adjustment to beginning accumulated deficit and statement of 
operations resulting from ASU No. 2016-01 
Modified retrospective adjustment to beginning accumulated deficit and deferred revenue 
resulting from ASU No. 2014-09 
Net loss 
Balance at December 31, 2018 

 24,930,979   $ 
 601,501  
 65,432  
 —  
 —  
 —  

 25,597,912   $ 
 573,803  
 158,677  
 66,652  

 5,175,000  
 —  
 —  
 —  

 31,572,044   $ 
 319,891  
 384,186  
 88,774  
 —  
 —  

 —  

 —  
 —  

 32,364,895   $ 

 17  
 514  
 6,310  
 —  
 —  

 25   $   219,122   $ 
 1  
 —  
 —  
 —  
 —  
 26   $   225,963   $ 
 1  
 —  
 —  

 12  
 1,363  
 563  

 57,989  
 9,129  
 —  
 —  

 5  
 —  
 —  
 —  
 32   $   295,019   $ 
 —  
 —  
 —  
 —  
 —  

 9  
 3,891  
 969  
 15,710  
 —  

 (251)  $ 
 —  
 —  
 —  
 199  
 —  
 (52)  $ 
 —  
 —  
 —  

 —  
 —  
 (235) 
 —  
 (287)  $ 
 —  
 —  
 —  
 —  
 34  

 (49,822)  $ 
 —  
 —  
 —  
 —  
 (40,193) 
 (90,015)  $ 
 —  
 —  
 —  

 —  
 —  
 —  
 (70,698) 

 (160,713)  $ 
 —  
 —  
 —  
 —  
 —  

 169,074  
 18  
 514  
 6,310  
 199  
 (40,193) 
 135,922  
 13  
 1,363  
 563  

 57,994  
 9,129  
 (235) 
 (70,698) 
 134,051  
 9  
 3,891  
 969  
 15,710  
 34  

 —  

 —  

 120  

 (120) 

 —  

 —  
 —  
 32   $   315,598   $ 

 —  
 —  

 —  
 —  
 (133)  $ 

 (19,930) 
 (88,288) 

 (269,051)  $ 

 (19,930) 
 (88,288) 
 46,446  

The accompanying notes are an integral part of these consolidated financial statements 

F-4 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
   
 
 
 
 
 
 
 
 
 
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
 
 
 
 
 
 
 
 
 
 
  
 
 
 
 
     
     
     
  
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
Voyager Therapeutics, Inc. 
Consolidated Statements of Cash Flows 
(amounts in thousands) 

Cash flow from operating activities 
Net loss 
Adjustments to reconcile net loss to net cash used in operating activities: 
Stock-based compensation expense 
Depreciation 
Amortization of premiums and discounts on marketable securities 
In-kind research and development expenses 
Other non-cash items 
Changes in operating assets and liabilities: 

Prepaid expenses and other current assets 
Other non-current assets 
Deferred revenue 
Accounts payable 
Accrued expenses 
Other non-current liabilities 
Lease incentive benefit 
Net cash used in operating activities 

Cash flow from investing activities 
Purchases of property and equipment 
Purchases of marketable securities 
Proceeds from maturities or sales of marketable securities 
Net cash provided by (used in) investing activities 

Cash flow from financing activities 
Proceeds from the issuance of stock net of discount and issuance costs 
Proceeds from the exercise of stock options 
Proceeds from the purchase of common stock under ESPP 

Net cash provided by financing activities 

Net increase (decrease) in cash and cash equivalents 
Cash, cash equivalents, and restricted cash beginning of period 
Cash, cash equivalents, and restricted cash end of period 
Supplemental disclosure of cash and non-cash activities 
Impact of adopting new accounting standards 
Capital expenditures incurred but not yet paid 

Year ended  
December 31,  
2017 

2018 

2016 

$ 

 (88,288) 

$ 

 (70,698) 

$ 

 (40,193) 

 15,710   
 2,117   
 (2,163) 
 176   
 859   

 (3,937) 
 (180) 
 61,380   
 (282) 
 (1,600) 
 —   
 321   
 (15,887) 

 9,238   
 1,595   
 (24) 
 113   
 46   

 1,630   
 —   
 (10,135) 
 470   
 4,900   
 1,000   
 515   
 (61,350) 

 6,310   
 612   
 696   
 1,182   
 709   

 (847) 
 7   
 (14,582) 
 (62) 
 2,636   
 —   
 1,050   
 (42,482) 

 (4,305) 
 (333,228) 
 364,000   
 26,467   

 (3,985) 
 (147,296) 
 147,600   
 (3,681) 

 (5,029) 
 (112,350) 
 165,100   
 47,721   

 —   
 3,889   
 860   
 4,749   
 15,329   
 32,265   
 47,594   

 20,050   
 300   

$ 

$ 
$ 

 57,994   
 1,363   
 563   
 59,920 
 (5,111) 
 37,376   
 32,265   

 —   
 —   

$ 

$ 
$ 

 —   
 514   
 —   
 514   
 5,753   
 31,623   
 37,376   

 —   
 242   

$ 

$ 
$ 

The accompanying notes are an integral part of these consolidated financial statements. 

F-5 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
     
     
     
  
 
 
     
 
 
     
 
 
     
 
 
 
 
 
 
 
 
 
 
 
 
 
  
  
  
 
  
  
  
 
 
 
 
 
 
 
 
 
  
  
  
 
 
 
 
 
 
 
 
 
 
  
  
  
 
  
  
  
 
  
  
  
 
  
  
  
 
  
  
  
 
  
  
  
 
  
  
  
 
  
  
  
 
 
 
 
 
 
 
 
 
 
 
  
  
  
 
  
  
  
 
  
  
  
 
  
  
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
  
  
 
 
 
 
 
  
  
 
  
  
  
 
  
  
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
VOYAGER THERAPEUTICS INC. 

NOTES TO CONSOLIDATED FINANCIAL STATEMENTS 

1. Nature of business 

Voyager Therapeutics, Inc. (the “Company”) is a clinical-stage gene therapy company focused on developing 

life-changing treatments for patients suffering from severe neurological diseases. The Company is focused on 
neurological diseases where it believes an adeno-associated virus (“AAV”) gene therapy approach that either increases 
or decreases the production of a specific protein can slow or reduce the symptoms experienced by patients, and therefore 
have a clinically meaningful impact. The Company has built a gene therapy platform that it believes positions itself to be 
a leading company at the intersection of AAV gene therapy and severe neurological disease. The Company’s gene 
therapy platform enables it to engineer, optimize, manufacture and deliver its AAV-based gene therapies that have the 
potential to provide durable efficacy following a single administration.  

Additionally, the Company is working to identify novel AAV capsids, which are the outer viral protein shells 

that enclose the genetic material of the virus payload. The Company’s team of experts in the fields of AAV gene therapy 
and neuroscience first identifies and selects severe neurological diseases that are well-suited for treatment using AAV 
gene therapy. The Company then engineers and optimizes AAV vectors for delivery of the virus payload to the targeted 
tissue or cells. The Company’s manufacturing process employs an established system that it believes will enable 
production of high quality AAV vectors at commercial-scale. In addition to the Company’s capsid optimization efforts, it 
leverages novel delivery paradigms, established routes of administration, and advances in dosing techniques to optimize 
delivery of its AAV gene therapies to target tissues, regions and cell types that are critical to the disease of interest. The 
Company believes it can achieve this directly, with targeted infusions to discrete regions of the brain or spinal cord, or 
systemically, in conjunction with its novel capsids. 

The Company’s business strategy focuses on discovering, developing, manufacturing and commercializing its 
gene therapy programs. As part of this strategy, the Company has developed core competencies specific to AAV gene 
therapy development and manufacturing and is beginning to build its commercial infrastructure. This business strategy 
also includes business development activities that may include in-licensing activities or partnering certain programs in 
certain geographies with collaborators, as the Company has demonstrated through its collaboration with Sanofi Genzyme 
(the “Sanofi Genzyme Collaboration”), its collaboration with AbbVie Biotechnology Ltd (the “AbbVie Tau 
Collaboration”), its collaboration with Neurocrine Biosciences, Inc. (the “Neurocrine Collaboration”), and its 
collaboration with AbbVie Ireland Unlimited Company (the “AbbVie Alpha-Synuclein Collaboration”). The Company is 
devoting substantially all of its efforts to product research and development, market development, and raising capital. 
The Company is subject to risks common to companies in the biotechnology and gene therapy industry, including but not 
limited to, the need to obtain sufficient capital to continue to fund its operations, risks of failure of preclinical studies and 
clinical trials, the need to obtain marketing approval for its product candidates, the need to successfully commercialize 
and gain market acceptance of its product candidates, dependence on key personnel, protection of proprietary 
technology, compliance with government regulations, development by competitors of technological innovations, and 
ability to transition from pilot-scale manufacturing to large-scale production of products.  

The Company has incurred annual net operating losses in every year since inception. As of December 31, 2018, 

the Company had an accumulated deficit of $269.1 million. The Company has not generated any product revenue and 
has financed its operations primarily through public offerings and private placements of its equity securities and funding 
from its collaborations with Sanofi Genzyme and AbbVie. Additionally, in January 2019, the Company entered into the 
Neurocrine Collaboration which is expected to commence in the first half of 2019. Under the Neurocrine Collaboration, 
the Company will receive an upfront payment of $165.0 million, including a $50.0 million investment in 4,179,728 
shares of common stock. The effectiveness of this agreement is subject to certain conditions, including the expirations or 
termination of the applicable waiting period under the Hart-Scott-Rodino Antitrust Improvements Act of 1976, as 
amended, and other customary closing conditions. In February 2019, the Company announced the AbbVie Alpha-
Synuclein Collaboration to develop and commercialize vectorized antibodies directed at pathological species of alpha-
synuclein for the potential treatment of Parkinson’s disease and other diseases (synucleinopathies) characterized by the 
abnormal accumulation of misfolded alpha-synuclein protein. Under the terms of the AbbVie Alpha-Synuclein 
Collaboration, the Company will receive an upfront payment of $65.0 million. Based upon the current operating plan, the 
Company expects that its existing cash, cash equivalents, and marketable debt securities, as well as amounts expected 

F-6 

 
from the upfront payment and expected reimbursement of development costs from the Neurocrine Collaboration 
Agreement and the upfront payment from the AbbVie Alpha-Synuclein Collaboration Agreement entered into in 2019, 
will enable the Company to fund its operating expenses and capital expenditure requirements into mid-2022. There can 
be no assurance that the Company will be able to obtain additional debt or equity financing or generate product revenue 
or revenue from collaborative partners on terms acceptable to the Company, on a timely basis or at all. The failure of the 
Company to obtain sufficient funds on acceptable terms when needed could have a material adverse effect on the 
Company’s business, results of operations, and financial condition. 

2. Summary of significant accounting policies 

The following is a summary of significant accounting policies followed in the preparation of these financial 

statements. 

Basis of presentation 

The accompanying consolidated financial statements include those of the Company and its subsidiary, Voyager 
Securities Corporation, after elimination of all intercompany accounts and transactions. The accompanying consolidated 
financial statements have been prepared in conformity with accounting principles generally accepted in the United States 
of America (“GAAP”). 

Public offerings 

On October 29, 2015, in preparation for the Company’s IPO, the Company’s Board of Directors and 
stockholders approved a 1-for-4.25 reverse split of the Company’s common stock, which became effective on October 
29, 2015. All share and per share amounts in the consolidated financial statements and notes thereto have been 
retroactively adjusted for all periods presented to give effect to this reverse split, including reclassifying an amount equal 
to the reduction in par value of common stock to additional paid-in capital. 

On November 16, 2015, the Company completed the sale of 5,750,000 shares of its common stock in its initial 

public offering (the “IPO”), at a price to the public of $14.00 per share, resulting in net proceeds to the Company of 
$72.9 million after deducting underwriting discounts, commissions and offering expenses payable by the Company.  

On November 7, 2017, the Company completed the sale of 5,175,000 shares of its common stock in a public 

offering at a price to the public of $12.00 per share, resulting in net proceeds to the Company of $58.0 million after 
deducting underwriting discounts, commissions, and offering expenses payable by the Company. 

Use of Estimates 

The preparation of consolidated financial statements in conformity with GAAP requires management to make 
estimates and assumptions that affect the amounts reported in the consolidated financial statements and accompanying 
notes. On an ongoing basis, the Company’s management evaluates its estimates, which include, but are not limited to, 
estimates related to revenue recognition, accrued expenses, stock-based compensation expense, and income taxes. The 
Company bases its estimates on historical experience and other market specific or other relevant assumptions that it 
believes to be reasonable under the circumstances. Actual results may differ from those estimates or assumptions. 

Fair Value of Financial Instruments 

ASC Topic 820, Fair Value Measurement (“ASC 820”), establishes a fair value hierarchy for instruments 

measured at fair value that distinguishes between assumptions based on market data (observable inputs) and the 
Company’s own assumptions (unobservable inputs). Observable inputs are inputs that market participants would use in 
pricing the asset or liability based on market data obtained from sources independent of the Company. Unobservable 
inputs are inputs that reflect the Company’s assumptions about the inputs that market participants would use in pricing 
the asset or liability, and are developed based on the best information available in the circumstances. 

ASC 820 identifies fair value as the exchange price, or exit price, representing the amount that would be 
received to sell an asset or paid to transfer a liability in an orderly transaction between market participants. As a basis for 

F-7 

considering market participant assumptions in fair value measurements, ASC 820 establishes a three-tier fair value 
hierarchy that distinguishes between the following: 

  Level 1—Quoted market prices in active markets for identical assets or liabilities. 

