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Autolus Therapeutics plc

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FY2020 Annual Report · Autolus Therapeutics plc
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UNITED STATES
SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

FORM  20-F

(Mark One)

    REGISTRATION STATEMENT PURSUANT TO SECTION 12(b) OR (g) OF THE SECURITIES EXCHANGE

ACT OF 1934

OR

x    ANNUAL REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 1934

For the fiscal year ended December 31, 2020

OR

    TRANSITION REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF

1934

for the transition period from                      to

OR

    SHELL COMPANY REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT

OF 1934

Date of event requiring this shell company report

Commission file number: 001-38547

Autolus Therapeutics plc

(Exact name of Registrant as specified in its charter)

N/A
(Translation of Registrant’s name into English)

England and Wales
(Jurisdiction of incorporation)

Forest House
58 Wood Lane
London W12 7RZ United Kingdom
(Address of principal executive offices)

Christian Itin
Chief Executive Officer
Autolus Therapeutics plc
58 Wood Lane
London W12 7RZ United Kingdom
+44 20 3829 6230
Email: ir@autolus.com

(Name, Telephone, E-mail and/or Facsimile number and Address of Company Contact Person)

Securities registered or to be registered, pursuant to Section 12(b) of the Act

Title of each class
American Depository Shares, each representing one ordinary
share, nominal value $0.000042 per share

Trading
Symbol

AUTL

Name of each exchange on which registered

The Nasdaq Stock Market LLC

Securities registered or to be registered pursuant to Section 12(g) of the Act: None

Securities for which there is a reporting obligation pursuant to Section 15(d) of the Act: None

Indicate the number of outstanding shares of each of the issuer’s classes of capital stock or common stock as of the close of business covered by the
annual report.

Ordinary shares, nominal value $0.000042 per share: 52,346,231 as of December 31, 2020

Indicate by check mark if the registrant is a well-known seasoned issuer, as defined in Rule 405 of the Securities Act.    Yes      No  x

If this report is an annual or transition report, indicate by check mark if the registrant is not required to file reports pursuant to Section 13 or
15(d) of the Securities Exchange Act of 1934.     Yes      No  x

Indicate by check mark whether the registrant (1) has filed all reports required to be filed by Section 13 or 15(d) of the Securities Exchange Act of
1934 during the preceding 12 months (or for such shorter period that the registrant was required to file such reports), and (2) has been subject to
such filing requirements for the past 90 days.     Yes  x    No  

Indicate by check mark whether the registrant has submitted every Interactive Data File required to be submitted pursuant to Rule 405 of
Regulation S-T during the preceding 12 months (or for such shorter period that the registrant was required to submit such files).     Yes  x    No  

Indicate by check mark whether the registrant is a large accelerated filer, an accelerated filer, a non-accelerated filer, or an emerging growth
company. See the definitions of “large accelerated filer,” “accelerated filer,” and “emerging growth company” in Rule 12b-2 of the Exchange Act.

Large accelerated filer             Accelerated filer  x           Non-accelerated filer             Emerging growth company  x

If an emerging growth company that prepares its financial statements in accordance with U.S. GAAP, indicate by check mark if the registrant has
elected not to use the extended transition period for complying with any new or revised financial accounting standards † provided pursuant to
Section 13(a) of the Exchange Act.  x

† The term “new or revised financial accounting standard” refers to any update issued by the Financial Accounting Standards Board to its
Accounting Standards Codification after April 5, 2012.

Indicate by check mark whether the registrant has filed a report on and attestation to its management’s assessment of the effectiveness of its
internal control over financial reporting under Section 404(b) of the Sarbanes-Oxley Act (15 U.S.C. 7262(b)) by the registered public accounting
firm that prepared or issued its audit report.    

Indicate by check mark which basis of accounting the registrant has used to prepare the financial statements included in this filing:

U.S. GAAP

x

International Financial Reporting Standards as issued
by the International Accounting Standards Board  

Other  

If “Other” has been checked in response to the previous question indicate by check mark which financial statement item the registrant has elected
to follow.    Item 17      Item 18  

If this is an annual report, indicate by check mark whether the registrant is a shell company (as defined in Rule 12b-2 of the Exchange
Act).    Yes      No x

PART I

ITEM 1.

ITEM 2.

ITEM 3.

ITEM 4.

TABLE OF CONTENTS

IDENTITY OF DIRECTORS, SENIOR MANAGEMENT AND ADVISERS

OFFER STATISTICS AND EXPECTED TIMETABLE

KEY INFORMATION
A. Selected financial data
B. Capitalization and indebtedness
C. Reasons for the offer and use of proceeds
D. Risk factors

INFORMATION ON THE COMPANY
A. History and development of the company
B. Business overview
C. Organizational structure
D. Property, plant and equipment

ITEM 4A.

UNRESOLVED STAFF COMMENTS

ITEM 5.

ITEM 6.

ITEM 7.

ITEM 8.

ITEM 9.

OPERATING AND FINANCIAL REVIEW AND PROSPECTS
A. Operating results
B. Liquidity and capital resources
C. Research and development, patents and licenses, etc.
D. Trend information
E. Off-balance sheet arrangements
F. Tabular disclosure of contractual obligations
G. Other disclosures

DIRECTORS, SENIOR MANAGEMENT AND EMPLOYEES
A. Directors and senior management
B. Compensation
C. Board practices
D. Employees
E. Share ownership

MAJOR SHAREHOLDERS AND RELATED PARTY TRANSACTIONS
A. Major shareholders
B. Related party transactions
C. Interests of experts and counsel

FINANCIAL INFORMATION
A. Consolidated statements and other financial information
B. Significant changes
THE OFFER AND LISTING
A. Offer and listing details
B. Plan of distribution

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ITEM 10.

ITEM 11.

ITEM 12.

PART II

ITEM 13.

ITEM 14.

ITEM 15.

ITEM 16
ITEM 16A
ITEM 16B
ITEM 16C
ITEM 16D
ITEM 16E
ITEM 16F
ITEM 16G
ITEM 16H

PART III

ITEM 17.

ITEM 18.

ITEM 19.

C. Markets
D. Selling shareholders
E. Dilution
F. Expense of the issue

ADDITIONAL INFORMATION
A. Share capital
B. Memorandum and articles of association
C. Material contracts
D. Exchange controls
E. Taxation
F. Dividends and paying agents
G. Statement by experts
H. Documents on display
I. Subsidiary information

QUANTITATIVE AND QUALITATIVE DISCLOSURES ABOUT MARKET RISK

DESCRIPTION OF SECURITIES OTHER THAN EQUITY SECURITIES
A. Debt securities
B. Warrants and rights
C. Other securities
D. American Depositary Shares

DEFAULTS, DIVIDEND ARREARAGES AND DELINQUENCIES

MATERIAL MODIFICATIONS TO THE RIGHTS OF SECURITY HOLDERS AND USE OF PROCEEDS

CONTROLS AND PROCEDURES

[Reserved].
Audit committee financial expert
Code of ethics
Principal accountant fees and services
Exemptions from the listing standards for audit committees
Purchases of equity securities by the issuer and affiliated purchasers
Changes in registrant’s certifying accountant
Corporate governance
Mine safety disclosure

FINANCIAL STATEMENTS

FINANCIAL STATEMENTS

EXHIBITS

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GENERAL INFORMATION
All references in this Annual Report on Form 20-F, or Annual Report, to “Autolus,” the “company,” “we,” “us” and “our” refer to Autolus Therapeutics plc
and its consolidated subsidiaries, except where the context otherwise requires.

PRESENTATION OF FINANCIAL AND OTHER INFORMATION

The consolidated financial statement data as of December 31, 2020 and 2019 and for the years ended December 31, 2020 and 2019, the three months ended
December 31, 2018, and the year ended September 30, 2018 have been derived from our consolidated financial statements, as presented elsewhere in this
Annual Report, which have been prepared in accordance with generally accepted accounting principles in the United States, or U.S. GAAP, as issued by the
Financial Accounting Standards Board, or FASB.

All references in this Annual Report to “$” are to U.S. dollars and all references to “£” are to pounds sterling. Solely for the convenience of the reader,
unless otherwise indicated, all pounds sterling amounts as of December 31, 2020 have been translated into U.S. dollars on the last business day of our fiscal
year ended December 31, 2020, using the exchange rate of £1.00 = $1.3663. All pound sterling amounts for the year ended December 31, 2020 have been
translated into U.S. dollars using the average annual exchange rate £1.00 = $1.2862. These translations should not be considered representations that any
such amounts have been, could have been or could be converted into U.S. dollars at that or any other exchange rate as at that or any other date.

5

CAUTIONARY STATEMENT REGARDING FORWARD-LOOKING STATEMENTS

This Annual Report contains forward-looking statements within the meaning of Section 27A of the Securities Act of 1933, as amended, or the Securities
Act,  and  Section  21E  of  the  Securities  Exchange  Act  of  1934,  as  amended,  or  the  Exchange  Act,  that  involve  substantial  risks  and  uncertainties.  The
forward-looking  statements  are  contained  principally  in  Part  I,  Item  4.B  “Business  Overview,”  Part  I,  Item  3.D.  “Risk  Factors,”  and  Part  I,  Item  5.
“Operating and Financial Review and Prospects,” but are also contained elsewhere in this Annual Report. In some cases, you can identify forward-looking
statements by the words “may,” “might,” “will,” “could,” “would,” “should,” “expect,” “intend,” “plan,” “objective,” “anticipate,” “believe,” “estimate,”
“predict,” “potential,” “continue” and “ongoing,” or the negative of these terms, or other comparable terminology intended to identify statements about the
future. These statements involve known and unknown risks, uncertainties and other important factors that may cause our actual results, levels of activity,
performance  or  achievements  to  be  materially  different  from  the  information  expressed  or  implied  by  these  forward-looking  statements.  The  forward-
looking statements contained in this Annual Report are based upon information available to us as of the date of this Annual Report and, while we believe
we  have  a  reasonable  basis  for  each  forward-looking  statement  contained  this  Annual  Report,  we  caution  you  that  these  statements  are  based  on  a
combination of facts and factors currently known by us and our expectations of the future, about which we cannot be certain. Forward-looking statements
include statements about:

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the development of our product candidates, including statements regarding the timing of initiation, completion and the outcome of clinical studies or trials and
related preparatory work, the period during which the results of the trials will become available and our research and development programs;

our ability to advance our product candidates into, and successfully complete, clinical trials;

our  ability  to  obtain  and  maintain  regulatory  approval  of  our  product  candidates  in  the  indications  for  which  we  plan  to  develop  them,  and  any  related
restrictions, limitations or warnings in the label of an approved drug or therapy;

the  interruption  of  key  clinical  trial  activities,  such  as  clinical  trial  site  monitoring,  due  to  limitations  on  travel,  quarantines  or  social  distancing  protocols
imposed or recommended by federal or state governments, employers and others in connection with the ongoing coronavirus 2019, or COVID-19, pandemic;

our ability to license additional intellectual property relating to our product candidates from third parties and to comply with our existing license agreement;

our plans to research, develop, manufacture and commercialize our product candidates;

the  timing  or  likelihood  of  regulatory  filings  and  approvals  for  our  product  candidates,  along  with  regulatory  developments  in  the  United  States,  European
Union, the United Kingdom and other foreign countries;

the size and growth potential of the markets for our product candidates, if approved, and the rate and degree of market acceptance of our product candidates,
including reimbursement that may be received from payors;

our ability to raise additional capital;

our commercialization, marketing and manufacturing capabilities and strategy;

our ability to attract collaborators with development, regulatory and commercialization expertise;

our expectations regarding our ability to obtain and maintain intellectual property protection;

our ability to attract and retain qualified employees and key personnel;

our ability to contract with third-party suppliers and manufacturers and their ability to perform adequately;

the scalability and commercial viability of our manufacturing methods and processes;

the success of competing therapies that are or may become available;

whether we are classified as a PFIC for current and future periods; and

our estimates regarding future expenses, revenues and needs for additional financing and the accuracy thereof.

You should refer to Item 3.D. "Risk Factors" for a discussion of important factors that may cause our actual results to differ materially from those expressed
or implied by our forward-looking statements. As a result of these factors, we cannot assure you that the forward-looking statements in this Annual Report
will prove to be accurate. Furthermore, if our forward-looking statements prove to be inaccurate, the inaccuracy may be material. In light of the significant
uncertainties in these forward-looking statements, you should not regard these statements as a representation or warranty by us or any other person that we
will achieve our objectives and plans in any specified time frame, or at all. We undertake no obligation to publicly update any forward-looking statements,
whether  as  a  result  of  new  information,  future  events  or  otherwise,  except  as  required  by  law.  You  should,  therefore,  not  rely  on  these  forward-looking
statements as representing our views as of any date subsequent to the date of this Annual Report.

6

Item 1. Identity of Directors, Senior Management and Advisers.

Not applicable.

PART I

Item 2. Offer Statistics and Expected Timetable.

Not applicable.

Item 3. Key Information.

A. Selected financial data.

We have elected to voluntarily comply with Item 3.A, as effective February 10, 2021 and are omitting this disclosure in reliance thereon.

B. Capitalization and indebtedness.

Not applicable.

C. Reasons for the offer and use of proceeds.

Not applicable.

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D. Risk factors.
An investment in our ADSs involves a high degree of risk. You should carefully consider the risks described below, and all other information appearing
elsewhere  in  this  Annual  Report,  including  our  consolidated  financial  statements  and  the  related  notes  hereto,  before  making  an  investment  decision
regarding our securities. The occurrence of any of the events or developments described below could harm our business, financial condition, results of
operations and growth prospects.

RISK FACTORS SUMMARY

Our business is subject to a number of risks and uncertainties, including those risks discussed at-length in the section below titled “Risk Factors.” These
risks include, among others, the following:

•

The ongoing COVID-19 pandemic could materially and adversely affect our business, results of operations and financial condition.

• We have incurred significant losses in every year since our inception. We expect to continue to incur losses over the next several years and may

never achieve or maintain profitability.

• Our limited operating history may make it difficult for you to evaluate the success of our business to date and to assess our future viability.

• We will need additional funding to complete the development of our product candidates, which may not be available on acceptable terms, if at all.

• We are early in our development efforts. All of our product candidates are in clinical development or in preclinical development. If we are unable
to advance our product candidates through clinical development, obtain regulatory approval and ultimately commercialize our product candidates,
or experience significant delays in doing so, our business will be materially harmed.

• Our proprietary, next-generation T cell programming technologies, our modular approach for engineering T cells and our manufacturing platform
for our programmed T cell product candidates, represent emerging approaches to cancer treatment that face significant challenges and hurdles.

• Our future success is highly dependent on the regulatory approval of our current clinical-stage programmed T cell product candidates and our

preclinical programs. All of our product candidates will require significant clinical or preclinical testing before we can seek regulatory approval
for and launch a product commercially.

• Adverse side effects or other safety risks associated with our product candidates could delay or preclude approval, cause us to suspend or

discontinue clinical trials, cause us to abandon product candidates, could limit the commercial profile of an approved label, or could result in
significant negative consequences following any potential marketing approval.

•

If the clinical trials of any of our product candidates fail to demonstrate safety and efficacy to the satisfaction of the FDA, the EMA or other
comparable regulatory authorities, or do not otherwise produce favorable results, we may incur additional costs or experience delays in
completing, or ultimately be unable to complete, the development and commercialization of our product candidates.

• We may not be able to successfully create our own manufacturing infrastructure for supply of our requirements of programmed T cell product

candidates for use in clinical trials and for commercial sale.

• Our product candidates are biologics and the manufacture of our product candidates is complex and we may encounter difficulties in production,
particularly with respect to process development or scaling-out of our manufacturing capabilities. If we encounter such difficulties, our ability to
provide supply of our product candidates for clinical trials or our products for patients, if approved, could be delayed or stopped.

• We operate in a rapidly changing industry and face significant competition, which may result in others discovering, developing or

commercializing products before or more successfully than we do.

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•

If we are unable to obtain and maintain patent protection for our T cell programming technologies and product candidates, or if the scope of the
patent protection obtained is not sufficiently broad, our competitors could develop and commercialize technology and biologics similar or
identical to ours, and our ability to successfully commercialize our technology and product candidates may be impaired.

• As an English public limited company, certain capital structure decisions will require shareholder approval, which may limit our flexibility to

manage our capital structure.

Risks Related to Our Financial Position and Need For Capital

We have incurred significant losses in every year since our inception. We expect to continue to incur losses over the next several years and may never
achieve or maintain profitability.

We are a clinical-stage biopharmaceutical company with a limited operating history and we have incurred significant net losses since our inception
in 2014. We have incurred losses of $142.1 million and $123.8 million for the years ended December 31, 2020 and 2019, $20.6 million for three months
ended  December  31,  2018,  and  $44.8  million  for  the  year  ended  September  30,  2018,  respectively.  As  of  December  31,  2020,  we  had  an  accumulated
deficit of $379.2 million. We have funded our operations to date primarily with proceeds from the sale of our equity securities.

We  have  no  products  approved  for  commercial  sale,  and  while  we  have  generated  a  small  amount  of  revenue  from  licensing,  we  are  devoting
substantially all of our financial resources and efforts to research and development of our programmed T cell product candidates as well as to building out
our manufacturing infrastructure, T cell programming technologies and management team. Investment in biopharmaceutical product development is highly
speculative  because  it  entails  substantial  upfront  capital  expenditures  and  significant  risk  that  any  potential  product  candidate  will  fail  to  demonstrate
adequate effect or an acceptable safety profile, gain regulatory approval and become commercially viable.

We expect that it will take at least several years until any of our product candidates receive marketing approval and are commercialized, and we
may never be successful in obtaining marketing approval and commercializing product candidates. We expect to continue to incur significant expenses and
increasing  operating  losses  for  the  foreseeable  future.  These  net  losses  will  adversely  impact  our  shareholders’  equity  and  net  assets  and  may  fluctuate
significantly from quarter to quarter and year to year. We anticipate that our expenses will increase substantially as we:

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continue  our  ongoing  and  planned  research  and  development  of  our  current  programmed  T  cell  product  candidates  for  the  treatment  of
hematological cancers and solid tumors;

initiate  preclinical  studies  and  clinical  trials  for  any  additional  product  candidates  that  we  may  pursue  in  the  future,  including  our  planned
development of additional T cell therapies for the treatment of hematological cancers and solid tumors;

seek to discover and develop additional product candidates and further expand our clinical product pipeline;

seek regulatory approvals for any product candidates that successfully complete clinical trials;

continue to scale up internal and external manufacturing capacity with the aim of securing sufficient quantities to meet our capacity requirements
for clinical trials and potential commercialization;

establish sales, marketing and distribution infrastructure to commercialize any product candidate for which we may obtain regulatory approval;

• make required milestone and royalty payments to UCL Business Ltd., or UCLB, the technology-transfer company of University College London,

or UCL, or other third parties, under license agreements pursuant to which we were granted some of our intellectual property rights;

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develop, maintain, expand and protect our intellectual property portfolio;

acquire or in-license other product candidates and technologies;

hire additional clinical, quality control and manufacturing personnel;

add clinical, operational, financial and management information systems and personnel, including personnel to support our product development
and planned future commercialization efforts;

expand our operations in the United States, Europe and other geographies; and

incur additional legal, accounting and other expenses associated with operating as a public company.

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To become and remain profitable, we must succeed in developing and eventually commercializing products that generate significant revenue. This
will require us to be successful in a range of challenging activities, including completing preclinical studies and clinical trials of our product candidates,
obtaining regulatory approval, manufacturing, marketing and selling any products for which we may obtain regulatory approval, as well as discovering and
developing additional product candidates. We may never succeed in these activities and, even if we do, may never generate revenues that are significant
enough to achieve profitability.

Because  of  the  numerous  risks  and  uncertainties  associated  with  the  development,  manufacturing,  delivery  and  commercialization  of  complex
autologous cell therapies, we are unable to accurately predict the timing or amount of expenses or when, or if, we will be able to achieve profitability. If we
are required by regulatory authorities to perform studies in addition to those currently expected, or if there are any delays in the initiation and completion of
our clinical trials or the development of any of our product candidates, our expenses could increase and profitability could be further delayed.

Even if we achieve profitability, we may not be able to sustain or increase profitability on a quarterly or annual basis. Our failure to become and
remain  profitable  would  depress  the  value  of  our  ADSs  and  could  impair  our  ability  to  raise  capital,  expand  our  business,  maintain  our  research  and
development efforts or continue our operations. A decline in the value of our ADSs could also cause you to lose all or part of your investment.

Our limited operating history may make it difficult for you to evaluate the success of our business to date and to assess our future viability.

        We  are  a  clinical-stage  biopharmaceutical  company  with  a  limited  operating  history.  As  an  organization,  we  have  not  demonstrated  an  ability  to
successfully complete late-stage clinical trials, obtain regulatory approvals, manufacture our product candidates at commercial scale or arrange for a third
party to do so on our behalf, conduct sales and marketing activities necessary for successful commercialization, or obtain reimbursement in the countries of
sale.  We  may  encounter  unforeseen  expenses,  difficulties,  complications,  and  delays  in  achieving  our  business  objectives.  Our  limited  history  as  an
operating company makes any assessment of our future success or viability subject to significant uncertainty. If we do not address these risks successfully
or are unable to transition at some point from a company with a research and development focus to a company capable of supporting commercial activities,
then our business will suffer.

We will need additional funding to complete the development of our product candidates, which may not be available on acceptable terms, if at all.

We will require substantial additional funding to meet our financial needs and to pursue our business objectives. If we are unable to raise capital

when needed, we could be forced to delay, reduce or altogether cease our product development programs or commercialization efforts.

Since  our  inception,  we  have  devoted  substantially  all  of  our  resources  to  fund  the  operating  expenses  and  capital  expenditure  requirements
associated with the research and development of our product candidates. These programs are described in greater detail under the heading “Our Pipeline” in
the  section  titled  “Business  Overview”  of  this  Annual  Report.  Our  current  funding  will  not  be  sufficient  for  us  to  fund  any  of  our  programmed  T  cell
product candidates through regulatory approval, and we will need to raise additional capital to complete the development and commercialization of our
programmed T cell product candidates, and in connection with our continuing operations and other planned activities. Our future capital requirements will
depend on many factors, including:

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the  progress,  results  and  costs  of  laboratory  testing,  manufacturing,  and  preclinical  and  clinical  development  of  our  current  and  future  product
candidates;

the timing and amounts of any milestone or royalty payments we may be required to make under current or future license agreements;

the  costs  of  leasing,  building  out,  equipping,  and  operating  the  facilities  necessary  to  research,  develop,  manufacture  and  commercialize  our
product candidates, as well as to support our continuing operations;

the costs of hiring additional clinical, quality control and manufacturing personnel;

the costs, timing and outcome of regulatory review of our product candidates;

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•

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the  costs  and  timing  of  future  commercialization  activities,  including  product  manufacturing,  marketing,  sales  and  distribution,  for  any  of  our
product candidates for which we receive marketing approval;

the revenue, if any, received from commercial sales of our product candidates for which we receive marketing approval;

the  costs  and  timing  of  preparing,  filing  and  prosecuting  patent  applications,  maintaining  and  enforcing  our  intellectual  property  rights  and
defending any intellectual property-related claims; and

the costs of operating as a public company.

Identifying potential product candidates and conducting preclinical testing and clinical trials is a time-consuming, expensive and uncertain process
that takes years to complete, and we may never generate the necessary data or results required to obtain regulatory approval and achieve product sales. In
addition, our product candidates, if approved, may not achieve commercial success. Our product revenues, if any, will be derived from sales of product
candidates  that  we  do  not  expect  to  be  commercially  available  for  several  years,  if  at  all.  Accordingly,  we  will  need  to  continue  to  rely  on  additional
financing to achieve our business objectives. Adequate additional financing may not be available to us on acceptable terms, or at all. In addition, we may
seek additional capital due to favorable market conditions or strategic considerations even if we believe we have sufficient funds for our current or future
operating plans.

To the extent that we raise additional capital through the sale of equity or convertible debt securities, your ownership interest will be diluted, and
the terms of these securities may include liquidation or other preferences that adversely affect your rights as a shareholder. Debt financing and preferred
equity financing, if available, could result in fixed payment obligations, and we may be required to accept terms that restrict our ability to incur additional
indebtedness, force us to maintain specified liquidity or other ratios or restrict our ability to pay dividends or make acquisitions.

If we raise additional funds through collaborations, strategic alliances or marketing, distribution or licensing arrangements with third parties, we
may be required to relinquish valuable rights to our technologies, future revenue streams, research programs or product candidates or to grant licenses on
terms that may not be favorable to us. In addition, we could also be required to seek funds through arrangements with collaborators or others at an earlier
stage than otherwise would be desirable. If we raise funds through research grants, we may be subject to certain requirements, which may limit our ability
to use the funds or require us to share information from our research and development. If we are unable to raise additional funds through equity or debt
financings  when  needed,  we  may  be  required  to  delay,  limit,  reduce  or  terminate  our  product  development  or  future  commercialization  efforts  or  grant
rights to a third party to develop and market product candidates that we would otherwise prefer to develop and market ourselves. Raising additional capital
through any of these or other means could adversely affect our business and the holdings or rights of our shareholders and may cause the market price of
our ADSs to decline.

Risks Related to the Development of Our Product Candidates

We are early in our development efforts. All of our product candidates are in clinical development or in preclinical development. If we are unable to
advance  our  product  candidates  through  clinical  development,  obtain  regulatory  approval  and  ultimately  commercialize  our  product  candidates,  or
experience significant delays in doing so, our business will be materially harmed.

We  have  established  clinical  proof-of-concept  for  only  one  of  our  product  candidates.  There  is  no  assurance  that  our  current  or  any  other  future
clinical trials of our product candidates will be successful or will generate positive clinical data and we may not receive marketing approval from the U.S.
Food  and  Drug  Administration,  or  FDA,  or  other  regulatory  agencies,  including  the  European  Medicines  Agency,  or  EMA,  for  any  of  our  product
candidates. In order to commence a clinical trial in the United States, we must submit an Investigational New Drug application, or IND, to the FDA and
have the IND application go into effect. Trials in the United States must be conducted pursuant to an active IND. An investigator may not administer an
investigational new drug to human subjects until the IND application goes into effect. Similar requirements apply to our conduct of trials in the United
Kingdom and European Union. We have active clinical trial applications in effect for two of our current clinical-stage product candidates, AUTO1 and
AUTO3. There can be no assurance that the FDA, EMA or other regulatory agencies will permit any future clinical trial application to go into effect in a
timely manner or at all.

Biopharmaceutical development is a long, expensive and uncertain process, and delay or failure can occur at any stage of any of our clinical trials.
Failure to obtain regulatory approval for our product candidates will prevent us from commercializing and marketing our product candidates. The success
in the development of our programmed T cell product candidates will depend on many factors, including:

•

completing preclinical studies and receiving regulatory approvals or clearance for conducting clinical trials for our preclinical-stage programs;

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obtaining positive results in our clinical trials demonstrating efficacy, safety, and durability of effect of our product candidates;

receiving approvals for commercialization of our product candidates from regulatory authorities;

• manufacturing our product candidates at an acceptable cost; and

• maintaining  and  growing  an  organization  of  scientists,  medical  professionals  and  business  people  who  can  develop  and  commercialize  our

products and technology.

Many  of  these  factors  are  beyond  our  control,  including  the  time  needed  to  adequately  complete  clinical  testing  and  the  regulatory  submission
process. It is possible that none of our product candidates will ever obtain regulatory approval, even if we expend substantial time and resources seeking
such approval. If we do not achieve one or more of these factors in a timely manner or at all, or any other factors impacting the successful development of
biopharmaceutical products, we could experience significant delays or an inability to successfully develop our product candidates, which would materially
harm our business.

Our proprietary, next-generation T cell programming technologies, our modular approach for engineering T cells and our manufacturing platform for
our programmed T cell product candidates, represent emerging approaches to cancer treatment that face significant challenges and hurdles.

We  have  concentrated  our  research  and  development  efforts  on  our  T  cell  technology  platform  using  our  expertise  in  tumor  biology  and  cell
programming, and our future success is highly dependent on the successful development and manufacture of our programmed T cell product candidates.
We do not currently have any approved or commercialized products. One of our most advanced product candidates employs a dual-targeting mechanism.
By  targeting  two  separate  antigens  on  the  cancer  cell  surface,  we  believe  this  product  candidate  has  the  potential  to  improve  durability  of  treatment
response and reduce the frequency of cancer relapse as compared to other currently available single-targeting T cell therapies. Our product candidate for the
treatment of T-cell lymphoma employs a novel approach to killing malignant T cells that aims to preserve approximately half of the normal, healthy T cells.
Some of our product candidates include a “safety switch” that is designed to allow for the elimination of the engineered T cells if a patient experiences
severe adverse side effects from the treatment. However, this “safety switch” technology has not been used to date in our clinical studies, and we do not
know  whether  it  would  have  the  intended  effect  if  used.  Additionally,  as  with  other  targeted  therapies,  off-tumor  or  off-target  activity  could  delay
development or require us to re-engineer or abandon a particular product candidate. Because programmed T cell therapies represent a relatively new field
of cellular immunotherapy and cancer treatment generally, developing and commercializing our product candidates subjects us to a number of risks and
challenges, including:

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•

•

•

•

obtaining  regulatory  approval  for  our  product  candidates,  as  the  FDA,  the  EMA  and  other  regulatory  authorities  have  limited  experience  with
programmed T cell therapies for cancer;

sourcing clinical and, if approved, commercial supplies of the materials used to manufacture our product candidates;

developing programming modules with the desired properties, while avoiding adverse reactions;

creating viral vectors capable of delivering multiple programming modules;

developing a reliable and consistent vector and cell manufacturing process;

establishing  manufacturing  capacity  suitable  for  the  manufacture  of  our  product  candidates  in  line  with  expanding  enrollment  in  our  clinical
studies and our projected commercial requirements;

achieving cost efficiencies in the scale-up of our manufacturing capacity;

developing protocols for the safe administration of our product candidates;

educating medical personnel regarding our programmed T cell therapies and the potential side effect profile of each of our product candidates,
such as potential adverse side effects related to cytokine release syndrome;

establishing integrated solutions in collaboration with specialty treatment centers in order to reduce the burdens and complex logistics commonly
associated with the administration of T cell therapies;

establishing sales and marketing capabilities to successfully launch and commercialize our product candidates if and when we obtain any required
regulatory approvals, and risks associated with gaining market acceptance of a novel therapy if we receive approval; and

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obtaining  coverage  and  adequate  reimbursement  from  third-party  payors  for  our  novel  and  personalized  therapies  in  connection  with
commercialization of any approved product candidates.

