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Arbutus Biopharma Corporation

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FY2022 Annual Report · Arbutus Biopharma Corporation
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UNITED STATES

SECURITIES AND EXCHANGE COMMISSION

Washington, D.C. 20549

Form 10-K

☒   ANNUAL REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934

For the Fiscal Year Ended December 31, 2022 

or 

☐   TRANSITION REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934

For the Transition Period from            to

Commission File Number: 001-34949 
Arbutus Biopharma Corporation
(Exact Name of Registrant as Specified in Its Charter)

British Columbia, Canada
(State or Other Jurisdiction of
Incorporation or Organization)

98-0597776
(I.R.S. Employer
Identification No.)

701 Veterans Circle
Warminster
PA
18974
(Address of Principal Executive Offices)

267-469-0914

Securities registered pursuant to Section 12(b) of the Act:

 (Registrant’s Telephone Number, Including Area Code)

Title of Each Class
Common shares, without par value

Trading Symbol(s)
ABUS

Name of Each Exchange on Which Registered
The Nasdaq Stock Market LLC

Securities registered pursuant to Section 12(g) of the Act: None.

Indicate by check mark if the registrant is a well-known seasoned issuer, as defined in Rule 405 of the Securities Act. Yes  No 

Indicate by check mark if the registrant is not required to file reports pursuant to Section 13 or Section 15(d) of the Act. Yes  No 

Indicate by check mark whether the registrant (1) has filed all reports required to be filed by Section 13 or 15(d) of the Securities Exchange Act of 1934
during the preceding 12 months (or for such shorter period that the registrant was required to file such reports), and (2) has been subject to such filing
requirements for the past 90 days. Yes  No 

Indicate by check mark whether the registrant has submitted electronically every Interactive Data File required to be submitted pursuant to Rule 405 of
Regulation S-T during the preceding 12 months (or for such shorter period that the registrant was required to submit such files). Yes  No 

Indicate by check mark whether the registrant is a large accelerated filer, an accelerated filer, a non-accelerated filer, a smaller reporting company, or an
emerging  growth  company.  See  the  definitions  of  “large  accelerated  filer,”  “accelerated  filer,”  “smaller  reporting  company,”  and  “emerging  growth
company” in Rule 12b-2 of the Exchange Act.

Large accelerated filer

Accelerated filer

Non-accelerated filer

Smaller reporting company

Emerging growth company

☐

☐

☒

☒

☐

 
 
 
If an emerging growth company, indicate by check mark if the registrant has elected not to use the extended transition period for complying with any new
or revised financial accounting standards provided pursuant to Section 13(a) of the Exchange Act. 
Indicate by check mark whether the registrant has filed a report on and attestation to its management’s assessment of the effectiveness of its internal control
over  financial  reporting  under  Section  404(b)  of  the  Sarbanes-Oxley  Act  (15  U.S.C.  7262(b))  by  the  registered  public  accounting  firm  that  prepared  or
issued its audit report. ☐

If securities are registered pursuant to Section 12(b) of the Act, indicate by check mark whether the financial statements of the registrant included in the
filing reflect the correction of an error to previously issued financial statements. ☐

Indicate by check mark whether any of those error corrections are restatements that required a recovery analysis of incentive-based compensation received
by any of the registrant’s executive officers during the relevant recovery period pursuant to § 240.10D-1(b). ☐

Indicate by check mark whether the registrant is a shell company (as defined in Rule 12b-2 of the Act). Yes ☐ No 

As of June 30, 2022, the last business day of the registrant’s most recently completed second fiscal quarter, the approximate aggregate market value of
voting and non-voting common equity held by non-affiliates of the registrant was $290,424,678 (based on the closing price of $2.71 per share as reported
on the Nasdaq Global Select Market as of that date).

As of February 28, 2023, the registrant had 162,570,989 common shares, without par value, outstanding.

DOCUMENTS INCORPORATED BY REFERENCE

Portions  of  the  registrant’s  definitive  proxy  statement  for  its  2023  Annual  Meeting  of  Shareholders,  which  the  registrant  intends  to  file  pursuant  to
Regulation  14A  with  the  Securities  and  Exchange  Commission  no  later  than  120  days  after  the  registrant’s  fiscal  year  ended  December  31,  2022,  are
incorporated by reference into Part III of this Form 10-K.

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ARBUTUS BIOPHARMA CORPORATION

TABLE OF CONTENTS

Cautionary Note Regarding Forward-looking Statements
Risk Factors Summary

PART I

Item 1.
Item 1A.
Item 1B.
Item 2.
Item 3.
Item 4.

PART II

Item 5.
Item 6.
Item 7.
Item 7A.
Item 8.
Item 9.
Item 9A.
Item 9B.
Item 9C.

PART III

Item 10.
Item 11.
Item 12.
Item 13.
Item 14.

PART IV

Business
Risk Factors
Unresolved Staff Comments
Properties
Legal Proceedings
Mine Safety Disclosures

Market for Registrant’s Common Equity, Related Stockholder Matters and Issuer Purchases of Equity Securities
Reserved
Management’s Discussion and Analysis of Financial Condition and Results of Operations
Quantitative and Qualitative Disclosures about Market Risk
Financial Statements and Supplementary Data
Changes in and Disagreements with Accountants on Accounting and Financial Disclosure
Controls and Procedures
Other Information
Disclosure Regarding Foreign Jurisdictions that Prevent Inspections

Directors, Executive Officers and Corporate Governance
Executive Compensation
Security Ownership of Certain Beneficial Owners and Management and Related Stockholder Matters
Certain Relationships and Related Transactions, and Director Independence
Principal Accountant Fees and Services

Item 15.

Exhibits and Financial Statement Schedules

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Cautionary Note Regarding Forward-looking Statements

This  Annual  Report  on  Form  10-K  (this  “Form  10-K”)  contains  “forward-looking  statements”  or  “forward-looking  information”  within  the  meaning  of
applicable United States and Canadian securities laws (we collectively refer to these items as “forward-looking statements”). Forward-looking statements
are  generally  identifiable  by  use  of  the  words  “believes,”  “may,”  “plans,”  “will,”  “anticipates,”  “intends,”  “budgets,”  “could,”  “estimates,”  “expects,”
“forecasts,” “projects” and similar expressions that are not based on historical fact or that are predictions of or indicate future events and trends, and the
negative of such expressions. Forward-looking statements in this Form 10-K, including the documents incorporated by reference, include statements about,
among other things:

•
•
•
•
•

•

•
•
•
•

•
•
•
•
•
•

•

our strategy, future operations, pre-clinical research, pre-clinical studies, clinical trials, prospects and the plans of management;
the potential for our product candidates to achieve their desired or anticipated outcomes;
the expected cost, timing and results of our clinical development plans and clinical trials, including our clinical collaborations with third parties;
the potential impact of the COVID-19 pandemic on our business and clinical trials;
the  discovery,  development  and  commercialization  of  a  curative  combination  regimen  for  chronic  hepatitis  B  infection,  a  disease  of  the  liver
caused by the hepatitis B virus (“HBV”);
the  potential  of  our  product  candidates  to  improve  upon  the  standard  of  care  and  contribute  to  a  functional  curative  combination  treatment
regimen;
obtaining necessary regulatory approvals;
obtaining adequate financing through a combination of financing activities and operations;
the potential for us to discover and/or develop new molecular entities for treating coronaviruses, including COVID-19;
the  expected  returns  and  benefits  from  strategic  alliances,  licensing  agreements,  and  research  collaborations  with  third  parties,  and  the  timing
thereof;
our expectations regarding our technology licensed to third parties, and the timing thereof;
our anticipated revenue and expense fluctuation and guidance;
our expectations regarding the timing of announcing data from our ongoing clinical trials;
our expectations regarding current patent disputes and litigation;
our expectation of a net cash burn between $95.0 million and $100.0 million in 2023;

our belief that we have sufficient cash resources to fund our operations into the fourth quarter of 2024; and

the possibility that our clinical development plans could be further delayed or suspended as a result of the military action by Russia in Ukraine,

as well as other statements relating to our future operations, financial performance or financial condition, prospects or other future events. Forward-looking
statements appear primarily in the sections of this Form 10-K entitled “Item 1-Business,” “Item 1A-Risk Factors,” “Item 7-Management’s Discussion and
Analysis of Financial Condition and Results of Operations,” “Item 7A-Quantitative and Qualitative Disclosures About Market Risk,” and “Item 8-Financial
Statements and Supplementary Data.”

Forward-looking statements are based upon current expectations and assumptions and are subject to a number of known and unknown risks, uncertainties
and other factors that could cause actual results to differ materially and adversely from those expressed or implied by such statements. Factors that could
cause  or  contribute  to  such  differences  include,  but  are  not  limited  to,  those  discussed  in  this  Form  10-K  and  in  particular  the  risks  and  uncertainties
discussed under “Item 1A-Risk Factors” of this Form 10-K. As a result, you should not place undue reliance on forward-looking statements.

Additionally, the forward-looking statements contained in this Form 10-K represent our views only as of the date of this Form 10-K (or any earlier date
indicated in such statement). While we may update certain forward-looking statements from time to time, we specifically disclaim any obligation to do so,
even if new information becomes available in the future. However, you are advised to consult any further disclosures we make on related subjects in the
periodic and current reports that we file with the Securities and Exchange Commission.

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The foregoing cautionary statements are intended to qualify all forward-looking statements wherever they may appear in this Form 10-K. For all forward-
looking statements, we claim protection of the safe harbor for the forward-looking statements contained in the Private Securities Litigation Reform Act of
1995.

This  Form  10-K  also  contains  estimates,  projections  and  other  information  concerning  our  industry,  our  business,  and  the  markets  for  certain  diseases,
including data regarding the estimated size of those markets, and the incidence and prevalence of certain medical conditions. Information that is based on
estimates, forecasts, projections, market research or similar methodologies is inherently subject to uncertainties and actual events or circumstances may
differ materially from events and circumstances reflected in this information. Unless otherwise expressly stated, we obtained this industry, business, market
and  other  data  from  reports,  research  surveys,  studies  and  similar  data  prepared  by  market  research  firms  and  other  third  parties,  industry,  medical  and
general publications, government data and similar sources.

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Risk Factors Summary

The following is a summary of the principal risks that could adversely affect our business, operations and financial results. For more information, see “Item
1A. Risk Factors” in this Annual Report on Form 10-K for the year ended December 31, 2022.

Risks Related to Our Business, Our Financial Results and Need for Additional Capital

• We are in the early stages of our development, and there is a limited amount of information about us upon which you can evaluate our product

candidates.

• We  will  require  substantial  additional  capital  to  fund  our  operations.  Additional  funds  may  be  dilutive  to  shareholders  or  impose  operational
restrictions.  Further,  if  additional  capital  is  not  available,  we  may  need  to  delay,  limit  or  eliminate  our  research,  development  and
commercialization programs and modify our business strategy.

• We have incurred losses in nearly every year since our inception and we anticipate that we will not achieve profits for the foreseeable future. To

date, we have had no product revenues.
The COVID-19 pandemic could adversely impact our business, including our clinical development plans.

•

Risks Related to Development, Clinical Testing, Regulatory Approval, Marketing, and Coverage and Reimbursement of our Product Candidates

• Our  product  candidates  are  in  early  stages  of  development  and  must  go  through  clinical  trials,  which  are  very  expensive,  time-consuming  and

•

•

•

difficult to design and implement. The timing and outcomes of clinical trials are uncertain.
Pre-clinical studies and preliminary and interim data from clinical trials of our product candidates are not necessarily predictive of the results or
success of ongoing or later clinical trials of our product candidates.
Because  we  have  limited  resources,  we  may  decide  to  pursue  a  particular  product  candidate  and  fail  to  advance  product  candidates  that  later
demonstrate a greater chance of clinical and commercial success.
Several of our current pre-clinical studies and clinical trials are being conducted outside the United States, and the FDA may not accept data from
trials conducted in locations outside the United States.

• We cannot guarantee how long it will take regulatory agencies to review our applications for product candidates.
•

If a particular product candidate causes undesirable side effects, then we may be unable to receive regulatory approval of or commercialize such
product candidate.

• We may find it difficult to enroll patients in our clinical trials, which could hinder such clinical trials.
•

Several of our and our collaboration partner’s current and planned clinical trials have been impacted and could be further delayed or suspended as
a result of the military action by Russia in Ukraine.
Even if our product candidates obtain regulatory approval, they will remain subject to ongoing regulatory requirements.

•
• We face significant competition from other biotechnology and pharmaceutical companies targeting HBV and coronaviruses, including COVID-19.
• We are largely dependent on the future commercial success of our HBV and coronavirus product candidates.
• We may incur substantial liabilities and may be required to limit commercialization of our products in response to product liability lawsuits.
•

Coverage and adequate reimbursement may not be available for our product candidates, which could make it difficult for us to sell our products
profitably.

• We are subject to United States and Canadian healthcare laws and regulations, which could expose us to adverse consequences such as criminal

•

sanctions, civil penalties, contractual damages or reputational harm, among others.
Failure to comply with the United States Foreign Corrupt Practices Act, and potentially other similar global laws could subject us to penalties and
other adverse consequences.

Risks Related to Our Dependence on Third Parties

• We depend on our license agreement with Alnylam Pharmaceuticals, Inc. for the commercialization of ONPATTRO™ (Patisiran).

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• We  expect  to  depend  in  part  on  our  licensing  agreements  for  a  significant  portion  of  our  revenues  for  the  foreseeable  future  and  to  develop,
conduct clinical trials with, obtain regulatory approvals for, and manufacture, market and sell some of our product candidates. If these licensing
agreements are unsuccessful, or anticipated milestone or royalty payments are not received, our business could be materially adversely affected.
• We  are  dependent  on  our  collaboration  and  licensing  partners  and,  therefore,  are  subject  to  the  efforts  of  these  parties  and  our  ability  to

successfully collaborate with them.

• We  will  depend  on  Qilu  Pharmaceutical  Co.,  Ltd.  for  the  development  and  commercialization  of  AB-729  in  China,  Hong  Kong,  Macau  and

•

Taiwan.
If conflicts arise between our collaboration or licensing partners and us, our collaboration or licensing partners may act in their best interest and
not in our best interest, which could adversely affect our business.

• We rely on third parties to conduct our clinical trials, and if they fail to fulfill their obligations, our development plans may be adversely affected.
• We  rely  exclusively  on  third  parties  to  formulate  and  manufacture  our  product  candidates,  which  exposes  us  to  risks  that  may  delay  or  hinder

development, regulatory approval and commercialization of our products.

Risks Related to Our Intellectual Property

• Other entities may assert patent rights that prevent us from developing or commercializing our products.
• Our patents and patent applications may be challenged and may be found to be invalid, which could adversely affect our business.
• We may incur substantial costs as a result of litigation or other proceedings relating to patent and other intellectual property rights which could

•

have a material adverse effect on our business and could cause the market value of our common shares to decline.
Confidentiality agreements with employees and others, including collaborators, may not adequately prevent disclosure of trade secrets and other
proprietary information.

Risks Related to the Ownership of our Common Shares

•

The concentration of common share ownership will likely limit the ability of the other shareholders to influence corporate matters. Further, our
articles and certain Canadian laws could delay or deter a change of control.

• We are incorporated in Canada, with our assets located both in Canada and the United States, with the result that it may be difficult for investors to

•

enforce judgments obtained against us or some of our officers.
If we are deemed to be a “passive foreign investment company,” investors who are subject to United States federal taxation would likely suffer
materially adverse United States federal income tax consequences.

General Risk Factors

If we are unable to attract and retain qualified key individuals, our ability to implement our business plan may be adversely affected.

•
• We could face liability from our controlled use of hazardous and radioactive materials.
• Our business, reputation, and operations could suffer in the event of information technology system failures.
• We may acquire other assets or businesses, or form strategic alliances or collaborations or make investments in other companies or technologies

that could harm our business.

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Item 1. Business 

Overview

PART I

Arbutus  Biopharma  Corporation  (“Arbutus”,  the  “Company”,  “we”,  “us”,  and  “our”)  is  a  clinical-stage  biopharmaceutical  company  leveraging  its
extensive virology expertise to develop novel therapeutics that target specific viral diseases. Our current focus areas include Hepatitis B virus (“HBV”),
SARS-CoV-2, and other coronaviruses. To address HBV, we are developing an RNA interference (“RNAi”) therapeutic, an oral PD-L1 inhibitor, and an
oral RNA destabilizer to potentially identify a combination regimen with the aim of providing a functional cure for patients with chronic HBV infection
(“cHBV”) by suppressing viral replication, reducing surface antigen and reawakening the immune system. We believe our lead compound, AB-729, is the
only RNAi therapeutic with evidence of immune re-awakening. AB-729 is currently being evaluated in multiple phase 2 clinical trials. We also have an
ongoing drug discovery and development program directed to identifying novel, orally active agents for treating coronaviruses, including SARS-CoV-2,
where we have nominated a compound and have begun IND-enabling pre-clinical studies. In addition, we are also exploring oncology applications for our
internal PD-L1 portfolio.

Strategy

The core elements of our strategy include:

•

Developing  a  broad  portfolio  of  compounds  that  target  cHBV.  Our  HBV  product  pipeline  includes  a  subcutaneously-delivered  RNAi
therapeutic, an oral HBV RNA destabilizer compound and an oral PD-L1 inhibitor. We believe that a combination of compounds that can suppress
HBV DNA replication and hepatitis B surface antigen (“HBsAg”) expression as well as reawaken patients’ HBV-specific immune response would
address the most important elements to achieving a functional cure. We define a functional cure as unquantifiable plasma HBV DNA and HBsAg
levels more than six months after discontinuation of all treatment, with or without quantifiable anti-HBsAg antibodies.

AB-729  is  our  proprietary  subcutaneously-delivered  RNAi  therapeutic  product  candidate  that  suppresses  all  HBV  antigens,  including  HBsAg
expression, which is thought to be a key prerequisite to enable reawakening of a patient’s immune system to respond to HBV. AB-729 is currently
in two Phase 2a proof-of-concept clinical trials in combination with other agents with potentially complementary mechanisms of action and we are
also  continuing  to  follow  subjects  from  our  Phase  1a/1b  clinical  trial  (“AB-729-001”).  Preliminary  data  from  AB-729-001  has  shown  that
treatment  with  AB-729  provided  robust  and  comparable  HBsAg  declines  regardless  of  dose,  dosing  interval  or  patient  characteristics  and  was
generally  safe  and  well-tolerated  after  completing  dosing  in  41  subjects.  Preliminary  data  also  suggests  that  treatment  with  AB-729  increased
HBV-specific immune responses and, in a small number of subjects who discontinued both AB-729 and nucleos(t)ide analogue (“NA”) therapy, a
sustained reduction in HBsAg and HBV DNA persisted after stopping AB-729. The clinical data for AB-729 continues to support its development
as a potential cornerstone agent for the treatment of cHBV infection.

AB-101 is our oral PD-L1 inhibitor that has the potential to reawaken patients’ HBV-specific immune response by inhibiting PD-L1. Preclinical
data in an HBV mouse model was presented at the 2022 AASLD Liver Meeting showing that combination treatment with AB-101 and an HBV-
targeting  GalNAc-siRNA  agent  resulted  in  activation  and  increased  frequency  of  HBV-specific  T-cells  and  greater  anti-HBsAg  antibody
production.  This  favorable  preclinical  profile  supports  further  development  of  AB-101  as  a  therapeutic  modality  for  cHBV  treatment  and  we
anticipate  initiating  a  Phase  1  healthy  subject  clinical  trial  with  AB-101  in  the  first  half  of  2023.  We  are  also  exploring  potential  oncology
applications for our internal PD-L1 portfolio.

AB-161 is our next-generation oral HBV specific RNA destabilizer. We have conducted extensive non-clinical safety evaluations with AB-161
that gives us confidence in this molecule’s ability to circumvent the peripheral neuropathy findings seen in non-clinical safety studies with our
first-generation oral RNA destabilizer, AB-452. We recently presented preclinical data at the Discovery on Target Conference showing that AB-
161 reduced HBV RNA and

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•

•

HBsAg in multiple preclinical models, with favorable liver centricity and lack of observed peripheral neuropathy. We anticipate initiating a Phase
1 healthy subject clinical trial with AB-161 in the first half of 2023.

Combining therapeutic product candidates with complementary mechanisms of action to find a functional cure for people with cHBV. We
believe that our proprietary product candidates AB-729, AB-101 and AB-161 may provide our first proprietary combination therapy for patients
with cHBV. In-line with our strategy to position AB-729 as a potential cornerstone therapeutic in future HBV combination regimens, and to help
guide future development of combination therapies of AB-729 with other compounds from our proprietary HBV portfolio, we are evaluating AB-
729 in combination with other agents with potentially complementary mechanisms of action, including the following:

• AB-729 in combination with ongoing standard-of-care NA therapy and short courses of Peg-IFNα-2a in subjects with cHBV in a Phase
2a proof-of-concept clinical trial (“AB-729-201”). Preliminary data from the lead-in phase of this clinical trial further validated AB-729’s
potential to reduce HBsAg in cHBV patients.

• AB-729 

in  combination  with  Vaccitech  plc’s 

(“Vaccitech”)  VTP-300,  a  proprietary  T-cell  stimulating  antigen-specific
immunotherapeutic,  and  NA  therapy  for  the  treatment  of  subjects  with  cHBV  in  a  Phase  2a  proof-of-concept  clinical  trial  (“AB-729-
202”).  We  recently  amended  the  clinical  trial  to  include  an  additional  arm  with  an  approved  PD-1  monoclonal  antibody  inhibitor,
nivolumab (Opdivo ).

®

Advancing small molecule antiviral product candidates to treat COVID-19 and future coronavirus outbreaks. This program is focused on
the  discovery  and  development  of  new  molecular  entities  for  treating  coronaviruses,  including  COVID-19,  that  address  specific  viral  targets
including the nsp5 viral protease (“M ”) and the nsp12 viral polymerase.

pro

•

In  the  fourth  quarter  of  2022,  we  nominated  AB-343  as  our  lead  coronavirus  drug  candidate  that  inhibits  the  SARS-CoV-2  M ,  a
validated target for the treatment of COVID-19 and potential future coronavirus outbreaks. In our pre-clinical research conducted to date,
AB-343  has  shown  pan-coronavirus  antiviral  activity,  no  reduction  in  potency  against  known  SARS-CoV-2  variants,  robust  activity
against  SARS-CoV-2  M   resistant  strains,  and  a  favorable  drug-drug  interaction  profile  with  no  need  for  ritonavir  boosting.  We  are
advancing AB-343 into IND-enabling studies. We are also continuing lead optimization activities for an nsp12 viral polymerase, which
could potentially be combined with AB-343 to achieve better patient treatment outcomes and for use in prophylactic settings.  

pro

pro

Background on HBV

Hepatitis B is a potentially life-threatening liver infection caused by HBV. HBV can cause chronic infection which leads to a higher risk of death from
cirrhosis and liver cancer. cHBV represents a significant unmet medical need. There are HBV vaccines approved by the FDA, which are indicated for the
prevention of infection caused by HBV. However,  the  World  Health  Organization  estimates  that  over  290  million  people  worldwide  suffer  from  cHBV,
while other estimates indicate that approximately 2.4 million people in the United States suffer from cHBV. Even with the availability of effective vaccines
and current treatment options, approximately 820,000 people die every year from complications related to cHBV. We believe there is a compelling market
opportunity for an HBV curative regimen. Currently, an estimated 30.4 million (10.5%) of a total of over 290 million people worldwide with cHBV are
diagnosed  and  approximately  6.6  million  (2.3%)  are  on  treatment.  We  believe  that  the  introduction  of  an  HBV  curative  regimen  with  a  finite  duration
would substantially increase diagnosis and treatment rates for people with cHBV.

Current treatments and their limitations

Today’s current treatment options for cHBV include pegylated interferon-α regimens (“Peg-IFNα”) and NA therapies. Peg-IFNα, a synthetic version of a
substance produced by the body to fight infection, is administered by injection and has numerous side effects including flu-like symptoms and depression.
NA therapies are oral antiviral medications which, when taken chronically, reduce HBV virus replication and inflammation and significantly reduce HBV
DNA in the blood. Oral NA

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therapies have become the standard-of-care for HBV treatment, mainly due to their ability to drive viral load to undetectable levels in the serum of patients,
their single pill once-a-day dosing and favorable safety profile. However, in most cases, once Peg-IFNα and NA therapies are stopped, virus replication
resumes and liver inflammation and fibrosis may still progress. While these treatments reduce viral load, less than 5% of patients are functionally cured
after a finite treatment duration. With such low cure rates, most patients with cHBV are required to take NA therapy daily for the rest of their lives.

Background on Coronaviruses

Coronaviruses are a large family of viruses that range from the common cold to more severe diseases such as severe acute respiratory syndrome (SARS),
Middle East respiratory syndrome (MERS), and COVID-19. COVID-19 has caused approximately 7.2 million deaths globally according to an analysis by
the Institute for Health Metrics and Evaluation (IHME). COVID-19 spreads when an infected person breathes out droplets and very small particles that
contain the virus. As we strive to identify and develop new antiviral small molecules to treat COVID-19 and future coronavirus outbreaks, we have focused
our research efforts on two essential targets critical for replication across all coronaviruses – nsp5 protease and nsp12 polymerase.

Current treatments and their limitations

Today’s current treatment options for COVID-19 include multiple vaccines, anti-viral drugs and antibodies to prevent or treat the disease. Despite the high
efficacy of the COVID-19 vaccines, it is estimated that more than 25% of the world’s population remains unvaccinated against COVID-19. Today’s anti-
viral  treatments  have  certain  limitations,  including  a  short  time  frame  to  begin  treatment,  potential  drug-drug  interactions  due  to  ritonavir  boosting  and
frequently  occurring  side  effects.  Patients  continue  to  be  hospitalized  and  die  from  COVID-19.  In  addition  to  the  availability  of  vaccines  and  other
treatments, new effective and safe therapies are needed to successfully combat the COVID-19 pandemic and any future coronavirus outbreaks.

Our Product Candidates

Our product pipeline includes multiple product candidates that target various steps in the HBV viral lifecycle and pan-coronavirus compounds that target
essential viral targets for replication.

Our product pipeline consists of the following programs:

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We  continue  to  explore  expansion  opportunities  for  our  pipeline  through  internal  discovery  and  development  activities  and  through  potential  strategic
alliances.

RNAi therapeutic (AB-729)

RNAi therapeutics represent a significant advancement in drug development. RNAi therapeutics utilize a natural pathway within cells to silence genes by
eliminating the disease-causing proteins that they code for. We are developing RNAi therapeutics that are designed to reduce HBsAg expression and other
HBV antigens in people with cHBV. Reducing HBsAg is widely believed to be a key prerequisite to enable a patient’s immune system to reawaken and
respond against the virus.

AB-729  is  a  subcutaneously-delivered  RNAi  single-trigger  therapeutic  targeted  to  hepatocytes  using  our  proprietary  covalently  conjugated  GalNAc
delivery technology. AB-729 reduces all HBV antigens and inhibits viral replication.

Phase 1a/1b single- and multiple-dose clinical trial (AB-729-001)

In this three-part clinical trial, we investigated the safety, tolerability, pharmacokinetics, and pharmacodynamics of single- and multi-doses of AB-729 in
healthy subjects and in cHBV patients with the goal of identifying the most appropriate doses and dosing intervals to take forward into Phase 2 clinical
development.

The first two parts evaluated single ascending doses of AB-729 in healthy subjects and in patients with cHBV, respectively. Data showed that a 60mg or
90mg  single  dose  of  AB-729  results  in  robust  HBsAg  and  HBV  DNA  declines  in  HBV  DNA  positive  patients.  Part  3  of  the  trial  dosed  HBV  DNA
negative/positive  patients  with  60mg  or  90mg  of  AB-729  every  4,  8  or  twelve  weeks.  Dosing  of  patients  in  Part  3  has  been  completed  and  we  are
continuing to follow these patients.

Data from Part 3 of the AB-729-001 clinical trial was presented at the 2022 European Association for the Study of the Liver (EASL) International Liver
Congress™ (ILC) in June 2022 and showed that repeat dosing of 60mg and 90mg of AB-729 in 41 patients resulted in robust and comparable HBsAg
declines  in  HBeAg  positive/negative  and  HBV  DNA  positive/negative  patients  at  week  48  (1.89  to  2.15  log10  decline  in  HBsAg).  Fifty  percent  of  the
patients (16 out of 32) maintained HBsAg levels below 100 IU/mL 24 weeks after their last dose of AB-729. Patients treated with AB-729 experienced an
increase in HBV-specific T-cells activation and a decrease in exhausted T-cells. In this trial, AB-729 was generally safe and well-tolerated.

At the AASLD Liver Meeting in November 2022, we presented additional data from Part 3 of the AB-729-001 clinical trial, which included nine patients
who had had previously completed 48 weeks of treatment with AB-729, and 24 weeks later met protocol-defined criteria to also stop NA therapy. These
nine patients had completed 12 to 44 weeks of follow-up after discontinuing their NA therapy. None had met the protocol-defined criteria to restart NA
therapy and there was no evidence of clinical or biochemical relapse. HBsAg levels remained at 1.05 log10 to 2.35 log10 below pre-trial levels in all nine
patients.  Three  patients  experienced  transient  HBV  DNA  elevations  that  spontaneously  resolved  without  intervention,  which  further  supports  AB-729’s
potential for immunological control. One patient restarted NA therapy at the investigator’s request after the week 20 visit; no alanine transaminase (“ALT”)
elevation or safety signals were observed. There were no adverse events (“AEs”) reported and no ALT flares were observed in the clinical trial. Recently,
one  of  the  eight  remaining  patients  met  the  protocol-defined  HBV  DNA  criteria  to  restart  NA  therapy  without  evidence  of  any  ALT  flare.  We  are
continuing to follow the seven patients who remain off NA therapy and anticipate reporting additional off-treatment data in the first half of 2023.

The new clinical data for AB-729 continues to support its development as a potential cornerstone agent for the treatment of cHBV infection. The efficacy
and safety data for AB-729, derived from up to one year of dosing, supported our view that 60 mg every 8 weeks was an appropriate dose to move forward
in our Phase 2a clinical trials. To advance our efforts to position AB-729 as a potential cornerstone therapeutic in future HBV combination regimens, we
are evaluating AB-729 in several Phase 2a proof-of-concept combination clinical trials with other agents with potentially complementary mechanisms of
action, some via clinical collaborations with other companies as described below.

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Phase 2a proof-of-concept clinical trial to evaluate AB-729 in combination with Peg-IFNα-2a (AB-729-201)

We  have  completed  enrollment  in  a  randomized,  open  label,  multicenter  Phase  2a  proof-of-concept  clinical  trial  investigating  the  safety  and  antiviral
activity of AB-729 in combination with ongoing NA therapy and short courses of Peg-IFNα-2a in 43 stably NA-suppressed, HBeAg negative, non-cirrhotic
patients with cHBV. After 24-weeks of dosing with AB-729 (60mg every 8 weeks), patients are randomized into one of four arms to receive ongoing NA
therapy plus Peg-IFNα-2a for either 12 or 24 weeks, with or without additional doses of AB-729. After completion of the assigned Peg-IFNα-2a treatment
period,  all  patients  will  remain  on  NA  therapy  for  the  initial  24-week  follow-up  period,  and  will  then  discontinue  NA  treatment,  provided  they  meet
protocol-defined stopping criteria. Patients who stop NA therapy will enter an intensive follow-up period for 48 weeks.

Preliminary data from the lead-in phase of the trial further validated AB-729’s potential to reduce HBsAg. For the first 15 patients who reached week 16 of
treatment and received two doses of AB-729 plus NA therapy, the mean HBsAg decline was 1.51 log10, comparable to the decline observed at the same
timepoint  in  the  Phase  1b  clinical  trial  AB-729-001  (1.56  log10),  while  continuing  to  exhibit  a  generally  safe  and  well-tolerated  profile.  We  anticipate
providing preliminary data from patients who have received doses of Peg-IFNα-2a in the first half of 2023.

Collaboration with Vaccitech (AB-729-202)

Through a clinical collaboration agreement with Vaccitech that we entered into in July 2021, we are enrolling patients in AB-729-202, a Phase 2a proof-of-
concept clinical trial evaluating the safety, antiviral activity and immunogenicity of Vaccitech’s VTP-300, a proprietary T-cell stimulating antigen-specific
immunotherapeutic, administered after AB-729 in NA-suppressed patients with cHBV. The trial is designed to enroll 40 NA-suppressed, HBeAg negative
or positive, non-cirrhotic cHBV patients. All patients will receive AB-729 (60mg every 8 weeks) plus NA therapy for 24 weeks. At week 24, treatment
with AB-729 will stop. Patients will continue only their NA therapy and will be randomized to receive VTP-300 or placebo at Week 26, Week 30 and at
Week 38 (if protocol-defined eligibility is met). At week 48, all patients will be evaluated for eligibility to discontinue NA therapy and will be followed for
an additional 24-48 weeks. We anticipate providing preliminary data from patients who received AB-729, NA therapy and VTP-300 in the second half of
2023.

We  recently  amended  the  AB-729-202  protocol  to  include  an  additional  arm  with  an  approved  PD-1  inhibitor,  nivolumab  (Opdivo®).  Upon  regulatory
approval of the amendment, 20 patients will receive AB-729 (60mg every 8 weeks) plus NA therapy for 24 weeks, followed by administration of VTP-300
plus a low dose of nivolumab in conjunction with the booster dose(s) only while remaining on their NA therapy. At week 48, all patients will be evaluated
for eligibility to discontinue NA therapy, and will be followed for an additional 24-48 weeks. We anticipate dosing the first patient in this arm in the first
half of 2023, subject to regulatory approval.

This clinical trial is being managed by us, subject to oversight by a joint development committee comprised of representatives from both companies. We
and Vaccitech retain full rights to our respective product candidates and will split all costs associated with the clinical trial. Pursuant to the agreement, the
parties intend to undertake a larger Phase 2b clinical trial depending on the results of the initial Phase 2a clinical trial.

Collaboration with Assembly

Through a clinical collaboration agreement with Assembly that we entered into in August 2020, Assembly conducted a clinical trial evaluating AB-729 in
combination with its first-generation HBV core inhibitor (capsid inhibitor) candidate VBR and standard-of-care NA therapy for the treatment of cHBV in
HBeAg negative patients with cHBV. The randomized, multi-center, open-label Phase 2a proof-of-concept clinical trial was designed to evaluate the safety,
pharmacokinetics, and antiviral activity of the triple combination of AB-729, VBR, and an NA (n=32) compared to the dual combinations of VBR with an
NA (n=16) and AB-729 with an NA (n=17). Patients were dosed for 48 weeks with AB-729 (60mg subcutaneously every 8 weeks) and/or VBR (300mg
orally once daily), with a 48-week follow-up period. At week 48, all patients were to be evaluated for eligibility to discontinue NA therapy. In July 2022,
Assembly  announced  its  plans  to  discontinue  development  of  VBR.  Despite  this,  in  consultation  with  Assembly,  we  continued  this  Phase  2a  proof-of-
concept  clinical  trial  in  order  to  fully  and  accurately  assess  the  results.  Assembly  completed  enrollment  in  the  clinical  trial  and  preliminary  data  were
presented at the 2022 AASLD Liver

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Meeting, which indicated that adding VBR to AB-729 and NA therapy does not positively or negatively impact the reduction of HBsAg compared to AB-
729 and NA therapy alone. Accordingly, we have mutually agreed to discontinue the clinical trial following completion of the final, on-treatment visit at
week  48.  All  regimens  were  generally  safe  and  well-tolerated  in  this  trial.  Both  parties  shared  in  the  costs  of  the  collaboration.  Except  to  the  extent
necessary to carry out Assembly’s responsibilities with respect to the collaboration trial, we have not provided any license grant to Assembly for use of
AB-729.

Oral PD-L1 Inhibitor (AB-101)

PD-L1  inhibitors  complement  our  pipeline  of  agents  and  could  potentially  be  an  important  part  of  a  combination  therapy  for  the  treatment  of  HBV  by
reawakening the immune system. Highly functional HBV-specific T cells within our immune system are believed to be required for long-term HBV viral
resolution.  However,  HBV-specific  T  cells  become  functionally  defective,  and  greatly  reduced  in  their  frequency  during  cHBV.  One  approach  to  boost
HBV-specific  T  cells  is  to  prevent  PD-L1  proteins  from  binding  to  PD-1  and  thus  inhibiting  the  HBV-specific  immune  function  of  T  cells.  Immune
checkpoints such as PD-1/PD-L1 play an important role in the induction and maintenance of immune tolerance and in T-cell activation.

AB-101 is our oral PD-L1 inhibitor candidate that we believe will allow for controlled checkpoint blockade while minimizing the systemic safety issues
typically seen with checkpoint antibody therapies. Preclinical data generated thus far indicates that AB-101 mediates activation and reinvigoration of HBV-
specific T-cells from cHBV patients. In June 2022, we presented a poster at the 2022 EASL ILC highlighting data from a study that was designed to assess
the  preclinical  activity  of  AB-101  and  the  compound’s  ability  to  reinvigorate  patient  HBV-specific  T-cells.  Studies  were  conducted  using  a  transgenic
MC38  tumor  mouse  model  and  peripheral  blood  mononuclear  cells  (PBMCs)  from  cHBV  patients.  The  data  presented  showed  that  once  daily  oral
administration of AB-101 resulted in profound tumor reduction that was associated with T-cell activation. In addition, AB-101 activates and reinvigorates
HBV-specific  T-cells  in  vitro.  Additionally,  preclinical  data  in  an  HBV  mouse  model  was  presented  at  the  2022  AASLD  Liver  Meeting  showing  that
monotherapy with AB-101 reduced PD-L1 in liver immune cells, confirming liver target engagement of the compound. Combination treatment with AB-
101 and an HBV-targeting GalNAc-siRNA agent resulted in activation and increased frequency of HBV-specific T-cells and greater anti-HBsAg antibody
production. This favorable preclinical profile supports further development of AB-101 as a therapeutic modality for cHBV treatment. We believe AB-101,
when used in combination with other approved and investigational agents, could potentially lead to a functional cure in HBV chronically infected patients.
We anticipate initiating a Phase 1 healthy subject clinical trial with AB-101 in the first half of 2023 with data from the single-ascending dose portion of the
clinical trial expected in the second half of 2023.

We  are  also  exploring  potential  oncology  applications  for  our  internal  PD-L1  portfolio.  Preclinical  data  was  selected  for  publication  at  the  American
Society of Clinical Oncology (ASCO) Annual Meeting in June 2022 showing that our oral small-molecule PD-L1 inhibitors in development, which possess
a  novel  mechanism  of  action,  have  the  ability  to  mediate  T-cell  activation  in  primary  human  immune  cells.  The  anti-tumor  efficacy  seen  in  vivo  was
comparable to anti-PD-L1 antibodies. The data is published in the Journal of Clinical Oncology.

Oral HBV RNA Destabilizer (AB-161)

HBV RNA destabilizers are small molecule orally available agents that cause the destabilization and ultimate degradation of HBV RNAs. Mechanistically,
RNA destabilizers target the host proteins PAPD5/7, which are involved in regulating the stability of HBV RNA transcripts.  In doing so, RNA destabilizers
lead to the selective degradation of HBV RNAs, thus reducing HBsAg levels and inhibiting viral replication. To provide a proprietary all-oral treatment
regimen  for  patients  with  cHBV,  we  believe  inclusion  of  a  small  molecule  RNA  destabilizer  is  key.  HBV  RNA  destabilizers  have  the  potential  to
complement or replace subcutaneously delivered RNAi agents, such as AB-729.

AB-161 is our next-generation oral small molecule RNA destabilizer specifically designed to target the liver. We have conducted extensive non-clinical
safety  evaluations  with  AB-161  that  provide  confidence  in  this  molecule’s  ability  to  circumvent  the  peripheral  neuropathy  findings  seen  in  non-clinical
safety studies with our first-generation oral RNA destabilizer, AB-452. We recently presented preclinical data at the 2022 Discovery on Target Conference
showing  that  AB-161  reduced  HBV  RNA  and  HBsAg  in  multiple  preclinical  models,  with  favorable  liver  centricity  and  lack  of  observed  peripheral
neuropathy. We anticipate initiating a Phase 1 healthy subject clinical trial with AB-161 in the first half of 2023 with single-ascending dose data expected in
the second half of 2023.

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Coronavirus Program

While our core mission is to find a cure for HBV, the magnitude of the coronavirus pandemic is undeniable. Given our science team’s proven expertise in
the discovery of new antiviral therapies, in 2020 we initiated a drug discovery effort for treating coronaviruses, including COVID-19. To that end, we have
assembled  an  internal  team  of  expert  scientists  under  the  direction  of  our  Chief  Scientific  Officer,  Dr.  Michael  Sofia,  to  identify  novel  small  molecule
therapies to treat COVID-19 and future coronavirus outbreaks. Dr. Sofia, who was awarded the Lasker-DeBakey Award for his discovery of sofosbuvir,
brings extensive antiviral drug discovery experience to this program. As we strive to identify and develop new antiviral small molecules to treat COVID-19
and  future  coronavirus  outbreaks,  we  have  focused  our  research  efforts  on  two  essential  targets  critical  for  replication  across  all  coronaviruses  –  nsp5
protease and nsp12 polymerase. These targets are essential viral proteins that our science team has experience in targeting.

Oral M  Inhibitor (AB-343)

pro

AB-343  is  our  lead  coronavirus  drug  candidate  that  inhibits  M .  In  our  pre-clinical  research  conducted  to  date,  AB-343  has  shown  pan-coronavirus
antiviral activity, no reduction in potency against known SARS-CoV-2 variants, robust activity against SARS-CoV-2 M  resistant strains, and a favorable
drug-drug interaction profile with no need for ritonavir boosting. We anticipate completing IND-enabling studies and initiating a Phase 1 clinical trial with
AB-343 in the second half of 2023. We also intend to nominate a nsp12 clinical candidate and initiate IND-enabling studies in the second half of 2023. An
nsp12 viral polymerase could potentially be combined with AB-343 to achieve better patient treatment outcomes and for use in prophylactic settings. 

pro

pro

Collaboration with X-Chem, Inc. and Proteros biostructures GmbH

In March 2021, we entered into a discovery research and license agreement, as amended, with X-Chem, Inc. (“X-Chem”) and Proteros biostructures GmbH
(“Proteros”) to focus on the discovery of novel inhibitors targeting the SARS-CoV-2 nsp5 main protease (M ). The agreement is designed to accelerate the
development of pan-coronavirus agents to treat COVID-19 and potential future coronavirus outbreaks. This collaboration brought together our expertise in
the discovery and development of antiviral agents with X-Chem’s industry leading DNA-encoded library (DEL) technology and Proteros’ protein sciences,
biophysics and structural biology capabilities and provides important synergies to potentially identify safe and effective therapies against coronaviruses,
including SARS-CoV-2. The collaboration allows for the rapid screening of one of the largest small molecule libraries against M  (an essential protein
required for the virus to replicate itself) and the use of state-of-the-art structure guided methods to rapidly optimize M  inhibitors to progress to clinical
candidates.  The  agreement  provides  for  payments  by  us  to  X-Chem  and  Proteros  upon  satisfaction  of  certain  development,  regulatory  and  commercial
milestones, as well as royalties on sales. Through this collaboration, we identified and obtained a worldwide exclusive license to several molecules that
inhibit M , a validated target for the treatment of COVID-19 and potential future coronavirus outbreaks.

pro

pro

pro

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COVID-19 Impact

The COVID-19 pandemic has resulted in and will likely continue to result in significant disruptions to businesses. Measures implemented around the world
in attempts to slow the spread of COVID-19 have had, and will likely continue to have, a major impact on clinical development, at least in the near-term,
including shortages and delays in the supply chain and prohibitions in certain countries on enrolling patients in new clinical trials. While we have been able
to progress with our clinical and pre-clinical activities to date, it is not possible to predict if the COVID-19 pandemic will materially impact our plans and
timelines in the future.

Other Collaborations, Royalty Entitlements and Intellectual Property Litigation

Collaboration with Qilu Pharmaceutical Co., Ltd. (“Qilu”)

In December 2021, we entered into a technology transfer and license agreement (the “License Agreement”) with Qilu, pursuant to which we granted Qilu a
sublicensable, royalty-bearing license, under certain intellectual property owned by us, which is non-exclusive as to development and manufacturing and
exclusive with respect to commercialization of AB-729, including

14

pharmaceutical products that include AB-729, for the treatment or prevention of hepatitis B in China, Hong Kong, Macau and Taiwan (the “Territory”).

In partial consideration for the rights granted by us, Qilu paid us a one-time upfront cash payment of $40 million on January 5, 2022 and agreed to pay us
milestone payments totaling up to $245 million, net of withholding taxes, upon the achievement of certain technology transfer, development, regulatory and
commercialization milestones. Qilu also agreed to pay us double digit royalties into the low twenties percent based upon annual net sales of AB-729 in the
Territory. The royalties are payable on a product-by-product and region-by-region basis, subject to certain limitations.

Qilu is responsible for all costs related to developing, obtaining regulatory approval for, and commercializing AB-729 for the treatment or prevention of
hepatitis B in the Territory. Qilu is required to use commercially reasonable efforts to develop, seek regulatory approval for, and commercialize at least one
AB-729  product  candidate  in  the  Territory.  A  joint  development  committee  has  been  established  between  us  and  Qilu  to  coordinate  and  review  the
development, manufacturing and commercialization plans. Both parties also have entered into a supply agreement and related quality agreement pursuant
to which we will manufacture or have manufactured and supply Qilu with all quantities of AB-729 necessary for Qilu to develop and commercialize in the
Territory until we have completed manufacturing technology transfer to Qilu and Qilu has received all approvals required for it or its designated contract
manufacturing organization to manufacture AB-729 in the Territory.

Concurrent with the execution of the License Agreement, we entered into a Share Purchase Agreement (the “Share Purchase Agreement”) with Anchor
Life Limited, a company established pursuant to the applicable laws and regulations of Hong Kong and an affiliate of Qilu (the “Investor”), pursuant to
which the Investor purchased 3,579,952 of our common shares, without par value (the “Common Shares”), at a purchase price of USD $4.19 per share,
which was a 15% premium on the thirty-day average closing price of the Common Shares as of the close of trading on December 10, 2021 (the “Share
Transaction”). We received $15.0 million of gross proceeds from the Share Transaction on January 6, 2022. The Common Shares sold to the Investor in the
Share  Transaction  represented  approximately  2.5%  of  the  Common  Shares  outstanding  immediately  prior  to  the  execution  of  the  Share  Purchase
Agreement.