  Level 2—Inputs other than Level 1 inputs that are either directly or indirectly observable, such as quoted 

market prices, interest rates, and yield curves. 

  Level 3—Unobservable inputs developed using estimates of assumptions developed by the Company, 

which reflect those that a market participant would use. 

To the extent that the valuation is based on models or inputs that are less observable or unobservable in the 

market, the determination of fair value requires more judgment. Accordingly, the degree of judgment exercised by the 
Company in determining fair value is greatest for instruments categorized in Level 3. A financial instrument’s level 
within the fair value hierarchy is based on the lowest level of any input that is significant to the fair value measurement. 

The carrying amounts reflected in the balance sheets for cash and cash equivalents, prepaid expenses and other 

current assets, accounts payable and accrued expenses approximate their fair values, due to their short-term nature. 

Cash and Cash Equivalents 

The Company considers all highly liquid investments purchased with original maturities of 90 days or less at 

acquisition to be cash equivalents. Cash and cash equivalents include cash held in banks and amounts held in money 
market funds. 

Marketable Securities 

The Company classifies marketable debt securities with a remaining maturity of greater than three months when 

purchased as available-for-sale. Marketable debt securities with a remaining maturity date greater than one year and 
marketable equity securities are classified as non-current where the Company has the intent and ability to hold these 
securities for at least the next 12 months. During 2016, the Company invested in a supplier and received common stock 
and warrants to purchase common stock in that entity. The common stock is considered an available-for-sale marketable 
equity security and is included in non-current marketable securities, and the warrants are included in non-current assets. 

All available for sale debt securities are carried at fair value with the unrealized gains and losses included in 

other comprehensive income (loss) as a component of stockholders’ equity until realized. Any premium or discount 
arising at purchase is amortized and/or accreted to interest income and/or expense. Realized gains and losses are 
determined using the specific identification method and are included in other income (expense). If any adjustment to fair 
value reflects a decline in value of the investment, the Company considers all available evidence to evaluate the extent to 
which the decline is “other than temporary” and, if so, recognizes the loss through a charge to the Company’s statement 
of operations and comprehensive loss. No other than temporary losses have been recognized. 

F-8 

Cash, cash equivalents, and marketable securities as of December 31, 2018 and 2017 consist of the following: 

Amortized   
Cost 

Unrealized   
Gains 

Unrealized   
Losses 

Fair 
Value 

(in thousands) 

 46,173   $ 

 —   $ 

 —   $ 

 46,173  

As of December 31, 2018 
Money market funds included in cash and cash equivalents   $ 
Marketable securities: 
U.S. Treasury notes 
Equity securities 

 108,951  
 1,220  

Total marketable securities 
Total money market funds and marketable securities 

  $   110,171   $ 
  $   156,344   $ 

As of December 31, 2017 
Money market funds included in cash and cash equivalents   $ 
Marketable securities: 
U.S. Treasury notes 
Equity securities 

 137,560  
 1,220  

Total marketable securities 
Total money market funds and marketable securities 

  $   138,780   $ 
  $   169,249   $ 

 30,469   $ 

 —   $ 

 —   $ 

 30,469  

 1  
 —  
 1   $ 
 1   $ 

 108,947  
 5  
 592  
 628  
 597   $   109,575  
 597   $   155,748  

 —  
 —  
 —   $ 
 —   $ 

 137,522  
 38  
 120  
 1,100  
 158   $   138,622  
 158   $   169,091  

All of the Company’s marketable debt securities at December 31, 2018 and 2017 have a contractual maturity of 

one year or less. 

Restricted Cash 

At December 31, 2018 and 2017, the Company maintained restricted cash totaling approximately $0.7 million 

held in the form of money market accounts as collateral for the Company’s facility lease obligation. The balance is 
included within deposits in other non-current assets in the accompanying consolidated balance sheets. The following 
table provides a reconciliation of cash, cash equivalents, and restricted cash within the condensed consolidated balance 
sheet that sum to the total of the same such amounts shown in the statement of cash flows: 

Cash and cash equivalents 
  $ 
Restricted cash included in deposits and other noncurrent assets    
  $ 
Total cash, cash equivalents, and restricted cash 

 46,859    $ 
 735   
 47,594    $ 

 31,530    $ 
 735   
 32,265    $ 

 36,641   
 735   
 37,376   

2018 

As of December 31,  

2017 
(in thousands) 

2016 

Property and Equipment 

Property and equipment consists of laboratory equipment, furniture and office equipment, and leasehold 

improvements and is stated at cost, less accumulated depreciation. Maintenance and repairs that do not improve or 
extend the lives of the respective assets are expensed to operations as incurred; while costs of major additions and 
betterments are capitalized. Depreciation is calculated over the estimated useful lives of the assets using the straight-line 
method. 

Impairment of Long-Lived Assets 

The Company evaluates long-lived assets for potential impairment when events or changes in circumstances 

indicate the carrying value of the assets may not be recoverable. Recoverability is measured by comparing the book 
values of the assets to the expected future net undiscounted cash flows that the assets are expected to generate. If such 
assets are considered to be impaired, the impairment to be recognized is measured by the amount by which the book 
values of the assets exceed their fair value. The Company has not recognized any impairment losses from inception 
through December 31, 2018. 

F-9 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
     
     
     
     
  
 
 
  
       
     
       
     
       
     
       
     
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
 
  
 
 
 
 
 
       
     
       
     
       
     
       
     
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
 
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
    
    
 
 
 
 
 
 
 
Revenue Recognition 

As of December 31, 2018, all of the Company’s revenue is generated from its collaboration agreements with 

Sanofi Genzyme Corporation, a Sanofi company (“Sanofi Genzyme”), and AbbVie Biotechnology Ltd. and its affiliates 
collectively, (“AbbVie”). 

The Company enters into collaboration agreements which are within the scope of ASC 606, Revenue from 

Contracts with Customers (“ASC 606”), under which the Company licenses rights to certain of the Company’s product 
candidates and performs research and development services. The terms of these arrangements typically include payment 
of one or more of the following: non-refundable, upfront fees; reimbursement of research and development costs; 
development, regulatory, and commercial milestone payments; and royalties on net sales of licensed products.  

Under ASC 606, an entity recognizes revenue when its customer obtains control of promised goods or services, 

in an amount that reflects the consideration which the entity expects to receive in exchange for those goods or services. 
To determine the appropriate amount of revenue to be recognized for arrangements determined to be within the scope of 
ASC 606, the Company performs the following five steps: (i) identification of the promised goods or services in the 
contract; (ii) determination of whether the promised goods or services are performance obligations including whether 
they are distinct in the context of the contract; (iii) measurement of the transaction price, including the constraint on 
variable consideration; (iv) allocation of the transaction price to the performance obligations; and (v) recognition of 
revenue when (or as) the Company satisfies each performance obligation. The Company only applies the five-step model 
to contracts when it is probable that the entity will collect consideration it is entitled to in exchange for the goods or 
services it transfers to the customer.  

The promised goods or services in the Company’s arrangements typically consist of license rights to the 

Company’s intellectual property and research and development services. The Company provides options to additional 
items in the contracts, which are accounted for as separate contracts when the customer elects to exercise such options, 
unless the option provides a material right to the customer. The Company evaluates the customer options for material 
rights, or options to acquire additional goods or services for free or at a discount. If the customer options are determined 
to represent a material right, the material right is recognized as a separate performance obligation at the outset of the 
arrangement. Performance obligations are promised goods or services in a contract to transfer a distinct good or service 
to the customer and are considered distinct when (i) the customer can benefit from the good or service on its own or 
together with other readily available resources and (ii) the promised good or service is separately identifiable from other 
promises in the contract. In assessing whether promised goods or services are distinct, the Company considers factors 
such as the stage of development of the underlying intellectual property, the capabilities of the customer to develop the 
intellectual property on its own or whether the required expertise is readily available and whether the goods or services 
are integral or dependent to other goods or services in the contract. 

The Company estimates the transaction price based on the amount expected to be received for transferring the 

promised goods or services in the contract. The consideration may include fixed consideration or variable consideration. 
At the inception of each arrangement that includes variable consideration, the Company evaluates the amount of 
potential payments and the likelihood that the payments will be received. The Company utilizes either the most likely 
amount method or expected amount method to estimate the amount expected to be received based on which method best 
predicts the amount expected to be received. The amount of variable consideration which is included in the transaction 
price may be constrained, and is included in the transaction price only to the extent that it is probable that a significant 
reversal in the amount of the cumulative revenue recognized will not occur in a future period.  

The Company’s contracts often include development and regulatory milestone payments which are assessed 
under the most likely amount method and constrained if it is probable that a significant revenue reversal would occur. 
Milestone payments that are not within the Company’s control or the licensee’s control, such as regulatory approvals, are 
not considered probable of being achieved until those approvals are received. At the end of each reporting period, the 
Company re-evaluates the probability of achievement of such development milestones and any related constraint, and if 
necessary, adjusts its estimate of the overall transaction price. Any such adjustments are recorded on a cumulative catch-
up basis, which would affect collaboration revenues in the period of adjustment. To date, the Company has not 
recognized any consideration related to the achievement of development, regulatory, or commercial milestone revenue 
resulting from any of the Company’s collaboration arrangements. 

F-10 

For arrangements that include sales-based royalties, including milestone payments based on the level of sales, 
and the license is deemed to be the predominant item to which the royalties relate, the Company recognizes revenue at 
the later of (i) when the related sales occur, or (ii) when the performance obligation to which some or all of the royalty 
has been allocated has been satisfied (or partially satisfied). To date, the Company has not recognized any consideration 
related to sales-based royalty revenue resulting from any of the Company’s collaboration arrangements. 

The Company allocates the transaction price based on the estimated stand-alone selling price of each of the 

performance obligations. The Company must develop assumptions that require judgment to determine the stand-alone 
selling price for each performance obligation identified in the contract. The Company utilizes key assumptions to 
determine the stand-alone selling price for service obligations, which may include other comparable transactions, pricing 
considered in negotiating the transaction and the estimated costs. Additionally, in determining the standalone selling 
price for material rights, the Company utilizes comparable transactions, clinical trial success probabilities, and estimates 
of option exercise likelihood. Variable consideration is allocated specifically to one or more performance obligations in a 
contract when the terms of the variable consideration relate to the satisfaction of the performance obligation and the 
resulting amounts allocated are consistent with the amounts the Company would expect to receive for the satisfaction of 
each performance obligation.  

The consideration allocated to each performance obligation is recognized as revenue when control is transferred 

for the related goods or services. For performance obligations which consist of licenses and other promises, the 
Company utilizes judgment to assess the nature of the combined performance obligation to determine whether the 
combined performance obligation is satisfied over time or at a point in time and, if over time, the appropriate method of 
measuring progress. The Company evaluates the measure of progress each reporting period and, if necessary, adjusts the 
measure of performance and related revenue recognition.  

Upfront payments and fees are recorded as deferred revenue upon receipt or when due until the Company 

performs its obligations under these arrangements. Amounts are recorded as accounts receivable when the Company’s 
right to consideration is unconditional 

Research and Development 

Research and development costs are charged to expense as incurred in performing research and development 

activities. The costs include employee compensation costs, external research, consultant costs, sponsored research, 
in-kind services provided under the Sanofi Genzyme agreement, license fees, process development and facilities costs. 
Facilities costs primarily include the allocation of rent, utilities and depreciation. 

Research Contract Costs and Accruals 

The Company has entered into various research and development contracts with research institutions and other 

companies. These agreements are generally cancelable. The Company records accruals for estimated ongoing research 
costs. When evaluating the adequacy of the accrued liabilities, the Company analyzes progress of the studies, including 
the phase or completion of events, invoices received and contracted costs. Significant judgments and estimates may be 
made in determining the accrued balances at the end of any reporting period. Actual results could differ from the 
Company’s estimates. The Company’s historical accrual estimates have not been materially different from the actual 
costs. 

Patent Costs 

The Company expenses patent application and related legal costs as incurred and classifies such costs as general 

and administrative expenses in the accompanying statements of operations. 

Stock-Based Compensation Expense 

The Company accounts for its stock-based compensation awards in accordance with ASC Topic 718 

Compensation—Stock Compensation (“ASC 718”). ASC 718 requires all stock-based payments to employees and 
directors, including grants of restricted stock and stock options, to be recognized as expense in the statements of 
operations based on their grant date fair values. Grants of restricted stock and stock options to other service providers, 

F-11 

referred to as non-employees, are required to be recognized as expense in the statements of operations based on their 
vesting date fair values. The Company estimates the fair value of options granted using the Black-Scholes option pricing 
model. The Company uses the fair value of its common stock to determine the fair value of restricted stock awards. 

The Black-Scholes option pricing model requires inputs based on certain subjective assumptions, including 

(a) the expected stock price volatility, (b) the calculation of expected term of the award, (c) the risk-free interest rate and 
(d) expected dividends. Due to a lack of company-specific historical and implied volatility data, the Company bases the 
estimate of expected volatility on the historical volatility of a group of similar companies that are publicly traded, 
blended with the most recent period of historic volatility of its common stock. The historical volatility is calculated based 
on a period of time commensurate with the expected term assumption. The computation of expected volatility is based 
on the historical volatility of a representative group of companies with similar characteristics to the Company, including 
stage of product development and life science industry focus. The Company uses the simplified method as prescribed by 
the SEC Staff Accounting Bulletin No. 107, Share-Based Payment, to calculate the expected term for stock options 
granted to employees as it does not have sufficient historical exercise data to provide a reasonable basis upon which to 
estimate the expected term. For stock options granted to non-employees, the Company utilizes the contractual term of the 
arrangement as the basis for the expected term assumption. The risk-free interest rate is based on a treasury instrument 
whose term is consistent with the expected term of the stock options. The expected dividend yield is assumed to be zero 
as the Company has never paid dividends and has no current plans to pay any dividends on its common stock. 