We may not be able to successfully develop our programmed T cell product candidates or our T cell programming technologies in a manner that

will yield products that are safe and effective, scalable or profitable.

Additionally, because our technology involves the genetic modification of patient cells ex vivo, we are subject to additional regulatory challenges
and risks, including regulatory requirements governing genetically modified organisms that have changed frequently and will likely continue to change in
the future, and that may limit or delay our ability to import our product candidates into certain countries for use in clinical trials or for commercial sale even
if we receive applicable marketing approvals.

Moreover,  public  perception  and  awareness  of  T  cell  therapy  safety  issues  may  adversely  influence  the  willingness  of  subjects  to  participate  in
clinical trials of our product candidates, or if approved, of physicians to prescribe our products. Physicians, hospitals and third-party payors often are slow
to adopt new products, technologies and treatment practices that require additional upfront costs and training. Treatment centers may not be willing or able
to devote the personnel and establish other infrastructure required for the administration of programmed T cell therapies. Physicians may not be willing to
undergo training to adopt this novel and personalized therapy, may decide the therapy is too complex to adopt without appropriate training and may choose
not to administer the therapy. Based on these and other factors, hospitals and payors may decide that the benefits of this new therapy do not or will not
outweigh its costs.

Our  future  success  is  highly  dependent  on  the  regulatory  approval  of  our  current  clinical-stage  programmed  T  cell  product  candidates  and  our
preclinical programs. All of our product candidates will require significant clinical or preclinical testing before we can seek regulatory approval for
and launch a product commercially.

    We do not have any products that have gained regulatory approval. Our business is substantially dependent on our ability to obtain regulatory approval
for, and, if approved, to successfully commercialize our programmed T cell product candidates. We cannot commercialize product candidates in the United
States without first obtaining regulatory approval for the product from the FDA; similarly, we cannot commercialize product candidates in countries outside
of the United States without obtaining regulatory approval from comparable regulatory authorities in relevant jurisdictions, such as the EMA in Europe.
Before obtaining regulatory approvals for the commercial sale of any product candidate for a particular indication, we must demonstrate with substantial
evidence gathered in preclinical and clinical studies, that the product candidate is safe and effective for that indication and that the manufacturing facilities,
processes and controls are adequate with respect to such product candidate. To date, we have had only limited interaction with both the FDA and the EMA
regarding our product candidates. Prior to seeking approval for any of our product candidates, we will need to confer with the FDA, the EMA and other
regulatory authorities regarding the design of our clinical trials and the type and amount of clinical data necessary to seek and gain approval for our product
candidates.

The  time  required  to  obtain  approval  by  the  FDA,  the  EMA  and  other  regulatory  authorities  is  unpredictable  but  typically  takes  many  years
following  the  commencement  of  preclinical  studies  and  clinical  trials  and  depends  upon  numerous  factors,  including  the  substantial  discretion  of  the
regulatory authorities. In addition, approval policies, regulations, or the type and amount of clinical data necessary to gain approval may change during the
course of a product candidate’s clinical development and may vary among jurisdictions. It is possible that none of our existing product candidates or any
future product candidates will ever obtain regulatory approval.

Our  product  candidates  could  fail  to  receive  regulatory  approval  from  the  FDA,  the  EMA  or  other  regulatory  authorities  for  many  reasons,

including:

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•

•

disagreement with the design, protocol or conduct of our clinical trials;

failure to demonstrate that a product candidate is safe and effective for its proposed indication;

failure of clinical trials to meet the level of statistical significance required for approval;

failure to demonstrate that a product candidate’s clinical and other benefits outweigh its safety risks;

disagreement with our interpretation of data from preclinical studies or clinical trials;

insufficiency  of  data  collected  from  clinical  trials  of  our  product  candidates  to  support  the  submission  and  filing  of  a  Biologics  License
Application, or BLA, or other submission or to obtain regulatory approval;

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failure to obtain approval of the manufacturing processes or our facilities;

changes in the approval policies or regulations that render our preclinical and clinical data insufficient for approval; or

lack of adequate funding to complete a clinical trial in a manner that is satisfactory to the applicable regulatory authority.

The FDA, the EMA or a comparable regulatory authority may require more information, including additional preclinical or clinical data to support
approval,  including  data  that  would  require  us  to  perform  additional  clinical  trials  or  modify  our  manufacturing  processes,  which  may  delay  or  prevent
approval and our commercialization plans, or we may decide to abandon the development program. If we change our manufacturing processes, we may be
required to conduct additional clinical trials or other studies, which also could delay or prevent approval of our product candidates. If we were to obtain
approval, regulatory authorities may approve any of our product candidates for fewer indications than we request (including failing to approve the most
commercially promising indications), may limit indications, may grant approval contingent on the performance of costly post-marketing clinical trials or
other post-marketing commitments, or may approve a product candidate with a label that does not include the labeling claims necessary or desirable for the
successful commercialization of that product candidate.

Depending on results we observe in our clinical trials, our development strategy may include the pursuit of expedited approvals from the FDA or the
EMA, such as through the accelerated approval pathway, and we may seek to achieve breakthrough therapy designation or regenerative medicine advanced
therapy, or RMAT, designation from the FDA or the PRIority MEdicines, or PRIME, designation from the EMA. Our product candidates may not qualify
for  such  designations,  and  the  clinical  data  obtained  from  trials  of  our  product  candidates  may  not  be  sufficient  to  qualify  for  any  expedited  approval
program.

Even  if  a  product  candidate  were  to  successfully  obtain  approval  from  the  FDA,  the  EMA  or  other  comparable  regulatory  authorities  in  other
jurisdictions,  any  approval  might  contain  significant  limitations  related  to  use  restrictions  for  specified  age  groups,  warnings,  precautions  or
contraindications, or may be subject to burdensome post-approval study or risk management requirements. If we are unable to obtain regulatory approval
for one of our product candidates in one or more jurisdictions, or any approval contains significant limitations, we may not be able to obtain sufficient
funding  to  continue  the  development  of  that  product  or  generate  revenues  attributable  to  that  product  candidate.  Also,  any  regulatory  approval  of  our
current or future product candidates, once obtained, may be withdrawn. See the risk factor titled “—Even if we complete the necessary preclinical studies
and  clinical  trials,  the  regulatory  approval  process  is  expensive,  time-consuming  and  uncertain  and  may  prevent  us  from  obtaining  approvals  for  the
commercialization of some or all of our product candidates. As a result, we cannot predict when or if, and in which territories, we will obtain marketing
approval to commercialize a product candidate.”

We may not be successful in our efforts to build a pipeline of product candidates.

A key element of our strategy is to use our expertise in tumor biology and cell programming and our proprietary and modular T cell programming
technologies  to  develop  what  we  believe  are  safer  and  more  effective  T  cell  therapies.  Our  initial  focus  is  on  the  development  of  a  pipeline  of  product
candidates  for  the  treatment  of  hematological  cancers  and  the  progression  of  these  product  candidates  through  clinical  development.  We  also  intend  to
develop follow-on, or next-generation, product candidates with additional elements of programming built into the programmed T cell product candidate to
offer  enhanced  characteristics  as  compared  to  the  earlier  product  generation,  such  as  pharmacological  control  or  insensitivity  to  checkpoint  inhibition.
However, we may not be able to develop product candidates that are safe and effective, or which compare favorably with our existing product candidates.
Even if we are successful in continuing to build our pipeline and developing next-generation product candidates or expanding into solid tumor indications,
the potential product candidates that we identify may not be suitable for clinical development, including as a result of lack of safety, lack of tolerability,
lack of anti-tumor activity, or other characteristics that indicate that they are unlikely to be products that will receive marketing approval, achieve market
acceptance or obtain reimbursements from third-party payors. If we do not successfully develop and commercialize product candidates or collaborate with
others to do so, we will not be able to obtain product revenue in future periods, which could significantly harm our financial position and adversely affect
the trading price of our ADSs.

Our preclinical programs may experience delays or may never advance to clinical trials, which would adversely affect our ability to obtain regulatory
approvals or to commercialize these programs on a timely basis or at all, which would have an adverse effect on our business.

Many  of  our  product  candidates  are  in  the  preclinical  development  stage.  The  risk  of  failure  of  preclinical  programs  is  high.  Before  we  can
commence clinical trials for a product candidate, we must complete extensive preclinical testing and studies to obtain regulatory clearance to initiate human
clinical trials, including based on IND applications in effect in the United States and clinical trial applications, or CTAs, in Europe. We cannot be certain of
the timely completion or outcome of our preclinical testing

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and studies and cannot predict if the FDA, the EMA or other regulatory authorities will accept our proposed clinical programs or if the outcome of our
preclinical testing and studies will ultimately support the further development of our programs. As a result, we cannot be sure that we will be able to submit
INDs or similar applications for our preclinical programs on the timelines we expect, if at all, and we cannot be sure that submission of INDs or similar
applications will result in the FDA, the EMA or other regulatory authorities allowing clinical trials to begin.

Clinical trials are difficult to design and implement, involve uncertain outcomes and may not be successful.

Human clinical trials are difficult to design and implement, in part because they are subject to rigorous regulatory requirements. The design of a
clinical trial can determine whether its results will support approval of a product, and flaws in the design of a clinical trial may not become apparent until
the  clinical  trial  is  well  advanced.  As  an  organization,  we  have  limited  experience  designing  clinical  trials  and  may  be  unable  to  design  and  execute  a
clinical trial to support regulatory approval. There is a high failure rate for biologic products proceeding through clinical trials, which may be higher for our
product candidates because they are based on new technology and engineered on a patient-by-patient basis. Many companies in the pharmaceutical and
biotechnology  industries  have  suffered  significant  setbacks  in  late-stage  clinical  trials  even  after  achieving  promising  results  in  preclinical  testing  and
earlier-stage clinical trials. Data obtained from preclinical and clinical activities are subject to varying interpretations, which may delay, limit or prevent
regulatory  approval.  In  addition,  we  may  experience  regulatory  delays  or  rejections  as  a  result  of  many  factors,  including  changes  in  regulatory  policy
during the period of our product candidate development. Any such delays could negatively impact our business, financial condition, results of operations
and prospects.

Success in preclinical studies or clinical trials may not be indicative of results in future clinical trials.

Results from preclinical studies are not necessarily predictive of future clinical trial results, and interim results of a clinical trial are not necessarily
indicative of final results. For example, we have treated only a small number of patients in all of our ongoing clinical trials. For that reason, we do not
know whether these candidates will be effective for the intended indications or safe in humans. Our product candidates may fail to show the desired safety
and efficacy in clinical development despite positive results in preclinical studies or having successfully advanced through initial clinical trials. This failure
to establish sufficient efficacy and safety could cause us to abandon clinical development of our product candidates.

We  depend  on  enrollment  of  patients  in  our  clinical  trials  for  our  product  candidates.  If  we  encounter  difficulties  enrolling  patients  in  our  clinical
trials, our clinical development activities could be delayed or otherwise adversely affected.

Identifying and qualifying patients to participate in clinical trials of our product candidates is critical to our success. We may experience difficulties
in  patient  enrollment  in  our  clinical  trials  for  a  variety  of  reasons.  The  timely  completion  of  clinical  trials  in  accordance  with  their  protocols  depends,
among other things, on our ability to enroll a sufficient number of patients who remain in the study until its conclusion. The enrollment of patients depends
on many factors, including:

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the patient eligibility criteria defined in the protocol;

the number of patients with the disease or condition being studied;

the perceived risks and benefits of the product candidate in the trial;

clinicians’ and patients’ perceptions as to the potential advantages of the product candidate being studied in relation to other available therapies,
including any new drugs that may be approved for the indications we are investigating or drugs that may be used off-label for these indications;

the size and nature of the patient population required for analysis of the trial’s primary endpoints;

the proximity of patients to study sites;

the design of the clinical trial;

our ability to recruit clinical trial investigators with the appropriate competencies and experience;

competing clinical trials for similar therapies or other new therapeutics not involving T cell-based immunotherapy;

our ability to obtain and maintain patient consents;

disruptions to health care systems caused by the coronavirus pandemic; and

the risk that patients enrolled in clinical trials will drop out of the clinical trials before completion of their treatment.

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In particular, some of our clinical trials will look to enroll patients with characteristics which are found in a very small population. For example, our
clinical trial for AUTO4 seeks to enroll patients with peripheral T-cell lymphoma, a rare and heterogeneous form of non-Hodgkin lymphoma, or NHL.
Other  companies  are  conducting  clinical  trials  with  their  redirected  T  cell  therapies  in  multiple  myeloma,  pediatric  relapsed  or  refractory  acute  B
lymphocytic leukemia, or pediatric ALL, and relapsed or refractory diffuse large B-cell lymphoma, or DLBCL, and seek to enroll patients in their studies
that may otherwise be eligible for our clinical trials, which could lead to slow recruitment and delays in our clinical programs. In addition, since the number
of qualified clinical investigators is limited, we expect to conduct some of our clinical trials at the same clinical trial sites that some of our competitors use,
which  could  further  reduce  the  number  of  patients  who  are  available  for  our  clinical  trials  in  these  clinical  trial  sites.  Moreover,  because  our  product
candidates represent a departure from more commonly used methods for cancer treatment, potential study participants and their doctors may be inclined to
use conventional therapies, such as chemotherapy and antibody therapy, rather than participate in our clinical trials.

Delays in patient enrollment may result in increased costs or may affect the timing or outcome of the planned clinical trials, which could prevent
completion of these clinical trials and adversely affect our ability to advance the development of our product candidates. In addition, many of the factors
that may lead to a delay in the commencement or completion of clinical trials may also ultimately lead to the denial of regulatory approval of our product
candidates.

The market opportunities for certain of our product candidates may be limited to those patients who are ineligible for or have failed prior treatments
and may be small, and our projections regarding the size of the addressable market may be incorrect.

Cancer therapies are sometimes characterized as first line, second line or third line, and the FDA often approves new therapies initially only for
third line use. When blood cancers are detected, they are treated with the first line of therapy with the intention of curing the cancer. This generally consists
of  chemotherapy,  radiation,  antibody  drugs,  tumor-targeted  small  molecules,  or  a  combination  of  these.  In  addition,  sometimes  a  bone  marrow
transplantation can be added to the first line therapy after the combination chemotherapy is given. If the patient’s cancer relapses, then they are given a
second line or third line therapy, which can consist of more chemotherapy, radiation, antibody drugs, tumor-targeted small molecules, or a combination of
these,  or  a  bone  marrow  transplant.  Generally,  the  higher  the  line  of  therapy,  the  lower  the  chance  of  a  cure.  With  third  or  higher  line,  the  goal  of  the
therapy in the treatment of lymphoma and myeloma is to control the growth of the tumor and extend the life of the patient, as a cure is unlikely to happen.
Patients are generally referred to clinical trials in these situations.

We  are  initially  developing  AUTO1  as  second  line  therapy  for  patients  with  ALL  who  are  considered  at  high  risk  for  relapse  and  as  third  line
therapy for other patients with ALL, AUTO3 as a third line therapy for DLBCL and AUTO4 as a second line therapy for TRBC1-positive T-cell lymphoma
patients. If AUTO4 is approved as a second line therapy, we may initiate a trial to position it as a consolidation therapy after first line chemotherapy in T-
cell lymphoma. There is no guarantee that any of our product candidates, even if approved, would be approved for an earlier line of therapy. In addition, we
may have to conduct additional large randomized clinical trials prior to gaining approval for the earlier line of therapy.

Our projections of both the number of people who have the cancers we are targeting, as well as the size of the patient population subset of people
with these cancers in a position to receive first, second, third and fourth line therapy and who have the potential to benefit from treatment with our product
candidates, are based on our beliefs and estimates. These estimates have been derived from a variety of sources, including scientific literature, surveys of
clinics, patient foundations, or market research and may prove to be incorrect. Further, new studies may change the estimated incidence or prevalence of
these cancers. The number of patients may turn out to be fewer than expected. Additionally, the potentially addressable patient population for our product
candidates may be limited or may not be amenable to treatment with our product candidates. For instance, in our clinical trial for AUTO4, we are initially
targeting a small patient population that suffers from peripheral T-cell lymphoma, a rare and heterogeneous form of NHL. Even if we obtain significant
market share for our product candidates, because the potential target populations are small, we may never achieve significant revenues without obtaining
regulatory approval for additional indications or as part of earlier lines of therapy.

Adverse side effects or other safety risks associated with our product candidates could delay or preclude approval, cause us to suspend or discontinue
clinical trials, cause us to abandon product candidates, limit the commercial profile of an approved label, or result in significant negative consequences
following any potential marketing approval.

In  clinical  trials  conducted  by  other  companies  involving  CAR  T  cells,  the  most  prominent  acute  toxicities  included  symptoms  thought  to  be
associated with cytokine release syndrome, or CRS, such as fever, low blood pressure and kidney dysfunction. Some patients also experienced toxicity of
the central nervous system, or neurotoxicity, such as confusion, tremor, cranial nerve dysfunction, seizures and speech impairment. Adverse events with the
worst grades and attributed to CAR T cells were severe and life threatening in some patients. The life threatening events were related to kidney dysfunction
and neurotoxicity. Severe and life threatening toxicities occurred mostly in the first two weeks after cell infusion and generally resolved within three weeks,
but several patients died in clinical trials involving CAR T cells developed by other companies and academic institutions.

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For example, as of the most recent data cut-off date of November 12, 2020, 20 patients in the ALLCAR19 clinical trial have received AUTO1. Of those 20
patients, three patients (15%) experienced Grade 3 neurotoxicity that resolved swiftly with steroids. We also observed Grade 3 or higher neutropenias in 7
of  20  patients  reporting  such  neutropenias  prior  to  treatment  with  AUTO1  and  8  of  17  patients  reporting  Grade  3  or  higher  neutropenias  28  days  after
treatment with AUTO1, with most resolving by month two or three post-dosing. Of the 20 patients, seven patients died while enrolled in the ALLCAR19
clinical  trial,  of  which  two  deaths  were  determined  to  be  due  to  progression  of  the  leukemia,  one  death  occurred  post  progression  following  infective
complication post-transplant and the four remaining deaths were determined to be due to infectious complications (a common complication of advanced
ALL). There can be no assurance that patients in ongoing or future trials of AUTO1, AUTO3, AUTO4 or any of our other product candidates will not
experience more severe CRS, unacceptable levels of neurotoxicity or other serious adverse events.

Our clinical trials include cancer patients who are very sick and whose health is deteriorating, and we expect that additional clinical trials of our
other product candidates will include similar patients with deteriorating health. It is possible that some of these patients may experience similar adverse
side effects as were observed in clinical trials conducted by other companies and academic institutions involving CAR T cells, and that additional patients
may  die  during  our  clinical  trials  for  various  reasons,  including  as  a  result  of  receiving  our  product  candidates,  because  the  patient’s  disease  is  too
advanced, or because the patient experiences medical problems that may not be related to our product candidate. Even if the deaths are not related to our
product candidate, the deaths could affect perceptions regarding the safety of our product candidate.

Patient deaths and severe side effects caused by our product candidates, or by products or product candidates of other companies that are thought to
have similarities with our therapeutic candidates, could result in the delay, suspension, clinical hold or termination of clinical trials by us, the FDA, the
EMA or other regulatory authorities for a number of reasons. If we elect or are required to delay, suspend or terminate any clinical trial of any product
candidates that we develop, the commercial prospects of such product candidates will be harmed and our ability to generate product revenues from any of
these product candidates would be delayed or eliminated. Serious adverse events observed in clinical trials could hinder or prevent market acceptance of
the product candidate at issue. Any of these occurrences may harm our business, prospects, financial condition and results of operations significantly.

If  the  clinical  trials  of  any  of  our  product  candidates  fail  to  demonstrate  safety  and  efficacy  to  the  satisfaction  of  the  FDA,  the  EMA  or  other
comparable regulatory authorities, or do not otherwise produce favorable results, we may incur additional costs or experience delays in completing, or
ultimately be unable to complete, the development and commercialization of our product candidates.

We may not commercialize, market, promote or sell any product candidate without obtaining marketing approval from the FDA, the EMA or other
comparable regulatory authority, and we may never receive such approvals. It is impossible to predict accurately when or if any of our product candidates
will  prove  effective  or  safe  in  humans  and  will  receive  regulatory  approval.  Before  obtaining  marketing  approval  from  regulatory  authorities  for  the
commercial sale of any of our product candidates, we must demonstrate through lengthy, complex and expensive preclinical testing and clinical trials that
our product candidates are both safe and effective for use in each target indication. Clinical testing is expensive, difficult to design and implement, can take
many years to complete and is uncertain as to outcome. A failure of one or more clinical trials can occur at any stage of testing.

We may experience numerous unforeseen events prior to, during, or as a result of, clinical trials that could delay or prevent our ability to receive

marketing approval or commercialize any of our product candidates, including:

•

•

the FDA, the EMA or other comparable regulatory authority may disagree as to the number, design or implementation of our clinical trials, or may
not interpret the results from clinical trials as we do;

regulators  or  institutional  review  boards  may  not  authorize  us  or  our  investigators  to  commence  a  clinical  trial  or  conduct  a  clinical  trial  at  a
prospective trial site;

• we may not reach agreement on acceptable terms with prospective clinical trial sites, the terms of which can be subject to extensive negotiation

and may vary significantly among different clinical trial sites;

•

clinical trials of our product candidates may produce negative or inconclusive results;

• we may decide, or regulators may require us, to conduct additional clinical trials or abandon product development programs;

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the number of patients required for clinical trials of our product candidates may be larger than we anticipate, enrollment in these clinical trials may
be slower than we anticipate, participants may drop out of these clinical trials at a higher rate than we anticipate or we may fail to recruit suitable
patients to participate in a trial;

our third-party contractors may fail to comply with regulatory requirements or meet their contractual obligations to us in a timely manner, or at all;

regulators  may  issue  a  clinical  hold,  or  regulators  or  institutional  review  boards  may  require  that  we  or  our  investigators  suspend  or  terminate
clinical research for various reasons, including noncompliance with regulatory requirements or a finding that the participants are being exposed to
unacceptable health risks;

the cost of clinical trials of our product candidates may be greater than we anticipate;

the FDA, the EMA or other comparable regulatory authorities may fail to approve our manufacturing processes or facilities;

the supply or quality of our product candidates or other materials necessary to conduct clinical trials of our product candidates may be insufficient
or inadequate;

our  product  candidates  may  have  undesirable  side  effects  or  other  unexpected  characteristics,  particularly  given  their  novel,  first-in-
human application, such as cytokine-induced toxicity and T cell aplasia, causing us or our investigators, regulators or institutional review boards
to suspend or terminate the clinical trials; and

the  approval  policies  or  regulations  of  the  FDA,  the  EMA  or  other  comparable  regulatory  authorities  may  significantly  change  in  a  manner
rendering our clinical data insufficient for approval.

To the extent that the results of the trials are not satisfactory for the FDA, the EMA or regulatory authorities in other countries or jurisdiction to
approve our BLA, Marketing Approval Application, or MAA, or other comparable application, the commercialization of our product candidates may be
significantly delayed, or we may be required to expend significant additional resources, which may not be available to us, to conduct additional trials in
support of potential approval of our product candidates.

We  may  not  be  able  to  successfully  create  our  own  manufacturing  infrastructure  for  supply  of  our  requirements  of  programmed  T  cell  product
candidates for use in clinical trials and for commercial sale.

Our manufacturing and commercialization strategy is based on establishing a fully integrated vein-to-vein product delivery cycle. Over time, we
expect  to  establish  regional  manufacturing  hubs  to  service  major  markets  to  meet  projected  needs  for  commercial  sale  quantities.  However,  we  do  not
currently own any facility that may be used as our clinical-scale manufacturing and processing facility and we currently use facilities and equipment at the
Cell and Gene Therapy Catapult, as well as third party vendors, for vector and cell manufacturing.

We are establishing our viral vector and cell manufacturing capacity by taking occupancy of manufacturing suites at the Cell and Gene Therapy
Catapult manufacturing center in Stevenage, United Kingdom, as well as several smaller facilities in the Stevenage, United Kingdom area. The Cell and
Gene Therapy Catapult manufacturing center provides shared infrastructure to collaborators working in segregated manufacturing suites. We have little to
no control over the actions of other collaborators and their actions could inadvertently damage or delay our ability to manufacture our product candidates.
In addition, we rely  on  external  vendors  to  manufacture  viral  vector  for  certain  of  our  product  candidates.  Our  long-term  plan  is  to  establish  additional
manufacturing sites in the United States and in Europe as needed. The implementation of this plan is subject to many risks. For example, the establishment
of a cell-therapy manufacturing facility is a complex endeavor requiring knowledgeable individuals. Creating an internal manufacturing infrastructure will
rely upon finding personnel with an appropriate background and training to staff and operate the facility. Should we be unable to find these individuals, we
may need to rely on external contractors or train additional personnel to fill the needed roles. There are a small number of individuals with experience in
cell therapy and the competition for these individuals is high.

We expect that the establishment of our own commercial cell manufacturing facilities will provide us with enhanced control of product supply for
both  clinical  trials  and  the  commercial  market,  enable  the  more  rapid  implementation  of  process  changes,  and  allow  for  better  long-term  cost  margins.
However, we have no experience as a company in designing and operating a commercial manufacturing facility and may never be successful in developing
our  own  manufacturing  facility  or  capability.  We  may  establish  additional  manufacturing  sites  as  we  expand  our  commercial  footprint  to  multiple
geographies,  which  may  lead  to  regulatory  delays  or  prove  costly.  Even  if  we  are  successful,  our  manufacturing  operations  could  be  affected  by  cost-
overruns, unexpected delays, equipment failures, labor shortages, natural disasters, power failures and numerous other factors, or we may not

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be successful in establishing sufficient capacity to produce our product candidates in sufficient quantities to meet the requirements for the potential launch
or to meet potential future demand, all of which could prevent us from realizing the intended benefits of our manufacturing strategy and have a material
adverse effect on our business.

We may not be successful in achieving cost of goods at commercial scale that provide for an attractive margin.

We believe that our current, fully enclosed manufacturing processes are fit for commercial scale and we anticipate they will enable commercial
supply at an economical cost. However, we have not yet established manufacturing capacity at commercial scale and may underestimate the cost and time
required to do so, or overestimate cost reductions from economies of scale that can be realized with our manufacturing processes. We may ultimately be
unable to manage the cost of goods for our product candidates to levels that will allow for a margin in line with our expectations and return on investment if
and when those product candidates are commercialized.

Our  product  candidates  are  biologics  and  the  manufacture  of  our  product  candidates  is  complex  and  we  may  encounter  difficulties  in  production,
particularly  with  respect  to  process  development  or  scaling-out  of  our  manufacturing  capabilities.  If  we  encounter  such  difficulties,  our  ability  to
provide supply of our product candidates for clinical trials or our products for patients, if approved, could be delayed or stopped.

We have developed a process for manufacturing programmed T cells in a fully enclosed system designed to minimize the risk of contamination, and
we  have  improved  the  viral  transduction  process  to  help  eliminate  processing  inconsistencies.  We  believe  that  our  current  processes  are  suitable  for
commercialization.  While  we  have  established  a  process  which  we  believe  is  scalable  for  commercial  production,  each  manufacturing  process  must  be
validated through the performance of process validation runs to guarantee that the facility, personnel, equipment, and process work as designed. We have
not yet manufactured or processed our product candidates on a commercial scale and may not be able to do so for any of our product candidates.

We,  like  other  manufacturers  of  biologic  products,  may  encounter  difficulties  in  production,  particularly  in  scaling  up  or  out,  validating  the
production process, and assuring high reliability of the manufacturing process. These problems include delays or break-downs in logistics and shipping,
difficulties with production costs and yields, quality control, and product testing, operator error, lack of availability of qualified personnel, as well as failure
to comply with strictly enforced federal, state and foreign regulations.

Furthermore, if microbial, viral or other contaminations are discovered in our supply of product candidates or in the manufacturing facilities, such
manufacturing facilities may need to be closed for an extended period of time to investigate and remedy the contamination. We cannot assure you that any
of these or other issues relating to the manufacture of our product candidates will not occur in the future. Any delay or interruption in the supply of clinical
trial supplies could delay the completion of clinical trials, increase the costs associated with maintaining clinical trial programs and, depending upon the
period of delay, require us to begin new clinical trials at additional expense or terminate clinical trials completely.