Alnylam Pharmaceuticals, Inc. (“Alnylam”) and Acuitas Therapeutics, Inc. (“Acuitas”)

We have two royalty entitlements to Alnylam’s global net sales of ONPATTRO.

In  2012,  we  entered  into  a  license  agreement  with  Alnylam  that  entitles  Alnylam  to  develop  and  commercialize  products  with  our  lipid  nanoparticle
(“LNP”) delivery technology. Alnylam’s ONPATTRO, which represents the first approved application of our LNP technology, was approved by the United
States FDA and the European Medicines Agency (“EMA”) during the third quarter of 2018 and was launched by Alnylam immediately upon approval in
the United States. Under the terms of this license agreement, we are entitled to tiered royalty payments on global net sales of ONPATTRO ranging from
1.00% - 2.33% after offsets, with the highest tier applicable to annual net sales above $500 million. This royalty interest was sold to the Ontario Municipal
Employees Retirement System (“OMERS”), effective as of January 1, 2019, for $20 million in gross proceeds before advisory fees. OMERS will retain this
entitlement until it has received $30 million in royalties, at which point 100% of this royalty entitlement on future global net sales of ONPATTRO will
revert to us. OMERS has assumed the risk of collecting up to $30 million of future royalty payments from Alnylam and we are not obligated to reimburse
OMERS if they fail to collect any such future royalties. If this royalty entitlement reverts to us, it has the potential to provide an active royalty stream or to
be  otherwise  monetized  again  in  full  or  in  part.  From  the  inception  of  the  royalty  sale  through  December  31,  2022,  an  aggregate  of  $18.9  million  of
royalties have been collected by OMERS.

We also have rights to a second royalty interest ranging from 0.75% to 1.125% on global net sales of ONPATTRO, with 0.75% applying to sales greater
than $500 million, originating from a settlement agreement and subsequent license agreement with Acuitas. This royalty entitlement from Acuitas has been
retained by us and was not part of the royalty entitlement sale to OMERS.

Genevant Sciences, Ltd.

In  April  2018,  we  entered  into  an  agreement  with  Roivant  Sciences  Ltd.  (“Roivant”),  our  largest  shareholder,  to  launch  Genevant  Sciences  Ltd.
(“Genevant”), a company focused on a broad range of RNA-based therapeutics enabled by our LNP

15

and  ligand  conjugate  delivery  technologies.  We  licensed  rights  to  our  LNP  and  ligand  conjugate  delivery  platforms  to  Genevant  for  RNA-based
applications outside of HBV, except to the extent certain rights had already been licensed to other third parties (the “Genevant License”). We retained all
rights to our LNP and conjugate delivery platforms for HBV.

Under the Genevant License, as amended, if a third party sublicensee of intellectual property licensed by Genevant from us commercializes a sublicensed
product, we become entitled to receive a specified percentage of certain revenue that may be received by Genevant for such sublicense, including royalties,
commercial milestones and other sales-related revenue, or, if less, tiered low single-digit royalties on net sales of the sublicensed product. The specified
percentage is 20% in the case of a mere sublicense (i.e., naked sublicense) by Genevant without additional contribution and 14% in the case of a bona fide
collaboration with Genevant.

Additionally,  if  Genevant  receives  proceeds  from  an  action  for  infringement  by  any  third  parties  of  our  intellectual  property  licensed  to  Genevant,  we
would be entitled to receive, after deduction of litigation costs, 20% of the proceeds received by Genevant or, if less, tiered low single-digit royalties on net
sales of the infringing product (inclusive of the proceeds from litigation or settlement, which would be treated as net sales).

In July 2020, Roivant recapitalized Genevant through an equity investment and conversion of previously issued convertible debt securities held by Roivant.
We  participated  in  the  recapitalization  of  Genevant  with  an  equity  investment  of  $2.5  million.  In  connection  with  the  recapitalization,  the  three  parties
entered into an Amended and Restated Shareholders Agreement that provides Roivant with substantial control of Genevant. We have a non-voting observer
seat on Genevant’s Board of Directors. As of December 31, 2022, we owned approximately 16% of the common equity of Genevant and the carrying value
of  our  investment  in  Genevant  was  zero.  Our  entitlement  to  receive  future  royalties  or  sublicensing  revenue  from  Genevant  was  not  impacted  by  the
recapitalization.

Moderna Inter Partes Review Petitions

On  February  21,  2018  and  March  5,  2018,  Moderna  Therapeutics,  Inc.  (“Moderna”)  filed  petitions  requesting  the  United  States  Patent  and  Trademark
Office (“USPTO”) to institute an Inter Partes Review of Arbutus United States Patents 9,404,127 (the “’127 Patent”) and 9,364,435 (the “’435 Patent”). In
its petitions, Moderna sought to invalidate all claims of each patent based on Moderna’s allegation that the claims are anticipated and/or obvious. We filed a
response  to  Moderna’s  petitions  on  June  14,  2018.  On  September  12,  2018,  the  Patent  Trial  and  Appeal  Board  (the  “PTAB”)  rendered  its  decision  to
institute Inter Partes Review of both the ‘127 Patent and the ‘435 Patent.

The status of these patents, which collectively represent only a fraction of our extensive LNP patent portfolio, is as follows:

‘127 Patent

With respect to the ‘127 Patent, the PTAB held all claims as invalid on September 10, 2019, by reason of anticipatory prior art. However this decision was
vacated and sent back (remanded) to the PTAB for a rehearing, pending the Supreme Court’s decision whether to grant certiorari in a different case, United
States v. Athrex, Inc. (“US v. Athrex”), the holding of which could impact the findings in the ‘127 Patent matter. The Supreme Court granted certiorari in
US v. Athrex on October 13, 2020 (i.e. agreed to review the decision appealed from a lower court). Until the Supreme Court rendered its opinion in US v.
Athrex, the ‘127 Patent hearing remained in abeyance, with no decision reached as to the validity of its claims. The Supreme Court decided on the US v.
Athrex  case  on  June  21,  2021,  following  which  the  Federal  Circuit  reinstated  the  appeal  sua  sponte,  requiring  the  parties  to  brief  how  the  case  should
proceed in light of the Supreme Court’s opinion or for the Appellant to waive the challenge. We elected to waive the challenge and proceed with the appeal
at the Federal Circuit. The opening brief was filed on October 25, 2021. Moderna’s responsive brief was filed on February 24, 2022 and our reply brief was
filed on April 26, 2022. An oral hearing for this matter was held on November 4, 2022.

‘435 Patent

With respect to the ‘435 Patent, the PTAB rendered its decision on September 11, 2019, holding certain claims invalid and upholding other claims as valid.
On November 13, 2019, we and Moderna both appealed the decision. Moderna filed its opening brief on May 4, 2020 and we provided our opening and
responsive brief on July 27, 2020. Moderna subsequently filed its reply and responsive brief on October 5, 2020, and we filed our reply brief on November
9, 2020. An oral hearing on the

16

‘435 Patent was held on October 7, 2021. On December 1, 2021, the Federal Circuit issued its opinion, leaving intact the PTAB’s holding regarding the
validity of certain claims in the ‘435 Patent and the invalidity of other claims in the ‘435 Patent. The decision in the ‘435 appeal was rendered final by
mandate on January 25, 2022.

‘069 Patent

On January 9, 2019, Moderna filed an additional petition requesting Inter Partes Review of Arbutus United States Patent 8,058,069 (the “’069 Patent”). The
PTAB instituted Inter Partes Review of the ‘069 Patent and, on July 23, 2020, issued a decision upholding all claims as valid. On September 23, 2020,
Moderna appealed the ‘069 Inter Partes Review decision to the Federal Circuit Court of Appeals. Moderna filed its opening brief in that appeal on February
23, 2021, we filed our responsive brief on May 11, 2021, and Moderna filed its reply brief on July 1, 2021. An oral hearing on the ‘069 Patent was held on
October 7, 2021, in a joint hearing with the hearing regarding the ‘435 patent, before the U.S. Court of Appeals for the Federal Circuit. On December 1,
2021,  the  Federal  Circuit  also  issued  its  ruling  with  respect  to  the  ‘069  Patent,  affirming  the  PTAB’s  finding  that  all  claims  were  valid.  The  Federal
Circuit’s decision in the ‘069 appeal was rendered final by mandate on January 10, 2022.

Moderna and Merck European Oppositions

On April 5, 2018, Moderna and Merck, Sharp & Dohme Corporation (“Merck”) filed Notices of Opposition to Arbutus’ European patent EP 2279254 (“the
’254  Patent”)  with  the  European  Patent  Office  (“EPO”),  requesting  that  the  ‘254  Patent  be  revoked  in  its  entirety  for  all  contracting  states.  We  filed  a
response to Moderna and Merck’s oppositions on September 3, 2018. A hearing was conducted before the Opposition Division of the EPO on October 10,
2019.  At  the  conclusion  of  the  hearing,  the  EPO  upheld  an  auxiliary  request  adopting  the  amendment,  as  put  forth  by  us,  of  certain  claims  of  the  ‘254
Patent. In February 2020 Moderna and Merck filed Notices of Appeal challenging the EPO’s grant of the auxiliary request. Merck filed its notice of appeal
on February 24, 2020 and Moderna on February 27, 2020. Both Merck and Moderna perfected their appeals by filing Grounds of Appeal on April 30, 2020.
We  filed  our  responses  to  the  appeals  on  September  18,  2020.  On  March  22,  2022,  Moderna  filed  further  written  submissions  to  which  Arbutus  and
Genevant responded in August 2022. The date for the oral proceedings has not been set.

While we are the patent holder, the ‘127 Patent, the ‘435 Patent, the ‘069 Patent and the ‘254 Patent have been licensed to Genevant and are included in the
rights licensed by us to Genevant under the Genevant License.

Patent Infringement Litigation vs. Moderna

On February 28, 2022, we and Genevant filed a lawsuit in the U.S. District Court for the District of Delaware against Moderna, Inc. and a Moderna affiliate
seeking damages for infringement of U.S. Patent Nos. 8,058,069, 8,492,359, 8,822,668, 9,364,435, 9,504,651, and 11,141,378 in the manufacture and sale
of  MRNA-1273,  Moderna’s  vaccine  for  COVID-19.  The  patents  relate  to  nucleic  acid-lipid  particles  and  lipid  vesicles,  as  well  as  compositions  and
methods for their use. The lawsuit does not seek an injunction or otherwise seek to impede the sale, manufacture or distribution of MRNA-1273. However,
we  seek  fair  compensation  for  Moderna’s  use  of  our  patented  technology  that  was  developed  with  great  effort  and  at  great  expense,  without  which
Moderna’s  COVID-19  vaccine  would  not  have  been  successful.  On  May  6,  2022,  Moderna  filed  a  partial  motion  to  dismiss  the  claims  “relating  to
Moderna’s sale and provision of COVID-19 vaccine doses to the U.S. Government.” On November 2, 2022, the Court issued an Order denying Moderna’s
motion. On November 30, 2022, Moderna filed its Answer to the Complaint and Counterclaims. Arbutus and Genevant filed their Answer to Moderna’s
Counterclaims on December 21, 2022. On February 14, 2023, the U.S. Dept. of Justice filed a Statement of Interest in the action. On February 16, 2023,
the Court held an Initial Pretrial Conference after which it issued an Order, dated February 16, 2023, ordering that within 14 days of the issuance of the
Order, the parties and the U.S. Government are to submit letters regarding the impact of the Governments’ Statement of Interest on the scheduling of the
matter.

Acuitas Declaratory Judgment Lawsuit

On March 18, 2022, Acuitas filed a lawsuit against us and Genevant in the Southern District of New York, asking the court to enter declaratory judgment
that Arbutus patent Nos. 8,058,069, 8,492,359, 8,822,668, 9,006,417, 9,364,435, 9,404,127,

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9,504,651,  9,518,272,  and  11,141,378  do  not  infringe  Pfizer  and  BioNTech’s  COVID-19  vaccine,  COMIRNATY,  which  uses  an  mRNA  lipid  provided,
under  license,  by  Acuitas.  Acuitas  also  seeks  a  declaration  that  each  of  the  listed  patents  is  invalid.  On  June  24,  2022,  we  and  Genevant  sought  a  pre-
motion conference concerning our anticipated motion to dismiss all of Acuitas’ claims due to lack of subject matter jurisdiction. The request for a pre-
motion conference was granted, but the case was subsequently re-assigned to a new judge who entered an order directing: (i) Acuitas to inform the court
whether it intended to file an amended complaint; (ii) that Acuitas must file any amended complaint by a certain date; and (iii) that if Acuitas did not file an
amended complaint, we and Genevant must file our motion to dismiss by a certain date. Acuitas filed its amended complaint on September 6, 2022. On
October  4,  2022,  we  and  Genevant  filed  our  motion  to  dismiss  the  Acuitas  action  for  lack  of  subject  matter  jurisdiction  based  on  the  lack  of  a  case  or
controversy. Acuitas filed its opposition to the motion to dismiss on November 1, 2022 and we and Genevant filed our reply brief on November 16, 2022.
The motion is now fully briefed. No case schedule is yet in place.

Potential Additional Payments Related to the Acquisition of Enantigen Therapeutics, Inc.

In October 2014, Arbutus Inc., our wholly-owned subsidiary, acquired all of the outstanding shares of Enantigen Therapeutics, Inc. (“Enantigen”) pursuant
to  a  stock  purchase  agreement.  The  amount  paid  to  Enantigen’s  selling  shareholders  could  be  up  to  an  additional  $102.5  million  in  sales  performance
milestones in connection with the sale of the first commercialized product by us for the treatment of HBV, regardless of whether such product is based upon
assets  acquired  under  this  agreement,  and  a  low  single-digit  royalty  on  net  sales  of  such  first  commercialized  HBV  product,  up  to  a  maximum  royalty
payment of $1.0 million that, if paid, would be offset against our performance milestone payment obligations.

Patents and Proprietary Rights

Our commercial success depends in part on our ability to obtain and maintain proprietary protection for our product candidates, novel discoveries, product
development technologies and other know-how, to operate without infringing on the proprietary rights of others and to prevent others from infringing our
proprietary rights. Our policy is to seek to protect our proprietary position by, among other methods, filing or in licensing United States and foreign patents
and patent applications related to our proprietary technology, inventions and improvements that are important to the development and implementation of
our business. We also rely on trademarks, trade secrets, know how, continuing technological innovation and potential in licensing opportunities to develop
and maintain our proprietary position.

In addition to our proprietary expertise, we own a portfolio of patents and patent applications directed to HBV core/capsid protein assembly inhibitors,
HBV  surface  antigens  secretion  inhibitors,  coronavirus  main  protease  inhibitors,  coronavirus  Nsp12  inhibitors,  LNP  inventions,  LNP  compositions  for
delivering  nucleic  acids  such  as  mRNA  and  RNAi,  the  formulation  and  manufacture  of  LNP-based  pharmaceuticals,  chemical  modification  of  RNAi
molecules, and RNAi drugs and processes directed at particular disease indications. In the United States our patents might be challenged by inter partes
review or opposition proceedings. In Europe, upon grant, a period of nine months is allowed for notification of opposition to such granted patents.  If our
patents are subjected to inter partes review or opposition proceedings, we would incur significant costs to defend them. Further, our failure to prevail in any
such  proceedings  could  limit  the  patent  protection  available  to  our  therapeutic  HBV  programs,  coronavirus  programs  or  RNAi  platform,  including  our
product candidates.

We  own  more  than  65  patent  families  related  to  our  compounds,  formulations,  and  technology,  but  we  cannot  be  certain  that  issued  patents  will  be
enforceable or provide adequate protection or that pending patent applications will result in issued patents.

The following table shows the estimated expiration dates, based on filing dates of pending patent applications, in the United States and the European Union
for the primary patents for our product candidates currently in clinical trials.

Product candidate

AB-729

Estimated Patent Expiration in US

Estimated Patent Expiration in EU

2038

2038

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Human Capital

Employee Composition

As of December 31, 2022, we had 98 employees (96 full-time and 2 part-time), 76 of whom were engaged in research and development, including three
medical doctors, 35 individuals with Doctors of Philosophy (PhDs) degrees, and another 10 individuals with Master of Science degrees. Our workforce is
49% female and 31% of our employees holding a position of vice president or higher are female. None of our employees are represented by a labor union
or covered by a collective bargaining agreement, nor have we experienced any work stoppages. We believe that relations with our employees are good. We
supplement our in-house expertise with outsourced capabilities when it would be cost prohibitive to build our own in-house capabilities. For example, we
outsource  a  substantial  portion  of  our  clinical  trial  work  to  clinical  research  organizations  and  a  majority  of  our  drug  manufacturing  is  out-sourced  to
contract manufacturers. Our in-house clinical development and manufacturing teams implement our development strategies and oversee the activities of
our outside vendors.

Employee Oversight, Training and Development

We  are  invested  in  the  professional  development  of  our  employees.  In  order  to  promote  long-term  retention  and  to  maximize  the  potential  of  our
employees, we provide individualized performance management programs. We also offer needs-based supplemental training to our employees. In order to
monitor  employee  satisfaction  and  as  well  to  identify  ways  in  which  employee  satisfaction  and  engagement  can  be  improved,  we  also  survey  our
employees  on  a  regular  basis,  reporting  the  results  of  the  surveys  to  management  and  to  our  board  of  directors.  In  2022,  we  experienced  our  lowest
employee  turnover  in  the  previous  seven  years,  while  many  other  companies  experienced  their  highest  in  the  midst  of  an  historically  competitive  job
market. Given our financial resources and our track record, we were able to hire 17 new employees in 2022 to support our expanding pipeline of research
programs and product candidates.

Compensation and Benefits

Drug development is a complex endeavor that requires deep expertise and attracting and retaining qualified employees for specialized biopharmaceutical
positions. Our compensation programs are designed to attract and retain top talent. We offer every employee a total compensation package consisting of
base  salary,  cash  target  bonus  targeting  the  50th  to  75th  percentile  of  market  based  on  company  size  and  industry,  a  comprehensive  benefit  package,
including  medical,  dental  and  vision  health  care  coverage,  a  401(k)  plan  with  an  employer  match,  tax-advantaged  savings  accounts  and  equity
compensation  for  every  employee,  which  includes  stock  options  and  restricted  stock  units.  We  also  provide  eligible  employees  the  opportunity  to
participate in our employee stock purchase plan and our employee rewards and recognition programs. In addition, we provide our employees with wellness
programs and we offer mental health support to our employees and dependents.

Work-life Balance

We aim to ensure our employees maintain a work-life balance by offering 25 paid days of time-off, 12 days of paid holidays, and we shut down in the last
week  of  December.  We  provide  paid  parental  leave  to  both  birth  and  adoptive  parents.  In  addition,  we  allow  our  employees  to  have  a  flexible  work
schedule and, to the extent possible, depending on the nature of the work, remote and hybrid work arrangements. We believe our focus on total rewards and
work-life balance contributed to our having been named one of Philadelphia Business Journal’s Best Places to Work in 2022, a prestigious award that is
based on employee survey results.

Environmental, Social and Governance

Environmental

We are a pre-commercial company of less than one hundred employees, engaged in research and development. Manufacturing activities to support these
activities is almost entirely outsourced and biohazardous and chemical waste disposal is handled by

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third party vendors. Although our environmental footprint is subsequently small, we regularly review and evaluate our energy use to identify ways in which
we can maximize efficiencies and minimize waste.

Social

The culture at Arbutus reflects our commitment to our employees, to our community, and to making a meaningful contribution to world health. We are
active in community outreach and participate in many local charities serving underserved communities in the Philadelphia area, including partnering with
Life Sciences Cares Philadelphia.

Safety in the Workplace

We strive to provide a productive and safe working environment for our employees. To protect the health and safety of our employees, we have a Health
and  Safety  Committee,  officially  certified  by  the  PA  Department  of  Labor  and  Industry  -  Bureau  of  Workers  Compensation,  which  is  committed  to  the
principles of leadership, responsibility, prevention, and compliance. We follow all recognized Environmental Health and Safety standards and management
systems. We have also established an Occupational Health and Safety policy and related standard operating procedures, all of which are used to train our
employees in the proper procedures for the workplace. We also solicit employee and contractor recommendations to improve on the safety of our working
conditions.

Diversity, Equity and Inclusion

Our commitment to diversity and inclusion is demonstrated by our placement of ultimate responsibility for diversity, equity and inclusion with our board of
directors, informed by the recommendations of management and the board’s Nominating and Governance Committee. Our Code of Business Conduct (the
“Code of Conduct”) prohibits discrimination and harassment of any kind, including discrimination or harassment based on age, race, ethnicity, religion,
gender,  sexual  preference  and  disability.  In  addition  to  our  anti-harassment  and  human  rights  policies,  we  also  require  mandatory  annual  training  in
unconscious  bias  and  anti-harassment.  Some  of  the  diversity  and  inclusion  initiatives  at  Arbutus  include  the  formation  of  a  Diversity  and  Inclusion
Committee comprised of Arbutus employees and the broadening of the geographical reach of our recruitment efforts. We also celebrate Juneteenth as a
corporate holiday.

Our Contribution to World Health

We are dedicated to meaningfully contributing to world health. We are pursuing the mission of finding a cure for Hepatitis B viral infections, an unmet
medical need affecting over 290 million people worldwide, and we are working to develop a treatment for coronaviruses, including COVID-19.

Governance

As stated in our Code of Conduct, we are committed to complying with all applicable laws, rules and regulations not just in the United States and Canada,
but in all the countries in which we operate. In addition to mandating training on our Code of Conduct on an annual basis, we also provide annual training
on insider trading, anti-bribery and anti-corruption, among other topics. In addition, we require our suppliers’ agreement to comply with anti-bribery and
anti-fraud provisions, and to comply with all applicable laws. All vendors also receive our Code of Conduct at the time of their engagement with us. We
comply with all applicable regulations in conducting clinical trials, including FDA ethical regulations, the Declaration of Helsinki and the International
Conference on Harmonisation - good Clinical Practices (ICH-GCP).

Competition

We face a broad range of current and potential competitors, from established global pharmaceutical companies with significant resources, to research-stage
companies.  In  addition,  we  face  competition  from  academic  and  research  institutions  and  government  agencies  for  the  discovery,  development  and
commercialization of novel therapeutics to treat HBV and coronaviruses. Many of our competitors, either alone or with their collaborative partners, have
significantly greater financial, product development, technical, manufacturing, sales, and marketing resources than we do. In addition, many of our direct
competitors are large pharmaceutical companies with internal research and development departments that have significantly

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greater experience in testing product candidates, obtaining FDA and other regulatory approvals of product candidates, and achieving widespread market
acceptance for those products.

As  a  significant  unmet  medical  need  exists  for  HBV,  there  are  several  large  and  small  pharmaceutical  companies  focused  on  delivering  singular  or
combinations of therapeutics for the treatment of HBV. These companies include, but are not limited to, Johnson & Johnson, Roche, Vir Biotechnology,
GlaxoSmithKline, Gilead Sciences, Assembly, Enanta Pharmaceuticals, Aligos Therapeutics and Vaccitech. These companies are developing products such
as  antisense  oligonucleotides,  capsid  inhibitors,  RNAi  therapeutics,  immune  modulators  and  surface  antigen  inhibitors.  These  product  candidates  are  in
various stages of pre-clinical and clinical development. Further, in addition to current investigational therapeutics in development, it is likely that additional
drugs will become available in the future for the treatment of HBV.

In addition, given the severity of the global coronavirus pandemic, several companies are developing or commercializing therapeutics for the treatment of
coronaviruses. These companies include, but are not limited to, Pfizer, Merck, Gilead, Vir Biotechnology, Shionogi, PardesBio, Enanta Pharmaceuticals,
Aligos Therapeutics and Cocrystal Pharma.

We  anticipate  that  we  will  face  competition  as  new  products  enter  the  marketplace.  Our  competitors’  products  may  be  safer,  more  effective,  or  more
effectively  marketed  and  sold  than  any  product  we  may  commercialize.  Competitive  singular  or  combination  products  may  render  one  or  more  of  our
product candidates obsolete or non-competitive before we can recover the expenses of developing and commercializing any of our product candidates. It is
also possible that the development of a cure or new treatment methods for HBV or coronaviruses could render one or more of our product candidates non-
competitive, obsolete, or reduce the demand for our product candidates.

We  believe  that  our  ability  to  compete  depends,  in  part,  upon  our  ability  to  develop  products,  successfully  complete  the  clinical  trials  and  regulatory
approval processes, and effectively market any approved products. Further, we need to attract and retain qualified personnel, obtain patent protection or
otherwise develop proprietary product candidates or processes, and secure sufficient capital resources for the substantial time period between the discovery
of lead compounds and their commercial sales, if any.

Manufacturing

We  currently  rely  on  third-party  manufacturers  to  supply  drug  substance  and  drug  products,  including  AB-729,  AB-101,  AB-161  and  AB-343,  for  our
ongoing and anticipated clinical trials and non-clinical studies. We currently have no plans to establish any large-scale internal manufacturing facilities for
our product candidates.

Government Regulation

Regulation by governmental authorities in the United States and in other countries is a significant consideration in our product development, manufacturing
and, if our product candidates are approved, marketing strategies. We expect that all our product candidates will require regulatory approval by the FDA
and  by  similar  regulatory  authorities  in  foreign  countries  prior  to  commercialization  and  will  be  subjected  to  rigorous  pre-clinical,  clinical,  and  post-
approval testing to demonstrate safety and effectiveness, as well as other significant regulatory requirements and restrictions in each jurisdiction in which
we would seek to market our products. In the United States, we are subject to extensive regulation by the FDA and other federal, state, and local regulatory
agencies.  United  States  federal  laws,  such  as  the  Federal  Food,  Drug,  and  Cosmetic  Act  (“FD&C  Act”),  and  regulations  issued  thereunder,  govern  the
testing, development, manufacture, quality control, safety, effectiveness, approval, storage, labeling, record keeping, reporting, distribution, import, export,
sale, and marketing of all biopharmaceutical products intended for therapeutic purposes. We believe that we and the third parties that work with us are in
compliance in all material respects with currently applicable laws, rules and regulations; however, any failure to comply could have a material negative
impact on our ability to successfully develop and commercialize our products, and therefore on our financial performance. In addition, the laws, rules and
regulations that apply to our business are subject to change and it is difficult to foresee whether, how, or when such changes may affect our business.

Obtaining governmental approvals to market our product candidates and maintaining ongoing compliance with applicable federal, state, local and foreign
statutes and regulations following any such approvals will require the expenditure of significant financial and human resources.

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Development and Approval

The process to develop and obtain approval for biopharmaceutical products for commercialization in the United States and many other countries is lengthy,
complex and expensive, and the outcome is far from certain. Although foreign requirements for conducting clinical trials and obtaining approval may differ
in certain respects from those in the United States, there are many similarities and they often are equally rigorous, and the outcome cannot be predicted
with confidence. A key component of any submission for approval in any jurisdiction is pre-clinical and clinical data demonstrating the product candidate’s
safety and effectiveness.

Pre-clinical Testing. Before testing any product candidate in humans in the United States, a company must develop pre-clinical data, generally including
laboratory evaluation of the product candidate’s chemistry and formulation, as well as toxicological and pharmacological studies in animal species to assess
safety and quality. Certain types of animal studies must be conducted in compliance with the FDA’s Good Laboratory Practice (“GLP”) regulations and the
Animal Welfare Act, which is enforced by the Department of Agriculture.

IND Application. A person or entity sponsoring clinical trials in the United States to evaluate a product candidate’s safety and effectiveness must submit to
the FDA, prior to commencing such trials, an investigational new drug (“IND”) application, which contains, among other data and information, pre-clinical
testing results and provides a basis for the FDA to conclude that there is an adequate basis for testing the drug in humans. If the FDA does not object to the
IND application within 30 days of submission, the clinical testing proposed in the IND may begin. Even after the IND has gone into effect and clinical
testing has begun, the FDA may put the clinical trials on “clinical hold,” suspending (or in some cases, ending) them because of safety concerns or for other
reasons.

Clinical  Trials.  Clinical  trials  involve  administering  a  product  candidate  to  human  volunteers  or  patients  under  the  supervision  of  a  qualified  clinical
investigator.  Clinical  trials  are  subject  to  extensive  regulation.  In  the  United  States,  this  includes  compliance  with  the  FDA’s  bioresearch  monitoring
regulations  and  current  good  clinical  practices  (“GCP”)  requirements,  which  establish  standards  for  conducting,  recording  data  from,  and  reporting  the
results of clinical trials, with the goals of assuring that the data and results are credible and accurate and that study participants’ rights, safety and well-
being  are  protected.  Each  clinical  trial  must  be  conducted  under  a  protocol  that  details,  among  other  things,  the  study  objectives  and  parameters  for
monitoring safety and the efficacy criteria, if any, to be evaluated. The protocol is submitted to the FDA as part of the IND and reviewed by the agency.
Additionally, each clinical trial must be reviewed, approved and conducted under the auspices of an Institutional Review Board (“IRB”). The sponsor of a
clinical trial, the investigators and IRBs each must comply with requirements and restrictions that govern, among other things, obtaining informed consent
from each study subject, complying with the protocol and investigational plan, adequately monitoring the clinical trial, and timely reporting AEs. Foreign
studies conducted under an IND must meet the same requirements applicable to studies conducted in the United States. However, if a foreign study is not
conducted under an IND, the data may still be submitted to the FDA in support of a product application, if the study was conducted in accordance with
GCP and the FDA is able to validate the data.

The  sponsor  of  a  clinical  trial  or  the  sponsor’s  designated  responsible  party  may  be  required  to  register  certain  information  about  the  trial  and  disclose
certain results on government or independent registry websites, such as clinicaltrials.gov.

Clinical testing is typically performed in three phases, which may overlap or be subdivided in some cases.

In  Phase  1  trials,  the  product  candidate  is  administered  to  a  small  number  of  human  subjects  to  assess  its  safety  and  to  develop  detailed  profiles  of  its
pharmacological  and  pharmacokinetic  actions  (i.e.,  absorption,  distribution,  metabolism  and  excretion),  assess  the  early  safety  profile,  determine  side
effects associated with increasing doses, and, if possible, gain early evidence of effectiveness. Although Phase 1 trials are typically conducted in healthy
human subjects, in some instances (including, for example, with some cancer therapies) the trial subjects are patients with the targeted disease or condition.

In  Phase  2  trials,  the  product  candidate  is  administered  to  a  relatively  small  sample  of  the  intended  patient  population  to  develop  initial  data  regarding
efficacy in the targeted disease, determine the optimal dose range, and generate additional information regarding the product candidate’s safety. Additional
animal toxicology studies may precede this phase.

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In Phase 3 trials, the product candidate is administered to a larger group of patients with the target disease or disorder, which may include patients with
concomitant diseases and medications. Typically, Phase 3 trials are conducted at multiple study sites and may be conducted concurrently for the sake of
time  and  efficiency.  The  purpose  of  Phase  3  clinical  trials  is  to  obtain  additional  information  about  safety  and  effectiveness  necessary  to  evaluate  the
product  candidate’s  overall  risk-benefit  profile  and  to  provide  a  basis  for  product  labeling.  Phase  3  data  often  form  the  core  basis  on  which  the  FDA
evaluates a product candidate’s safety and effectiveness when considering the product application.

The  study  sponsor,  the  FDA  or  an  IRB  may  suspend  or  terminate  a  clinical  trial  at  any  time  on  various  grounds,  including  a  determination  that  study
subjects  are  being  exposed  to  an  unacceptable  health  risk.  Success  in  early-stage  clinical  trials  does  not  assure  success  in  later-stage  clinical  trials.
Moreover, data from clinical trials are not always conclusive and may be subject to alternative interpretations that could delay, limit or prevent approval.

NDA Submission and Review. After completing the clinical studies, a sponsor seeking approval to market a product candidate in the United States submits
to  the  FDA  a  New  Drug  Application  (“NDA”).  The  NDA  is  a  comprehensive  application  intended  to  demonstrate  the  product  candidate’s  safety  and
effectiveness and includes, among other things, pre-clinical and clinical data, information about the product candidate’s composition, the sponsor’s plans
for manufacturing and packaging and proposed labeling. When an NDA is submitted, the FDA makes an initial determination as to whether the application
is  sufficiently  complete  to  be  accepted  for  review.  If  the  application  is  not,  the  FDA  may  refuse  to  accept  the  NDA  for  filing  and  request  additional
information. A refusal to file, which requires resubmission of the NDA with the requested additional information, delays review of the application.

FDA performance goals generally provide for action on an NDA within 10 months of the 60-day filing date, or within 12 months of the NDA submission.
That deadline can be extended under certain circumstances, including by the FDA’s requests for additional information. The targeted action date can also be
shortened  to  6  months  of  the  60-day  filing  date,  or  8  months  after  NDA  submission  for  product  candidates  that  are  granted  priority  review  designation
because they are intended to treat serious or life-threatening conditions and demonstrate the potential to address unmet medical needs. The FDA has other
programs  to  expedite  development  and  review  of  product  candidates  that  address  serious  or  life-threatening  conditions.  For  example,  the  Fast  Track
program is intended to facilitate the development and review of new drugs that demonstrate the potential to address unmet medical needs involving serious
or life-threatening diseases or conditions. If a product candidate receives Fast Track designation, the FDA may review sections of the NDA on a rolling
basis, rather than requiring the entire application to be submitted to begin the review. Product candidates with Fast Track designation also may be eligible
for  more  frequent  meetings  and  correspondence  with  the  FDA  about  the  product  candidate’s  development.  Another  FDA  program  intended  to  expedite
development is the Accelerated Approval pathway, which allows approval on the basis of a surrogate endpoint that is reasonably likely to predict clinical
benefit or on an intermediate clinical endpoint. To qualify for review under the Accelerated Approval pathway, a product candidate must treat a serious
condition, provide a meaningful advantage over available therapies, and demonstrate an effect on a surrogate endpoint that is reasonably likely to predict
clinical  benefit  or  on  an  intermediate  clinical  endpoint.  On  December  29,  2022,  Congress  enacted  the  Consolidated  Appropriations  Act  of  2023,  which
included several changes to the Accelerated Approval pathway within the Food and Drug Omnibus Reform Act (“FDORA”). Under FDORA, the FDA
must  specify  the  conditions  for  any  post-approval  studies  before  granting  an  Accelerated  Approval.  FDORA  gives  the  agency  significant  flexibility  in
setting  forth  such  conditions,  which  may  include  enrollment  targets,  study  protocol  and  milestones—including  the  target  date  of  study  completion.  The
FDA may also require, as appropriate, that certain post-approval studies be underway prior to Accelerated Approval or within a specified time from the
date of approval. Accelerated Approval sponsors are required to report progress every six months on required post-approval trials. Breakthrough Therapy
designation, which is available for product candidates under development for serious or life-threatening conditions and where preliminary clinical evidence
shows that the product candidate may have substantial improvement on at least one clinically significant endpoint over available therapies, means that a
product candidate will be eligible for all of the benefits of Fast Track designation, as well as more intensive guidance from the FDA on an efficient drug
development program and a commitment from the agency to involve senior FDA managers in such guidance. Even if a product candidate qualifies for Fast
Track designation or Breakthrough Therapy designation, the FDA may later decide that the product no longer meets the conditions for designation and may
rescind the designation, and/or may determine that the product does not meet the standards for approval. As applicable, we anticipate seeking to utilize
these programs to expedite the development and review of our product candidates, but we cannot ensure that our product candidates will qualify for such
programs, or that we will be able to maintain such designations if we qualify for such programs.

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The FDA reviews applications to determine, among other things, whether a product is safe and effective for its intended use and whether the manufacturing
controls  are  adequate  to  assure  and  preserve  the  product’s  identity,  strength,  quality,  and  purity.  For  some  NDAs,  the  FDA  may  convene  an  advisory
committee to seek insights and recommendations on issues relevant to approval of the application. Although the FDA is not bound by the recommendation
of an advisory committee, the agency considers such recommendations carefully when making decisions. Before approving a new drug product, the FDA
also  requires  that  the  facilities  at  which  the  product  will  be  manufactured  or  advanced  through  the  supply  chain  be  in  compliance  with  current  good
manufacturing practices (“GMP”) requirements and regulations governing, among other things, the manufacture, shipment, and storage of the product. The
FDA  also  can  conduct  audits  to  determine  if  the  clinical  trials  were  conducted  in  compliance  with  GCP.  After  review  of  an  NDA,  the  FDA  may  grant
marketing approval, request additional information, or issue a complete response letter (“CRL”) communicating the reasons for the agency’s decision not to
approve  the  application.  The  CRL  may  request  additional  information,  including  additional  preclinical  or  clinical  data,  for  the  FDA  to  reconsider  the
application. An NDA may be resubmitted with the deficiencies addressed, but resubmission does not guarantee approval. Data from clinical trials are not
always conclusive, and the FDA’s interpretation of data may differ from the sponsor’s. Obtaining approval can take years, requires substantial resources
and depends on a number of factors, including the severity of the targeted disease or condition, the availability of alternative treatments, and the risks and
benefits  demonstrated  in  clinical  trials.  Additionally,  as  a  condition  of  approval,  the  FDA  may  impose  restrictions  that  could  affect  the  commercial
prospects of a product and increase our costs, such as a Risk Evaluation and Mitigation Strategy (“REMS”), and/or post-approval commitments to conduct
additional clinical trials or non-clinical studies or to conduct surveillance programs to monitor the product’s effects. Under the Pediatric Research Equity
Act (“PREA”), certain applications for approval must also include an assessment, generally based on clinical study data, of the safety and effectiveness of
the subject product in relevant pediatric populations, unless a waiver or deferral is granted.

Moreover,  once  a  product  is  approved,  information  about  its  safety  or  effectiveness  from  broader  clinical  use  may  limit  or  prevent  successful
commercialization  because  of  regulatory  action,  market  forces  or  for  other  reasons.  Post-approval  modifications  to  a  drug  product,  such  as  changes  in
indications,  labeling  or  manufacturing  processes  or  facilities,  may  require  development  and  submission  of  additional  information  or  data  in  a  new  or
supplemental NDA, which would also require prior FDA approval.

Competition. The  Drug  Price  Competition  and  Patent  Term  Restoration  Act  of  1984  (the  “Hatch-Waxman  Act”)  establishes  two  abbreviated  approval
pathways for product candidates that are in some way follow-on versions of already approved branded NDA products: (i) generic versions of the approved
reference listed drug (“RLD”), which may be approved under an abbreviated new drug application (“ANDA”) by showing that the generic product is the
“same as” the approved product in key respects; and (ii) a product that is similar but not identical to a listed drug, which may be approved under a 505(b)
(2) NDA, in which the sponsor relies to some degree on information from investigations that were not conducted by or for the applicant and for which the
applicant has not obtained a right of reference, and submits its own product-specific data to support the differences between the product and the listed drug.

The sponsor of an ANDA or 505(b)(2) application seeking to rely on an approved product as the RLD or listed drug must make one of several certifications
regarding  each  patent  for  the  RLD  that  is  listed  in  the  FDA  publication,  Approved  Drug  Products  with  Therapeutic  Equivalence  Evaluations,  which  is
referred  to  as  the  Orange  Book.  A  “Paragraph  I”  certification  is  the  sponsor’s  statement  that  patent  information  has  not  been  filed  for  the  RLD.  A
“Paragraph II” certification is the sponsor’s statement that the RLD’s patents have expired. A “Paragraph III” certification is the sponsor’s statement that it
will wait for the patent to expire before obtaining approval for its product. A “Paragraph IV” certification is an assertion that the patent does not block
approval of the later product, either because the patent is invalid or unenforceable or because the patent, even if valid, is not infringed by the new product.
Once the FDA accepts for filing an ANDA or 505(b)(2) application containing a Paragraph IV certification, the applicant must within 20 days provide
notice  to  the  RLD  or  listed  drug  NDA  holder  and  patent  owner  that  the  application  has  been  submitted  and  provide  the  factual  and  legal  basis  for  the
applicant’s assertion that the patent is invalid or not infringed. If the NDA holder or patent owner files suit against the ANDA or 505(b)(2) applicant for
patent infringement within 45 days of receiving the Paragraph IV notice, the FDA is prohibited from approving the ANDA or 505(b)(2) application for a
period of 30 months or the resolution of the underlying suit, whichever is earlier.

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Exclusivity  and  Patent  Protection.  In  the  United  States  and  elsewhere,  certain  regulatory  exclusivities  and  patent  rights  can  provide  an  approved  drug
product with protection from certain competitors’ products for a period of time and within a certain scope. In the United States, those protections include
regulatory exclusivity under the Hatch-Waxman Act, which provides periods of exclusivity for a branded drug product that would serve as an RLD for a
generic drug applicant filing and an ANDA under section 505(j) of the FD&C Act or as a listed drug for an applicant filing an NDA under section 505(b)
(2) of the FD&C Act. If such a product is a “new chemical entity” (“NCE”) generally meaning that the active moiety has never before been approved in
any drug, there is a period of five years from the product’s approval during which the FDA may not accept for filing any ANDA or 505(b)(2) application
for a drug with the same active moiety. An ANDA or 505(b)(2) application may be submitted after four years, however, if the sponsor of the application
makes a Paragraph IV certification (as described above). Such a product that is not an NCE may qualify for a three-year period of exclusivity if its NDA
contains new clinical data (other than bioavailability studies), derived from studies conducted by or for the sponsor, that were necessary for approval. In
this instance, the three-year exclusivity period does not preclude filing or review of an ANDA or 505(b)(2) application; rather, the FDA is precluded from
granting final approval to the ANDA or 505(b)(2) application until three years after approval of the RLD. This three-year exclusivity applies only to the
conditions of approval that required submission of the clinical data.

The Hatch-Waxman Act also provides for the restoration of a portion of the patent term lost during product development and FDA review of an NDA if
approval  of  the  application  is  the  first  permitted  commercial  marketing  of  a  drug  containing  the  active  ingredient.  The  patent  term  restoration  period  is
generally one-half the time between the effective date of the IND or the date of patent grant (whichever is later) and the date of submission of the NDA,
plus the time between the date of submission of the NDA and the date of FDA approval of the product. The maximum period of restoration is five years,
and the patent cannot be extended to more than 14 years from the date of FDA approval of the product. Only one patent claiming each approved product is
eligible for restoration and the patent holder must apply for restoration within 60 days of approval. The USPTO, in consultation with the FDA, reviews and
approves the application for patent term restoration.

Emergency Use Authorization (“EUA”). The Secretary of Health and Human Services may authorize unapproved medical products to be marketed in the
context of an actual or potential emergency that has been designated by the U.S. government. The COVID-19 pandemic has been designated as such a
national emergency. After an emergency has been announced, the Secretary of Health and Human Services may authorize the issuance of and the FDA
Commissioner  may  issue  EUAs  for  the  use  of  specific  products  based  on  criteria  established  by  the  FDCA,  including  that  the  product  at  issue  may  be
effective  in  diagnosing,  treating,  or  preventing  serious  or  life-threatening  diseases  when  there  are  no  adequate,  approved,  and  available  alternatives.
Although the criteria of an EUA differ from the criteria for approval of an NDA, EUAs nevertheless require the development and submission of data to
satisfy the relevant FDA standards, and a number of ongoing compliance obligations. The FDA expects EUA holders to work toward submission of full
applications,  such  as  an  NDA,  as  soon  as  possible. An  EUA  is  also  subject  to  additional  conditions  and  restrictions  and  is  product-specific.  An  EUA
terminates  when  the  emergency  determination  underlying  the  EUA  terminates.  An  EUA  is  not  a  long-term  alternative  to  obtaining  FDA  approval,
licensure, or clearance for a product. The FDA may revoke an EUA for a variety of reasons, including where it is determined that the underlying health
emergency no longer exists or warrants such authorization, so it is not possible to predict how long an EUA may remain in place.

Post-Approval Regulation

Once approved, drug products are subject to continuing extensive regulation by the FDA, including ongoing monitoring for safety information, maintaining
appropriate registrations and licenses, and hosting periodic inspections. If ongoing regulatory requirements are not met, or if safety problems occur after a
product reaches market, the FDA may take actions to change the conditions under which the product is marketed, such as requiring labeling modifications,
restricting  distribution,  or  even  withdrawing  approval.  In  addition  to  FDA  regulation,  our  business  is  also  subject  to  extensive  federal,  state,  local  and
foreign regulation.

Good Manufacturing Practices. Companies engaged in manufacturing drug products or their components must comply with applicable GMP requirements,
which include requirements regarding organization and training of personnel, building and facilities, equipment, control of components and drug product
containers,  closures,  production  and  process  controls,  packaging  and  labeling  controls,  holding  and  distribution,  laboratory  controls  and  records  and
reports. The FDA inspects equipment, facilities and manufacturing processes before approval and conducts periodic re-inspections after approval. If, after
receiving

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approval,  a  company  makes  a  material  change  in  manufacturing  equipment,  location,  or  process  (all  of  which  are,  to  some  degree,  incorporated  in  the
NDA), additional regulatory review and approval may be required. Failure to comply with applicable GMP requirements or the conditions of the product’s
approval may lead the FDA to take enforcement actions, such as issuing a warning letter, or to seek sanctions, including fines, civil penalties, injunctions,
suspension  of  manufacturing  operations,  imposition  of  operating  restrictions,  withdrawal  of  FDA  approval,  seizure  or  recall  of  products,  and  criminal
prosecution. Although we periodically monitor FDA compliance of the third parties on which we rely for manufacturing our product candidates, we cannot
be certain that our present or future third-party manufacturers will consistently comply with GMP or other applicable FDA regulatory requirements.

Sales and Marketing. Once a product is approved, the advertising, promotion and marketing of the product will be subject to close regulation, including
with  regard  to  promotion  to  healthcare  practitioners,  direct-to-consumer  advertising,  communications  regarding  unapproved  uses,  industry-sponsored
scientific  and  educational  activities  and  promotional  activities  involving  the  internet.  In  addition  to  FDA  restrictions  on  marketing  of  pharmaceutical
products, state and federal fraud and abuse laws have been applied to restrict certain marketing practices in the pharmaceutical industry. Failure to comply
with applicable requirements in this area may subject a company to adverse publicity, investigations and enforcement action by the FDA, the Department of
Justice, the Office of the Inspector General of the Department of Health and Human Services, and/or state authorities. This could subject a company to a
range of penalties that could have a significant commercial impact, including civil and criminal fines and agreements that materially restrict the manner in
which a company promotes or distributes drug products.

New Legislation. New legislation is passed periodically in Congress, or at the state level, that could significantly change the statutory provisions governing
the  approval,  manufacturing  and  marketing  of  products  regulated  by  the  FDA.  Further,  the  FDA  revises  its  regulations  and  guidance  in  light  of  new
legislation  in  ways  that  may  affect  our  business  or  product  candidates.  It  is  impossible  to  predict  whether  other  changes  to  legislation,  regulation,  or
guidance will be enacted, or what the impact of such changes, if any, may be.