The Company expenses the fair value of its stock-based compensation awards to employees on a straight-line 

basis over the associated service period, which is generally the period in which the related services are received. 
Stock-based compensation awards to non-employees are adjusted through stock-based compensation expense at each 
reporting period end to reflect the current fair value of such awards and are expensed on a straight-line basis. 

The Company records the expense for stock-based compensation awards subject to performance conditions over 

the remaining service period when management determines that achievement of the performance condition is probable. 
Management evaluates when the achievement of a performance condition is probable based on the expected satisfaction 
of the performance conditions as of the reporting date. 

Income Taxes 

Income taxes are recorded in accordance with ASC Topic 740, Income Taxes (“ASC 740”), which provides for 
deferred taxes using an asset and liability approach. Under this method, deferred tax assets and liabilities are determined 
based on the difference between the financial reporting and the tax reporting basis of assets and liabilities and are 
measured using the enacted tax rates and laws that are expected to be in effect when the differences are expected to 
reverse. The Company provides a valuation allowance against net deferred tax assets unless, based upon the weight of 
available evidence, it is more likely than not that the deferred tax assets will be realized. 

The Company accounts for uncertain tax positions in accordance with the provisions of ASC 740. When 

uncertain tax positions exist, the Company recognizes the tax benefit of tax positions to the extent that the benefit will 
more likely than not be realized. The determination as to whether the tax benefit will more likely than not be realized is 
based upon the technical merits of the tax position as well as consideration of the available facts and circumstances. As 
of December 31, 2018, the Company does not have any significant uncertain tax positions. 

Comprehensive Loss 

Comprehensive loss is comprised of net loss and other comprehensive income or loss. Other comprehensive 

income or loss consists of unrealized gains or losses on marketable securities. 

Net Loss Per Share 

Basic net loss per share is calculated by dividing the net loss by the weighted-average number of shares of 

common stock outstanding during the period, without consideration for potentially dilutive securities. Diluted net loss 
per share is computed by dividing the net loss by the weighted-average number of shares of common stock and 
potentially dilutive securities outstanding for the period determined using the treasury-stock and if-converted methods. 

F-12 

For purposes of the diluted net loss per share, unvested restricted common stock and outstanding stock options 

are considered to be potentially dilutive securities, but are excluded from the calculation of diluted net loss per share 
because their effect would be anti-dilutive and therefore, basic and diluted net loss per share were the same for all 
periods presented. 

The following table sets forth the outstanding potentially dilutive securities that have been excluded in the 

calculation of diluted net loss per share because to do so would be anti-dilutive: 

Unvested restricted common stock 
Outstanding stock options 

Total 

Concentrations of Credit Risk and Off-Balance Sheet Risk 

As of December 31,  

2018 

2017 

 235,294   
 4,225,152   
 4,460,446   

 557,979   
 3,143,566   
 3,701,545   

2016 
 1,167,984  
 1,871,237  
 3,039,221  

The Company has no financial instruments with off-balance sheet risk such as foreign exchange contracts, 
option contracts or other foreign currency hedging arrangements. Financial instruments that potentially subject the 
Company to a concentration of credit risk are cash and cash equivalents. The Company’s cash is held in accounts at a 
financial institution that may exceed federally insured limits. The Company has not experienced any credit losses in such 
accounts and does not believe it is exposed to any significant credit risk on these funds. 

Concentration of Suppliers 

The Company is dependent on a third-party manufacturer to supply certain products for research and 
development activities in its programs. In particular, the Company relies on a sole manufacturer to supply it with specific 
vectors related to the Company’s research and development programs. 

Segment Information 

Operating segments are defined as components of an enterprise about which separate discrete information is 

available for evaluation by the chief operating decision maker in deciding how to allocate resources and assess 
performance. The Company and the Company’s chief operating decision maker, the Company’s Chief Executive Officer, 
views the Company’s operations and manages its business as a single operating segment, which is the business of 
developing and commercializing gene therapies. 

Recently Adopted Accounting Pronouncements 

In January 2016, the Financial Accounting Standards Board (“FASB”) issued ASC Update No. 2016-01, 

Financial Instruments - Overall (Subtopic 825-10): Recognition and Measurement of Financial Assets and Financial 
Liabilities (“Update No. 2016-01”). The purpose of Update No. 2016-01 is to improve financial reporting for financial 
instruments by reducing the number of items recorded to other comprehensive income. The Company adopted Update 
No. 2016-01 in the first quarter of 2018, using the modified retrospective method. Unrealized gains and losses previously 
recorded to other comprehensive income (loss) were reclassified to accumulated deficit and all future fair value changes 
will be recorded to other income (loss). The adoption of the standard on January 1, 2018 did not have a material impact 
on the Company’s consolidated financial statements. 

In August 2016, the FASB issued ASU No. 2016-15, Statement of Cash Flows (“ASU 2016-15”), which 
simplifies certain elements of cash flow classification and is intended to reduce diversity in practice in how certain 
transactions are classified in the statement of cash flows. The update is effective for fiscal years, and interim periods 
within those years, beginning after December 15, 2017. The Company adopted ASU 2016-15 on January 1, 2018, and 
such adoption did not have a material impact on the Company's consolidated financial statements. 

In November 2016, the FASB issued ASU No. 2016-18, Restricted Cash (“ASU 2016-18”). The amendments in 
ASU 2016-18 require an entity to reconcile and explain the period-over-period change in total cash, cash equivalents and 
restricted cash within its statements of cash flows and was adopted utilizing a full retrospective approach. The Company 
F-13 

 
 
 
 
 
 
 
 
 
 
 
 
     
     
     
 
  
  
  
adopted the new standard on January 1, 2018. The Company has included the necessary reconciliation within Note 2 
“Restricted Cash”. 

In May 2014, the FASB issued ASU No. 2014-09, Revenue from Contracts with Customers (“ASU 2014-09”), 

which supersedes the revenue recognition requirements in ASC 605, Revenue Recognition (“ASC 605”), and most 
industry-specific guidance. The new standard requires that an entity recognize revenue to depict the transfer of promised 
goods or services to customers in an amount that reflects the consideration to which the entity expects to be entitled in 
exchange for those goods or services. The update also requires additional disclosure about the nature, amount, timing, 
and uncertainty of revenue and cash flows arising from customer contracts, including significant judgments and changes 
in judgments and assets recognized from costs incurred to obtain or fulfill a contract. Thereafter, a series of clarifying 
ASUs, narrow scope improvements and practical expedients were issued. This collective guidance resulted in the new 
revenue standard, ASC 606.  

The Company adopted the new standard effective January 1, 2018 using the modified retrospective approach. 

The Company had one open contract, relating to the Sanofi Genzyme Collaboration, on the adoption date and has 
assessed it under the new revenue standard. The adoption of ASC 606 resulted in the changes to (i) the allocation of 
arrangement consideration, including the determination of estimated selling price and the allocation of variable 
consideration to specific performance obligations and (ii) the application of proportional performance as a measure of 
progress on service-related deliverables. 

The Company has accounted for the impact of adopting ASC 606 as a cumulative catch-up under the modified 

retrospective approach, which is represented as an increase of $20.0 million to deferred revenue with an offset to 
accumulated deficit, effective January 1, 2018. The following financial statement line items have been shown to reflect 
comparative balances under ASC 606 and ASC 605 for the year ended December 31, 2018, for both of the Sanofi 
Genzyme Collaboration and AbbVie Tau Collaboration, collectively. 

Condensed Consolidated Statements of Operations and Comprehensive Loss 

Year ended December 31, 2018 

Collaboration revenue 
Loss before income taxes 
Net loss 
Net loss per share, basic and diluted 

Condensed Consolidated Balance Sheets 

Deferred revenue, current 
Deferred revenue, non-current 
Accumulated deficit 

$ 

$ 

Under ASC 606 

 Effect of change 

Under ASC 605 
(in thousands, except per share data) 
 7,619   $ 
 (88,468)   
 (88,288)   
 (2.75)   

 11,095   $ 
 (84,992)   
 (84,812)   
 (2.64)   

 (3,476) 
 (3,476) 
 (3,476) 
 (0.11) 

Under ASC 606 

As of December 31, 2018 

Under ASC 605 
(in thousands) 

 Effect of change 

 20,847   $ 
 92,199    
 (269,051)   

 19,111   $ 
 70,529    
 (245,645)   

 1,736  
 21,670  
 (23,406) 

Condensed Consolidated Statements of Cash Flows 

$ 
Net loss 
Adjustments to reconcile net loss to net cash used in operating activities:   

Deferred revenue 

 (88,288) $ 

 (84,812) $ 

 (3,476) 

 61,380    

 57,904    

 3,476  

Year ended December 31, 2018 

Under ASC 606 

  Under ASC 605     Effect of change   
(in thousands) 

F-14 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
    
    
 
 
Recent Accounting Pronouncements 

In February 2016, the FASB issued ASU No. 2016-02, a comprehensive new lease accounting standard, which 

provides revised guidance on accounting for lease arrangements by both lessors and lessees and requires lessees to 
recognize a lease liability and a right-of-use asset for most leases. In July 2018, the FASB issued ASU No. 2018-11, 
Leases (Topic 842), Targeted Improvements, which provides an additional transition method that allows entities to 
initially apply the new lease requirements at the adoption date, not the earliest period presented, and recognize a 
cumulative effect adjustment to the opening balance of retained earnings in the period of adoption. The Company 
expects to elect this transition method at the adoption date of January 1, 2019. The Company also expects to elect a 
package of practical expedients, under which an entity need not reassess whether any expired or existing contracts are or 
contain leases, the lease classification for any expired or existing leases, or initial direct costs for any existing leases. The 
Company is currently in the process of evaluating the impact that the adoption of this guidance will have on its 
consolidated financial statements and related disclosures. 

On December 22, 2017, the SEC staff issued Staff Accounting Bulletin No. 118, to address the application of 

GAAP in situations when a registrant does not have the necessary information available, prepared, or analyzed 
(including computations) in reasonable detail to complete the accounting for certain income tax effects of the Tax Cuts 
and Jobs Act (the “Tax Reform Act”). The Company has recognized the provisional tax impacts related to the 
revaluation of the deferred tax assets and liabilities and included these amounts in its consolidated financial statements 
for the year ended December 31, 2017. The ultimate impact may differ from these provisional amounts due to, among 
other things, additional analysis, changes in interpretations and assumptions the Company has made, additional 
regulatory guidance that may be issued, and actions the Company may take as a result of the Tax Reform Act. The 
accounting was completed when the Company’s 2017 U.S. corporate income tax return was filed in 2018, and no 
material differences arose as compared to provisional amounts initially reflected in the consolidated financial statements 
for the year ended December 31, 2017.  

In June 2018, the FASB issued ASU No. 2018-07, Compensation-Stock Compensation: Improvements to 

Nonemployee Share-Based Payment Accounting (“ASC 718”). The new standard largely aligns the accounting for share-
based payment awards issued to employees and nonemployees by expanding the scope of ASC 718 to apply to 
nonemployee share-based transactions, as long as the transaction is not effectively a form of financing. The new 
guidance will be effective for the Company on January 1, 2019. The Company is currently evaluating the potential 
impact that this guidance may have on its consolidated financial statements. 

In August 2018, the FASB issued ASU No. 2018-13, Fair Value Measurement (Topic 820): Disclosure 
Requirements for Fair Value Measurement (“ASU 2018-13”). The new standard added, modified or removed disclosure 
requirements under Topic 820 for clarity and consistency. ASU 2018-13 is effective for all entities for fiscal years, and 
interim periods within those fiscal years, beginning after December 15, 2019. The Company is currently evaluating the 
potential impact that this guidance may have on its consolidated financial statements.  

F-15 

 
 
3. Fair value measurements 

Assets and liabilities measured at fair value on a recurring basis as of December 31, 2018 and 2017 are as 

follows: 

Assets 

Total 

  Quoted Prices   
in Active 
  Markets for   
Identical Assets  
(Level 1) 

Significant 
Other 
Observable 
Inputs 
(Level 2) 

(in thousands) 

Significant 
Unobservable    
Inputs 
(Level 3) 

December 31, 2018 
Money market funds included in cash and cash 
equivalents 
Marketable securities: 
U.S. Treasury notes 
Equity securities 

Total marketable securities 
Warrants to purchase equity securities 

Total 

December 31, 2017 
Money market funds included in cash and cash 
equivalents 
Marketable securities: 
U.S. Treasury notes 
Equity securities 

Total marketable securities 
Warrants to purchase equity securities 

Total 

  $ 

 46,173 

$ 

 46,173 

$ 

 — 

$ 

 108,947 
 628 
  $   109,575 
 234 
  $   155,982 

 108,947 
 628 
 109,575 
 — 
 155,748 

$ 

$ 

$ 

$ 

 — 
 — 
 — 
 234 
 234 

$ 

$ 

     $ 

 30,469 

$ 

 30,469 

$ 

 — 

$ 

 137,522 
 1,100 
  $   138,622 
 569 
  $   169,660 

 137,522 
 1,100 
 138,622 
 — 
 169,091 

$ 

$ 

$ 

$ 

 — 
 — 
 — 
 569 
 569 

$ 

$ 

 —  

 —  
 —  
 —  
 —  
 — 

 —  

 —  
 —  
 —  
 —  
 —  

The Company measures the fair value of money market funds, U.S. Treasuries and equity securities based on 
quoted prices in active markets for identical securities. The Level 2 equity securities include warrants used to purchase 
equity securities that are valued using the Black-Scholes model. The Black-Scholes option pricing model requires inputs 
based on certain subjective assumptions, including (a) the expected stock price volatility, (b) the calculation of expected 
term of the awards, (c) the risk-free interest rate, and (d) expected dividends. The assumptions utilized to value the 
warrants to purchase equity securities as of December 31, 2018, 2017, and 2016 are as follows: 

Risk-free interest rate 
Expected dividend yield 
Expected term (in years) 
Expected volatility 

As of December 31,  

2018 

2017 

2016 

 2.5  %   
 —  %   
 2.7 
 112.7  %   

 2.0  %   
 —  %   
 3.7 
 103.5  %   

 1.8  %   
 —  %   
 4.7 
 97.5  %   

The expected volatility is based on the historic volatility for the equity securities underlying the warrants and is 

calculated based on a period of time commensurate with the expected term assumption. The expected term is based on 
the remaining contractual life of the warrants on each measurement date. The risk-free interest rate is based on a treasury 
instrument whose term is consistent with the expected term of the warrants. The expected dividend yield is assumed to 
be zero as the entity that issued the warrants has never paid and has not indicated any intention to pay dividends. 