The manufacture and delivery of programmed T cell therapies to patients involves complex, integrated processes, including harvesting T cells from
patients, programming the T cells ex vivo, multiplying the T cells to obtain the desired dose, and ultimately infusing the T cells back into a patient’s body.
As a result of the complexities, the cost to manufacture biologics in general, and our programmed T cell product candidates in particular, is generally higher
than  traditional  small  molecule  chemical  compounds,  and  the  manufacturing  process  is  less  reliable  and  is  more  difficult  and  costly  to  reproduce.  In
addition, our manufacturing process will be susceptible to product loss or failure due to logistical issues associated with the collection of white blood cells
from the patient, shipping such patient material to the manufacturing site, storing and processing such patient material, shipping the patient material with
the programmed T cells back to the patient, and infusing the patient with the final product. Other manufacturing issues include the differences in patient
starting  materials,  inconsistency  in  cell  growth,  variability  in  product  characteristics,  interruptions  in  the  manufacturing  process,  equipment  or  reagent
failure, improper installation or operation of equipment, and vendor or operator error. Even minor deviations from normal manufacturing processes could
result in reduced production yields, product defects, and other supply disruptions. For  example,  in  clinical  trials  of  AUTO1  conducted  by  UCL  using  a
manufacturing process that differs from our semi-automated manufacturing process, UCL experienced product failures for three patients enrolled in the
CARPALL trial and produced only a partial dose for one patient in the ALLCAR19 trial. If we lose, destroy or otherwise impair the patient materials at any
point in the vein-to-vein supply chain, the manufacturing process for that patient will need to be restarted and the resulting delay may adversely affect that
patient’s outcome due to the risk of disease progression. In addition, because our product candidates are manufactured for each particular patient, we will
be required to maintain a chain of identity with respect to materials as they move from the patient to the manufacturing facility, through the manufacturing
process, and back to the patient. Maintaining such a chain of identity is difficult and complex, and failure to do so could result in adverse patient outcomes,
loss of product, or regulatory action including withdrawal of our products from the market. Further, as product candidates are developed through preclinical
to late stage clinical trials towards approval and commercialization, it is common that various aspects of the development program, such as manufacturing
methods, are altered along the way in an effort to optimize processes and results.

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Such changes carry the risk that they will not achieve these intended objectives, and any of these changes could cause our product candidates to perform
differently and affect the results of planned clinical trials or other future clinical trials.

Our manufacturing facilities also require commissioning and validation activities to demonstrate that they operate as designed, and are subject to
government inspections by the FDA, the EMA and other comparable regulatory authorities. If we are unable to reliably produce products to specifications
acceptable to the regulatory authorities, we may not obtain or maintain the approvals we need to manufacture our products. Further, our facilities may fail
to pass government inspections prior to or after the commercial launch of our product candidates, which would cause significant delays and additional costs
required to remediate any deficiencies identified by the regulatory authorities. Any of these challenges could delay completion of clinical trials, require
bridging  clinical  trials  or  the  repetition  of  one  or  more  clinical  trials,  increase  clinical  trial  costs,  delay  approval  of  our  product  candidate,  impair
commercialization  efforts,  increase  our  cost  of  goods,  and  have  an  adverse  effect  on  our  business,  financial  condition,  results  of  operations  and  growth
prospects.

Prior treatments can alter the cancer and negatively impact chances for achieving clinical activity with our programmed T cells.

Patients with hematological cancers receive highly toxic lympho-depleting chemotherapy as their initial treatments. These therapies can impact the
viability of the T cells collected from the patient and can contribute to highly variable responses to programmed T cell therapies. Patients could also have
received prior therapies that target the same target antigen on the cancer cells as our intended programmed T cell product candidate and thereby lead to a
selection of cancer cells with low or no expression of the target. As a result, our programmed T cell product candidates may not recognize the cancer cell
and may fail to achieve clinical activity. Both of our most advanced product candidates, AUTO1 and AUTO3, may face this challenge. For example, ALL
patients could have received blinatumomab or Kymriah, or a CD19 ADC, or a CD22 targeting CAR T, or CD22 ADC, like inotuzomab, or similar products
or  product  candidates  prior  to  receiving  AUTO1;  and  DLBCL  patients  could  have  received  Yescarta,  Kymriah,  JCAR-17,  inotuzomab,  CD22-targeting
CAR or blinatumomab, or similar products or product candidates prior to receiving AUTO3. If any of our product candidates do not achieve a sufficient
level of clinical activity, we may discontinue the development of that product candidate, which could have an adverse effect on the value of our ADSs.

We may expend our resources to pursue a particular product candidate or indication and fail to capitalize on product candidates or indications that
may be more profitable or have a greater likelihood of success.

Because we have limited financial and management resources, we focus on research programs and product candidates that we identify for specific
indications.  As  a  result,  we  may  forego  or  delay  pursuit  of  opportunities  with  other  product  candidates  or  for  other  indications  that  later  prove  to  have
greater  commercial  potential.  Our  resource  allocation  decisions  may  cause  us  to  fail  to  capitalize  on  viable  commercial  products  or  profitable  market
opportunities. Our spending on current and future research and development programs and product candidates for specific indications may not yield any
commercially  viable  products.  If  we  do  not  accurately  evaluate  the  commercial  potential  or  target  market  for  a  particular  product  candidate,  we  may
relinquish valuable rights to that product candidate through collaboration, licensing or other royalty arrangements in cases in which it would have been
more advantageous for us to retain sole development and commercialization rights to such product candidate.

We plan to seek, but may fail to obtain “breakthrough therapy” designation or “regenerative medicine advanced therapy” (RMAT) designation from
the  FDA  and  “PRIME”  designation  from  the  EMA,  and  may  pursue  accelerated  approval  for  some  or  all  of  our  programmed  T  cell  product
candidates, which may prolong the regulatory approval process for our product candidates.

In 2012, the FDA established a breakthrough therapy designation which is intended to expedite the development and review of product candidates
that treat serious or life-threatening diseases when “preliminary clinical evidence indicates that the drug may demonstrate substantial improvement over
existing  therapies  on  one  or  more  clinically  significant  endpoints,  such  as  substantial  treatment  effects  observed  early  in  clinical  development.”  The
designation  of  a  product  candidate  as  a  breakthrough  therapy  provides  potential  benefits  that  include  more  frequent  meetings  with  FDA  to  discuss  the
development plan for the product candidate and ensure collection of appropriate data needed to support approval; more frequent written correspondence
from the FDA about such things as the design of the proposed clinical trials and use of biomarkers; guidance on an efficient drug development program,
beginning as early as Phase 1; organizational commitment involving senior managers; and eligibility for rolling review and priority review. The frequency
of communication from the FDA is intended to allow for questions and issues to be resolved quickly, which often leads to earlier drug approval and access
by patients.

RMAT was introduced as a new designation under the 21st Century Cures Act for the development and review of certain regenerative medicine
therapies. To receive RMAT designation, a regenerative medicine product candidate must be intended to treat, modify, reverse, or cure a serious or life-
threatening  disease  or  condition  with  preliminary  clinical  evidence  indicating  that  the  drug  has  the  potential  to  address  unmet  medical  need.  RMAT
designation does not require evidence to indicate that the drug may offer a substantial improvement over available therapies, as breakthrough designation
requires. In November 2017, the FDA released draft guidance that clarified that gene therapies, including genetically modified cells, that lead to a durable
modification of

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cells or tissues, may meet the definition of a regenerative medicine therapy for RMAT designation. Similar to breakthrough designation, an RMAT product
candidate  receives:  intensive  guidance  on  an  efficient  drug  development  program;  intensive  involvement  of  senior  managers  and  experienced  staff  on  a
proactive, collaborative and cross-disciplinary review; and a rolling review. Regenerative medicine therapies that qualify for RMAT designation may also
qualify for other FDA expedited programs, if they meet the criteria for such programs.

Similarly,  the  EMA  has  established  the  PRIME  scheme  to  expedite  the  development  and  review  of  product  candidates  that  show  a  potential  to

address to a significant extent an unmet medical need, based on early clinical data.

We intend to seek breakthrough therapy designation, RMAT designation or PRIME designation for some or all of our programmed T cell product
candidates that may qualify. There is no assurance that we will obtain breakthrough therapy designation or RMAT designation, or that we will obtain access
to PRIME for any of our product candidates. Breakthrough therapy designation and PRIME eligibility do not change the standards for product approval,
and there is no assurance that such designation or eligibility will result in expedited review or approval. Additionally, breakthrough therapy designation and
access to PRIME can each be revoked if the criteria for eligibility cease to be met as clinical data emerges.

We may also seek accelerated approval for certain of our product candidates. Under the FDA’s fast track and accelerated approval programs, the
FDA  may  approve  a  drug  or  biologic  for  a  serious  or  life-threatening  illness  that  provides  meaningful  therapeutic  benefit  to  patients  over  existing
treatments based upon a surrogate endpoint that is reasonably likely to predict clinical benefit, or on a clinical endpoint that can be measured earlier than
irreversible morbidity or mortality, that is reasonably likely to predict an effect on irreversible morbidity or mortality or other clinical benefit, taking into
account the severity, rarity, or prevalence of the condition and the availability or lack of alternative treatments. For drugs granted accelerated approval,
post-marketing  confirmatory  trials  have  been  required  to  describe  the  anticipated  effect  on  irreversible  morbidity  or  mortality  or  other  clinical  benefit.
These  confirmatory  trials  must  be  completed  with  due  diligence.  Moreover,  the  FDA  may  withdraw  approval  of  our  indication  approved  under  the
accelerated approval pathway if, for example:

•

•

the  trial  or  trials  required  to  verify  the  predicted  clinical  benefit  of  our  product  candidates  fail  to  verify  such  benefit  or  do  not  demonstrate
sufficient clinical benefit to justify the risks associated with the drug;

other evidence demonstrates that our product candidates are not shown to be safe or effective under the conditions of use;

• we fail to conduct any required post-approval trial of our product candidates with due diligence; or

• we disseminate false or misleading promotional materials relating to the relevant product candidate.

 Risks Related to our Business Operations

The effects of health epidemics, including the ongoing global coronavirus COVID-19 pandemic, in regions where we, or the third parties on which we
rely, have business operations could adversely impact our business, including our clinical trials, preclinical studies and supply chains, depending on
the location, duration and severity of disruptions to the systems affecting our business.

Beginning in late 2019, the outbreak of a novel strain of coronavirus, COVID-19, has evolved into a global pandemic and in March 2020, the
World  Health  Organization  characterized  COVID-19  as  a  pandemic.  The  COVID-19  pandemic,  which  has  continued  to  spread,  and  the  related  adverse
public  health  developments,  including  orders  to  shelter-in-place,  travel  restrictions,  and  the  imposition  of  additional  requirements  on  businesses,  have
adversely affected workforces, organizations, healthcare communities, economies, and financial markets globally, leading to an economic downturn and
increased  market  volatility.  The  U.S.  government  also  has  imposed  travel  restrictions  on  travel  between  the  United  States,  United  Kingdom  and  certain
other  countries.  It  has  also  disrupted  the  normal  operations  of  businesses  across  industries,  including  ours,  and  caused  significant  disruption  in  the
operations of third party manufacturers and contract research organizations, or CROs, upon whom we rely.
    In response to the spread of COVID-19 as well as public health directives and orders, we have implemented work-from-home policies to support the
community efforts to reduce the transmission of COVID-19 and protect employees, complying with guidance from government authorities in the countries
within which we operate. We have implemented a number of measures to ensure employee safety and business continuity, including limiting access to our
laboratory and manufacturing facilities to only those individuals required to execute their job responsibilities and restricted the number of staff working
concurrently  in  any  given  laboratory.  Our  company  headquarters  is  located  in  London,  our  U.S.  headquarters  is  located  in  the  Baltimore-Washington
metropolitan area, and our CROs and CMOs are located in the United States, the United Kingdom, and European Union.

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    We are currently conducting clinical trials in the United States, United Kingdom and Spain. Timely enrollment in our clinical trials is dependent upon
capacity at our clinical trial sites, some of which have been and are currently adversely affected by COVID-19. Due to the uneven geographic impact of the
pandemic, these localized disruptions are difficult to predict. Shutdowns or other restrictions related to COVID-19 or other infectious diseases could impact
personnel  at  third-party  manufacturing  facilities,  which,  in  turn,  could  impact  the  availability  or  cost  of  materials  and  disrupt  our  supply  chain.
Additionally, many of our clinical trials involve immunocompromised patients who are at higher risk for COVID-19 and who are therefore more likely to
avoid hospitals or other high-risk areas.

        The  effects  of  the  governmental  restrictions  and  guidelines,  and  the  measures  we  have  implemented  to  comply  with  them,  may  negatively  impact
productivity, disrupt our business and delay our clinical programs and timelines (for example, our timelines for AUTO1 and AUTO3). The magnitude of
these potential disruptions will depend, in part, on the length and severity of the restrictions and other limitations on our ability to conduct our business in
the ordinary course.

    As a result of the COVID-19 outbreak, or similar pandemics, we may experience disruptions that could severely impact our business, clinical trials and
preclinical studies, including:

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•

•

delays  or  difficulties  in  enrolling  patients  in  our  clinical  trials,  including  travel  restrictions  on  patients  and  constraints  on  the  capacity  of  our
clinical trial sites;
delays or difficulties in clinical site initiation, including difficulties in training clinical site investigators and clinical site staff;
delays or disruptions in non-clinical experiments and investigational new drug application-enabling good laboratory practice standard toxicology
studies due to unforeseen circumstances in supply chain;
delays or disruptions in our ability to manufacture programmed T cell therapies due to supply chain and transportation system disruptions or lack
of manufacturing staff;
increased  rates  of  patients  withdrawing  from  our  clinical  trials  following  enrollment  as  a  result  of  contracting  COVID-19,  being  forced  to
quarantine, or being unable to visit clinical trial locations;
diversion or prioritization of healthcare resources away from the conduct of clinical trials and towards the COVID-19 pandemic, including the
diversion of hospitals serving as our clinical trial sites and hospital staff supporting the conduct of our clinical trials, particularly for clinical trials
that require in-patient monitoring following administration of the product candidate;
delays or disruptions in the availability of clinical site staff, who, as healthcare providers, may have heightened exposure to COVID-19, which, in
turn, could adversely impact our clinical trial operations;
interruption of our key clinical trial activities, such as clinical assessments at pre-specified time points during the trial and clinical trial site data
monitoring, due to limitations on travel imposed or recommended by governmental entities, employers and others or interruption of clinical trial
subject visits and study procedures (particularly any procedures that may be deemed non-essential), which may impact the integrity of subject data
and clinical study endpoints;
interruption  or  delays  in  the  operations  of  the  U.S.  Food  and  Drug  Administration,  European  Medicines  Agency  and  comparable  foreign
regulatory agencies or their refusal to accept data from clinical trials in affected geographies, which may impact approval timelines;
delays  in  necessary  interactions  with  local  regulators,  ethics  committees  and  other  important  agencies  and  contractors  due  to  limitations  in
employee resources or forced furlough of government employees;
limitations on employee resources that would otherwise be focused on the conduct of our preclinical studies and clinical trials, including because
of sickness of employees or their families, the desire of employees to avoid contact with large groups of people, an increased reliance on working
from home or mass transit disruptions; and
reduced ability to engage with the medical and investor communities due to the cancellation of conferences scheduled throughout the year.

    These and other factors arising from the COVID-19 pandemic could worsen in countries that are already afflicted with COVID-19, could continue to
spread to additional countries, or could return to countries where the pandemic has been partially contained, each of which could continue to adversely
impact our ability to conduct clinical trials and our business generally, and could have a material adverse impact on our operations and financial condition
and results.

    In addition, the trading price for our ADSs as well as trading price for the publicly traded securities of other biopharmaceutical companies, as well as the
broader global financial markets, have been highly volatile as a result of the COVID-19 pandemic and the resulting impact on U.K. and U.S. economic
activities. As a result, we may face difficulties raising capital when needed, and any such sales may be on unfavorable terms to us. Further, to the extent we
raise additional capital through the sale of equity or convertible debt securities, the ownership interest of existing shareholders will be diluted.

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The ultimate impact of the COVID-19 pandemic on our business operations is highly uncertain and subject to change and will depend on future
developments which are difficult to predict, including the duration of the pandemic, the ultimate geographic spread of the disease, additional or modified
government actions, new information that will emerge concerning the severity and impact of COVID-19 and other actions taken to contain or address its
impact in the short and long term, among others. We do not yet know the full extent of potential delays or impacts on our business, our commercialization
efforts, our clinical studies, our research programs, healthcare systems or the global economy, and if the ultimate impact of the COVID-19 pandemic and
the resulting uncertain economic and healthcare environment is more severe than we anticipated, we may not be able to execute on our current operating
plan or on our strategy. If the duration of the COVID-19 pandemic and the associated period of business and social restrictions and economic uncertainty is
longer than we anticipated, our cash, cash equivalents, and marketable securities may not be sufficient to fund the activities under our operating plan for the
time period that we anticipated, and we may be required to revise our operating plan further. To the extent the COVID-19 pandemic adversely affects our
business and financial results, it may also have the effect of heightening many of the other risks described in this “Risk Factors” section.

As a company based outside of the United States, our business is subject to economic, political, regulatory and other risks associated with international
operations.

Our business is subject to risks associated with conducting business outside of the United States, as our company is based in the United Kingdom
and conducts operations internationally. Many of our suppliers and clinical trial relationships are located outside the United States. Accordingly, our future
results could be harmed by a variety of factors, including:

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economic weakness, including inflation, or political instability in particular non-U.S. economies and markets;

differing and changing regulatory requirements for product approvals;

differing jurisdictions could present different issues for securing, maintaining or obtaining freedom to operate in such jurisdictions;

potentially reduced protection for intellectual property rights;

difficulties in compliance with different, complex and changing laws, regulations and court systems of multiple jurisdictions and compliance with
a wide variety of foreign laws, treaties and regulations;

changes in non-U.S. regulations and customs, tariffs and trade barriers;

changes in non-U.S. currency exchange rates of the pound sterling, U.S. dollar, euro and currency controls;

changes  in  a  specific  country’s  or  region’s  political  or  economic  environment,  including  the  implications  of  the  United  Kingdom's  withdrawal
from the European Union;

trade protection measures, import or export licensing requirements or other restrictive actions by governments;

differing reimbursement regimes and price controls in certain non-U.S. markets;

negative consequences from changes in tax laws;

compliance with tax, employment, immigration and labor laws for employees living or traveling abroad, including, for example, the variable tax
treatment in different jurisdictions of options granted under our share option schemes or equity incentive plans;

• workforce uncertainty in countries where labor unrest is more common than in the United States;

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•

•

litigation or administrative actions resulting from claims against us by current or former employees or consultants individually or as part of class
actions, including claims of wrongful terminations, discrimination, misclassification or other violations of labor law or other alleged conduct;

difficulties associated with staffing and managing international operations, including differing labor relations;

production shortages resulting from any events affecting raw material supply or manufacturing capabilities abroad; and

business interruptions resulting from geo-political actions, including war and terrorism, natural disasters--including earthquakes, typhoons, floods
and fires--or health epidemics, such as the coronavirus pandemic.

Exchange rate fluctuations may materially affect our results of operations and financial condition.

Our functional currency and that of our subsidiaries is the pound sterling and our reporting currency is the U.S. dollar. Given that our functional
currency and that of our subsidiaries is the pound sterling, but our reporting currency is the U.S. dollar, fluctuations in currency exchange rates between the
U.S.  dollar  and  the  pound  sterling  could  materially  and  adversely  affect  our  business.  There  may  be  instances  in  which  costs  and  revenue  will  not  be
matched with respect to currency denomination. Currently, we do not have any exchange rate hedging arrangements in place.

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Additionally, although we are based in the United Kingdom, we source research and development, manufacturing, consulting and other services
from the United States and other countries. Further, potential future revenue may be derived from the United States, countries within the euro zone, and
various other countries around the world. As a result, our business and the price of our ADSs may be affected by fluctuations in foreign exchange rates not
only  between  the  pound  sterling  and  the  U.S.  dollar,  but  also  the  euro  and  other  currencies,  which  may  have  a  significant  impact  on  our  results  of
operations and cash flows from period to period. As a result, to the extent we continue our expansion on a global basis, we expect that increasing portions
of  our  revenue,  cost  of  revenue,  assets  and  liabilities  will  be  subject  to  fluctuations  in  currency  valuations.  We  may  experience  economic  loss  and  a
negative impact on earnings or net assets solely as a result of currency exchange rate fluctuations.

We will need to manage the size of our organization, and we may experience difficulties.

As of December 31, 2020, we had 384 employees, 376 of whom are full-time. In January 2021, we announced adjustments to our workforce and
infrastructure footprint, which will involve an overall reduction in headcount of approximately 20%. As our development and commercialization plans and
strategies  develop,  and  as  we  further  develop  as  a  public  company,  we  may  need  additional  managerial,  operational,  financial  and  other  personnel,
including  personnel  to  support  our  product  development  and  planned  future  commercialization  efforts.  Future  growth  will  impose  significant  added
responsibilities on members of management, including:

•

identifying, recruiting, integrating, maintaining and motivating additional employees;

• managing our internal development efforts effectively, including the clinical, FDA and EMA review processes for our product candidates; and

•

improving our operational, financial and management controls, reporting systems and procedures.

          There  are  a  small  number  of  individuals  with  experience  in  cell  therapy  and  the  competition  for  these  individuals  is  high.  Our  future  financial
performance and our ability to commercialize our product candidates will depend, in part, on our ability to effectively manage any future growth, and our
management may also have to divert a disproportionate amount of its attention away from day-to-day activities in order to devote a substantial amount of
time to managing these growth activities.

If we are not able to effectively manage the size of our organization, we may not be able to successfully implement the tasks necessary to further

develop and commercialize our product candidates and, accordingly, may not achieve our research, development and commercialization goals.

In addition to expanding our organization, we are building out our development and manufacturing capabilities, which requires significant capital
expenditures. If these capital expenditures are higher than expected, it may adversely affect our financial condition and capital resources. In addition, if the
availability of manufacturing capacity is delayed, it may limit our ability to rapidly expand the size of our organization in order to meet our corporate goals.

Our future success depends on our ability to retain key members of senior management and to attract, retain and motivate qualified personnel.

Our  ability  to  compete  in  the  highly  competitive  biopharmaceutical  industry  depends  upon  our  ability  to  attract  and  retain  highly  qualified
management, research and development, clinical, financial and business development personnel. We are highly dependent on our management, scientific
and  medical  personnel,  including  Dr.  Christian  Itin,  our  Chief  Executive  Officer  and  Dr.  Martin  Pulé,  our  scientific  founder,  Senior  Vice  President  and
Chief Scientific Officer. Each member of our senior management may terminate their employment with us at any time. We do not maintain ‘‘key person’’
insurance for any of our employees.

Recruiting  and  retaining  qualified  scientific  and  clinical  personnel  and,  if  we  progress  the  development  of  any  of  our  product  candidates,
commercialization, manufacturing and sales and marketing personnel, will be critical to our success. The loss of the services of members of our senior
management or other key employees could impede the achievement of our research, development and commercialization objectives and seriously harm our
ability to successfully implement our business strategy. Furthermore, replacing members of our senior management and key employees may be difficult and
may take an extended period of time because of the limited number of individuals in our industry with the breadth of skills and experience required to
successfully develop, gain regulatory approval of and commercialize our product candidates. Our success also depends on our ability to continue to attract,
retain  and  motivate  highly  skilled  junior,  mid-level  and  senior  managers,  as  well  as  junior,  mid-level  and  senior  scientific  and  medical  personnel.
Competition to hire from this limited candidate pool is intense, and we may be unable to hire, train, retain or motivate these key personnel on acceptable
terms given the competition among numerous pharmaceutical and biotechnology companies for similar personnel. We also experience competition for the
hiring of scientific and clinical personnel from universities and research institutions. In addition, we rely on consultants and advisors, including scientific
and  clinical  advisors,  to  assist  us  in  formulating  our  research  and  development  and  commercialization  strategy.  Our  consultants  and  advisors  may  have
commitments under consulting or advisory contracts with other entities that may limit their availability to us. If we are unable to continue to attract and
retain high-quality personnel, our ability to pursue our growth strategy will be limited.

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We  will  require  additional  funding  to  continue  our  planned  operations.  If  we  engage  in  future  acquisitions  or  strategic  collaborations,  this  may
increase our capital requirements, dilute our shareholders, cause us to incur debt or assume contingent liabilities and subject us to other risks.

We  will  need  additional  capital  in  the  future  to  continue  our  planned  operations.  To  the  extent  we  raise  additional  capital  by  issuing  equity
securities, our shareholders may experience substantial dilution. We may sell ordinary shares or ADSs, convertible securities or other equity securities in
one or more transactions at prices and in a manner we determine from time to time. If we sell ordinary shares or ADSs, convertible securities or other
equity securities in more than one transaction, investors may be materially diluted by subsequent sales. These sales may also result in material dilution to
our  existing  shareholders,  and  new  investors  could  gain  rights  superior  to  our  existing  shareholders.  In  addition,  we  have  an  Open  Market  Sale
Agreement
, or Sales Agreement, with Jefferies LLC, or Jefferies, to sell up to $100.0 million worth of our ADSs, from time to time, through an “at the
market” equity offering program under which Jefferies will act as sales agent. As of March 1, 2021, $84.1 million worth of ADSs remained available for
sale under the “at the market” equity offering program. If we raise additional capital through our “at the market” equity offering program, or other public or
private  equity  offerings,  the  ownership  interest  of  our  existing  shareholders  will  be  diluted  and  may  cause  the  market  price  of  our  ADSs  to  decline.
Furthermore,  new  investors  purchasing  securities  that  we  may  issue  and  sell  in  the  future  could  obtain  rights  superior  to  the  rights  of  our  existing
shareholders.

SM

From  time  to  time,  we  may  also  evaluate  various  acquisitions  and  strategic  collaborations,  including  licensing  or  acquiring  complementary
products, intellectual property rights, technologies or businesses, as we may deem appropriate to carry out our business plan. Any potential acquisition or
strategic collaboration may entail numerous risks, including:

•

•

•

•

•

•

•

increased operating expenses and cash requirements;

the assumption of additional indebtedness or contingent liabilities;

assimilation  of  operations,  intellectual  property  and  products  of  an  acquired  company,  including  difficulties  associated  with  integrating  new
personnel;

the  diversion  of  our  management’s  attention  from  our  existing  programs  and  initiatives  in  pursuing  such  a  strategic  partnership,  merger  or
acquisition;

retention of key employees, the loss of key personnel and uncertainties in our ability to maintain key business relationships;

risks and uncertainties associated with the other party to such a transaction, including the prospects of that party and their existing products or
product candidates and regulatory approvals; and

our inability to generate revenue from acquired technology sufficient to meet our objectives in undertaking the acquisition or even to offset the
associated acquisition and maintenance costs.

Additionally, if we undertake acquisitions, we may issue dilutive securities, assume or incur debt obligations, incur large one-time expenses and
acquire  intangible  assets  that  could  result  in  significant  future  amortization  expenses.  Moreover,  we  may  not  be  able  to  locate  suitable  acquisition
opportunities and this inability could impair our ability to grow or obtain access to technology or products that may be important to the development of our
business.

Our internal computer systems, or those of our future collaborators or other contractors or consultants, may fail or suffer security breaches, which
could result in a significant disruption of our product development programs and our ability to operate our business effectively.

Our internal computer systems and those of our current and any future collaborators and other contractors or consultants are vulnerable to damage
from computer viruses, unauthorized access, natural disasters, terrorism, war and telecommunication and electrical failures. While we have not experienced
any significant system failure, accident or security breach to date, if such an event were to occur and cause interruptions in our operations, it could result in
a disruption of our development programs and our business operations, whether due to a loss of our trade secrets or other proprietary information or other
similar  disruptions.  For  example,  the  loss  of  clinical  trial  data  from  completed  or  future  clinical  trials  could  result  in  delays  in  our  regulatory  approval
efforts and significantly increase our costs to recover or reproduce the data. To the extent that any disruption or security breach were to result in a loss of, or
damage to, our data or applications, or inappropriate disclosure of confidential or proprietary information, we could incur liability, our competitive position
could be harmed and the further development and commercialization of our product candidates could be delayed.

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European data collection is governed by restrictive regulations governing the use, processing, and cross-border transfer of personal information.

The  collection,  use,  disclosure,  transfer,  or  other  processing  of  personal  data  (including  data  concerning  health  and/or  genetic  data)  regarding
individuals in the European Economic Area, or EEA, or otherwise carried out in the context of any establishment in the EEA, is subject to the provisions of
the EU General Data Protection Regulation, or the GDPR, which became effective and enforceable across all then-current member states of the EEA on
May 25, 2018. Further, by operation of the UK GDPR, as defined below, the GDPR continues to apply in substantially equivalent form in the context of
UK establishments and UK-related processing operations. Where we refer to the GDPR in this risk factor, we are also making reference to the UK GDPR
in the context of the UK, unless we make a specific distinction between the UK and EU GDPR or the context requires otherwise.

Both  the  GDPR  and  UK  GDPR  have  “extra-territorial”  reach  in  that  they  apply  to  processing  of  personal  data  carried  out  in  the  context  of  a
controller’s or processor’s establishment in the EEA/UK (as applicable), and also to the processing of personal data by a controller or processor with no
establishment in the EEA/UK (as applicable) where the processing in question concerns the offering of goods or services to, and/or the monitoring of the
behavior of, individuals in the EEA/UK (as applicable).