Other  Requirements.  Companies  that  manufacture  or  distribute  drug  products  pursuant  to  approved  NDAs  must  meet  numerous  other  regulatory
requirements, including adverse event reporting, submission of periodic reports, and record-keeping obligations.

Fraud and Abuse Laws. At such time as we market, sell and distribute any products for which we obtain marketing approval, it is possible that our business
activities could be subject to scrutiny and enforcement under one or more federal or state health care fraud and abuse laws and regulations, which may
constrain the business or financial arrangements and relationships through which we market, sell and distribute any products for which we obtain marketing
approval.  These  restrictions  under  applicable  federal  and  state  health  care  fraud  and  abuse  laws  and  regulations  that  may  affect  our  ability  to  operate
include:

•

•

The  U.S.  federal  Anti-Kickback  Law,  which  prohibits,  among  other  things,  knowingly  or  willingly  offering,  paying,  soliciting  or  receiving
remuneration, directly or indirectly, in cash or in kind, to induce or reward the purchasing, leasing, ordering or arranging for or recommending the
purchase, lease or order of any health care items or service for which payment may be made, in whole or in part, by federal healthcare programs
such as Medicare and Medicaid. This statute has been interpreted to apply to arrangements between pharmaceutical companies on one hand and
prescribers,  purchasers  and  formulary  managers  on  the  other.  Liability  may  be  established  under  the  U.S.  federal  Anti-Kickback  Law  without
proving  actual  knowledge  of  the  statute  or  specific  intent  to  violate  it.  In  addition,  the  government  may  assert  that  a  claim  including  items  or
services resulting from a violation of the U.S. federal Anti-Kickback Law constitutes a false or fraudulent claim for purposes of the U.S. federal
civil False Claims Act. Although there are a number of statutory exemptions and regulatory safe harbors to the U.S. federal Anti-Kickback Law
protecting  certain  common  business  arrangements  and  activities  from  prosecution  or  regulatory  sanctions,  the  exemptions  and  safe  harbors  are
drawn narrowly, and practices that do not fit squarely within an exemption or safe harbor, or for which no exception or safe harbor is available,
may be subject to scrutiny.

The U.S. federal civil False Claims Act, which prohibits, among other things, individuals or entities from knowingly presenting, or causing to be
presented, a false or fraudulent claim for payment of government funds or knowingly making, using or causing to be made or used, a false record
or  statement  material  to  an  obligation  to  pay  money  to  the  government  or  knowingly  concealing  or  knowingly  and  improperly  avoiding,
decreasing or concealing an obligation to

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pay money to the federal government. Actions under the False Claims Act may be brought by the United States Attorney General or as a qui tam
action  by  a  private  individual  (a  whistleblower)  in  the  name  of  the  government  and  the  individual,  and  the  whistleblower  may  share  in  any
monetary  recovery.  Many  pharmaceutical  and  other  healthcare  companies  have  been  investigated  and  have  reached  substantial  financial
settlements  with  the  federal  government  under  the  civil  False  Claims  Act  for  a  variety  of  alleged  improper  marketing  activities,  including:
providing  free  product  to  customers  with  the  expectation  that  the  customers  would  bill  federal  programs  for  the  product;  providing  sham
consulting  fees,  grants,  free  travel  and  other  benefits  to  physicians  to  induce  them  to  prescribe  the  company’s  products;  and  inflating  prices
reported  to  private  price  publication  services,  which  are  used  to  set  drug  payment  rates  under  government  healthcare  programs.  In  addition,  in
recent years the government has pursued civil False Claims Act cases against a number of pharmaceutical companies for causing false claims to be
submitted as a result of the marketing of their products for unapproved, and thus non-reimbursable, uses. Because of the threat of treble damages
and  mandatory  penalties  per  false  or  fraudulent  claim  or  statement,  healthcare  and  pharmaceutical  companies  often  resolve  allegations  without
admissions of liability for significant and material amounts. Pharmaceutical and other healthcare companies also are subject to other federal false
claim laws, including, among others, federal criminal healthcare fraud and false statement statutes that extend to non-government health benefit
programs.

•

The  fraud  provisions  of  the  Health  Insurance  Portability  and  Accountability  Act  of  1996  (“HIPAA”),  which  impose  criminal  liability  for
knowingly  and  willfully  executing  a  scheme  to  defraud  any  healthcare  benefit  program,  including  private  third-party  payors,  and  prohibit
knowingly and willfully falsifying, concealing or covering up a material fact or making any materially false, fictitious or fraudulent statement or
representation,  or  making  or  using  any  false  writing  or  document  knowing  the  same  to  contain  any  materially  false  fictitious  or  fraudulent
statement or entry, in connection with the delivery of or payment for healthcare benefits, items or services.

• Analogous  state  and  local  laws  and  regulations,  such  as  state  anti-kickback  and  false  claims  laws,  which  may  apply  to  sales  or  marketing
arrangements and claims involving healthcare items or services reimbursed by non-governmental third-party payors, including private insurers;
state and foreign laws that require pharmaceutical companies to comply with the pharmaceutical industry’s voluntary compliance guidelines and
the  relevant  compliance  guidance  promulgated  by  the  federal  government  or  otherwise  restrict  payments  that  may  be  made  to  healthcare
providers;  state  laws  that  restrict  the  ability  of  manufacturers  to  offer  co-pay  support  to  patients  for  certain  prescription  drugs;  and  state  and
foreign laws that require drug manufacturers to report information related to clinical trials, or information related to payments and other transfers
of  value  to  physicians  and  other  healthcare  providers  or  marketing  expenditures;  state  laws  and  local  ordinances  that  require  identification  or
licensing of sales representatives.

•

•

The U.S. federal Physician Payment Sunshine Act, being implemented as the Open Payments Program, which requires manufacturers of drugs,
devices, biologics, and medical supplies for which payment is available under Medicare, Medicaid or the Children’s Health Insurance Program
(with certain exceptions) to report annually to the Centers for Medicare and Medicaid Services (“CMS”) information related to direct or indirect
payments and other transfers of value to physicians and teaching hospitals, as well as ownership and investment interests held in the company by
physicians and their immediate family members. As of 2022, applicable manufacturers are also required to report information regarding payments
and transfers of value provided (starting in 2021) to physician assistants, nurse practitioners, clinical nurse specialists, certified nurse anesthetists,
and certified nurse-midwives.

The federal Foreign Corrupt Practices Act of 1997 and other similar anti-bribery laws in other jurisdictions generally prohibit companies and their
intermediaries  from  providing  money  or  anything  of  value  to  officials  of  foreign  governments,  foreign  political  parties  or  international
organizations  with  the  intent  to  obtain  or  retain  business  or  seek  a  business  advantage.  Recently,  there  has  been  a  substantial  increase  in  anti-
bribery law enforcement activity by United States regulators, with more frequent and aggressive investigations and enforcement proceedings by
both the Department of Justice and the United States Securities and Exchange Commission (the “SEC”). Violations of United States or foreign
laws  or  regulations  could  result  in  the  imposition  of  substantial  fines,  interruptions  of  business,  loss  of  supplier,  vendor  or  other  third-party
relationships, termination of necessary licenses and permits and other legal or equitable sanctions. Other internal or government investigations or
legal or regulatory proceedings, including lawsuits brought by private litigants, may also follow as a consequence.

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Violations  of  any  of  the  laws  described  above  or  any  other  governmental  regulations  are  punishable  by  significant  civil,  criminal  and  administrative
penalties, damages, fines and exclusion from government-funded healthcare programs, such as Medicare and Medicaid. Although  compliance  programs
can  mitigate  the  risk  of  investigation  and  prosecution  for  violations  of  these  laws,  the  risks  cannot  be  entirely  eliminated.  Moreover,  achieving  and
sustaining compliance with applicable federal and state privacy, security and fraud laws may prove costly.

Privacy Laws. We are also subject to federal, state and foreign laws and regulations governing data privacy and security of health information, and the
collection, use and disclosure, and protection of health-related and other personal information. The legislative and regulatory landscape for privacy and data
protection continues to evolve, and there has been an increasing focus on privacy and data protection issues that may affect our business, including recently
enacted  laws  in  all  jurisdictions  where  we  operate.    Numerous  federal  and  state  laws,  including  state  security  breach  notification  laws,  state  health
information privacy laws, state genetic privacy laws, and federal and state consumer protection and privacy laws, (including, for example, Section 5 of the
Federal Trade Commission Act (“FTC Act”), and the California Consumer Privacy Act (“CCPA”)) govern the collection, use and disclosure of personal
information. These laws may differ from each other in significant ways, thus complicating compliance efforts. Federal regulators, state attorneys general,
and  plaintiffs’  attorneys  have  been  and  will  likely  continue  to  be  active  in  this  space.  Activities  outside  of  the  U.S.  implicate  local  and  national  data
protection standards, impose additional compliance requirements and generate additional risks of enforcement for non-compliance. The European Union’s
General  Data  Protection  Regulation  (“GDPR”)  and  other  data  protection,  privacy  and  similar  national,  state/provincial  and  local  laws  may  restrict  the
access, use and disclosure of patient health information abroad. Compliance efforts will likely be an increasing and substantial cost in the future.

Failure to comply with such laws and regulations could result in government enforcement actions and create liability for us (including the imposition of
significant penalties), private litigation and/or adverse publicity that could negatively affect our business. In addition, if we successfully commercialize our
product candidates, we may obtain patient health information from healthcare providers who prescribe our products and research institutions we collaborate
with, and they are subject to privacy and security requirements under HIPAA.  Although we are not directly subject to HIPAA other than potentially with
respect to providing certain employee benefits, we could potentially be subject to criminal penalties if we, or our affiliates or our agents knowingly receive
individually identifiable health information maintained by a HIPAA-covered entity in a manner that is not authorized or permitted by HIPAA.

The  Federal  Trade  Commission  (“FTC”)  also  sets  expectations  for  failing  to  take  appropriate  steps  to  keep  consumers’  personal  information  secure,  or
failing to provide a level of security commensurate to promises made to individual about the security of their personal information (such as in a privacy
notice)  may  constitute  unfair  or  deceptive  acts  or  practices  in  violation  of  Section  5(a)  of  the  FTC  Act.  The  FTC  expects  a  company’s  data  security
measures to be reasonable and appropriate in light of the sensitivity and volume of consumer information it holds, the size and complexity of its business,
and the cost of available tools to improve security and reduce vulnerabilities. Individually identifiable health information is considered sensitive data that
merits stronger safeguards. With respect to privacy, the FTC also sets expectations for failing to honor the privacy promises made to individuals about how
the company handles consumers’ personal information; such failure may also constitute unfair or deceptive acts or practices in violation of the FTC Act.
Enforcement by the FTC under the FTC Act can result in civil penalties or enforcement actions.

In California, the CCPA establishes certain requirements for data use and sharing transparency and provides California residents certain rights concerning
the use, disclosure, and retention of their personal information. The CCPA and its implementing regulations have already been amended multiple times
since  their  enactment.  In  November  2020,  California  voters  approved  the  California  Privacy  Rights  Act  (“CPRA”)  ballot  initiative  which  introduced
significant  amendments  to  the  CCPA  and  established  and  funded  a  dedicated  California  privacy  regulator,  the  California  Privacy  Protection  Agency
(“CPPA”). The amendments introduced by the CPRA went into effect on January 1, 2023, and new implementing regulations are expected to be introduced
by the CPPA. Failure to comply with the CCPA may result in, among other things, significant civil penalties and injunctive relief, or statutory or actual
damages. In addition, California residents have the right to bring a private right of action in connection with certain types of incidents. These claims may
result in significant liability and damages. Similarly, there are a number of legislative proposals in the United States, at both the federal and state level, that
could impose new obligations or limitations in areas affecting our business. For example, other states, including Virginia, Colorado, Utah, and Connecticut
have enacted privacy laws similar to the CCPA that impose new obligations or limitations in

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areas affecting our business and we continue to assess the impact of these state legislations on our business as additional information and guidance becomes
available. These laws and regulations are evolving and subject to interpretation, and may impose limitations on our activities or otherwise adversely affect
our business.

Activities outside of the U.S. implicate local and national data protection standards, impose additional compliance requirements and generate additional
risks of enforcement for non-compliance. The European Union’s GDPR, which imposes fines of up to EUR 20 million or 4% of the annual global revenue
of a noncompliant company, whichever is greater, and other data protection, privacy and similar national, state/provincial and local laws may also restrict
the access, use and disclosure of patient health information abroad. We may be required to expend significant capital and other resources to ensure ongoing
compliance  with  applicable  privacy  and  data  security  laws,  to  protect  against  security  breaches  and  hackers,  or  to  alleviate  problems  caused  by  such
breaches.  Compliance  with  these  laws  is  difficult,  constantly  evolving,  time  consuming,  and  requires  a  flexible  privacy  framework  and  substantial
resources.  Compliance  efforts  will  likely  be  an  increasing  and  substantial  cost  in  the  future.  There  are  also  a  number  of  legislative  proposals  in  the
European Union, the United States, at both the federal and state level, as well as other jurisdictions that could impose new obligations or limitations in
areas affecting our business. In addition, some countries are considering or have passed legislation implementing data protection requirements or requiring
local storage and processing of data or similar requirements that could increase the cost and complexity of delivering our services and research activities.
These  laws  and  regulations,  as  well  as  any  associated  claims,  inquiries,  or  investigations  or  any  other  government  actions  may  lead  to  unfavorable
outcomes including increased compliance costs, delays or impediments in the development of new products, negative publicity, increased operating costs,
diversion of management time and attention, and remedies that harm our business, including fines or demands or orders that we modify or cease existing
business practices.

With regard to transfer of personal data, the GDPR restricts the ability of companies to transfer personal data from the European Economic Area to the
United States and other countries, which may adversely affect our ability to transfer personal data or otherwise may cause us to incur significant costs to
come into compliance with applicable data transfer impact assessments and implementation of legal data transfer mechanisms. One mechanism previously
relied  upon  by  companies  for  such  transfers  was  the  EU-U.S.  Privacy  Shield  Framework  (the  “Privacy  Shield”).  However,  in  July  2020,  the  European
Court of Justice ruled the Privacy Shield to be an invalid data transfer mechanism and confirmed that the European Commission’s Standard Contractual
Clauses (the “Model Clauses”) remain valid and in June 2021, the European Commission published updated versions of the Model Clauses, which must be
incorporated into new and existing agreements within prescribed timeframes in order to continue to lawfully transfer personal data outside of the European
Union.  As  a  result,  companies  may  no  longer  rely  on  the  Privacy  Shield  as  a  basis  on  which  to  transfer  personal  data  from  the  European  Union  to  the
United States. U.S.-based companies are permitted to rely on other authorized means and procedures to transfer personal data provided by the GDPR. The
Model Clauses may also come under increased scrutiny as a result of the European Court of Justice’s judgement in July 2020, though they remain the most
common  authorized  procedure  to  transfer  personal  data  out  of  the  European  Union.  On  December,  13  2022,  the  European  Commission  adopted  a  draft
adequacy decision for the EU-U.S. Data Privacy Framework. The draft decision concludes that the United States ensures an adequate level of protection for
personal data transferred from the European Union to the United States. The draft adequacy decision text will also have to be approved by a committee
composed of representatives of the European Union Member States and the European Parliament can exercise its right of scrutiny. After this process, the
European Commission is then expected to adopt the final adequacy decision, which will allow data to flow freely from the European Union to the United
States. After one year from the notification date of the adequacy decision to the Member States and subsequently at least every four years, the European
Commission will carry out a new evaluation and could conclude that an adequate level of protection is no longer ensured and decide to suspend, amend or
repeal the adequacy decision, or limit its scope.

Coverage and Reimbursement

Significant uncertainty exists as to the coverage and reimbursement status of any product candidates for which we may obtain regulatory approval. The
regulations that govern marketing approvals, pricing and reimbursement for new drug products vary widely from country to country. Current  and  future
legislation may significantly change the approval requirements in ways that could involve additional costs and cause delays in obtaining approvals. Some
countries require approval of the sale price of a drug before it can be marketed. In many countries, the pricing review period begins after marketing or
product licensing approval is granted. In  some  foreign  markets,  prescription  pharmaceutical  pricing  remains  subject  to  continuing  governmental  control
even after initial approval is granted. As a result, we might obtain marketing approval for a product in a particular country, but then be subject to price
regulations that delay our commercial launch of the product, possibly for lengthy time

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periods,  which  could  negatively  impact  the  revenues  we  are  able  to  generate  from  the  sale  of  the  product  in  that  particular  country.  Adverse  pricing
limitations may hinder our ability to recoup our investment in one or more product candidates even if our product candidates obtain marketing approval.

Our  ability  to  commercialize  any  products  successfully  also  will  depend  in  part  on  the  extent  to  which  coverage  and  adequate  reimbursement  for  these
products and related treatments will be available in a timely manner from third-party payors, including government healthcare programs such as Medicare
and  Medicaid,  commercial  health  insurers  and  managed  care  organizations.  Government  authorities  and  other  third-party  payors,  such  as  private  health
insurers and health maintenance organizations, determine which medications they will cover and establish reimbursement levels. Third-party payors may
limit coverage to specific products on an approved list, or formulary, which may not include all of the FDA-approved products for a particular indication.
The process for determining whether a payor will provide coverage for a product may be separate from the process for setting the price or reimbursement
rate that the payor will pay for the product once coverage is approved.

A primary trend in the United States healthcare industry and elsewhere is cost containment. Government healthcare programs and other third-party payors
are increasingly challenging the prices charged for medical products and services and examining the medical necessity and cost-effectiveness of medical
products and services, in addition to their safety and efficacy, and have attempted to control costs by limiting coverage and the amount of reimbursement
for particular medications. Increasingly, third-party payors are requiring that drug companies provide them with predetermined discounts from list prices
and are challenging the prices charged for medical products. We cannot be sure that coverage and reimbursement will be available promptly or at all for
any  product  that  we  commercialize  and,  if  reimbursement  is  available,  what  the  level  of  reimbursement  will  be.  Moreover,  eligibility  for  coverage  and
reimbursement  does  not  imply  that  any  drug  will  be  paid  for  in  all  cases.  Limited  coverage  may  impact  the  demand  for,  or  the  price  of,  any  product
candidate for which we obtain marketing approval. If coverage and reimbursement are not available or reimbursement is available only to limited levels,
we may not successfully commercialize any product candidate for which we obtain marketing approval.

Obtaining  coverage  and  adequate  reimbursement  is  a  time-consuming  and  costly  process.  There  may  be  significant  delays  in  obtaining  coverage  and
reimbursement for newly approved drugs, and coverage may be more limited than the purposes for which the drug is approved by the FDA or comparable
foreign regulatory authorities. Moreover, eligibility for coverage and reimbursement does not imply that a drug will be paid for in all cases or at a rate that
covers our costs, including research, development, manufacture, sale and distribution. Interim reimbursement levels for new drugs, if applicable, may also
not be sufficient to cover our costs and may only be temporary. Reimbursement rates may vary according to the use of the drug and the clinical setting in
which it is used, may be based on reimbursement levels already set for lower cost drugs and may be incorporated into existing payments for other services.
Net prices for drugs may be reduced by mandatory discounts or rebates required by government healthcare programs or private payors and by any future
relaxation of laws that presently restrict imports of drugs from countries where they may be sold at lower prices than in the United States. Limited coverage
may impact the demand for, or the price of, any product candidate for which we obtain marketing approval. Third-party payors also may seek additional
clinical evidence, including expensive pharmacoeconomic studies, beyond the data required to obtain marketing approval, demonstrating clinical benefits
and value in specific patient populations, before covering our products for those patients. If reimbursement is available only for limited indications, we may
not be able to successfully commercialize any product candidate for which we obtain marketing approval. Our inability to promptly obtain coverage and
profitable reimbursement rates from both government-funded and private payors for any approved products that we develop could have a material adverse
effect on our operating results, our ability to raise capital needed to commercialize products and our overall financial condition.

If we successfully commercialize any of our products, we may participate in the Medicaid Drug Rebate Program. Participation is required for federal funds
to be available for our products under Medicaid and Medicare Part B. Under the Medicaid Drug Rebate Program, we would be required to pay a rebate to
each state Medicaid program for our covered outpatient drugs that are dispensed to Medicaid beneficiaries and paid for by a state Medicaid program as a
condition of having federal funds being made available to the states for our drugs under Medicaid and under Part B of the Medicare program. Rebates owed
by manufacturers under the Medicaid Drug Rebate Program are currently capped at 100 percent of average manufacturer price, but, effective January 1,
2024, this cap will be lifted, which could adversely affect our rebate liability.

Federal law requires that any company that participates in the Medicaid Drug Rebate Program also participate in the Public Health Service’s 340B drug
pricing program in order for federal funds to be available for the manufacturer’s drugs under

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Medicaid and Medicare Part B. The 340B program requires participating manufacturers to agree to charge statutorily-defined covered entities no more than
the 340B “ceiling price” for the manufacturer’s covered outpatient drugs. These 340B covered entities include a variety of community health clinics and
other  entities  that  receive  health  services  grants  from  the  Public  Health  Service,  as  well  as  hospitals  that  serve  a  disproportionate  share  of  low-income
patients.

Medicare is a federal program that is administered by the federal government that covers individuals age 65 and over or that are disabled as well as those
with certain health conditions. Medicare Part B generally covers drugs that must be administered by physicians or other health care practitioners; among
others.  Medicare  Part  B  generally  pays  for  such  drugs  under  a  payment  methodology  based  on  the  average  sales  price  of  the  drugs.  Manufacturers  are
required  to  report  average  sales  price  information  to  CMS  on  a  quarterly  basis.  The  manufacturer-submitted  information  is  used  by  CMS  to  calculate
Medicare payment rates. Effective January 1, 2023, manufacturers will be obligated to pay refunds to Medicare for single source drugs or biologicals, or
biosimilar biological products, reimbursed under Medicare Part B and packaged in single-dose containers or single-use packages, for units of discarded
drug reimbursed by Medicare Part B in excess of 10 percent of total allowed charges under Medicare Part B for that drug. Manufacturers that fail to pay
refunds could be subject to civil monetary penalties of 125 percent of the refund amount. Further, starting in January 2023, the Inflation Reduction Act of
2022 (“IRA”) establishes a Medicare Part B inflation rebate scheme, under which, generally speaking, manufacturers will owe rebates if the average sales
price of a Part B drug increases faster than the pace of inflation. Failure to timely pay a Part B inflation rebate is subject to a civil monetary penalty.

Medicare  Part  D  generally  provides  coverage  to  enrolled  Medicare  patients  for  self-administered  drugs  (i.e.,  drugs  that  are  not  administered  by  a
physician). Medicare Part D is administered by private prescription drug plans approved by the U.S. government and, subject to detailed program rules and
government oversight, each drug plan establishes its own Medicare Part D formulary for prescription drug coverage and pricing, which the drug plan may
modify from time to time. The prescription drug plans negotiate pricing with manufacturers and pharmacies, and may condition formulary placement on
the availability of manufacturer discounts. In addition, under the coverage gap discount program, manufacturers are required to provide a 70% discount on
brand  name  prescription  drugs  utilized  by  Medicare  Part  D  beneficiaries  when  those  beneficiaries  are  in  the  coverage  gap  phase  of  the  Part  D  benefit
design. Civil monetary penalties could be due if a manufacturer were to fail to offer discounts under the coverage gap discount program. The IRA sunsets
the coverage gap discount program starting in 2025 and replaces it with a new manufacturer discount program. Failure to pay a discount under this new
program will be subject to a civil monetary penalty. In addition, starting in October 2022, the IRA established a Medicare Part D inflation rebate scheme,
under which, generally speaking, manufacturers will owe additional rebates if the average manufacturer price of a Part D drug increases faster than the pace
of inflation. Failure to timely pay a Part D inflation rebate is subject to a civil monetary penalty.

The IRA also creates a drug price negotiation program under which the prices for Medicare units of certain high Medicare spend drugs and biologicals
without generic or biosimilar competition will be capped by reference to, among other things, a specified non-federal average manufacturer price starting in
2026. Failure to comply with requirements under the drug price negotiation program is subject to an excise tax and/or a civil monetary penalty. This or any
other legislative change could impact the market conditions for our product candidates.

In addition, in order to be eligible to have its products paid for with federal funds under the Medicaid and Medicare Part B programs and purchased by the
Department of Veterans Affairs (the “VA”), Department of Defense (“DoD”), Public Health Service, and Coast Guard (the “Big Four agencies”) and certain
federal  grantees,  a  manufacturer  also  must  participate  in  the  VA  Federal  Supply  Schedule  (“FSS”)  pricing  program,  established  by  Section  603  of  the
Veterans Health Care Act of 1992 (the “VHCA”). Under this program, the manufacturer is obligated to make its covered drugs (innovator multiple source
drugs, single source drugs, and biologics) available for procurement on an FSS contract and charge a price to the Big Four agencies that is no higher than
the Federal Ceiling Price (“FCP”), which is a price calculated pursuant to a statutory formula. The FCP is derived from a calculated price point called the
“non-federal average manufacturer price” (“Non-FAMP”), which we will be required to calculate and report to the VA on a quarterly and annual basis.
Moreover, pursuant to Defense Health Agency (“DHA”) regulations, manufacturers must provide rebates on utilization of their innovator and single source
products that are dispensed to TRICARE beneficiaries by TRICARE network retail pharmacies. The formula for determining the rebate is established in the
regulations and is based on the difference between the annual non-federal average manufacturer price and the Federal Ceiling Price, each required to be
calculated  by  us  under  the  VHCA.  The  requirements  under  the  Medicaid  Drug  Rebate  Program,  340B  program,  FSS,  and  TRICARE  programs  could
reduce the revenue we may generate from any products that are commercialized in the future and could adversely affect our business and operating results.

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United States Healthcare Reform

The United States and many foreign jurisdictions have enacted or proposed legislative and regulatory changes affecting the healthcare system that could
prevent  or  delay  marketing  approval  of  our  product  candidates,  restrict  or  regulate  post-approval  activities  and  affect  our  ability  to  profitably  sell  any
product  candidate  for  which  we  obtain  marketing  approval.  The  United  States  government,  state  legislatures  and  foreign  governments  also  have  shown
significant  interest  in  implementing  cost-containment  programs  to  limit  the  growth  of  government-paid  healthcare  costs,  including  price  controls,
restrictions on reimbursement and requirements for substitution of generic products for branded prescription drugs.

In recent years, Congress has considered reductions in Medicare reimbursement levels for drugs administered by physicians. The Centers for Medicare &
Medicaid Services (CMS), the agency that administers the Medicare and Medicaid programs, has authority to revise reimbursement rates and to implement
coverage restrictions. Cost reduction initiatives and changes in coverage implemented through legislation or regulation could decrease utilization of and
reimbursement for any approved products, which in turn would affect the price we can receive for those products. Any reduction in reimbursement from
Medicare and other government programs may result in a similar reduction in payment from commercial payers. The implementation of cost containment
measures or other healthcare reforms may prevent us from being able to generate revenue, attain profitability, or commercialize our products.

The Affordable Care Act, as amended (the “Affordable Care Act”), has substantially changed the way healthcare is financed by both governmental and
private insurers, and has significantly impacted the pharmaceutical industry. The Affordable Care Act was intended to broaden access to health insurance,
reduce  or  constrain  the  growth  of  healthcare  spending,  enhance  remedies  against  healthcare  fraud  and  abuse,  add  new  transparency  requirements  for
healthcare and health insurance industries, impose new taxes and fees on pharmaceutical and medical device manufacturers, and impose additional health
policy reforms.

Certain provisions of the Affordable Care Act have been subject to judicial challenges as well as efforts to modify them or to alter their interpretation and
implementation.  For  example,  the  Tax  Cuts  and  Jobs  Act,  enacted  on  December  22,  2017,  eliminated  the  tax-based  shared  responsibility  payment  for
individuals who fail to maintain minimum essential coverage under section 5000A of the Internal Revenue Code of 1986, commonly referred to as the
individual mandate, effective January 1, 2019. Additional legislative changes, regulatory changes, and judicial challenges related to the Affordable Care
Act remain possible. It is unclear how efforts to modify or invalidate the Affordable Care Act or its implementing regulations, or portions thereof, will
affect  our  business.  Any  such  changes  could  decrease  the  number  of  individuals  with  health  coverage.  It  is  possible  that  the  Affordable  Care  Act,  as
currently enacted or as it may be amended in the future, and other healthcare reform measures, including those that may be adopted in the future, could
have a material adverse effect on our industry generally and on our ability to successfully commercialize our product candidates, if approved.

In addition, other legislative changes have been proposed since the Affordable Care Act was enacted. For example, the Budget Control Act of 2011, among
other  things,  created  the  Joint  Select  Committee  on  Deficit  Reduction  to  recommend  to  Congress  proposals  for  spending  reductions.  The  Joint  Select
Committee did not achieve a targeted deficit reduction, which triggered the legislation’s automatic reductions. In concert with subsequent legislation, this
has resulted in aggregate reductions to Medicare payments to providers of, on average, 2% per fiscal year through 2031. Sequestration is currently set at
2%  and  will  increase  to  2.25%  for  the  first  half  of  fiscal  year  2030,  to  3%  for  the  second  half  of  fiscal  year  2030,  and  to  4%  for  the  remainder  of  the
sequestration period that lasts through the first six months of fiscal year 2031. As long as these cuts remain in effect, they could adversely impact payment
for any of our products that are reimbursed under Medicare, once commercialized.

We expect that the Affordable Care Act, as well as other healthcare reform measures that have been adopted and may be adopted in the future, may result in
more rigorous coverage criteria and new payment methodologies, and in additional downward pressure on coverage and payment and the price that we
receive  for  any  approved  product,  and  could  seriously  harm  our  future  revenues.  Any  reduction  in  reimbursement  from  Medicare,  Medicaid  or  other
government  programs  may  result  in  a  similar  reduction  in  payments  from  private  payors.  The  implementation  of  cost  containment  measures  or  other
healthcare reforms may prevent us from being able to generate revenue, attain profitability or commercialize our products.

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Foreign Regulation

In addition to regulations in the United States, we will be subject to a number of significant regulations in other jurisdictions regarding research, clinical
trials, approval, manufacturing, distribution, marketing and promotion, safety reporting, privacy and pricing and reimbursement. These requirements and
restrictions vary from country to country, but in many instances are similar to the United States requirements, and failure to comply with them could have
similar negative effects as noncompliance in the United States.

Corporate Information

Tekmira Pharmaceuticals Corporation (“Tekmira”) was incorporated pursuant to the British Columbia Business Corporations Act (“BCBCA”) on October
6,  2005,  and  commenced  active  business  on  April  30,  2007,  when  Tekmira  and  its  parent  company,  Inex  Pharmaceuticals  Corporation  (“Inex”),  were
reorganized under a statutory plan of arrangement (the “Plan of Arrangement”) completed under the provisions of the BCBCA. Pursuant to the Plan of
Arrangement, all of Inex’s business was transferred to Tekmira.

Protiva Biotherapeutics Inc. (“Protiva”) was acquired on May 30, 2008.

On March 4, 2015, we completed a business combination pursuant to which OnCore Biopharma, Inc. (“OnCore”) became our wholly-owned subsidiary of
Tekmira.

On July 31, 2015, we changed our corporate name from Tekmira Pharmaceuticals Corporation to Arbutus Biopharma Corporation and OnCore changed its
corporate name to Arbutus Biopharma, Inc.

On January 1, 2018, Protiva was amalgamated with Arbutus Biopharma Corporation.

We had one wholly-owned subsidiary as of December 31, 2022: Arbutus Biopharma, Inc.

Our principal executive office is located at 701 Veterans Circle, Warminster, Pennsylvania, USA, 18974, and our telephone number is (267) 469-0914.

Unless stated otherwise or the context otherwise requires, references herein to “Arbutus”, “we”, “us” and “our” refer to Arbutus Biopharma Corporation,
and, unless the context requires otherwise, the subsidiaries through which we conduct business.

Investor Information

We are a reporting issuer in Canada under the securities laws of each of the Provinces of Canada. Our common shares trade on the Nasdaq Global Select
Market under the symbol “ABUS”. We maintain a website at http://www.arbutusbio.com. The information on our website is not incorporated by reference
into this Annual Report on Form 10-K and should not be considered to be a part of this Annual Report on Form 10-K. Our website address is included in
this Annual Report on Form 10-K as an inactive technical reference only. Copies of this Annual Report on Form 10-K, and our other annual reports on
Form  10-K,  proxy  statements,  quarterly  reports  on  Form  10-Q,  current  reports  on  Form  8-K  and,  if  applicable,  amendments  to  those  reports  filed  or
furnished pursuant to Section 13(a) or 15(d) of the Securities Exchange Act of 1934, as amended, are available free of charge through our website under
“Investors – Financial Information – SEC Filings” as soon as reasonably practicable after we electronically file these materials with, or otherwise furnish
them to, the SEC. The SEC maintains an Internet site that contains reports, proxy and information statements and other information regarding issuers that
file electronically with the SEC at www.sec.gov.

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Item 1A. Risk Factors

Our business is subject to substantial risks and uncertainties. The occurrence of any of the following risks and uncertainties, either alone or taken together,
could materially and adversely affect our business, financial condition, results of operations or prospects. In these circumstances, the market price of our
common shares could decline and you may lose all or part of your investment. The risks and uncertainties described below are not the only ones we face.
Risks and uncertainties of general applicability and additional risks and uncertainties not currently known to us or that we currently deem to be immaterial
may also materially and adversely affect our business, financial condition, results of operations or prospects.

Risks Related to Our Business, Our Financial Results and Need for Additional Capital

We  are  in  the  early  stages  of  our  development,  and  there  is  a  limited  amount  of  information  about  us  upon  which  you  can  evaluate  our  product
candidates.

We  have  not  begun  to  market  or  generate  revenues  from  the  commercialization  of  any  of  our  product  candidates.  We  have  only  a  limited  history  upon
which you can evaluate our business and prospects as our product candidates are still at an early stage of development and thus we have limited experience
and have not yet demonstrated an ability to successfully overcome many of the risks and uncertainties frequently encountered by companies in new and
rapidly evolving fields, particularly in the biopharmaceutical area. For example, to execute our business plan, we will need to successfully:

execute research and development activities using technologies involved in the development of our product candidates;
build, maintain and protect a strong intellectual property portfolio;
gain regulatory approval and market acceptance for the commercialization of any product candidates we develop;
conduct sales and marketing activities if any of our product candidates are approved;
develop and maintain successful strategic relationships; and

•
•
•
•
•
• manage our spending and cash requirements as our expenses are expected to continue to increase due to research and pre-clinical work, clinical

trials, regulatory approvals, commercialization and maintaining our intellectual property portfolio.

If  we  are  unsuccessful  in  accomplishing  these  objectives,  we  may  not  be  able  to  develop  our  product  candidates,  raise  capital,  expand  our  business  or
continue  our  operations. The  approach  we  are  taking  to  discover  and  develop  novel  product  candidates  is  unproven  and  may  never  lead  to  marketable
products.

We are concentrating and intend to continue to concentrate our internal research and development efforts primarily on the discovery and development of
product candidates targeting cHBV in order to ultimately develop a functional curative combination regimen, as well as on therapies to treat coronaviruses,
including  COVID-19.  Our  future  success  depends  in  part  on  the  successful  development  of  these  product  candidates. Our  approach  to  the  treatment  of
HBV is unproven, and we do not know whether we will be able to develop any products of commercial value.

There is no known functional cure for HBV. Any compounds that we develop may not effectively address HBV persistence. Even if we are able to develop
compounds  that  address  one  or  more  of  the  key  factors  in  the  HBV  life  cycle  (e.g.,  HBV  replication,  HBsAg  expression  and  immune  reactivation),
targeting these key factors has not been proven to functionally cure HBV. If we cannot develop compounds to achieve our goal of functionally curing HBV
internally, we may be unable to acquire additional product candidates on terms acceptable to us, or at all. Even if we are able to acquire or develop product
candidates that address one of these mechanisms of action in pre-clinical studies, we may not succeed in demonstrating safety and efficacy of the product
candidate in clinical trials. If we are unable to identify suitable compounds for pre-clinical and clinical development, we will not succeed in realizing our
goal of a functional curative combination regimen for HBV.

We  will  require  substantial  additional  capital  to  fund  our  operations.  Additional  funds  may  be  dilutive  to  shareholders  or  impose  operational
restrictions. Further, if additional capital is not available, we may need to delay, limit or eliminate our research, development and commercialization
programs and modify our business strategy.

Our principal sources of liquidity are cash, cash equivalents and investments in marketable securities, which were $184.3 million as of December 31, 2022.
We believe that our $184.3 million of cash, cash equivalents and investments in marketable securities as of December 31, 2022 will be sufficient to fund
our operations into the fourth quarter of 2024. However, changing circumstances may cause us to consume capital faster than we currently anticipate, and
we may need to spend more money than

34

currently expected because of such circumstances. Within the next several years, substantial additional funds will be required to continue with the active
development of our pipeline product candidates and technologies. In particular, our funding needs may vary depending on a number of factors including:

•
•
•
•

revenues earned from our licensing partners, including Alnylam, Qilu, Acuitas and Gritstone Oncology, Inc. (“Gritstone”);
the extent to which we continue the development of our product candidates or form licensing arrangements to advance our product candidates;
our decisions to in-license or acquire additional products, additional product candidates or technology for development;
our ability to attract and retain development or commercialization partners, and their effectiveness in carrying out the development and ultimate
commercialization of one or more of our product candidates;

• whether  batches  of  product  candidates  that  we  manufacture  fail  to  meet  specifications  resulting  in  clinical  trial  delays  and  investigational  and

remanufacturing costs;
the decisions, and the timing of decisions, made by health regulatory agencies regarding our technology and product candidates;
competing products, product candidates and technological and market developments; and
prosecuting and enforcing our patent claims and other intellectual property rights.

•
•
•

We  will  seek  to  obtain  funding  to  maintain  and  advance  our  business  from  a  variety  of  sources  including  equity  financings,  debt  financings,  licensing
agreements, partnerships, government grants and contracts and other strategic transactions and funding opportunities. There can be no assurance that we
will be able to complete any such transaction on acceptable terms or otherwise.

If we are able to raise additional capital through the issuance of equity securities, the percentage ownership of our current shareholders will be reduced. In
addition, we may issue equity as part of the consideration to our licensors, to compensate consultants or to settle outstanding payables, all of which could
cause our shareholders to experience additional dilution in net book value per share. Any such additional equity securities may have rights, preferences and
privileges senior to those of the holders of our common shares.

Debt financing, if available, will result in increased fixed payment obligations and may involve agreements that include covenants limiting or restricting
our ability to take specific actions, such as incurring additional debt, making capital expenditures or declaring dividends. Any debt financing or additional
equity that we raise may contain terms, such as liquidation and other preferences, which are not favorable to us or our existing shareholders. If we raise
additional  funds  through  corporate  collaborations,  partnerships  or  other  strategic  transactions,  it  may  be  necessary  to  relinquish  valuable  rights  to  our
product candidates, our technologies or future revenue streams or to grant licenses or sell assets on terms that may not be favorable to us.

If we are unable to raise additional capital in sufficient amounts or on terms acceptable to us, we will need to curtail and reduce our operations and costs,
and modify our business strategy which may require us to, among other things:

•

•

•

•

significantly delay, scale back or discontinue the development or commercialization of one or more of our product candidates or one or more of
our research and development initiatives;
seek collaborators for one or more of our product candidates or one or more of our research and development initiatives at an earlier stage than
otherwise would be desirable or on terms that are less favorable than might otherwise be available;
sell or license on unfavorable terms our rights to one or more of our technologies, product candidates or research and development initiatives that
we otherwise would seek to develop or commercialize ourselves; or
cease operations.

35

We have incurred losses in nearly every year since our inception and we anticipate that we will not achieve profits for the foreseeable future. To date,
we have had no product revenues.

With the exception of the years ended December 31, 2006 and December 31, 2012, we have incurred losses each fiscal year since inception through the
year ended December 31, 2022 and have not received any revenues other than from research and development collaborations, royalties, license fees and
milestone  payments.  From  inception  to  December  31,  2022,  we  have  an  accumulated  net  deficit  of  approximately  $1.2  billion.  Investment  in  drug
development is highly speculative because it entails substantial upfront capital expenditures and significant risk that a product candidate will fail to gain
regulatory  approval  or  become  commercially  viable.  We  continue  to  incur  significant  research,  development  and  other  expenses  related  to  our  ongoing
operations, including development of our product candidates. We do not expect to achieve profits until such time as product sales, milestone payments and
royalty payments, if any, generate sufficient revenues to fund our continuing operations. We cannot predict if we will ever achieve profitability and, if we
do, we may not be able to remain consistently profitable or increase our profitability.

We  expect  to  continue  to  incur  significant  expenses  and  operating  losses  for  the  foreseeable  future.  We  anticipate  that  our  expenses  will  continue  to  be
significant if and as we:

continue our research and pre-clinical and clinical development of our product candidates;
•
initiate additional pre-clinical, clinical or other studies or trials for our product candidates;
•
continue or expand our licensing arrangements with our licensing partners;
•
change or add additional manufacturers or suppliers;
•
seek regulatory approvals for our product candidates that successfully complete clinical trials;
•
establish a sales, marketing and distribution infrastructure to commercialize any product candidates for which we may obtain regulatory approval;
•
seek to identify and validate additional product candidates;
•
•
acquire or in-license other product candidates and technologies;
• maintain, protect and expand our intellectual property portfolio;
•
•
•

attract and retain skilled personnel;
create additional infrastructure to support our research, product development and planned future commercialization efforts; and
experience any delays or encounter issues with any of the above.

The  net  losses  we  incur  may  fluctuate  significantly  from  quarter  to  quarter  and  year  to  year,  such  that  a  period-to-period  comparison  of  our  results  of
operations may not be a good indication of our future performance.

The COVID-19 pandemic could adversely impact our business, including our clinical development plans.

We continue to monitor the effects of COVID-19, which has caused significant disruptions around the world. We may continue to experience disruptions as
a result of the COVID-19 pandemic that could severely impact our business, including:

•

•

•

•

interruption of key manufacturing, research and clinical development activities due to limitations on work and travel imposed or recommended by
federal or state governments, employers and others;
delays  or  difficulties  in  clinical  trial  site  operations,  including  difficulties  in  recruiting  clinical  site  investigators  and  clinical  site  staff  and
difficulties in enrolling subjects or treating subjects in active trials;
the  diversion  of  healthcare  resources  away  from  the  conduct  of  clinical  trial  matters  to  focus  on  COVID-19  pandemic  concerns,  including  the
administration  of  COVID-19  vaccines,  which  could  negatively  affect  the  attention  of  physicians  serving  as  our  clinical  trial  investigators,  the
hospitals serving as our clinical trial sites and the hospital staff supporting the conduct of our clinical trials;
limitations on travel and quarantine requirements that interrupt key clinical trial activities, such as clinical trial site initiations, our ability and the
ability of our clinical research organizations (“CROs”) to access and monitor clinical trial sites, and new clinical trial site policies resulting from
the  COVID-19  pandemic  that  determine  essential  and  non-essential  functions  and  staff,  which  may  impact  the  ability  of  site  staff  to  conduct
assessments or result in delays to the conduct of the assessments as part of our clinical trial protocols, or impact the ability to enter assessment
results into clinical trial databases in a timely manner, or limit the ability of a subject to participate in a clinical trial or delay access to product
candidate dosing or assessments;

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interruption of key business activities due to illness and/or quarantine of key individuals and delays associated with recruiting, hiring and training
new temporary or permanent replacements for such key individuals, both internally and at our third party service providers;
delays in research and clinical trial sites receiving the supplies and materials needed to conduct preclinical studies and clinical trials, due to work
stoppages, travel and shipping interruptions or restrictions or other reasons;
potential clinical trial subjects may be unable or unwilling to participate further (or may have to limit participation) in our clinical trials due to
risks related to the COVID-19 pandemic;
difficulties  in  raising  additional  capital  needed  to  pursue  the  development  of  our  programs  due  to  the  slowing  of  our  economy  and  near  term
and/or long term negative effects of the pandemic on the financial, banking and capital markets;
changes in local regulations as part of a response to the COVID-19 outbreak that may require us to change the ways in which research, including
clinical development, is conducted, which may result in unexpected costs; and
delays  in  necessary  interactions  with  regulators  and  other  important  agencies  and  contractors  due  to  limitations  in  employee  resources,  travel
restrictions or forced furlough of government employees.

If a subject participating in one of our clinical trials contracts COVID-19, this could negatively impact the data readouts from these trials; for example, the
subject  may  be  unable  to  participate  further  (or  may  have  to  limit  participation)  in  our  clinical  trial,  the  subject  may  show  a  different  clinical  trial
assessment than if the subject had not contracted the COVID-19, or the subject could experience an AE that could be attributed to our product candidate.

The global outbreak of COVID-19 continues to evolve, including with the emergence of new COVID-19 variants in 2022. The extent to which the COVID-
19 pandemic may further impact our business will depend on future developments, which are highly uncertain and cannot be predicted with confidence,
such  as  the  duration  of  the  pandemic,  travel  restrictions  and  social  distancing  in  the  United  States  and  other  countries,  business  closures  or  business
disruptions and the effectiveness of actions taken in the United States and other countries to contain and treat the virus.

We do not generate revenues from product sales and may never be profitable.

Our ability to generate revenue and achieve profitability depends on our ability, alone or with strategic partners, to successfully complete the development
of, and obtain the regulatory approvals necessary for, the manufacture and commercialization of our product candidates. We do not anticipate generating
significant revenues from product sales for the foreseeable future, if ever. Our ability to generate future revenues from product sales depends heavily on our
success in:

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completing research and pre-clinical and clinical development of our product candidates;
seeking and obtaining regulatory approvals for product candidates for which we complete clinical trials;
developing a sustainable, scalable, reproducible, and transferable manufacturing process for our product candidates;
establishing and maintaining supply and manufacturing relationships with third parties that can provide adequate (in amount and quality) products
and services to support clinical development and the market demand for our product candidates for which we obtain regulatory approval;
launching and commercializing product candidates for which we obtain regulatory approval, either by collaborating with partners or, if launched
independently, by establishing a sales force, marketing, sales operations and distribution infrastructure;
obtaining market acceptance of our product candidates for which we obtain regulatory approval as viable treatment options;
addressing any competing technological and market developments;
implementing additional internal systems and infrastructure, as needed;
identifying and validating new product candidates;
negotiating favorable terms in any collaboration, licensing or other arrangements into which we may enter;

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• maintaining, protecting and expanding our portfolio of intellectual property rights, including patents, trade secrets and know-how; and
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attracting, hiring and retaining qualified personnel.