F-16 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
 
 
 
 
 
 
  
 
 
 
 
 
 
 
 
 
 
 
 
     
     
     
     
  
 
 
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
   
4. Prepaid expenses and other current assets 

Prepaid expenses and other current assets consist of the following: 

Prepaid research and development contracts 
Other current assets 
Prepaid insurance 
Accrued interest receivable 

Total 

5. Property and equipment, net 

Property and equipment, net consists of the following: 

Leasehold improvements 
Laboratory equipment 
Furniture and office equipment 
Other 
Construction in progress 
Total property and equipment 
Less: accumulated depreciation 
Property and equipment, net 

As of December 31,  

2018 

2017 

(in thousands) 

     $ 

  $ 

 4,497   $ 
 1,360  
 617  
 201  
 6,675   $ 

 1,330  
 766  
 520  
 122  
 2,738  

As of December 31,  

2018 

2017 

(in thousands) 
 7,035      $ 
 8,843  
 1,675  
 306  
 19  
 17,878  
 (5,107)  
 12,771   $ 

 6,421  
 5,262  
 1,565  
 25  
 —  
 13,273  
 (2,990) 
 10,283  

  $ 

  $ 

The Company recorded $2.1 million, $1.6 million, and $0.6 million in depreciation expense during the years 

ended December 31, 2018, 2017, and 2016, respectively. 

6. Accrued expenses 

Accrued expenses consist of the following: 

Employee compensation costs 
Research and development costs 
Professional services 
Accrued goods and services 
Patent costs 
Other 

Total 

7. Commitments and contingencies 

Operating Leases 

As of December 31,  

2018 

2017 

(in thousands) 
 3,780      $ 
 3,555  
 1,448  
 784  
 120  
 101  
 9,788   $ 

 3,383  
 5,780  
 1,762  
 388  
 120  
 64  
 11,497  

  $ 

$ 

During April 2014, the Company entered into an agreement to lease the 75 Sidney Street facility under a 

non-cancelable operating lease that would have expired December 15, 2019. The lease includes two renewal options, 
each for five-year terms and at fair market value upon exercise. The lease contains escalating rent clauses which require 
higher rent payments in future years. The Company expenses rent on a straight-line basis over the term of the lease, 
including any rent-free periods. 

F-17 

 
 
 
 
 
 
 
 
 
 
  
 
     
     
  
 
 
 
 
 
 
 
  
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
     
     
  
 
 
 
 
  
 
 
  
  
 
 
 
 
 
  
 
  
  
 
  
  
 
 
 
 
 
 
 
 
 
 
 
 
     
     
  
 
 
 
 
  
  
 
  
  
 
  
  
 
 
 
 
  
  
 
 
In December 2015, the Company executed an amendment to the 75 Sidney Street lease to extend its term, and 
executed an agreement to lease a facility at 64 Sidney Street until December 31, 2024. The facility at 64 Sidney Street 
includes laboratory and office space, and was ready for occupancy in early 2017.  

In February 2018, the Company executed a second amendment to the 75 Sidney Street lease to lease additional 

space to support its continued growth. The additional facility includes laboratory and office space, and was ready for 
occupancy in mid-2018.  

In June 2018, the Company executed a third amendment to the 75 Sidney Street lease to lease additional space 

to further support its continued growth. The additional facility includes laboratory and office space, and was ready for 
occupancy in late 2018. The third amendment extended the term of the 75 Sidney Street lease until November 30, 2026. 
Additionally, the Company executed a second amendment to the 64 Sidney Street lease to extend that lease until 
November 30, 2026. 

The Company received leasehold improvement incentives from the landlord totaling $1.3 million and $3.5 

million for 75 Sidney Street and 64 Sidney Street, respectively. The Company recorded these incentives as a component 
of deferred rent and is amortizing these incentives as a reduction of rent expense over the life of the lease. The leasehold 
improvements have been recorded as fixed assets. The Company is entitled to receive approximately $0.3 million of 
leasehold improvements for the additional space at 75 Sidney Street.  

Rent expense of approximately $4.0 million, $2.9 million, and $2.0 million was incurred during the years ended 

December 31, 2018, 2017, and 2016, respectively. 

Future annual minimum lease payments at December 31, 2018 are as follows: 

2019 
2020 
2021 
2022 
2023 
2024+ 

. 

Significant Agreements 

Sanofi Genzyme Collaboration Agreement 

Summary of Agreement 

      Total Minimum 
      Lease Payments 

(in thousands)   
 4,325  
 4,731  
 4,863  
 5,001  
 5,141  
 16,783  
 40,844  

  $ 

In February 2015, the Company entered into an agreement with Sanofi Genzyme (the “Sanofi Genzyme 

Collaboration Agreement”) which included a non-refundable upfront payment of $65.0 million. In addition, 
contemporaneous with entering into the Sanofi Genzyme Collaboration Agreement, Sanofi Genzyme entered into a 
Series B Stock Purchase Agreement, under which Sanofi Genzyme purchased 10,000,000 shares of Series B Preferred 
Stock for $30.0 million. The fair value of the Series B Preferred Stock at the time of issuance was approximately 
$25.0 million. The $5.0 million premium over the fair value is accounted for as additional consideration under the Sanofi 
Genzyme Collaboration Agreement. 

Under the Sanofi Genzyme Collaboration Agreement, the Company granted Sanofi Genzyme an exclusive 

option to license, develop and commercialize (i) ex-U.S. rights to the following programs, which are referred to as Split 
Territory Programs; VY-AADC (“Parkinson’s Program”), VY-FXN01 (“Friedreich’s ataxia Program”), a future program 
to be designated by Sanofi Genzyme (“Future Program), and VY-HTT01 (“Huntington’s Program”), with an incremental 
option to co-commercialize VY-HTT01 in the United States and (ii) worldwide rights to VY-SMN101 (“Spinal 
Muscular Atrophy Program”). Sanofi Genzyme’s option for the Split Territory Programs and the Spinal Muscular 

F-18 

 
 
 
 
 
 
  
 
  
 
 
 
  
 
  
 
  
 
 
 
 
 
 
 
Atrophy Program is triggered following the completion of the first proof-of-principle human clinical study (“POP 
Study”), on a program by program basis. 

Prior to any option exercise by Sanofi Genzyme, the Company will collaborate with Sanofi Genzyme in the 
development of products under each Split Territory Program and the Spinal Muscular Atrophy Program pursuant to a 
written development plan and under the guidance of an Alliance Joint Steering Committee (“AJSC”), comprised of an 
equal number of employees from the Company and Sanofi Genzyme. 

The Company is required to use commercially reasonable efforts to develop products under each Split Territory 

Program and the Spinal Muscular Atrophy Program through the completion of the applicable POP Study. During the 
development of these joint programs, the activities are guided by a Development Advisory Committee (“DAC”). The 
DAC may elect to utilize certain Sanofi Genzyme technology relating to the Parkinson’s Program, the Huntington’s 
Program or generally with the manufacture of Split Territory Program products. 

The Company is solely responsible for all costs incurred in connection with the development of the Split 

Territory Programs and the Spinal Muscular Atrophy Program products prior to the exercise of an option by Sanofi 
Genzyme with the exception of the following: (i) at the Company’s request and upon mutual agreement, Sanofi 
Genzyme will provide “in-kind” services valued at up to $5.0 million and (ii) Sanofi Genzyme shall be responsible for 
the costs and expenses of activities under the Huntington’s Program development plan to the extent such activities are 
covered by financial support Sanofi Genzyme is entitled to receive from a patient advocacy group, collectively Sanofi 
Genzyme “in-kind” and other funding. 

Other than the Parkinson’s Program (for which a POP Study has already been completed), if the Company does 

not initiate a POP Study for a given Split Territory Program by December 31, 2026 (or for the Future Program by the 
tenth anniversary of the date the Future Program is nominated by Sanofi Genzyme), and Sanofi Genzyme has not 
terminated the Sanofi Genzyme Collaboration Agreement with respect to the collaboration program, then Sanofi 
Genzyme shall be entitled, as its sole and exclusive remedy, to a credit of $10.0 million for each such program against 
other milestone or royalty payments payable by Sanofi Genzyme under the Sanofi Genzyme Collaboration Agreement. 
However, if the POP Study is not initiated due to a regulatory delay or a force majeure event, such time period shall be 
extended for so long as such delay continues. 

With the exception of the Parkinson’s Program, Sanofi Genzyme is required to pay an option exercise payment 

of $20.0 million or $30.0 million for each Split Territory Program, as well as the Spinal Muscular Atrophy Program. 

Upon Sanofi Genzyme’s exercise of its option to license a given product in a Split Territory Program (“Split 

Territory Licensed Product”), the Company will have sole responsibility for the development of such Split Territory 
Licensed Product in the United States and Sanofi Genzyme shall have sole responsibility for development of such Split 
Territory Licensed Product in the rest of the world. The Company and Sanofi Genzyme will have shared responsibility 
for execution of ongoing development of such Split Territory Licensed Product that is not specific to either territory, 
including costs associated therewith. The Company is responsible for all commercialization activities relating to Split 
Territory Licensed Products in the United States, including all of the associated costs. Sanofi Genzyme is responsible for 
all commercialization activities relating to the Split Territory Licensed Products in the rest of the world, including all of 
the associated costs. If Sanofi Genzyme exercised its co-commercialization rights, Sanofi Genzyme will be the lead party 
responsible for all commercialization activities related to the Huntington’s Licensed Product in the United States. 

Upon exercise of the option, Sanofi Genzyme shall have the sole right to develop the Spinal Muscular Atrophy 

Product worldwide. Sanofi Genzyme shall be responsible for all of the development costs that occur after the option 
exercise date for the Spinal Muscular Atrophy Program. Sanofi Genzyme is also responsible for commercialization 
activities relating to the Spinal Muscular Atrophy Product worldwide. 

Sanofi Genzyme is required to pay the Company for specified regulatory and commercial milestones, if 

achieved, up to $540.0 million across all programs. The Company is no longer entitled to receive a total of $105.0 
million related to regulatory and commercial milestone payments for VY-AADC as a result of Sanofi Genzyme’s 
decision to not exercise its option for the Parkinson’s Program (the “PD Opt-Out”). The regulatory approval milestones 
are payable upon either regulatory approval in the United States or regulatory and reimbursement approval in the 
European Union and range from $40.0 million to $50.0 million per milestone, with an aggregate total of $220.0 million, 
after accounting for the PD Opt-Out. The commercial milestones are payable upon achievement of specified annual net 

F-19 

sales in each program and range from $50.0 million to $100.0 million per milestone, with an aggregate total of 
$320.0 million, after accounting for the PD Opt-Out. 

In addition, to the extent any Split Territory Licensed Products or the Spinal Muscular Atrophy Licensed 

Product are commercialized, the Company is entitled to tiered royalty payments ranging from the mid-single digits to 
mid-teens based on a percentage of net sales by Sanofi Genzyme. Sanofi Genzyme is entitled to receive tiered royalty 
payments related to sales of the Split Territory Licensed Product ranging from the low-single digits to mid-single digits 
based on a percentage of net sales by the Company depending on whether the Company uses Sanofi Genzyme 
technology in the Split Territory Licensed Product. If Sanofi Genzyme elects to co-commercialize VY-HTT01 in the 
United States, the Company and Sanofi Genzyme will share in any profits or losses from VY-HTT01 product sales. 

The Sanofi Genzyme Collaboration Agreement will continue in effect until the later of (i) the expiration of the 

last to expire of the option rights and (ii) the expiration of all payment obligations unless sooner terminated by the 
Company or Sanofi Genzyme. The Company and Sanofi Genzyme have customary termination rights including the right 
to terminate for an uncured material breach of the agreement committed by the other party and Sanofi Genzyme has the 
right to terminate for convenience. 

Accounting Analysis 

At inception, the Sanofi Genzyme Collaboration Agreement included the following performance obligations: (i) 

research and development services for each of the Split Territory Programs and the Spinal Muscular Atrophy Program, 
(ii) participation in the AJSC, (iii) participation in the DAC and (iv) a material right associated with an option to obtain a 
development and commercial license in the Parkinson’s Program (“PD Material Right”). The Company determined that 
the option to obtain a development and commercial license in the Parkinson’s Program was a material right under ASC 
606 primarily because there were no additional option exercise payments payable by Sanofi Genzyme at the time of 
option exercise. Therefore, the PD Material Right was considered a performance obligation at the inception of the 
arrangement. The options in the other Split Territory Programs and the Spinal Muscular Atrophy Program do not provide 
a material right to the customer that it would receive without entering into the contract principally because the option 
fees are at least equal to the standalone selling price for the underlying goods. Therefore, the other Split Territory 
Programs and the Spinal Muscular Atrophy Program options are not performance obligations at inception. 