The  GDPR  sets  out  a  number  of  requirements  that  must  be  complied  with  when  handling  personal  data  (i.e.,  data  relating  to  an  identified  or
identifiable living individual) such as: limiting permitted processing of personal data to only that which is necessary for specified, explicit and legitimate
purposes; requiring the establishment a legal basis for processing personal data; expressly confirming that “pseudonymized” or key‑coded data constitutes
personal data to which the GDPR applies; creating obligations for controllers and processors to appoint data protection officers in certain circumstances;
increasing  transparency  obligations  to  data  subjects  for  controllers  (including  presentation  of  certain  information  in  a  concise,  intelligible  and  easily
accessible form about how their personal data is used and their rights vis-à-vis that data and its use); introducing the obligation to carry out so-called data
protection impact assessments in certain circumstances; establishing limitations on collection and retention of personal data through “data minimization”
and “storage limitation” principles; establishing obligations to implement “privacy by design”; introducing obligations to honor increased rights for data
subjects (such as rights for individuals to be ‘forgotten,’ rights to data portability, rights to object etc. in certain circumstances); formalizing a heightened
and codified standard of data subject consent; establishing obligations to implement certain technical and organizational safeguards to protect the security
and  confidentiality  of  personal  data;  introducing  obligations  to  agree  to  certain  specific  contractual  terms  and  to  take  certain  measures  when  engaging
third‑party  processors  and  joint  controllers;  introducing  the  obligation  to  provide  notice  of  certain  significant  personal  data  breaches  to  the  relevant
supervisory authority(ies) and affected individuals; and mandating the appointment of representatives in the UK and/or EU in certain circumstances. The
processing of “special category personal data”, such as health information, may also impose heightened compliance burdens under the GDPR and is a topic
of active interest among relevant regulators.

The GDPR provides that EEA Member States may make their own further laws and regulations to introduce specific requirements related to the
processing of “special categories of personal data”, including personal data related to health, biometric data used for unique identification purposes and
genetic information; as well as personal data related to criminal offences or convictions – in the UK, the Data Protection Act 2018 complements the UK
GDPR in this regard. This fact may lead to greater divergence on the law that applies to the processing of such data types across the EEA and/or UK,
compliance with which, as and where applicable, may increase our costs and could increase our overall compliance risk. Such country-specific regulations
could  also  limit  our  ability  to  collect,  use  and  share  data  in  the  context  of  our  EEA  and/or  UK  operations,  and/or  could  cause  our  compliance  costs  to
increase, ultimately having an adverse impact on our business, and harming our business and financial condition.

The  GDPR  also  provides  for  more  robust  regulatory  enforcement  and  greater  penalties  for  noncompliance  than  previous  data  protection  laws,
including fines of up to €20 million or 4% of global annual revenue of any noncompliant company for the preceding financial year, whichever is higher. In
addition  to  administrative  fines,  a  wide  variety  of  other  potential  enforcement  powers  are  available  to  competent  supervisory  authorities  in  respect  of
potential and suspected violations of the GDPR, including extensive audit and inspection rights, and powers to order temporary or permanent bans on all or
some  processing  of  personal  data  carried  out  by  noncompliant  actors.  The  GDPR  also  confers  a  private  right  of  action  on  data  subjects  and  consumer
associations to lodge complaints with supervisory authorities, seek judicial remedies, and obtain compensation for damages resulting from violations of the
GDPR.

A particular issue presented by certain European data protection laws, including the GDPR, is that they generally restrict transfers of personal data
from Europe, including the EEA, United Kingdom and Switzerland, to the United States and most other countries unless the parties to the transfer have
implemented specific safeguards to protect the transferred personal data. One of the primary safeguards allowing U.S. companies to import personal data
from Europe had been certification to the EU-U.S. Privacy

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Shield  and  Swiss  U.S.  Privacy  Shield  frameworks  administered  by  the  U.S.  Department  of  Commerce.  However,  the  EU  U.S.  Privacy  Shield  was
invalidated in July 2020 by the Court of Justice of the European Union, or CJEU, in a case known colloquially as “Schrems II.” Following this decision,
the United Kingdom government has similarly invalidated use of the EU U.S. Privacy Shield as a mechanism for lawful personal data transfers from the
United  Kingdom  to  the  United  States  under  the  UK  GDPR.  Also,  the  Swiss  Federal  Data  Protection  and  Information  Commissioner,  or  the  FDPIC,
announced that the Swiss-U.S. Privacy Shield does not provide adequate safeguards for the purposes of personal data transfers from Switzerland to the
United  States.  The  CJEU’s  decision  in  Schrems  II  also  raised  questions  about  whether  one  of  the  primary  alternatives  to  the  EU-U.S.  Privacy  Shield,
namely, the European Commission’s Standard Contractual Clauses, can lawfully be used for personal data transfers from Europe to the United States or
other third countries that are not the subject of an adequacy decision of the European Commission. While the CJEU upheld the adequacy of the Standard
Contractual Clauses in principle in Schrems II, it made clear that reliance on those Clauses alone may not necessarily be sufficient in all circumstances. Use
of  the  Standard  Contractual  Clauses  must  now  be  assessed  on  a  case-by-case  basis  taking  into  account  the  legal  regime  applicable  in  the  destination
country, in particular regarding applicable surveillance laws and relevant rights of individuals with respect to the transferred data. In the context of any
given transfer, where the legal regime applicable in the destination country may or does conflict with the intended operation of the Standard Contractual
Clauses and/or applicable European law, the decision in Schrems II and subsequent draft guidance from the European Data Protection Board, or EDPB,
would require the parties to that transfer to implement certain supplementary technical, organizational and/or contractual measures to rely on the Standard
Contractual Clauses as a compliant transfer mechanism. However, the aforementioned draft guidance from the EDPB on such supplementary technical,
organizational and/or contractual measures appears to conclude that no combination of such measures could be sufficient to allow effective reliance on the
Standard Contractual Clauses in the context of transfers of personal data in the clear to recipients in countries where the power granted to public authorities
to  access  the  transferred  data  goes  beyond  that  which  is  ‘necessary  and  proportionate  in  a  democratic  society’  –  which  may,  following  the  CJEU’s
conclusions in Schrems II on relevant powers of United States public authorities and commentary in that draft EDPB guidance, include the United States in
certain circumstances (e.g., where Section 702 of the US Foreign Intelligence Surveillance Act applies). At present, there are few, if any, viable alternatives
to the EU-U.S. Privacy Shield and the Standard Contractual Clauses. As such, if we are unable to implement a valid solution for personal data transfers
from  Europe,  including,  for  example,  obtaining  individuals’  explicit  consent  to  transfer  their  personal  data  from  Europe  to  the  United  States  or  other
countries, we will face increased exposure to regulatory actions, substantial fines and injunctions against processing personal data from Europe. Inability to
import  personal  data  from  Europe,  including  the  EEA,  United  Kingdom  or  Switzerland,  may  also:  restrict  our  activities  in  Europe;  limit  our  ability  to
collaborate with partners as well as other service providers, contractors and other companies subject to European data protection laws; and/or require us to
increase our data processing capabilities in Europe at significant expense or otherwise cause us to change the geographical location or segregation of our
relevant  systems  and  operations  –  any  or  all  of  which  could  adversely  affect  our  financial  results. Additionally,  other  countries  outside  of  Europe  have
enacted or are considering enacting similar cross-border data transfer restrictions and laws requiring local data residency, which could increase the cost and
complexity of delivering our services and operating our business. The type of challenges we face in Europe will likely also arise in other jurisdictions that
adopt laws similar in construction to the GDPR or regulatory frameworks of equivalent complexity.

Further, Brexit has created uncertainty regarding data protection regulation in the United Kingdom. Following the United Kingdom’s withdrawal
from the European Union on January 31, 2020, pursuant to the transitional arrangements agreed to between the United Kingdom and European Union, the
GDPR  continued  to  have  effect  in  law  in  the  United  Kingdom,  and  continued  to  do  so  until  December  31,  2020  as  if  the  United  Kingdom  remained  a
Member  State  of  the  European  Union  for  such  purposes.  Following  December  31,  2020,  and  the  expiry  of  those  transitional  arrangements,  the  data
protection obligations of the GDPR continue to apply to United Kingdom-related processing of personal data in substantially unvaried form under the so-
called “UK GDPR” (i.e., the GDPR as it continues to form part of United Kingdom laws by virtue of section 3 of the European Union (Withdrawal) Act
2018,  as  amended  (including  by  the  various  Data  Protection,  Privacy  and  Electronic  Communications  (Amendments  etc)  (EU  Exit)  Regulations)).
However, going forward, there will be increasing scope for divergence in application, interpretation and enforcement of the data protection law as between
the United Kingdom and EEA. Furthermore, the relationship between the United Kingdom and the EEA in relation to certain aspects of data protection law
remains somewhat uncertain. For example, it is unclear whether transfers of personal data from the EEA to the United Kingdom will be permitted to take
place  on  the  basis  of  a  future  adequacy  decision  of  the  European  Commission,  or  whether  a  “transfer  mechanism,”  such  as  the  Standard  Contractual
Clauses, will be required. For the meantime, under the Trade and Cooperation Agreement, it has been agreed that transfers of personal data to the United
Kingdom from European Union Member States will not be treated as “restricted transfers” to a non-EEA country for a period of up to four months from
January 1, 2021, plus a potential further two months extension, or the extended adequacy assessment period. This will also apply to transfers to the United
Kingdom from EEA Member States, assuming those Member States accede to the relevant provision of the Trade and Cooperation Agreement. Although
the current maximum duration of the extended adequacy assessment period is six months it may end sooner, for example, in the event that the European
Commission adopts an adequacy decision in respect of the United Kingdom, or the United Kingdom amends the UK GDPR and/or makes certain changes
regarding data transfers under the UK GDPR / Data Protection Act 2018 without the consent

27

of the European Union (unless those amendments or decisions are made simply to keep relevant United Kingdom laws aligned with the European Union’s
data protection regime). If the European Commission does not adopt an ‘adequacy decision’ in respect of the United Kingdom prior to the expiry of the
extended adequacy assessment period, from that point onwards the United Kingdom will be an “inadequate third country” under the GDPR and transfers of
data from the EEA to the United Kingdom will require a “transfer mechanism,” such as the Standard Contractual Clauses.

Additionally, as noted above, the United Kingdom has transposed the GDPR into United Kingdom domestic law by way of the UK GDPR with
effect from January 2021, which could expose us to two parallel regimes, each of which potentially authorizes similar fines and other potentially divergent
enforcement  actions  for  certain  violations  (each  regime  separately  having  the  ability  to  fine  up  to  the  higher  of €20,000,000/£17,000,000  or  4%  of  an
undertaking’s  total  global  annual  turnover).  Also,  following  the  expiry  of  the  post-Brexit  transitional  arrangements,  the  United  Kingdom  Information
Commissioner’s Office is not able to be our “lead supervisory authority” in respect of any “cross border processing” for the purposes of the GDPR. For so
long as we are unable to, and/or do not, designate a lead supervisory authority in an EEA member state, with effect from January 1, 2021, we are not able to
benefit from the GDPR’s “one stop shop” mechanism. Amongst other things, this would mean that, in the event of a violation of the GDPR affecting data
subjects across the United Kingdom and the EEA, we could be investigated by, and ultimately fined by the United Kingdom Information Commissioner’s
Office and the supervisory authority in each and every EEA member state where data subjects have been affected by such violation.

Business disruptions could seriously harm our future revenue and financial condition and increase our costs and expenses.

Our  operations,  and  those  of  our  vendors  and  suppliers,  could  be  subject  to  earthquakes,  power  shortages,  telecommunications  failures,  water
shortages,  floods,  hurricanes,  typhoons,  fires,  extreme  weather  conditions,  medical  epidemics  and  other  natural  or  man-made  disasters  or  business
interruptions, for which we are predominantly self-insured. The occurrence of any of these business disruptions could seriously harm our operations and
financial  condition  and  increase  our  costs  and  expenses.  We  currently  rely  on  third-party  suppliers  to  produce  and  process  our  product  candidates  on
a patient-by-patient basis. Our ability to obtain clinical supplies of our product candidates could be disrupted if the operations of these suppliers are affected
by a man-made or natural disaster or other business interruption.

Risks Related to Our Dependence on Third Parties

We  are  dependent  on  intellectual  property  obtained  or  licensed  from  third  parties,  and  if  we  were  to  fail  to  comply  with  our  obligations  under  our
existing and any future intellectual property licenses with third parties, we could lose intellectual property rights that are important to our business and
we may not be able to continue developing or commercializing our product candidates, if approved.

    We are party to an exclusive intellectual property license agreement with UCL Business Ltd., or UCLB, the technology-transfer company of University
College London, or UCL, which is important to our business and under which we have acquired or licensed patent rights related to 25 patent families and
other intellectual property related to our business. We expect to enter into additional license agreements in the future. Our existing license agreement with
UCLB  imposes,  and  we  expect  that  future  license  agreements  will  impose,  various  due  diligence,  milestone  payment,  royalty,  insurance  and  other
obligations on us. Any uncured, material breach under the UCLB license agreement could result in our loss of rights to practice the patent rights (including
those that have been assigned to us) and other intellectual property licensed to us, and could compromise our development and commercialization efforts
for our current or any future product candidates.

Licensing of intellectual property is of critical importance to our business and involves complex legal, business and scientific issues. For example,
under our license agreement with UCLB, our exclusive rights under certain of the patents is subject to specified exclusions. Our right to enforce any patents
that may issue from such patent rights similarly excludes enforcing them in such excluded fields, and obligates us to coordinate our enforcement efforts
with a third-party licensee, if any, with rights in that excluded field. If a third party-licensee has the right to enforce those patents in their field, it could put
a patent that may issue from this family at risk of being invalidated or construed narrowly, in which case we would no longer have the benefit of the patents
for our own exclusivity. Disputes may arise between us and our licensors regarding intellectual property subject to a license agreement, including disputes
regarding:

•

the scope of rights granted under the license agreement and other interpretation-related issues;

• whether and the extent to which our technology and processes infringe on intellectual property of the licensor that is not subject to the licensing

agreement;

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•

•

•

•

•

our rights to third parties;

our diligence obligations with respect to the use of the licensed technology in relation to our development and commercialization of our product
candidates, and what activities satisfy those diligence obligations;

the ownership of inventions and know-how resulting from the joint creation or use of intellectual property by our licensors and us;

our right to transfer or assign the license; and

the effects of termination.

If disputes over intellectual property that we have licensed prevent or impair our ability to maintain our current licensing arrangement on acceptable
terms, we may be unable to successfully develop and commercialize the affected product candidates. See the section of this Annual Report titled “Business
Overview - Our License Agreement with UCL Business Ltd.” for a more detailed description of our license agreement with UCLB, as well as our rights
and obligations under the agreement.

We rely, and expect to continue to rely, on third parties to conduct the preclinical and clinical trials for our product candidates, and those third parties
may  not  perform  satisfactorily,  including  failing  to  meet  deadlines  for  the  completion  of  such  trials  or  failing  to  comply  with  applicable  regulatory
requirements.

We depend and will continue to depend upon independent investigators and collaborators, such as universities, medical institutions, and strategic
partners to conduct our preclinical and clinical trials. Agreements with such third parties might terminate for a variety of reasons, including a failure to
perform by the third parties. If we need to enter into alternative arrangements, our product development activities would be delayed.

Our reliance on these third parties for research and development activities will reduce our control over these activities but will not relieve us of our
responsibilities.  For  example,  we  will  remain  responsible  for  ensuring  that  each  of  our  clinical  trials  is  conducted  in  accordance  with  the  general
investigational  plan  and  protocols  for  the  trial.  Moreover,  the  FDA  requires  us  to  comply  with  regulatory  standards,  commonly  referred  to  as  good
laboratory practices, or GLP, and good clinical practices, or GCP, for conducting, recording and reporting the results of preclinical and clinical trials to
assure that data and reported results are credible and accurate and that the rights, integrity and confidentiality of trial participants are protected. Similar
regulatory  requirements  apply  outside  the  United  States,  including  the  International  Council  for  Harmonisation  of  Technical  Requirements  for  the
Registration  of  Pharmaceuticals  for  Human  Use,  or  ICH.  We  are  also  required  to  register  certain  ongoing  clinical  trials  and  post  the  results  of  certain
completed clinical trials on a government-sponsored database, ClinicalTrials.gov, within specified timeframes. Failure to do so by us or third parties can
result in FDA refusal to approve applications based on the clinical data, enforcement actions, adverse publicity and civil and criminal sanctions.

Furthermore, these third parties may also have relationships with other entities, some of which may be our competitors. If these third parties do not
successfully  carry  out  their  contractual  duties,  meet  expected  deadlines  or  conduct  our  clinical  trials  in  accordance  with  regulatory  requirements  or  our
stated protocols, we will not be able to obtain, or may be delayed in obtaining, marketing approvals for our product candidates and will not be able to, or
may be delayed in our efforts to, successfully commercialize our product candidates.

In addition, principal investigators for our clinical trials may serve as scientific advisors or consultants to us from time to time and may receive cash
or  equity  compensation  in  connection  with  such  services.  If  these  relationships  and  any  related  compensation  result  in  perceived  or  actual  conflicts  of
interest,  or  the  FDA  concludes  that  the  financial  relationship  may  have  affected  the  interpretation  of  the  trial,  the  integrity  of  the  data  generated  at  the
applicable clinical trial site may be questioned and the utility of the clinical trial itself may be jeopardized, which could result in the delay or rejection by
the FDA. Any such delay or rejection could prevent us from commercializing our clinical-stage product candidates or any future product candidates.

Cell-based  therapies  rely  on  the  availability  of  reagents,  specialized  equipment,  and  other  specialty  materials,  which  may  not  be  available  to  us  on
acceptable terms or at all. For some of these reagents, equipment, and materials, we rely or may rely on sole source vendors or a limited number of
vendors, which could impair our ability to manufacture and supply our products.

Manufacturing  our  product  candidates  will  require  many  reagents,  which  are  substances  used  in  our  manufacturing  processes  to  bring  about
chemical  or  biological  reactions,  and  other  specialty  materials  and  equipment,  some  of  which  are  manufactured  or  supplied  by  small  companies  with
limited resources and experience to support commercial biologics production. We currently depend on a limited number of vendors for access to facilities
and supply of certain materials and equipment used in the manufacture of our product candidates. For example, we currently use facilities and equipment at
the Cell and Gene Therapy Catapult, as well as third party vendors, for vector and cell manufacturing. In addition, we purchase equipment and reagents
critical

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for the manufacture of our product candidates from Miltenyi Biotec GmbH and other suppliers on a purchase order basis. Some of our suppliers may not
have the capacity to support commercial products manufactured under cGMP by biopharmaceutical firms or may otherwise be ill-equipped to support our
needs. We also do not have supply contracts with many of these suppliers, and may not be able to obtain supply contracts with them on acceptable terms or
at all. Accordingly, we may not be able to obtain key materials and equipment to support clinical or commercial manufacturing.

For some of these reagents, equipment, and materials, we rely and may in the future rely on sole source vendors or a limited number of vendors. An
inability to continue to source product from any of these suppliers, which could be due to regulatory actions or requirements affecting the supplier, adverse
financial or other strategic developments experienced by a supplier, labor disputes or shortages, unexpected demands, or quality issues, could adversely
affect our ability to satisfy demand for our product candidates, which could adversely and materially affect our product sales and operating results or our
ability to conduct clinical trials, either of which could significantly harm our business.

As we continue to develop and scale our manufacturing process, we may need to obtain rights to and supplies of certain materials and equipment to
be used as part of that process. We may not be able to obtain rights to such materials on commercially reasonable terms, or at all, and if we are unable to
alter our process in a commercially viable manner to avoid the use of such materials or find a suitable substitute, it would have a material adverse effect on
our business.

Risks Related to Regulatory Approval of Our Product Candidates and Other Legal Compliance Matters

Even if we complete the necessary preclinical studies and clinical trials, the regulatory approval process is expensive, time-consuming and uncertain
and may prevent us from obtaining approvals for the commercialization of some or all of our product candidates. As a result, we cannot predict when
or if, and in which territories, we will obtain marketing approval to commercialize a product candidate.

Our  product  candidates  and  the  activities  associated  with  their  development  and  commercialization,  including  their  design,  research,  testing,
manufacture,  safety,  efficacy,  quality  control,  recordkeeping,  labeling,  packaging,  storage,  approval,  advertising,  promotion,  sale,  distribution,  import,
export, and reporting of safety and other post-market information, are subject to comprehensive regulation by the FDA, the EMA and other comparable
regulatory authorities in other jurisdictions. Failure to obtain marketing approval for a product candidate will prevent us from commercializing the product
candidate. We have not received approval to market any of our product candidates from regulatory authorities in any jurisdiction. We have only limited
experience in filing and supporting the applications necessary to gain marketing approvals and may rely on third-party contract research organizations, or
CROs,  to  assist  us  in  this  process.  Securing  marketing  approval  requires  the  submission  of  extensive  preclinical  and  clinical  data  and  supporting
information to regulatory authorities for each therapeutic indication to establish the product candidate’s safety and efficacy. Securing marketing approval
also  requires  the  submission  of  information  about  the  product  manufacturing  process  to,  and  inspection  of  manufacturing  facilities  by,  the  regulatory
authorities. Our product candidates may not be effective, may be only moderately effective or may prove to have undesirable or unintended side effects,
toxicities or other characteristics that may preclude our obtaining marketing approval or prevent or limit commercial use. If any of our product candidates
receives marketing approval, the accompanying label may limit its approved use, which could limit sales of the product.

The process of obtaining marketing approvals, both in the United States and abroad, is expensive and may take many years, if approval is obtained
at all, and can vary substantially based upon a variety of factors, including the type, complexity and novelty of the product candidates involved. Securing
marketing  approval  requires  the  submission  of  extensive  preclinical  and  clinical  data  and  supporting  information  to  regulatory  authorities  for  each
therapeutic  indication  to  establish  the  product  candidate’s  safety  and  efficacy.  Securing  marketing  approval  also  requires  the  submission  of  information
about  the  product  manufacturing  process  to,  and  inspection  of  manufacturing  facilities  by,  the  regulatory  authorities.  The  FDA,  the  EMA  or  other
regulatory authorities may determine that our product candidates are not safe and effective, only moderately effective or have undesirable or unintended
side effects, toxicities or other characteristics that preclude our obtaining marketing approval or prevent or limit commercial use.

In addition, we are developing a proprietary diagnostic test for use with our AUTO4 and AUTO5 product candidates. This test will require separate
regulatory approval in addition to the regulatory approval of AUTO4 and AUTO5, respectively. Failure to obtain marketing approval for the diagnostic test
could prevent us from commercializing either AUTO4 or AUTO5 unless another similar diagnostic test for distinguishing TRBC1-positive and TRBC2-
positive T-cell lymphomas is commercially available.

In  addition,  changes  in  marketing  approval  policies  during  the  development  period,  changes  in  or  the  enactment  of  additional  statutes  or
regulations,  or  changes  in  regulatory  review  for  each  submitted  product  application,  may  cause  delays  in  the  approval  or  rejection  of  an  application.
Regulatory  authorities  have  substantial  discretion  in  the  approval  process  and  may  refuse  to  accept  any  application  or  may  decide  that  our  data  is
insufficient for approval and require additional preclinical, clinical or other studies. In addition, varying interpretations of the data obtained from preclinical
and clinical testing could delay, limit or prevent

30

marketing  approval  of  a  product  candidate.  Any  marketing  approval  we  ultimately  obtain  may  be  limited  or  subject  to  restrictions  or  post-approval
commitments that render the approved product not commercially viable.

If we experience delays in obtaining approval or if we fail to obtain approval of our product candidates, the commercial prospects for our product

candidates may be harmed and our ability to generate revenues will be impaired.

In  order  to  market  and  sell  our  products  in  the  European  Union  and  any  other  jurisdictions,  we  must  obtain  separate  marketing  approvals  and
comply with numerous and varying regulatory requirements. The approval procedure varies among countries and can involve additional testing. The time
required  to  obtain  approval  may  differ  substantially  from  that  required  to  obtain  approval  from  the  FDA.  The  regulatory  approval  process  outside  the
United States generally includes all of the risks associated with obtaining approval from the FDA. In addition, in many countries outside the United States,
it is required that the product be approved for reimbursement before the product can be approved for sale in that country. We may not obtain approvals from
regulatory authorities outside the United States on a timely basis, if at all. Approval by the FDA does not ensure approval by regulatory authorities in other
countries or jurisdictions, and approval by one regulatory authority outside the United States does not ensure approval by regulatory authorities in other
countries or jurisdictions or by the FDA. However, failure to obtain approval in one jurisdiction may impact our ability to obtain approval elsewhere. We
may not be able to file for marketing approvals and may not receive necessary approvals to commercialize our products in any market.

Obtaining and maintaining regulatory approval of our product candidates in one jurisdiction does not mean that we will be successful in obtaining
regulatory approval of our product candidates in other jurisdictions.

Obtaining and maintaining regulatory approval of our product candidates in one jurisdiction does not guarantee that we will be able to obtain or
maintain regulatory approval in any other jurisdiction, but a failure or delay in obtaining regulatory approval in one jurisdiction may have a negative effect
on  the  regulatory  approval  process  in  others.  For  example,  even  if  the  FDA  grants  marketing  approval  of  a  product  candidate,  comparable  regulatory
authorities  in  other  jurisdictions  must  also  approve  the  manufacturing,  marketing  and  promotion  of  the  product  candidate  in  those  countries.  Approval
procedures vary among jurisdictions and can involve requirements and administrative review periods different from those in the United States, including
additional  preclinical  studies  or  clinical  trials  as  clinical  studies  conducted  in  one  jurisdiction  may  not  be  accepted  by  regulatory  authorities  in  other
jurisdictions. In many jurisdictions outside the United States, a product candidate must be approved for reimbursement before it can be approved for sale in
that jurisdiction. In some cases, the price that we intend to charge for our products is also subject to approval.

Obtaining foreign regulatory approvals and compliance with foreign regulatory requirements could result in significant delays, difficulties and costs
for us and could delay or prevent the introduction of our products in certain countries. If we fail to comply with the regulatory requirements in international
markets  and/or  to  receive  applicable  marketing  approvals,  our  target  market  will  be  reduced  and  our  ability  to  realize  the  full  market  potential  of  our
product candidates will be harmed.

The  withdrawal  of  the  United  Kingdom  from  the  European  Union,  commonly  referred  to  as  “Brexit,”  may  adversely  impact  our  ability  to  obtain
regulatory  approvals  of  our  product  candidates  in  the  European  Union,  result  in  restrictions  or  imposition  of  taxes  and  duties  for  importing  our
product candidates into the European Union, and may require us to incur additional expenses in order to develop, manufacture and commercialize our
product candidates in the European Union.

Following the result of a referendum in 2016, the United Kingdom left the European Union on January 31, 2020, commonly referred to as Brexit.
Pursuant  to  the  formal  withdrawal  arrangements  agreed  between  the  United  Kingdom  and  the  European  Union,  the  United  Kingdom  was  subject  to  a
transition period until December 31, 2020, or the Transition Period, during which European Union rules continued to apply. The Trade and Cooperation
Agreement, which outlines the future trading relationship between the United Kingdom and the European Union, was agreed in December 2020.

Since a significant proportion of the regulatory framework in the United Kingdom applicable to our business and our product candidates is derived
from  EU  directives  and  regulations,  Brexit  has  had,  and  may  continue  to  have,  a  material  impact  upon  the  regulatory  regime  with  respect  to  the
development,  manufacture,  importation,  approval  and  commercialization  of  our  product  candidates  in  the  United  Kingdom  or  the  European  Union.  For
example, Great Britain is no longer covered by the centralized procedures for obtaining European Union-wide marketing authorization from the EMA and a
separate  marketing  authorization  will  be  required  to  market  our  product  candidates  in  Great  Britain.  It  is  currently  unclear  whether  the  Medicines  and
Healthcare  products  Regulatory  Agency,  or  MHRA,  in  the  United  Kingdom  is  sufficiently  prepared  to  handle  the  increased  volume  of  marketing
authorization applications that it is likely to receive. Any delay in obtaining, or an inability to obtain, any marketing approvals, would delay or prevent us
from commercializing our product candidates in the United Kingdom.

While the Trade and Cooperation Agreement provides for the tariff-free trade of medicinal products between the United Kingdom and the European

Union there may be additional non-tariff costs to such trade which did not exist prior to the end of the

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Transition Period. Further, should the United Kingdom diverge from the European Union from a regulatory perspective in relation to medicinal products,
tariffs could be put into place in the future. We could therefore, both now and in the future, face significant additional expenses (when compared to the
position prior to the end of the Transition Period) to operate our business, which could significantly and materially harm or delay our ability to generate
revenues or achieve profitability of our business. Any further changes in international trade, tariff and import/export regulations as a result of Brexit or
otherwise may impose unexpected duty costs or other non-tariff barriers on us. These developments, or the perception that any of them could occur, may
significantly  reduce  global  trade  and,  in  particular,  trade  between  the  impacted  nations  and  the  United  Kingdom.  It  is  also  possible  that  Brexit  may
negatively affect our ability to attract and retain employees, particularly those from the European Union.

In  the  short  term  there  is  a  risk  of  disrupted  import  and  export  processes  due  to  a  lack  of  administrative  processing  capacity  by  the  respective

United Kingdom and European Union customs agencies that may delay time-sensitive shipments and may negatively impact our product supply chain.