Even if one or more of the product candidates that we develop is approved for commercial sale, we anticipate incurring significant costs associated with
commercializing  any  approved  product  candidate.  Our  expenses  could  increase  beyond  expectations  if  we  are  required  by  the  FDA  or  other  regulatory
authorities outside the United States to perform clinical trials or

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other studies in addition to those that we currently anticipate. Even if we are able to generate revenues from the sale of any approved product candidates,
we may not become profitable and may need to obtain additional funding to continue operations.

Risks Related to Development, Clinical Testing, Regulatory Approval, Marketing, and Coverage and Reimbursement of our Product Candidates

Our product candidates are in early stages of development and must go through clinical trials, which are very expensive, time-consuming and difficult
to design and implement. The outcomes of clinical trials are uncertain, and delays in the completion of or the termination of any clinical trial of our
product candidates could harm our business, financial condition and prospects.

Our research and development programs are at an early stage of development. We must demonstrate our product candidates’ safety and efficacy in humans
through extensive clinical testing, which is expensive and time-consuming and requires specialized knowledge and expertise.

Clinical trials are also expensive and difficult to design and implement, in part because they are subject to rigorous regulatory requirements. The clinical
trial process is also time-consuming, and the outcome is not certain. We estimate that clinical trials of our product candidates will take multiple years to
complete.  Failure  can  occur  at  any  stage  of  a  clinical  trial,  and  we  could  encounter  problems  that  cause  us  to  abandon  or  repeat  clinical  trials.  The
commencement and completion of clinical trials may be delayed or precluded by a number of factors, including:

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delay or failure in reaching agreement with the FDA or other regulatory authority outside the United States on the design of a given trial, or in
obtaining authorization to commence a trial;
delay or failure in reaching agreement on acceptable terms with prospective CROs and clinical trial sites;
delay or failure in obtaining approval of an IRB or ethics committees before a clinical trial can be initiated at a given site;
any  shelter-in-place  orders  from  local,  state  or  federal  governments  or  clinical  trial  site  policies  resulting  from  the  COVID-19  pandemic  that
determine essential and non-essential functions and staff, which may impact the ability of the staff to conduct assessments or result in delays to the
conduct of the assessments as part of our clinical trial protocols, or impact the ability to enter assessment results into clinical trial databases in a
timely manner;

• withdrawal  of  clinical  trial  sites  from  our  clinical  trials,  including  as  a  result  of  changing  standards  of  care  or  the  ineligibility  of  a  site  to

participate;
delay or failure in recruiting and enrolling subjects in our clinical trials;
delay or failure in having subjects complete a clinical trial or return for post-treatment follow up;
clinical sites or investigators deviating from trial protocol, failing to conduct the trial in accordance with applicable regulatory requirements, or
dropping out of a trial;
inability to identify and maintain a sufficient number of trial sites;
failure of CROs to meet their contractual obligations or deadlines;
the need to modify a trial protocol;
unforeseen safety issues;
emergence of dosing issues;
lack of effectiveness data during clinical trials;
changes in the standard of care of the indication being studied;
reliance  on  third-party  suppliers  for  the  clinical  trial  supply  of  product  candidates  and  failure  by  our  third-party  suppliers  to  comply  with
regulatory requirements or meet their contractual obligations to us in a timely manner, or at all;
inability to monitor subjects adequately during or after treatment;
limitations  on  our  or  our  CROs’  ability  to  access  and  verify  clinical  trial  data  captured  at  clinical  trial  sites  through  monitoring  and  source
document verification;
lack of sufficient funding to finance the clinical trials; and
changes in governmental regulations or administrative action.

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We, the FDA, other regulatory authorities outside the United States, or an IRB may suspend a clinical trial at any time for various reasons, including if it
appears that the clinical trial is exposing participants to unacceptable health risks or if the FDA

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or one or more other regulatory authorities outside the United States find deficiencies in our IND or similar application outside the United States or the
conduct of the trial. If we experience delays in the completion of, or the termination of, any clinical trial of any of our product candidates, the commercial
prospects of such product candidate will be harmed, and our ability to generate product revenues from such product candidate will be delayed or rendered
impossible. In addition, any delays in completing our clinical trials will increase our costs, slow down our product candidate development and approval
process, and jeopardize our ability to commence product sales and generate revenues.

Even  if  our  clinical  trials  are  successfully  completed  as  planned,  the  results  may  not  support  approval  of  our  product  candidates  under  the  laws  and
regulations  of  the  FDA  or  other  regulatory  authorities  outside  the  United  States.  The  clinical  trial  process  may  fail  to  demonstrate  that  our  product
candidates are both safe and effective for their intended uses. Pre-clinical and clinical data and analyses are often able to be interpreted in different ways.
Even  if  we  view  our  results  favorably,  if  a  regulatory  authority  has  a  different  view,  we  may  still  fail  to  obtain  regulatory  approval  of  our  product
candidates.

Any of these occurrences may harm our business, financial condition, results of operations, cash flows and prospects significantly.

Pre-clinical  studies  and  preliminary  and  interim  data  from  clinical  trials  of  our  product  candidates  are  not  necessarily  predictive  of  the  results  or
success of ongoing or later clinical trials of our product candidates. If we cannot replicate the results from our pre-clinical studies and initial clinical
trials of our product candidates in later clinical trials, we may be unable to successfully develop, obtain regulatory approval for and commercialize our
product candidates.

Pre-clinical studies and any positive preliminary and interim data from our clinical trials of our product candidates may not necessarily be predictive of the
results  of  ongoing  or  later  clinical  trials.  A  number  of  companies  in  the  pharmaceutical  and  biotechnology  industries,  including  us  and  many  other
companies with greater resources and experience than we, have suffered significant setbacks in clinical trials, even after seeing promising results in prior
pre-clinical  studies  and  clinical  trials.  Even  if  we  are  able  to  complete  our  planned  clinical  trials  of  our  product  candidates  according  to  our  current
development  timeline,  initial  positive  results  from  pre-clinical  studies  and  clinical  trials  of  our  product  candidates  may  not  be  replicated  in  subsequent
clinical  trials.  The  design  of  our  later  stage  clinical  trials  could  differ  in  significant  ways  (e.g.,  inclusion  and  exclusion  criteria,  endpoints,  statistical
analysis plan) from our earlier stage clinical trials, which could cause the outcomes of the later stage trials to differ from those of our earlier stage clinical
trials. If we fail to produce positive results in our planned clinical trials of any of our product candidates, the development timeline and regulatory approval
and  commercialization  prospects  for  our  product  candidates,  and,  correspondingly,  our  business  and  financial  prospects,  would  be  materially  adversely
affected.

Because  we  have  limited  resources,  we  may  decide  to  pursue  a  particular  product  candidate  and  fail  to  advance  product  candidates  that  later
demonstrate a greater chance of clinical and commercial success.

We are an early-stage company with limited resources and revenues. The product candidates we currently have under development will require significant
development, pre-clinical and clinical testing and investment of significant funds before their commercialization. Because of this, we must make strategic
decisions regarding resource allocations and which product candidates to pursue. There can be no assurance that we will be able to develop all potentially
promising product candidates that we may identify. Based on preliminary results, we may choose to advance a particular product candidate that later fails to
be successful, and simultaneously forgo or defer further investment in other product candidates that later are discovered to demonstrate greater promise in
terms of clinical and commercial success. If we make resource allocation decisions that later are shown to be inaccurate, our business and prospects could
be harmed.

Several of our current pre-clinical studies and clinical trials are being conducted outside the United States, and the FDA may not accept data from
trials conducted in locations outside the United States.

Several of our current pre-clinical studies and clinical trials are being conducted outside the United States and we may conduct further pre-clinical studies
and  clinical  trials  outside  the  United  States  in  the  future.  We  are  currently  conducting  clinical  trials  in  Moldova,  Thailand,  Taiwan,  South  Korea,  Hong
Kong, the United Kingdom, Australia and New Zealand, among other countries. To the extent we do not conduct these clinical trials under an IND, the
FDA  may  not  accept  data  from  such  trials.  Although  the  FDA  may  accept  data  from  clinical  trials  conducted  outside  the  United  States  that  are  not
conducted under an IND, the FDA’s acceptance of these data is subject to certain conditions. For example, the clinical trial must be well designed

39

and  conducted  and  performed  by  qualified  investigators  in  accordance  with  ethical  principles.  The  trial  population  must  also  adequately  represent  the
United States population, and the data must be applicable to the United States population and United States medical practice in ways that the FDA deems
clinically  meaningful.  In  general,  the  patient  population  for  any  clinical  trials  conducted  outside  of  the  United  States  must  be  representative  of  the
population for whom we intend to label the product in the United States. In addition, while these clinical trials are subject to the applicable local laws, FDA
acceptance of the data will be dependent upon its ability to verify the data and its determination that the trials complied with all applicable United States
laws and regulations. We cannot assure you that the FDA will accept data from trials conducted outside of the United States that are not conducted under an
IND.  If  the  FDA  does  not  accept  the  data  from  such  clinical  trials,  we  likely  would  need  to  conduct  additional  trials,  which  would  be  costly  and  time-
consuming and could delay or permanently halt our development of our product candidates.

We  cannot  guarantee  how  long  it  will  take  regulatory  agencies  to  review  our  applications  for  product  candidates,  and  we  may  fail  to  obtain  the
necessary regulatory approvals to market our product candidates.

Before we can commercialize our product candidates in the United States, we must obtain approval from the FDA. We must similarly obtain approvals
from comparable regulatory authorities to commercialize our product candidates in jurisdictions outside the United States.

To obtain marketing approval, United States laws require:

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controlled research and human clinical testing that comply with GLP and GCP, as applicable;
establishment of the safety and efficacy of the product for each use sought;
government review and approval of a submission containing, among other things, manufacturing, pre-clinical and clinical data; and
compliance with GMP regulations.

The process of reviewing and approving a drug is time-consuming, unpredictable, and dependent on a variety of factors outside of our control. The FDA
and corresponding regulatory authorities in jurisdictions outside the United States have a significant amount of discretion in deciding whether or not to
approve  a  marketing  application.  Our  product  candidates  could  fail  to  receive  regulatory  approval  from  the  FDA  or  comparable  regulatory  authorities
outside the United States for several reasons, including:

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disagreement with the design or implementation of our clinical trials;
failure to demonstrate that our product candidate is safe and effective for the proposed indication;
failure of clinical trial results to meet the level of statistical significance required for approval;
failure to demonstrate that the product candidate’s benefits outweigh its risks;
disagreement with our interpretation of pre-clinical or clinical data; and
inadequacies in the manufacturing facilities or processes of third-party manufacturers.

The FDA or comparable regulatory authorities outside the United States may require us to conduct additional pre-clinical and clinical testing, which may
delay or prevent approval of a product candidate and our commercialization plans, or cause us to abandon the development program. Further, any approval
we receive may be for fewer or more limited indications than we request, may not include labeling claims necessary for successful commercialization of
the  product  candidate,  or  may  be  contingent  upon  our  conducting  costly  post-marketing  studies.  Any  of  these  scenarios  could  materially  harm  the
commercial prospects of a product candidate, and our operations will be adversely effected.

If  a  particular  product  candidate  causes  undesirable  side  effects,  then  we  may  be  unable  to  receive  regulatory  approval  of  or  commercialize  such
product candidate.

We may experience numerous unforeseen events during, or as a result of, the testing process that could delay or prevent commercialization of any of our
product  candidates,  including  the  occurrence  of  undesirable  side  effects.  Such  side  effects  could  lead  to  clinical  trial  challenges,  such  as  difficulties  in
subject recruitment, retention, and adherence, potential product liability claims, and possible termination by health authorities. These types of clinical trial
challenges could in turn, delay or prevent regulatory approval of our product candidate. Side effects may also lead regulatory authorities to require stronger
product warnings on the product label, costly post-marketing studies, and/or a Risk Evaluation and Mitigation Strategy

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(“REMS”), among other possible requirements. If the product candidate has already been approved, such approval may be withdrawn. Any delay in, denial,
or withdrawal of marketing approval for one of our product candidates will adversely affect our business, including our results of operations and financial
position.  Even  if  one  or  more  of  our  product  candidates  receives  marketing  approval,  undesirable  side  effects  may  limit  such  product’s  commercial
viability.  Patients  may  not  wish  to  use  our  product,  physicians  may  not  prescribe  our  product,  and  our  reputation  may  suffer.  Any  of  these  events  may
significantly harm our business and financial prospects.

We may find it difficult to enroll patients in our clinical trials, which could delay or prevent clinical trials of our product candidates.

Identifying  and  qualifying  patients  to  participate  in  clinical  trials  of  our  product  candidates  is  critical  to  our  success.  The  timing  of  our  clinical  trials
depends in part on the speed at which we can recruit patients to participate in testing our product candidates.

We may not be able to identify, recruit and enroll a sufficient number of patients, or those with required or desired characteristics to achieve diversity in a
clinical trial, or complete our clinical trials in a timely manner. Subject enrollment is affected by a variety factors including, among others:

severity of the disease under investigation;
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design of the trial protocol;
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prevalence of the disease/size of the patient population;
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eligibility criteria for the clinical trial in question;
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perceived risks and benefits of the product candidate under study;
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• willingness or availability of patients to participate in the clinical trials;
proximity and availability of clinical trial sites for prospective patients;
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ability to recruit clinical trial investigators with the appropriate competencies and experience;
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availability of competing therapies and clinical trials;
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efforts to facilitate timely enrollment in clinical trials;
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ability to obtain and maintain subject consents;
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patient referral practices of physicians;
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risk that patients enrolled in clinical trials will drop out of the trials before completion; and
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ability to monitor patients adequately during and after treatment.
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If patients are unwilling to participate in our clinical trials, the timeline for recruiting patients, conducting clinical trials and obtaining regulatory approval
of potential products may be delayed. These delays could result in increased costs, delays in advancing or testing our product candidates or termination of
the clinical trials altogether.

Several of our and our collaboration partner’s current and planned clinical trials have been impacted and could be further delayed or suspended as a
result of the military action by Russia in Ukraine.

In February 2022, Russia commenced a military invasion of Ukraine. A portion of our clinical trial evaluating AB-836 and a cohort of Antios Therapeutics,
Inc.’s  (“Antios”)  clinical  trial  evaluating  a  triple  combination  including  AB-729  were  being  conducted  in  Ukraine  at  that  time.  We  had  also  planned  to
conduct a portion of the following clinical trials in Ukraine: (i) our Phase 2a clinical trial evaluating AB-729 in combination with ongoing NA therapy and
short courses of PEG-IFNα-2a in cHBV patients and (ii) our planned Phase 2a clinical trial to evaluate a triple combination of AB-729 with Vaccitech’s
VTP-300 and an NA therapy. As a result of such military invasion, we intend to utilize alternative clinical trial sites for our ongoing and planned clinical
trials impacted by the military action in Ukraine.

Russia’s  invasion  and  the  ensuing  response  by  Ukraine  has  disrupted  our  and  our  collaboration  partners’  current  clinical  trials  in  such  jurisdictions  and
could  increase  our  costs  and  disrupt  future  planned  clinical  development  activities.  For  example,  enrollment  was  completed  in  a  cohort  of  patients  in
Antios’ ongoing Phase 2a proof-of-concept clinical trial evaluating a triple combination of AB-729, Antios’ proprietary Active Site Polymerase Inhibitor
Nucleotide  (ASPIN),  ATI-2173,  and  Viread  (tenofovir  disoproxil  fumarate),  a  nucleos(t)ide  reverse  transcriptase  inhibitor.  However,  the  majority  of
patients in this cohort

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were enrolled in Ukraine and, as a result, these patients have been lost to follow-up before completing the clinical trial. Antios terminated this clinical trial
and we have terminated our clinical collaboration agreement with Antios.

Although the length and impact of Russia’s military action is highly unpredictable, actions by Russia, or potentially other countries, against Ukraine and
surrounding areas may adversely affect our ability to adequately conduct or complete certain clinical trials and maintain compliance with relevant protocols
due to, among other reasons, the prioritization of hospital resources away from clinical trials, reallocation or evacuation of site staff and subjects, or as a
result of government-imposed curfews, warfare, violence, or other governmental action or events that restrict movement. These developments may also
result in our inability to access sites for monitoring or to obtain data from affected sites or patients going forward. We could also experience disruptions in
our supply chain or limits to our ability to provide sufficient investigational materials in Ukraine and surrounding regions. Alternative sites to fully and
timely compensate for our clinical trial activities in Ukraine may not be available and we may need to find other countries to conduct these clinical trials. If
these clinical trials are further interrupted, our clinical development plans for these product candidates could be significantly delayed, which would increase
our  costs,  slow  down  our  product  candidate  development  and  approval  process  and  jeopardize  our  ability  to  commence  product  sales  and  generate
revenues.

Even if our product candidates obtain regulatory approval, they will remain subject to ongoing regulatory requirements and oversight.

Approved drug products are subject to ongoing regulatory requirements and oversight, including requirements related to manufacturing, quality control,
further development, labeling, packaging, storage, distribution, safety surveillance, import, export, advertising, promotion, recordkeeping and reporting. In
addition, we will be subject to continued compliance with GMP and GCP requirements for any clinical trials that we conduct post-approval. If we or any of
the third parties on which we rely fail to meet those requirements, the FDA or comparable regulatory authorities outside the United States could initiate
enforcement actions. Other potential consequences include the issuance of fines, warning letters, untitled letters or holds on clinical trials, product seizure
or detention or refusal to permit the import or export of our product candidates, permanent injunctions and consent decrees, or the imposition of civil or
criminal  penalties,  any  of  which  could  significantly  impair  our  ability  to  successfully  commercialize  a  given  product.  If  the  FDA  or  a  comparable
regulatory  authority  outside  the  United  States  becomes  aware  of  new  safety  information,  it  can  impose  additional  restrictions  on  how  the  product  is
marketed or may seek to withdraw marketing approval altogether.

Further, the U.S. and state governments have shown significant interest in establishing cost containment measures to limit the growth of government-paid
health care costs, including price controls, restrictions on reimbursement, and requirements for substitution of generic products for branded prescription
drugs. For example, in March 2010, the Patient Protection and Affordable Care Act, as amended (the “ACA”), became law in the United States. A primary
goal  of  the  ACA  is  to  reduce  the  cost  of  health  care,  and  it  has  substantially  changed  the  way  health  care  is  financed  by  both  government  and  private
insurers. While we cannot predict with certainty what impact on federal and other reimbursement policies this legislation will have in general or on our
business specifically, the ACA may result in downward pressure on pharmaceutical reimbursement, which could negatively affect market acceptance of,
and the price we may charge for, any products we develop that receives regulatory approval. Legislative changes to and regulatory changes under the ACA
remain  possible,  but  the  nature  and  extent  of  such  potential  additional  changes  are  uncertain  at  this  time.  We  expect  that  the  ACA,  its  implementation,
efforts  to  modify  or  invalidate  the  ACA,  or  portions  thereof,  or  its  implementation,  and  other  healthcare  reform  measures,  including  those  that  may  be
adopted  in  the  future,  could  have  a  material  adverse  effect  on  our  industry  generally  and  on  our  ability  to  successfully  commercialize  our  product
candidates, if approved.

The marketability of any products for which we receive regulatory approval for commercial sale may suffer if the government and third-party payors fail to
provide adequate coverage and reimbursement. In addition, cost containment measures in the United States has been an area or increasing emphasis, and
we expect they will continue to increase the pressure on pharmaceutical pricing. Coverage policies and third-party reimbursement rates may change at any
time. Even if favorable coverage and reimbursement status is attained for one or more products for which we receive regulatory approval, less favorable
coverage policies and reimbursement rates may be adopted in the future.

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We face significant competition from other biotechnology and pharmaceutical companies targeting HBV and coronaviruses, including COVID-19.

As a significant unmet medical need exists for HBV, there are several large and small pharmaceutical companies focused on delivering therapeutics for
treatment  of  HBV.  These  companies  include,  but  are  not  limited  to  Roche,  Vir  Biotechnology,  GlaxoSmithKline,  Gilead  Sciences,  Assembly,  Enanta
Pharmaceuticals, Aligos Therapeutics and Vaccitech. Further, in addition to current investigational therapeutics in development, it is likely that additional
drugs will become available in the future for the treatment of HBV.

In addition, given the severity of the global coronavirus pandemic, several companies are developing or commercializing therapeutics for the treatment of
coronaviruses. These companies include, but are not limited to, Pfizer, Merck, Gilead, Vir Biotechnology, Shionogi, PardesBio, Enanta Pharmaceuticals,
Aligos Therapeutics and Cocrystal Pharma.

Many  of  our  existing  or  potential  competitors  have  substantially  greater  financial,  technical  and  human  resources  than  we  do  and  significantly  greater
experience in the discovery and development of product candidates, as well as in obtaining regulatory approvals of those product candidates in the United
States and other countries. Many of our current and potential future competitors also have significantly more experience commercializing drugs that have
been approved for marketing.

We anticipate significant competition in the HBV and coronavirus markets, with several early and late phase product candidates announced. We will also
face  competition  for  other  product  candidates  that  we  expect  to  develop  in  the  future.  Competition  may  increase  further  as  a  result  of  advances  in  the
commercial applicability of technologies and greater availability of capital for investment in these industries. Our competitors may succeed in developing,
acquiring or licensing, on an exclusive basis, product candidates that are more effective or less costly than any product candidate that we may develop.

If  we  successfully  develop  product  candidates,  and  obtain  approval  for  them,  we  will  face  competition  based  on  many  different  factors,  including  the
following:

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safety and effectiveness of our products;
ease with which our products can be administered and the extent to which patients and physicians accept new routes of administration;
timing and scope of regulatory approvals for these products;
availability and cost of manufacturing, marketing and sales capabilities;
price;
reimbursement coverage; and
patent position.

Our  competitors  may  develop  or  commercialize  products  with  significant  advantages  over  any  products  we  develop  based  on  any  of  the  factors  listed
above, or on other factors. Our competitors may therefore be more successful in commercializing their products than we are, which could adversely affect
our competitive position and business. Competitive products may make any products we develop and commercialize obsolete or uncompetitive before we
can recover the expenses of developing and commercializing such products. Such competitors could also recruit our employees, which could negatively
impact our level of expertise and the ability to execute on our business plan.

We are largely dependent on the future commercial success of our HBV and coronavirus product candidates.

Our ability to generate revenues and become profitable will depend in large part on the future commercial success of our HBV and coronavirus product
candidates, if they are approved for marketing. If any product that we commercialize in the future does not gain an adequate level of acceptance among
physicians, patients and third parties, or our estimates of the number of people who have cHBV or are infected with coronaviruses are lower than expected,
we may not generate significant product revenues or become profitable. Market acceptance by physicians, patients and third party payors of the products
we may commercialize will depend on a number of factors, some of which are beyond our control, including:

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their efficacy, safety and other potential advantages in relation to alternative treatments;
their relative convenience and ease of administration in relation to alternative treatments;

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the  availability  of  adequate  coverage  or  reimbursement  by  third  parties,  such  as  insurance  companies  and  other  healthcare  payors,  and  by
government healthcare programs, including Medicare and Medicaid;
the prevalence and severity of adverse events;
their cost of treatment in relation to alternative treatments, including generic products;
the extent and strength of our third party manufacturer and supplier support;
the extent and strength of marketing and distribution support;
the limitations or warnings contained in a product’s approved labeling; and
distribution  and  use  restrictions  imposed  by  the  FDA  or  other  regulatory  authorities  outside  the  United  States  or  that  are  part  of  a  REMS  or
voluntary risk management plan.

For example, even if our products have been approved by the FDA, physicians and patients may not immediately be receptive to them and may be slow to
adopt  them.  If  our  products  do  not  achieve  an  adequate  level  of  acceptance  among  physicians,  patients  and  third  party  payors,  we  may  not  generate
meaningful revenues and we may not become profitable.

We may incur substantial liabilities and may be required to limit commercialization of our products in response to product liability lawsuits.

The testing and marketing of medical products entail an inherent risk of product liability. Product liability claims may be brought against us by patients,
healthcare providers or others using, administering or selling our products. Large judgments have been awarded in class action lawsuits based on drugs that
had unanticipated side effects, which is an example of just one possible product liability claim that may be brought against us. If we cannot successfully
defend  ourselves  against  product  liability  claims,  we  may  incur  substantial  liabilities  or  be  required  to  limit  commercialization  of  our  products.  Our
inability to obtain sufficient product liability insurance at an acceptable cost to protect against potential product liability claims could prevent or inhibit the
commercialization of products we develop, alone or with partners. Although we currently have product liability insurance coverage for our clinical trials
for expenses or losses, our insurance coverage is limited to $10 million per occurrence, and $10 million in the aggregate, and may not reimburse us or may
not be sufficient to reimburse us for any or all expenses or losses we may suffer. Moreover, insurance coverage is becoming increasingly expensive and in
the future, we may not be able to maintain insurance coverage at a reasonable cost or in sufficient amounts to protect us against losses due to liability. We
intend  to  expand  our  insurance  coverage  to  include  the  sale  of  commercial  products  if  we  obtain  marketing  approval  for  our  product  candidates  in
development, but we may be unable to obtain commercially reasonable product liability insurance for any products approved for marketing. Further, even if
our  agreements  with  any  current  or  future  partners  entitle  us  to  indemnification  against  losses,  such  indemnification  may  not  be  available  or  adequate
should any claims arise. A successful product liability claim or series of claims brought against us could cause our share price to fall and, if judgments
exceed our insurance coverage, could decrease our cash and adversely affect our business.

Coverage  and  adequate  reimbursement  may  not  be  available  for  our  product  candidates,  which  could  make  it  difficult  for  us  to  sell  our  products
profitably.

Market  acceptance  and  sales  of  any  products  that  we  develop  will  depend  in  part  on  the  extent  to  which  reimbursement  for  these  products  and  related
treatments will be available from third party payors, including government health administration authorities and private health insurers. Third party payors
decide  which  drugs  they  will  pay  for  and  establish  reimbursement  levels.  Third  party  payors  often  rely  upon  Medicare  coverage  policy  and  payment
limitations in setting their own reimbursement policies. However, decisions regarding the extent of coverage and amount of reimbursement to be provided
for  each  of  our  products  will  be  made  on  a  plan  by  plan  basis.  One  payor’s  determination  to  provide  coverage  for  a  product  does  not  assure  that  other
payors will also provide coverage, and adequate reimbursement, for the product. Additionally, a third party payor’s decision to provide coverage for a drug
does  not  imply  that  an  adequate  reimbursement  rate  will  be  approved.  Each  plan  determines  whether  or  not  it  will  provide  coverage  for  a  drug,  what
amount it will pay the manufacturer for the drug, and on what tier of its formulary the drug will be placed. The position of a drug on a formulary generally
determines the copayment that a patient will need to make to obtain the drug and can strongly influence the adoption of a drug by patients and physicians.
Patients  who  are  prescribed  treatments  for  their  conditions  and  providers  performing  the  prescribed  services  generally  rely  on  third  party  payors  to
reimburse all or part of the associated healthcare costs. Patients are unlikely to use our products unless coverage is provided and reimbursement is adequate
to cover a significant portion of the cost of our products.

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A primary trend in the United States healthcare industry and elsewhere is cost containment. Third party payors have attempted to control costs by limiting
coverage  and  the  amount  of  reimbursement  for  particular  medications.  We  cannot  be  sure  that  coverage  and  reimbursement  will  be  available  for  any
product that we commercialize and, if reimbursement is available, what the level of reimbursement will be. Inadequate coverage and reimbursement may
impact the demand for, or the price of, any product for which we obtain marketing approval. If coverage and adequate reimbursement is not available, or is
available only to limited levels, we may not be able to successfully commercialize any product candidates that we develop.

Additionally,  there  have  been  a  number  of  legislative  and  regulatory  proposals  to  change  the  healthcare  system  in  the  United  States  and  in  some
jurisdictions  outside  the  United  States  that  could  affect  our  ability  to  sell  any  future  products  profitably.  These  legislative  and  regulatory  changes  may
negatively impact the reimbursement for any future products, following approval.

We are subject to United States and Canadian healthcare laws and regulations, which could expose us to criminal sanctions, civil penalties, contractual
damages, reputational harm, fines, disgorgement, exclusion from participation in government healthcare programs, curtailment or restricting of our
operations and diminished profits and future earnings.

Healthcare providers, physicians and others will play a primary role in the recommendation and prescription of any products for which we obtain marketing
approval.  Our  future  arrangements  with  healthcare  providers,  patients  and  third  party  payors  will  expose  us  to  broadly  applicable  United  States  and
Canadian fraud and abuse and other healthcare laws and regulations that may constrain the business or financial arrangements and collaborative partners
through  which  we  market,  sell  and  distribute  any  products  for  which  we  obtain  marketing  approval.  Restrictions  under  applicable  federal  and  state
healthcare laws and regulations are described in further detail in the section entitled Government Regulation – Post-Approval Regulation and include the
following:

•

•

the  U.S.  federal  Anti-Kickback  Law  prohibits  persons  from,  among  other  things,  knowingly  and  willfully  soliciting,  offering,  receiving  or
providing  remuneration,  directly  or  indirectly,  in  cash  or  in  kind,  to  induce  or  reward,  or  in  return  for,  the  referral  of  an  individual  for  the
furnishing or arranging for the furnishing, or the purchase, lease or order, or arranging for or recommending purchase, lease or order, any good or
service for which payment may be made under a federal healthcare program such as Medicare and Medicaid;

the U.S. federal civil False Claims Act imposes civil penalties, sometimes pursued through whistleblower or qui tam actions, against individuals or
entities  for,  among  other  things,  knowingly  presenting,  or  causing  to  be  presented  claims  for  payment  of  government  funds  that  are  false  or
fraudulent  or  making  a  false  statement  material  to  an  obligation  to  pay  money  to  the  government  or  knowingly  concealing  or  knowingly  and
improperly avoiding, decreasing, or concealing an obligation to pay money to the federal government;

• HIPAA imposes criminal liability for knowingly and willfully executing a scheme to defraud any healthcare benefit program, knowingly and

willfully falsifying, concealing or covering up a material fact or making any materially false, fictitious or fraudulent statement or representation, or
making or using any false writing or document knowing the same to contain any materially false fictitious or fraudulent statement or entry, in
connection with the delivery of or payment for healthcare benefits, items or services;

• HIPAA  and  its  implementing  regulations  also  impose  obligations  on  certain  covered  entity  health  care  providers,  health  plans  and  health  care
clearinghouses as well as their business associates that perform certain services involving the use or disclosure of individually identifiable health
information, including mandatory contractual terms, with respect to safeguarding the privacy, security and transmission of individually identifiable
health information. We may obtain health information from third parties (including research institutions from which we obtain clinical trial data)
that are subject to privacy and security requirements under HIPAA. Although we are not directly subject to HIPAA - other than with respect to
providing certain employee benefits - we could potentially be subject to criminal penalties if we, our affiliates, or our agents knowingly receive
individually identifiable health information maintained by a HIPAA-covered entity in a manner that is not authorized or permitted by HIPAA;

•

numerous federal and state laws and regulations that address privacy and data security, including state data breach notifications laws, state health
information and/or genetic privacy laws, and federal and state consumer protection laws (e.g., Section 5 of the Federal Trade Commission Act, or
FTC Act, and the CCPA), govern the collection, use, disclosure and protection of health-related and other personal information, many of which
differ from each other in significant ways, thus complicating the compliance efforts. Compliance with these laws is difficult, constantly

45

•

•

•

•

•

evolving, and time-consuming, and companies that do not comply with these laws may face government enforcement actions, civil and/or criminal
penalties, or private action, as well as adverse publicity that could negatively affect our operating results and business;

activities  outside  of  the  U.S.  implicate  local  and  national  data  protection  standards,  impose  additional  compliance  requirements  and  generate
additional  risks  of  enforcement  for  non-compliance.  The  European  Union’s  GDPR  and  other  data  protection,  privacy  and  similar  national,
state/provincial and local laws may restrict the access, use and disclosure of patient health information abroad. Compliance efforts will likely be an
increasing and substantial cost in the future;

the  U.S.  federal  Physician  Payment  Sunshine  Act,  being  implemented  as  the  Open  Payments  Program,  which  requires  manufacturers  of  drugs,
devices, biologics, and medical supplies for which payment is available under Medicare, Medicaid or the Children’s Health Insurance Program
(with certain exceptions) to report annually to CMS information related to direct or indirect payments and other transfers of value to physicians
and teaching hospitals (and certain other practitioners beginning as of 2022), as well as ownership and investment interests held in the company by
physicians and their immediate family members;

price reporting requirements under the Medicaid Drug Rebate Program and the 340B Program and with respect to average sales price reporting
under the Medicare Part B program, and rebate or discount liability under the Medicaid Drug Rebate Program, the 340B Program, and Medicare
Part  D,  with  respect  to  which  we  could  be  subject  to  civil  monetary  penalties  for  a  failure  to  comply  with  our  reporting  or  rebate  or  discount
obligations, or termination from the Medicaid Drug Rebate Program or 340B program, which, in turn, could jeopardize the availability of federal
funds for our products under Medicaid and Medicare Part B;

the IRA, which, among other things, requires the U.S. Secretary of Health and Human Services to negotiate, with respect to Medicare units and
subject  to  a  specified  cap,  the  price  of  a  set  number  of  certain  high  Medicare  spend  drugs  and  biologicals  per  year  starting  in  2026,  penalizes
manufacturers of certain Medicare Parts B and D drugs for price increases above inflation, and makes several changes to the Medicare Part D
benefit, including a limit on annual out-of-pocket costs, and a change in manufacturer liability under the program which could negatively affect
our business and financial condition; and

analogous state laws and laws and regulations outside the United States, such as state anti-kickback and false claims laws, which may apply to
sales or marketing arrangements and claims involving healthcare items or services reimbursed by non-governmental third-party payors, including
private insurers; state laws and laws outside the United States that require pharmaceutical companies to comply with the pharmaceutical industry’s
voluntary compliance guidelines and the relevant compliance guidance promulgated by the federal government or otherwise restrict payments that
may be made to certain healthcare providers; state laws and laws outside the United States that require drug manufacturers to report information
related to clinical trials, or information related to payments and other transfers of value to physicians and other healthcare providers or marketing
expenditures; state laws that restrict the ability of manufacturers to offer co-pay support to patients for certain prescription drugs; and state laws
and local ordinances that require identification or licensing of sales representatives.

Efforts to ensure that our collaborations with third parties, and our business generally, will comply with applicable United States and Canadian healthcare
laws and regulations will involve substantial costs. It is possible that governmental authorities will conclude that our business practices may not comply
with current or future statutes, regulations or case law involving applicable fraud and abuse or other healthcare laws and regulations. If our operations are
found to be in violation of any of these laws or any other governmental laws and regulations that may apply to us, we may be subject to significant civil,
criminal  and  administrative  penalties,  damages,  fines,  imprisonment,  exclusion  of  products  from  government  funded  healthcare  programs,  contractual
damages,  reputational  harm,  disgorgement,  curtailment  or  restricting  of  our  operations,  any  of  which  could  substantially  disrupt  our  operations  and
diminish our profits and future earnings. If any of the physicians or other providers or entities with whom we expect to do business is found not to be in
compliance  with  applicable  laws,  they  may  be  subject  to  criminal,  civil  or  administrative  sanctions,  including  exclusions  from  government  funded
healthcare programs. The risk of our being found in violation of these laws is increased by the fact that many of them have not been fully interpreted by the
regulatory authorities or the courts, and their provisions are open to a variety of interpretations.

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Failure to comply with the United States Foreign Corrupt Practices Act (“FCPA”), and potentially other global anti-corruption and anti-bribery laws
such as the Canadian Corruption of Foreign Public Officials Act, could subject us to penalties and other adverse consequences.

We are subject to the FCPA, and potentially other applicable domestic or foreign anti-corruption or anti-bribery laws, which generally prohibit companies
from engaging in bribery or other prohibited payments to foreign officials for the purpose of obtaining or retaining business and requires companies to
maintain accurate books and records and internal controls, including at foreign-controlled subsidiaries.

Compliance  with  these  anti-corruption  laws  and  anti-bribery  laws  may  be  expensive  and  difficult,  particularly  in  countries  in  which  corruption  is  a
recognized problem. In addition, these laws present particular challenges in the pharmaceutical industry, because, in many countries, hospitals are operated
by the government, and physicians and other hospital employees are considered to be foreign officials. Certain payments to hospitals in connection with
clinical trials and other work have been deemed to be improper payments to governmental officials and have led to FCPA enforcement actions.

We can make no assurance that our employees or other agents will not engage in prohibited conduct under our policies and procedures and anti-corruption
laws and anti-bribery laws such as FCPA for which we might be held responsible. If our employees or other agents are found to have engaged in such
practices, we could suffer severe penalties and other consequences that may have a material adverse effect on our business, financial condition and results
of operations.

Risks Related to Our Dependence on Third Parties

We depend on our license agreement with Alnylam for the commercialization of ONPATTRO™ (Patisiran).

In  2012,  we  entered  into  a  license  agreement  with  Alnylam  that  entitles  Alnylam  to  develop  and  commercialize  products  with  our  LNP  technology.
Alnylam received FDA approval in August 2018 and launched ONPATTRO immediately upon approval. We are entitled to low to mid-single-digit royalty
payments escalating based on sales performance and received our first royalty payment in the fourth quarter of 2018. In July 2019, we sold this royalty
entitlement  to  OMERS,  the  defined  benefit  pension  plan  for  municipal  employees  based  in  the  Province  of  Ontario,  Canada,  effective  as  of  January  1,
2019, for $20 million in gross proceeds before advisory fees. OMERS will retain this royalty entitlement until it has received $30 million in royalties, at
which point 100% of this royalty entitlement on future global net sales of ONPATTRO will revert to us. From the inception of the royalty sale through
December 31, 2022, an aggregate of $18.9 million of royalties have been collected by OMERS. The possibility and timing of any possible reversion of the
royalty entitlement is affected by many factors including:

• Alnylam’s and its distributors’ and sublicensees’ ability to effectively market and sell ONPATTRO in each country where sold;
•
•
•
•
•

the manner of sale, whether directly by Alnylam or by sublicensees or distributors, and the terms of sublicensing and distribution agreements;
the amount and timing of sales of Alnylam in each country;
regulatory approvals, appropriate labeling, and desirable pricing, insurance coverage and reimbursement;
competition; and
commencement of marketing in additional countries.

If Alnylam’s commercialization of ONPATTRO does not continue to be successful, the royalty entitlement may never revert back to us.

We expect to depend in part on our licensing agreements for a significant portion of our revenues for the foreseeable future and to develop, conduct
clinical trials with, obtain regulatory approvals for, and manufacture, market and sell some of our product candidates. If these licensing agreements
are unsuccessful, or anticipated milestone or royalty payments are not received, our business could be materially adversely affected.

We expect that we will depend in part on our licensing agreements with Alnylam, Qilu and Gritstone to provide revenue to partially fund our operations,
especially in the near term. Furthermore, our strategy is to enter into various additional arrangements with corporate and academic collaborators, licensors,
licensees and others for the research, development, clinical testing, manufacturing, marketing and commercialization of our product candidates or other
products based upon our

47

technology. We may be unable to continue to establish such licensing agreements, and any licensing agreements we do establish may be unsuccessful, or
we may not receive milestone payments or royalties as anticipated.

Should any licensing partner fail to develop or ultimately successfully commercialize any of the product candidates or technology to which it has obtained
rights,  our  business  may  be  adversely  affected.  In  addition,  once  initiated,  there  can  be  no  assurance  that  any  of  these  licensing  agreements  will  be
continued or result in successfully commercialized products. Failure of a licensing partner to continue funding any particular program could delay or halt
the development or commercialization of any products arising out of such program. In addition, there can be no assurance that the licensing partners will
not pursue alternative technologies or develop alternative products either on their own or in collaboration with others, including our competitors.

We are dependent on our collaboration and licensing partners and, therefore, are subject to the efforts of these parties and our ability to successfully
collaborate with them.

We have entered into a number of clinical collaboration agreements, including with Assembly and Vaccitech. We are responsible for managing the clinical
trial under the collaboration agreement with Vaccitech, while Assembly is responsible for managing the clinical trials under the collaboration we have with
them. The success of our collaborations depend on not only our efforts, but also on the efforts of our counterparties. Because we are not responsible for
managing  the  clinical  trials  with  Assembly,  the  success  of  those  collaborations  also  depend  on  whether  Assembly  is  successful  in  performance  of  its
activities, to the extent it is responsible for performance of collaboration activities. Additionally, these counterparties could change their strategic focus or
pursue alternative technologies, which could materially and adversely affect our business. Similarly, we are dependent on X-Chem and Proteros pursuant to
our  discovery  and  research  agreement  to  work  toward  the  development  of  pan-coronavirus  agents  to  treat  COVID-19  and  potential  future  coronavirus
outbreaks.

For  example,  through  the  clinical  collaboration  agreement  with  Assembly  that  we  entered  into  in  August  2020,  Assembly  conducted  a  clinical  trial
evaluating AB-729 in combination with its first-generation HBV core inhibitor (capsid inhibitor) candidate VBR and standard-of-care NA therapy for the
treatment of cHBV in HBeAg negative patients with cHBV to evaluate the safety, pharmacokinetics, and antiviral activity of the triple combination of AB-
729,  VBR,  and  an  NA  (n=32).  In  July  2022,  Assembly  announced  its  plans  to  discontinue  development  of  VBR.  Despite  this,  in  consultation  with
Assembly, we continued this Phase 2a proof-of-concept clinical trial in order to fully and accurately assess the results.

In addition, if we have a dispute or enter into litigation with any of these parties in the future, it could delay development programs, distract management
from other business activities, and generate substantial expense.

We will depend on Qilu for the development and commercialization of AB-729 in China, Hong Kong, Macau and Taiwan.

In December 2021, we entered into the License Agreement with Qilu, pursuant to which we granted Qilu an exclusive (except as to certain retained rights),
sublicensable, royalty-bearing license, under certain intellectual property owned by us, to develop, manufacture and commercialize AB-729 in the Territory.
The timing and amount of any milestone and royalty payments we may receive under the License Agreement will depend, in part, on the efforts of Qilu.
We  will  depend  on  Qilu  to  comply  with  all  applicable  laws  relative  to  the  development  and  commercialization  of  AB-729  in  the  Territory.  Under  the
License Agreement, Qilu is required to use commercially reasonable efforts to develop, seek regulatory approval for, and commercialize at least one AB-
729 product candidate in the Territory. Any failure by Qilu to use such commercially reasonable efforts could have a material adverse impact on financial
results  and  operations.  Additionally,  if  Qilu  were  to  violate,  or  was  alleged  to  have  violated,  any  laws  or  regulations  during  the  performance  of  its
obligations  to  us,  we  could  suffer  financial  and  reputational  harm  or  other  negative  outcomes.  Any  termination,  breach  or  expiration  of  the  License
Agreement  could  also  have  a  material  adverse  impact  on  our  business  by  reducing  or  eliminating  the  potential  for  us  to  receive  milestone  and  royalty
payments.  If  that  were  to  occur,  we  may  be  required  to  devote  additional  time,  costs  and  attention  to  pursue  the  manufacture,  development  and
commercialization  of  AB-729  in  the  Territory.  In  certain  situations,  Qilu  has  the  ability  to  terminate  the  License  Agreement  and  retain  all  rights  to
manufacture, develop and commercialize AB-729 in the Territory with no obligation to make any additional milestone or royalty payments to us.

48

If conflicts arise between our collaboration or licensing partners and us, our collaboration or licensing partners may act in their best interest and not in
our best interest, which could adversely affect our business.

Conflicts  may  arise  with  our  collaboration  or  licensing  partners,  including  Alnylam,  Qilu,  Gritstone,  Assembly  and  Vaccitech  if  they  pursue  alternative
therapies for the diseases that we have targeted or develop alternative products either on their own or in collaboration with others. Competing products,
either developed by our present collaboration or licensing partners or any future partners or to which our present partners or any future partners have rights,
may result in development delays or the withdrawal of their support for one or more of our product candidates.

Additionally, conflicts may arise if there is a dispute about the progress of, or other activities related to, the clinical development of a product candidate, the
achievement and payment of a milestone amount, the payment of royalties or the ownership of intellectual property that is developed during the course of
the  collaborative  arrangement.  Similarly,  the  parties  to  a  licensing  agreement  may  disagree  as  to  which  party  owns  newly  developed  products.  If  an
agreement is terminated as a result of a dispute and before we have realized the benefits of the collaboration or licensing arrangement, our reputation could
be harmed and we might not obtain revenues that we anticipated receiving.

We  rely  on  third  parties  to  conduct  our  clinical  trials,  and  if  they  fail  to  fulfill  their  obligations,  perform  services  in  a  satisfactory  manner,  and/or
comply with applicable legal or regulatory requirements, our development plans may be adversely affected.

We rely on independent clinical investigators, CROs and other third-party service providers to assist us in managing, monitoring and otherwise carrying out
our  clinical  trials.  We  have  contracted  with,  and  we  plan  to  continue  to  contract  with,  certain  third  parties  to  provide  certain  services,  including  site
selection,  enrollment,  monitoring  and  data  management.  Although  we  depend  heavily  on  these  parties  and  have  contractual  agreements  governing  their
activities, we do not control them and therefore, we cannot be assured that these third parties will adequately perform all of their contractual obligations to
us. If our third-party service providers cannot adequately fulfill their obligations to us on a timely and satisfactory basis or if the quality or accuracy of our
clinical  trial  data  is  compromised  due  to  failure  to  adhere  to  our  protocols  or  regulatory  requirements,  or  if  such  third  parties  otherwise  fail  to  meet
deadlines or follow legal or regulatory requirements, our development plans may be delayed or terminated.

If any of our relationships with these third-parties terminate, we may not be able to enter into arrangements with alternative third parties on commercially
reasonable terms or at all. Switching or adding additional third-party service providers involves additional cost and requires management time and focus. In
addition,  there  is  a  natural  transition  period  when  a  new  third-party  service  provider  begins  work.  As  a  result,  delays  may  occur,  which  can  materially
impact our ability to meet our desired development timelines.

We  rely  exclusively  on  third  parties  to  formulate  and  manufacture  our  product  candidates,  which  exposes  us  to  a  number  of  risks  that  may  delay
development, regulatory approval and commercialization of our products or result in higher product costs.

We have limited experience in drug formulation or manufacturing and we lack the resources and expertise to formulate or manufacture our own product
candidates  internally.  Therefore,  we  rely  on,  and  expect  to  continue  to  rely  on,  third-party  expertise  to  support  us  in  this  area.  We  have  entered  into
contracts with third-party manufacturers to manufacture, supply, store and distribute supplies of our product candidates for our clinical trials. If any of our
product  candidates  receive  FDA  approval,  we  expect  to  rely  on  third-party  contractors  to  manufacture  our  products.  We  have  no  current  plans  to  build
internal manufacturing capacity for any product candidate, and we have no long-term supply arrangements.