The Company has identified $74.6 million of total transaction price consisting of the $65.0 million upfront fee, 

the $5.0 million premium paid in excess of fair value of the Series B Preferred Stock and $4.6 million of Sanofi 
Genzyme “in-kind” funding, which represents the transaction price at adoption. Additional consideration to be paid to 
the Company upon the exercise of the license options by Sanofi Genzyme or upon reaching certain milestones are 
excluded from the transaction price as they relate to option fees and milestones that can only be achieved subsequent to 
the option exercise or are outside of the initial contract term. 

The Company has allocated the transaction price to the separate performance obligations based on their relative 

standalone selling price. For all performance obligations, the Company determined the standalone selling price at 
contract inception based on each obligation’s estimated standalone selling price (“ESP”). The Company determined the 
ESP for the service related deliverable for the research and development activities based on internal estimates of the 
costs to perform the services, including expected internal expenses and expenses with third parties for services and 
supplies, marked up to include a reasonable profit margin and adjusted for the scope of the potential license. Significant 
inputs used to determine the total expense of the research and development activities include, the length of time required 
and the number and costs of various studies that will be performed to complete the applicable POP Study. The ESP for 
the AJSC and DAC have been estimated based on the costs incurred to participate in the committees, marked up to 
include a reasonable profit margin. The ESP for the PD Material Right was determined based on the estimated value of 
the license adjusted for the estimated probability that the option would be exercised by Sanofi Genzyme. 

F-20 

Based on the relative standalone selling price allocation, the allocation of the transaction price to the separate 

performance obligations was as follows: 

Unit of Accounting 

Research and Development Services for: 

Huntington’s Program 
Parkinson’s Program 
Friedreich’s ataxia Program 
Spinal Muscular Atrophy Program 
Future Program 
Committee Obligations: 

AJSC 
DAC 

PD Material Right 

Total 

Amount 
  (in thousands) 

  $ 

  $ 

 14,228  
 6,040  
 14,821  
 29,116  
 2,239  

 133  
 207  
 7,855  
 74,639  

The Company recognizes the amounts associated with research and development services and committee 
obligations on a proportional performance basis over the period of service using input-based measurements such as costs 
incurred to date, to estimate proportion performed, and remeasures its progress towards completion at the end of each 
reporting period. The amount allocated to the PD Material Right was initially deferred and recognized in full prior to the 
adoption of ASC 606. 

In October 2017, Sanofi Genzyme decided not to exercise the PD Material Right. Therefore, in the year ended 
December 31, 2017, the Company recognized revenue of $7.8 million of consideration which had been allocated to the 
PD Material Right. In addition, revenue recognized during the years ended December 31, 2017 and 2016 include 
amounts recognized related to consideration allocated to research and development services for various programs under 
the Sanofi Genzyme Collaboration Agreement. During 2017 the Company reassessed the estimated period of 
performance for each of the units of accounting and determined that the estimated period would be extended for two 
units of accounting. During 2016 the Company deprioritized the development of VY-SMN101 for the treatment of 
Spinal Muscular Atrophy. As a result, the Company ceased recognizing the revenue allocated to this program. These 
adjustments were made on a prospective basis and resulted in decreases in revenue recognized by $2.1 million and $9.5 
million, respectively, for the year ended December 31, 2017.  

During the years ended December 31, 2018, 2017, and 2016, the Company recognized $0.7 million, $10.1 

million, and $14.2 million, respectively, of revenue associated with its collaboration with Sanofi Genzyme related to 
research and development services performed during the period and for consideration allocated to the PD Material Right, 
which was recognized during 2017. As of December 31, 2018, there is $50.9 million of deferred revenue related to the 
Sanofi Genzyme Collaboration Agreement, which is classified as either current or noncurrent in the accompanying 
balance sheet based on the period the services are expected to be delivered. 

Costs incurred relating to the programs that Sanofi Genzyme has the option to license under the Sanofi 

Genzyme Collaboration Agreement consist of internal and external research and development costs, which primarily 
include: salaries and benefits, lab supplies and preclinical research studies. All costs are included in research and 
development expenses in the Company’s statement of operations during the years ended December 31, 2018, 2017, and 
2016. 

AbbVie Tau Collaboration Agreement 

Summary of Agreement 

In February 2018, the Company entered into an exclusive collaboration and option agreement (the “AbbVie Tau 

Collaboration Agreement”) with AbbVie for the research, development and commercialization of AAV and other virus-
based gene therapy products for the treatment of diseases of the central nervous system and other neurodegenerative 
diseases related to defective or excess aggregation of tau protein in the human brain, including Alzheimer’s disease. 

F-21 

 
 
 
 
 
 
    
  
 
 
 
 
 
 
  
 
  
 
  
 
  
 
 
 
 
 
  
 
  
 
  
 
Under the AbbVie Tau Collaboration Agreement, the Company and AbbVie have agreed to collaborate on the research 
and development of specified vectorized antibody compounds comprised of an AAV or other viral capsid and a virus 
vector genome that encodes one or more antibodies that target and bind to a tau protein. The collaboration is comprised 
of a research period (the “Research Period”), a development period (the “Development Period”), and an exclusive license 
option (the “License Option”). The AbbVie Tau Collaboration Agreement included a non-refundable upfront payment of 
$69.0 million for services during the Research Period. 

During the Research Period, each party has agreed to identify up to five antibodies for inclusion in the 
collaboration. Subject to certain conditions and exceptions, the parties will select up to three antibodies (each, a 
“Research Antibody”) as candidates for creation of research compounds (each, a “Research Compound”), with AbbVie 
having the right to select two of the three Research Antibodies. The Company is required to use diligent efforts to 
conduct antibody engineering and other research activities to create Research Compounds and to develop product 
candidates containing or comprised of such Research Compounds (“Product Candidates”). The Company is solely 
responsible for its costs and expenses during the Research Period. During a specified portion of the Research Period, 
AbbVie may exercise one or more of its exclusive development options (each, a “Development Option”) to select up to a 
total of three Research Compounds (the “Selected Research Compounds”) and their corresponding Product Candidates 
(the “Selected Product Candidates”) to proceed to the Development Period. 

Upon AbbVie’s exercise of a Development Option, AbbVie will pay the Company $80.0 million for the first 

Selected Research Compound and $30.0 million each for up to two additional Selected Research Compounds. During the 
Development Period, the Company is obligated to use diligent efforts to conduct development activities, including 
Investigational New Drug application-enabling and Phase 1 clinical trial activities, for the Selected Research Compounds 
and corresponding Selected Product Candidates. The Company will be solely responsible for the costs and expenses 
during the Development Period. During a specified portion of the Development Period (the “License Option Period”), 
AbbVie may exercise its License Option to further develop and commercialize all of the Research Compounds (the 
“Licensed Compounds”), and corresponding product candidates (the “Licensed Products”). Upon AbbVie’s exercise of 
its License Option, AbbVie will provide a one-time payment of $75.0 million to the Company, and the Company will 
grant to AbbVie an exclusive, worldwide license, with the right to sublicense, under certain of the Company’s 
intellectual property rights to develop and commercialize the Licensed Compounds and the Licensed Products for all 
human diagnostic, prophylactic and therapeutic uses. In addition, after AbbVie’s exercise of the License Option, the 
Company has certain obligations to complete any remaining research and development activities that have not been 
completed for any Research Compounds and Product Candidates. 

The Company’s research and development activities will be conducted pursuant to the plans agreed to by the 

parties and overseen by a joint governance committee (“JGC”) as detailed in the AbbVie Tau Collaboration Agreement. 
Any material amendment to the research or development plans must be mutually agreed to by the Company and AbbVie, 
which may be through the JGC. 

Under the AbbVie Tau Collaboration Agreement, AbbVie is required to use commercially reasonable efforts to 

develop and commercialize at least one Licensed Product in each of the United States, Japan, the United Kingdom, 
Germany, France, Italy, and Spain. After exercise of the License Option, AbbVie is solely responsible for all 
development and commercialization activities relating to Licensed Compounds and Licensed Products at its sole cost and 
expense, subject to the agreed-upon research and development plans. The Company may elect to share in AbbVie’s 
development costs relating to a Licensed Product on an indication-by-indication basis in exchange for a specified 
increase in royalties (a “Cost-Sharing Option”). If the Company exercises a Cost-Sharing Option, the Company may 
either reimburse AbbVie for AbbVie’s applicable development costs or, in the case of certain budget overruns, AbbVie 
may instead deduct applicable development costs, up to a specified cap, from milestone and royalty payments owed by 
AbbVie to the Company. 

Under the AbbVie Tau Collaboration, the Company is eligible to receive specified development and first-sale 
milestone payments for each Licensed Compound of up to an aggregate of $550.0 million in the case of an Alzheimer’s 
disease indication, up to $230.0 million in the case of the first indication other than Alzheimer’s disease and up to $115.0 
million for a subsequent non-Alzheimer’s disease indication. Additionally, the Company is eligible to receive tiered, 
escalating royalties, in a range from a high-single digit to a mid-to-high teen (or, if the Company has exercised its Cost-
Sharing Option, low-twenties) percentage of aggregate net sales of Licensed Products on a Licensed Compound by 
Licensed Compound basis, subject to potential reductions in certain circumstances. For each Licensed Product, AbbVie 

F-22 

  
 
  
  
  
also has the right to decrease or eliminate its royalty payments on such Licensed Product in exchange for a one-time 
payment by AbbVie at a fair market value to be negotiated by the parties or determined pursuant to dispute resolution 
procedures specified in the AbbVie Tau Collaboration Agreement. 

Unless earlier terminated, the AbbVie Tau Collaboration Agreement will expire on the earliest to occur of the 
expiration of (i) the Development Option Period, without AbbVie’s exercise of a Development Option; (ii) the License 
Option Period, without AbbVie’s exercise of its License Option; and (iii) the last-to-expire royalty term with respect to 
all Licensed Products in all countries. The Company and AbbVie have customary termination rights including the right 
to terminate for an uncured material breach of the agreement committed by the other party, and AbbVie has the right to 
terminate for convenience.  

Accounting Analysis 

The Company assessed the promised goods and services under the AbbVie Tau Collaboration Agreement, in 

accordance with ASC 606, and determined that the AbbVie Tau Collaboration Agreement includes the following 
performance obligations: (i) research services during the Research Period (through the delivery of the final research 
report) including the identification of the Research Antibodies, conduct of research activities and provision of 
information to AbbVie to allow AbbVie to determine whether to exercise up to three Development Options to be 
rendered (collectively, the “Research Services”), and (ii) a material right associated with the Development Option on the 
first Research Compound and associated Product Candidates (“First Development Option Material Right”). The first 
Development Option provides AbbVie with (i) additional development services on a selected Research Compound and 
(ii) the ability to exercise the License Option. The Company has concluded the option provides a material right as the 
consideration paid by AbbVie upon exercise of the first Development Option is less than the amount that the Company 
would otherwise expect to receive outside the context of the contract.  

The Company has concluded that the First Development Option Material Right is a separate performance 

obligation under ASC 606 as AbbVie is provided additional services and a License Option for additional consideration 
that represents a significant discount from amounts that would otherwise be offered outside the context of the contract. 
The First Development Option Material Right is distinct from the other performance obligations in the arrangement as it 
is an option in the contract that is not required for AbbVie to obtain the benefit of the other promised goods or services in 
the arrangement. The First Development Option Material Right does not include the underlying goods or services that 
are delivered upon exercise of the option, but rather represents the value to the customer of having the right to obtain 
development services and the right to the License Option at an advantageous price. 

The Company received a nonrefundable, upfront payment of $69.0 million as consideration under the AbbVie 

Tau Collaboration Agreement, which represents the transaction price at inception. Additional consideration to be paid to 
the Company upon the exercise of the Development and License Options by AbbVie or upon reaching certain milestones 
are excluded from the transaction price as they relate to option fees and milestones that can only be achieved subsequent 
to the option exercise or are outside of the initial contact term. 

The Company has allocated the transaction price to the separate performance obligations based on their relative 

standalone selling price. The Company determined the standalone selling price at contract inception based on each 
obligation’s ESP. The Company determined the ESP for the research services obligation based on internal estimates of 
the costs to perform the services, including expected internal expenses and expenses with third parties for services and 
supplies, inclusive of a reasonable profit margin. Significant inputs used to determine the total expense of the research 
services include the length of time required, the internal hours expected to be incurred on the services and the number 
and costs of various studies that will be performed to complete the Research Plan. The ESP for the First Development 
Option Material Right was determined based on the fees AbbVie would pay to exercise the Development and License 
Options, the estimated costs to perform the development services, inclusive of a reasonable profit margin, the estimated 
value of the License Option using comparable transactions, and the probability that the Development and License 
Options would be exercised by AbbVie. 

Based on the relative standalone selling price, the allocation of the transaction price to the separate performance 

obligations was as follows: 

Performance Obligation 

Amount 

F-23 

  
 
 
 
 
 
 
 
 
Research Services 
First Development Option Material Right 

Total 

  $ 

  $ 

(in thousands) 

 34,482  
 34,518  
 69,000  

The Company recognizes the amounts associated with Research Services on a proportional performance basis 

over the period of service using input-based measurements of total cost of research incurred to estimate proportion 
performed and remeasures its progress towards completion at the end of each reporting period. The amount allocated to 
the First Development Option Material Right is recorded as deferred revenue and will be recognized either over the 
period in which goods and services underlying the option are transferred or upon expiry of the option. 