Even if we obtain marketing approvals for our product candidates, the terms of approvals and ongoing regulation of our products may limit how we
manufacture and market our products and compliance with such requirements may involve substantial resources, which could materially impair our
ability to generate revenue.

Even if marketing approval of a product candidate is granted, an approved product and its manufacturer and marketer are subject to ongoing review
and  extensive  regulatory  requirements  for  manufacturing  processes,  labeling,  packaging,  distribution,  adverse  event  reporting,  storage,  advertising,
promotion, sampling, and recordkeeping, including the potential requirements to implement a risk evaluation and mitigation strategy, or REMS, program or
to  conduct  costly  post-marketing  studies  or  clinical  trials  and  surveillance  to  monitor  the  safety  or  efficacy  of  the  product.  We  must  also  comply  with
requirements  concerning  advertising  and  promotion  for  any  of  our  product  candidates  for  which  we  obtain  marketing  approval.  Promotional
communications with respect to prescription drugs are subject to a variety of legal and regulatory restrictions and must be consistent with the information in
the product’s approved labeling. Thus, we will not be able to promote any products we develop for indications or uses for which they are not approved. In
addition,  manufacturers  of  approved  products  and  those  manufacturers’  facilities  are  required  to  comply  with  extensive  regulatory  requirements  of  the
FDA,  the  EMA  and  other  regulatory  authorities,  including  ensuring  that  quality  control  and  manufacturing  procedures  conform  to  cGMP  and  other
comparable  regulations  and  standards,  which  include  requirements  relating  to  quality  control  and  quality  assurance  as  well  as  the  corresponding
maintenance of records and documentation and reporting requirements. We or our suppliers could be subject to periodic unannounced inspections by the
FDA, the EMA, or other regulatory authorities to monitor and ensure compliance with cGMP.

Accordingly, assuming we receive marketing approval for one or more of our product candidates, we and suppliers will continue to expend time,
money and effort in all areas of regulatory compliance, including manufacturing, production, product surveillance and quality control. If we are not able to
comply with post-approval regulatory requirements, we could have the marketing approvals for our products withdrawn by regulatory authorities and our
ability to market any future products could be limited, which could adversely affect our ability to achieve or sustain profitability.

Thus, the cost of compliance with post-approval regulations may have a negative effect on our operating results and financial condition.

Any product candidate for which we obtain marketing approval could be subject to post-marketing restrictions or recall or withdrawal from the market,
and  we  may  be  subject  to  penalties  if  we  fail  to  comply  with  regulatory  requirements  or  if  we  experience  unanticipated  problems  with  our  product
candidates, when and if any of them are approved.

The FDA and other federal and state agencies, including the U.S. Department of Justice, or DOJ, closely regulate compliance with all requirements
governing prescription drug products, including requirements pertaining to marketing and promotion of products in accordance with the provisions of the
approved  labeling  and  manufacturing  of  products  in  accordance  with  cGMP  requirements.  The  FDA  and  DOJ  impose  stringent  restrictions  on
manufacturers’ communications regarding off-label use and if we do not market our products for their approved indications, or if other of our marketing
claims are deemed false or misleading, we may be subject to enforcement action. Physicians, on the other hand, may prescribe products for off-label uses.
The  FDA  and  other  regulatory  agencies  do  not  regulate  a  physician’s  choice  of  drug  treatment  made  in  the  physician’s  independent  medical  judgment.
However,  companies  may  only  share  truthful  and  not  misleading  information  that  is  otherwise  consistent  with  a  product’s  FDA  approved  labeling.
Violations of such requirements may lead to investigations alleging violations of the Food, Drug and Cosmetic Act, or FDCA, and other statutes, including
the U.S. federal False Claims Act and other federal and state healthcare fraud and abuse laws as well as state consumer protection laws.

Our  failure  to  comply  with  all  regulatory  requirements,  and  later  discovery  of  previously  unknown  adverse  events  or  other  problems  with  our

products, manufacturers or manufacturing processes, may yield various results, including:

32

•

•

•

•

•

litigation involving patients taking our products;

restrictions on such products, manufacturers or manufacturing processes;

restrictions on the labeling or marketing of a product;

restrictions on product distribution or use;

requirements to conduct post-marketing studies or clinical trials;

• warning or untitled letters;

• withdrawal of the products from the market;

•

•

•

•

•

•

•

•

•

•

refusal to approve pending applications or supplements to approved applications that we submit;

recall of products;

fines, restitution or disgorgement of profits or revenues;

suspension or withdrawal of marketing approvals;

suspension of any ongoing clinical trials;

damage to relationships with any potential collaborators;

unfavorable press coverage and damage to our reputation;

refusal to permit the import or export of our products;

product seizure; or

injunctions or the imposition of civil or criminal penalties.

Non-compliance  by  us  or  any  future  collaborator  with  regulatory  requirements  regarding  safety  monitoring  or  pharmacovigilance,  and  with
requirements  related  to  the  development  of  products  for  the  pediatric  population,  can  also  result  in  significant  financial  penalties.  Similarly,  failure  to
comply with regulatory requirements regarding the protection of personal information can also lead to significant penalties and sanctions.

Non-compliance  with  EU  requirements  regarding  safety  monitoring  or  pharmacovigilance,  and  with  requirements  related  to  the  development  of
products for the pediatric population, also can result in significant financial penalties. Similarly, failure to comply with the European Union’s requirements
regarding the protection of personal information can also lead to significant penalties and sanctions.

If any of these events occurs, our ability to sell such product may be impaired, and we may incur substantial additional expense to comply with

regulatory requirements, which could adversely affect our business, financial condition and results of operations.

33

 
 
Our  employees,  independent  contractors,  principal  investigators,  consultants,  commercial  partners  and  vendors  may  engage  in  misconduct  or  other
improper activities, including non-compliance with regulatory standards and requirements.

We are exposed to the risk of employee fraud or other misconduct or failure to comply with applicable regulatory requirements. Misconduct by
employees and independent contractors, such as principal investigators, consultants, commercial partners, and vendors, could include failures to comply
with regulations of the FDA, the EMA and other comparable regulatory authorities, to provide accurate information to such regulators, to comply with
manufacturing  standards  we  have  established,  to  comply  with  healthcare  fraud  and  abuse  laws,  to  report  financial  information  or  data  accurately  or  to
disclose  unauthorized  activities  to  us.  In  particular,  sales,  marketing  and  other  business  arrangements  in  the  healthcare  industry  are  subject  to  extensive
laws and regulations intended to prevent fraud, misconduct, kickbacks, self-dealing and other abusive practices. These laws and regulations may restrict or
prohibit  a  wide  range  of  business  activities,  including,  but  not  limited  to,  research,  manufacturing,  distribution,  pricing,  discounting,  marketing  and
promotion, sales commission, customer incentive programs and other business arrangements. Employee and independent contractor misconduct could also
involve the improper use of individually identifiable information, including, without limitation, information obtained in the course of clinical trials, which
could result in regulatory sanctions and serious harm to our reputation.

In  addition,  federal  procurement  laws  impose  substantial  penalties  for  misconduct  in  connection  with  government  contracts  and  require  certain

contractors to maintain a code of business ethics and conduct.

It is not always possible to identify and deter employee and independent contractor misconduct, and any precautions we take to detect and prevent
improper activities may not be effective in controlling unknown or unmanaged risks or losses or in protecting us from governmental investigations or other
actions or lawsuits stemming from a failure to be in compliance with such laws. If any such actions are instituted against us, those actions could have a
significant  impact  on  our  business,  including  the  imposition  of  civil,  criminal  and  administrative  penalties,  damages,  monetary  fines,  disgorgement  of
profits,  possible  exclusion  from  participation  in  Medicare,  Medicaid  and  other  federal  healthcare  programs,  National  Health  Service  in  the  United
Kingdom, or other government supported healthcare in other jurisdictions, contractual damages, reputational harm, diminished profits and future earnings,
additional  reporting  or  oversight  obligations  if  we  become  subject  to  a  corporate  integrity  agreement  or  other  agreement  to  resolve  allegations  of  non-
compliance with the law and curtailment or restructuring of our operations, any of which could adversely affect our ability to operate.

Our business operations and current and future relationships with healthcare professionals, principal investigators, consultants, customers and third-
party  payors  in  the  United  States  and  elsewhere  may  be  subject,  directly  or  indirectly,  to  applicable  anti-kickback,  fraud  and  abuse,  false  claims,
physician  payment  transparency,  health  information  privacy  and  security  and  other  healthcare  laws  and  regulations,  which  could  expose  us  to
substantial penalties.

Healthcare  providers,  physicians  and  third-party  payors  in  the  United  States  and  elsewhere  will  play  a  primary  role  in  the  recommendation  and
prescription  of  any  product  candidates  for  which  we  obtain  marketing  approval.  Our  current  and  future  arrangements  with  healthcare  professionals,
principal  investigators,  consultants,  customers  and  third-party  payors  may  expose  us  to  broadly  applicable  fraud  and  abuse  and  other  healthcare  laws,
including, without limitation, the U.S. federal Anti-Kickback Statute and the U.S. federal False Claims Act, that may constrain the business or financial
arrangements and relationships through which we sell, market and distribute any product candidates for which we obtain marketing approval. In addition,
we may be subject to physician payment transparency laws and patient privacy and security regulation by the U.S. federal government and by the states and
foreign  jurisdictions  in  which  we  conduct  our  business.  The  applicable  federal,  state  and  foreign  healthcare  laws  that  may  affect  our  ability  to  operate
include the following:

•

the U.S. federal Anti-Kickback Statute, which prohibits, among other things, persons or entities from knowingly and willfully soliciting, offering,
receiving  or  providing  remuneration,  directly  or  indirectly,  in  cash  or  in  kind,  to  induce  or  reward,  or  in  return  for,  either  the  referral  of  an
individual for, or the purchase, lease, order or recommendation of, any good, facility, item or service, for which payment may be made, in whole
or in part, under federal and state healthcare programs such as Medicare and Medicaid. The term ‘‘remuneration’’ has been broadly interpreted to
include anything of value. This statute has been interpreted to apply to arrangements between pharmaceutical manufacturers on the one hand and
prescribers,  purchasers  and  formulary  managers  on  the  other  hand.  Although  there  are  a  number  of  statutory  exceptions  and  regulatory  safe
harbors protecting certain common activities from prosecution or other regulatory sanctions, the exceptions and safe harbors are drawn narrowly,
and practices that involve remuneration that are alleged to be intended to induce prescribing, purchases or recommendations may be subject to
scrutiny if they do not qualify for an exception or safe harbor. Failure to meet all of the requirements of a particular applicable statutory exception
or  regulatory  safe  harbor  does  not  make  the  conduct  per  se  illegal  under  the  U.S.  federal  Anti-Kickback  Statute.  Instead,  the  legality  of  the
arrangement  will  be  evaluated  on  a  case-by-case  basis  based  on  a  cumulative  review  of  all  its  facts  and  circumstances.  Several  courts  have
interpreted the statute’s intent requirement to mean that if any one purpose of an arrangement

34

involving remuneration is to induce referrals of federal healthcare covered business, the U.S. federal Anti-Kickback Statute has been violated;

• U.S. federal civil and criminal false claims laws, including the U.S. federal False Claims Act, which can be enforced though civil whistleblower or
qui  tam  actions,  and  civil  monetary  penalty  laws,  which  prohibit,  among  other  things,  individuals  or  entities  from  knowingly  presenting,  or
causing to be presented, to the federal government, including the Medicare and Medicaid programs, claims for payment that are false or fraudulent
or  making  a  false  statement  to  avoid,  decrease  or  conceal  an  obligation  to  pay  money  to  the  federal  government.  Pharmaceutical  and  other
healthcare  companies  have  been  prosecuted  under  these  laws  for,  among  other  things,  allegedly  inflating  drug  prices  they  report  to  pricing
services, which in turn were used by the government to set Medicare and Medicaid reimbursement rates, and for allegedly providing free product
to  customers  with  the  expectation  that  the  customers  would  bill  federal  programs  for  the  product.  In  addition,  certain  marketing  practices,
including off-label promotion, may also violate false claims laws. Further, pharmaceutical manufacturers can be held liable under the U.S. federal
False Claims Act even when they do not submit claims directly to government payors if they are deemed to “cause” the submission of false or
fraudulent claims;

•

the U.S. federal Health Insurance Portability and Accountability Act of 1996, or HIPAA, which created new federal criminal statutes that prohibit
knowingly and willfully executing, or attempting to execute, a scheme to defraud any healthcare benefit program or obtain, by means of false or
fraudulent  pretenses,  representations  or  promises,  any  of  the  money  or  property  owned  by,  or  under  the  custody  or  control  of,  any  healthcare
benefit program, regardless of whether the payor is public or private, knowingly and willfully embezzling or stealing from a healthcare benefit
program, willfully obstructing a criminal investigation of a healthcare offense and knowingly and willfully falsifying, concealing or covering up
by  any  trick  or  device  a  material  fact  or  making  any  materially  false  statements  in  connection  with  the  delivery  of,  or  payment  for,  healthcare
benefits, items or services relating to healthcare matters;

• HIPAA,  as  amended  by  the  Health  Information  Technology  for  Economic  and  Clinical  Health  Act  of  2009,  or  HITECH,  and  their  respective
implementing  regulations,  which  impose  obligations  on  “covered  entities,”  including  certain  healthcare  providers,  health  plans,  and  healthcare
clearinghouses,  as  well  as  their  respective  “business  associates”  that  create,  receive,  maintain  or  transmit  individually  identifiable  health
information  for  or  on  behalf  of  a  covered  entity,  and  their  covered  subcontractors,  with  respect  to  safeguarding  the  privacy,  security  and
transmission  of  individually  identifiable  health  information.  Additionally,  HITECH  also  created  four  new  tiers  of  civil  monetary  penalties,
amended HIPAA to make civil and criminal penalties directly applicable to business associates, and gave state attorneys general new authority to
file civil actions for damages or injunctions in U.S. federal courts to enforce HIPAA and seek attorneys’ fees and costs associated with pursuing
federal civil actions;

•

•

•

the FDCA, which prohibits, among other things, the adulteration or misbranding of drugs, biologics and medical devices;

the U.S. federal Physician Payments Sunshine Act, created under Section 6002 of Patient Protection and Affordable Care Act, as amended by the
Health Care and Education Reconciliation Act, or collectively, the ACA, and its implementing regulations, created annual reporting requirements
for  certain  manufacturers  of  drugs,  devices,  biologicals  and  medical  supplies  for  which  payment  is  available  under  Medicare,  Medicaid  or  the
Children’s Health Insurance Program (with certain exceptions), to annually report to the Centers for Medicare and Medicaid Services, or CMS,
information  related  to  certain  payments  and  “transfers  of  value”  provided  to  physicians  (defined  to  include  doctors,  dentists,  optometrists,
podiatrists and chiropractors) and teaching hospitals, as well as ownership and investment interests held by physicians and their immediate family
members. Beginning in 2022, applicable manufacturers also will be required to report information regarding payments and “transfers of value”
provided during the previous year to physician assistants, nurse practitioners, clinical nurse specialists, certified nurse anesthetists, anesthesiologist
assistants, and certified nurse-midwives;

analogous state laws and regulations and foreign laws, such as state anti-kickback and false claims laws, which may apply to sales or marketing
arrangements and claims involving healthcare items or services reimbursed by non-governmental third-party payors, including private insurers;
state and foreign laws that require pharmaceutical companies to comply with the pharmaceutical industry’s voluntary compliance guidelines and
the  relevant  compliance  guidance  promulgated  by  the  federal  government  or  to  adopt  compliance  programs  as  prescribed  by  state  laws  and
regulations, or that otherwise restrict payments that may be made to healthcare providers; state and foreign laws that require drug manufacturers to
report information related to payments and other transfers of value to physicians and other healthcare providers or marketing expenditures; state
and local laws that require the registration of pharmaceutical sales representatives; and state and foreign laws governing the privacy and security
of health information in certain circumstances, many of which differ from each other in significant ways and often are not preempted by HIPAA,
thus complicating compliance efforts; and

•

similar healthcare laws and regulations in the European Union and other jurisdictions, including reporting requirements detailing interactions with
and payments to healthcare providers and laws governing the privacy and security of certain

35

protected information, such as GDPR, which imposes obligations and restrictions on the collection and use of personal data relating to individuals
located in the European Union (including health data).

Further,  the  ACA,  among  other  things,  amended  the  intent  requirement  of  the  U.S.  federal  Anti-Kickback  Statute  and  certain  criminal  statutes
governing healthcare fraud. A  person  or  entity  no  longer  needs  to  have  actual  knowledge  of  the  statute  or  specific  intent  to  violate  it  in  order  to  have
committed a violation. In addition, the ACA provided that the government may assert that a claim including items or services resulting from a violation of
the U.S. federal Anti-Kickback Statute constitutes a false or fraudulent claim for purposes of the U.S. federal False Claims Act.

Because  of  the  breadth  of  these  laws  and  the  narrowness  of  their  exceptions  and  safe  harbors,  it  is  possible  that  our  business  activities  can  be
subject  to  challenge  under  one  or  more  of  such  laws.  The  scope  and  enforcement  of  each  of  these  laws  is  uncertain  and  subject  to  rapid  change  in  the
current environment of healthcare reform. Federal and state enforcement bodies have continued their scrutiny of interactions between healthcare companies
and healthcare providers, which has led to a number of significant investigations, prosecutions, convictions and settlements in the healthcare industry.

Efforts to ensure that our internal operations and future business arrangements with third parties will comply with applicable healthcare laws and
regulations will involve substantial costs. If our operations are found to be in violation of any of these laws or any other governmental regulations that may
apply  to  us,  we  may  be  subject  to  significant  civil,  criminal  and  administrative  penalties,  including,  without  limitation,  damages,  monetary  fines,
imprisonment,  disgorgement  of  profits,  possible  exclusion  from  participation  in  Medicare,  Medicaid  and  other  federal  healthcare  programs,  contractual
damages,  reputational  harm,  diminished  profits  and  future  earnings,  additional  reporting  or  oversight  obligations  if  we  become  subject  to  a  corporate
integrity agreement or other agreement to resolve allegations of non-compliance with the law and curtailment or restructuring of our operations, any of
which could adversely affect our ability to operate our business and pursue our strategy. If any of the physicians or other healthcare providers or entities
with  whom  we  expect  to  do  business,  including  future  collaborators,  are  found  not  to  be  in  compliance  with  applicable  laws,  they  may  be  subject  to
criminal,  civil  or  administrative  sanctions,  including  exclusions  from  participation  in  government  healthcare  programs,  which  could  also  affect  our
business.

Our product candidates are subject to government price controls in certain jurisdictions that may affect our revenue.

There has been heightened governmental scrutiny in the United Kingdom, United States, European Union and other jurisdictions of pharmaceutical
pricing  practices  in  light  of  the  rising  cost  of  prescription  drugs  and  biologics.  In  the  United  States,  such  scrutiny  has  resulted  in  several  recent
Congressional inquiries and proposed and enacted federal and state legislation designed to, among other things, bring more transparency to product pricing,
review the relationship between pricing and manufacturer patient programs, and reform government program reimbursement methodologies for products.
At  the  federal  level,  the  Trump  administration  used  several  means  to  propose  or  implement  drug  pricing  reform,  including  through  federal  budget
proposals, executive orders and policy initiatives. For example, on July 24, 2020 and September 13, 2020, the Trump administration announced several
executive orders related to prescription drug pricing that seek to implement several of the administration’s proposals. As a result, the FDA released a final
rule on September 24, 2020, effective November 30, 2020, providing guidance for states to build and submit importation plans for drugs from Canada.
Further, on November 20, 2020, the Department of Health and Human Services finalized a regulation removing safe harbor protection for price reductions
from  pharmaceutical  manufacturers  to  plan  sponsors  under  Medicare  Part  D,  either  directly  or  through  pharmacy  benefit  managers,  unless  the  price
reduction is required by law. The implementation of the rule has been delayed by the Biden administration from January 1, 2022 to January 1, 2023 in
response to ongoing litigation. The rule also creates a new safe harbor for price reductions reflected at the point-of-sale, as well as a new safe harbor for
certain  fixed  fee  arrangements  between  pharmacy  benefit  managers  and  manufacturers,  the  implementation  of  which  have  also  been  delayed  pending
review  by  the  Biden  administration  until  March  22,  2021.  On  November  20,  2020,  CMS  issued  an  interim  final  rule  implementing  the  Trump
administration’s Most Favored Nation executive order, which would tie Medicare Part B payments for certain physician-administered drugs to the lowest
price  paid  in  other  economically  advanced  countries,  effective  January  1,  2021.  On  December  28,  2020,  the  U.S.  District  Court  in  Northern  California
issued a nationwide preliminary injunction against implementation of the interim final rule. It is unclear whether the Biden administration will work to
reverse these measures or pursue similar policy initiatives.

At the state level, legislatures have increasingly enacted legislation and implemented regulations designed to control pharmaceutical and biological
product pricing, including price or patient reimbursement constraints, discounts, restrictions on certain product access and marketing cost disclosure and
transparency measures, and, in some cases, designed to encourage importation from other countries and bulk purchasing.

Outside of the United States, particularly in the European Union, the pricing of prescription pharmaceuticals is subject to governmental control. In

these countries, pricing negotiations with governmental authorities can take considerable time after the

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receipt of marketing approval for a product. To obtain coverage and reimbursement or pricing approval in some countries, we may be required to conduct a
clinical trial that compares the cost-effectiveness of our product candidate to other available therapies. If reimbursement of our products is unavailable or
limited in scope or amount, or if pricing is set at unsatisfactory levels, our business could be harmed.

Recently enacted and future legislation in the United States and other countries may affect the prices we may obtain for our product candidates and
increase the difficulty and cost for us to commercialize our product candidates.

In  the  United  States  and  many  other  countries,  rising  healthcare  costs  have  been  a  concern  for  governments,  patients  and  the  health  insurance
sector,  which  has  resulted  in  a  number  of  changes  to  laws  and  regulations,  and  may  result  in  further  legislative  and  regulatory  action  regarding  the
healthcare and health insurance systems that could affect our ability to profitably sell any product candidates for which we obtain marketing approval. For a
detailed discussion of healthcare reform initiatives of importance to the pharmaceutical industry, see the section titled “Business—Government Regulation
and Product Approval—Healthcare Reform Efforts.”

For example, the ACA was enacted in the United States in March 2010 with the stated goals of containing healthcare costs, improving quality and
expanding access to healthcare, and includes measures to change healthcare delivery, increase the number of individuals with insurance, ensure access to
certain basic healthcare services, and contain the rising cost of care. There have been executive, judicial and Congressional challenges to certain aspects of
the ACA. While Congress has not passed repeal legislation, several bills affecting the implementation of certain taxes under the ACA have been signed into
law. H.R. 1: An Act to provide for reconciliation pursuant to titles II and V of the concurrent resolution on the budget for fiscal year 2018, or the Tax Cuts
and  Jobs  Act  of  2017,  includes  a  provision  repealing,  effective  January  1,  2019,  the  tax-based  shared  responsibility  payment  imposed  by  the  ACA  on
certain  individuals  who  fail  to  maintain  qualifying  health  coverage  for  all  or  part  of  a  year  that  is  commonly  referred  to  as  the  “individual  mandate”.
Additionally,  the  2020  federal  spending  package  permanently  eliminated,  effective  January  1,  2020,  the  ACA-mandated  “Cadillac”  tax  on  high-cost
employer-sponsored health coverage and medical device tax and, effective January 1, 2021, also eliminated the health insurer tax. Further, the Bipartisan
Budget Act of 2018, among other things, amended the ACA, effective January 1, 2019, to increase from 50% to 70% the point-of-sale discount that is owed
by pharmaceutical manufacturers who participate in Medicare Part D and to close the coverage gap in most Medicare drug plans, commonly referred to as
the “donut hole”. On December 14, 2018, a Texas U.S. District Court Judge ruled that the ACA is unconstitutional in its entirety because the “individual
mandate” was repealed by Congress as part of the Tax Act. Additionally, on December 18, 2019, the U.S. Court of Appeals for the 5  Circuit upheld the
District Court ruling that the individual mandate was unconstitutional and remanded the case back to the District Court to determine whether the remaining
provisions of the ACA are invalid as well. The U.S. Supreme Court is currently reviewing the case, although it is unknown when a decision will be made.
Further, although the U.S. Supreme Court has not yet ruled on the constitutionality of the ACA, on January 28, 2021, President Biden issued an executive
order to initiate a special enrollment period from February 15, 2021 through May 15, 2021 for purposes of obtaining health insurance coverage through the
ACA  marketplace.  The  executive  order  also  instructs  certain  governmental  agencies  to  review  and  reconsider  their  existing  policies  and  rules  that  limit
access  to  healthcare,  including  among  others,  reexamining  Medicaid  demonstration  projects  and  waiver  programs  that  include  work  requirements,  and
policies that create unnecessary barriers to obtaining access to health insurance coverage through Medicaid or the ACA. It is unclear how the Supreme
Court ruling, other such litigation, and the healthcare reform measures of the Biden administration will impact the ACA and our business.

th

In  addition,  other  federal  health  reform  measures  have  been  proposed  and  adopted  in  the  United  States.  For  example,  as  a  result  of  the  Budget
Control  Act  of  2011,  providers  are  subject  to  Medicare  payment  reductions  of  2%  per  fiscal  year  through  2030  with  the  exception  of  a  temporary
suspension from May 1, 2020 through March 31, 2021 unless additional Congressional action is taken. Further, the American Taxpayer Relief Act of 2012
reduced Medicare payments to several providers and increased the statute of limitations period for the government to recover overpayments from providers
from three to five years. The Medicare Access and CHIP Reauthorization Act of 2015 also introduced a quality payment program, or Quality Payment
Program, under which certain individual Medicare providers will be subject to certain incentives or penalties based on new program quality standards. The
Quality Payment Program provides clinicians with two ways to participate, including through the Advanced Alternative Payment Models, or APMs, and
the Merit-based Incentive Payment System, or MIPS. In November 2019, CMS issued a final rule finalizing the changes to the Quality Payment Program.
At this time, it is unclear how the introduction of the Quality Payment Program will impact overall physician reimbursement under the Medicare program.
Any reduction in reimbursement from Medicare or other government programs may result in a similar reduction in payments from private payors. It is also
possible that additional governmental action is taken in response to the COVID-19 pandemic.

The  combination  of  healthcare  cost  containment  measures,  increased  health  insurance  costs,  reduction  of  the  number  of  people  with  health

insurance coverage, as well as future legislation and regulations focused on reducing healthcare costs by

37

reducing  the  cost  of  or  reimbursement  and  access  to  pharmaceutical  products,  may  limit  or  delay  our  ability  to  generate  revenue,  attain  profitability,  or
commercialize our products.

We are subject to the U.K. Bribery Act 2010, or the Bribery Act, the U.S. Foreign Corrupt Practices Act of 1977, as amended, or the FCPA, and other
anti-corruption  laws,  as  well  as  export  control  laws,  import  and  customs  laws,  trade  and  economic  sanctions  laws  and  other  laws  governing  our
operations.

Our operations are subject to anti-corruption laws, including the Bribery Act, the FCPA, the U.S. domestic bribery statute contained in 18 U.S.C.
§201, the U.S. Travel Act, and other anti-corruption laws that apply in countries where we do business. The Bribery Act, the FCPA and these other laws
generally  prohibit  us  and  our  employees  and  intermediaries  from  authorizing,  promising,  offering,  or  providing,  directly  or  indirectly,  improper  or
prohibited payments, or anything else of value, to government officials or other persons to obtain or retain business or gain some other business advantage.

Under the Bribery Act, we may also be liable for failing to prevent a person associated with us from committing a bribery offense. We and those
acting  on  our  behalf  operate  in  a  number  of  jurisdictions  that  pose  a  high  risk  of  potential  Bribery  Act  or  FCPA  violations,  and  we  participate  in
collaborations and relationships with third parties whose corrupt or illegal activities could potentially subject us to liability under the Bribery Act, FCPA or
local anticorruption laws, even if we do not explicitly authorize or have actual knowledge of such activities. In addition, we cannot predict the nature, scope
or  effect  of  future  regulatory  requirements  to  which  our  international  operations  might  be  subject  or  the  manner  in  which  existing  laws  might  be
administered or interpreted.

Compliance  with  the  Bribery  Act,  the  FCPA  and  these  other  laws  is  expensive  and  difficult,  particularly  in  countries  in  which  corruption  is  a
recognized problem. In addition, anti-corruption laws present particular challenges in the pharmaceutical industry, because, in many countries, hospitals are
operated by the government, and doctors and other hospital employees are considered foreign officials. Certain payments to hospitals in connection with
clinical trials and other work have been deemed to be improper payments to government officials and have led to enforcement actions.

We are also subject to other laws and regulations governing our international operations, including regulations administered by the governments of
the United States and the United Kingdom, and authorities in the European Union, including applicable export control regulations, economic sanctions and
embargoes on certain countries and persons, anti-money laundering laws, import and customs requirements and currency exchange regulations, collectively
referred to as the Trade Control laws.