Our reliance on third-party manufacturers exposes us to potential risks, such as the following:

• we  may  be  unable  to  contract  with  third-party  manufacturers  on  acceptable  terms,  or  at  all,  because  the  number  of  potential  manufacturers  is
limited.  Potential  manufacturers  of  any  product  candidate  that  is  approved  will  be  subject  to  FDA  compliance  inspections  and  any  new
manufacturer would have to be qualified to produce our products;

•

our third-party manufacturers might be unable to formulate and manufacture our product candidates and products in the volume and of the quality
required to meet our clinical and commercial needs, if any;

49

•

•

•

•

our third-party manufacturers may not perform as agreed or may not remain in the contract manufacturing business for the time required to supply
our clinical trials through completion or to successfully produce, store and distribute our commercial products, if approved;
drug  manufacturers  are  subject  to  ongoing  periodic  unannounced  inspection  by  the  FDA  and  other  government  agencies  to  ensure  compliance
with  cGMP  and  other  government  regulations  and  corresponding  foreign  standards.  We  do  not  have  control  over  third-party  manufacturers’
compliance with these regulations and standards, but we may ultimately be responsible for any of their failures;
if any third-party manufacturer makes improvements in the manufacturing process for our product candidates, we may not own, or may have to
share, the intellectual property rights to such improvements; and
a third-party manufacturer may gain knowledge from working with us that could be used to supply one of our competitors with a product that
competes with ours.

Each  of  these  risks  could  delay  or  have  other  adverse  impacts  on  our  clinical  trials  and  the  approval  and  commercialization  of  our  product  candidates,
potentially resulting in higher costs, reduced revenues or both.

Risks Related to Our Intellectual Property

Other companies or organizations may assert patent rights that prevent us from developing or commercializing our products.

RNAi, capsid inhibitors and RNA destabilizer, as well as our other novel HBV assets, have generated many different patent applications from organizations
and  individuals  seeking  to  obtain  patents  in  the  field.  These  applications  claim  many  different  methods,  compositions  and  processes  relating  to  the
discovery, development and commercialization of these therapeutic products. It is likely that there could be litigation and other proceedings, such as inter
partes review and opposition proceedings in various patent offices, relating to patent rights in RNAi, capsid inhibitors, RNA destabilizer and other small
molecule compounds targeted at HBV. We are aware of patents and patent applications owned by third parties that may in the future be alleged by such
third parties to cover the use of one or more of our products. We may need to acquire or obtain a license from such third parties to any such issued patents
to  market  or  sell  any  such  products,  which  may  not  be  available  on  commercially  acceptable  terms  or  at  all.  If  such  third  parties  obtain  valid  and
enforceable patents and successfully prove infringement of an approved Arbutus product, and we are not able to acquire such issued patents or negotiate a
license  on  acceptable  terms,  and  if  such  approved  Arbutus  product  is  determined  to  infringe  any  such  issued  patents,  then  we  may  be  forced  to  pay
royalties,  damages  and  costs,  or  we  may  be  prevented  from  commercializing  such  approved  Arbutus  product  altogether,  which  could  have  a  material
adverse impact on our business.

Our patents and patent applications may be challenged and may be found to be invalid, which could adversely affect our business.

Certain United States, Canadian and international patents and patent applications we own involve complex legal and factual questions for which important
legal principles are largely unresolved. For example, no consistent policy has emerged for the breadth of biotechnology patent claims that are granted by
the USPTO or enforced by the United States federal courts. In addition, the coverage claimed in a patent application can be significantly reduced before a
patent is issued. Also, we face at least the following intellectual property risks:

•
•
•
•
•

some or all patent applications may not result in the issuance of a patent;
patents issued to us may not provide us with any competitive advantages;
patents could be challenged by third parties;
competitors may find ways to design around our patents; and
competitors could independently develop products which duplicate our products.

A number of industry competitors and institutions have developed technologies, filed patent applications or received patents on various technologies that
may be related to or affect our business. Some of these technologies, applications or patents may conflict with our technologies or patent applications. Such
conflict could limit the scope of the patents, if any, that we may be able to obtain or result in the denial of our patent applications. In addition, we could
incur substantial costs in filing suits against others to have such patents declared invalid. As publication of discoveries in the scientific or patent literature
often lags behind

50

actual discoveries, we cannot be certain we or any licensor was the first creator of inventions covered by pending patent applications or that we or such
licensor was the first to file patent applications for such inventions. Any future proceedings could result in substantial costs, even if the eventual outcomes
are favorable. There can be no assurance that our patents, if issued, will be held valid or enforceable by a court or that a competitor’s technology or product
would be found to infringe such patents.

We may incur substantial costs as a result of litigation or other proceedings relating to patent and other intellectual property rights which could have a
material  adverse  effect  on  our  business,  financial  condition  and  results  of  operations  and  could  cause  the  market  value  of  our  common  shares  to
decline.

There has been significant litigation in the biotechnology industry over contractual obligations, patents and other proprietary rights, and we may become
involved  in  various  types  of  litigation  that  arise  from  time  to  time.  Involvement  in  litigation  could  consume  a  substantial  portion  of  our  resources,
regardless of the outcome of the litigation. Counterparties in litigation may be better able to sustain the costs of litigation because they have substantially
greater resources. If claims against us are successful, in addition to any potential liability for damages, we could be required to obtain a license, grant cross-
licenses,  and  pay  substantial  milestones  or  royalties  in  order  to  continue  to  develop,  manufacture  or  market  the  affected  products.  Involvement  and
continuation of involvement in litigation may result in significant and unsustainable expense, and divert management’s attention from ongoing business
concerns and interfere with our normal operations. Litigation is also inherently uncertain with respect to the time and expenses associated therewith, and
involves risks and uncertainties in the litigation process itself, such as discovery of new evidence or acceptance of unanticipated or novel legal theories,
changes in interpretation of the law due to decisions in other cases, the inherent difficulty in predicting the decisions of judges and juries and the possibility
of appeals. Ultimately we could be prevented from commercializing a product or be forced to cease some aspect of our business operations as a result of
claims of patent infringement or violation of other intellectual property rights and the costs associated with litigation, which could have a material adverse
effect on our business, financial condition, and operating results and could cause the market value of our common shares to decline.

Confidentiality  agreements  with  employees  and  others,  including  collaborators,  may  not  adequately  prevent  disclosure  of  trade  secrets  and  other
proprietary information.

Much of our know-how and technology may constitute trade secrets. There can be no assurance, however, that we will be able to meaningfully protect our
trade secrets. In order to protect our proprietary technology and processes, we rely in part on confidentiality agreements with our collaborators, employees,
vendors, consultants, outside scientific collaborators and sponsored researchers, and other advisors. These agreements offer only limited protection, and as
such  may  not  effectively  prevent  disclosure  of  confidential  information  and  also  may  not  provide  an  adequate  remedy  in  the  event  of  unauthorized
disclosure of confidential information. In addition, others may independently discover trade secrets and proprietary information, and in such cases we could
not assert any trade secret rights against such party. Costly and time consuming litigation could continue to be necessary to enforce and determine the scope
of our proprietary rights, and failure to obtain or maintain trade secret protection could adversely affect our competitive business position.

Risks Related to the Ownership of our Common Shares

The concentration of common share ownership will likely limit the ability of the other shareholders to influence corporate matters.

As of February 28, 2023, executive officers, directors, five percent or greater shareholders, and their respective affiliated entities beneficially owned, in the
aggregate, approximately 26% of our outstanding common shares.

Entities  associated  with  Roivant  Sciences  Ltd.  (“Roivant”)  collectively  held  as  a  group  approximately  25%  of  our  outstanding  common  shares  as  of
February 28, 2023.

As a result, Roivant can significantly influence the outcome of matters requiring shareholder approval, including the election of directors, amendments of
our organizational documents, or approval of any merger, sale of assets or other major corporate transaction. This may prevent or discourage unsolicited
acquisition proposals or offers for our common shares that you may feel are in your best interest. The interests of Roivant may not always coincide with
your interests or the interests of other

51

shareholders and they may act in a manner that advances their best interests and not necessarily those of other shareholders, including seeking a premium
value for their common shares. These actions might affect the prevailing market price for our common shares. In addition, Roivant and certain of our other
principal shareholders that have held their shares for several years may be more interested in selling our company to an acquirer than other investors, or
they may want us to pursue strategies that deviate from the interests of other shareholders. Such concentration of ownership control may also:

•
•
•

delay, defer or prevent a change in control;
entrench our management and/or the board of directors; or
impede a merger, consolidation, takeover or other business combination involving us that other shareholders may desire.

We are incorporated in Canada, with our assets located both in Canada and the United States, with the result that it may be difficult for investors to
enforce judgments obtained against us or some of our officers.

We  are  incorporated  under  the  laws  of  the  Province  of  British  Columbia  and  some  of  our  assets  are  located  outside  the  United  States.  While  we  have
appointed National Registered Agents, Inc. as our agent for service of process to effect service of process within the United States upon us, it may not be
possible  for  you  to  enforce  against  us  or  our  insiders  in  the  United  States,  judgments  obtained  in  United  States  courts  based  upon  the  civil  liability
provisions of the United States federal securities laws or other laws of the United States. In addition, there is doubt as to whether original action could be
brought  in  Canada  against  us  or  our  directors  or  officers  based  solely  upon  United  States  federal  or  state  securities  laws  and  as  to  the  enforceability  in
Canadian courts of judgments of United States courts obtained in actions based upon the civil liability provisions of United States federal or state securities
laws.

Conversely, all of our directors and officers reside outside Canada, and the majority of our physical assets are also located outside Canada. While we have
appointed Farris LLP as our agent for service of process in Canada, it may not be possible for you to enforce in Canada against our assets or those directors
and officers residing outside Canada, judgments obtained in Canadian courts based upon the civil liability provisions of the Canadian securities laws or
other laws of Canada.

If we are deemed to be a “passive foreign investment company” for the current or any future taxable year, investors who are subject to United States
federal taxation would likely suffer materially adverse United States federal income tax consequences.

We generally will be a “passive foreign investment company” under the meaning of Section 1297 of the Code (a “PFIC”) if (a) 75% or more of our gross
income  is  “passive  income”  (generally,  dividends,  interest,  rents,  royalties,  and  gains  from  the  disposition  of  assets  producing  passive  income)  in  any
taxable year, or (b) if at least 50% or more of the quarterly average value of our assets produce, or are held for the production of, passive income in any
taxable year. We have determined that we have not been a PFIC for the three taxable years ended December 31, 2022, however recent changes to Treasury
regulations under the Code have made this determination more challenging for us, and we cannot provide any assurances that we will not become a PFIC in
the future. If we are a PFIC for any taxable year during which a United States person holds our common shares, it would likely result in materially adverse
United States federal income tax consequences for such United States person, including, but not limited to, any gain from the sale of our common shares
would  be  taxed  as  ordinary  income,  as  opposed  to  capital  gain,  and  such  gain  and  certain  distributions  on  our  common  shares  would  be  subject  to  an
interest charge, except in certain circumstances. It may be possible for United States persons to fully or partially mitigate such tax consequences by making
a “qualifying electing fund election,” as defined in the Code (a “QEF Election”), but although we have provided this information in the past, there is no
requirement that we do so.

Our articles and certain Canadian laws could delay or deter a change of control.

Our preferred shares are available for issuance from time to time at the discretion of our board of directors, without shareholder approval. Our articles allow
our board, without shareholder approval, to determine the special rights to be attached to our preferred shares, and such rights may be superior to those of
our common shares.

In addition, limitations on the ability to acquire and hold our common shares may be imposed by the Competition Act in Canada. This legislation permits
the Commissioner of Competition of Canada to review any acquisition of a significant interest in us. This legislation grants the Commissioner jurisdiction
to challenge such an acquisition before the Canadian Competition

52

Tribunal if the Commissioner believes that it would, or would be likely to, result in a substantial lessening or prevention of competition in any market in
Canada. The Investment Canada Act subjects an acquisition of control of a Canadian-company by a non-Canadian to government review if the value of our
assets,  as  calculated  pursuant  to  the  legislation,  exceeds  a  threshold  amount.  A  reviewable  acquisition  may  not  proceed  unless  the  relevant  minister  is
satisfied that the investment is likely to result in a net benefit to Canada. Any of the foregoing could prevent or delay a change of control and may deprive
or limit strategic opportunities for our shareholders to sell their shares.

53

General Risk Factors

If we are unable to attract and retain qualified key management, scientific staff, consultants and advisors, our ability to implement our business plan
may be adversely affected.

We depend upon our senior executive officers as well as key scientific, management and other personnel. The competition for qualified personnel in the
biotechnology field is intense. We rely heavily on our ability to attract and retain qualified managerial, scientific and technical staff. The loss of the service
of any of the members of our senior management, including William H. Collier, our President and Chief Executive Officer, and Michael J. Sofia, our Chief
Scientific  Officer,  may  adversely  affect  our  ability  to  develop  our  technology,  add  to  our  pipeline,  advance  our  product  candidates  and  manage  our
operations. We do not carry key person life insurance on any of our employees.

We  rely  on  consultants  and  advisors,  including  scientific  and  clinical  advisors,  to  assist  us  in  formulating  our  research  and  development  and
commercialization  strategy.  Our  consultants  and  advisors  may  be  employed  by  other  entities  and  may  have  commitments  under  consulting  or  advisory
contracts with those entities that may limit their availability to us. If we are unable to continue to attract and retain highly qualified personnel, our ability to
develop and commercialize our product candidates will be limited.

We could face liability from our controlled use of hazardous and radioactive materials in our research and development processes.

We use certain radioactive materials, biological materials and chemicals, including organic solvents, acids and gases stored under pressure, in our research
and  development  activities.  Our  use  of  radioactive  materials  is  regulated  by  the  United  States  Nuclear  Regulatory  Commission  and  Pennsylvania
Department of Environmental Protection for the possession, transfer, import, export, use, storage, handling and disposal of radioactive materials. Our use of
biological materials and chemicals, including the use, manufacture, storage, handling and disposal of such materials and certain waste products is regulated
by a number of federal, state and local laws and regulations. Although we believe that our safety procedures for handling such materials comply with the
standards prescribed by such laws and regulations, the risk of accidental contamination or injury from these materials cannot be completely eliminated. In
the event of such an accident, we could be held liable for any damages that result or penalized with fines, and any such liability could exceed our resources.
We are not specifically insured with respect to this liability.

Our business, reputation, and operations could suffer in the event of information technology system failures, such as a cybersecurity breach.

We are increasingly dependent on sophisticated software applications and computing infrastructure to conduct critical operations. We depend on both our
own  systems,  networks,  and  technology  as  well  as  the  systems,  networks  and  technology  of  our  contractors,  consultants,  vendors  and  other  business
partners. Disruption, degradation, or manipulation of systems, networks or technology through intentional or accidental means could materially adversely
impact key business processes. Despite the implementation of security measures, our systems, networks and technology and those of our contractors and
consultants are vulnerable to damage from computer viruses (including ransomware), cybersecurity breaches and other forms of unauthorized access, as
well as natural disasters, terrorism, war, telecommunication and electrical failures, cyberattacks, phishing or other fraudulent schemes, persons inside our
organization, or persons with access to systems inside our organization or those with whom we do business. The risk of a cyberattack or other cybersecurity
incidents  has  generally  increased  as  the  number,  intensity  and  sophistication  of  attempted  attacks  and  intrusions  from  around  the  world  have  increased.
Although to date the cybersecurity incidents we have experienced have not resulted in any material losses, such events impacting either our own systems,
networks  and  technology,  or  those  of  our  contractors,  consultants,  vendors,  or  other  business  partners  could  threaten  the  confidentiality,  integrity  and
availability of regulated personal information, confidential information or intellectual property. This could result in the modification of critical data, the loss
of Company funds and/or the failure or interruption of critical operations. For example, the loss of pre-clinical trial data or data from completed or ongoing
clinical trials for our product candidates could result in delays in our regulatory filings and development efforts and significantly increase our costs. There
can be no assurance that our efforts to protect data and systems will prevent service interruption or the loss of critical or sensitive information from our or
third party providers’ databases or systems. Additionally, while we have implemented security measures that we believe are appropriate and continue to
enhance cybersecurity protections, a regulator could deem our

54

security  measures  not  to  be  appropriate  given  the  lack  of  prescriptive  measures  in  certain  data  protection  laws.  To  the  extent  that  any  disruption  or
cybersecurity  incident  results  or  appears  to  result  in  such  interruption  or  loss,  we  could  incur  material  financial,  legal,  business  or  reputational  harm,
including  regulatory  fines,  penalties  or  intervention,  or  claims  by  third  parties  that  we  have  breached  privacy-  or  confidentiality-related  obligations.
Furthermore, the development of our product candidates could be delayed, and our insurance may not provide any or adequate coverage of any such losses.

We may acquire other assets or businesses, or form strategic alliances or collaborations or make investments in other companies or technologies that
could harm our financial condition, results of operations or cash flows, dilute our shareholders’ ownership, incur debt or cause us to incur significant
expense.

As  part  of  our  business  strategy,  we  may  pursue  acquisitions  of  assets  or  businesses,  or  strategic  alliances  or  collaborations,  to  expand  our  existing
technologies and operations. We may not identify or complete these transactions in a timely manner, on a cost-effective basis, or at all, and we may not
realize the anticipated benefits of any such transaction, any of which could have a detrimental effect on our financial condition, results of operations or cash
flows.  We  may  not  be  able  to  find  suitable  acquisition  candidates,  and  if  we  make  any  acquisitions,  we  may  not  be  able  to  integrate  these  acquisitions
successfully  into  our  existing  business  and  we  may  incur  debt  or  assume  unknown  or  contingent  liabilities  in  connection  therewith.  Integration  of  an
acquired company or assets may also disrupt ongoing operations, require the hiring of additional personnel and the implementation of additional internal
systems and infrastructure, especially the acquisition of commercial assets, and require management resources that would otherwise focus on developing
our existing business. We may not be able to find suitable collaboration partners or identify other investment opportunities, and we may experience losses
related to any such investments.

To finance any acquisitions or collaborations, we may choose to issue debt or equity securities as consideration. Any such issuance of shares would dilute
the ownership of our shareholders. If the price of our common shares is low or volatile, we may not be able to acquire other assets or businesses or fund a
transaction using our equity securities as consideration. Alternatively, it may be necessary for us to raise additional capital for acquisitions through public
or private financings. Additional capital may not be available on terms that are favorable to us, or at all.

55

Item 1B. Unresolved Staff Comments

There are currently no unresolved staff comments.

Item 2. Properties

Since  November  1,  2016,  we  have  had  a  lease  agreement  for  our  headquarters  at  701  Veterans  Circle,  Warminster,  Pennsylvania.  The  building  has
approximately 35,000 square feet of laboratory facilities and office space. The lease expires on April 30, 2027. We also have the option of extending the
lease for two further five-year terms.

From January 2019 through June 2021, we leased approximately 8,500 square feet of office space at 626 Jacksonville Rd, Warminster, Pennsylvania. In
mid-2021, we amended the contract to relet a portion of the leased space and , as the initial three-year lease term was set to expire on December 31, 2021,
we extended the lease through December 31, 2022. On August 31, 2022, we terminated the lease early in full.

We believe that the total space available to us under our current leases will meet our needs for the foreseeable future and that additional space would be
available to us on commercially reasonable terms if required.

Item 3. Legal Proceedings

Patent Infringement Litigation vs. Moderna

On February 28, 2022, we and Genevant filed a lawsuit in the U.S. District Court for the District of Delaware against Moderna, Inc. and a Moderna affiliate
(collectively, “Moderna”) seeking damages for infringement of U.S. Patent Nos. 8,058,069, 8,492,359, 8,822,668, 9,364,435, 9,504,651, and 11,141,378 in
the manufacture and sale of MRNA-1273, Moderna’s vaccine for COVID-19. The patents relate to nucleic acid-lipid particles and lipid vesicles, as well as
compositions  and  methods  for  their  use.  The  lawsuit  does  not  seek  an  injunction  or  otherwise  seek  to  impede  the  sale,  manufacture  or  distribution  of
MRNA-1273. However, the Company seeks fair compensation for Moderna’s use of its patented technology that was developed with great effort and at
great expense, without which Moderna’s COVID-19 vaccine would not have been successful. On May 6, 2022, Moderna filed a partial motion to dismiss
the claims “relating to Moderna’s sale and provision of COVID-19 vaccine doses to the U.S. Government.” On November 2, 2022, the Court issued an
Order denying Moderna’s motion. On November 30, 2022, Moderna filed its Answer to the Complaint and Counterclaims. Arbutus and Genevant filed
their  Answer  to  Moderna’s  Counterclaims  on  December  21,  2022.  On  February  14,  2023,  the  U.S.  Dept.  of  Justice  filed  a  Statement  of  Interest  in  the
action. On February 16, 2023, the Court held an Initial Pretrial Conference after which it issued an Order, dated February 16, 2023, ordering that within 14
days of the issuance of the Order, the parties and the U.S. Government are to submit letters regarding the impact of the Governments’ Statement of Interest
on the scheduling of the matter.

Acuitas Declaratory Judgment Lawsuit

On March 18, 2022, Acuitas Therapeutics Inc. (“Acuitas”) filed a lawsuit against us and Genevant in the U.S. District Court for the Southern District of
New  York,  asking  the  court  to  enter  declaratory  judgment  that  Arbutus  patent  Nos.  8,058,069,  8,492,359,  8,822,668,  9,006,417,  9,364,435,  9,404,127,
9,504,651,  9,518,272,  and  11,141,378  do  not  infringe  Pfizer  and  BioNTech’s  COVID-19  vaccine,  COMIRNATY,  which  uses  an  mRNA  lipid  provided,
under  license,  by  Acuitas.  Acuitas  also  seeks  a  declaration  that  each  of  the  listed  patents  is  invalid.  On  June  24,  2022,  we  and  Genevant  sought  a  pre-
motion conference concerning our anticipated motion to dismiss all of Acuitas’ claims due to lack of subject matter jurisdiction. The request for a pre-
motion conference was granted, but the case was subsequently re-assigned to a new judge who entered an order directing: (i) Acuitas to inform the court
whether it intended to file an amended complaint; (ii) that Acuitas must file any amended complaint by a certain date; and (iii) that if Acuitas did not file an
amended complaint, we and Genevant must file our motion to dismiss by a certain date. Acuitas filed its amended complaint on September 6, 2022. On
October  4,  2022,  we  and  Genevant  filed  our  motion  to  dismiss  the  Acuitas  action  for  lack  of  subject  matter  jurisdiction  based  on  the  lack  of  a  case  or
controversy. Acuitas filed its opposition to the motion to dismiss on November 1, 2022, and we and Genevant filed our reply brief on November 16, 2022.
The motion is now fully briefed. No case schedule is yet in place.

56

 
University of British Columbia

Certain early work on lipid nanoparticle delivery systems and related inventions was undertaken at the University of British Columbia (“UBC”), as well as
by us that was subsequently assigned to UBC. These inventions are licensed to us by UBC under a license agreement, initially entered into in 1998 and as
amended in 2001, 2006 and 2007. We granted sublicenses under the UBC license to certain third parties, including Alnylam. In November 2014, UBC filed
a demand for arbitration against us which alleged entitlement to unpaid royalties. In August 2019, the arbitrator issued his decision for the second phase of
the arbitration, awarding UBC $5.9 million, which included interest of approximately $2.6 million. We paid the $5.9 million award to UBC in September
2019 and paid an additional $0.2 million award for costs and attorneys’ fees in March 2021, and this matter is now fully resolved.

On December 18, 2020, UBC delivered to us a notice of arbitration alleging that under its cross license with us, it is due royalties of $2.0 million plus
interest arising from our sale to OMERS of part of our royalty interest on future global net sales of ONPATTRO, currently being sold by Alnylam. Oral
hearings for this matter were held in April 2022 and, on July 11, 2022, the arbitrator issued his decision fully dismissing UBC’s claim for royalties. As a
result, no payments are owed to UBC. In September 2022, the arbitrator awarded the Company $0.5 million for reimbursement of costs and attorneys’ fees,
which the Company received from UBC in October 2022. This matter is now fully resolved.

We are also involved with various legal matters arising in the ordinary course of business. We make provisions for liabilities when it is both probable that a
liability has been incurred and the amount of the loss can be reasonably estimated. Such provisions are reviewed at least quarterly and adjusted to reflect
the  impact  of  any  settlement  negotiations,  judicial  and  administrative  rulings,  advice  of  legal  counsel,  and  other  information  and  events  pertaining  to  a
particular case. Litigation is inherently unpredictable. Although the ultimate resolution of these various matters cannot be determined at this time, we do
not believe that such matters, individually or in the aggregate, will have a material adverse effect on our consolidated results of operations, cash flows, or
financial condition.

Item 4. Mine Safety Disclosures

Not applicable.

57

PART II

Item 5. Market for Registrant’s Common Equity, Related Stockholder Matters and Issuer Purchases of Equity Securities

Our common shares trade on the Nasdaq Global Select Market under the symbol “ABUS” following our name change to Arbutus Biopharma Corporation
on July 31, 2015. As of February 28, 2023, there were 103 registered holders of common shares and 162,570,989 common shares issued and outstanding.

Securities Authorized for Issuance under Equity Compensation Plans

Information regarding securities authorized for issuance under equity compensation plans is incorporated by reference into the information in Part III, Item
12 of this Form 10-K.

Recent Sales of Unregistered Securities

Other  than  as  previously  disclosed  on  our  Current  Reports  on  Form  8-K  or  Quarterly  Reports  on  Form  10-Q,  we  did  not  issue  any  unregistered  equity
securities during the twelve months ended December 31, 2022.

Purchases of Equity Securities by the Issuer and Affiliated Purchasers

We did not repurchase any of our equity securities during the year ended December 31, 2022.

Item 6. Reserved

58

 
Item 7. Management’s Discussion and Analysis of Financial Condition and Results of Operations

Overview

Arbutus  Biopharma  Corporation  (“Arbutus”,  the  “Company”,  “we”,  “us”,  and  “our”)  is  a  clinical-stage  biopharmaceutical  company  leveraging  its
extensive virology expertise to develop novel therapeutics that target specific viral diseases. Our current focus areas include Hepatitis B virus (“HBV”),
SARS-CoV-2, and other coronaviruses. To address HBV, we are developing an RNA interference (“RNAi”) therapeutic, an oral PD-L1 inhibitor, and an
oral RNA destabilizer to potentially identify a combination regimen with the aim of providing a functional cure for patients with chronic HBV infection
(“cHBV”) by suppressing viral replication, reducing surface antigen and reawakening the immune system. We believe our lead compound, AB-729, is the
only RNAi therapeutic with evidence of immune re-awakening. AB-729 is currently being evaluated in multiple phase 2 clinical trials. We also have an
ongoing drug discovery and development program directed to identifying novel, orally active agents for treating coronaviruses, including SARS-CoV-2,
where we have nominated a compound and have begun IND-enabling pre-clinical studies. In addition, we are also exploring oncology applications for our
internal PD-L1 portfolio.

Our product pipeline consists of the following programs:

AB-729, our proprietary subcutaneously-delivered RNAi therapeutic product candidate that suppresses HBsAg expression, which is thought to be a key
prerequisite  to  enable  reawakening  of  a  patient’s  immune  system  to  respond  to  HBV,  is  currently  in  two  Phase  2a  proof-of-concept  clinical  trials  in
combination  with  other  agents  with  potentially  complementary  mechanisms  of  action  and  we  are  continuing  to  follow  patients  from  our  Phase  1a/1b
clinical trial (“AB-729-001”). Preliminary data from AB-729-001 has shown that treatment with AB-729 resulted in meaningful declines in HBsAg while
being  well  tolerated  with  no  serious  adverse  events  (SAEs)  noted  after  both  single  and  repeat  dosing.  Preliminary  data  also  suggests  that  long-term
suppression  of  HBsAg  with  AB-729  results  in  increased  HBV-specific  immune  response.  The  clinical  data  for  AB-729  continues  to  support  its
development as a potential cornerstone agent for the treatment of cHBV infection.

AB-101 is our oral PD-L1 inhibitor that has the potential to reawaken patients’ HBV-specific immune response by inhibiting PD-L1. Preclinical data in an
HBV mouse model was presented at the 2022 AASLD Liver Meeting showing that combination treatment with AB-101 and an HBV-targeting GalNAc-
siRNA  agent  resulted  in  activation  and  increased  frequency  of  HBV-specific  T-cells  and  greater  anti-HBsAg  antibody  production.  This  favorable
preclinical  profile  supports  further  development  of  AB-101  as  a  therapeutic  modality  for  cHBV  treatment.  We  are  also  exploring  potential  oncology
applications for our internal PD-L1 portfolio.

59

AB-161 is our next-generation oral HBV specific RNA destabilizer. We have conducted extensive non-clinical safety evaluations with AB-161 that gives
us confidence in this molecule’s ability to circumvent the peripheral neuropathy findings seen in non-clinical safety studies with our first-generation oral
RNA destabilizer, AB-452. We recently presented preclinical data at the Discovery on Target Conference showing that AB-161 reduced HBV RNA and
HBsAg in multiple preclinical models, with favorable liver centricity and lack of observed peripheral neuropathy.

AB-343 is our lead candidate that inhibits the SARS-CoV-2 nsp5 M . We also intend to nominate a nsp12 clinical candidate and initiate IND-enabling
studies in the second half of 2023. An nsp12 viral polymerase could potentially be combined with AB-343 to achieve better patient treatment outcomes and
for use in prophylactic settings. 

pro

COVID-19 Impact

We continue to monitor the effects of COVID-19, which has caused significant disruptions around the world. Measures implemented around the world in
attempts to slow the spread of COVID-19 have had, and will likely continue to have, a major impact on clinical development, at least in the near-term,
including shortages and delays in the supply chain, and prohibitions in certain countries on enrolling patients in new clinical trials. While we have been
able to progress with our clinical and pre-clinical activities to date, it is not possible to predict if the COVID-19 pandemic will materially impact our plans
and timelines in the future.

Collaborations and Royalty Entitlements

Qilu Pharmaceutical Co., Ltd. (“Qilu”)

In December 2021, we entered into a technology transfer and license agreement (the “License Agreement”) with Qilu, pursuant to which we granted Qilu a
sublicensable, royalty-bearing license, under certain intellectual property owned by us, which is non-exclusive as to development and manufacturing and
exclusive with respect to commercialization of AB-729, including pharmaceutical products that include AB-729, for the treatment or prevention of hepatitis
B in China, Hong Kong, Macau and Taiwan (the “Territory”).

In partial consideration for the rights granted by us, Qilu paid us a one-time upfront cash payment of $40 million on January 5, 2022 and agreed to pay us
milestone payments totaling up to $245 million, net of withholding taxes, upon the achievement of certain technology transfer, development, regulatory and
commercialization milestones. Qilu also agreed to pay us double digit royalties into the low twenties percent based upon annual net sales of AB-729 in the
Territory. The royalties are payable on a product-by-product and region-by-region basis, subject to certain limitations.

Qilu is responsible for all costs related to developing, obtaining regulatory approval for, and commercializing AB-729 for the treatment or prevention of
hepatitis B in the Territory. Qilu is required to use commercially reasonable efforts to develop, seek regulatory approval for, and commercialize at least one
AB-729  product  candidate  in  the  Territory.  A  joint  development  committee  has  been  established  between  us  and  Qilu  to  coordinate  and  review  the
development, manufacturing and commercialization plans. Both parties also have entered into a supply agreement and related quality agreement pursuant
to which we will manufacture or have manufactured and supply Qilu with all quantities of AB-729 necessary for Qilu to develop and commercialize in the
Territory until we have completed manufacturing technology transfer to Qilu and Qilu has received all approvals required for it or its designated contract
manufacturing organization to manufacture AB-729 in the Territory.

Concurrent with the execution of the License Agreement, we entered into a Share Purchase Agreement (the “Share Purchase Agreement”) with Anchor
Life Limited, a company established pursuant to the applicable laws and regulations of Hong Kong and an affiliate of Qilu (the “Investor”), pursuant to
which the Investor purchased 3,579,952 of our common shares, without par value (the “Common Shares”), at a purchase price of USD $4.19 per share,
which was a 15% premium on the thirty-day average closing price of the Common Shares as of the close of trading on December 10, 2021 (the “Share
Transaction”). We received $15.0 million of gross proceeds from the Share Transaction on January 6, 2022. The Common Shares sold to the Investor in the
Share  Transaction  represented  approximately  2.5%  of  the  Common  Shares  outstanding  immediately  prior  to  the  execution  of  the  Share  Purchase
Agreement.

60

Alnylam Pharmaceuticals, Inc. and Acuitas Therapeutics, Inc

We  have  a  royalty  entitlement  on  ONPATTRO®  (Patisiran)  (“ONPATTRO”),  a  drug  developed  by  Alnylam  Pharmaceuticals,  Inc.  (“Alnylam”)  under  a
license agreement with us that incorporates our lipid nanoparticle delivery (“LNP”) technology. In July 2019, we received $20 million in gross proceeds
before advisory fees from the sale of this royalty interest to Ontario Municipal Employees Retirement System (“OMERS”), effective as of January 1, 2019.
The royalty interest will revert back to us after OMERS receives $30 million in royalty payments from Alnylam. We also have rights to a second, lower
royalty interest on global net sales of ONPATTRO originating from a settlement agreement and subsequent license agreement with Acuitas Therapeutics,
Inc. (“Acuitas”). The royalty entitlement from Acuitas has been retained by us and was not part of the royalty entitlement sale to OMERS.

Genevant Sciences, Ltd.

As of December 31, 2022, we owned approximately 16% of the common equity of Genevant Sciences Ltd. (“Genevant”), a company we launched with
Roivant  Sciences,  Ltd.  and  to  which  we  licensed  rights  to  our  lipid  nanoparticle  ("LNP")  and  ligand  conjugate  delivery  platforms  for  RNA-based
applications outside of HBV, except to the extent certain rights had already been licensed to other third parties (the “Genevant License”). We retained all
rights to our LNP and conjugate delivery platforms for HBV. Under the Genevant License, as amended, if a third party sublicensee of intellectual property
licensed by Genevant from us commercializes a sublicensed product, we become entitled to receive a specified percentage of certain revenue that may be
received by Genevant for such sublicense, including royalties, commercial milestones and other sales-related revenue, or, if less, tiered low single-digit
royalties on net sales of the sublicensed product. The specified percentage is 20% in the case of a mere sublicense (i.e., naked sublicense) by Genevant
without additional contribution and 14% in the case of a bona fide collaboration with Genevant.

Additionally,  if  Genevant  receives  proceeds  from  an  action  for  infringement  by  any  third  parties  of  our  intellectual  property  licensed  to  Genevant,  we
would be entitled to receive, after deduction of litigation costs, 20% of the proceeds received by Genevant or, if less, tiered low single-digit royalties on net
sales of the infringing product (inclusive of the proceeds from litigation or settlement, which would be treated as net sales).

Refer  to  “Item  1.  Business.”  and  Note  9  of  the  Consolidated  Financial  Statements  for  a  discussion  of  our  clinical  collaborations  and  other  royalty
entitlements.

61

CRITICAL ACCOUNTING POLICIES AND ESTIMATES

The accounting for our contingent consideration and our License Agreement with Qilu are significant accounting policies that we believe are critical in
fully understanding and evaluating our financial results. These accounting policies require us to make certain estimates and assumptions. We believe that
the estimates and assumptions upon which we rely are reasonable, based upon information available to us at the time that these estimates and assumptions
are made.  Actual results may differ from our estimates. Our critical accounting estimates affect the calculation of our net income or loss.

Contingent Consideration

In connection with the acquisition of Enantigen Therapeutics, Inc. (“Enantigen”) in October 2014, we have obligations to make potential future payments
of  up  to  $102.5  million  upon  the  achievement  of  certain  commercial  milestones. The  sales  milestones  are  tied  to  the  first  commercial  sales  by  us  of  a
product  indicated  for  the  treatment  of  cHBV.  These  potential  contingent  payments  are  recorded  as  a  liability  and  remeasured  to  fair  value  as  of  each
reporting  date.    In  assessing  the  fair  value  of  the  liability,  significant  judgments  are  required  to  be  made  by  management  to  estimate  the  probability  of
program success, the timing and extent of future product sales, appropriate discount rates, and other estimates and assumptions that could materially affect
the determination of fair value.

In order to estimate the probability of program success, we evaluate the status and progress of our clinical trials with our lead product candidate, AB-729,
in comparison to actual historical success rates for other clinical trials. We update our assumptions related to probability of success as AB-729 advances
through clinical trials. For the timing and extent of future product sales, we also consider the status and progress of AB-729, future revenue forecasts and
other macroeconomic indicators that forecast market conditions. The discount rate at which we calculate the present value of our potential future liability is
based on consideration of market-comparative data, market-based discount rates, and company-specific risk premiums.

As  assumptions  related  to  the  probability  of  program  success  and  timing  and  amount  of  potential  future  product  sales  are  highly  uncertain  due  to  the
unpredictable nature of product development, we assessed the sensitivity of the fair value measurement to changes in assumptions, and determined that
changes within a reasonable range would not result in a materially different assessment of fair value.  

Revenue from collaborations and licenses

We generate revenue primarily through collaboration agreements and license agreements. Such agreements may require us to deliver various rights and/or
services, including intellectual property rights or licenses and research, development and manufacturing services. Under such agreements, we are generally
eligible to receive non-refundable upfront payments, funding for research, development and manufacturing services, milestone payments, and royalties.

Our collaboration agreements fall under the scope of ASC Topic 808, Collaborative Arrangements, (“ASC 808”) when both parties are active participants
in the arrangement and are exposed to significant risks and rewards. For certain arrangements under the scope of ASC 808, we analogize to ASC 606 for
some aspects, including for the delivery of a good or service (i.e., a unit of account).

ASC 606, Revenue From Contracts with Customers (“ASC 606”) requires an entity to recognize the amount of revenue to which it expects to be entitled
for the transfer of promised goods or services to customers under a five-step model: (i) identify contract(s) with a customer; (ii) identify the performance
obligations in the contract; (iii) determine the transaction price; (iv) allocate the transaction price to the performance obligations in the contract; and (v)
recognize revenue when or as a performance obligation is satisfied.

In contracts where we have more than one performance obligation to provide its customer with goods or services, each performance obligation is evaluated
to determine whether it is distinct based on whether (i) the customer can benefit from the good or service either on its own or together with other resources
that are readily available and (ii) the good or service is separately identifiable from other promises in the contract. The consideration under the contract is
then allocated between the distinct performance obligations based on their respective relative stand-alone selling prices. The estimated stand-alone selling
price of each deliverable reflects our best estimate of what the selling price would be if the deliverable was regularly sold on a

62

stand-alone basis and is determined by reference to market rates for the good or service when sold to others or by using an adjusted market assessment
approach if the selling price on a stand-alone basis is not available.

The consideration allocated to each distinct performance obligation is recognized as revenue when control is transferred to the customer for the related
goods  or  services.  Consideration  associated  with  at-risk  substantive  performance  milestones,  including  sales-based  milestones,  is  recognized  as  revenue
when  it  is  probable  that  a  significant  reversal  of  the  cumulative  revenue  recognized  will  not  occur.  Sales-based  royalties  received  in  connection  with
licenses of intellectual property are subject to a specific exception in the revenue standards, whereby the consideration is not included in the transaction
price and recognized in revenue until the customer’s subsequent sales or usages occur.

Prior  to  recognizing  revenue,  we  make  estimates  of  the  transaction  price,  including  variable  consideration  that  is  subject  to  a  constraint.  Amounts  of
variable consideration are included in the transaction price to the extent that it is probable that a significant reversal in the amount of cumulative revenue
recognized will not occur and when the uncertainty associated with the variable consideration is subsequently resolved. These estimates are re-assessed
each reporting period as required.

For performance obligations satisfied over time, we estimate the efforts needed to complete the performance obligation and recognize revenue by
measuring the progress towards complete satisfaction of the performance obligation using an input measure.

Management may be required to exercise considerable judgment in estimating revenue to be recognized. Judgment is required in identifying performance
obligations, estimating the transaction price, estimating the stand-alone selling price of identified performance obligations, and estimating the progress
towards satisfaction of performance obligations.

63

RESULTS OF OPERATIONS

The following summarizes our results of operations for the year ended December 31, 2022 compared to the year ended December 31, 2021:
Year Ended December 31,

Revenue
Operating expenses

Loss from operations

Other income (loss)

Loss before income taxes

Income tax expense

Net loss

Dividend accretion of convertible preferred shares

Net loss attributable to common shares

2022

2021

(in thousands)

$

$

39,019  $
104,475 
(65,456)
444 
(65,012)
(4,444)
(69,456)
— 
(69,456) $

10,988 
84,510 
(73,522)
(2,725)
(76,247)
— 
(76,247)
(12,139)
(88,386)

For  the  fiscal  year  ended  December  31,  2022,  our  net  loss  attributable  to  common  shares  was  $69.5  million,  or  a  loss  of  $0.46  per  basic  and  diluted
common share, as compared to a net loss of $88.4 million, or a loss of $0.83 per basic and diluted common share, for the year ended December 31, 2021.

Revenue

Revenue for the years ended December 31, 2022 and 2021 is summarized in the following table:

Revenue from collaborations and licenses

Royalties from sales of Onpattro
Qilu Pharmaceutical Co., Ltd.
Other milestone and royalty payments

Non-cash royalty revenue

Royalties from sales of Onpattro
Total revenue

Year ended December 31,

2022

2021

(in thousands, except percentages)

$

$

5,316 
26,015 
35 

7,653 
39,019 

14 % $
67 %
— %

20 %
101 % $

4,675 
— 
205 

6,108 
10,988 

43 %
— %
2 %

56 %
100 %

Revenue consists mainly of royalties received from other companies for sales of products that utilize our licensed technologies.

Total  revenue  increased  $28.0  million  for  the  year  ended  December  31,  2022  compared  to  2021,  due  primarily  to  $26.0  million  in  license  revenue
recognized related to our progress towards the satisfaction of our performance obligations with respect to our technology transfer and licensing agreement
with  Qilu,  which  closed  in  January  2022,  as  well  as  a  $2.2  million  increase  in  license  royalty  revenue  from  Alnylam  and  Acuitas  due  to  the  growth  of
Alnylam’s sales of ONPATTRO.

The royalty interest for ONPATTRO from Alnylam was sold to OMERS, effective as of January 1, 2019, for $20 million in gross proceeds before advisory
fees. OMERS will retain this entitlement until it has received $30 million in royalties, at which point 100% of such royalty interest on future global net
sales of ONPATTRO will revert back to us. OMERS has assumed the risk of collecting up to $30 million of future royalty payments from Alnylam and we
are not obligated to reimburse OMERS if they fail to collect any such future royalties. During the term of this agreement, we recognize non-cash royalty
revenue related to the sales of ONPATTRO. From the inception of the royalty sale through December 31, 2022, we have recorded an aggregate

64

 
 
 
 
of $18.9 million of non-cash royalty revenue for royalties earned by OMERS. The royalty interest for ONPATTRO from Acuitas was not part of the royalty
sale to OMERS and we have retained the rights to receive those royalties. Revenue contracts are described in more detail in “Item 1. Business.”

Operating expenses

Operating expenses for the years ended December 31, 2022 and 2021 are summarized in the following table:

Research and development
General and administrative
Change in fair value of contingent consideration

Total operating expenses

Research and development

Year ended December 31,

2022

2021

(in thousands, except percentages)

$

$

84,408 
17,834 
2,233 
104,475 

81 % $
17 %
2 %
100 % $

65,502 
17,136 
1,872 
84,510 

78 %
20 %
2 %
100 %

Research  and  development  expenses  consist  primarily  of  personnel  expenses,  fees  paid  to  clinical  research  organizations  and  contract  manufacturers,
consumables and materials, consulting, and other third party expenses to support our clinical and pre-clinical activities, as well as a portion of stock-based
compensation and general overhead costs.

Research and development expenses increased $18.9 million in 2022 compared to 2021 due primarily to an increase in expenses for our ongoing AB-729
Phase 2a clinical trials, an increase in expenses for our early-stage development programs, including AB-101 and AB-161, and an increase in compensation
costs due to hiring several new employees for our research and development team in early 2022, partially offset by a decrease in expenses for our AB-836
Phase 1a/1b clinical trial, which we discontinued during the fourth quarter of 2022.

A  significant  portion  of  our  research  and  development  expenses  are  not  tracked  by  project,  as  they  benefit  multiple  projects  or  our  overall  technology
platform.

General and administrative

General  and  administrative  expenses  increased  $0.7  million  in  2022  compared  to  2021,  due  primarily  to  increases  in  employee  compensation  costs  and
non-cash stock-based compensation expense.

Change in fair value of contingent consideration

In October 2014, Arbutus Inc., our wholly-owned subsidiary, acquired all of the outstanding shares of Enantigen pursuant to a stock purchase agreement.
The amount paid to Enantigen’s selling shareholders could be up to an additional $102.5 million in sales performance milestones in connection with the
sale  of  the  first  commercialized  product  by  us  for  the  treatment  of  HBV,  regardless  of  whether  such  product  is  based  upon  assets  acquired  under  this
agreement, and a low single-digit royalty on net sales of such first commercialized HBV product, up to a maximum royalty payment of $1.0 million.

In  general,  increases  in  the  fair  value  of  the  contingent  consideration  are  related  to  the  progress  of  our  programs  as  they  get  closer  to  triggering  these
contingent payments. In 2022 and 2021, the fair value of our contingent consideration liability increased $2.2 million and $1.9 million, respectively, related
to  fair  value  adjustments  for  the  passage  of  time  and  the  progression  of  our  programs  through  clinical  trials  and  our  assessment  of  the  probability  of
commercialization.

65

 
 
 
 
 
Other income (losses)

Other income (losses) for the years ended December 31, 2022 and 2021 are summarized in the following table:

Interest income
Interest expense
Foreign exchange (loss) gain
Total other income (loss)

Interest income

2022

2,192 
(1,726)
(22)
444 

Year ended December 31,

2021

(in thousands, except percentages)

494 % $
(389)%
(5)%
100 % $

127 
(2,857)
5 
(2,725)

$

$

(5)%
105 %
— %
100 %

Interest income increased $2.1 million in 2022 compared to 2021 due primarily to a general increase in market interest rates related to our investments in
marketable securities.

Interest expense

Interest expense decreased $1.1 million in 2022 compared to 2021 due primarily to a decrease in the non-cash amortization of discount and issuance costs
related to the sale of a portion of our ONPATTRO royalty interest to OMERS in July 2019.