During the year ended December 31, 2018, the Company recognized $6.9 million of revenue associated with 

the AbbVie Tau Collaboration related to the Research Services performed during the period. As of December 31, 2018, 
there is $62.1 million of deferred revenue related to the AbbVie Tau Collaboration Agreement, which is classified as 
either current or noncurrent in the accompanying balance sheet based on the period the goods or services are expected to 
be delivered. 

Costs incurred relating to the AbbVie Tau Collaboration Agreement consist of internal and external research 

and development costs, which primarily include salaries and benefits, lab supplies, and preclinical research studies. All 
of these costs are included in research and development expenses in the Company’s statement of operations during the 
year ended December 31, 2018. 

MRI Interventions License and Securities Purchase Agreements 

Summary of Agreement 

In September 2016, the Company entered into a securities purchase agreement (the “Securities Purchase 
Agreement”) and a license agreement (the “MRIC License Agreement”) with MRI Interventions, Inc. (“MRIC”). MRIC 
is the primary supplier of the ClearPoint® System, which is being used by the Company in ongoing development and 
clinical trials. Under the Securities Purchase Agreement, the Company paid $2.0 million for shares of MRIC common 
stock and a warrant to purchase additional shares of MRIC common stock. The Company also entered into the MRIC 
License Agreement, which provided for certain rights to MRIC technology and for MRIC to transfer the rights and 
know-how to manufacture the ClearPoint System to enable the Company to utilize an alternative supplier for the 
ClearPoint System for use in the Company’s development and clinical trials. During 2017, the Company terminated the 
MRIC License Agreement and all prior and future commitments and obligations under such agreement became null and 
void. The Company continues to hold the common stock and warrants to purchase additional shares of common stock as 
an available-for-sale security and non-current asset, respectively. 

In May 2018, the Company entered into a master services and supply agreement with MRIC (the “MRIC 

Supply Agreement”) which provides for MRIC to perform certain manufacturing, supply, development and services as 
requested by the Company, including the supply of the ClearPoint System and cannula devices.  

As of December 31, 2018, the Company continued to hold the common stock and warrants to purchase 

additional shares of common stock as an available-for-sale security and non-current asset, respectively. 

Other Agreements 

During 2018, 2017, and 2016, the Company entered into various agreements with contract research 

organizations and institutions to license intellectual property. In consideration for the licensed rights the Company 
generally made upfront payments, which were recorded as research and development expense as the acquired 
technologies were considered in-process research and development. The license agreements obligate the Company to 
make additional payments that are contingent upon specific clinical trial and regulatory approval milestones being 
achieved as well as royalties on future product sales. The agreements to license intellectual property include potential 
milestone payments that are dependent upon the development of products licensed under the agreements and contingent 
upon the achievement of clinical trial or regulatory approval milestones. As of December 31, 2018 and 2017, there have 

F-24 

 
 
 
 
 
 
been no milestones achieved. The Company can generally terminate the license agreements upon 30-90 days prior 
written notice. 

Additionally, certain license agreements require the Company to reimburse the licensor for certain past and 

ongoing patent related expenses. During the year ended December 31, 2018, 2017, and 2016, the Company incurred $0.6 
million, $0.8 million, and $1.8 million of expense, respectively, related to these reimbursable patent costs which are 
recorded as general and administrative expense 

During the year ended December 31, 2016, the Company entered into a research and development funding 

arrangement with a non-profit organization that provides up to $4.0 million in funding to the Company upon the 
achievement of clinical and development milestones. The agreement provides that the Company repay amounts received 
under certain circumstances including termination of the agreement, and to pay an amount up to 2.6 times the funding 
received upon successful development and commercialization of any products developed. During the year ended 
December 31, 2017, the Company earned a milestone payment of $1.0 million. The Company has evaluated the 
arrangement and has concluded that it represents a research and development financing arrangement as it is probable that 
the Company will repay amounts received under the arrangement. As a result, the $1.0 million earned through the year 
ended December 31, 2017 is recorded as a non-current liability in the consolidated balance sheet. 

Litigation 

The Company is not a party to any material legal matters or claims and does not have contingency reserves established 
for any litigation liabilities as of December 31, 2018 or 2017. 

8. Preferred stock 

The Company has authorized preferred stock amounting to 5,000,000 shares as of December 31, 2018 and 

2017. The authorized preferred stock was classified under stockholders’ equity at December 31, 2018. 

9. Common stock 

As of December 31, 2018 and 2017, the Company had authorized 120,000,000 shares of common stock, at 

$0.001 par value per share. 

General 

The voting, dividend and liquidation rights of the holders of the common stock are subject to and qualified by 

the rights, powers and preferences of the holders of preferred stock. The common stock has the following characteristics: 

Voting 

The holders of shares of common stock are entitled to one vote for each share of common stock held at all 

meetings of stockholders and written actions in lieu of meetings. 

Dividends 

The holders of shares of common stock are entitled to receive dividends, if and when declared by the Board of 

Directors. No dividends have been declared or paid by the Company since its inception. 

Liquidation 

The holders of shares of common stock are entitled to share ratably in the Company’s remaining assets 

available for distribution to its stockholders in the event of any voluntary or involuntary liquidation, dissolution or 
winding up of the Company or upon occurrence of a deemed liquidation event. 

F-25 

Shares Reserved For Future Issuance 

Shares reserved for vesting of restricted stock awards under the Founder 
Agreements 
Shares reserved for vesting of restricted stock awards under the 2014 Option and 
Stock Plan 
Shares reserved for exercise of outstanding stock options 
Shares reserved for issuances under the 2015 Stock Option Plan 
Shares reserved for issuances under the 2015 Employee Stock Purchase Plan 

As of December 31,  

2018 

2017 

 235,294  

 366,914  

 —  
 4,225,152  
 1,973,227  
 963,386  
 7,397,059  

 191,065   
 3,143,566   
 1,501,005  
 730,860  
 5,933,410   

10. Stock-based compensation 

2014 Stock Option and Grant Plan 

In January 2014, the Company adopted the 2014 Stock Option and Grant Plan (the “2014 Plan”), under which it 

could grant incentive stock options, non-qualified stock options, restricted stock awards, unrestricted stock awards, or 
restricted stock units to purchase up to 823,529 shares of Common Stock to employees, officers, directors and 
consultants of the Company. 

In April 2014, the Company amended the Plan to allow for the issuance of up to 1,411,764 shares of Common 
Stock. In August 2014, April 2015, August 2015 and October 2015 the Company further amended the Plan to allow for 
the issuance of up to 2,000,000, 2,047,058, 2,669,411 and 2,998,823 shares of Common Stock, respectively. During 
2014 the Company issued only restricted stock awards under the Plan and during 2015 the Company only granted stock 
options. 

The terms of stock option agreements, including vesting requirements, were determined by the Board of 
Directors and were subject to the provisions of the 2014 Plan. Restricted stock awards granted by the Company generally 
vest based on each grantee’s continued service with the Company during a specified period following grant. Stock 
options granted to employees generally vest over four years, with 25% vesting on the one year anniversary and 75% 
vesting ratably, on a monthly basis, over the remaining three years. Stock options granted to non-employee consultants 
generally vest monthly over a period of one to four years. 

Founder Awards 

In January 2014, the Company issued 1,188,233 shares of restricted stock to its Founders at an original issuance 
price of $0.0425 per share. Of the total restricted shares awarded to the Founders, 835,292 shares generally vest over one 
to four years, based on each Founder’s continued service to the Company in varying capacity as a Scientific Advisory 
Board member, consultant, director, officer or employee, as set forth in each grantee’s individual restricted stock 
purchase agreement.  

The remaining 352,941 of the shares issued will begin vesting upon the achievement of certain performance 

objectives as well as continued service to the Company, as set forth in the agreements. These performance conditions are 
tied to certain milestone events specific to the Company’s corporate goals, including but not limited to preclinical and 
clinical development milestones related to the Company’s product candidates. Stock-based compensation expense 
associated with these performance-based awards will be recognized when the achievement of the performance condition 
is considered probable, using management’s best estimates. Management concluded that the achievement of the 
performance milestone for one of the three performance-based awards had been met during 2016. Accordingly, 
stock-based compensation expense in the amount of $0.3 million, $1.4 million, and $1.1 million was recorded in the 
years ended December 31, 2018, 2017, and 2016, respectively. 

2015 Stock Option Plan  

In October 2015, the Company’s board of directors and stockholders approved the 2015 Stock Option and 

Incentive Plan “(2015 Stock Option Plan”), which became effective upon the completion of the IPO. The 2015 Stock 
Option Plan provides the Company with the flexibility to use various equity-based incentive and other awards as 
compensation tools to motivate its workforce. These tools include stock options, stock appreciation rights, restricted 
stock, restricted stock units, unrestricted stock, performance share awards and cash-based awards. The 2015 Stock 

F-26 

 
 
 
 
 
 
 
 
 
 
     
     
  
  
  
  
 
 
 
  
 
 
 
 
Option Plan replaced the 2014 Plan. Any options or awards outstanding under the 2014 Plan remained outstanding and 
effective. The number of shares initially reserved for issuance under the 2015 Stock Option Plan is the sum of 
(i) 1,311,812 shares of common stock and (ii) the number of shares under the 2014 Plan that are not needed to fulfill the 
Company’s obligations for awards issued under the 2014 Plan as a result of forfeiture, expiration, cancellation, 
termination or net issuances of awards thereunder. The number of shares of common stock that may be issued under the 
2015 Stock Option Plan is also subject to increase on the first day of each fiscal year by up to 4% of the Company’s 
issued and outstanding shares of common stock on the immediately preceding December 31.  

Effective January 1, 2016, 2017, 2018 and 2019, an additional 1,069,971, 1,070,635, 1,285,200, and 1,302,830 
shares of common stock, respectively, were added to the Company’s 2015 Stock Option Plan pursuant to its “evergreen” 
provision, for future issuance. During the year ended December 31, 2018, the Company granted options to purchase 
2,215,891 shares of common stock to employees and directors. As of December 31, 2018, there were 1,973,227 shares 
available for future issuance under the 2015 Stock Option Plan. 

2015 Employee Stock Purchase Plan  

In October 2015, the Company’s board of directors and stockholders approved the 2015 Employee Stock 

Purchase Plan (the “2015 ESPP”). Under the 2015 ESPP, all full-time employees of the Company are eligible to 
purchase common stock of the Company twice per year, at the end of each six-month payment period. During each 
payment period, eligible employees who so elect, may authorize payroll deductions in an amount of 1% to 10% (whole 
percentages only) of the employee’s base pay for each payroll period. At the end of each payment period, the 
accumulated deductions are used to purchase shares of common stock from the Company at a discount. A total of 
262,362 shares of common stock were initially authorized for issuance under this plan. The 2015 ESPP became effective 
upon the completion of the IPO. Effective January 1, 2016, 2017, 2018, and 2019, a total of 267,492, 267,658, 321,300, 
and 325,707 shares of common stock, respectively, were added to the 2015 ESPP, pursuant to its evergreen provision, 
for future issuance. The Company issued 88,774 shares of common stock under the 2015 ESPP in the year ended 
December 31, 2018.  

Stock-based Compensation Expense 

Total compensation cost recognized for all stock-based compensation awards in the statements of operations 

and comprehensive loss is as follows: 

2018 

Year ended December 31,  
2017 
(in thousands) 

2016 

Research and development 
General and administrative 

Total stock-compensation expense 

  $ 

$ 

 4,717   $ 
 10,993  
 15,710   $ 

 5,367   $ 
 3,871  
 9,238   $ 

 4,296  
 2,014  
 6,310  

Restricted Stock 

A summary of the status of and changes in unvested restricted stock was as follows: 

Unvested restricted common stock as of December 31, 2017   
Issued 
Vested 
Repurchased 
Unvested restricted common stock as of December 31, 2018  

      Weighted 
Average 
Grant Date 
Fair Value 
Per Share 

Shares 
 557,979   $ 
 —  
 (319,891)  $ 
 (2,794)  $ 
 235,294   $ 

 0.70  

 0.84  
 1.11  
 0.51  

The expense related to awards granted to employees and non-employees was $0.1 million and $0.4 million, 

respectively, for the year ended December 31, 2018. The expense related to awards granted to employees and non-

F-27 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
     
    
    
  
 
 
  
 
  
  
  
 
 
 
 
 
 
 
 
     
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
     
     
 
 
 
 
 
 
employees was $0.5 million and $2.7 million, respectively, for the year ended December 31, 2017. The expense related 
to awards granted to employees and non-employees was $0.5 million and $2.6 million, respectively, for the year ended 
December 31, 2016. 

Stock Options 

A summary of the status of, and changes in, stock options was as follows: 

Outstanding at December 31, 2017 

Granted 
Exercised 
Cancelled or forfeited 

Outstanding at December 31, 2018 
Exercisable at December 31, 2018 
Vested and expected to vest at December 31, 2018 

     Weighted       Remaining       Aggregate 
Intrinsic 
  Contractual   
Value 
     (in thousands)  

Life 
(in years) 

Shares 

 3,143,566   $ 
 2,215,891   $ 
 (384,186)  $ 
 (750,119)  $ 
 4,225,152   $ 
 1,531,832   $ 
 4,225,152   $ 

Average 
Exercise 
Price 
 11.82   
 20.78  
 10.13  
 16.13  
 15.91  
 12.55   
 15.91  

 8.4   $ 
 7.3   $ 
 8.4   $ 

 578 
 507 
 578 

Using the Black-Scholes option pricing model, the weighted average fair value of options granted to employees 

and directors during the year ended December 31, 2018 was $13.87. The stock-based compensation expense related to 
stock option awards granted to employees and directors was $14.9 million, $5.5 million, and $3.0 million for the years 
ended December 31, 2018, 2017, and 2016, respectively.  