There  is  no  assurance  that  we  will  be  completely  effective  in  ensuring  our  compliance  with  all  applicable  anti-corruption  laws,  including  the
Bribery Act, the FCPA or other legal requirements, including Trade Control laws. If we are not in compliance with the Bribery Act, the FCPA and other
anti-corruption laws or Trade Control laws, we may be subject to criminal and civil penalties, disgorgement and other sanctions and remedial measures, and
legal expenses, which could have an adverse impact on our business, financial condition, results of operations and liquidity. Likewise, any investigation of
any  potential  violations  of  the  Bribery  Act,  the  FCPA,  other  anti-corruption  laws  or  Trade  Control  laws  by  United  States,  United  Kingdom  or  other
authorities could also have an adverse impact on our reputation, our business, results of operations and financial condition. Further, the failure to comply
with laws governing international business practices may result in substantial civil and criminal penalties and suspension or debarment from government
contracting.

If we fail to comply with environmental, health and safety laws and regulations, we could become subject to fines or penalties or incur costs that could
harm our business.

We  are  subject  to  numerous  environmental,  health  and  safety  laws  and  regulations,  including  those  governing  laboratory  procedures  and  the
handling,  use,  storage,  treatment  and  disposal  of  hazardous  materials  and  wastes.  Our  operations  involve  the  use  of  hazardous  materials,  including
chemicals and biological materials. Our operations also produce hazardous waste products. We generally contract with third parties for the disposal of these
materials and wastes. We cannot eliminate the risk of contamination or injury from these materials. In the event of contamination or injury resulting from
our  use  of  hazardous  materials,  we  could  be  held  liable  for  any  resulting  damages,  and  any  liability  could  exceed  our  resources.  We  also  could  incur
significant costs associated with civil or criminal fines and penalties for failure to comply with such laws and regulations.

Although  we  maintain  insurance  to  cover  us  for  costs  and  expenses  we  may  incur  due  to  injuries  to  our  employees  resulting  from  the  use  of
hazardous  materials,  this  insurance  may  not  provide  adequate  coverage  against  potential  liabilities.  We  do  not  maintain  insurance  for  environmental
liability or toxic tort claims that may be asserted against us in connection with our storage or disposal of biological or hazardous materials.

In addition, we may incur substantial costs in order to comply with current or future environmental, health and safety laws and regulations. These
current or future laws and regulations may impair our research, development or production efforts. Our failure to comply with these laws and regulations
also may result in substantial fines, penalties or other sanctions.

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Risks Related to the Commercialization of Our Product Candidates

If we are unable to establish sales, marketing and distribution capabilities for our product candidates, or enter into sales, marketing and distribution
agreements with third parties, we may not be successful in commercializing our product candidates, if and when they are approved.

We  currently  plan  to  work  to  build  our  global  commercialization  capabilities  internally  over  time  such  that  we  are  able  to  commercialize  any
product candidate for which we may obtain regulatory approval. However, we currently have no sales, marketing or distribution capabilities and have no
experience  in  marketing  or  distributing  pharmaceutical  products.  To  achieve  commercial  success  for  any  product  candidate  for  which  we  may  obtain
marketing approval, we will need to establish a sales and marketing organization and establish logistics and distribution processes to commercialize and
deliver  our  product  candidates  to  patients  and  healthcare  providers.  The  development  of  sales,  marketing  and  distribution  capabilities  will  require
substantial resources, will be time-consuming and could delay any product launch.

If  we  are  unable  or  decide  not  to  establish  internal  sales,  marketing  and  distribution  capabilities,  we  would  have  to  pursue  collaborative
arrangements regarding the sales and marketing of our products. However, we may not be successful in entering into arrangements with third parties to sell,
market and distribute our product candidates or may be unable to do so on terms that are favorable to us, or if we are able to do so, that they would be
effective and successful in commercializing our products. Our product revenues and our profitability, if any, would likely to be lower than if we were to
sell, market and distribute any product candidates that we develop ourselves. In addition, we would have limited control over such third parties, and any of
them may fail to devote the necessary resources and attention to sell and market our product candidates effectively.

If we do not establish sales, marketing and distribution capabilities successfully, either on our own or in collaboration with third parties, we will not

be successful in commercializing our product candidates in the United States or elsewhere.

We  operate  in  a  rapidly  changing  industry  and  face  significant  competition,  which  may  result  in  others  discovering,  developing  or  commercializing
products before or more successfully than we do.

The  development  and  commercialization  of  new  biopharmaceutical  products  is  highly  competitive  and  subject  to  rapid  and  significant
technological  advancements.  We  face  competition  from  major  multi-national  pharmaceutical  companies,  biotechnology  companies  and  specialty
pharmaceutical  companies  with  respect  to  our  current  and  future  product  candidates  that  we  may  develop  and  commercialize  in  the  future.  There  are  a
number  of  large  pharmaceutical  and  biotechnology  companies  that  currently  market  and  sell  products  or  are  pursuing  the  development  of  product
candidates for the treatment of cancer. Smaller or early-stage companies may also prove to be significant competitors, particularly through collaborative
arrangements with large, established companies. Potential competitors also include academic institutions, government agencies and other public and private
research  organizations.  Due  to  their  promising  clinical  therapeutic  effect  in  clinical  exploratory  trials,  engineered  T  cell  therapies,  redirected  T  cell
therapies in general and antibody-drug conjugates are being pursued by multiple biotechnology and pharmaceutical companies, including Novartis AG, or
Novartis, Gilead Sciences, Inc., or Gilead, Bristol-Myers Squibb, or BMS, Janssen Biotech Inc., bluebird bio, Inc., or bluebird bio, Roche Holding AG,
Seattle Genetics, and Amgen Inc. Our competitors may succeed in developing, acquiring or licensing technologies and products that are more effective,
more  effectively  marketed  and  sold  or  less  costly  than  any  product  candidates  that  we  may  develop,  which  could  render  our  product  candidates  non-
competitive and obsolete.

We  are  developing  AUTO1,  our  CD19-targeting  programmed  T  cell  product  candidate  for  the  treatment  of  adult  ALL,  and  AUTO3,  our  dual-
targeting  CD19/CD22  programmed  T  cell  product  candidate  for  the  treatment  of  relapsed  or  refractory  DLBCL.  Novartis  and  Gilead  have  received
marketing approval for their anti-CD19 CAR T cell therapy, and BMS submitted their Biologics License Application in December 2019 to the FDA for
approval  of  another  anti-CD19  CAR  T  cell  therapy.  AUTO1  is  expected  to  compete  directly  with  these  companies  and  therapies.  In  addition,  some
companies,  such  as  Cellectis,  Inc.,  Les  Laboratoires  Servier  SAS  and  Allogene  Therapeutics  Inc.,  are  pursuing  allogeneic  T  cell  products  that  could
compete with our programmed T cell product candidates.

Novartis and Gilead may be successful in establishing a strong market position for their CD19-targeted CAR T cell products, and we may not be
able to compete effectively against these therapies once they have been established. In addition, our competitors with development-stage programs may
obtain marketing approval from the FDA, the EMA or other comparable regulatory authorities for their product candidates more rapidly than we do, and
they could establish a strong market position before we are able to enter the market.

Many of our competitors, either alone or with their strategic collaborators, have substantially greater financial, technical and human resources than
we  do.  Accordingly,  our  competitors  may  be  more  successful  than  we  are  in  obtaining  approval  for  treatments  and  achieving  widespread  market
acceptance, which may render our treatments obsolete or non-competitive. Mergers

39

and acquisitions in the biotechnology and pharmaceutical industries may result in even more resources being concentrated among a smaller number of our
competitors. These competitors also compete with us in recruiting and retaining qualified scientific and management personnel and establishing clinical
study sites and patient registration for clinical studies, as well as in acquiring technologies complementary to, or necessary for, our programs. Smaller or
early-stage companies may also prove to be significant competitors, particularly through collaborative arrangements with large and established companies.

Our commercial opportunity could be reduced or eliminated if our competitors develop and commercialize products that are safer, more effective,
have fewer or less severe side effects, are more convenient or are less expensive or better reimbursed than any products that we may commercialize. Our
competitors also may obtain FDA, EMA or other regulatory approval for their products more rapidly than we may obtain approval for ours, which could
result in our competitors establishing a strong market position for either the product or a specific indication before we are able to enter the market.

 Even  if  any  of  our  product  candidates  receives  marketing  approval,  it  may  fail  to  achieve  the  degree  of  market  acceptance  by  physicians,  patients,
third-party payors and others in the medical community necessary for commercial success.

Even if we obtain approvals from the FDA, the EMA or other comparable regulatory agencies and are able to initiate commercialization of our
clinical-stage product candidates or any other product candidates we develop, the product candidate may not achieve market acceptance among physicians,
patients,  hospitals,  including  pharmacy  directors,  and  third-party  payors  and,  ultimately,  may  not  be  commercially  successful.  The  degree  of  market
acceptance of our product candidates, if approved for commercial sale, will depend on a number of factors, including:

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the clinical indications for which our product candidates are approved;

physicians, hospitals, cancer treatment centers, and patients considering our product candidates as a safe and effective treatment;

hospitals and cancer treatment centers establishing the infrastructure required for the administration of redirected T cell therapies;

the potential and perceived advantages of our product candidates over alternative treatments;

the prevalence and severity of any side effects;

product labeling or product insert requirements of the FDA, the EMA or other regulatory authorities;

limitations or warnings contained in the labeling approved by the FDA or the EMA;

the timing of market introduction of our product candidates as well as competitive products;

the cost of treatment in relation to alternative treatments;

the amount of upfront costs or training required for physicians to administer our product candidates;

the availability of coverage, adequate reimbursement, and pricing by third-party payors and government authorities;

the willingness of patients to pay out-of-pocket in the absence of comprehensive coverage and adequate reimbursement by third-party payors and
government authorities;

relative convenience and ease of administration, including as compared to alternative treatments and competitive therapies; and

the effectiveness of our sales and marketing efforts and distribution support.

Our efforts to educate physicians, patients, third-party payors and others in the medical community on the benefits of our products, if approved,
may require significant resources and may never be successful. Such efforts may require more resources than are typically required due to the complexity
and  uniqueness  of  our  product  candidates.  Because  we  expect  sales  of  our  product  candidates,  if  approved,  to  generate  substantially  all  of  our  product
revenue for the foreseeable future, the failure of our product candidates to find market acceptance would harm our business and could require us to seek
additional financing.

In  addition,  although  we  are  not  utilizing  embryonic  stem  cells  or  replication  competent  vectors,  adverse  publicity  due  to  the  ethical  and  social
controversies surrounding the therapeutic use of such technologies, and reported side effects from any clinical trials using these technologies or the failure
of such trials to demonstrate that these therapies are safe and effective, may limit market acceptance our product candidates. If our product candidates are
approved but fail to achieve market acceptance among physicians, patients, hospitals, cancer treatment centers or others in the medical community, we will
not be able to generate significant revenue.

40

 
Even if our products achieve market acceptance, we may not be able to maintain that market acceptance over time if new products or technologies

are introduced that are more favorably received than our products, are more cost effective or render our products obsolete.

Coverage and adequate reimbursement may not be available for our current or any future product candidates, which could make it difficult for us to
sell profitably, if approved.

Market  acceptance  and  sales  of  any  product  candidates  that  we  commercialize,  if  approved,  will  depend  in  part  on  the  extent  to  which
reimbursement for these products and related treatments will be available from third-party payors, including government health administration authorities,
managed care organizations and private health insurers. Third-party payors decide which therapies they will pay for and establish reimbursement levels.
Third-party payors in the United States often rely upon Medicare coverage policy and payment limitations in setting their own coverage and reimbursement
policies. However, decisions regarding the extent of coverage and amount of reimbursement to be provided for any product candidates that we develop will
be made on a payor-by-payor basis. One payor’s determination to provide coverage for a drug does not assure that other payors will also provide coverage
for the drug. Additionally,  a  third-party  payor’s  decision  to  provide  coverage  for  a  therapy  does  not  imply  that  an  adequate  reimbursement  rate  will  be
approved.  Third-party  payors  are  increasingly  challenging  the  price,  examining  the  medical  necessity  and  reviewing  the  cost-effectiveness  of  medical
products,  therapies  and  services,  in  addition  to  questioning  their  safety  and  efficacy.  We  may  incur  significant  costs  to  conduct  expensive  pharmaco-
economic studies in order to demonstrate the medical necessity and cost-effectiveness of our product candidates, in addition to the costs required to obtain
FDA approvals. Our product candidates may not be considered medically necessary or cost-effective.

Each payor determines whether or not it will provide coverage for a therapy, what amount it will pay the manufacturer for the therapy, and on what
tier of its list of covered drugs, or formulary, it will be placed. The position on a payor’s formulary, generally determines the co-payment that a patient will
need  to  make  to  obtain  the  therapy  and  can  strongly  influence  the  adoption  of  such  therapy  by  patients  and  physicians.  Patients  who  are  prescribed
treatments  for  their  conditions  and  providers  prescribing  such  services  generally  rely  on  third-party  payors  to  reimburse  all  or  part  of  the  associated
healthcare  costs.  Patients  are  unlikely  to  use  our  products,  and  providers  are  unlikely  to  prescribe  our  products,  unless  coverage  is  provided  and
reimbursement  is  adequate  to  cover  a  significant  portion  of  the  cost  of  our  products  and  their  administration.  Therefore,  coverage  and  adequate
reimbursement is critical to new medical product acceptance.

A primary trend in the U.S. healthcare industry and elsewhere is cost containment. Third-party payors have attempted to control costs by limiting
coverage and the amount of reimbursement for particular medications. We cannot be sure that coverage and reimbursement will be available for any drug
that we commercialize and, if reimbursement is available, what the level of reimbursement will be. Even if favorable coverage and reimbursement status is
attained for one or more product candidates for which we receive regulatory approval, less favorable coverage policies and reimbursement rates may be
implemented in the future. Inadequate coverage and reimbursement may impact the demand for, or the price of, any drug for which we obtain marketing
approval.  If  coverage  and  adequate  reimbursement  are  not  available,  or  are  available  only  to  limited  levels,  we  may  not  be  able  to  successfully
commercialize our current and any future product candidates that we develop.

Additionally, we are developing a proprietary diagnostic test for use with certain of our product candidates. We will be required to obtain coverage
and  reimbursement  for  this  test  separate  and  apart  from  the  coverage  and  reimbursement  we  seek  for  our  product  candidates,  if  approved.  There  is
significant uncertainty regarding our ability to obtain coverage and adequate reimbursement for this proprietary diagnostic test for reasons similar to those
applicable to our product candidates.

Product liability lawsuits against us could cause us to incur substantial liabilities and to limit commercialization of any products that we may develop.

We face an inherent risk of product liability exposure related to the testing of our product candidates in human clinical trials and will face an even
greater  risk  if  we  commercially  sell  any  products  that  we  may  develop.  If  we  cannot  successfully  defend  ourselves  against  claims  that  our  product
candidates or products caused injuries, we will incur substantial liabilities. Regardless of merit or eventual outcome, liability claims may result in:

•

•

•

reduced resources of our management to pursue our business strategy;

decreased demand for any product candidates or products that we may develop;

injury to our reputation and significant negative media attention;

• withdrawal of clinical trial participants;

•

•

initiation of investigations by regulators;

product recalls, withdrawals or labeling, marketing or promotional restrictions;

41

•

•

•

•

significant costs to defend the resulting litigation;

substantial monetary awards paid to clinical trial participants or patients;

loss of revenue; and

the inability to commercialize any products that we may develop.

We currently hold £1.0 million in product liability insurance coverage in the aggregate, with a per incident limit of £1.0 million, which may not be
adequate to cover all liabilities that we may incur. We may need to increase our insurance coverage as we expand our clinical trials or if we commence
commercialization  of  our  product  candidates.  Insurance  coverage  is  increasingly  expensive.  We  may  not  be  able  to  maintain  insurance  coverage  at  a
reasonable cost or in an amount adequate to satisfy any liability that may arise.

Risks Related to Our Intellectual Property

If we are unable to obtain and maintain patent protection for our T cell programming technologies and product candidates, or if the scope of the patent
protection obtained is not sufficiently broad, our competitors could develop and commercialize technology and biologics similar or identical to ours,
and our ability to successfully commercialize our technology and product candidates may be impaired.

Our success depends, in large part, on our ability to obtain and maintain patent protection in the United States, the European Union, the United
Kingdom and other countries with respect to our product candidates. We seek to protect our proprietary position by filing patent applications related to our
technology and product candidates in the major pharmaceutical markets, including the United States, major countries in Europe and Japan. If we do not
adequately protect our intellectual property, competitors may be able to use our technologies and erode or negate any competitive advantage that we may
have, which could harm our business and ability to achieve profitability.

To  protect  our  proprietary  positions,  we  file  patent  applications  in  the  United  States  and  other  countries  related  to  our  novel  technologies  and
product candidates that are important to our business. The patent application and prosecution process is expensive and time-consuming. We may not be
able to file and prosecute all necessary or desirable patent applications at a reasonable cost or in a timely manner. We may also fail to identify patentable
aspects of our research and development before it is too late to obtain patent protection. It is possible that defects of form in the preparation or filing of our
patents or patent applications may exist, or may arise in the future, such as with respect to proper priority claims, inventorship, claim scope or patent term
adjustments. If any current or future licensors or licensees are not fully cooperative or disagree with us as to the prosecution, maintenance or enforcement
of any patent rights, such patent rights could be compromised and we might not be able to prevent third parties from making, using and selling competing
products.  If  there  are  material  defects  in  the  form  or  preparation  of  our  patents  or  patent  applications,  such  patents  or  applications  may  be  invalid  and
unenforceable. Moreover, our competitors may independently develop equivalent knowledge, methods and know-how. Any of these outcomes could impair
our ability to prevent competition from third parties.

Prosecution of our owned and in-licensed patent portfolio is at an early stage. We currently have nine patents which have been issued from our
pending  applications  in  the  United  States,  and  eleven  patents  which  have  been  issued  from  our  pending  applications  in  Europe.  Much  of  our  patent
portfolio consists of pending priority applications that are not examined and pending applications under the Patent Cooperation Treaty, or PCT. Neither
priority applications nor PCT applications can themselves give rise to issued patents. Rather, protection for the inventions disclosed in these applications
must  be  further  pursued  by  applicable  deadlines  via  applications  that  are  subject  to  examination.  As  applicable  deadlines  for  the  priority  and  PCT
applications  become  due,  we  will  need  to  decide  whether  and  in  which  countries  or  jurisdictions  to  pursue  patent  protection  for  the  various  inventions
claimed in these applications, and we will only have the opportunity to pursue and obtain patents in those jurisdictions where we pursue protection.

It  is  also  possible  that  we  will  fail  to  identify  patentable  aspects  of  our  research  and  development  output  before  it  is  too  late  to  obtain  patent
protection.  The  patent  applications  that  we  own  or  in-license  may  fail  to  result  in  issued  patents  with  claims  that  cover  our  current  and  future  product
candidates  in  the  United  States  or  in  other  foreign  countries.  Our  patent  applications  cannot  be  enforced  against  third  parties  practicing  the  technology
claimed in such applications unless, and until, a patent issues from such applications, and then only to the extent the issued claims cover the technology.

If the patent applications we hold or have in-licensed with respect to our development programs and product candidates fail to issue, if their breadth

or strength of protection is threatened, or if they fail to provide meaningful exclusivity for our current and

42

future product candidates, it could threaten our ability to commercialize our product candidates. Any such outcome could have a negative effect on our
business.

The  patent  position  of  biotechnology  and  pharmaceutical  companies  generally  is  highly  uncertain.  Changes  in  either  the  patent  laws  or
interpretation of the patent laws in the United States and other countries may diminish the value of our patents or narrow the scope of our patent protection.
In addition, the protections offered by laws of different countries vary. No consistent policy regarding the breadth of claims allowed in biotechnology and
pharmaceutical patents has emerged to date in the United States or in many foreign jurisdictions. In addition, the determination of patent rights with respect
to  pharmaceutical  compounds  and  technologies  commonly  involves  complex  legal  and  factual  questions,  which  has  in  recent  years  been  the  subject  of
much litigation. As a result, the issuance, scope, validity, enforceability and commercial value of our patent rights, whether owned or in-licensed, are highly
uncertain.  Furthermore,  recent  changes  in  patent  laws  in  the  United  States,  may  affect  the  scope,  strength  and  enforceability  of  our  patent  rights  or  the
nature of proceedings that may be brought by or against us related to our patent rights. Additionally, the U.S. Supreme Court has ruled on several patent
cases in recent years either narrowing the scope of patent protection available in certain circumstances or weakening the rights of patent owners in certain
situations. In addition to increasing uncertainty with regard to our ability to obtain patents in the future, this combination of events has created uncertainty
with respect to the value of patents, once obtained. Depending on decisions by the U.S. Congress, the U.S. federal courts, and the USPTO, the laws and
regulations governing patents could change in unpredictable ways that could weaken our ability to obtain patents or to enforce any patents that we might
obtain in the future.

We may not be aware of all third-party intellectual property rights potentially relating to our current and future our product candidates. Publications
of  discoveries  in  the  scientific  literature  often  lag  behind  the  actual  discoveries,  and  patent  applications  in  the  United  States  and  other  jurisdictions  are
typically not published until 18 months after filing, or in some cases not at all. Therefore, we cannot be certain that we or our licensors were the first to
make the inventions claimed in our patents or pending patent applications, or that we or our licensors were the first to file for patent protection of such
inventions. Similarly, should we own or in-license any patents or patent applications in the future, we may not be certain that we or the applicable licensor
were the first to file for patent protection for the inventions claimed in such patents or patent applications. As a result, the issuance, scope, validity and
commercial value of our patent rights cannot be predicted with any certainty. Moreover, we may be subject to a third-party pre-issuance submission of prior
art to the U.S. Patent and Trademark Office, or USPTO, or become involved in opposition, derivation, reexamination, inter partes review or interference
proceedings,  in  the  United  States  or  elsewhere,  challenging  our  patent  rights  or  the  patent  rights  of  others.  An  adverse  determination  in  any  such
submission,  proceeding  or  litigation  could  reduce  the  scope  of,  or  invalidate,  our  patent  rights,  allow  third  parties  to  commercialize  our  technology  or
product  candidates  and  compete  directly  with  us,  without  payment  to  us,  or  result  in  our  inability  to  manufacture  or  commercialize  products  without
infringing third-party patent rights, which could significantly harm our business and results of operations.

Our pending and future patent applications, whether owned or in-licensed, may not result in patents being issued that protect our technology or
product candidates, in whole or in part, or which effectively prevent others from commercializing competitive technologies and products. Even if our patent
applications issue as patents, they may not issue in a form that will provide us with any meaningful protection against competing products or processes
sufficient to achieve our business objectives, prevent competitors from competing with us or otherwise provide us with any competitive advantage. Our
competitors may be able to circumvent our owned or licensed patents, should they issue, by developing similar or alternative technologies or products in
a non-infringing manner. Our competitors may seek approval to market their own products similar to or otherwise competitive with our products. In these
circumstances,  we  may  need  to  defend  and/or  assert  our  patents,  including  by  filing  lawsuits  alleging  patent  infringement.  In  any  of  these  types  of
proceedings, a court or other agency with jurisdiction may find our patents invalid and/or unenforceable.

The  issuance  of  a  patent  is  not  conclusive  as  to  its  inventorship,  scope,  validity  or  enforceability,  and  our  owned  and  licensed  patents  may  be
challenged in the courts or patent offices in the United States and abroad. Such challenges may result in loss of exclusivity or freedom to operate or in
patent  claims  being  narrowed,  invalidated  or  held  unenforceable,  in  whole  or  in  part,  which  could  limit  our  ability  to  stop  others  from  using  or
commercializing similar or identical technology and products, or limit the duration of the patent protection of our technology and products. In addition,
given the amount of time required for the development, testing and regulatory review of new product candidates, patents protecting such candidates might
expire before or shortly after such candidates are commercialized.

Third parties may initiate legal proceedings alleging that we are infringing their intellectual property rights, the outcome of which would be uncertain
and could significantly harm our business.

Our commercial success depends, in part, on our ability to develop, manufacture, market and sell our product candidates and use our proprietary
and modular T cell programming technology without infringing the intellectual property and other proprietary rights of third parties. Numerous third-party
U.S. and non-U.S. issued patents exist in the area of biotechnology,

43

including in the area of programmed T cell therapies and including patents held by our competitors. If any third-party patents cover our product candidates
or technologies, we may not be free to manufacture or commercialize our product candidates as planned.

There is a substantial amount of intellectual property litigation in the biotechnology and pharmaceutical industries, and we may become party to, or
threatened  with,  litigation  or  other  adversarial  proceedings  regarding  intellectual  property  rights  with  respect  to  our  technology  or  product  candidates,
including interference proceedings before the USPTO. Intellectual property disputes arise in a number of areas including with respect to patents, use of
other proprietary rights and the contractual terms of license arrangements. Third parties may assert claims against us based on existing or future intellectual
property  rights  and  claims  may  also  come  from  competitors  against  whom  our  own  patent  portfolio  may  have  no  deterrent  effect.  The  outcome  of
intellectual  property  litigation  is  subject  to  uncertainties  that  cannot  be  adequately  quantified  in  advance.  Other  parties  may  allege  that  our  product
candidates or the use of our technologies infringes patent claims or other intellectual property rights held by them or that we are employing their proprietary
technology without authorization. As we continue to develop and, if approved, commercialize our current and future product candidates, competitors may
claim that our technology infringes their intellectual property rights as part of business strategies designed to impede our successful commercialization.
There are and may in the future be additional third-party patents or patent applications with claims to, for example, materials, compositions, formulations,
methods of manufacture or methods for treatment related to the use or manufacture of any one or more of our product candidates. For example, we are
aware  of  third-party  U.S.  patents  that  claim  technology  related  to  AUTO1.  These  U.S.  patents  will  expire  between  2023  and  2025,  and  there  are  no
counterpart patents in Europe or the rest of the world that extend beyond the earliest expected regulatory approval date of AUTO1. If regulatory approval is
received for AUTO1, unless we are able to obtain a license or licenses to the third-party U.S. patent or patents on commercially reasonable terms or any
applicable  patent  or  patents  are  invalidated,  held  to  be  unenforceable,  or  deemed  uninfringed  by  our  activities,  we  currently  intend  to  launch  AUTO1
outside the United States first, and delay the commercial launch of AUTO1 in the United States until the expiration of any applicable third-party patent or
patents covering AUTO1. As a result, the future commercial opportunity of AUTO1 in the United States could be adversely impacted. Moreover, we may
fail to identify relevant third party patents or patent applications, or we may incorrectly conclude that the claims of an issued patent are invalid or are not
infringed by our activities. Because patent applications can take many years to issue, third parties may have currently pending patent applications which
may later result in issued patents that any of our product candidates may infringe, or which such third parties claim are infringed by our technologies.

If  we  are  found  to  infringe  a  third  party’s  intellectual  property  rights,  we  could  be  forced,  including  by  court  order,  to  cease  developing,
manufacturing or commercializing the infringing product candidate or product. Alternatively, we may be required or may choose to obtain a license from
such third party in order to use the infringing technology and continue developing, manufacturing or marketing the infringing product candidate. However,
we may not be able to obtain any required license on commercially reasonable terms or at all. Even if we were able to obtain a license, it could be non-
exclusive,  thereby  giving  our  competitors  access  to  the  same  technologies  licensed  to  us.  In  addition,  we  could  be  found  liable  for  monetary  damages,
including  treble  damages  and  attorneys’  fees  if  we  are  found  to  have  willfully  infringed  a  patent.  A  finding  of  infringement  could  prevent  us  from
commercializing  our  product  candidates  or  force  us  to  cease  some  of  our  business  operations.  Claims  that  we  have  misappropriated  the  confidential
information  or  trade  secrets  of  third  parties  could  have  a  similar  negative  effect  on  our  business.  Even  if  successful,  the  defense  of  any  claim  of
infringement or misappropriation is time-consuming, expensive and diverts the attention of our management from our ongoing business operations.

 We  may  need  to  license  intellectual  property  from  third  parties,  and  such  licenses  may  not  be  available  or  may  not  be  available  on  commercially
reasonable terms.

A third party may hold intellectual property rights, including patent rights, that are important or necessary to the development or manufacture of our
product candidates. It may be necessary for us to use the patented or proprietary technology of third parties to commercialize our product candidates, in
which case we would be required to obtain a license from these third parties. Such a license may not be available on commercially reasonable terms, or at
all,  and  we  could  be  forced  to  accept  unfavorable  contractual  terms.  If  we  are  unable  to  obtain  such  licenses  on  commercially  reasonable  terms,  our
business could be harmed.

We may become involved in lawsuits to protect or enforce our intellectual property, which could be expensive, time-consuming and unsuccessful.

Competitors may infringe our patents, if issued, trademarks, copyrights or other intellectual property. To counter infringement or unauthorized use,
we may be required to file infringement claims, which can be expensive and time-consuming and divert the time and attention of our management and
scientific  personnel.  Any  claims  we  assert  against  perceived  infringers  could  provoke  these  parties  to  assert  counterclaims  against  us  alleging  that  we
infringe their patents, trademarks, copyrights or other intellectual property. In addition, in a patent infringement proceeding, there is a risk that a court will
decide that a patent of ours is invalid or unenforceable, in whole or in part, and that we do not have the right to stop the other party from using the invention
at issue. There is also a risk that, even if the validity of such patents is upheld, the court will construe the patent’s claims narrowly or decide that we do not
have the right to stop the other party from using the invention at issue on the grounds that our

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patents  do  not  cover  the  invention.  An  adverse  outcome  in  a  litigation  or  proceeding  involving  our  patents  could  limit  our  ability  to  assert  our  patents
against those parties or other competitors and may curtail or preclude our ability to exclude third parties from making and selling similar or competitive
products. Similarly, if we assert trademark infringement claims, a court may determine that the marks we have asserted are invalid or unenforceable, or that
the party against whom we have asserted trademark infringement has superior rights to the marks in question. In this case, we could ultimately be forced to
cease use of such trademarks.