Dividend accretion of convertible preferred shares

Dividend accretion of convertible preferred shares decreased to zero in 2022 compared to $12.1 million in 2021. The dividend accretion on the convertible
preferred shares previously held by Roivant was equal to 8.75% per annum, compounded annually. All convertible preferred shares mandatorily converted
into 22,833,922 common shares on October 18, 2021.

Income tax expense

Income tax expense for the years ended December 31, 2022 and 2021 are summarized in the following table:

Income tax expense

$

4,444 

100 % $

Year ended December 31,

2022

(in thousands, except percentages)

2021

— 

— %

We  recognized  income  tax  expense  of  $4.4  million  during  2022  for  withholding  taxes  paid  to  the  Chinese  taxing  authority  by  Qilu  on  our  behalf  in
connection with the upfront license fee Qilu paid us.

66

 
LIQUIDITY AND CAPITAL RESOURCES 

Since our incorporation, we have financed our operations through the sales of equity, debt, revenues from research and development collaborations and
licenses  with  corporate  partners,  a  royalty  monetization,  interest  income  on  funds  available  for  investment,  and  government  contracts,  grants  and  tax
credits.

As of December 31, 2022, we had cash and cash equivalents of $30.8 million and investments in marketable securities of $153.5 million, totaling $184.3
million. We had no outstanding debt as of December 31, 2022.

Sources of Liquidity

Sale Agreement

We  have  an  Open  Market  Sale  Agreement
  with  Jefferies  dated  December  20,  2018,  as  amended  by  Amendment  No.  1,  dated  December  20,  2019,
Amendment No. 2, dated August 7, 2020 and Amendment No. 3, dated March 4, 2021 (as amended, the “Sale Agreement”), under which we may offer and
sell common shares, from time to time.

SM

On  December  23,  2019,  we  filed  a  shelf  registration  statement  on  Form  S-3  with  the  SEC  (File  No.  333-235674)  and  accompanying  base  prospectus,
declared  effective  by  the  SEC  on  January  10,  2020  (the  “January  2020  Registration  Statement”),  for  the  offer  and  sale  of  up  to  $150  million  of  our
securities.

On August 28, 2020, we filed a shelf registration statement on Form S-3 with the SEC (File No. 333-248467) and accompanying base prospectus, declared
effective by the SEC on October 22, 2020 (the “October 2020 Registration Statement”), for the offer and sale of up to $200 million of our securities. On
March 4, 2021, we filed a prospectus supplement with the SEC in connection with the offering of up to an additional $75.0 million of our common shares
pursuant  to  the  Sale  Agreement  under  the  October  2020  Registration  Statement,  which  we  fully  utilized  during  2021.  On  October  8,  2021,  we  filed  a
prospectus supplement with the SEC for the offer and sale of up to an additional $75.0 million of our common shares pursuant to the Sale Agreement under
the October 2020 Registration Statement.

On  November  4,  2021,  we  filed  a  shelf  registration  statement  on  Form  S-3  with  the  SEC  (File  No.  333-248467)  and  accompanying  base  prospectus,
declared effective by the SEC on November 18, 2021 (the “November 2021 Registration Statement”), for the offer and sale of up to $250 million of our
securities.

On March 3, 2022, we filed a prospectus supplement with the SEC (the “March 2022 Prospectus Supplement”) for the offer and sale of up to an additional
$100.0  million  of  our  common  shares  pursuant  to  the  Sale  Agreement  under:  (i)  the  January  2020  Registration  Statement;  (ii)  the  October  2020
Registration Statement; and (iii) the November 2021 Registration Statement.

During  the  years  ended  December  31,  2022  and  2021,  we  issued  8,645,426  and  31,571,036  common  shares,  respectively,  under  the  Sale  Agreement
resulting in net proceeds of approximately $20.3 million and $134.7 million, respectively. As of December 31, 2022, we had an aggregate of $131.1 million
remaining available under the October 2021 Prospectus Supplement and the March 2022 Prospectus Supplement.

Royalty Entitlements

Additionally, we have a royalty entitlement on ONPATTRO, a drug developed by Alnylam that incorporates our LNP technology and was approved by the
FDA and the EMA during the third quarter of 2018 and was launched by Alnylam immediately upon approval in the United States. In July 2019, we sold a
portion of this royalty interest to OMERS, effective as

67

of January 1, 2019, for $20 million in gross proceeds before advisory fees. OMERS will retain this entitlement until it has received $30 million in royalties,
at which point 100% of such royalty interest on future global net sales of ONPATTRO will revert to us. OMERS has assumed the risk of collecting up to
$30 million of future royalty payments from Alnylam and Arbutus is not obligated to reimburse OMERS if they fail to collect any such future royalties.
From  the  inception  of  the  royalty  sale  through  December  31,  2022,  we  have  recorded  an  aggregate  of  $18.9  million  of  non-cash  royalty  revenue  for
royalties earned by OMERS. If this royalty entitlement reverts to us, it has the potential to provide an active royalty stream or to be otherwise monetized
again in full or in part. In addition to the royalty from the Alnylam LNP license agreement, we are also receiving a second, lower royalty interest on global
net  sales  of  ONPATTRO  originating  from  a  settlement  agreement  and  subsequent  license  agreement  with  Acuitas.  The  royalty  from  Acuitas  has  been
retained by us and was not part of the royalty sale to OMERS.

In December 2021, we entered into a technology transfer and exclusive licensing agreement with Qilu pursuant to which we granted Qilu an exclusive
(with  certain  exceptions),  sublicensable,  royalty-bearing  license,  under  certain  intellectual  property  owned  by  us,  to  develop,  manufacture  and
commercialize AB-729 for the treatment or prevention of cHBV in the Territory. In partial consideration for the rights granted by us, Qilu paid us a one-
time upfront cash payment of $40 million and made an equity investment of $15.0 million, both received in January 2022, and agreed to pay us milestone
payments  totaling  up  to  $245  million,  net  of  withholding  taxes,  upon  the  achievement  of  certain  technology  transfer,  development,  regulatory  and
commercialization milestones. Qilu also agreed to pay us double digit royalties into the low twenties percent based upon annual net sales of AB-729 in the
Territory.

Cash requirements

We believe that our $184.3 million of cash, cash equivalents and investments in marketable securities as of December 31, 2022 will be sufficient to fund
our operations into the fourth quarter of 2024 based on our expectation of a net cash burn between $95.0 million and $100.0 million in 2023. In the future,
substantial additional funds will be required to continue with the active development of our pipeline products and technologies. In particular, our funding
needs may vary depending on a number of factors including:

•

•

•

•

•

•

•

•

•

•

•

•

the effects of the COVID-19 pandemic on our business, the medical community and the global economy;

revenue  earned  from  our  legacy  collaborative  partnerships  and  licensing  agreements,  including  potential  royalty  payments  from  Alnylam’s
ONPATTRO;

revenue earned from ongoing collaborative partnerships, including milestone and royalty payments;

the potential requirement to make milestone payments related to our legacy agreements;

the extent to which we continue the development of our product candidates, add new product candidates to our pipeline, or form collaborative
relationships or licensing arrangements to advance our product candidates;

delays in the development of our product candidates due to pre-clinical and clinical findings;

our decisions to in-license or acquire additional products, product candidates or technology for development;

our ability to attract and retain development or commercialization partners, and their effectiveness in carrying out the development and ultimate
commercialization of one or more of our product candidates;

whether  batches  of  product  candidates  that  we  manufacture  fail  to  meet  specifications  resulting  in  clinical  trial  delays  and  investigational  and
remanufacturing costs;

the decisions, and the timing of decisions, made by health regulatory agencies regarding our technology and product candidates;

competing products, product candidates and technological and market developments; and

costs associated with prosecuting and enforcing our patent claims and other intellectual property rights, including litigation and arbitration arising
in the course of our business activities.

68

We intend to seek funding to maintain and advance our business from a variety of sources including public or private equity or debt financing, potential
monetization transactions, collaborative or licensing arrangements with pharmaceutical companies and government grants and contracts. There can be no
assurance that funding will be available at all or on acceptable terms to permit further development of our research and development programs. Further, the
COVID-19 pandemic has also led to severe disruption and volatility in the global capital markets, which could increase our cost of capital and adversely
affect our ability to access the capital markets in the future.

If adequate funding is not available, we may be required to delay, reduce or eliminate one or more of our research or development programs or reduce
expenses associated with our non-core activities. We may need to obtain funds through arrangements with collaborators or others that may require us to
relinquish most or all of our rights to product candidates at an earlier stage of development or on less favorable terms than we would otherwise seek if we
were better funded. Insufficient financing may also mean failing to prosecute our patents or relinquishing rights to some of our technologies that we would
otherwise develop or commercialize.

Cash Flows

The following table summarizes our cash flow activities for the periods indicated:

Net loss
Non-cash items
Change in deferred license revenue
Net change in operating items

Net cash used in operating activities
Net cash used in investing activities

Issuance of common shares pursuant to Share Purchase Agreement
Issuance of common shares pursuant to exercise of ESPP

Net cash provided by other financing activities
Net cash provided by financing activities

Effect of foreign exchange rate changes on cash and cash equivalents

(Decrease) increase in cash and cash equivalents

Cash and cash equivalents, beginning of period
Cash and cash equivalents, end of period

Year ended December 31,

2022

2021

(in thousands)

$

$

$

$

(69,456) $
4,857 
22,455 
6,788 
(35,356) $
(74,942)
10,973 
395 
20,446 
31,814 
(22)
(78,506) $
109,282 
30,776  $

(76,247)
7,790 
— 
925 
(67,532)
(12,678)
— 
461 
136,775 
137,236 
5 
57,031 
52,251 
109,282 

Net cash used in operating activities in 2022 decreased $32.2 million compared to 2021 due primarily to a January 2022 upfront cash payment of $40.0
million from Qilu in connection with the License Agreement and a $4.0 million premium paid by Qilu as part of their $15.0 million equity investment.
These cash inflows were offset by $79.4 million of cash used in operations.

Net cash used in investing activities in 2022 increased by $62.3 million compared to 2021 due primarily to the timing of acquisitions and maturities of
investments in marketable securities.

Net  cash  provided  by  financing  activities  in  2022  decreased  $105.4  million  compared  to  2021.  Cash  provided  by  financing  activities  in  2022  consisted
primarily of $20.3 million of proceeds from sales of common shares under the Sale Agreement and $11.0 million for the fair value of shares purchased by
Qilu as part of their $15.0 million equity investment, of which the remaining $4.0 million was a premium paid by Qilu on the equity investment and was
allocated to deferred revenue. Cash

69

 
 
provided by financing activities in 2021 consisted primarily of $134.7 million of proceeds from sales of common shares under the Sale Agreement.

RECENT ACCOUNTING PRONOUNCEMENTS

From time to time, new accounting pronouncements are issued by the Financial Accounting Standards Board or other standard setting bodies that we adopt
as of the specified effective date. Unless otherwise discussed, we believe that the impact of recently issued standards that are not yet effective will not have
a material impact on our financial position or results of operations upon adoption.

Please refer to note 2 to our consolidated financial statements included in Part II, Item 8, “Financial Statements and Supplementary Data,” of this Annual
Report on Form 10-K for a description of recent accounting pronouncements applicable to our business. 

Item 7A. Quantitative and Qualitative Disclosures about Market Risk

Not applicable.

70

 
Item 8. Financial Statements and Supplementary Data

INDEX TO CONSOLIDATED FINANCIAL STATEMENTS

Report of Ernst & Young, LLP, Independent Registered Public Accounting Firm - PCAOB ID: 42
Consolidated Balance Sheets at December 31, 2022 and 2021
Consolidated Statements of Operations and Comprehensive Loss for the Years Ended December 31, 2022 and 2021
Consolidated Statements of Stockholders’ Equity for the Years Ended December 31, 2022 and 2021
Consolidated Statements of Cash Flows for the Years Ended December 31, 2022 and 2021
Notes to Consolidated Financial Statements

Page
72
76
77
78
79
80

71

 
Report of Independent Registered Public Accounting Firm

To the Shareholders and the Board of Directors of Arbutus Biopharma Corporation

Opinion on the Financial Statements

We have audited the accompanying consolidated balance sheets of Arbutus Biopharma Corporation (the Company) as of December 31, 2022 and 2021, and
the related consolidated statements of operations and comprehensive loss, stockholders' equity, and cash flows for the years then ended, and the related
notes (collectively referred to as the “consolidated financial statements”). In our opinion, the consolidated financial statements present fairly, in all material
respects, the financial position of the Company at December 31, 2022 and 2021, and the results of its operations and its cash flows for the years then ended,
in conformity with U.S. generally accepted accounting principles.

Basis for Opinion

These financial statements are the responsibility of the Company’s management. Our responsibility is to express an opinion on the Company’s financial
statements  based  on  our  audits.  We  are  a  public  accounting  firm  registered  with  the  Public  Company  Accounting  Oversight  Board  (United  States)
(PCAOB) and are required to be independent with respect to the Company in accordance with the U.S. federal securities laws and the applicable rules and
regulations of the Securities and Exchange Commission and the PCAOB.

We conducted our audits in accordance with the standards of the PCAOB. Those standards require that we plan and perform the audit to obtain reasonable
assurance about whether the financial statements are free of material misstatement, whether due to error or fraud. The Company is not required to have, nor
were we engaged to perform, an audit of its internal control over financial reporting. As part of our audits we are required to obtain an understanding of
internal  control  over  financial  reporting  but  not  for  the  purpose  of  expressing  an  opinion  on  the  effectiveness  of  the  Company's  internal  control  over
financial reporting. Accordingly, we express no such opinion.

Our  audits  included  performing  procedures  to  assess  the  risks  of  material  misstatement  of  the  financial  statements,  whether  due  to  error  or  fraud,  and
performing procedures that respond to those risks. Such procedures included examining, on a test basis, evidence regarding the amounts and disclosures in
the  financial  statements.  Our  audits  also  included  evaluating  the  accounting  principles  used  and  significant  estimates  made  by  management,  as  well  as
evaluating the overall presentation of the financial statements. We believe that our audits provide a reasonable basis for our opinion.

Critical Audit Matters

The  critical  audit  matters  communicated  below  are  matters  arising  from  the  current  period  audit  of  the  financial  statements  that  were  communicated  or
required  to  be  communicated  to  the  audit  committee  and  that:  (1)  relate  to  accounts  or  disclosures  that  are  material  to  the  financial  statements  and  (2)
involved our especially challenging, subjective or complex judgments. The communication of critical audit matters do not alter in any way our opinion on
the consolidated financial statements, taken as a whole, and we are not, by communicating the critical audit matters below, providing separate opinions on
the critical audit matters or on the accounts or disclosures to which they relate.

72

Description of the Matter

Valuation of contingent consideration liability
As discussed in Note 10 to the consolidated financial statements, the Company’s contingent consideration liability, which
consists of sales-based milestones and royalties, resulting from the acquisition of Enantigen in 2014, is remeasured to its
estimated  fair  value  each  reporting  period.  As  of  December  31,  2022,  the  contingent  consideration  liability  was  $7.5
million.

Auditing the valuation of the contingent consideration liability was complex and highly judgmental due to the significant
estimation  required  in  determining  the  fair  value.  In  particular,  the  fair  value  estimate  was  sensitive  to  significant
assumptions such as the probability of successfully commercializing a treatment for the hepatitis B virus, the timing and
amount  of  future  revenues  related  to  commercial  sales,  and  the  discount  rate.  These  assumptions  are  affected  by
expectations  about  future  industry,  regulatory,  market  or  economic  conditions  and  are  forward-looking  and  inherently
uncertain.

How We Addressed the
Matter in Our Audit

To  test  the  estimated  fair  value  of  the  contingent  consideration  liability,  we  performed  audit  procedures  that  included,
among  others,  assessing  the  terms  of  the  arrangement,  evaluating  the  methodology  used,  and  testing  the  significant
assumptions  discussed  above  used  by  the  Company  in  its  analysis.  We  also  compared  the  significant  assumptions  to
current industry, market and economic trends to corroborate the Company’s estimates and performed sensitivity analyses
of significant assumptions to evaluate the changes in the contingent consideration liability that would result from changes
in the significant assumptions. We also involved our valuation specialists to assist us in testing the discount rate. 

73

 
 
Description of the Matter

How We Addressed the
Matter in Our Audit

Collaboration and License Agreement with Qilu

As  discussed  in  Note  11  to  the  consolidated  financial  statements,  in  December  2021,  the  Company  entered  into  a
technology  transfer  and  license  agreement  with  Qilu  Pharmaceuticals  Co.,  Ltd.  (Qilu).  Under  the  agreement,  the
Company  granted  Qilu  an  exclusive  right  to  develop  and  commercialize  AB-729  for  the  treatment  and  prevention  of
hepatitis B in the People's Republic of China, Hong Kong, Macau, and Taiwan. The Company agreed to provide clinical
supply of the licensed product to Qilu until the Company has completed the manufacturing technology transfer to Qilu.
The Company received a $40.0 million up-front payment, net of withholding taxes, during 2022 in connection with this
arrangement  and  is  also  eligible  to  receive  additional  development  and  regulatory  milestone  payments,  sales-based
milestones  and  royalties  as  well  as  additional  payments  for  clinical  supply  under  the  arrangement.  The  Company
identified two commitments under the arrangement: (i) rights to develop, use, sell, have sold, offer for sale and import any
product  comprised  of  Licensed  Product  (the  “Qilu  License”)  and  (ii)  drug  supply  obligations  and  manufacturing
technology  transfer  (the  “Manufacturing  Obligations”).  The  Company  determined  that  these  two  commitments  are  not
distinct performance obligations for purposes of recognizing revenue as the manufacturing process is highly specialized
and Qilu would not be able to benefit from the Qilu License without the Company’s involvement in the manufacturing
activities  until  the  transfer  of  the  manufacturing  know-how  is  complete.  As  such,  the  Company  combined  these
commitments into one performance obligation to which the transaction price is allocated and recognized over time using
an inputs method based on labor hours expended by the Company on its Manufacturing Obligations. 

Auditing  the  Company's  revenue  recognition  for  the  Qilu  collaboration  and  license  agreement  was  challenging,  as
significant  judgment  was  required  to  apply  the  authoritative  accounting  guidance  to  the  arrangement.  The  Company
exercised significant judgment in determining the revenue recognition for this arrangement, including as it relates to the
identification of performance obligations, as well as estimating the total number of labor hours that will be expended to
complete the Manufacturing Obligations.

Our audit procedures to test the Company's determination of revenue recognition for the Qilu collaboration and license
agreement included, among others, reading the contractual agreement, testing management's identification of significant
terms  for  completeness,  including  identification  of  performance  obligations,  and  evaluating  the  appropriateness  of
management's  application  of  authoritative  guidance  and  existing  accounting  policies.  We  also  discussed  the  judgments
inherent  in  the  Company's  determination  of  revenue  recognition,  including  the  identification  of  the  performance
obligations and estimating the total number of expected hours required to complete the Manufacturing Obligations, with
research  and  development  personnel  responsible  for  overseeing  the  satisfaction  of  the  Company's  Manufacturing
Obligations.  We  also  tested  a  sample  of  actual  hours  expended  during  2022  on  the  Manufacturing  Obligations  and
performed  a  lookback  analysis,  comparing  the  total  actual  hours  expended  throughout  the  year  to  the  total  number  of
future expected hours as of December 31, 2022, based on the progress to date and the nature of the future activities to be
performed.

/s/ Ernst & Young LLP

We have served as the Company's auditor since 2019.

Philadelphia, Pennsylvania

74

 
 
March 2, 2023

75

ARBUTUS BIOPHARMA CORPORATION

Consolidated Balance Sheets

(Expressed in thousands of US Dollars, except share and per share amounts)

Assets
Current assets:

Cash and cash equivalents
Investments in marketable securities, current
Accounts receivable
Prepaid expenses and other current assets

Total current assets
Property and equipment, net of accumulated depreciation
Investments in marketable securities, non-current
Right of use asset
Other non-current assets

Total assets
Liabilities and stockholders' equity
Current liabilities:

Accounts payable and accrued liabilities
Deferred license revenue, current
Lease liability, current
Total current liabilities
Liability related to sale of future royalties
Deferred license revenue, non-current
Contingent consideration
Lease liability, non-current
Total liabilities
Stockholders’ equity
Common shares

Authorized: unlimited number without par value
Issued and outstanding: 157,455,363 (December 31, 2021: 144,987,736)
Additional paid-in capital
Deficit
Accumulated other comprehensive loss
Total stockholders' equity

Total liabilities and stockholders' equity

December 31, 2022

December 31, 2021

$

$

$

$

30,776  $
116,137 
1,352 
2,874 
151,139 
5,070 
37,363 
1,744 
103 
195,419  $

16,029  $
16,456 
372 
32,857 
10,365 
5,999 
7,531 
1,815 
58,567 

109,282 
46,035 
899 
4,445 
160,661 
5,983 
35,688 
2,092 
61 
204,485 

10,838 
— 
383 
11,221 
16,296 
— 
5,298 
2,231 
35,046 

1,318,737 
72,406 
(1,203,803)
(50,488)
136,852 
195,419  $

1,286,636 
65,485 
(1,134,347)
(48,335)
169,439 
204,485 

See accompanying notes to the consolidated financial statements.

76

 
 
 
 
 
 
 
 
 
 
 
ARBUTUS BIOPHARMA CORPORATION

Consolidated Statements of Operations and Comprehensive Loss

(Expressed in thousands of US Dollars, except share and per share amounts)

Revenue

Collaborations and licenses
Non-cash royalty revenue

Total revenue
Operating expenses

Research and development
General and administrative
Change in fair value of contingent consideration

Total operating expenses
Loss from operations
Other income (loss)
Interest income
Interest expense
Foreign exchange (loss) gain

Total other income (loss)
Loss before income taxes
Income tax expense
Net loss
Items applicable to preferred shares

Dividend accretion of convertible preferred shares

Net loss attributable to common shares
Loss per share

Basic and diluted

Weighted average number of common shares

Basic and diluted
Comprehensive loss

Unrealized loss on available-for-sale securities

Comprehensive loss

Year ended December 31,

2022

2021

$

$

$

$

$
$

$

31,366 
7,653 
39,019 

84,408 
17,834 
2,233 
104,475 
(65,456)

2,192 
(1,726)
(22)
444 
(65,012)
(4,444)
(69,456)

— 
(69,456)

(0.46)

150,939,337 

(2,153)
(71,609)

$

$

$

$
$

4,880 
6,108 
10,988 

65,502 
17,136 
1,872 
84,510 
(73,522)

127 
(2,857)
5 
(2,725)
(76,247)
— 
(76,247)

(12,139)
(88,386)

(0.83)

106,242,452 

(164)
(76,411)

See accompanying notes to the consolidated financial statements.

77

 
 
 
 
 
 
 
 
 
 
 
 
ARBUTUS BIOPHARMA CORPORATION

Consolidated Statement of Stockholders’ Equity

(Expressed in thousands of US Dollars, except share and per share amounts)

Convertible Preferred Shares

Common Shares

Number of
shares
1,164,000 

Share
capital

$

149,408 

Number of
shares
89,678,722 

Share capital

Additional
paid-in
capital

Deficit

$

985,939 

$

60,751 

$

(1,045,961)

Accumulated other
comprehensive loss
(48,171)
$

Total
stockholders'
equity

$

101,966 

— 

12,139 

— 

— 

— 

(12,139)

(1,164,000)
— 

(161,547)
— 

22,833,922 
— 

161,547 
— 

— 

— 

31,571,036 

134,665 

— 
6,385 

263 

— 

(356)

— 
— 

— 

— 

— 

— 

196,335 

707,721 

— 
— 

817 

3,668 

(1,558)

— 
— 

— 
— 

— 
(76,247)

— 

— 
— 

— 

— 

— 

— 

(164)

— 

144,987,736 

$

1,286,636 

$

65,485 

$

(1,134,347)

$

(48,335)

$

— 

— 

— 

— 

8,645,426 

20,324 

7,182 

26 

— 

171,224 

588 

(193)

3,579,952 

10,973 

71,025 

— 
— 

216 

— 
— 

— 

(94)

— 
— 

— 

— 

— 

— 

— 

— 

— 

— 

— 

— 

— 

— 

— 
(69,456)

(2,153)
— 

157,455,363 

$

1,318,737 

$

72,406 

$

(1,203,803)

$

(50,488)

$

Balance at December 31, 2020
Accretion of accumulated dividends on
Preferred Shares
Conversion of Preferred Shares into
Common Shares
Stock-based compensation
Certain fair value adjustments to
liability stock option awards
Issuance of common shares pursuant to
the Open Market Sales Agreement
Issuance of common shares pursuant to
exercise of ESPP
Issuance of common shares pursuant to
exercise of stock options
Unrealized loss on available-for-sale
securities
Net loss

Balance at December 31, 2021

Stock-based compensation
Certain fair value adjustments to
liability stock option awards
Issuance of common shares pursuant to
the Open Market Sales Agreement
Issuance of common shares pursuant to
exercise of ESPP
Issuance of common shares pursuant to
Share Purchase Agreement
Issuance of common shares pursuant to
exercise of stock options
Unrealized loss on available-for-sale
securities
Net loss

Balance at December 31, 2022

— 

— 

— 

— 

— 
— 
— 

— 

— 

— 

— 

— 

— 

— 
— 

— 

$

$

— 

— 

— 

— 

— 
— 
— 

— 

— 

— 

— 

— 

— 

— 
— 

— 

— 

— 
6,385 

263 

134,665 

461 

2,110 

(164)

(76,247)

169,439 

7,182 

26 

20,324 

395 

10,973 

122 

(2,153)
(69,456)

136,852 

See accompanying notes to the consolidated financial statements.

78

 
 
ARBUTUS BIOPHARMA CORPORATION

Consolidated Statements of Cash Flows

(Expressed in thousands of US Dollars, except share and per share amounts)

OPERATING ACTIVITIES

Net loss
Non-cash items:
Depreciation
Stock-based compensation expense
Change in fair value of contingent consideration
Non-cash royalty revenue
Non-cash interest expense
Net accretion and amortization of investments in marketable securities

Net change in operating items:

Accounts receivable
Prepaid expenses and other assets
Accounts payable and accrued liabilities
Deferred license revenue
Other liabilities

Net cash used in operating activities
INVESTING ACTIVITIES

Purchase of investments in marketable securities
Disposition of investments in marketable securities
Acquisition of property and equipment

Net cash used in investing activities
FINANCING ACTIVITIES

Issuance of common shares pursuant to Share Purchase Agreement
Issuance of common shares pursuant to the ATM
Issuance of common shares pursuant to exercise of stock options
Issuance of common shares pursuant to exercise of ESPP

Net cash provided by financing activities
Effect of foreign exchange rate changes on cash and cash equivalents
(Decrease) increase in cash and cash equivalents
Cash and cash equivalents, beginning of period
Cash and cash equivalents, end of period
Supplemental cash flow information
Preferred shares dividends accrued

Year ended December 31,

2022

2021

$

(69,456) $

(76,247)

1,427 
7,182 
2,233 
(7,653)
1,722 
(54)

(453)
2,430 
5,216 
22,455 
(405)
(35,356)

(130,430)
56,000 
(512)
(74,942)

10,973 
20,324 
122 
395 
31,814 
(22)
(78,506) $
109,282  $
30,776  $

1,753 
6,424 
1,872 
(6,108)
2,850 
999 

413 
(1,025)
1,911 
— 
(374)
(67,532)

(82,219)
70,350 
(809)
(12,678)

— 
134,665 
2,110 
461 
137,236 
5 
57,031 
52,251 
109,282 

—  $

(12,139)

$
$
$

$

 See accompanying notes to the consolidated financial statements.

79

 
 
 
 
 
 
 
 
 
 
ARBUTUS BIOPHARMA CORPORATION

Notes to Consolidated Financial Statements

(Tabular amounts in thousands of US Dollars, except share and per share amounts) 

1.    Organization

Description of the Business

Arbutus Biopharma Corporation (“Arbutus” or the “Company”) is a clinical-stage biopharmaceutical company leveraging its extensive virology expertise
to develop novel therapeutics that target specific viral diseases. The Company’s current focus areas include Hepatitis B virus (“HBV”), SARS-CoV-2 and
other coronaviruses. To address HBV, the Company is developing an RNA interference (“RNAi”) therapeutic, an oral PD-L1 inhibitor, and an oral RNA
destabilizer to potentially identify a combination regimen with the aim of providing a functional cure for patients with chronic HBV infection (“cHBV”) by
suppressing  viral  replication,  reducing  surface  antigen  and  reawakening  the  immune  system.  The  Company  believes  its  lead  compound,  AB-729,  is  the
only RNAi therapeutic with evidence of immune re-awakening. AB-729 is currently being evaluated in multiple phase 2 clinical trials. The Company also
has an ongoing drug discovery and development program directed to identifying novel, orally active agents for treating coronaviruses, including SARS-
CoV-2,  where  the  Company  has  nominated  a  compound  and  has  begun  IND-enabling  pre-clinical  studies.  In  addition,  the  Company  is  also  exploring
oncology applications for its internal PD-L1 portfolio.

Liquidity

At December 31, 2022, the Company had an aggregate of $184.3 million in cash, cash equivalents and investments in marketable securities. The Company
had no outstanding debt as of December 31, 2022. The Company believes it has sufficient cash resources to fund its operations for at least the next 12
months.

The success of the Company is dependent on obtaining the necessary regulatory approvals to bring its products to market and achieve profitable operations.
The Company’s research and development activities and the commercialization of its products are dependent on its ability to successfully complete these
activities and to obtain adequate financing through a combination of financing activities and operations. It is not possible to predict either the outcome of
the Company’s existing or future research and development programs or the Company’s ability to continue to fund these programs in the future.

COVID-19 Impact

The Company continues to monitor the effects of COVID-19, which has caused significant disruptions around the world. Measures implemented around
the world in attempts to slow the spread of COVID-19 have had, and will likely continue to have, a major impact on clinical development, at least in the
near-term, including shortages and delays in the supply chain, and prohibitions in certain countries on enrolling patients in new clinical trials. While the
Company has been able to progress with its clinical and pre-clinical activities to date, it is not possible to predict if the COVID-19 pandemic will materially
impact the Company’s plans and timelines in the future.

2.    Significant accounting policies 

Basis of presentation and principles of consolidation

These consolidated financial statements have been prepared in accordance with U.S. generally accepted accounting principles (“GAAP”) and include the
accounts of Arbutus Biopharma Corporation and its one wholly-owned subsidiary, Arbutus Biopharma, Inc. All intercompany balances and transactions
have been eliminated. Certain prior year amounts have been reclassified to conform to the current year presentation. In February 2021, Arbutus Biopharma
US Holdings, Inc., which was another wholly-owned subsidiary, merged into Arbutus Biopharma, Inc. with Arbutus Biopharma, Inc. continuing its legal
existence and Arbutus Biopharma US Holdings, Inc. ceasing to exist.

80

 
Use of estimates

The preparation of the consolidated financial statements in conformity with GAAP requires management to make estimates and assumptions about future
events that affect the reported amounts of assets, liabilities, revenue, expenses and contingent liabilities as of the end or during the reporting period. Actual
results  could  significantly  differ  from  those  estimates.  Significant  estimates  in  the  accompanying  consolidated  financial  statements  impact  contingent
consideration, income tax recoveries, stock-based compensation, clinical trial accruals and the sale of future royalties liability.

Cash and cash equivalents

Cash and cash equivalents are all highly liquid instruments with an original maturity of three months or less when purchased. Cash equivalents are recorded
at cost plus accrued interest. The carrying value of these cash equivalents approximates their fair value.

Investments in marketable securities

The Company’s short-term investments consist of marketable securities that have original maturities exceeding three months and remaining maturities of
less than one year. The Company classifies investments with remaining maturities of one year or longer as non-current. These investments are accounted
for  as  available-for-sale  securities  and  are  reported  at  fair  value,  with  unrealized  gains  and  losses  reported  in  other  comprehensive  loss  until  their
disposition.  Realized  gains  and  losses  from  the  sale  of  marketable  securities,  if  any,  are  calculated  using  the  specific-identification  method,  and  are
recorded as a component of other income or loss. The Company reviews its available-for-sale securities at each period end to determine if they remain
available-for-sale based on the Company’s current intent and ability to sell the security if it is required to do so. Declines in value judged to be other-than-
temporary are included in interest expense in the Company’s statements of operations and comprehensive loss. As of December 31, 2022, the recorded
value of the Company’s investments in marketable securities was deemed to be recoverable in all respects.

All investments are governed by the Company’s Investment Policy approved by the Company’s board of directors.

Foreign currency translation and functional currency conversion

The Company’s functional currency is the United States dollar. Monetary assets and liabilities denominated in foreign currencies are translated into United
States dollars using exchange rates in effect at the balance sheet date. Opening balances related to non-monetary assets and liabilities are based on prior
period translated amounts, and non-monetary assets and non-monetary liabilities are translated at the approximate exchange rate prevailing at the date of
the transaction. Revenue and expense transactions are translated at the approximate exchange rate in effect at the time of the transaction. Foreign exchange
gains and losses are included in the statement of operations and comprehensive loss as foreign exchange gains or losses.

Investment in Genevant

Arbutus accounts for its interest in Genevant as equity securities without readily determinable fair values. Accordingly, an estimate of the fair value of the
securities is based on the original cost less previously recognized equity method losses, less impairments, plus or minus changes resulting from observable
price  changes  in  orderly  transactions  for  identical  or  similar  Genevant  securities.  As  of  December  31,  2022,  Arbutus  owned  approximately  16%  of  the
common equity of Genevant and the carrying value of Arbutus’ investment in Genevant was zero.

See note 5 for more information.

81

Property and equipment

Property and equipment is recorded at cost less impairment losses and accumulated depreciation. The Company records depreciation using the straight-line
method over the estimated useful lives of the capital assets as follows:

Laboratory equipment
Computer and office equipment
Furniture and fixtures

Useful Life (Years)
5
to
5

5

2

Leasehold  improvements  are  depreciated  over  their  estimated  useful  lives  but  in  no  case  longer  than  the  lease  term,  except  where  lease  renewal  is
reasonably assured.

Property and equipment is reviewed for impairment whenever events or changes in circumstances indicate that the carrying amount of such assets may not
be recoverable. If such a review should indicate that the carrying amount of long-lived assets is not recoverable, then such assets are written down to their
fair values.

Revenue from collaborations and licenses

The Company generates revenue primarily through collaboration agreements and license agreements. Such agreements may require the Company to deliver
various  rights  and/or  services,  including  intellectual  property  rights  or  licenses  and  research,  development  and  manufacturing  services.  Under  such
agreements, the Company is generally eligible to receive non-refundable upfront payments, funding for research, development and manufacturing services,
milestone payments, and royalties.

The Company’s collaboration agreements fall under the scope of ASC Topic 808, Collaborative Arrangements, (“ASC 808”) when both parties are active
participants  in  the  arrangement  and  are  exposed  to  significant  risks  and  rewards.  For  certain  arrangements  under  the  scope  of  ASC  808,  the  Company
analogizes to ASC 606 for some aspects, including for the delivery of a good or service (i.e., a unit of account).

ASC 606, Revenue From Contracts with Customers (“ASC 606”) requires an entity to recognize the amount of revenue to which it expects to be entitled
for the transfer of promised goods or services to customers under a five-step model: (i) identify contract(s) with a customer; (ii) identify the performance
obligations in the contract; (iii) determine the transaction price; (iv) allocate the transaction price to the performance obligations in the contract; and (v)
recognize revenue when or as a performance obligation is satisfied.

In contracts where the Company has more than one performance obligation to provide its customer with goods or services, each performance obligation is
evaluated to determine whether it is distinct based on whether: (i) the customer can benefit from the good or service either on its own or together with other
resources that are readily available; and (ii) the good or service is separately identifiable from other promises in the contract. The consideration under the
contract is then allocated between the distinct performance obligations based on their respective relative stand-alone selling prices. The estimated stand-
alone selling price of each deliverable reflects the Company’s best estimate of what the selling price would be if the deliverable was regularly sold on a
stand-alone basis and is determined by reference to market rates for the good or service when sold to others or by using an adjusted market assessment
approach if the selling price on a stand-alone basis is not available.

The consideration allocated to each distinct performance obligation is recognized as revenue when control is transferred to the customer for the related
goods  or  services.  Consideration  associated  with  at-risk  substantive  performance  milestones,  including  sales-based  milestones,  is  recognized  as  revenue
when  it  is  probable  that  a  significant  reversal  of  the  cumulative  revenue  recognized  will  not  occur.  Sales-based  royalties  received  in  connection  with
licenses of intellectual property are subject to a specific exception in the revenue standards, whereby the consideration is not included in the transaction
price and recognized in revenue until the customer’s subsequent sales or usages occur.

82

 
 
Leases

The  Company  accounts  for  its  lease  under  ASC  842,  Leases,  which  generally  requires  the  recognition  of  operating  and  financing  lease  liabilities  with
corresponding right-of-use assets on the balance sheet. See note 6 for more information.

Research and development costs

Research and development costs include compensation and benefits for research and development employees, an allocation of overhead expenses and costs
associated with materials and supplies used in clinical trials and research and development, outside contracted services including clinical and pre-clinical
study  costs,  legal,  regulatory  compliance  and  fees  paid  to  consultants  or  outside  parties  for  research  and  development  activities  performed  on  the
Company’s behalf. Such costs are charged to expense in the period in which they are incurred.

Research and development costs that are paid in advance of performance or receipt are recorded as prepaid expense and are amortized over the period that
the services are performed.

Net loss attributable to common shareholders per share

Net loss attributable to common shareholders per share is calculated based on the weighted average number of common shares outstanding. Diluted net loss
attributable  to  common  shareholders  per  share  does  not  differ  from  basic  net  loss  attributable  to  common  shareholders  per  share  for  the  years  ended
December  31,  2022  and  2021,  since  the  effect  of  including  potential  common  shares  would  be  anti-dilutive.  For  the  year  ended  December  31,  2022,
potential common shares of 15.5 million pertaining to outstanding stock options were excluded from the calculation of net loss attributable to common
shareholders  per  share.  A  total  of  approximately  11.4  million  outstanding  stock  options  were  excluded  from  the  calculation  for  the  year  ended
December 31, 2021.

On October 18, 2021, the Company’s outstanding Series A participating convertible preferred shares (“ Preferred Shares”) were converted into 22,833,922
common shares. Prior to that date, the Company followed the two-class method when computing net loss attributable to common shareholders per share as
the Preferred Shares, as further described in note 12, met the definition of participating securities. The Company’s Preferred Shares entitled the holders to
participate  in  dividends  but  did  not  require  the  holders  to  participate  in  losses  of  the  Company.  Accordingly,  net  losses  attributable  to  holders  of  the
Company’s common shares were not allocated to holders of the Preferred Shares.

See note 12 and note 13 for more information about the Company’s common shares.

Deferred income taxes

Income  taxes  are  accounted  for  using  the  asset  and  liability  method  of  accounting.  Deferred  income  taxes  are  recognized  for  the  future  income  tax
consequences  attributable  to  differences  between  the  carrying  values  of  assets  and  liabilities  and  their  respective  income  tax  bases  and  for  loss  carry-
forwards.  Deferred  income  tax  assets  and  liabilities  are  measured  using  enacted  income  tax  rates  expected  to  apply  to  taxable  income  in  the  periods  in
which temporary differences are expected to be recovered or settled. The effect on deferred income tax assets and liabilities of a change in tax laws or rates
is included in earnings in the period that includes the enactment date. When realization of deferred income tax assets does not meet the more-likely-than-
not criterion for recognition, a valuation allowance is provided.

83

  
 
Stock-based compensation

The  Company  measures  and  recognizes  compensation  expense  for  all  share-based  compensation  arrangements  based  on  estimated  fair  values.  The
Company uses the Black-Scholes option valuation model to estimate the fair value of stock options at the date of grant. The Black-Scholes option valuation
model requires the input of subjective assumptions to calculate the value of stock options. For those assumptions, the Company uses historical data and
other information to estimate the expected price volatility and risk free interest rate for all awards. The expected life of stock options granted are estimated
to be five years for employees and six years for directors and executives, based on the Company’s historical experience. Assumptions on the dividend yield
are based on the fact that the Company has never paid cash dividends and has no present intention to pay cash dividends. Expense is recognized over the
vesting period for all awards and commences at the grant date for time-based awards and upon the Company’s determination that the achievement of such
performance conditions is probable for performance-based awards. Forfeitures are recognized as they occur.

For the Company’s Employee Stock Purchase Plan, the fair value of the right to acquire stock at a discounted price under the plan is calculated using the
Black-Scholes valuation model. Expense is recognized over the period the employee contributes to the plan through payroll deductions.

The  Company  accounts  for  liability-classified  stock  option  awards  (“liability  options”)  under  ASC  718  -  Compensation  -  Stock  Compensation  (“ASC
718”),  under  which  awards  of  options  that  provide  for  an  exercise  price  that  is  not  denominated  in:  (a)  the  currency  of  a  market  in  which  a  substantial
portion of the Company’s equity securities trades, (b) the currency in which the employee’s pay is denominated, or (c) the Company’s functional currency,
are required to be classified as liabilities. As of January 1, 2016, the Company changed its functional currency to US dollars, which resulted in certain stock
option awards with exercise prices denominated in Canadian dollars having an exercise price that is not denominated in the Company’s functional currency.
As  such,  the  historic  equity  classification  of  these  stock  option  awards  changed  to  liability  classification  effective  January  1,  2016.  The  change  in
classification resulted in reclassification of these awards from additional paid-in capital to a liability.

Liability options are re-measured to their fair values at each reporting date with changes in the fair value recognized in share-based compensation expense
or additional paid-in capital until settlement or cancellation. Under ASC 718, when an award is reclassified from equity to liability, if at the reclassification
date the original vesting conditions are expected to be satisfied, then the minimum amount of compensation cost to be recognized is based on the grant date
fair value of the original award. Fair value changes below this minimum amount are recorded in additional paid-in capital.

Preferred Shares

The Company accounted for its Preferred Shares under ASC 480 – Distinguishing Liabilities from Equity (“ASC 480”), which provides guidance for equity
instruments with conversion features. The Company classified the Preferred Shares in its consolidated balance sheet wholly as equity, with no bifurcation
of conversion feature from the host contract, given that the Preferred Shares could not be cash-settled and the redemption features, which included a fixed
conversion  ratio  with  predetermined  timing  and  proceeds,  were  within  the  Company’s  control.  The  Company  accrued  for  the  8.75%  per  annum
compounding accrual at each reporting period-end date as an increase to share capital, and an increase to deficit. The Company’s Preferred Shares were
converted into 22,833,922 common shares on October 18, 2021.

Segment information

As of December 31, 2022, the Company viewed its operations and managed its business as one operating segment consistent with how its chief operating
decision-maker, the Chief Executive Officer, makes decisions regarding resource allocation and assessing performance. Substantially all of the Company’s
premises, property and equipment are located in the United States.

Comprehensive loss

Comprehensive loss is comprised of net loss and adjustments for the change in unrealized gains and losses on investments in available-for-sale marketable
securities. The Company includes comprehensive loss and its components in the consolidated statements of operations and comprehensive loss, net of tax
effects if any.

84

 
Concentrations of Credit Risk

Financial  instruments  which  potentially  subject  the  Company  to  credit  risk  consist  primarily  of  cash,  cash  equivalents  and  marketable  securities.  The
Company holds these investments in highly rated financial institutions, and, by policy, limits the amounts of credit exposure to any one financial institution.
These amounts at times may exceed federally insured limits. The Company has not experienced any credit losses in such accounts and does not believe it is
exposed  to  any  significant  credit  risk  on  these  funds.  The  Company  has  no  off-balance  sheet  concentrations  of  credit  risk,  such  as  foreign  currency
exchange contracts, option contracts or other hedging arrangements.

Recent accounting pronouncements

In June 2016, the Financial Accounting Standards Board (“FASB”) issued Accounting Standards Update (“ASU”) No. 2016-13, Financial Instruments -
Credit Losses: Measurement of Credit Losses on Financial Instruments (ASC 326). The guidance is effective for the Company beginning January 1, 2023
and  it  changes  how  entities  account  for  credit  losses  on  financial  assets  and  other  instruments  that  are  not  measured  at  fair  value  through  net  income,
including available-for-sale debt securities. The Company does not anticipate that the new guidance will have a material impact on its results of operations
or financial position.

3.    Fair value measurements 

The Company measures certain financial instruments and other items at fair value.

To determine the fair value, the Company uses the fair value hierarchy for inputs used in measuring fair value that maximizes the use of observable inputs
and minimizes the use of unobservable inputs by requiring that the most observable inputs be used when available. Observable inputs are inputs market
participants would use to value an asset or liability and are developed based on market data obtained from independent sources. Unobservable inputs are
inputs based on assumptions about the factors market participants would use to value an asset or liability. The three levels of inputs that may be used to
measure fair value are as follows:

•

•

•

Level 1 inputs are quoted market prices for identical instruments available in active markets. The Company’s cash and cash equivalents are
measured using Level 1 inputs.

Level  2  inputs  are  inputs  other  than  quoted  prices  included  within  Level  1  that  are  observable  for  the  asset  or  liability  either  directly  or
indirectly. If the asset or liability has a contractual term, the input must be observable for substantially the full term. An example includes
quoted  market  prices  for  similar  assets  or  liabilities  in  active  markets.  The  Company’s  investments  in  marketable  securities  are  measured
using Level 2 inputs.

Level  3  inputs  are  unobservable  inputs  for  the  asset  or  liability  and  will  reflect  management’s  assumptions  about  market  assumptions  that
would be used to price the asset or liability. The Company’s liability-classified options and contingent consideration are measured using Level
3 inputs.

Assets and liabilities are classified based on the lowest level of input that is significant to the fair value measurements. Changes in the observability of
valuation inputs may result in a reclassification of levels for certain securities within the fair value hierarchy.

The carrying values of cash and cash equivalents, accounts receivable, accounts payable and accrued liabilities approximate their fair values due to the
immediate or short-term maturity of these financial instruments.

To determine the fair value of the contingent consideration (note 10), the Company uses a probability weighted assessment of the likelihood the milestones
would  be  met  and  the  estimated  timing  of  such  payments,  and  then  the  potential  contingent  payments  were  discounted  to  their  present  value  using  a
probability adjusted discount rate that reflects the early stage nature of the development program, time to complete the program development, and overall
biotech indices. The Company determined that the fair value of the contingent consideration was $7.5 million as of December 31, 2022 and the increase of
$2.2 million has been recorded within operating expenses in the statement of operations and comprehensive loss for the year ended December 31, 2022.
The assumptions used in the discounted cash flow model are level 3 inputs as defined above. The

85

 
Company assessed the sensitivity of the fair value measurement to changes in these unobservable inputs, and determined that changes within a reasonable
range would not result in a materially different assessment of fair value.  