The fair value of each option issued to employees and directors was estimated at the date of grant using the 

Black-Scholes option pricing model with the following weighted-average assumptions: 

Risk-free interest rate 
Expected dividend yield 
Expected term (in years) 
Expected volatility 

Year ended December 31,  

2018 

2017 

2016 

 2.8 %     
 — %    
 6.0  
 74.4 %     

 2.0 %     
 — %    
 6.0  

 73.7 %     

 1.5 %    
 — %   
 6.0  

 73.1 %    

There were no new options granted to non-employees during the year ended December 31, 2018. Unvested 

options granted to non-employees are revalued at each measurement period until they vest. The expense related to stock 
option awards granted to non-employees was $0.3 million, $0.4 million, and $0.2 million for the years ended 
December 31, 2018, 2017, and 2016, respectively. As of December 31, 2018, the Company had unrecognized 
stock-based compensation expense related to its unvested stock options of $26.1 million which is expected to be 
recognized over the remaining weighted average vesting period of 2.78 years. 

The fair value of each option issued to non-employees was estimated at each vesting and reporting date using 

the Black-Scholes option pricing model. The reporting date fair value was determined using the following 
weighted-average assumptions: 

Risk-free interest rate 
Expected dividend yield 
Expected term (in years) 
Expected volatility 

As of December 31,  

2018 

2017 

2016 

 2.6 %   
 — %   
 7.6  
 73.2 %   

 2.4 %   
 — %   
 8.5  
 76.2 %   

 2.1 %  
 — %  
 9.1  
 83.3 %  

F-28 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
     
    
    
  
  
 
 
  
 
 
  
 
 
  
 
 
  
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
    
    
     
 
  
  
  
  
  
 
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
     
    
     
 
  
  
  
  
  
 
  
 
 
11. 401(k) Savings plan  

The Company has a defined-contribution savings plan under Section 401(k) of the Internal Revenue Code (the 

“401(k) Plan”). The 401(k) Plan covers all employees who meet defined minimum age and service requirements, and 
allows participants to defer a portion of their annual compensation on a pretax basis. The Company expensed 
approximately $0.8 million, $0.5 million, and $0.3 million related to employer contributions made during the years 
ended December 31, 2018, 2017, and 2016, respectively. 

12. Income taxes  

The Company accounts for income taxes using the asset and liability method. Under the asset and liability 
method, deferred tax assets and liabilities are recognized for the future tax consequences attributable to temporary 
differences between the financial statement carrying amounts of existing assets and liabilities and their respective tax 
bases and for tax carryforwards, such as net operating losses. Deferred tax assets and liabilities are measured using 
enacted tax rates expected to apply to taxable income in the years in which those temporary differences are expected to 
be recovered or settled. The effect on deferred tax assets and liabilities of a change in tax rates is recognized in the 
provision for income taxes in the period that includes the enactment date. The Company records a valuation allowance to 
reduce the carrying amount of deferred tax assets if it is more likely than not that such asset will not be realized. The 
evaluation of uncertain tax positions is based on factors including, but not limited to, changes in the law, the 
measurement of tax positions taken or expected to be taken in tax returns, the effective settlement of matters subject to 
audit, new audit activity, and changes in facts or circumstances related to a tax position. The Company evaluates its tax 
positions on an annual basis.  

On December 22, 2017, the U.S. federal government enacted comprehensive tax legislation with the U.S. Tax 

Cuts and Jobs Act (“Tax Act”) that made changes to the U.S. tax code impacting the year ended December 31, 2017 and 
future years. Effective January 1, 2018, the Tax Act reduced the U.S. federal corporate tax rate from 35% to 21%.  

For the year ended December 31, 2017, the Tax Act required a one-time transition tax on certain unrepatriated 
earnings of foreign subsidiaries that is payable over eight years. At December 31, 2018, the Company does not have any 
foreign subsidiaries and the international aspects of the Tax Act are not applicable.  

In December 2017, the SEC staff issued Staff Accounting Bulletin No. 118 (“SAB 118”), which provided 
guidance on accounting for the tax effects of the Tax Act. SAB 118 provided a measurement period that should not 
extend beyond one year from the date of the Tax Act enactment for companies to complete the accounting under 
Accounting Standards Codification 740—Income Taxes. In accordance with SAB 118, to the extent that a company’s 
accounting for certain income tax effects of the Tax Act is incomplete, but the company is able to determine a reasonable 
estimate, it must record a provisional estimate in its financial statements. The Company’s accounting for the Tax Act was 
completed in the fourth quarter of 2018 and resulted in no adjustments to the Company’s prior provisional estimates 
recorded in the period ended December 31, 2017. 

The benefit for incomes taxes is as follows: 

Current 

Federal 
State 
Total current 
Deferred 

Federal 
State 

Total deferred 
Total tax expense 

Year ended December 31,  

2018 

2017 

(in thousands) 

 180 
 — 
 180 

 — 
 — 
 — 
 180 

 $ 

 $ 

 —  
 —  
 —  

 —  
 —  
 —  
 —  

$ 

$ 

F-29 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
  
 
  
 
 
  
 
  
 
  
A reconciliation of the expected income tax (benefit) computed using the federal statutory income tax rate at the 

Company’s effective tax rate is as follows: 

Income tax computed at federal statutory tax rate 
State taxes, net of federal benefit 
General business credit carryovers 
Non-deductible expenses 
Deferred rate change 
Change in valuation allowance 
Total 

Year ended December 31,  

2018 

2017 

2016 

 21.0 %   
 6.3 %   
 3.1 %   
 (2.1) %   
 — %   
 (28.1) %   
 0.2 %   

 34.0 %   
 6.1 %   
 5.0 %   
 (4.1)%   
 (21.8)%   
 (19.2)%   
 — %   

 34.0 %   
 5.6 %   
 4.2 %   
 (4.0)%   
 — %   
 (40.2)%   
 (0.4)%   

The Company has incurred net operating losses (“NOLs”) since June 2013. At December 31, 2018, the 

Company had federal and state net operating loss carryforwards of $162.9 million and $163.8 million, respectively. 
During 2018, the company generated federal and state NOLs carryforwards of $73.1 million and $72.1 million, 
respectively. The federal net operating loss carryforward generated in 2018 is limited to 80% of taxable income and has 
an indefinite carryforward period. The Company’s federal net operating loss carryforward generated in the period ended 
December 31, 2017 and prior, as well as the Company’s state NOL carryforwards begin to expire in 2033. As of 
December 31, 2018, the Company also had federal and state research and development tax credit carryforwards of $9.4 
million and $3.0 million, respectively, which expire beginning in 2028. As of December 31, 2018, the Company had 
state investment credits of $0.4 million, which expire beginning in 2019. 

Under the provisions of the Internal Revenue Code, certain substantial changes in the Company’s ownership 

may result in a limitation on the amount of NOL carryforwards and research and development credit carryforwards that 
may be utilized annually to offset future taxable income and taxes payable. In general, an ownership change, as defined 
by Section 382, results from transactions that increase the ownership of 5% stockholders or public groups in the stock of 
a corporation by more than 50 percent in the aggregate over a three-year period. During 2016, the Company completed a 
study through June 30, 2016, to determine whether any ownership change has occurred since the Company’s formation 
and has determined that transactions have resulted in three ownership changes, as defined by Section 382. There could be 
additional ownership changes in the future that could further limit the amount of NOLs and tax credit carryforwards that 
the Company can utilize.  

The significant components of the Company’s deferred tax assets and (liabilities) as of December 31, 2018 and 

2017 are as follows: 

Deferred tax assets: 

Net operating loss carryforwards 
Tax credit carryforwards 
Deferred rent 
Deferred revenue 
Non-deductible accruals and reserves 
Intangibles 
Stock compensation 
Total deferred tax assets 
Less valuation allowance 
Net deferred tax assets 

Deferred tax liabilities 

Depreciation and amortization 

Net deferred taxes 

F-30 

Year ended December 31,    

2018 

2017 

(in thousands) 

$   44,556  $   24,642  
 8,832  
 1,458  
 8,622  
 817  
 832  
 1,361  
    46,564  
    (44,953) 
 1,611  

 12,021 
 1,560 
 13,926 
 1,105 
 930 
 2,802 
 76,900 
    (75,213)
 1,687 

 (1,687)

$ 

 —  $ 

 (1,611) 
 —  

 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
 
 
 
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
  
 
 
 
 
   
 
 
  
  
  
  
 
 
  
  
  
  
 
 
  
  
  
   
 
   
 
 
  
As required by ASC 740, management has evaluated the positive and negative evidence bearing upon the 

realizability of its deferred tax assets, which principally comprise NOL carryforwards, research and development credit 
carryforwards, deferred revenue. Management has determined that it is more likely than not that the Company will not 
recognize the benefits of its federal and state deferred tax assets, and as a result, a valuation allowance of $75.2 million 
and $45.0 million has been established at December 31, 2018 and 2017, respectively. The change in valuation allowance 
was $30.2 million for the year ended December 31, 2018, primarily due to additional operating losses incurred by the 
Company for the year ended December 31, 2018. The primary reason for the difference between the income tax expense 
recorded by the Company and the amount of income tax expense at statutory income tax rates was the change in the 
valuation allowance.  

At December 31, 2018 and 2017, the Company had no unrecognized tax benefits. The Company has not as yet 

conducted a study of its research and development credit carryforwards. This study may result in an adjustment to the 
Company’s research and development credit carryforwards; however, until a study is completed, and any adjustment is 
known, no amounts are being presented as an uncertain tax position. A full valuation allowance has been provided 
against the Company’s research and development credits, and if an adjustment is required, this adjustment would be 
offset by an adjustment to the valuation allowance. Thus, there would be no impact to the balance sheets or statements of 
operations if an adjustment were required.  

Interest and penalty charges, if any, related to unrecognized tax benefits would be classified as income tax 

expense in the accompanying statements of operations. As of December 31, 2018 and 2017, the Company has no accrued 
interest related to uncertain tax positions. Since the Company is in a loss carryforward position, it is generally subject to 
examination by the U.S. federal, state, and local income tax authorities for all tax years in which a loss carryforward is 
available. 

13. Related-party transactions 

Since inception, the Company received consulting and management services from one of its investors. The total 

amount of consulting and management services provided by this investor was de minimis during the years ended 
December 31, 2018 and 2017.  

14. Selected quarterly financial data (unaudited)  

The following table contains quarterly financial information for 2018 and 2017. The Company believes that the 
following information reflects all normal recurring adjustments necessary for a fair statement of the information for the 
periods presented. The operating results for any quarter are not necessarily indicative of results for any future period.  

First 
Quarter 

Second 
  Quarter 

      Third 

  Quarter 

      Fourth 
  Quarter 

Total 

2018 

Collaboration revenue 
Total operating expenses 
Loss from operations 
Net loss 
Net loss per share 

Collaboration revenue 
Total operating expenses 
Loss from operations 
Net loss 
Net loss per share 

(amounts in thousands, except per share data) 
 2,008   $ 
 25,169  
    (23,161) 
 (22,532) 

 2,575   $ 
 28,269  
    (25,694) 
  (25,541) 

 2,094   $ 
 23,241  
    (21,147) 
 (20,289) 

$ 

 942   $ 

 22,035  
    (21,093) 
    (19,926) 
$ 

 (0.63)  $ 

 (0.70)  $ 

 7,619  
 98,714  
    (91,095) 
    (88,288) 
 (2.75) 

 (0.80)  $ 

 (0.63)  $ 

2017 

First 
Quarter 

      Second 
  Quarter 

      Third 

  Quarter 

      Fourth 
  Quarter 

Total 

(amounts in thousands, except per share data) 

$ 

 1,464   $ 
 18,986  
    (17,522) 
    (16,648) 
$ 

 (0.65)  $ 

 1,177   $ 
 19,816  
    (18,639) 
 (18,876) 

 1,148   $ 
 24,503  
    (23,355) 
 (23,346) 

 (0.73)  $ 

 (0.89)  $ 

 6,346   $   10,135  
 81,998  
 18,693  
    (71,863) 
    (12,347) 
    (70,698) 
 (11,828) 
 (2.64) 

 (0.40)  $ 

F-31 

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
    
     
 
 
 
 
 
 
 
  
  
  
  
  
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
     
 
 
 
 
 
 
 
  
  
  
  
  
 
 
 
15. Subsequent events 

Neurocrine Collaboration  

In January 2019, the Company entered into a collaboration and license agreement with Neurocrine for the 

research, development and commercialization of four programs including the Company’s Parkinson’s disease program, 
the Company’s Friedreich’s ataxia program, and two programs to be determined by the Company and Neurocrine at a 
later date. Under the terms of the agreement, the Company will receive an upfront payment of $165.0 million, inclusive 
of $50.0 million for the sale of 4,179,728 shares of its common stock, funding of development costs under the 
Parkinson’s and Friedreich’s ataxi programs, and may receive future development and regulatory milestones and 
royalties. 

AbbVie Alpha-Synuclein Collaboration  

In February 2019, the Company entered into a separate collaboration agreement with AbbVie, the AbbVie 

Alpha-Synuclein Collaboration Agreement, for the research, development, and commercialization of AAV and other 
virus based gene therapy products directed against pathological species of alpha-synuclein for the potential treatment of 
Parkinson’s disease and other synucleinopathies. Under the terms of the AbbVie Alpha-Synuclein Collaboration 
Agreement, the Company will receive an upfront payment of $65.0 million and may receive option exercise payments, 
future regulatory and commercial milestone payments and royalties. 