In  any  infringement  litigation,  any  award  of  monetary  damages  we  receive  may  not  be  commercially  valuable.  Furthermore,  because  of  the
substantial amount of discovery required in connection with intellectual property litigation, there is a risk that some of our confidential information could
be  compromised  by  disclosure  during  litigation.  In  addition,  there  could  be  public  announcements  of  the  results  of  hearings,  motions  or  other  interim
proceedings or developments and if securities analysts or investors perceive these results to be negative, it could have a substantial adverse effect on the
price  of  our  ADSs.  Moreover,  there  can  be  no  assurance  that  we  will  have  sufficient  financial  or  other  resources  to  file  and  pursue  such  infringement
claims, which typically last for years before they are concluded. Some of our competitors may be able to sustain the costs of such litigation or proceedings
more  effectively  than  we  can  because  of  their  greater  financial  resources  and  more  mature  and  developed  intellectual  property  portfolios.  Even  if  we
ultimately prevail in such claims, the monetary cost of such litigation and the diversion of the attention of our management and scientific personnel could
outweigh any benefit we receive as a result of the proceedings. Accordingly, despite our efforts, we may not be able to prevent third parties from infringing,
misappropriating or successfully challenging our intellectual property rights. Uncertainties resulting from the initiation and continuation of patent litigation
or other proceedings could have a negative impact on our ability to compete in the marketplace.

 We may be subject to claims by third parties asserting that we or our employees have misappropriated their intellectual property, or claiming ownership
of what we regard as our own intellectual property.

Many of our employees were previously employed at universities or other biotechnology or pharmaceutical companies, and our founder and Chief
Scientific Officer, Dr. Martin Pulé, is currently employed both by us and UCL. Although we try to ensure that our employees do not use the proprietary
information or know-how of third parties in their work for us, we may be subject to claims that these employees or we have inadvertently or otherwise used
intellectual  property,  including  trade  secrets  or  other  proprietary  information,  of  any  such  employee’s  former  employer.  We  may  also  in  the  future  be
subject  to  claims  that  we  have  caused  an  employee  to  breach  the  terms  of  his  or  her  non-competition  or  non-solicitation  agreement.  Litigation  may  be
necessary to defend against these potential claims.

In addition, while it is our policy to require our employees and contractors who may be involved in the development of intellectual property to
execute  agreements  assigning  such  intellectual  property  to  us,  such  employees  and  contractors  may  breach  the  agreement  and  claim  the  developed
intellectual property as their own.

    As of December 31, 2020, our patent portfolio is comprised of 96 patent families, of which 25 patent families originated from UCLB, the technology-
transfer company of UCL, 3 patent families are in-licensed from Noile-Immune Biotech, Inc., and 68 patent families we own and have originated from our
own research. Of the 25 patent families that were originally in-licensed from UCL, 24 have now been assigned to Autolus under a Deed of Assignment
dated 15 October 2020. Because we have acquired or licensed certain of our patents from UCLB and Noile-Immune Biotech, Inc, we must rely on their
prior  practices  with  regard  to  the  assignment  of  such  intellectual  property.  Our  and  their  assignment  agreements  may  not  be  self-executing  or  may  be
breached, and we may be forced to bring claims against third parties or defend claims they may bring against us, to determine the ownership of what we
regard as our intellectual property.

If we fail in prosecuting or defending any such claims, in addition to paying monetary damages, we may lose valuable intellectual property rights or
personnel.  A  court  could  prohibit  us  from  using  technologies  or  features  that  are  essential  to  our  products  if  such  technologies  or  features  are  found  to
incorporate  or  be  derived  from  the  trade  secrets  or  other  proprietary  information  of  the  former  employers.  Even  if  we  are  successful  in  prosecuting  or
defending  against  such  claims,  litigation  could  result  in  substantial  costs  and  could  be  a  distraction  to  management.  In  addition,  any  litigation  or  threat
thereof may adversely affect our ability to hire employees or contract with independent service providers. Moreover, a loss of key personnel or their work
product could hamper or prevent our ability to commercialize our products.

We may be subject to claims challenging the inventorship or ownership of our owned or in-licensed patent rights and other intellectual property.

We  generally  enter  into  confidentiality  and  intellectual  property  assignment  agreements  with  our  employees,  consultants,  outside  scientific
collaborators,  sponsored  researchers  and  other  advisors.  However,  these  agreements  may  not  be  honored  and  may  not  effectively  assign  intellectual
property rights to us. For example, disputes may arise from conflicting obligations of consultants or others who are involved in developing our technology
and  product  candidates.  Litigation  may  be  necessary  to  defend  against  these  and  other  claims  challenging  inventorship  or  ownership.  The  owners  of
intellectual property in-licensed to us could also face such claims. If we or our licensors fail in defending any such claims, in addition to paying monetary
damages, we may lose

45

valuable intellectual property rights, such as exclusive ownership of, or right to use, valuable intellectual property. Such an outcome could have a material
adverse effect on our business. Even if we or our licensors are successful in defending against such claims, litigation could result in substantial costs and be
a distraction to management and other employees.

Any trademarks we may obtain may be infringed or successfully challenged, resulting in harm to our business.

We expect to rely on trademarks as one means to distinguish any of our product candidates that are approved for marketing from the products of our
competitors. We have not yet selected trademarks for our product candidates and have not yet begun the process of applying to register trademarks for our
product candidates. Once we select trademarks and apply to register them, our trademark applications may not be approved. Third parties may oppose our
trademark applications, or otherwise challenge our use of the trademarks. In the event that our trademarks are successfully challenged, we could be forced
to rebrand our products, which could result in loss of brand recognition and could require us to devote resources to advertising and marketing new brands.
Our competitors may infringe our trademarks and we may not have adequate resources to enforce our trademarks.

In addition, any proprietary name we propose to use with our clinical-stage product candidates or any other product candidate in the United States
must be approved by the FDA, regardless of whether we have registered it, or applied to register it, as a trademark. The FDA typically conducts a review of
proposed  product  names,  including  an  evaluation  of  the  potential  for  confusion  with  other  product  names.  If  the  FDA  objects  to  any  of  our  proposed
proprietary product names, we may be required to expend significant additional resources in an effort to identify a suitable proprietary product name that
would qualify under applicable trademark laws, not infringe the existing rights of third parties and be acceptable to the FDA.

If we are unable to protect the confidentiality of our trade secrets, our business and competitive position would be harmed.

In  addition  to  seeking  patent  and  trademark  protection  for  our  product  candidates,  we  also  rely  on  trade  secrets,  including  unpatented  know-
how, technology and other proprietary information, to maintain our competitive position. We seek to protect our trade secrets, in part, by entering into non-
disclosure and confidentiality agreements with parties who have access to them, such as our employees, consultants, advisors and other third parties. We
also enter into confidentiality and invention or patent assignment agreements with our employees and consultants. Despite these efforts, any of these parties
may  breach  the  agreements  and  disclose  our  proprietary  information,  including  our  trade  secrets.  Monitoring  unauthorized  uses  and  disclosures  of  our
intellectual property is difficult, and we do not know whether the steps we have taken to protect our intellectual property will be effective. In addition, we
may not be able to obtain adequate remedies for any such breaches. Enforcing a claim that a party illegally disclosed or misappropriated a trade secret is
difficult, expensive and time-consuming, and the outcome is unpredictable. In addition, some courts inside and outside the United States are less willing or
unwilling to protect trade secrets.

Moreover,  our  competitors  may  independently  develop  knowledge,  methods  and  know-how  equivalent  to  our  trade  secrets.  Competitors  could
purchase our products and replicate some or all of the competitive advantages we derive from our development efforts for technologies on which we do not
have patent protection. If any of our trade secrets were to be lawfully obtained or independently developed by a competitor, we would have no right to
prevent them, or those to whom they communicate it, from using that technology or information to compete with us. If any of our trade secrets were to be
disclosed to or independently developed by a competitor, our competitive position would be harmed.

We may not be able to protect our intellectual property rights throughout the world.

Filing, prosecuting and defending patents on product candidates in all countries throughout the world would be prohibitively expensive, and our
intellectual property rights in some countries outside the United States could be less extensive than those in the United States. In some cases, we may not
be  able  to  obtain  patent  protection  for  certain  technology  outside  the  United  States.  In  addition,  the  laws  of  some  foreign  countries  do  not  protect
intellectual property rights to the same extent as federal and state laws in the United States, even in jurisdictions where we do pursue patent protection.
Consequently, we may not be able to prevent third parties from practicing our inventions in all countries outside the United States, even in jurisdictions
where we do pursue patent protection or from selling or importing products made using our inventions in and into the United States or other jurisdictions.

Competitors may use our technologies in jurisdictions where we have not pursued and obtained patent protection to develop their own products
and, further, may export otherwise infringing products to territories where we have patent protection, but enforcement is not as strong as that in the United
States. These products may compete with our product candidates and preclinical programs and our patents or other intellectual property rights may not be
effective or sufficient to prevent them from competing.

Many companies have encountered significant problems in protecting and defending intellectual property rights in foreign jurisdictions. The legal
systems of certain countries, particularly certain developing countries, do not favor the enforcement of patents, trade secrets and other intellectual property
protection, particularly those relating to biotechnology products, which could make it difficult for us to stop the infringement of our patents, if pursued and
obtained,  or  marketing  of  competing  products  in  violation  of  our  proprietary  rights  generally.  Proceedings  to  enforce  our  patent  rights  in  foreign
jurisdictions could result in

46

 
substantial costs and divert our efforts and attention from other aspects of our business, could put our patents at risk of being invalidated or interpreted
narrowly and our patent applications at risk of not issuing and could provoke third parties to assert claims against us. We may not prevail in any lawsuits
that  we  initiate,  and  the  damages  or  other  remedies  awarded,  if  any,  may  not  be  commercially  meaningful.  Accordingly,  our  efforts  to  enforce  our
intellectual property rights around the world may be inadequate to obtain a significant commercial advantage from the intellectual property that we develop
or license.

Obtaining  and  maintaining  our  patent  protection  depends  on  compliance  with  various  procedural,  document  submission,  fee  payment  and  other
requirements  imposed  by  governmental  patent  agencies,  and  our  patent  protection  could  be  reduced  or  eliminated  for  non-compliance  with  these
requirements.

Periodic  maintenance  and  annuity  fees  on  any  issued  patent  are  due  to  be  paid  to  the  USPTO  and  patent  agencies  outside  the  United  States  in
several  stages  over  the  lifetime  of  the  patent.  The  USPTO  and  various  foreign  governmental  patent  agencies  require  compliance  with  a  number  of
procedural, documentary, fee payment and other similar provisions during the patent application process. While an inadvertent lapse can in many cases be
cured  by  payment  of  a  late  fee  or  by  other  means  in  accordance  with  the  applicable  rules,  there  are  situations  in  which  noncompliance  can  result  in
abandonment  or  lapse  of  the  patent  or  patent  application,  resulting  in  partial  or  complete  loss  of  patent  rights  in  the  relevant  jurisdiction.  Non-
compliance events that could result in abandonment or lapse of a patent or patent application include failure to respond to official actions within prescribed
time limits, non-payment of fees and failure to properly legalize and submit formal documents. If we or our licensors fail to maintain the patents and patent
applications covering our products or product candidates, our competitors might be able to enter the market, which would harm our business. In addition, to
the extent that we have responsibility for taking any action related to the prosecution or maintenance of patents or patent application in-licensed from a
third party, any failure on our part to maintain the in-licensed rights could jeopardize our rights under the relevant license and may expose us to liability.

Risks Related to Ownership of Our Securities and Our Status as a Public Company

An active trading market for our ADSs may not continue to develop or be sustained.

Prior to our IPO in June 2018, there was no public market for our ordinary shares or our ADSs. Although our ADSs are listed on The Nasdaq Global
Select Market, we cannot assure you that an active trading market for our ADSs will continue to develop or be sustained. If an active market for our ADSs
does not continue to develop or is not sustained, it may be difficult for investors to sell ADSs without depressing the market price for the ADSs or to sell
the ADSs at all.

The trading price of our ADSs has been and may continue to be highly volatile and may fluctuate due to factors beyond our control.

    We completed our initial public offering in June 2018, and there has been a public market for the ADSs for only a short period of time. From January 1,
2020 to February 26, 2021, the closing sale price of our ADSs ranged from a high of $16.30 to a low of $4.20 per ADS. The trading price of our ADSs is
likely to continue to be subject to wide fluctuations in response to various factors, some of which are beyond our control, including limited trading volume.
In addition to the factors discussed in this “Risk Factors” section and elsewhere in this Annual Report, these factors include:

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the commencement, enrollment or results of our planned and future clinical trials;

positive or negative results from, or delays in, testing and clinical trials by us, collaborators or competitors;

the loss of any of our key scientific or management personnel;

regulatory or legal developments in the United States, United Kingdom and other countries;

the success of competitive products or technologies;

adverse actions taken by regulatory agencies with respect to our clinical trials or manufacturers;

changes or developments in laws or regulations applicable to our product candidates and preclinical program;

changes to our relationships with collaborators, manufacturers or suppliers;

concerns regarding the safety of our product candidates or programmed T cells in general;

announcements concerning our competitors or the pharmaceutical industry in general;

actual or anticipated fluctuations in our operating results;

changes in financial estimates or recommendations by securities analysts;

potential acquisitions, financing, collaborations or other corporate transactions;

the results of our efforts to discover, develop, acquire or in-license additional product candidates;

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the trading volume of our ADSs on The Nasdaq Global Select Market;

sales of our ADSs or ordinary shares by us, members of our senior management and directors or our shareholders or the anticipation that such
sales may occur in the future;

general economic, political, and market conditions and overall fluctuations in the financial markets in the United States or the United Kingdom;

price  and  volume  fluctuations  of  the  listed  securities  comparable  companies  and,  in  particular,  those  that  operate  in  the  biopharmaceutical
industry;

investors’ general perception of us and our business; and

other events and factors, many of which are beyond our control.

These  and  other  market  and  industry  factors  may  cause  the  market  price  and  demand  for  our  ADSs  to  fluctuate  substantially,  regardless  of  our
actual operating performance, which may limit or prevent investors from selling their ADSs and may otherwise negatively affect the liquidity of our ADSs.
In addition, the stock market in general, and biopharmaceutical companies in particular, have experienced extreme price and volume fluctuations that have
often been unrelated or disproportionate to the operating performance of these companies.

Broad market and industry factors, including the effects of the COVID-19 pandemic on the global economy, may negatively affect the market price
of our ADSs, regardless of our actual operating performance. For example, negative publicity regarding drug pricing and price increases by pharmaceutical
companies  has  negatively  impacted,  and  may  continue  to  negatively  impact,  the  markets  for  biotechnology  and  pharmaceutical  stocks.  In  the  past,
following  periods  of  volatility  in  the  market,  securities  class-action  litigation  has  often  been  instituted  against  companies.  Such  litigation,  if  instituted
against  us,  could  result  in  substantial  costs  and  diversion  of  management’s  attention  and  resources,  which  could  materially  and  adversely  affect  our
business, financial condition, results of operations and growth prospects.

Future sales of our ADSs in the public market could cause our share price to decline

    As of March 1, 2021, approximately 70.5 million of our ordinary shares (including ordinary shares in the form of ADSs) were issued and outstanding.
Sales of a substantial number of shares of our ADSs in the public market, or the perception that these sales might occur, could depress the market price of
our ADSs and could impair our ability to raise capital through the sale of additional equity securities.

    We previously filed a registration statement on Form S-8 under the Securities Act to register ordinary shares (including in the form of ADSs) subject to
options or other equity awards issued or reserved for future issuance under our equity incentive plans, and we have also filed a "shelf" registration
statement on Form F-3 under the Securities Act to register securities having an aggregate offering price not to exceed $300 million. We have also entered
into a sales agreement with Jefferies pursuant to which we may issue and sell ADSs with aggregate offering proceeds of up to $100 million, on an "at-the-
market" basis from time to time, and as of March 1, 2021, $84.1 million worth of ADSs remained available for sale under the program. In addition, in the
future, we may issue ordinary shares, ADS or other securities if we need to raise additional capital. The number of new ordinary shares or ADSs, or
securities convertible into our ordinary shares or ADSs, issued in connection with raising additional capital could represent a material portion of our then-
outstanding ordinary shares.

Additionally,  the  holders  of  an  aggregate  of  approximately  12.2  million  of  our  ordinary  shares,  or  their  transferees,  have  rights,  subject  to  some
conditions, to require us to file one or more registration statements covering their shares or to include their shares in registration statements that we may file
for  ourselves  or  other  shareholders.  If  we  were  to  register  the  resale  of  these  shares  or  ADSs,  they  could  be  freely  sold  in  the  public  market.  If  these
additional shares or ADSs are sold, or if it is perceived that they will be sold, in the public market, the trading price of our ADSs could decline.

If we fail to implement and maintain effective internal controls over financial reporting, our ability to produce accurate financial statements on

a timely basis could be impaired.

    We are subject to the reporting requirements of the Securities Exchange Act of 1934, the Sarbanes-Oxley Act of 2002 and the rules and regulations of
The Nasdaq Global Market. Pursuant to Section 404 of the Sarbanes-Oxley Act of 2002, beginning with this Annual Report, we were required to perform
system and process evaluation and testing of our internal control over financial reporting to allow our management to report on the effectiveness of our
internal control over financial reporting. However, while we remain an emerging growth company, we will not be required to include an attestation report
on internal control over financial reporting issued by our independent registered public accounting firm. We will incur substantial additional professional
fees and internal costs to expand our accounting and finance functions and that we expend significant management efforts.

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Under  standards  established  by  the  Public  Company  Accounting  Oversight  Board,  a  material  weakness  is  a  deficiency,  or  combination  of
deficiencies, in internal control over financial reporting such that there is a reasonable possibility that a material misstatement of a company’s annual or
interim financial statements will not be prevented or detected and corrected on a timely basis. In the course of auditing our financial statements as of and
for  the  years  ended  September  30,  2017  and  2016  in  preparation  for  our  IPO,  our  independent  registered  public  accounting  firm  identified  a  material
weakness  related  to  our  financial  statement  closing  process.  Although  we  made  significant  progress  to  enhance  our  in-house  accounting  and  finance
function,  in  connection  with  the  audit  of  our  financial  statements  as  of  and  for  the  year  ended  September  30,  2018,  our  independent  registered  public
accounting  firm  concluded  that  the  material  weakness  had  not  yet  been  fully  remediated  as  of  September  30,  2018.  This  material  weakness  primarily
related to our lack of controls over the review of new complex accounting issues involving significant judgment or estimates in the financial statement
closing  process,  and  insufficient  management  review  controls  over  identifying  the  accounting  impact  of  changes  to  contractual  arrangements  in  the
financial statement closing process, including the impact on our financial statements and disclosures.

In 2018 and 2019, we have hired additional finance and accounting personnel with appropriate expertise to perform specific functions and build
out our financial infrastructure, implemented improved processes and internal controls and further developed and documented our accounting policies and
financial reporting procedures, including ongoing senior management review and audit committee oversight. Our independent registered public accounting
firm concluded that the material weakness was remediated as of December 31, 2019.

If other material weaknesses are identified in the future or we are not able to comply with the requirements of Section 404 in a timely manner, the
accuracy  and  timing  of  our  financial  reporting  may  be  adversely  affected.  If  we  are  unable  to  maintain  effective  internal  controls,  we  may  not  have
adequate,  accurate  or  timely  financial  information,  and  we  may  be  unable  to  meet  our  reporting  obligations  as  a  public  company,  including  the
requirements of the Sarbanes-Oxley Act, we may be unable to accurately report our financial results in future periods, or report them within the timeframes
required  by  the  requirements  of  the  SEC,  Nasdaq  or  the  Sarbanes-Oxley  Act.  Failure  to  comply  with  the  Sarbanes-Oxley  Act,  when  and  as  applicable,
could also potentially subject us to sanctions or investigations by the SEC or other regulatory authorities. Any failure to maintain or implement required
new  or  improved  controls,  or  any  difficulties  we  encounter  in  their  implementation,  could  result  in  identification  of  additional  material  weaknesses  or
significant deficiencies, cause us to fail to meet our reporting obligations or result in material misstatements in our financial statements. Furthermore, if we
cannot provide reliable financial reports or prevent fraud, our business and results of operations could be harmed, and investors could lose confidence in
our reported financial information.

Our senior management, directors and principal shareholders, if they choose to act together, have the ability to control or significantly influence all
matters submitted to our shareholders for approval.

As of December 31, 2020, members of our senior management, directors and current beneficial owners of 5% or more of our ordinary shares and
their respective affiliates beneficially owned, in the aggregate, approximately 70% of our outstanding ordinary shares (including ordinary shares in the form
of ADSs). As a result, if these shareholders were to choose to act together, they would be able to control or significantly influence all matters submitted to
our  shareholders  for  approval,  as  well  as  our  management  and  affairs.  For  example,  these  persons,  if  they  choose  to  act  together,  would  control  or
significantly influence the election of directors and approval of any merger, consolidation or sale of all or substantially all of our assets. This concentration
of ownership control may harm the market price of our ADSs by:

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delaying, deferring, or preventing a change in control;
entrenching our management and/or the board of directors;
impeding a merger, scheme of arrangement, consolidation, takeover, or other business combination involving us; or
discouraging a potential acquirer from making a tender offer or otherwise attempting to obtain control of us.

In addition, some of these persons or entities may have interests different than yours. For example, because many of these shareholders purchased their
shares at prices substantially lower than our current trading price and have held their shares for a longer period, they may be more interested in selling our
company to an acquirer than other investors, or they may want us to pursue strategies that deviate from the interests of other shareholders.

The rights of our shareholders may differ from the rights typically offered to shareholders of a U.S. corporation.

We are incorporated under English law. The rights of holders of ordinary shares and, therefore, certain of the rights of holders of our ADSs, are
governed by English law, including the provisions of the U.K. Companies Act 2006, or the Companies Act, and by our Articles of Association. These rights
differ in certain respects from the rights of shareholders in typical U.S. corporations. See the section titled “Differences in Corporate Law” set below for a
description of the principal differences between the provisions of the Companies Act applicable to us and, for example, the Delaware General Corporation
Law relating to shareholders’ rights and protections.

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Holders of our ADSs may not have the same voting rights as the holders of our ordinary shares and may not receive voting materials in time to be able
to exercise their right to vote.

    Holders of the ADSs do not have the same rights as our shareholders and in accordance with the provisions of the deposit agreement, will not be able to
exercise  voting  rights  attaching  to  the  ordinary  shares  evidenced  by  the  ADSs  on  an  individual  basis.  The  Depositary  or  its  nominee  will  act  as  the
representative for the holders of the ADSs and will exercise the voting rights attached to the ordinary shares represented by the ADSs. Holders of our ADSs
may not receive voting materials in time to instruct the Depositary to vote, and it is possible that they, or persons who hold their ADSs through brokers,
dealers or other third parties, will not have the opportunity to exercise a right to vote. Furthermore, the Depositary will not be liable for any failure to carry
out any instructions to vote, for the manner in which any vote is cast or for the effect of any such vote. As a result, holders of our ADSs may not be able to
exercise  voting  rights  and  may  lack  recourse  if  their  ADSs  are  not  voted  as  requested.  In  addition,  holders  of  our  ADSs  will  not  be  able  to  call  a
shareholders’ meeting.

Holders of our ADSs may not receive distributions on our ordinary shares represented by the ADSs or any value for them if it is illegal or impractical to
make them available to holders of ADSs.

    Although we do not have any present plans to declare or pay any dividends, in the event we declare and pay any dividend, the Depositary for the ADSs
has agreed to pay to holders of our ADSs the cash dividends or other distributions it or the custodian receives on our ordinary shares or other deposited
securities after deducting its fees and expenses. Holders of our ADSs will receive these distributions in proportion to the number of our ordinary shares
their  ADSs  represent.  However,  in  accordance  with  the  limitations  set  forth  in  the  deposit  agreement,  it  may  be  unlawful  or  impractical  to  make  a
distribution available to holders of the ADSs. We have no obligation to take any other action to permit distribution on the ADSs, ordinary shares, rights or
anything else to holders of the ADSs. This means that holders of our ADSs may not receive the distributions we make on our ordinary shares or any value
from them if it is unlawful or impractical to make them available to you. These restrictions may have an adverse effect on the value of the ADSs.

Because we do not anticipate paying any cash dividends on our ADSs in the foreseeable future, capital appreciation, if any, will be our ADS holders’
and shareholders’ sole source of gains and they may never receive a return on their investment.

Under current English law, a company’s accumulated realized profits must exceed its accumulated realized losses (on a non-consolidated basis)
before dividends can be paid. Therefore, we must have distributable profits before issuing a dividend. We have never declared or paid a dividend on our
ordinary shares in the past, and we currently intend to retain our future earnings, if any, to fund the development and growth of our business. As a result,
capital  appreciation,  if  any,  on  our  ADSs  will  be  our  ADS  holders’  sole  source  of  gains  for  the  foreseeable  future,  and  they  will  suffer  a  loss  on  their
investment if they are unable to sell their ADSs at or above the price at which they purchased the ADSs.

If we are a passive foreign investment company, there could be adverse U.S. federal income tax consequences to U.S. Holders.

Under the Internal Revenue Code of 1986, as amended, or the Code, we will be a passive foreign investment company, or PFIC, for any taxable
year in which (1) 75% or more of our gross income consists of passive income or (2) 50% or more of the average quarterly value of our assets consists of
assets  that  produce,  or  are  held  for  the  production  of,  passive  income,  including  cash.  For  purposes  of  these  tests,  passive  income  includes  dividends,
interest, gains from the sale or exchange of investment property and certain rents and royalties. In addition, for purposes of the above calculations, a non-
U.S. corporation that directly or indirectly owns at least 25% by value of the shares of another corporation is treated as if it held its proportionate share of
the assets and received directly its proportionate share of the income of such other corporation. If we are a PFIC for any taxable year during which a U.S.
Holder (as defined in Item 10.E. “Taxation - Material U.S. Federal Income Tax Considerations for U.S. Holders”) holds our ADSs, the U.S. Holder may be
subject to adverse tax consequences regardless of whether we continue to qualify as a PFIC, including ineligibility for any preferred tax rates on capital
gains or on actual or deemed dividends, interest charges on certain taxes treated as deferred, and additional reporting requirements.

We do not believe we were a PFIC for our taxable year ended December 31, 2020. Based on our current estimates of expected gross assets and
income, we do not believe we will be a PFIC for our taxable year ending December 31, 2021. However, no assurances regarding our PFIC status can be
provided for any past, current or future taxable years. The determination of whether we are a PFIC is a fact-intensive determination made on an annual
basis and the applicable law is subject to varying interpretation. In particular, the characterization of our assets as active or passive may depend in part on
our current and intended future business plans, which are subject to change. In addition, for our current and future taxable years, the total value of our
assets for PFIC testing purposes may be determined in part by reference to the market price of our ordinary shares or ADSs from time to time, which may
fluctuate considerably. Under the income test, our status as a PFIC depends on the composition of our income which will depend on the transactions we
enter into in the future and our corporate structure. The composition of our income and assets is also affected by how, and how quickly, we spend the cash
we raise in any offering. Our U.S. counsel expresses no opinion with

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respect to our PFIC status for our taxable year ended December 31, 2020, and also expresses no opinion with regard to our expectations regarding our PFIC
status in the future.

If we are a PFIC, U.S. Holders (as defined in Item 10.E. “Taxation - Material U.S. Federal Income Tax Considerations for U.S. Holders) of our
ADSs would be subject to adverse U.S. federal income tax consequences, such as ineligibility for any preferred tax rates on capital gains or on actual or
deemed  dividends,  interest  charges  on  certain  taxes  treated  as  deferred,  and  additional  reporting  requirements  under  U.S.  federal  income  tax  laws  and
regulations. For further discussion of the PFIC rules and the adverse U.S. federal income tax consequences in the event we are classified as a PFIC, see
Item 10.E. “Taxation - Material U.S. Federal Income Tax Considerations for U.S. Holders” in this Annual Report.

If a United States person is treated as owning at least 10% of our ordinary shares, including ordinary shares represented by ADSs, such holder may be
subject to adverse U.S. federal income tax consequences.

If a U.S. Holder is treated as owning (directly, indirectly or constructively through the application of attribution rules) at least 10% of the value or
voting power of our ordinary shares, including ordinary shares represented by ADSs, such U.S. Holder may be treated as a “United States shareholder”
with respect to each “controlled foreign corporation” in our group (if any). Because our group includes at least one U.S. subsidiary (Autolus Inc.), certain
of our non-U.S. subsidiaries may be treated as controlled foreign corporations (regardless of whether Autolus Therapeutics plc is treated as a controlled
foreign corporation). A United States shareholder of a controlled foreign corporation may be required to annually report and include in its U.S. taxable
income its pro rata share of “Subpart F income,” “global intangible low-taxed income” and investments in U.S. property by controlled foreign corporations,
regardless of whether we make any distributions. An individual that is a United States shareholder with respect to a controlled foreign corporation generally
would  not  be  allowed  certain  tax  deductions  or  foreign  tax  credits  that  would  be  allowed  to  a  United  States  shareholder  that  is  a  U.S.  corporation.  We
cannot  provide  any  assurances  that  we  will  assist  investors  in  determining  whether  any  of  our  non-U.S.  subsidiaries,  if  any,  are  treated  as  a  controlled
foreign corporation or whether such investor is treated as a United States shareholder with respect to any of such controlled foreign corporations. Further,
we cannot provide any assurances that we will furnish to any U.S. shareholder information that may be necessary to comply with the reporting and tax
paying obligations discussed above. Failure to comply with these reporting obligations may subject you to significant monetary penalties and may prevent
the statute of limitations with respect to your U.S. federal income tax return for the year for which reporting was due from starting. U.S. Holders should
consult their tax advisors regarding the potential application of these rules to their investment in our ADSs.