The following tables present information about the Company’s assets and liabilities that are measured at fair value on a recurring basis, and indicates the
fair value hierarchy of the valuation techniques used to determine such fair value:

As of December 31, 2022
Assets
Cash and cash equivalents
Investments in marketable securities, current
Investments in marketable securities, non-current

Total
Liabilities
Liability-classified options
Contingent consideration

Total

As of December 31, 2021
Assets
Cash and cash equivalents
Investments in marketable securities, current
Investments in marketable securities, non-current

Total
Liabilities
Liability-classified options
Contingent consideration
Total

Level 1

Level 2

Level 3

Total

30,776  $
— 
— 
30,776  $

—  $
— 
—  $

(in thousands)

—  $

116,137 
37,363 
153,500  $

—  $
— 
—  $

—  $
— 
— 
—  $

1  $

7,531 
7,532  $

30,776 
116,137 
37,363 
184,276 

1 
7,531 
7,532 

Level 1

Level 2

Level 3

Total

109,282  $
— 
— 
109,282  $

—  $
— 
—  $

(in thousands)

—  $

46,035 
35,688 
81,723  $

—  $
— 
—  $

—  $
— 
— 
—  $

26  $

5,298 
5,324  $

109,282 
46,035 
35,688 
191,005 

26 
5,298 
5,324 

$

$

$

$

$

$

$

$

86

The following table presents the changes in fair value of the Company’s liability-classified stock option awards:

Year ended December 31, 2022
Year ended December 31, 2021

$
$

26  $
250  $

(in thousands)
—  $
(96) $

(25) $
(128) $

1 
26 

Liability at beginning of
the period

Fair value of liability-classified
options exercised in the period

Decrease in fair value of
liability

Liability at end of the
period

The following table presents the changes in fair value of the Company’s contingent consideration:

Year ended December 31, 2022
Year ended December 31, 2021

$
$

(in thousands)

5,298  $
3,426  $

2,233  $
1,872  $

7,531 
5,298 

Liability at beginning of the period

Increase in fair value of liability

Liability at end of the period

87

 
4.    Investments in marketable securities 

Investments in marketable securities and cash equivalents consisted of the following:

As of December 31, 2022
Cash equivalents

Money market fund
Total

Investments in marketable short-term securities

US government agency bonds
US corporate bonds
US treasury bills
US government bonds
Total

Investments in marketable long-term securities

US government agency bonds
US corporate bonds
US government bonds
Total

Amortized Cost

Gross Unrealized Gain

(1)

Gross Unrealized Loss

(1)

Fair Value

(in thousands)

$
$

$

$

$

$

23,218  $
23,218  $

26,686  $
27,144 
8,483 
55,361 
117,674  $

3,724  $

25,433 
8,972 
38,129  $

—  $
—  $

—  $
— 
— 
— 
—  $

—  $
— 
— 
—  $

—  $
—  $

(424) $
(303)
(16)
(794)
(1,537) $

(130) $
(336)
(300)
(766) $

23,218 
23,218 

26,262 
26,841 
8,467 
54,567 
116,137 

3,594 
25,097 
8,672 
37,363 

(1) 
  Gross unrealized gain (loss) is pre-tax and is reported in accumulated other comprehensive loss.

Amortized Cost

Gross Unrealized Gain

(1)

Gross Unrealized Loss

(1)

Fair Value

As of December 31, 2021
Cash equivalents

Money market fund
Total

Investments in marketable short-term securities

US government agency bonds
US government bonds
Total

Investments in marketable long-term securities

US government agency bonds
US treasury bills

Total

$
$

$

$

$

$

93,211  $
93,211  $

8,131  $

37,968 
46,099  $

13,068  $
22,707 
35,775  $

(in thousands)

—  $
—  $

—  $
— 
—  $

—  $
— 
—  $

—  $
—  $

(11) $
(53)
(64) $

(29) $
(58)
(87) $

93,211 
93,211 

8,120 
37,915 
46,035 

13,039 
22,649 
35,688 

(1) 

Gross unrealized gain (loss) is pre-tax and is reported in accumulated other comprehensive loss.

The contractual maturity of the $116.1 million of short-term marketable securities held by the Company as of December 31, 2022 is less than one year. As
of December 31, 2022, the Company held $37.4 million of long-term marketable securities with contractual maturities of more than one year, but less than
five years. As of December 31, 2021, the Company’s $46.0 million of short-term marketable securities had contractual maturities of less than one year,
while the Company’s $35.7 million of long-term marketable securities had maturities of more than one year, but less than five years.

The Company had realized gains on investments of less than $0.1 million and zero for the years ended December 31, 2022 and 2021, respectively.

88

5.    Investment in Genevant

In April 2018, the Company entered into an agreement with Roivant Sciences Ltd. (“Roivant”), its largest shareholder, to launch Genevant Sciences Ltd.
(“Genevant”), a company focused on a broad range of RNA-based therapeutics enabled by the Company’s LNP and ligand conjugate delivery technologies.
The Company licensed rights to its LNP and ligand conjugate delivery platforms to Genevant for RNA-based applications outside of HBV, except to the
extent certain rights had already been licensed to other third parties (the “Genevant License”). The Company retained all rights to its LNP and conjugate
delivery platforms for HBV.

Under the Genevant License, as amended, if a third party sublicensee of intellectual property licensed by Genevant from the Company commercializes a
sublicensed  product,  the  Company  becomes  entitled  to  receive  a  specified  percentage  of  certain  revenue  that  may  be  received  by  Genevant  for  such
sublicense,  including  royalties,  commercial  milestones  and  other  sales-related  revenue,  or,  if  less,  tiered  low  single-digit  royalties  on  net  sales  of  the
sublicensed product. The specified percentage is 20% in the case of a mere sublicense (i.e., naked sublicense) by Genevant without additional contribution
and 14% in the case of a bona fide collaboration with Genevant.

Additionally,  if  Genevant  receives  proceeds  from  an  action  for  infringement  by  any  third  parties  of  the  Company’s  intellectual  property  licensed  to
Genevant, the Company would be entitled to receive, after deduction of litigation costs, 20% of the proceeds received by Genevant or, if less, tiered low
single-digit royalties on net sales of the infringing product (inclusive of the proceeds from litigation or settlement, which would be treated as net sales).

The Company accounts for its interest in Genevant as equity securities without readily determinable fair values. Accordingly, an estimate of the fair value
of  the  securities  is  based  on  the  original  cost  less  previously  recognized  equity  method  losses,  less  impairments,  plus  or  minus  changes  resulting  from
observable  price  changes  in  orderly  transactions  for  identical  or  a  similar  Genevant  securities.  As  of  December  31,  2022,  the  carrying  value  of  the
Company’s investment in Genevant was zero and the Company owned approximately 16% of the common equity of Genevant.

6.    Leases

The Company had one operating lease for its office and laboratory space as of December 31, 2022. The Company’s corporate headquarters is located at 701
Veterans Circle, Warminster, Pennsylvania. The lease expires on April 30, 2027, and the Company has the option of extending the lease for two further
five-year  terms.  The  Company  also  previously  leased  office  space  located  at  626  Jacksonville  Road,  Warminster,  Pennsylvania  under  a  lease  that
terminated on August 31, 2022.

The Company accounts for its leases under ASC 842, Leases. Leases with an initial term of 12 months or less are not recorded on the balance sheet. The
Company determines if an arrangement is a lease at inception. Right-of-use assets represent the Company’s right to use an underlying asset for the lease
term and lease liabilities represent the Company’s obligation to make lease payments arising from the lease. Operating lease right-of-use assets and lease
liabilities are recognized based on the present value of lease payments over the lease term. The leases do not provide an implicit rate so in determining the
present value of lease payments, the Company utilized its incremental borrowing rate for the applicable lease, which was 9.0% for the 701 Veterans Circle
lease and 7.6% for the 626 Jacksonville Road lease. The Company recognizes lease expense on a straight-line basis over the remaining lease term.
During each of the years ended December 31, 2022 and 2021, the Company incurred total operating lease expenses of $0.7 million, which included lease
expenses associated with fixed lease payments of $0.6 million, and variable payments associated with common area maintenance and similar expenses of
$0.1 million.

89

Weighted average remaining lease term and discount rate were as follows:

Weighted-average remaining lease term (years)
Weighted average discount rate

The Company did not include options to extend its lease terms as part of its ROU asset and lease liabilities.

Supplemental cash flow information related to the Company’s operating leases was as follows:

As of December 31, 2022
4.3
9.0%

Cash paid for amounts included in the measurement of lease liabilities

$

2022

2021

(in thousands)
641  $

650 

Future minimum lease payments under operating leases that have remaining terms as of December 31, 2022 are as follows:

2023
2024
2025
2026
2027
Thereafter

Total lease payments

Less: interest

Present value of lease payments

7.    Property and equipment

The Company’s property and equipment balances as of the years ended December 31, 2022 and 2021 are as follows:

As of December 31, 2022

(in thousands)

598 
616 
635 
654 
134 
— 
2,637 
(450)
2,187 

$

$

$

December 31, 2022
Lab equipment
Leasehold improvements
Computer hardware and software

December 31, 2021
Lab equipment
Leasehold improvements
Computer hardware and software

Cost

Cost

Accumulated
depreciation
(in thousands)

Net book value

6,890  $
8,590 
391 
15,871  $

(5,679) $
(4,749)
(373)
(10,801) $

1,211 
3,841 
18 
5,070 

Accumulated
depreciation

(in thousands)

Net book value

6,408  $
8,563 
386 
15,357  $

(5,178) $
(3,883)
(313)
(9,374) $

1,230 
4,680 
73 
5,983 

$

$

$

$

90

 
 
Depreciation expense for the years ended December 31, 2022 and 2021 was $1.4 million and $1.8 million, respectively.

8.    Accounts payable and accrued liabilities

Accounts payable and accrued liabilities are comprised of the following:

Trade accounts payable
Payroll accruals
Research and development accruals
Professional fee accruals
Other accrued liabilities

Total

9.    Sale of future royalties

December 31, 2022

December 31, 2021

(in thousands)

3,520  $
3,730 
8,261 
512 
6 
16,029  $

3,174 
4,279 
2,371 
983 
31 
10,838 

$

$

On July 2, 2019, the Company entered into a Purchase and Sale Agreement (the “Agreement”) with the Ontario Municipal Employees Retirement System
(“OMERS”), pursuant to which the Company sold to OMERS part of its royalty interest on future global net sales of ONPATTRO, an RNA interference
therapeutic currently being sold by Alnylam.

ONPATTRO utilizes Arbutus’s LNP technology, which was licensed to Alnylam pursuant to the Cross-License Agreement, dated November 12, 2012, by
and  between  the  Company  and  Alnylam  (the  “LNP  License  Agreement”).  Under  the  terms  of  the  LNP  License  Agreement,  the  Company  is  entitled  to
tiered royalty payments on global net sales of ONPATTRO ranging from 1.00% to 2.33% after offsets, with the highest tier applicable to annual net sales
above  $500  million.  This  royalty  interest  was  sold  to  OMERS,  effective  as  of  January  1,  2019,  for  $20  million  in  gross  proceeds  before  advisory  fees.
OMERS will retain this entitlement until it has received $30 million in royalties, at which point 100% of such royalty interest on future global net sales of
ONPATTRO  will  revert  to  the  Company.  OMERS  has  assumed  the  risk  of  collecting  up  to  $30  million  of  future  royalty  payments  from  Alnylam  and
Arbutus is not obligated to reimburse OMERS if they fail to collect any such future royalties. From the inception of the royalty sale through December 31,
2022, an aggregate of $18.9 million of royalties have been collected by OMERS.

The $30 million in royalties to be paid to OMERS is accounted for as a liability, with the difference between the liability and the gross proceeds received
accounted for as a discount. The discount, as well as $1.5 million of transaction costs, will be amortized as interest expense based on the projected balance
of the liability as of the beginning of each period. Management estimated an effective annual interest rate of approximately 8%. Over the course of the
Agreement, the actual interest rate will be affected by the amount and timing of royalty revenue recognized and changes in the timing of forecasted royalty
revenue. On a quarterly basis, the Company will reassess the expected timing of the royalty revenue, recalculate the amortization and effective interest rate
and adjust the accounting prospectively as needed.

The  Company  recognizes  non-cash  royalty  revenue  related  to  the  sales  of  ONPATTRO  during  the  term  of  the  Agreement.  As  royalties  are  remitted  to
OMERS from Alnylam, the balance of the recognized liability is effectively repaid over the life of the Agreement. There are a number of factors that could
materially affect the amount and timing of royalty payments from Alnylam, none of which are within the Company’s control.

During the year ended December 31, 2022, the Company recognized non-cash royalty revenue of $7.7 million and $1.7 million of related non-cash interest
expense.  During  the  year  ended  December  31,  2021,  the  Company  recognized  non-cash  royalty  revenue  of  $6.1  million  and  related  non-cash  interest
expense of $2.9 million.

91

 
 
 
The table below shows the activity related to the net liability for the years ended December 31, 2022 and December 31, 2021:

Net liability related to sale of future royalties - beginning balance
Non-cash royalty revenue
Non-cash interest expense
Net liability related to sale of future royalties - ending balance

Twelve Months Ended December 31,

2022

2021

(in thousands)

16,296  $
(7,653)
1,722 
10,365  $

19,554 
(6,108)
2,850 
16,296 

$

$

In addition to the royalty from the LNP License Agreement, the Company is also receiving a second royalty interest ranging from 0.75% to 1.125% on
global net sales of ONPATTRO, with 0.75% applying to sales greater than $500 million, originating from a settlement agreement and subsequent license
agreement with Acuitas Therapeutics, Inc. (“Acuitas”). The royalty from Acuitas has been retained by the Company and was not part of the royalty sale to
OMERS.

10.    Contingencies and commitments

Arbitration with the University of British Columbia

Certain early work on lipid nanoparticle delivery systems and related inventions was undertaken at the University of British Columbia (“UBC”), as well as
by  the  Company  that  was  subsequently  assigned  to  UBC.  These  inventions  are  licensed  to  the  Company  by  UBC  under  a  license  agreement,  initially
entered  into  in  1998  and  as  amended  in  2001,  2006  and  2007.  The  Company  has  granted  sublicenses  under  the  UBC  license  to  certain  third  parties,
including Alnylam.

In November 2014, UBC filed a demand for arbitration against the Company which alleged entitlement to unpaid royalties. In August 2019, the arbitrator
issued  its  decision  for  the  second  phase  of  the  arbitration,  awarding  UBC  $5.9  million,  which  included  interest  of  approximately  $2.6  million.  The
Company paid the $5.9 million award to UBC in September 2019 and paid an additional $0.2 million for costs and attorneys’ fees in March 2021, and this
matter is now fully resolved.

On December 18, 2020, UBC delivered to the Company a notice of arbitration alleging that under the cross license between UBC and Arbutus, it was due
royalties of $2.0 million plus interest arising from the Company’s sale to OMERS of part of its royalty interest on future global net sales of ONPATTRO,
currently  being  sold  by  Alnylam.  Oral  hearings  for  this  matter  were  held  in  April  2022  and,  on  July  11,  2022,  the  arbitrator  issued  his  decision  fully
dismissing UBC’s claim for royalties. As a result, no payments are owed to UBC. In September 2022, the arbitrator awarded the Company $0.5 million for
reimbursement of costs and attorneys’ fees, which the Company received from UBC in October 2022. This matter is now fully resolved.

Stock Purchase Agreement with Enantigen

In  October  2014,  Arbutus  Inc.,  the  Company’s  wholly-owned  subsidiary,  acquired  all  of  the  outstanding  shares  of  Enantigen  Therapeutics,  Inc.
(“Enantigen”) pursuant to a stock purchase agreement. The amount paid to Enantigen’s selling shareholders could be up to an additional $102.5 million in
sales performance milestones in connection with the sale of the first commercialized product by Arbutus for the treatment of HBV, regardless of whether
such product is based upon assets acquired under this agreement, and a low single-digit royalty on net sales of such first commercialized HBV product, up
to a maximum royalty payment of $1.0 million that, if paid, would be offset against Arbutus’ milestone payment obligations. Certain other development
milestones related to the acquisition were tied to programs which are no longer under development by Arbutus, and therefore the contingency related to
those development milestones is zero.

The  contingent  consideration  is  a  financial  liability  and  is  measured  at  its  fair  value  at  each  reporting  period,  with  any  changes  in  fair  value  from  the
previous reporting period recorded in the statement of operations and comprehensive loss (note 3).

The fair value of the contingent consideration was $7.5 million as of December 31, 2022.

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11.    Collaborations and royalty entitlements

Collaborations

Qilu Pharmaceuticals Co, Ltd.

In December 2021, the Company entered into a technology transfer and exclusive licensing agreement (the “License Agreement”) with Qilu, pursuant to
which  the  Company  granted  Qilu  an  exclusive  (except  as  to  certain  retained  rights),  sublicensable,  royalty-bearing  license,  under  certain  intellectual
property  owned  by  the  Company,  to  develop,  manufacture  and  commercialize  AB-729,  including  pharmaceutical  products  that  include  AB-729,  for  the
treatment or prevention of hepatitis B in China, Hong Kong, Macau and Taiwan (the “Territory”).

In partial consideration for the rights granted by the Company, Qilu paid the Company a one-time upfront cash payment of $40.0 million on January 5,
2022 and agreed to pay the Company milestone payments totaling up to $245 million, net of withholding taxes, upon the achievement of certain technology
transfer,  development,  regulatory  and  commercialization  milestones  (the  “Milestone  Payments”).  Qilu  paid  $4.4  million  of  withholding  taxes  to  the
Chinese taxing authority on the Company’s behalf, related to the upfront cash payment. In  addition,  Qilu  also  agreed  to  pay  the  Company  double  digit
royalties  into  the  low  twenties  percent  based  upon  annual  net  sales  of  AB-729  in  the  Territory.  The  royalties  are  payable  on  a  product-by-product  and
region-by-region basis, subject to certain limitations.

Qilu is responsible for all costs related to developing, obtaining regulatory approval for, and commercializing AB-729 for the treatment or prevention of
hepatitis B in the Territory. Qilu is required to use commercially reasonable efforts to develop, seek regulatory approval for, and commercialize at least one
AB-729 product candidate in the Territory. A joint development committee has been established between the Company and Qilu to coordinate and review
the  development,  manufacturing  and  commercialization  plans.  Both  parties  also  have  entered  into  a  supply  agreement  and  related  quality  agreement
pursuant to which the Company will manufacture or have manufactured and supply Qilu with all quantities of AB-729 necessary for Qilu to develop and
commercialize in the Territory until the Company has completed manufacturing technology transfer to Qilu and Qilu has received all approvals required for
it or its designated contract manufacturing organization to manufacture AB-729 in the Territory.

Concurrent with the execution of the license agreement, the Company entered into a Share Purchase Agreement (the “Share Purchase Agreement”) with
Anchor  Life  Limited,  a  company  established  pursuant  to  the  applicable  laws  and  regulations  of  Hong  Kong  and  an  affiliate  of  Qilu  (the  “Investor”),
pursuant to which the Investor purchased 3,579,952 of the Company’s common shares, without par value (the “Common Shares”), at a purchase price of
USD $4.19 per share, which was a 15% premium on the thirty-day average closing price of the Common Shares as of the close of trading on December 10,
2021  (the  “Share  Transaction”).  The  Company  received  $15.0  million  of  gross  proceeds  from  the  Share  Transaction  on  January  6,  2022.  The  Common
Shares sold to the Investor in the Share Transaction represented approximately 2.5% of the Common Shares outstanding immediately prior to the execution
of the Share Purchase Agreement.

The License Agreement falls under the scope of ASC 808 as both parties are active participants in the arrangement and are exposed to significant risks and
rewards. While this arrangement is in the scope of ASC 808, the Company analogizes to ASC 606 for some aspects of this arrangement, including for the
delivery  of  a  good  or  service  (i.e.,  a  unit  of  account).  In  accordance  with  the  guidance,  the  Company  identified  the  following  commitments  under  the
arrangement: (i) rights to develop, use, sell, have sold, offer for sale and import any product comprised of Licensed Product (the “Qilu License”); and (ii)
drug supply obligations and manufacturing technology transfer (the “Manufacturing Obligations”). The Company determined that these two commitments
are not distinct performance obligations for purposes of recognizing revenue as the manufacturing process is highly specialized and Qilu would not be able
to benefit from the Qilu License without the Company’s involvement in the manufacturing activities until the transfer of the manufacturing know-how is
complete. As such, the Company will combine these commitments into one performance obligation to which the transaction price will be allocated to and
will recognize this transaction price associated with the bundled performance obligation over time using an inputs method based on labor hours expended
by the Company on its Manufacturing Obligations.

The  Company  determined  the  initial  transaction  price  of  the  combined  performance  obligation  to  be  $49.3  million,  which  includes  the  $40.0  million
upfront fee, $4.4 million of withholding taxes paid by Qilu on behalf of the Company, the premium

93

 
paid  for  the  Share  Transaction  of  $4.1  million,  and  $0.8  million  associated  with  certain  manufacturing  costs  expected  to  be  reimbursed  by  Qilu.  The
Company  determined  the  Milestone  Payments  to  be  variable  consideration  subject  to  constraint  at  inception.  At  the  end  of  each  subsequent  reporting
period, the Company will reevaluate the probability of achievement of the future development, regulatory, and sales milestones subject to constraint and, if
necessary,  will  adjust  its  estimate  of  the  overall  transaction  price.  Any  such  adjustments  will  be  recorded  on  a  cumulative  catch-up  basis,  which  would
affect revenues and earnings in the period of adjustment.

The following table outlines the transaction price and the changes to the related asset and liability balances during the twelve months ended December 31,
2022:

Combined performance obligation
Less contract asset
Total deferred license revenue
Less current portion of deferred license revenue
Non-current deferred license revenue

Transaction Price

Twelve Months Ended December 31, 2022

Cumulative Collaboration Revenue
Recognized
(in thousands)

$

49,270  $

26,015  $

$

Deferred License Revenue

23,255 
(800)
22,455 
16,456 
5,999 

The Company recognized $26.0 million of revenue based on labor hours expended by the Company on its Manufacturing Obligations during the twelve
months ended December 31, 2022.

As of December 31, 2022, the balance of the deferred license revenue was $23.3 million, which, in accordance with ASC 210-20, was partially offset by
the  contract  asset  associated  with  the  manufacturing  cost  reimbursement  of  $0.8  million,  resulting  in  a  net  deferred  license  revenue  liability  of  $22.5
million. The $4.4 million of withholding taxes paid by Qilu on behalf of the Company was recorded as income tax expense during the twelve months ended
December 31, 2022.

The Company incurred $0.6 million of incremental costs in obtaining the Qilu License, which the Company capitalized in other current assets and other
assets and amortizes as a component of general and administrative expense commensurate with the recognition of the combined performance obligation.
The Company recognized $0.3 million of related amortization expense for the twelve months ended December 31, 2022.
The  Company  reevaluates  the  transaction  price  and  the  total  estimated  labor  hours  expected  to  be  incurred  to  satisfy  the  performance  obligations  and
adjusts the deferred revenue at the end of each reporting period. Such changes will result in a change to the amount of collaboration revenue recognized and
deferred revenue.

Assembly Biosciences, Inc.

In  August  2020,  the  Company  entered  into  a  clinical  collaboration  agreement  with  Assembly  Biosciences,  Inc.  (“Assembly”)  to  evaluate  AB-729  in
combination with Assembly’s first-generation HBV core inhibitor (capsid inhibitor) candidate vebicorvir (“VBR”) and standard-of-care NA therapy for the
treatment  of  patients  with  HBV  infection.  Assembly  has  completed  enrollment  in  the  clinical  trial.  In  July  2022,  Assembly  announced  its  plan  to
discontinue development of VBR. Despite this, in consultation with Assembly, the Company continued dosing patients in this Phase 2a proof-of-concept
clinical trial in order to fully and accurately assess the results. Preliminary data from 65 patients indicated that adding VBR to AB-729 and NA therapy
does not positively or negatively impact the reduction of HBsAg compared to AB-729 and NA therapy alone. Accordingly, the Company and Assembly
mutually agreed to discontinue the clinical trial following completion of the final, on-treatment visit at week 48. The Company and Assembly shared in the
costs of the collaboration. The Company incurred $2.8 million and $2.6 million of costs related to the collaboration during the years ended December 31,
2022 and 2021, respectively, and reflected those costs in research and development in the statements of operations and comprehensive loss. Except to the
extent necessary

94

to carry out Assembly’s responsibilities with respect to the collaboration trial, the Company has not provided any license grant to Assembly for use of the
Company’s AB-729 compound.

Vaccitech plc

In July 2021, the Company entered into a clinical collaboration agreement with Vaccitech plc (“Vaccitech”) to evaluate AB-729 followed by Vaccitech’s
VTP-300, a proprietary T-cell stimulating antigen-specific immunotherapeutic, in NrtI-suppressed patients with cHBV.

The Company is responsible for managing this Phase 2a proof-of-concept clinical trial, subject to oversight by a joint development committee comprised of
representatives from the Company and Vaccitech. The Company and Vaccitech retain full rights to their respective product candidates and will split all
costs associated with the clinical trial. The Company incurred $0.8 million and $0.5 million of costs related to the collaboration, net of Vaccitech’s 50%
share, during the years ended December 31, 2022 and 2021, respectively, and reflected those net costs in research and development in the statements of
operations and comprehensive loss.

X-Chem, Inc. and Proteros biostructures GmbH

In  March  2021,  the  Company  entered  into  a  discovery  research  and  license  agreement,  as  amended,  with  X-Chem,  Inc.  (“X-Chem”)  and  Proteros
biostructures  GmbH  (“Proteros”)  to  focus  on  the  discovery  of  novel  inhibitors  targeting  the  SARS-CoV-2  nsp5  main  protease  (M ). The agreement is
designed  to  accelerate  the  development  of  pan-coronavirus  agents  to  treat  COVID-19  and  potential  future  coronavirus  outbreaks.  This  collaboration
brought  together  the  Company’s  expertise  in  the  discovery  and  development  of  antiviral  agents  with  X-Chem’s  industry  leading  DNA-encoded  library
(DEL)  technology  and  Proteros’  protein  sciences,  biophysics  and  structural  biology  capabilities  and  provides  important  synergies  to  potentially  identify
safe  and  effective  therapies  against  coronaviruses  including  SARS-CoV-2.  The  collaboration  allows  for  the  rapid  screening  of  one  of  the  largest  small
molecule libraries against M   (an  essential  protein  required  for  the  virus  to  replicate  itself)  and  the  use  of  state-of-the-art  structure  guided  methods  to
rapidly  optimize  M   inhibitors  to  progress  to  clinical  candidates.  Through  this  collaboration,  the  Company  has  identified  and  obtained  a  worldwide
exclusive license to several molecules that inhibit M , a validated target for the treatment of COVID-19 and potential future coronavirus outbreaks. In the
fourth  quarter  of  2022,  the  Company  nominated  AB-343  as  its  lead  candidate  that  inhibits  M   and  the  Company  is  also  continuing  lead  optimization
activities for an nsp12 viral polymerase candidate.

pro

pro

pro

pro

pro

The  agreement  provides  for  payments  by  the  Company  to  X-Chem  and  Proteros  upon  satisfaction  of  certain  development,  regulatory  and  commercial
milestones, as well as royalties on sales. The agreement with X-Chem and Proteros was amended effective March 31, 2022 primarily to extend the term of
the collaboration and update the funding and fee structure. The Company incurred $1.3 million and $1.9 million of costs related to the collaboration during
the  years  ended  December  31,  2022  and  2021,  respectively,  and  reflected  those  costs  in  research  and  development  in  the  statements  of  operations  and
comprehensive loss.

Royalty Entitlements

Alnylam Pharmaceuticals, Inc. and Acuitas Therapeutics, Inc.

The Company has two royalty entitlements to Alnylam’s global net sales of ONPATTRO.

95

In  2012,  the  Company  entered  into  a  license  agreement  with  Alnylam  Pharmaceuticals,  Inc.  (“Alnylam”)  that  entitles  Alnylam  to  develop  and
commercialize products with the Company’s LNP technology. Alnylam’s ONPATTRO, which represents the first approved application of the Company’s
LNP  technology,  was  launched  by  Alnylam  in  2018.  Under  the  terms  of  this  license  agreement,  the  Company  is  entitled  to  tiered  royalty  payments  on
global net sales of ONPATTRO ranging from 1.00% - 2.33% after offsets, with the highest tier applicable to annual net sales above $500 million. This
royalty  interest  was  sold  to  OMERS,  effective  as  of  January  1,  2019,  for  $20  million  in  gross  proceeds  before  advisory  fees.  OMERS  will  retain  this
entitlement until it has received $30 million in royalties, at which point 100% of this royalty entitlement on future global net sales of ONPATTRO will
revert back to the Company. OMERS has assumed the risk of collecting up to $30.0 million of future royalty payments from Alnylam and the Company is
not obligated to reimburse OMERS if they fail to collect any such future royalties. If this royalty entitlement reverts to the Company, it has the potential to
provide an active royalty stream or to be otherwise monetized again in full or in part. From the inception of the royalty sale through December 31, 2022, an
aggregate of $18.9 million of royalties have been earned by OMERS. See note 9 for further details.

The Company also has rights to a second royalty interest ranging from 0.75% to 1.125% on global net sales of ONPATTRO, with 0.75% applying to sales
greater  than  $500  million,  originating  from  a  settlement  agreement  and  subsequent  license  agreement  with  Acuitas  Therapeutics,  Inc.  (“Acuitas”).  This
royalty entitlement from Acuitas has been retained by the Company and was not part of the royalty entitlement sale to OMERS.

Gritstone Oncology, Inc.

On October 16, 2017, the Company entered into a license agreement with Gritstone that granted them worldwide access to its portfolio of proprietary and
clinically  validated  LNP  technology  and  associated  intellectual  property  to  deliver  Gritstone’s  self-replicating,  non-mRNA,  RNA-based  neoantigen
immunotherapy  products.  Gritstone  paid  the  Company  an  upfront  payment,  and  will  make  payments  for  achievement  of  development,  regulatory,  and
commercial milestones and royalties. As a result of the Company’s agreement with Genevant (see note 5 for details), from April 11, 2018 going forward,
Genevant is entitled to 50% of the revenues earned by the Company from Gritstone. The Company is the agent in this arrangement and records revenue on
a net basis. Milestone  payments  that  are  not  within  the  control  of  the  Company  or  the  licensee,  such  as  those  that  require  regulatory  approvals,  are  not
considered  probable  of  being  achieved  until  those  approvals  are  received.  The  Company  did  not  receive  any  payments  from  Gritstone  during  the  years
ended December 31, 2022 or 2021.

Revenues from the Company’s royalty entitlements are summarized in the following table:

Revenue from collaborations and licenses

Royalties from sales of Onpattro
Qilu Pharmaceutical Co., Ltd.
Other milestone and royalty payments

Non-cash royalty revenue

Royalties from sales of Onpattro

Total revenue

12.    Shareholders’ equity

Authorized share capital

Year ended December 31,

2022

2021

(in thousands)

$

$

5,316  $

26,015 
35 

7,653 
39,019  $

4,675 
— 
205 

6,108 
10,988 

The Company’s authorized share capital consists of an unlimited number of common shares and preferred shares, without par value, and 1,164,000 Series
A participating convertible preferred shares, without par value.

96

  
 
Open Market Sale Agreement

The Company has an Open Market Sale Agreement with Jefferies LLC (“Jefferies”) dated December 20, 2018, as amended by Amendment No. 1, dated
December 20, 2019, Amendment No. 2, dated August 7, 2020 and Amendment No. 3, dated March 4, 2021 (as amended, the “Sale Agreement”), under
which the Company may issue and sell common shares, from time to time.

On December 23, 2019, the Company filed a shelf registration statement on Form S-3 with the Securities and Exchange Commission (the “SEC”) (File No.
333-235674)  and  accompanying  base  prospectus,  which  was  declared  effective  by  the  SEC  on  January  10,  2020  (the  “January  2020  Registration
Statement”),  for  the  offer  and  sale  of  up  to  $150.0  million  of  the  Company’s  securities.  The  January  2020  Registration  Statement  also  contained  a
prospectus  supplement  in  connection  with  the  offering  of  up  to  $50.0  million  of  the  Company’s  common  shares  pursuant  to  the  Sale  Agreement.  This
prospectus supplement was fully utilized during 2020. On August 7, 2020, the Company filed a prospectus supplement with the SEC (the “August 2020
Prospectus Supplement”) in connection with the offering of up to an additional $75.0 million of its common shares pursuant to the Sale Agreement under
the January 2020 Registration Statement. The August 2020 Prospectus Supplement was fully utilized during 2020.

On  August  28,  2020,  the  Company  filed  a  shelf  registration  statement  on  Form  S-3  with  the  SEC  (File  No.  333-248467)  and  accompanying  base
prospectus, which was declared effective by the SEC on October 22, 2020 (the “October 2020 Registration Statement”), for the offer and sale of up to
$200.0 million of the Company’s securities. On March 4, 2021, the Company filed a prospectus supplement with the SEC (the “March 2021 Prospectus
Supplement”) in connection with the offering of up to an additional $75.0 million of its common shares pursuant to the Sale Agreement under October
2020 Registration Statement. The March 2021 Prospectus Supplement was fully utilized during 2021. On October 8, 2021, the Company filed a prospectus
supplement with the SEC (the “October 2021 Prospectus Supplement”) in connection with the offering of up to an additional $75.0 million of its common
shares pursuant to the Sale Agreement under the October 2020 Registration Statement.

On  November  4,  2021,  the  Company  filed  a  shelf  registration  statement  on  Form  S-3  with  the  SEC  (File  No.  333-260782)  and  accompanying  base
prospectus,  declared  effective  by  the  SEC  on  November  18,  2021  (the  “November  2021  Registration  Statement”),  for  the  offer  and  sale  of  up  to
$250.0 million of the Company’s securities.

On March 3, 2022, the Company filed a prospectus supplement with the SEC (the “March 2022 Prospectus Supplement”) in connection with the offering of
up to an additional $100.0 million of its common shares pursuant to the Sale Agreement under: (i) the January 2020 Registration Statement; (ii) the October
2020 Registration Statement; and (iii) the November 2021 Registration Statement.

During  the  years  ended  December  31,  2022  and  2021,  the  Company  issued  8,645,426  and  31,571,036  common  shares,  respectively,  under  the  Sale
Agreement, resulting in net proceeds of approximately $20.3 million and $134.7 million, respectively.

As of December 31, 2022, there was approximately $131.1 million remaining available in aggregate under the October 2021 Prospectus Supplement and
the March 2022 Prospectus Supplement.

Series A Preferred Shares

In October 2017, the Company entered into a subscription agreement with Roivant for the sale of Preferred Shares to Roivant for gross proceeds of $116.4
million. The Preferred Shares were non-voting and were convertible into common shares at a conversion price of $7.13 per share (which represented a 15%
premium to the closing price of $6.20 per share). The purchase price for the Preferred Shares plus an amount equal to 8.75% per annum, compounded
annually,  was  subject  to  mandatory  conversion  into  common  shares  on  October  18,  2021,  at  which  time  the  Preferred  Shares  were  converted  into
22,833,922 common shares and both the lockup and standstill periods that Roivant had previously agreed to expired. As of December 31, 2022, Roivant
owned approximately 25% of the Company’s outstanding common shares.

The Company recorded the Preferred Shares wholly as equity with no bifurcation of conversion feature from the host contract, given that the Preferred
Shares  could  not  be  cash  settled  and  the  redemption  features  were  within  the  Company’s  control,  which  included  a  fixed  conversion  ratio  with
predetermined timing and proceeds. The Company accrued for the 8.75% per annum

97

compounding coupon at each reporting period end date as an increase to share capital, and an increase to deficit (see statement of stockholder’s equity).

13.    Stock-based compensation

Awards outstanding and available for issuance

During  the  year  ended  December  31,  2022,  the  Company  had  stock  options  outstanding  under  the  following  plans  (collectively,  the  “Plans”):  the  2016
Omnibus Share and Incentive Plan (the “2016 Plan”), the 2011 Omnibus Share Compensation Plan (the “2011 Plan”); the 2019 inducement grant; and the
OnCore Option Plan.

As of December 31, 2022, the aggregate number of shares authorized for awards under all Plans was 28,290,202. As of December 31, 2022, the Company
had 15,450,598 options outstanding and 8,842,931 awards available for issuance under the Plans. 

The Company issues new common shares of stock to settle options exercised.

The 2011 Plan expired in June 2021. Under the 2016 Plan, the Company’s board of directors may grant options, and other types of awards, to employees,
directors and consultants of the Company.  The exercise price of the options is determined by the Company’s board of directors but will be at least equal to
the closing market price of the common shares on the date of grant and the term may not exceed 10 years.  Options granted generally vest over four years
for employees and for directors’ initial grants, and immediately for directors’ annual grants.

In June 2019, the Company provided an inducement grant of 1,112,000 options to its newly hired Chief Executive Officer. These options were awarded in a
separate plan as non-qualified awards and are governed by the substantially the same terms as the 2016 Plan.

Hereafter,  information  on  options  governed  by  the  2016  Plan,  the  2011  Plan  and  the  2019  inducement  grant  (the  “Arbutus  Plans”)  is  presented  on  a
consolidated basis as the terms of the plans are similar. Information on the OnCore Option Plan is presented separately.

98

Stock options under the Arbutus Plans

Equity-classified stock options under the Arbutus Plans

The following table summarizes activity related to the Company’s equity-classified stock options, including its performance options, for the year ended
December 31, 2022:

Stock Options Outstanding

Vested Stock Options

Non-Vested Stock Options

Number

Weighted-Average
Exercise Price

Number

Number

Balance as of December 31, 2021
Options granted
Options exercised
Options forfeited, canceled or expired
Options vested
Balance as of December 31, 2022

11,309,974  $
4,808,295  $
(71,025) $
(697,246) $
—  $
15,349,998  $

4.14 
2.77 
1.70 
3.31 
— 
3.76 

6,544,348 
— 
(71,025)
(100,399)
3,258,855 
9,631,779 

Weighted-Average
Grant-Date Fair Value
2.71 
2.09 
— 
2.44 
2.42 
2.39 

4,765,626  $
4,808,295  $
—  $
(596,847) $
(3,258,855) $
5,718,219  $

The intrinsic value of options exercised under the Arbutus Plans during 2022 and 2021 are $0.1 million and $0.2 million, respectively.

The following table summarizes additional information related to the Company’s equity-classified stock options, including its performance options, as of
December 31, 2022:

As of December 31, 2022

Options outstanding and expected to vest
Number of stock options outstanding
Weighted-average exercise price
Intrinsic value (in $000s)
Weighted-average term remaining

Vested stock options

Number of vested stock options
Weighted-average exercise price
Intrinsic value (in $000s)
Weighted-average term remaining

$
$

$
$

15,349,998 
3.76 
380 
7.3 years

9,631,779 
4.07 
327 
6.6 years

The assumptions used in the Black-Scholes option-pricing for grants made during the years ended December 31, 2022 and 2021 are as follows:

Expected average option term
Expected volatility
Expected dividends
Risk-free interest rate

December 31, 2022

December 31, 2021

5.5 years
97.0 %
— %
1.77 %

5.6 years
93.4 %
— %
0.67 %

The Company considers all available information when estimating the fair value of its stock option grants.

99

Stock options under the other plans

As of December 31, 2022, the Company also has 20,000 liability option awards outstanding with a weighted average exercise price of $12.10 and 80,600
stock option awards outstanding under the OnCore Option Plan with a weighted average exercise price of $0.56.

Employee Stock Purchase Plan

In May 2020, the Company’s stockholders approved the 2020 Employee Stock Purchase Plan (the “ESPP”) which became effective on May 28, 2020. A
total of 1,500,000 common shares were reserved for issuance under the ESPP. Company employees contribute funds via payroll deductions, which are used
to buy Company common shares at a discount of up to 15% based on the lower of the price at the start of the offering period and at the end of the relevant
purchase period within such offering period. The initial offering period under the ESPP was September 1, 2020 through August 31, 2021 with purchase
dates set on February 26, 2021 and August 31, 2021, with subsequent offering periods beginning on September 1 and ending on August 31. The Company
issued 171,224 and 196,335 shares under its ESPP for the years ended December 31, 2022 and 2021, respectively. As of December 31, 2022, there were
1,132,441 shares remaining for issuance under the ESPP. For the years ended December 31, 2022 and 2021, the Company recognized $0.2 million and $0.3
million, respectively, of stock-based compensation expense related to the ESPP. The fair value of the right to acquire stock at a discounted price under the
ESPP  is  calculated  using  the  Black-Scholes  valuation  model  and  recorded  as  stock-based  compensation.  Expense  is  recognized  over  the  period  the
employee contributes to the plan through payroll deductions.

Stock-based compensation expense

Total stock-based compensation expense was comprised of: (1) vesting of options awarded to employees under the Arbutus and OnCore Plans calculated in
accordance  with  the  fair  value  method  as  described  above;  (2)  fair  value  adjustments  for  the  Company’s  liability-classified  stock  options;  and  (3)
amortization of compensation cost related to the ESPP.

The Company recognizes forfeitures as they occur, and the effects of forfeitures are reflected in stock-based compensation expense.

Stock-based compensation has been recorded in the consolidated statement of operations and comprehensive loss as follows:

Research and development
General and administrative

Total

Year Ended December 31,

2022

2021

(in thousands)

2,912  $
4,270 
7,182  $

2,777 
3,647 
6,424 

$

$

At  December  31,  2022,  there  remained  $13.2  million  of  unrecognized  compensation  expense  related  to  unvested  equity  employee  stock  options  to  be
recognized as expense over a weighted-average period of approximately 2.4 years.

For each of the years ended December 31, 2022 and 2021, the Company had zero performance based stock compensation expense.

14.    Income taxes

The Company is subject to taxation and files income tax returns in Canadian federal and provincial, United States federal and several state jurisdictions. In
December 2022, the United States Internal Revenue service completed its examination of the Company’s federal tax return for 2018. In May 2022, The
Canada Revenue Agency completed its examination of the Company’s Canadian tax returns for 2018 and 2019, with no adjustments proposed.

100

 
 
Income tax expense varies from the amounts that would be computed by applying the combined Canadian federal and provincial income tax rate of 27%
(2021 - 27%) to the loss before income taxes as shown in the following tables:

Computed taxes (benefits) at Canadian federal and provincial tax rates
Withholding taxes
Other
Permanent and other differences
Foreign tax credit applied
Federal and Provincial ITCs applied
Change in valuation allowance
Difference due to income taxed at foreign rates
Stock-based compensation
Income tax expense

Year ended December 31,

2022

2021

(in thousands)

$

$

(17,554) $
4,444 
761 
869 
(4,444)
(324)
14,563 
5,625 
504 
4,444  $

(23,864)
— 
(1,041)
4,292 
— 
(611)
15,928 
4,840 
456 
— 

As of December 31, 2022, the Company had investment tax credits available to reduce Canadian federal income taxes of $7.2 million, versus $7.4 million
as of December 31, 2021, which expire between 2030 and 2037, and provincial income taxes of $2.0 million, versus $2.1 million as of December 31, 2021,
which expire between 2024 and 2027. The investment tax credits are accounted for under a flow-through method. In addition, the Company had research
and development credits of $3.7 million as of December 31, 2022, and $3.8 million as of December 31, 2021, which expire between 2031 and 2038 and
which can be used to reduce future taxable income in the United States.

As of December 31, 2022, the Company had scientific research and experimental development expenditures of $62.2 million available for indefinite carry-
forward, versus $62.8 million as of December 31, 2021. The Company also had net operating losses of $148.1 million as of December 31, 2022 and $177.7
million as of December 31, 2021, which are due to expire between 2028 and 2038 and which can be used to offset future taxable income in Canada.

As of December 31, 2022 and December 31, 2021, the Company had $11.7 million of net operating losses due to expire in 2035 which can be used to offset
future taxable income in the United States. Future use of a portion of the United States loss carryforwards are subject to limitations under Internal Revenue
Code  Section  382.  United  States  net  operating  loss  carryforwards  arising  in  2019  and  future  periods  have  an  indefinite  carryforward  period.  As  of
December 31, 2022 and December 31, 2021, the Company had $203.9 million and $197.8 million, respectively, of total regular net operating losses which
can be used to offset future taxable income in the United States.

As  a  result  of  ownership  changes  occurring  on  October  1,  2014  and  March  4,  2015,  the  Company’s  ability  to  use  these  losses  may  be  limited.  Losses
incurred to date may be further limited if a subsequent change in control occurs.

The Company generated $28.7 million and $93.7 million in pre-tax domestic and foreign losses, respectively, for the year ended December 31, 2022. The
Company generated $7.7 million and $80.7 million in pre-tax domestic and foreign losses, respectively, for the year ended December 31, 2021.

As required by the 2017 Tax Cuts and Jobs Act and effective in 2022, the deferred tax asset as of December 31, 2022 included $16.5 million related to the
mandatory capitalization and amortization of research and development expenses.

101

 
 
Significant components of the Company’s deferred tax assets and liabilities are shown below:

Deferred tax assets (liabilities):

Non-capital losses carryforwards
Canadian research and development deductions
Book amortization in excess of tax
Revenue recognized for tax purposes in excess of revenue recognized for accounting purposes
Tax value in excess of accounting value in lease inducements
Deferred revenue
Canadian Federal investment tax credits
Canadian Provincial investment tax credits
Equity accounted for investment
U.S. Federal research and development credits
Deductible stock options
U.S. research and experimental expenditures capitalization
Accrued interest payable
Amortization
Other

Total deferred tax assets
Valuation allowance

Net deferred tax assets (liabilities)

15.    Related party transactions

As of December 31,

2022

2021

(in thousands)

$

$

$

83,564  $
16,791 
(461)
2,799 
93 
6,063 
5,278 
1,953 
3,375 
3,633 
3,681 
16,471 
1,507 
387 
153 
145,287  $
(145,287)

—  $

90,255 
16,968 
(634)
4,400 
549 
— 
5,301 
2,119 
3,375 
3,741 
3,309 
— 
— 
— 
1,341 
130,724 
(130,724)
— 

Pursuant to a financing and related subscription agreement, the Company issued Roivant the Preferred Shares in October 2017. On October 18, 2021, the
Preferred  Shares  were  converted  into  22,833,922  common  shares.  As  of  December  31,  2022,  Roivant  owned  approximately  25%  of  the  Company’s
outstanding common shares. See note 12 for further details.