F-32 

 
 
EXHIBIT INDEX 

Description 

Form or 
Schedule       

Incorporated by Reference to: 
Filing 
Date with 
SEC 

SEC File 
Number 

Exhibit 
No. 

Filed  
Herewith 

Exhibit 
No. 

1.1 

  Sales Agreement by and between the 
Registrant and Cowen and Company, 
LLC, dated as of December 1, 2016. 

S-3 

1.2 

  12/01/2016   

3.1 

  Amended and Restated Certificate of 

8-K 

3.1 

  11/16/2015   

Incorporation of the Registrant. 

3.2 

  Amended and Restated By-Laws of the 

8-K 

3.2 

  11/16/2015   

Registrant. 

4.1 

  Specimen Common Stock Certificate of 

10-K 

4.1 

3/14/18 

the Registrant. 

4.2 

4.3 

  Form of Indenture to be entered into 
between the Registrant and Trustee. 

  S-3/A 

4.2 

  12/19/2016   

  Second Amended and Restated Investors’ 
Rights Agreement, by and among the 
Registrant and certain of its stockholders, 
dated as of April 10, 2015. 

  S-1/A 

4.2 

  10/28/2015   

10.1# 

  2014 Stock Option and Grant Plan and 
forms of award agreements thereunder. 

  S-1/A 

10.1 

  10/28/2015   

10.2# 

  2015 Stock Option and Incentive Plan and 
forms of award agreements thereunder. 

  S-1/A 

10.2 

  10/28/2015   

10.3† 

  Collaboration Agreement by and between 

  S-1/A 

10.3 

  11/06/2015   

the Registrant and Sanofi Genzyme 
Corporation, dated February 11, 2015. 

10.4† 

  Exclusive License Agreement by and 

S-1 

10.4 

  10/09/2015   

between the Registrant and the University 
of Massachusetts, dated January 30, 2014. 

10.5 

  Lease Agreement by and between the 

  S-1/A 

10.5 

  10/28/2015   

Registrant and UP 45/75 Sidney Street, 
LLC, dated as of April 1, 2014. 

10.6 

  First Amendment to Lease Agreement by 
and between the Registrant and 45/75 
Sidney Street, LLC, dated as of December 
23, 2015. 

10-Q 

10.5 

  05/12/2016   

10.7# 

  Offer Letter by and between the 

  S-1/A 

10.6 

  10/28/2015   

Registrant and Bernard Ravina, M.D., 
dated January 15, 2014. 

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10.9# 

  Offer Letter by and between the 

  S-1/A 

10.8 

  10/28/2015   

Registrant and Steven Paul, M.D., dated 
July 24, 2014. 

10.10 

  Form of Indemnification Agreement to be 
entered into between the Registrant and 
its directors. 

10.11 

  Form of Indemnification Agreement to be 
entered into between the Registrant and 
its executive officers. 

  S-1/A 

10.9 

  10/28/2015   

  S-1/A 

10.10 

  10/28/2015   

10.12†    License Agreement, by and between the 

  S-1/A 

10.11 

  11/04/2015   

Registrant and ReGenX Biosciences, 
LLC, dated May 28, 2014. 

10.13#    2015 Employee Stock Purchase Plan. 

  S-1/A 

10.12 

  10/28/2015   

10.14#    Employment Agreement by and between 
the Registrant and Steven M. Paul, Dated 
May 11, 2016. 

10.16#    Employment Agreement by and between 
the Registrant and Dinah Sah, dated May 
11, 2016. 

10-Q 

10.1 

  05/12/2016   

10-Q 

10.3 

  05/12/2016   

10.17 

  Lease Agreement by and between the 

10-Q 

10.6 

  05/12/2016   

Registrant and UP 64 Sidney Street, LLC, 
dated as of December 23, 2015. 

10.18 

  First Amendment to Lease Agreement, by 
and between Voyager Therapeutics, Inc. 
and UP 64 Sidney Street, LLC, dated June 
1, 2018. 

10.19#    Employment Agreement by and between 
the Registrant and Jane Pritchett 
Henderson, dated January 1, 2017. 

10.20#    Employment Agreement by and between 
the Registrant and Matthew Ottmer, dated 
September 11, 2017. 

8-K 

10.2 

  06/05/2018   

8-K 

10.1 

  01/03/2017   

8-K 

10.1 

  9/18/2017 

10.21 

  Second Amendment to the Lease 

8-K 

10.1 

  02/07/2018   

Agreement by and between the Registrant 
and UP 45/75 Sidney Street, LLC, dated 
as of February 5, 2018. 

10.22#    Amendment No. 1 to 2015 Employee 

10-K 

10.21 

  03/14/2018   

Stock Purchase Plan. 

10.23††    Collaboration Agreement by and between 
the Registrant and AbbVie Biotechnology 
Ltd, dated as of February 16, 2018. 

10-K 

10.22 

  03/14/2018   

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10.24#    Retirement Agreement, by and between 

8-K 

10.1 

  06/29/2018   

Voyager Therapeutics, Inc. and Steven M. 
Paul, dated June 28, 2018. 

10.25#    Employee Agreement, by and between 

8-K 

10.2 

  06/29/2018   

Voyager Therapeutics, Inc. and G. Andre 
Turenne, dated June 28, 2018. 

10.26#    Consulting Agreement, by and between 

10-Q 

10.5 

  08/07/2018   

Voyager Therapeutics, Inc. and Steven M. 
Paul, M.D., dated August 2, 2018. 

10.27#    Form of Non-Qualified Stock Option 

Agreement for Inducement Grant. 

10.28††    Collaboration and License Agreement, by 
and between the Registrant and 
Neurocrine Biosciences, Inc., dated 
January 28, 2019. 

10.29 

  Stock Purchase Agreement, by and 

between the Registrant and Neurocrine 
Biosciences, Inc., dated January 28, 2019. 

10.30 

Investor Agreement, by and between the 
Registrant and Neurocrine Biosciences, 
Inc., dated January 28, 2019. 

10.31††    Collaboration and Option Agreement, by 
and between the Registrant and AbbVie 
Ireland Unlimited Company, dated 
February 21, 2019. 

10.32#    Employment Agreement, by and between 

10-Q 

10.3 

  11/07/2018   

Voyager Therapeutics, Inc. and Allison 
Dorval, dated November 7, 2018. 

10.33#    Form of Restricted Stock Unit Agreement 

for Inducement Grant. 

10.34 

  Third Amendment to Lease Agreement, 
by and between Voyager Therapeutics, 
Inc. and UP 45/75 Sidney Street, LLC, 
dated June 1, 2018. 

21.1 

  Subsidiaries of the Registrant. 

23.1 

  Consent of Ernst & Young, Independent 
Registered Public Accounting Firm. 

24.1 

  Power of Attorney (see signature page of 

this Annual Report on Form 10-K). 

8-K 

10.1 

  06/05/2018   

F-35 

001-
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X 

X 

X 

X 

X 

X 

X 

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X 

X 

X 

31.1 

31.2 

  Certification of Principal Executive 
Officer pursuant to Exchange Act 
Rules 13a-14 or 15d-14. 

  Certification of Principal Financial 
Officer pursuant to Exchange Act 
Rules 13a-14 or 15d-14. 

32.1+ 

  Certifications of Principal Executive 

Officer and Principal Financial Officer 
pursuant to Exchange Act Rules 13a-
14(b) or 15d-14(b) and 18 U.S.C. 
Section 1350. 

101.INS    XBRL Instance Document. 

101.SCH   XBRL Taxonomy Extension Schema 

Document. 

101.CAL   XBRL Taxonomy Extension Calculation 

Document. 

101.LAB   XBRL Taxonomy Extension Definition 

Linkbase Document. 

101.PRE   XBRL Taxonomy Extension Labels 

Linkbase Document. 

101.DEF   XBRL Taxonomy Extension Presentation 

Link Document. 

#     Management contract or compensatory plan or arrangement filed in response to Item 15(a)(3) of the Instructions to 

the Annual Report on Form 10-K. 

†     Confidential treatment has been granted as to certain portions, which portions have been omitted and separately filed 

with the Securities and Exchange Commission. 

††   Confidential treatment has been requested as to certain portions, which portions have been omitted and separately 

filed with the Securities and Exchange Commission. 

+     The certification furnished in Exhibit 32.1 hereto is deemed to be furnished with this Annual Report on Form 10-K 

and will not be deemed to be “filed” for purposes of Section 18 of the Securities Exchange Act of 1934, as amended, 
except to the extent that the Registrant specifically incorporates it by reference. 

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SIGNATURES  

Pursuant to the requirements of Section 13 or 15(d) of the Securities Exchange Act of 1934, the registrant has 

duly caused this Form 10-K to be signed on its behalf by the undersigned, thereunto duly authorized. 

 February 26, 2019 

VOYAGER THERAPEUTICS, INC. 
/s/ G. Andre Turenne
By: 

G. Andre Turenne 
Chief Executive Officer, President, and 
Director 

SIGNATURES AND POWER OF ATTORNEY  

We, the undersigned directors and officers of Voyager Therapeutics, Inc. (the “Company”), hereby severally 
constitute and appoint G. Andre Turenne and Allison Dorval, and each of them singly, our true and lawful attorneys, 
with full power to them, and to each of them singly, to sign for us and in our names in the capacities indicated below, 
any and all amendments to this Annual Report on Form 10-K, and to file or cause to be filed the same, with all exhibits 
thereto and other documents in connection therewith, with the Securities and Exchange Commission, granting unto said 
attorneys, and each of them, full power and authority to do and perform each and every act and thing requisite and 
necessary to be done in connection therewith, as fully to all intents and purposes as each of us might or could do in 
person, and hereby ratifying and confirming all that said attorneys, and each of them, or their substitute or substitutes, 
shall do or cause to be done by virtue of this Power of Attorney. 

Pursuant to the requirements of the Securities Exchange Act of 1934, this Annual Report on Form 10-K has 

been signed by the following persons in the capacities and on the dates indicated. 

Name 

/s/ G. Andre Turenne 
G. Andre Turenne 

/s/Allison Dorval 
Allison Dorval 

/s/Mark Levin 
Mark Levin 

/s/Jim Geraghty 
Jim Geraghty 

/s/Michael Higgins 
Michael Higgins 

/s/Perry A. Karsen 
Perry A. Karsen 

/s/Steven Hyman, M.D. 
Steven Hyman, M.D. 

/s/Wendy Dixon, Ph.D. 
Wendy Dixon, Ph. D. 

/s/Steve Paul, M.D. 
Steve Paul, M.D. 

/s/Glenn Pierce, M.D., Ph.D. 
Glenn Pierce, M.D., Ph.D. 

Date 
February 26, 2019 

February 26, 2019 

February 26, 2019 

February 26, 2019 

February 26, 2019 

February 26, 2019 

February 26, 2019 

February 26, 2019 

February 26, 2019 

February 26, 2019 

4 

4 

Title 
Chief Executive Officer, President, and Director   
(Principal Executive Officer) 
Chief Financial Officer  
(Principal Financial and Accounting Officer) 
Director 

Director 

Director 

Director 

Director 

Director 

Director 

Director 

F-37 

 
  
  
  
 
 
 
 
 
 
 
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
  
 
  
  
  
  
  
  
 
 
  
  
  
  
  
  
 
  
  
  
  
  
  
 
 
 
 
 
 
 
Management Team

Board of Directors

Mark Levin
Chairman; Partner, Third Rock Ventures

Wendy Dixon, Ph.D.
Formerly of Bristol-Myers Squibb Company

Jim Geraghty
Chairman of the Board, Idera Pharmaceuticals  
and Orchard Therapeutics

Michael Higgins
Entrepreneur-in-Residence, Polaris Partners

Steven Hyman, M.D.
Director, Stanley Center for Psychiatric Research  
at the Broad Institute

Perry A. Karsen
Former Chief Executive Officer,  
Celgene Cellular Therapeutics

Steven Paul, M.D.
President, CEO and Chairman of the Board,  
Karuna Therapeutics

Glenn Pierce, M.D., Ph.D.
Entrepreneur-in-Residence at Third Rock Ventures

Andre Turenne
President and Chief Executive Officer

Omar Khwaja, M.D., Ph.D.
Chief Medical Officer

Matthew P. Ottmer
Chief Operating Officer

Allison Dorval
Chief Financial Officer

Dinah Sah, Ph.D.
Chief Scientific Officer

Luis Maranga, Ph.D. 
Chief Technical Operations Officer

Robert Hesslein   
General Counsel

Kathleen Hayes 
VP of Human Resources

Robert Pietrusko, Pharm.D. 
SVP of Regulatory Affairs and Quality Assurance

Matthew Osborne 
VP of Corporate Affairs,  
Communications and Investor Relations

Allen Nunnally  
VP of Corporate and Business Development

Legal Counsel
Wilmer Cutler Pickering Hale and Dorr LLP 
New York, NY

Independent Auditors
Ernst & Young LLP
Boston, MA

Transfer Agent and Registrar
Computershare Trust Company, N.A.
Canton, MA

Annual Meeting
The Annual Meeting of Stockholders will be held
Thursday, June 13, 2019 at 8:00 a.m. EDT
Voyager Therapeutics, Inc.,
64 Sidney Street, Cambridge, MA 02139

®

Voyager Therapeutics

75 Sidney Street
Cambridge, MA 02139
info@voyagertherapeutics.com
857-259-5340