As  of  December  31,  2022,  the  carrying  value  of  the  Company’s  investment  in  Genevant  was  zero  and  the  Company  owned  approximately  16%  of  the
common equity of Genevant. See note 5 for further details.

During  each  of  the  years  ended  December  31,  2022  and  2021,  Genevant  purchased  certain  administrative  and  transitional  services  from  the  Company
totaling less than $0.1 million. These services were billed at agreed hourly rates and reflective of market rates for such services and these costs were netted
in research and development in the income statement.

102

 
 
 
 
Item 9. Changes in and Disagreements with Accountants on Accounting and Financial Disclosure

None.

Item 9A. Controls and Procedures

Disclosure Controls and Procedures

Our management, including our principal executive officer and principal financial officer, has evaluated the effectiveness of the design and operation of our
disclosure controls and procedures (as defined in Rules 13a-15(e) and 15d-15(e) under the Securities Exchange Act of 1934, as amended (the “Exchange
Act”)), as of the end of the period covered by this Annual Report on Form 10-K. Based upon that evaluation, our Chief Executive Officer (our principal
executive  officer)  and  Chief  Financial  Officer  (our  principal  financial  officer),  concluded  that,  as  of  December  31,  2022,  our  disclosure  controls  and
procedures were effective to provide reasonable assurance that (a) the information required to be disclosed by us in the reports that we file or submit under
the Exchange Act is recorded, processed, summarized and reported within the time periods specified in the SEC’s rules and forms, and (b) such information
is accumulated and communicated to our management, including our Chief Executive Officer and Chief Financial Officer, as appropriate, to allow timely
decisions regarding required disclosure.

In  designing  and  evaluating  our  disclosure  controls  and  procedures,  our  management  recognized  that  any  controls  and  procedures,  no  matter  how  well
designed and operated, can provide only reasonable assurance of achieving the desired control objectives, and our management necessarily was required to
apply its judgment in evaluating the cost-benefit relationship of possible controls and procedures.

Management’s Annual Report on Internal Control over Financial Reporting

Management is responsible for establishing and maintaining adequate internal control over financial reporting, as such term is defined in Exchange Act
Rules  13a-15(f)  and  15d-15(f).  Under  the  supervision  and  with  the  participation  of  our  management,  including  our  principal  executive  officer  and  our
principal financial officer, we conducted an evaluation of the effectiveness of our internal control over financial reporting based on the framework in the
Internal Control - Integrated Framework (2013) issued by the Committee of Sponsoring Organizations of the Treadway Commission (“COSO 2013”).

Our internal control over financial reporting is a process designed to provide reasonable assurance regarding the reliability of financial reporting and the
preparation of financial statements for external purposes in accordance with generally accepted accounting principles in the United States of America. Our
internal  control  over  financial  reporting  includes  those  policies  and  procedures  that  (i)  pertain  to  the  maintenance  of  records  that,  in  reasonable  detail,
accurately and fairly reflect the transactions and dispositions of our assets; (ii) provide reasonable assurance that transactions are recorded as necessary to
permit preparation of financial statements in accordance with generally accepted accounting principles, and our receipts and expenditures are being made
only in accordance with authorizations of our management and directors; and (iii) provide reasonable assurance regarding prevention or timely detection of
unauthorized acquisition, use, or disposition of our assets that could have a material effect on the financial statements.

Because of its inherent limitations, internal control over financial reporting may not prevent or detect misstatements. Also, projections of any evaluation of
effectiveness  to  future  periods  are  subject  to  the  risk  that  controls  may  become  inadequate  because  of  changes  in  conditions,  or  that  the  degree  of
compliance with the policies or procedures may deteriorate. Based on our evaluation under the framework in COSO 2013, our management concluded that
our internal control over financial reporting was effective as of December 31, 2022.

Changes in Internal Control over Financial Reporting

There have not been changes in our internal control over financial reporting during the quarter ended December 31, 2022 that have materially affected or
are reasonably likely to materially affect the Company’s internal control over financial reporting.

103

 
 
 
Item 9B. Other Information

None.

Item 9C. Disclosure Regarding Foreign Jurisdictions that Prevent Inspections.    

Not applicable.

104

Item 10. Directors, Executive Officers and Corporate Governance

PART III

The information required by this item is incorporated herein by reference to our Proxy Statement for the 2023 Annual General Meeting of the Stockholders
to be filed with the SEC within 120 days of the fiscal year ended December 31, 2022.

We  have  adopted  a  code  of  business  conduct  for  directors,  officers  and  employees  (the  “Code  of  Conduct”),  which  is  available  on  our  website  at
http://investor.arbutusbio.com/corporate-governance-0 and also at www.sedar.com. We intend to satisfy the disclosure requirement under Item 5.05 of Form
8-K regarding any amendment to, or waiver from, a provision of this Code of Conduct by posting such information on the website address and location
specified above.

Item 11. Executive Compensation

The information required by this item is incorporated herein by reference to our Proxy Statement for the 2023 Annual General Meeting of the Stockholders
to be filed with the SEC within 120 days of the fiscal year ended December 31, 2022.

Item 12. Security Ownership of Certain Beneficial Owners and Management and Related Stockholder Matters

The information required by this item is incorporated herein by reference to our Proxy Statement for the 2023 Annual General Meeting of the Stockholders
to be filed with the SEC within 120 days of the fiscal year ended December 31, 2022.

Item 13. Certain Relationships and Related Transactions, and Director Independence

The information required by this item is incorporated herein by reference to our Proxy Statement for the 2023 Annual General Meeting of the Stockholders
to be filed with the SEC within 120 days of the fiscal year ended December 31, 2022.

Item 14. Principal Accounting Fees and Services

The information required by this item is incorporated herein by reference to our Proxy Statement for the 2023 Annual General Meeting of the Stockholders
to be filed with the SEC within 120 days of the fiscal year ended December 31, 2022.

105

Item 15. Exhibits and Financial Statement Schedules

PART IV

Exhibit

2.1*

3.1*

3.2*

4.1**

10.1†*

10.2†*

10.3†*

10.4*#

10.5†*

10.6†*

10.7†*

10.8†*

10.9*#

  Description

Agreement and Plan of Merger and Reorganization, dated January 11, 2015, by and among Tekmira Pharmaceuticals Corporation, TKM
Acquisition Corporation and OnCore Biopharma, Inc. (incorporated herein by reference to Exhibit 2.1 to the Registrant’s Current
Report on Form 8-K/A, filed with the SEC on January 26, 2015).

Notice of Articles and Articles of the Company, as amended (incorporated herein by reference to Exhibit 3.1 to the Registrant’s Annual
Report on Form 10-K for the year ended December 31, 2017, filed with the SEC on March 16, 2018).

Amendment to Articles of the Company (incorporated herein by reference to Exhibit 3.1 to the Registrant’s Quarterly Report on Form
10-Q for the quarter ended September 30, 2018, filed with the SEC on November 7, 2018).

Description of Securities Registered Pursuant to Section 12 of the Securities Exchange Act of 1934.

Amended and Restated License Agreement, between Inex Pharmaceuticals Corporation and Hana Biosciences, Inc., dated April 30,
2007 (incorporated herein by reference to Exhibit 4.2 to the Registrant’s Amendment No. 1 to Form 20-F for the year ended December
31, 2010, filed with the SEC on January 31, 2012).

Amendment No. 1 to the Amended and Restated Agreement, between the Company (formerly Inex Pharmaceuticals Corporation) and
Hana Biosciences, Inc., effective as of May 27, 2009 (incorporated herein by reference to Exhibit 4.1 to the Registrant’s Annual Report
on Form 20-F for the year ended December 31, 2010, filed with the SEC on June 3, 2011).

Amendment No. 2 to the Amended and Restated Agreement, between the Company (formerly Inex Pharmaceuticals Corporation) and
Hana Biosciences, Inc., effective as of September 20, 2010 (incorporated herein by reference to Exhibit 4.21 to the Registrant’s Annual
Report on Form 20-F for the year ended December 31, 2010, filed with the SEC on June 3, 2011).

Form of Arbutus Biopharma Corporation Indemnity Agreement (incorporated herein by reference to Exhibit 10.4 to the Registrant’s
Annual Report on Form 10-K for the year ended December 31, 2022, filed with the SEC on March 3, 2022).

License Agreement between the University of British Columbia and Inex Pharmaceuticals Corporation executed on July 30, 2001
(incorporated herein by reference to Exhibit 4.17 to the Registrant’s Annual Report on Form 20-F for the year ended December 31,
2010, filed with the SEC on June 3, 2011).

Amendment Agreement between the University of British Columbia and Inex Pharmaceuticals Corporation dated July 11, 2006
(incorporated herein by reference to Exhibit 4.18 to the Registrant’s Annual Report on Form 20-F for the year ended December 31,
2010, filed with the SEC on June 3, 2011).

Second Amendment Agreement between the University of British Columbia and Inex Pharmaceuticals Corporation dated January 8,
2007 (incorporated herein by reference to Exhibit 4.19 to the Registrant’s Annual Report on Form 20-F for the year ended December
31, 2010, filed with the SEC on June 3, 2011).

Consent Agreement of the University of British Columbia to Inex/Alnylam Sublicense Agreement dated January 8, 2007 (incorporated
herein by reference to Exhibit 4.20 to the Registrant’s Annual Report on Form 20-F for the year ended December 31, 2010, filed with
the SEC on June 3, 2011).

Tekmira 2011 Omnibus Share Compensation Plan approved by shareholders on June 22, 2011 (incorporated herein by reference to
Exhibit 4.25 to the Registrant’s Annual Report on Form 20-F for the year ended December 31, 2011, filed with the SEC on March 27,
2012).

10.10†*

Settlement Agreement and General Release, by and among Tekmira Pharmaceuticals Corporation, Protiva Biotherapeutics Inc.,
Alnylam Pharmaceuticals, Inc., and AlCana Technologies, Inc., dated November 12, 2012 (incorporated herein by reference to Exhibit
4.26 to the Registrant’s Annual Report on Form 20-F for the year ended December 31, 2012, filed with the SEC on March 27, 2013).

106

10.11†*

10.12*

10.13*#

10.14*#

10.15†*

10.16*#

10.17*#

10.18†*

10.19†*

10.20*

10.21*

10.22*

10.23*

10.24*

10.25*

10.26*#

Cross-License Agreement by and among Alnylam Pharmaceuticals, Inc., Tekmira Pharmaceuticals Corporation and Protiva
Biotherapeutics Inc., dated November 12, 2012 (incorporated herein by reference to Exhibit 4.27 to the Registrant’s Annual Report on
Form 20-F for the year ended December 31, 2012, filed with the SEC on March 27, 2013).

Form of Standstill Agreement (incorporated herein by reference to Exhibit 2.1 to the Registrant’s Current Report on Form 8-K/A, filed
with the SEC on January 26, 2015).

Executive Employment Agreement, dated effective as of February 25, 2016, between Arbutus Biopharma, Inc. and Elizabeth Howard
(incorporated herein by reference to Exhibit 10.78 to the Registrant’s Annual Report on Form 10-K for the year ended December 31,
2015, filed with the SEC on March 9, 2016).

Amending Agreement, dated as of November 2, 2015, among Arbutus Biopharma Corporation, Roivant Sciences Ltd., Patrick T.
Higgins, Michael J. McElhaugh, Michael J. Sofia and Bryce A. Roberts (incorporated herein by reference to Exhibit 10.3 to the
Registrant’s Quarterly Report on Form 10-Q for the quarter ended September 30, 2015, filed with the SEC on November 5, 2015).

Stock Purchase Agreement by and among OnCore Biopharma, Inc. and each of the stockholders of Enantigen Therapeutics, Inc., dated
as of October 1, 2014 (incorporated herein by reference to Exhibit 10.3 to the Registrant’s Quarterly Report on Form 10-Q for the
quarter ended March 31, 2015, filed with the SEC on May 6, 2015).

Executive Employment Agreement, dated effective as of July 11, 2015, between OnCore Biopharma, Inc. and Michael J. Sofia
(incorporated herein by reference to Exhibit 10.8 to the Registrant’s Quarterly Report on Form 10-Q for the quarter ended June 30,
2015, filed with the SEC on August 7, 2015).

Amended 2011 Omnibus Share Compensation Plan (incorporated herein by reference to Exhibit 10.1 to the Registrant’s Quarterly
Report on Form 10-Q for the quarter ended June 30, 2016, filed with the SEC on August 4, 2016).

Lease Agreement between the Company and ARE-PA Region No. 7, LLC dated August 9, 2016 (incorporated herein by reference to
Exhibit 10.1 to the Registrant’s Quarterly Report on Form 10-Q for the quarter ended September 30, 2016, filed with the SEC on
November 3, 2016).

First Amendment to Lease Agreement between Arbutus Biopharma, Inc. and ARE-PA Region No. 7, LLC dated October 7, 2016
(incorporated herein by reference to Exhibit 10.2 to the Registrant’s Quarterly Report on Form 10-Q for the quarter ended September
30, 2016, filed with the SEC on November 3, 2016).

Acknowledgment of Commencement Date in connection with Lease Agreement between the Company and ARE-PA Region No. 7,
LLC dated August 9, 2016 and as amended on October, 7, 2016 (incorporated herein by reference to Exhibit 10.3 to the Registrant’s
Quarterly Report on Form 10-Q for the quarter ended September 30, 2016, filed with the SEC on November 3, 2016).

Master Contribution And Share Subscription Agreement, by and between the Company, Genevant Sciences Ltd. and Roivant Sciences
LTD. (incorporated herein by reference Exhibit 10.1 to the Registrant's Quarterly Report on Form 10-Q for the quarter ended March 31,
2018, filed with the SEC on May 4, 2018).

Open Market Sale AgreementSM, dated December 20, 2018, by and between the Company and Jefferies LLC (incorporated herein by
reference to Exhibit 1.1 of the Current Report on Form 8-K, filed with the SEC on December 20, 2018).

Amendment No. 1 to the Open Market Sale AgreementSM, dated December 20, 2019, by and between the Company and Jefferies LLC
(incorporated herein by reference to Exhibit 1.3 to the Registrant’s Registration Statement on Form S-3, filed with the SEC on
December 20, 2019).

Amendment No. 2 to the Open Market Sale AgreementSM, dated August 7, 2020, by and between the Company and Jefferies LLC
(incorporated herein by reference to Exhibit 1.1 to the Registrant’s Current Report on Form 8-K, filed with the SEC on August 7, 2020).

Amendment No. 3 to the Open Market Sale AgreementSM, dated March 4, 2021, by and between Arbutus Biopharma Corporation and
Jefferies LLC (incorporated herein by reference to Exhibit 10.1 to the Registrant’s Current Report on Form 8-K, filed with the SEC on
March 4, 2021.)

Executive Employment Agreement, dated June 11, 2018, by and between the Company and David Hastings (incorporated herein by
reference to Exhibit 10.52 of the Registrant’s Annual Report on Form 10-K for the year end December 31, 2018, filed with the SEC on
March 7, 2019).

107

10.27*#

10.28*#

10.29*

10.30*#

10.31*#

10.32*#

10.33†*

10.34†*

10.35†*

10.36†*

10.37†*

10.38*

10.39*#

10.41**

21.1**

23.1**

31.1**

Employment Agreement, dated June 13, 2019, by and between the Company and William H. Collier (incorporated herein by reference
to Exhibit 10.3 to the Registrant’s Current Report on Form 8-K, filed with the SEC on June 18, 2019).

Executive Employment Agreement, dated July 10, 2015, by and between the Company and Michael McElhaugh, as amended by the
First Amendment to Executive Employment Agreement, dated April 20, 2016, and the Second Amendment to Executive Employment
Agreement dated December 11, 2018 (incorporated herein by reference to Exhibit 10.5 to the Registrant’s Quarterly Report on Form
10-Q for the quarter ended June 30, 2019, filed with the SEC on August 5, 2019).

Purchase and Sale Agreement, dated July 2, 2019, by and between the Company and OCM IP Healthcare Portfolio LP (incorporated
herein by reference to Exhibit 10.6 to the Registrant’s Quarterly Report on Form 10-Q for the quarter ended June 30, 2019, filed with
the SEC on August 5, 2019).

Arbutus Biopharma Corporation 2020 Employee Stock Purchase Plan (incorporated herein by reference to Exhibit 10.2 to the
Registrant’s Current Report on Form 8-K, filed with the SEC on June 1, 2020).

Form of Arbutus Biopharma Corporation Option Agreement (incorporated herein by reference to Exhibit 10.8 to the Registrant’s
Quarterly Report on Form 10-Q for the quarter ended June 30, 2019, filed with the SEC on August 5, 2019).

Option Agreement, dated June 24, by and between the Company and William H. Collier (incorporated herein by reference to Exhibit
10.9 to the Registrant’s Quarterly Report on Form 10-Q for the quarter ended June 30, 2019, filed with the SEC on August 5, 2019).

Cross License Agreement, dated April 11, 2018, by and between the Company and Genevant Sciences Ltd. (incorporated herein by
reference to Exhibit 10.3 to the Registrant’s Quarterly Report on Form 10-Q for the quarter ended June 30, 2020, filed with the SEC on
August 7, 2020).

First Amendment to Cross License Agreement, dated June 27, 2018, by and among the Company, Genevant Sciences Ltd and Genevant
Sciences GmbH. (incorporated herein by reference to Exhibit 10.4 to the Registrant’s Quarterly Report on Form 10-Q for the quarter
ended June 30, 2020, filed with the SEC on August 7, 2020).

Second Amendment to Cross License Agreement, dated June 27, 2018, by and among the Company, Genevant Sciences Ltd. and
Genevant Sciences GmbH. (incorporated herein by reference to Exhibit 10.5 to the Registrant’s Quarterly Report on Form 10-Q for the
quarter ended June 30, 2020, filed with the SEC on August 7, 2020).

License Agreement, dated December 9, 2021, by and between the Company and Genevant Sciences GmbH (incorporated herein by
reference to Exhibit 10.1 to the Registrant’s Current Report on Form 8-K, filed with the SEC on December 10, 2021).

Technology Transfer and Exclusive License Agreement, dated December 13, 2021, by and between the Company and Qilu
Pharmaceutical Co., Ltd. (incorporated herein by reference to Exhibit 10.41 to the Registrant’s Annual Report on Form 10-K for the
year ended December 31, 2022, filed with the SEC on March 3, 2022).

Arbutus Biopharma Corporation 2016 Omnibus Share and Incentive Plan, as supplemented and amended (incorporated herein by
reference to Exhibit 10.1 to the Registrant’s Current Report on Form 8-K, filed with the SEC on May 31, 2022).

Third Amendment to Executive Employment Agreement, dated November 1, 2022, by and between the Company and Michael
McElhaugh (incorporated herein by reference to Exhibit 10.1 to the Registrant’s Current Report on Form 8-K, filed with the SEC on
October 24, 2022)

Form of Arbutus Biopharma Corporation Restricted Stock Unit Agreement.

List of Subsidiaries.

Consent of Ernst and Young LLP, an Independent Registered Public Accounting Firm.

Certification of Chief Executive Officer pursuant to Rule 13a-14 or 15d-14 of the Securities Exchange Act of 1934, as adopted pursuant
to Section 302 of the Sarbanes-Oxley Act of 2002.

108

31.2**

32.1**

32.2**

Certification of Chief Financial Officer pursuant to Rule 13a-14 or 15d-14 of the Securities Exchange Act of 1934, as adopted pursuant
to Section 302 of the Sarbanes-Oxley Act of 2002.

Certification of Chief Executive Officer pursuant to 18 U.S.C. Section 1350, as adopted pursuant to Section 906 of the Sarbanes-Oxley
Act of 2002.

Certification of Chief Financial Officer pursuant to 18 U.S.C. Section 1350, as adopted pursuant to Section 906 of the Sarbanes-Oxley
Act of 2002.

101.INS**

  XBRL Instance Document

101.SCH**

  XBRL Taxonomy Extension Schema Document

101.CAL**

  XBRL Taxonomy Extension Calculation Linkbase Document

101.DEF**

  XBRL Taxonomy Extension Definition Linkbase Document

101.LAB**

  XBRL Taxonomy Extension Label Linkbase Document

101.PRE**

  XBRL Taxonomy Extension Presentation Linkbase Document

104**

Cover Page Interactive Data File (formatted as Inline XBRL and Contained in Exhibit 101).

*     Previously filed

**    Filed or furnished herewith, as applicable

†     Certain confidential portions of the agreement were omitted by means of marking such portions with brackets (due to the registrant customarily and

actually treating such information as private or confidential and such omitted information not being material) pursuant to Item 601 of Regulation
S-K promulgated by the SEC. Arbutus agrees to supplementally furnish a copy of any confidential portions to the SEC upon request.

#    Management Contract or Compensatory Arrangement.

Financial Statements

 See Index to Consolidated Financial Statements under Item 8 of Part II.

Financial Statement Schedules

 None.

Item 16.     Form 10-K Summary

    None.

109

Pursuant to the requirements of Section 13 or 15(d) of the Securities Exchange Act of 1934, the Registrant has duly caused this report to be signed on its
behalf by the undersigned, thereunto duly authorized on March 2, 2023.

SIGNATURES

ARBUTUS BIOPHARMA CORPORATION

By:

/s/ William H. Collier
William H. Collier
President and Chief Executive Officer

Pursuant to the requirements of the Securities Exchange Act of 1934, this report has been signed below by the following persons on behalf of the Registrant
and in the capacities indicated on March 2, 2023.

Signatures

/s/ Frank Torti, M.D.
Frank Torti, M.D.

/s/ William H. Collier
William H. Collier

/s/ David C. Hastings
David C. Hastings

/s/ Daniel Burgess
Daniel Burgess

/s/ Richard C. Henriques
Richard C. Henriques

/s/ Keith Manchester, M.D.
Keith Manchester, M.D.

/s/ James Meyers
James Meyers

Capacity in Which Signed

Director (Chairman)

President and Chief Executive Officer and Director
(Principal Executive Officer)

Chief Financial Officer
(Principal Financial Officer and Principal Accounting Officer)

Director

Director

Director

Director

110

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
DESCRIPTION OF THE REGISTRANT’S SECURITIES
REGISTERED PURSUANT TO SECTION 12 OF THE
SECURITIES EXCHANGE ACT OF 1934

Exhibit 4.1 

As of the date of the Annual Report on Form 10-K of which this exhibit forms a part, the only class of securities of Arbutus Biopharma Corporation
(“we,” “us” and “our”) registered under Section 12 of the Securities Exchange Act of 1934, as amended (the “Exchange Act”), is our common shares,
without par value.

CAPITAL STOCK

The  following  description  of  our  capital  stock  summarizes  provisions  of  our  Notice  of  Articles  and  Articles,  as  amended,  or  our  Articles,  the
Investment Canada Act (Canada), the Competition Act (Canada) and the Business Corporations Act (British Columbia). The following description does not
purport to be complete and is subject to, and qualified in its entirety by, our Articles, which are incorporated by reference as exhibits to the Annual Report
on  Form  10-K  of  which  this  exhibit  is  a  part,  and  to  the  applicable  provisions  of  the  Investment  Canada  Act,  the  Competition  Act  and  the  Business
Corporations Act.

Authorized and Outstanding Shares

Our authorized share capital consists of (i) an unlimited number of common shares, without par value, (ii) an unlimited number of preferred shares, without
par value, and (iii) 1,164,000 Series A Participating Convertible Preferred Shares. As of February 28, 2023 there were (a) 162,570,989 common shares
outstanding and (b) 0 Series A Participating Convertible Preferred Shares outstanding. None of our common shares or preferred shares are held by us or on
behalf of us.

Voting Rights

The holders of our common shares are entitled to receive notice of any meeting of our shareholders and to attend and vote thereat, except those meetings at
which only the holders of shares of another class or of a particular series are entitled to vote. Each common share entitles its holder to one vote. There are
no cumulative voting rights.

Dividends

Subject to the rights of the holders of preferred shares, the holders of common shares are entitled to receive on a pro rata basis such dividends as our Board
of Directors may declare out of funds legally available for payment of dividends.

Liquidation Rights

In the event of the dissolution, liquidation, winding-up or other distribution of our assets, those holders are entitled to receive on a pro rata basis all of our
assets remaining after payment of all of our liabilities, subject to the rights of holders of preferred shares.

Other Rights and Preferences.

The terms of our common shares do not include any preemptive, conversion or subscription rights, nor any redemption or sinking fund provisions. The
common shares are not subject to future calls or assessments by us.

Limitations to Control due to Certain Provisions of Canadian and British Columbian Law and our Articles

Unless such offer constitutes an exempt transaction, an offer made by a person, or an offeror, to acquire outstanding shares of a Canadian entity that, when
aggregated with the offeror’s holdings (and those of persons or companies acting jointly with the offeror), would constitute 20% or more of the outstanding
shares,  would  be  subject  to  the  take-over  provisions  of  Canadian  securities  laws.  The  foregoing  is  a  limited  and  general  summary  of  certain  aspects  of
applicable securities law in the provinces and territories of Canada, all in effect as of the date hereof.

In addition to the take-over bid requirements noted above, the acquisition of shares may trigger the application of additional statutory regimes including
amongst others, the Investment Canada Act (Canada) and the Competition Act (Canada).

As well, under the Business Corporations Act (British Columbia), unless otherwise stated in our Articles, certain corporate actions require the approval of a
special majority of shareholders, meaning holders of shares representing 66 2/3% of those votes cast in respect of a shareholder vote addressing such
matter. Those items requiring the approval of a special majority generally relate to fundamental changes with respect to our business, and include amongst
others, resolutions: (i) removing a

director prior to the expiry of his or her term; (ii) altering our Articles, (iii) approving an amalgamation; (iv) approving a plan of arrangement; and (v)
providing for a sale of all or substantially all of our assets.

The Nasdaq Global Select Market

Our common shares are listed on the Nasdaq Global Select Market under the symbol “ABUS.”

Transfer Agent and Registrar

The transfer agent and registrar for our common shares is TSX Trust Company.

Exhibit 10.41

ARBUTUS BIOPHARMA CORPORATION
2016 OMNIBUS SHARE AND INCENTIVE PLAN

RESTRICTED STOCK UNIT AGREEMENT

COVER SHEET

Arbutus Biopharma Corporation, a corporation incorporated under the laws of British Columbia, Canada (the “Company”),  hereby  grants
Restricted Stock Units (the “RSUs”) representing the right to receive common shares, without par value, of the Company (the “Shares”), to
the individual named below as Participant, subject to the vesting and other conditions set forth below. The terms and conditions of the RSUs
are set forth in this cover sheet, in the attachment (collectively, the “Agreement”) and in the Company’s 2016 Omnibus Share and Incentive
Plan (as it may be amended from time to time, the “Plan”).

Grant Date: ______________________     

Name of Participant: ______________________

Number of Shares Covered by RSUs: ______________________

Vesting Schedule:

The  RSUs  vest  in  three  equal  annual  installments  beginning  one  year  from  the  grant  date,  subject  to  the  Reporting  Person's  continuous
service as of each vesting date.

By signing this cover sheet, you agree to all of the terms and conditions described in this Agreement and in the Plan, a
copy of which is also attached. You acknowledge that you have carefully reviewed the Plan, and you agree that the Plan will
control in the event any provision of this Agreement should appear to be inconsistent. Certain capitalized terms used in this
Agreement are defined in the Plan, and have the respective meanings set forth in the Plan.

Grantee:        

(Signature)

Company:    ______________________________________________________

(Signature)

    Title: Chief Financial Officer

Attachment

This document is not a stock certificate or a negotiable instrument.

Restricted Stock Units

Vesting

Change in Control

Leaves of Absence

Issuance

ARBUTUS BIOPHARMA CORPORATION
2016 OMNIBUS SHARE AND INCENTIVE PLAN

RESTRICTED STOCK UNIT AGREEMENT

This Agreement evidences an award of RSUs in the number set forth on the cover sheet. Each RSU
represents the right to receive one Share, subject to the vesting and other conditions set forth in this
Agreement and in the Plan.
The RSUs shall vest in accordance with the vesting schedule set forth on the cover sheet of this
Agreement; provided, however, that for purposes of vesting, fractional numbers of Shares shall be
rounded to the nearest whole number, and the number of RSUs that shall vest on the final vesting
date  shall  be  rounded  up  or  down  as  necessary  such  that  the  total  number  of  RSUs  that  vest
pursuant to the vesting schedule shall be equal to the number of RSUs covered by this grant as set
forth on the cover sheet of this Agreement.

Unless  the  termination  of  your  service  as  an  Eligible  Person  (“Service”)  triggers  accelerated
vesting or other treatment of the RSUs pursuant to the terms of this Agreement or the Plan, you
shall immediately and automatically forfeit the unvested RSUs to the Company in the event your
Service terminates for any reason. No RSUs shall vest after your termination of Service.

In the event of a Change in Control, the RSUs will be treated in the manner provided in Section
7(b) of the Plan.
This section of the Agreement applies solely to Participants who are employees of the Company
or any of its subsidiaries. For purposes of the RSUs, your Service does not terminate when you go
on a bona fide  employee  leave  of  absence  that  was  approved  by  the  Company  in  writing,  if  the
terms of the leave provide for continued service crediting, or when continued service crediting is
required by applicable law. However, in all other cases, your Service will be treated as terminating
ninety (90) days after you went on employee leave, unless your right to return to active work is
guaranteed by law or by a contract. Your Service terminates in any event when the approved leave
ends, unless you immediately return to active employee work.

The  Company  determines,  in  its  sole  discretion,  which  leaves  count  for  this  purpose,  and  when
your Service terminates for all purposes under the Plan.

The  issuance  of  the  Shares  underlying  any  RSUs  that  become  vested  hereunder  shall  be  made
within thirty (30) days after the applicable vesting date. Any  such  issuance  shall  be  evidenced  in
such  a  manner  as  the  Company,  in  its  discretion,  will  deem  appropriate,  including,  without
limitation, book-entry, direct registration or issuance of one or more Share certificates.

2

Withholding Taxes

Transfer of RSUs

Retention Rights

Shareholder Rights

Clawback

Adjustments

In  order  to  comply  with  all  applicable  federal,  state,  local  or  foreign  tax  laws  or  regulations,  the
Company may take such actions as it deems appropriate to ensure that all applicable federal, state,
local or foreign payroll, withholding, income or other taxes that may be due relating to the RSUs
and  the  issuance  of  Shares  with  respect  to  the  RSUs,  which  are  your  sole  and  absolute
responsibility, are withheld or collected from you. If you are an employee of the Company or any
of its subsidiaries as of the Grant Date, then you hereby agree as a condition of this Agreement that
you  will  enter  into  a  side  letter  agreement  with  the  Company  prior  to  or  as  soon  as  practicable
following  the  Grant  Date  (or  at  such  other  time  as  directed  by  the  Company),  in  a  form  that  is
acceptable  to  the  Company,  pursuant  to  which  you  will  make  an  election  to  satisfy  all  tax
withholding obligations that arise hereunder pursuant to the “sell to cover” tax withholding method.
The RSUs are not transferable by you other than to a designated beneficiary upon your death or by
will or the laws of descent and distribution. No assignment or transfer of the RSUs, or the rights
represented thereby, whether voluntary or involuntary, by operation of law or otherwise (except to a
designated  beneficiary  upon  death  by  will  or  the  laws  of  descent  or  distribution)  will  vest  in  the
assignee  or  transferee  any  interest  or  right  herein  whatsoever,  but  immediately  upon  such
assignment or transfer the RSUs will terminate and become of no further effect.
This section of the Agreement applies solely to Participants who are employees of the Company
or any of its subsidiaries. Neither the RSUs nor this Agreement gives you the right to be retained
or  employed  by  the  Company  (or  any  subsidiary  of  the  Company)  in  any  capacity.  Unless
otherwise specified in any written employment or other agreement between the Company and you,
the Company reserves the right to terminate your Service at any time and for any reason.

You, or your estate or heirs, have no rights as a shareholder of the Company until the Shares have
been  issued  to  you  upon  vesting  of  the  RSUs  and  either  a  certificate  evidencing  your  Shares  has
been issued or an appropriate entry has been made on the Company’s books. No  adjustments  are
made for dividends or other rights if the applicable record date occurs before your share certificate
is issued (or an appropriate book entry has been made).
The RSUs are subject to mandatory repayment by you to the Company to the extent you are or in
the  future  become  subject  to  (i)  any  “clawback”  or  recoupment  policy  that  is  adopted  by  the
Company or a subsidiary of the Company to comply with the requirements of any applicable laws,
or (ii) any applicable laws which impose mandatory recoupment, under circumstances set forth in
such applicable laws.
The  number  of  Shares  subject  to  issuance  upon  vesting  of  the  RSUs  is  subject  to  adjustment  in
accordance with Section 4(c) of the Plan. The RSUs shall be subject to the terms of any applicable
agreement  of  merger,  liquidation  or  reorganization  in  the  event  the  Company  is  subject  to  such
corporate activity.

3

Applicable Law

The Plan 

Data Privacy

Consent to Electronic Delivery

This  Agreement  will  be  interpreted  and  enforced  under  the  laws  of  the  Province  of  British
Columbia and the laws of Canada applicable therein, other than any conflicts or choice of law rule
or  principle  that  might  otherwise  refer  construction  or  interpretation  of  this  Agreement  to  the
substantive law of another jurisdiction.
The text of the Plan is incorporated into this Agreement by reference.

This Agreement and the Plan constitute the entire understanding between you and the Company
regarding the RSUs. Any prior agreements, commitments or negotiations concerning this grant are
superseded; except that any written employment, consulting, confidentiality, non-solicitation, non-
competition, and/or severance agreement between you and the Company or any subsidiary of the
Company shall supersede this Agreement with respect to its subject matter.

In  order  to  administer  the  Plan,  the  Company  may  process  personal  data  about  you.  Such  data
includes, but is not limited to the information provided in this Agreement and any changes thereto,
other  appropriate  personal  and  financial  data  about  you  such  as  home  address  and  business
addresses and other contact information, payroll information and any other information that might
be deemed appropriate by the Company to facilitate the administration of the Plan.

By accepting this grant of RSUs, you give explicit consent to the Company to process any such
personal data. You also give explicit consent to the Company to transfer any such personal data
outside  the  country  in  which  you  work  or  are  employed,  including,  with  respect  to  non-U.S.
resident  grantees,  to  the  United  States,  to  transferees  who  shall  include  the  Company  and  other
persons who are designated by the Company to administer the Plan.

By  accepting  this  grant  of  RSUs,  you  consent  to  receive  documents  related  to  the  RSUs  by
electronic  delivery  (including  e-mail  or  reference  to  a  website  or  other  URL)  and,  if  requested,
agree to participate in the Plan through an on-line or electronic system established and maintained
by the Company or another third party designated by the Company, and your consent shall remain
in  effect  throughout  your  term  of  Service  and  thereafter  until  you  withdraw  such  consent  in
writing to the Company.

4

Code Section 409A

Successors and Assigns

Severability

The  RSUs  are  intended  to  be  exempt  from,  Code  Section  409A  except  to  the  extent  subject
thereto, in which case the RSUs are intended to comply with Code 409A, and, accordingly, to the
maximum  extent  permitted,  this  Agreement  will  be  interpreted  and  administered  to  be  in
compliance with Code Section 409A. Notwithstanding anything to the contrary in the Plan or this
Agreement, neither the Company, any subsidiaries of the Company, the Board, nor the Committee
will have any obligation to take any action to prevent the assessment of any excise tax or penalty
on you under Code Section 409A, and neither the Company, any subsidiaries of the Company, the
Board, nor the Committee will have any liability to you for such tax or penalty.

For  purposes  of  this  Agreement,  a  termination  of  Service  only  occurs  upon  an  event  that
constitutes  a  “separation  from  service”  (within  the  meaning  of  Code  Section  409A  and  the
regulations thereunder). Notwithstanding anything in this Agreement to the contrary, if at the time
of your separation from service, (i) you are a “specified employee” (within the meaning of Code
Section 409A and the regulations thereunder, and using the identification methodology selected by
the Company from time to time), and (ii) the Company makes a good faith determination that an
amount  payable  to  you  on  account  of  such  separation  from  service  constitutes  deferred
compensation (within the meaning of Code Section 409A) the payment of which is required to be
delayed pursuant to the six (6)-month delay rule set forth in Code Section 409A in order to avoid
taxes or penalties under Section 409A (the “Delay Period”), then the Company will not pay such
amount on the otherwise scheduled payment date but will instead pay it in a lump sum on the first
payroll date after such Delay Period (or upon your death, if earlier), without interest thereupon.

This  Agreement  shall  inure  to  the  successors  and  assigns  of  the  parties;  provided,  however,  that
neither  this  Agreement  nor  any  rights  hereunder  may  be  assigned  by  you,  except  to  the  extent
expressly permitted herein.
If  any  provision  of  this  Agreement  is  held  invalid  or  unenforceable  by  any  court  of  competent
jurisdiction,  the  other  provisions  of  this  Agreement  will  remain  in  full  force  and  effect.  Any
provision of this Agreement held invalid or unenforceable only in part or degree will remain in full
force and effect to the extent not held invalid or unenforceable.

5

Arbutus Biopharma Corporation

List of Subsidiaries

Arbutus Biopharma Inc.

Delaware, United States of America

Name

Jurisdiction

Exhibit 21.1 

 
 
 
Exhibit 23.1

We consent to the incorporation by reference in the following Registration Statements:

Consent of Independent Registered Public Accounting Firm

1. Registration Statement (Form S-8 No. 333-266527) pertaining to the Arbutus Biopharma Corporation 2016 Omnibus Share and Incentive Plan,

2. Registration  Statement  (Form  S-3  No.  333-260782)  pertaining  to  the  offering,  issuance  and  sale  of  up  to  (a)  $250,000,000  of  common  shares,
preferred shares, warrants, debt securities and units of Arbutus Biopharma Corporation and (b) 38,847,462 common shares offered by the selling
shareholder named therein,

3. Registration  Statement  (Form  S-3  No.  333-248467)  pertaining  to  the  offering,  issuance  and  sale  of  up  to  $200,000,000  of  common  shares,

preferred shares, warrants, debt securities and units of Arbutus Biopharma Corporation,

4. Registration Statement (Form S-8 No. 333-258494) pertaining to the Arbutus Biopharma Corporation 2016 Omnibus Share and Incentive Plan,

5. Registration Statement (Form S-8 No. 333-239407) pertaining to the Arbutus Biopharma Corporation 2016 Omnibus Share and Incentive Plan and

the Arbutus Biopharma Corporation 2020 Employee Stock Purchase Plan,

6. Registration Statement (Form S-8 No. 333-233192) pertaining to the Inducement Stock Option Award of Arbutus Biopharma Corporation,

7. Registration Statement (Form S-8 No. 333-228919) pertaining to the Arbutus Biopharma Corporation 2011 Omnibus Share Compensation Plan,

8. Registration Statement (Form S-8 No. 333-212115) pertaining to the Arbutus Biopharma Corporation 2016 Omnibus Share and Incentive Plan,

9. Registration Statement (Form S-8 No. 333-202762) pertaining to the OnCore Biopharma, Inc. 2014 Equity Incentive Plan, and

10. Registration Statement (Form S-8 No. 333-186185) pertaining to the Tekmira 2011 Omnibus Share Compensation Plan, the Tekmira Share Option

Plan and the Protiva 2000 Incentive Stock Option Plan,

of our report dated March 2, 2023, with respect to the consolidated financial statements of Arbutus Biopharma Corporation included in this Annual Report
(Form 10-K) of Arbutus Biopharma Corporation for the year ended December 31, 2022.

/s/ Ernst & Young LLP

Philadelphia, Pennsylvania
March 2, 2023

                
Exhibit 31.1

CERTIFICATION PURSUANT TO RULE 13a-14 AND 15d-14 OF THE SECURITIES

EXCHANGE ACT OF 1934, AS ADOPTED PURSUANT TO SECTION 302 OF THE

SARBANES-OXLEY ACT OF 2002

I, William Collier, President and Chief Executive Officer of Arbutus Biopharma Corporation, certify that:

1.

I have reviewed this Annual Report on Form 10-K of Arbutus Biopharma Corporation;

2. Based on my knowledge, this report does not contain any untrue statement of a material fact or omit to state a material fact necessary to make
the statements made, in light of the circumstances under which such statements were made, not misleading with respect to the period covered
by this report;

3. Based on my knowledge, the financial statements, and other financial information included in this report, fairly present in all material respects

the financial condition, results of operations and cash flows of the registrant as of, and for, the periods presented in this report;

4. The registrant’s other certifying officer and I are responsible for establishing and maintaining disclosure controls and procedures (as defined in
Exchange Act Rules 13a-15(e) and 15d-15(e)) and internal control over financial reporting (as defined in Exchange Act Rules 13a-15(f) and
15d-15(f)) for the registrant and have:

(a) Designed such disclosure controls and procedures, or caused such disclosure controls and procedures to be designed under our

supervision, to ensure that material information relating to the registrant, including its consolidated subsidiaries, is made known to us by
others within those entities, particularly during the period in which this report is being prepared;

(b) Designed such internal control over financial reporting, or caused such internal control over financial reporting to be designed under our
supervision, to provide reasonable assurance regarding the reliability of financial reporting and the preparation of financial statements for
external purposes in accordance with generally accepted accounting principles;

(c) Evaluated the effectiveness of the registrant’s disclosure controls and procedures and presented in this report our conclusions about the
effectiveness of the disclosure controls and procedures, as of the end of the period covered by this report based on such evaluation; and

(d) Disclosed in this report any change in the registrant’s internal control over financial reporting that occurred during the registrant’s most
recent fiscal quarter (the registrant’s fourth fiscal quarter in the case of an annual report) that has materially affected, or is reasonably
likely to materially affect, the registrant’s internal control over financial reporting; and

5. The registrant’s other certifying officer and I have disclosed, based on our most recent evaluation of internal control over financial reporting, to

the registrant’s auditors and the audit committee of the registrant’s board of directors (or persons performing the equivalent functions):

(a) All significant deficiencies and material weaknesses in the design or operation of internal control over financial reporting which are
reasonably likely to adversely affect the registrant’s ability to record, process, summarize and report financial information; and

(b) Any fraud, whether or not material, that involves management or other employees who have a significant role in the registrant’s internal

control over financial reporting.

Date: March 2, 2023

/s/ William Collier

Name: William Collier

Title: President and Chief Executive Officer
(Principal Executive Officer)

Exhibit 31.2

CERTIFICATION PURSUANT TO RULE 13a-14 AND 15d-14 OF THE SECURITIES

EXCHANGE ACT OF 1934, AS ADOPTED PURSUANT TO SECTION 302 OF THE

SARBANES-OXLEY ACT OF 2002

I, David C. Hastings, Chief Financial Officer of Arbutus Biopharma Corporation, certify that:

1.

I have reviewed this Annual Report on Form 10-K of Arbutus Biopharma Corporation;

2. Based on my knowledge, this report does not contain any untrue statement of a material fact or omit to state a material fact necessary to make
the statements made, in light of the circumstances under which such statements were made, not misleading with respect to the period covered
by this report;

3. Based on my knowledge, the financial statements, and other financial information included in this report, fairly present in all material respects

the financial condition, results of operations and cash flows of the registrant as of, and for, the periods presented in this report;

4. The registrant’s other certifying officer and I are responsible for establishing and maintaining disclosure controls and procedures (as defined in
Exchange Act Rules 13a-15(e) and 15d-15(e)) and internal control over financial reporting (as defined in Exchange Act Rules 13a-15(f) and
15d-15(f)) for the registrant and have:

a. Designed such disclosure controls and procedures, or caused such disclosure controls and procedures to be designed under our

supervision, to ensure that material information relating to the registrant, including its consolidated subsidiaries, is made known to us by
others within those entities, particularly during the period in which this report is being prepared;

b. Designed such internal control over financial reporting, or caused such internal control over financial reporting to be designed under our
supervision, to provide reasonable assurance regarding the reliability of financial reporting and the preparation of financial statements for
external purposes in accordance with generally accepted accounting principles;

c. Evaluated the effectiveness of the registrant’s disclosure controls and procedures and presented in this report our conclusions about the
effectiveness of the disclosure controls and procedures, as of the end of the period covered by this report based on such evaluation; and

d. Disclosed in this report any change in the registrant’s internal control over financial reporting that occurred during the registrant’s most
recent fiscal quarter (the registrant’s fourth fiscal quarter in the case of an annual report) that has materially affected, or is reasonably
likely to materially affect, the registrant’s internal control over financial reporting; and

5. The registrant’s other certifying officer and I have disclosed, based on our most recent evaluation of internal control over financial reporting, to

the registrant’s auditors and the audit committee of the registrant’s board of directors (or persons performing the equivalent functions):

a. All significant deficiencies and material weaknesses in the design or operation of internal control over financial reporting which are
reasonably likely to adversely affect the registrant’s ability to record, process, summarize and report financial information; and

b. Any fraud, whether or not material, that involves management or other employees who have a significant role in the registrant’s internal

control over financial reporting.

Date: March 2, 2023

/s/ David C. Hastings

Name: David C. Hastings

Title: Chief Financial Officer

(Principal Financial Officer)

Exhibit 32.1

CERTIFICATION PURSUANT TO 18 U.S.C. SECTION 1350,

AS ADOPTED PURSUANT TO SECTION 906

OF THE SARBANES-OXLEY ACT OF 2002

In connection with the Annual Report of Arbutus Biopharma Corporation (the “Company”) on Form 10-K for the year ended December 31, 2022, as filed
with the Securities and Exchange Commission on the date hereof (the “Report”), I William Collier, President and Chief Executive Officer of the Company,
certify that to the best of my knowledge:

1. The Report fully complies with the requirements of Section 13(a) or 15(d) of the Securities Exchange Act of 1934, as amended; and

2. The information contained in the Report fairly presents, in all material respects, the financial condition and results of the operations of the

Company.

Date: March 2, 2023

/s/ William Collier

Name: William Collier

Title: President and Chief Executive Officer
(Principal Executive Officer)

Exhibit 32.2

CERTIFICATION PURSUANT TO 18 U.S.C. SECTION 1350,

AS ADOPTED PURSUANT TO SECTION 906

OF THE SARBANES-OXLEY ACT OF 2002

In connection with the Annual Report of Arbutus Biopharma Corporation (the “Company”) on Form 10-K for the year ended December 31, 2022, as filed
with the Securities and Exchange Commission on the date hereof (the “Report”), I David C. Hastings, Chief Financial Officer of the Company, certify that
to the best of my knowledge:

1. The Report fully complies with the requirements of Section 13(a) or 15(d) of the Securities Exchange Act of 1934, as amended; and

2. The information contained in the Report fairly presents, in all material respects, the financial condition and results of the operations of the

Company.

Date: March 2, 2023

/s/ David C. Hastings

Name: David C. Hastings

Title: Chief Financial Officer

(Principal Financial Officer)