Future changes to tax laws could materially adversely affect our company and reduce net returns to our shareholders.

The  tax  treatment  of  the  company  is,  and  our  ADSs  and  ordinary  shares  are,  subject  to  changes  in  tax  laws,  regulations  and  treaties,  or  the
interpretation thereof, tax policy initiatives and reforms under consideration and the practices of tax authorities in jurisdictions in which we operate, as well
as  tax  policy  initiatives  and  reforms  related  to  the  Organization  for  Economic  Co-Operation  and  Development’s,  or  OECD,  Base  Erosion  and  Profit
Shifting, or BEPS, Project, the European Commission’s state aid investigations and other initiatives.

Such  changes  may  include  (but  are  not  limited  to)  the  taxation  of  operating  income,  investment  income,  dividends  received  or  (in  the  specific
context of withholding tax) dividends paid, or the stamp duty of stamp duty reserve tax treatment of our ADSs or ordinary shares. We are unable to predict
what tax reform may be proposed or enacted in the future or what effect such changes would have on our business, but such changes, to the extent they are
brought into tax legislation, regulations, policies or practices, could affect our financial position and overall or effective tax rates in the future in countries
where we have operations, reduce post-tax returns to our shareholders, and increase the complexity, burden and cost of tax compliance.

Tax authorities may disagree with our positions and conclusions regarding certain tax positions, or may apply existing rules in an unforeseen manner,
resulting in unanticipated costs, taxes or non-realization of expected benefits.

A  tax  authority  may  disagree  with  tax  positions  that  we  have  taken,  which  could  result  in  increased  tax  liabilities.  For  example,  Her  Majesty’s
Revenue & Customs, or HMRC, the U.S. Internal Revenue Service or another tax authority could challenge our allocation of income by tax jurisdiction and
the amounts paid between our affiliated companies pursuant to our intercompany arrangements and transfer pricing policies, including amounts paid with
respect to our intellectual property development. Similarly, a tax authority could assert that we are subject to tax in a jurisdiction where we believe we have
not established a taxable connection, often referred to as a ‘‘permanent establishment’’ under international tax treaties, and such an assertion, if successful,
could increase our expected tax liability in one or more jurisdictions. A tax authority may take the position that material income tax liabilities, interest and
penalties are payable by us, in which case, we expect that we might contest such assessment. Contesting such an assessment may be lengthy and costly and
if we were unsuccessful in disputing the assessment, the implications could increase our anticipated effective tax rate, where applicable.

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We may be unable to use net operating loss and tax credit carryforwards and certain built-in losses to reduce future tax payments or benefits from
favorable U.K. tax legislation.

As  a  U.K.  resident  trading  entity,  we  are  subject  to  U.K.  corporate  taxation.  Due  to  the  nature  of  our  business,  we  have  generated  losses  since
inception.  As  of  December  31,  2020  we  had  cumulative  carryforward  tax  losses  of  $211.7  million.  Subject  to  any  relevant  utilization  criteria  and
restrictions (including those that limit the percentage of profits that can be reduced by carried forward losses and those that can restrict the use of carried
forward losses where there is a change of ownership of more than half the ordinary shares of the company and a major change in the nature, conduct or
scale of the trade), we expect these to be eligible for carry forward and utilization against future operating profits. As a company that carries out extensive
research  and  development  activities,  we  benefit  from  the  U.K.  research  and  development  tax  relief  programs,  being  the  Small  and  Medium-sized
Enterprises R&D tax relief program, or SME Program, and, to the extent that our projects are grant funded or relate to work subcontracted to the company
by third parties, the Research and Development Expenditure Credit program, or RDEC Program. Under the SME Program, we are able to surrender some
of our trading losses that arise from our qualifying research and development activities for a cash rebate of up to 33.35% of such qualifying research and
development  expenditures.  The  majority  of  our  pipeline  research,  clinical  trials  management  and  manufacturing  development  activities  are  eligible  for
inclusion within these tax credit cash rebate claims. We may not be able to continue to claim payable research and development tax credits in the future if
we cease to qualify as a SME, based on size criteria concerning employee headcount, turnover and gross assets. The U.K. Government has published draft
legislation by which it intends to introduce a cap on payable credit claims in excess of £20,000 with effect from April 2021 by reference to, broadly, three
times the total PAYE and NICs liability of the company, subject to an exception. If such cap comes into force, and if such exception does not apply, this
could restrict the amount of payable credit that we claim.

We may benefit in the future from the United Kingdom’s “patent box” regime, which allows certain profits attributable to revenues from patented
products (and other qualifying income) to be taxed at an effective rate of 10% by giving an additional tax deduction. We are the exclusive licensee or owner
of one patent and several patent applications which, if issued, would cover our product candidates, and accordingly, future upfront fees, milestone fees,
product  revenues  and  royalties  could  be  eligible  for  this  deduction.  When  taken  in  combination  with  the  enhanced  relief  available  on  our  research  and
development  expenditures,  we  expect  a  long-term  rate  of  corporation  tax  lower  than  statutory  to  apply  to  us.  If,  however,  there  are  unexpected  adverse
changes  to  the  U.K.  research  and  development  tax  credit  regime  or  the  “patent  box”  regime,  or  for  any  reason  we  are  unable  to  qualify  for  such
advantageous  tax  legislation,  or  we  are  unable  to  use  net  operating  loss  and  tax  credit  carryforwards  and  certain  built-in  losses  to  reduce  future  tax
payments  then  our  business,  results  of  operations  and  financial  condition  may  be  adversely  affected.  This  may  impact  our  ongoing  requirement  for
investment and the timeframes within which additional investment is required.

We  will  incur  significantly  increased  costs  and  demands  upon  management  as  a  result  of  being  a  public  company,  and  our  management  will  be
required to devote substantial time to new compliance initiatives.

As a public company listed in the United States, we have and will continue to incur significant legal, accounting and other expenses that we did not
incur previously. These expenses will likely be even more significant after we no longer qualify as an emerging growth company. The Sarbanes-Oxley Act,
the Dodd-Frank Wall Street Reform and Consumer Protection Act, the listing requirements of Nasdaq and other applicable securities rules and regulations
impose various requirements on public companies in the United States, including the establishment and maintenance of effective disclosure and financial
controls  and  corporate  governance  practices.  Our  senior  management  and  other  personnel  will  need  to  devote  a  substantial  amount  of  time  to  these
compliance initiatives. Moreover, these rules and regulations will increase our legal and financial compliance costs and will make some activities more
time-consuming  and  costly.  For  example,  we  expect  that  these  rules  and  regulations  may  make  it  more  difficult  and  more  expensive  for  us  to  maintain
director  and  officer  liability  insurance,  which  in  turn  could  make  it  more  difficult  for  us  to  attract  and  retain  qualified  senior  management  personnel  or
members for our board of directors.

We  are  an  “emerging  growth  company”  and  as  a  result  of  the  reduced  disclosure  and  governance  requirements  applicable  to  emerging  growth
companies, our ADSs may be less attractive to investors.

We are an “emerging growth company” as defined in the Jumpstart Our Business Startups Act of 2012, or the JOBS Act. For as long as we continue
to  be  an  emerging  growth  company,  we  may  take  advantage  of  exemptions  from  various  reporting  requirements  that  are  applicable  to  other  public
companies  that  are  not  emerging  growth  companies,  including  not  being  required  to  comply  with  the  auditor  attestation  requirements  of  Section  404,
exemptions from the requirements of holding a nonbinding advisory vote on executive compensation and shareholder approval of any golden parachute
payments not previously approved. As an emerging growth company, we are able to report only two years of financial results and selected financial data
compared to three and five years, respectively, for comparable data reported by other public companies. We may take advantage of these exemptions until
we are no longer an emerging growth company. We could be an emerging growth company for up to five years, although circumstances could cause us to
lose  that  status  earlier,  including  if  the  aggregate  market  value  of  our  ordinary  shares,  including  ordinary  shares  represented  by  ADSs,  held  by  non-
affiliates  exceeds  $700  million  as  of  the  end  of  our  second  fiscal  quarter  before  that  time,  in  which  case  we  would  no  longer  be  an  emerging  growth
company as of the following December 31 (the last day of our fiscal year). We cannot predict if investors will find our ADSs less attractive because we
may rely on these

52

exemptions. If some investors find our ADSs less attractive as a result, there may be a less active trading market for our ADSs and the price of our ADSs
may be more volatile.

Under  Section  107(b)  of  the  JOBS  Act,  emerging  growth  companies  can  delay  adopting  new  or  revised  accounting  standards  until  such  time  as
those standards apply to private companies. We have irrevocably elected not to avail ourselves of this exemption from new or revised accounting standards
and, therefore, we are subject to the same new or revised accounting standards as other public companies that are not emerging growth companies.

As  a  foreign  private  issuer,  we  are  permitted  to  and  follow  certain  home  country  practices  in  relation  to  corporate  governance  matters  that  differ
significantly from Nasdaq corporate governance listing standards. These practices may afford less protection to shareholders than they would enjoy if
we complied fully with Nasdaq corporate governance listing standards.

As a foreign private issuer, we are permitted to and follow certain home country corporate governance practices as opposed to those requirements
that would otherwise be required by the Nasdaq Stock Market for domestic U.S. issuers. Following our home country governance practices allows us to
follow  English  corporate  law  and  the  Companies  Act  with  regard  to  certain  corporate  governance  matters  as  opposed  to  the  requirements  that  would
otherwise apply to U.S. companies listed on Nasdaq may provide less protection to our shareholders than what is accorded to investors under the Nasdaq
rules applicable to domestic U.S. issuers.

As a foreign private issuer, we are exempt from the rules and regulations under the Exchange Act related to the furnishing and content of proxy
statements. Our officers, directors and principal shareholders are also exempt from the reporting and short-swing profit recovery provisions contained in
Section 16 of the Exchange Act. In addition, we are not required under the Exchange Act to file reports and financial statements with the SEC as frequently
or as promptly as U.S. domestic companies whose securities are registered under the Exchange Act and we are exempt from filing quarterly reports with
the  SEC  under  the  Exchange  Act.  We  also  intend  to  continue  to  follow  English  corporate  governance  practices  in  lieu  of  the  following  corporate
governance requirements of Nasdaq: (i) disclosure requirement within four business days of any determination to grant a waiver of the code of business
conduct and ethics to directors and officers and (ii) requirement to obtain shareholder approval for certain issuances of securities, including shareholder
approval of option plans. Moreover, we are not required to comply with Regulation FD, which restricts the selective disclosure of material information,
although we have voluntarily adopted a corporate disclosure policy substantially similar to Regulation FD. These exemptions and leniencies will reduce the
frequency and scope of information and protections to which you may otherwise have been eligible in relation to a U.S. domestic issuer.

We may lose our foreign private issuer status in the future, which could result in significant additional costs and expenses.

    As discussed above, we are a foreign private issuer, and therefore, we are not required to comply with all of the periodic disclosure and current reporting
requirements of the Exchange Act. The determination of foreign private issuer status is made annually on the last business day of an issuer’s most recently
completed second fiscal quarter, and, accordingly, the next determination will be made with respect to us on June 30, 2021. We would lose our foreign
private issuer status if, for example, more than 50% of our ordinary shares are directly or indirectly held by residents of the United States and we fail to
meet additional requirements necessary to maintain our foreign private issuer status.

        If  we  lose  our  foreign  private  issuer  status  on  this  determination  date,  we  will  be  required  to  file  with  the  SEC  periodic  reports  and  registration
statements on U.S. domestic issuer forms beginning on January 1, 2022, which are more detailed and extensive than the forms available to a foreign private
issuer. We will also have to mandatorily comply with U.S. federal proxy requirements, and our officers, directors and principal shareholders will become
subject to the short-swing profit disclosure and recovery provisions of Section 16 of the Exchange Act. In addition, we will lose our ability to rely upon
exemptions from certain corporate governance requirements under the Nasdaq listing rules. As a U.S. listed public company that is not a foreign private
issuer,  we  would  incur  significant  additional  legal,  accounting  and  other  expenses  that  we  do  not  currently  incur  as  a  foreign  private  issuer,  as  well  as
increased accounting, reporting and other expenses in order to maintain a listing on a U.S. securities exchange.

Provisions in the U.K. City Code on Takeovers and Mergers that may have anti-takeover effects do not currently apply to us.

The  U.K.  City  Code  on  Takeovers  and  Mergers,  or  the  Takeover  Code,  applies  to  an  offer  for,  among  other  things,  a  public  company  whose
registered office is in the United Kingdom if the company is considered by the Panel on Takeovers and Mergers, or the Takeover Panel, to have its place of
central management and control in the United Kingdom (or the Channel Islands or the Isle of Man). This is known as the “residency test.” The test for
central management and control under the Takeover Code is different from that used by the U.K. tax authorities. Under the Takeover Code, the Takeover
Panel will determine whether we have our place of central management and control in the United Kingdom by looking at various factors, primarily where
the directors are resident.

53

In  June  2019,  the  Takeover  Panel  Executive  confirmed  that,  based  on  our  current  circumstances,  we  are  not  subject  to  the  Takeover  Code.  As  a
result, our shareholders are not entitled to the benefit of certain takeover offer protections provided under the Takeover Code. We believe that this position
is unlikely to change at any time in the near future but, in accordance with good practice, we will review the situation on a regular basis and consult with
the Takeover Panel if there is any change in our circumstances which may have a bearing on whether the Takeover Panel would determine our place of
central management and control to be in the United Kingdom.

You  may  face  difficulties  in  protecting  your  interests,  and  your  ability  to  protect  your  rights  through  the  U.S.  federal  courts  may  be  limited,
because we are incorporated under the laws of England and Wales, conduct most of our operations outside the United States and most of our directors
and senior management reside outside the United States.

We are incorporated and have our registered office in, and are currently existing under the laws of, England and Wales. In addition, most of our
tangible assets are located, and most of our senior management and directors reside, outside of the United States. As a result, it may not be possible to serve
process  within  the  United  States  on  certain  directors  or  us  or  to  enforce  judgments  obtained  in  U.S.  courts  against  such  directors  or  us  based  on  civil
liability provisions of the securities laws of the United States. As a result, it may not be possible for investors to effect service of process within the United
States  upon  such  persons  or  to  enforce  judgments  obtained  in  U.S.  courts  against  them  or  us,  including  judgments  predicated  upon  the  civil  liability
provisions of the U.S. federal securities laws.

The  United  States  and  the  United  Kingdom  do  not  currently  have  a  treaty  providing  for  recognition  and  enforcement  of  judgments  (other  than
arbitration  awards)  in  civil  and  commercial  matters.  Consequently,  a  final  judgment  for  payment  given  by  a  court  in  the  United  States,  whether  or  not
predicated solely upon U.S. securities laws, would not automatically be recognized or enforceable in the United Kingdom. In addition, uncertainty exists as
to  whether  English  courts  would  entertain  original  actions  brought  in  the  United  Kingdom  against  us  or  our  directors  or  senior  management  predicated
upon the securities laws of the United States or any state in the United States. Any final and conclusive monetary judgment for a definite sum obtained
against us in U.S. courts would be treated by English courts as a cause of action in itself and sued upon as a debt at common law so that no retrial of the
issues would be necessary, provided that certain requirements are met.

Whether these requirements are met in respect of a judgment based upon the civil liability provisions of the U.S. securities laws, including whether
the award of monetary damages under such laws would constitute a penalty, is subject to determination by the court making such decision. If an English
court gives judgment for the sum payable under a U.S. judgment, the English judgment will be enforceable by methods generally available for this purpose.
These methods generally permit the English court discretion to prescribe the manner of enforcement.

As  a  result,  U.S.  investors  may  not  be  able  to  enforce  against  us  or  certain  of  our  directors  any  judgments  obtained  in  U.S.  courts  in  civil  and

commercial matters, including judgments under the U.S. federal securities laws.

As an English public limited company, certain capital structure decisions will require shareholder approval, which may limit our flexibility to

manage our capital structure.

On June 18, 2018, we altered the legal status of our company under English law from a private limited company by re-registering as a public limited
company and changing our name from Autolus Therapeutics Limited to Autolus Therapeutics plc. English law provides that a board of directors may only
allot  shares  (or  rights  to  subscribe  for  or  convertible  into  shares)  with  the  prior  authorization  of  shareholders,  such  authorization  stating  the  aggregate
nominal amount of shares that it covers and valid for a maximum period of five years, each as specified in the articles of association or relevant shareholder
resolution. We have obtained authority from our shareholders to allot additional shares for a period of five years from June 2018 (being the date on which
we adopted our articles of association containing the relevant authorization), up to a maximum nominal amount of $8,400, which authorization will need to
be renewed upon expiration (i.e., at least every five years) but may be sought more frequently for additional five-year terms (or any shorter period).

English law also generally provides shareholders with preemptive rights when new shares are issued for cash. However, it is possible for the articles
of association, or for shareholders to pass a special resolution at a general meeting, being a resolution passed by at least 75% of the votes cast, to disapply
preemptive rights. Such a disapplication of preemptive rights may be for a maximum period of up to five years from the date of adoption of the articles of
association, if the disapplication is contained in the articles of association, or from the date of the shareholder special resolution, if the disapplication is by
shareholder special resolution. In either case, this disapplication would need to be renewed by our shareholders upon its expiration (i.e., at least every five
years). We have obtained authority from our shareholders to disapply preemptive rights for a period of five years from June 2018 (being the date at which
we adopted our articles of association containing the relevant disapplication) up to a maximum nominal amount of $8,400, which disapplication will need
to be renewed upon expiration (i.e., at least every five years) to remain effective, but may be sought more frequently for additional five-year terms (or any
shorter period).

54

English  law  also  generally  prohibits  a  public  company  from  repurchasing  its  own  shares  without  the  prior  approval  of  shareholders  by  ordinary
resolution, being a resolution passed by a simple majority of votes cast, and other formalities. Such approval may be for a maximum period of up to five
years.

Our articles of association designates that the U.S. federal district courts will be the exclusive forum for the resolution of any complaint asserting a
cause of action arising under the Securities Act.

Our articles of association provides that the U.S. federal district courts will be the exclusive forum for resolving any complaint asserting a cause of
action arising under the Securities Act. This choice of forum provision may limit a shareholder’s ability to bring a claim in a judicial forum that it finds
favorable for disputes with us or our directors, officers or other employees, which may discourage such lawsuits. If a court were to find either choice of
forum provision contained in our articles of association to be inapplicable or unenforceable in an action, we may incur additional costs associated with
resolving such action in other jurisdictions, which could adversely affect our results of operations and financial condition.

If equity research analysts do not publish research or reports, or publish unfavorable research or reports, about us, our business or our market, the
price and trading volume of our ADSs could decline.

The trading market for our ADSs is influenced by the research and reports that equity research analysts publish about us and our business. As a
newly  public  company,  we  have  only  limited  research  coverage  by  equity  research  analysts.  Equity  research  analysts  may  elect  not  to  provide  research
coverage of our ADSs, and such lack of research coverage may adversely affect the market price of our ADSs. Even if we have equity research analyst
coverage, we will not have any control over the analysts, or the content and opinions included in their reports. The price of our ADSs could decline if one
or more equity research analysts downgrade our ADSs or issue other unfavorable commentary or research about us. If one or more equity research analysts
ceases coverage of us or fails to publish reports on us regularly, demand for our ADSs could decrease, which in turn could cause the trading price or trading
volume of our ADSs to decline.

55

Item 4. Information on the Company.

A. History and development of the company.

    We are a public limited company, originally incorporated pursuant to the laws of England and Wales in February 2018 as a private company with limited
liability called Autolus Therapeutics Limited. Autolus Limited was originally incorporated under the laws of England and Wales in July 2014. Pursuant to
the terms of a corporate reorganization, the shareholders of Autolus Limited exchanged each of the shares held by them in Autolus Limited for the same
number and class of newly issued shares of Autolus Therapeutics Limited and, as a result, Autolus Limited became a wholly owned subsidiary of Autolus
Therapeutics Limited, a holding company incorporated in February 2018 with nominal assets and liabilities, which had not conducted any operations prior
to the share exchange other than actions incidental to the exchange and its incorporation. On June 18, 2018, Autolus Therapeutics Limited re-registered as a
public  limited  company  and  was  renamed  Autolus  Therapeutics  plc.  Following  the  re-registration  of  Autolus  Therapeutics  Limited  as  a  public  limited
company, Autolus Limited reduced its issued share capital pursuant to Part 17 of the Companies Act by way of the cancellation of all of its issued series A
preferred shares, C ordinary shares, deferred shares and all but 100 B ordinary shares. On June 22, 2018, the different classes of our issued share capital
were  converted  into  a  single  class  of  ordinary  shares  and  various  classes  of  deferred  shares,  and  we  completed  our  IPO  on  the  Nasdaq  Global  Select
Market. Our ADSs trade under the symbol AUTL. Our ordinary shares are not listed.

    Our registered office and principal executive offices are located at Forest House, 58 Wood Lane, White City, London W12 7RZ, United Kingdom and
our  telephone  number  is  +44  20  3829  6230.  Our  website  address  is  www.autolus.com.  Information  contained  in,  or  that  can  be  accessed  through,  our
website is not a part of, and shall not be incorporated by reference into, this document.  We have included our website address in this document solely as an
inactive  textual  reference.  The  SEC  maintains  a  website  (www.sec.gov)  that  contains  reports,  proxy  and  information  statements  and  other  information
regarding registrants, such as us, that file electronically with the SEC.

    Our agent for service of process in the United States is Corporation Service Company, 1180 Avenue of the Americas, Suite 210, New York, New York
10036.

Our capital expenditures for the years ended December 31, 2020 and 2019, the three months ended December 31, 2018, and year ended September
30, 2018 amounted to $14.7 million, $18.3 million, $4.4 million, and $9.5 million, respectively. These capital expenditures primarily consisted of leasehold
improvements,  laboratory  equipment,  and  computer  and  office  equipment  in  the  United  Kingdom.  We  expect  our  capital  expenditures  to  increase  in
absolute terms in the near term as we continue to advance our research and development programs, to expand our internal manufacturing capabilities, and
otherwise  to  grow  our  operations.  We  anticipate  our  capital  expenditures  in  2021  to  be  financed  from  the  proceeds  from  our  existing  cash  and  cash
equivalents, including the net proceeds from the sale of ADSs under our at-the-market offering program in January 2021 and our follow-on capital raise in
February 2021 .

B. Business overview.

We are a biopharmaceutical company developing next-generation programmed T cell therapies for the treatment of cancer. Using our broad suite of
proprietary  and  modular  T  cell  programming  technologies,  we  are  engineering  precisely  targeted,  controlled  and  highly  active  T  cell  therapies  that  are
designed to better recognize cancer cells, break down their defense mechanisms and attack and kill these cells. We believe our programmed T cell therapies
have  the  potential  to  be  best-in-class  and  offer  cancer  patients  substantial  benefits  over  the  existing  standard  of  care,  including  the  potential  for  cure  in
some patients.

Cancers  thrive  on  their  ability  to  fend  off  T  cells  by  evading  recognition  by  T  cells  and  by  establishing  other  defense  mechanisms,  such  as
checkpoint inhibition, and creating a hostile microenvironment. Our T cell programming technologies allow us to tailor our therapies to address the specific
cancer  we  are  targeting  and  introduce  new  programming  modules  into  a  patient’s  T  cells  to  give  those  T  cells  improved  properties  to  better  recognize
cancer cells and overcome fundamental cancer defense mechanisms. We believe our leadership in T cell programming technologies will provide us with a
competitive  advantage  as  we  look  to  develop  future  generations  of  T  cell  therapies  targeting  both  hematological  cancers  and  solid  tumors,  including
potential products that could have a tolerability profile such to make them amenable to be used in outpatient settings.

Our  current  clinical-stage  pipeline  comprises  five  programs  being  developed  in  eight  hematological  and  solid  tumor  indications.  We  have

worldwide commercial rights to all of our programmed T cell therapies.

Our current clinical-stage programs are:

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AUTO1:

A  CD19-targeting  programmed  T  cell  investigational  therapy  with  a  CD19  binder  designed  to  improve  the  efficacy  and  safety
profile,  as  compared  to  other  CD19  CAR  T  therapies.  In  December  2020,  we  announced  updated  data  that  supports  AUTO1’s
anti-leukemia activity in the absence of severe cytokine release syndrome, or CRS, in ALLCAR19, an ongoing Phase 1 clinical
trial in adult patients with relapsed or refractory acute B lymphocytic leukemia, or Adult ALL. Data presented showed AUTO1
was well tolerated, despite patients having high disease burden and being heavily pre-treated. High level of sustained complete
response  rate,  or  CR,  were  achieved  without  need  for  subsequent  stem  cell  transplant  and  durability  of  remissions  was  highly
encouraging. We initiated a Phase 1b/2 clinical trial of AUTO1 for the treatment of adult ALL in 2020. This trial may potentially
be a registrational trial. In addition, we are exploring activity for AUTO1 in other B-Cell malignancies. The AUTO1 ALLCAR19
Phase 1 trial has been extended in B cell lymphomas including indolent NHL, or iNHL, and chronic lymphocytic leukemia, or
CLL.  We  presented  initial  data  from  four  patients  with  iNHL  at  the  2020  Annual  Meeting  of  the  American  Society  of
Hematology, or ASH. Furthermore, AUTO1 will be investigated in primary CNS lymphoma, or PCNSL, in collaboration with our
academic partner University College London, or UCL, in an exploratory Phase 1 clinical trial called CAROUSEL, expected to
start in the first quarter of 2021. We expect to provide updates from these additional studies in 2021. Furthermore, we may pursue
a pediatric label through an investigational program in pediatric ALL.

AUTO1/22:

AUTO3:

We  commenced  a  Phase  1  clinical  trial  in  pediatric  patients  with  relapsed  or  refractory  ALL  with  our  next-generation  product
candidate, AUTO1/22, (previously designated AUTO1NG) in the fourth quarter of 2020. AUTO1/22 is a dual-targeting CAR-T
which builds on the AUTO1 approach utilizing the same CD19 CAR, alongside a novel CD22 CAR designed to reduce antigen
negative relapse of disease. We expect to report initial data from this trial in the fourth quarter of 2021.

The  first  dual-targeting,  bicistronic,  or  having  two  chimeric  antigen  receptors  within  one  vector,  programmed  T  cell
investigational therapy for the treatment of relapsed or refractory diffuse large B-cell lymphoma, or r/r DLBCL, independently
targeting B-lymphocyte antigens CD19 and CD22.

DLBCL  (ALEXANDER  Trial):  We  initiated  a  Phase  1/2  clinical  trial  of  AUTO3  in  DLBCL  in  the  third  quarter  of  2017  and
reported  initial  data  from  the  dose-escalation  phase  of  the  trial  in  the  fourth  quarter  of  2018.  Since  then,  we  have  periodically
reported  updated  safety  and  efficacy  results  indicating  that  AUTO3  was  generally  well  tolerated.  Data  presented  in  December
2020 from the cohort of patients receiving the recommended Phase 2 dose (doses of greater than or equal to 150 x 106 CAR T
cells) and pre-conditioning pembrolizumab on day minus 1, showed an objective response rate, or ORR, of 66% and CR rate, of
55%. For all patients on study, across all dose levels, that were evaluable in the trial (n = 49), the ORR was 65% and CR rate was
51%. As of the data cut-off date of October 30, 2020, 49 patients in the Phase 1/2 clinical trial of AUTO3 have been treated and
were evaluable for safety. Across all 49 patients, only three cases of neurotoxicity, or NT, have been reported, with two patients
experiencing Grade 3 or higher NT. None of the patients achieving a CR experienced any NT and all cases of NT reported have
been atypical in nature and seen in a setting with disease progression and confounding factors. A majority of patients receiving
AUTO3 in the outpatient setting did not require hospital admission. Those patients admitted were managed without requiring ICU
care. In January 2021, we announced our intent to seek a partner for the AUTO3 program before progressing the program into the
next phase of development.

AUTO4:

A programmed T cell investigational therapy for the treatment of peripheral T-cell lymphoma targeting TRBC1. Unique targeting
of  TRBC1  potentially  opens  a  new  therapeutic  approach.  The  preclinical  study  package  suggested  selective  binding  and  anti-
tumor activity of TRBC1 and TRBC2 CARs in vitro and in vivo. We initiated a Phase 1/2 clinical trial in the fourth quarter of
2018 and we expect to report Phase 1 interim data in 2021.

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AUTO6:

A programmed T cell investigational therapy targeting GD2 in development for the treatment of neuroblastoma utilizing a new
binder designed to minimize on-target, off-tumor toxicity, humanized to reduce immunogenicity, including RQR8 safety switch.
Findings from a Phase 1 clinical trial with AUTO6 were published in November 2020 and provide evidence that AUTO6 induces
clinical  activity  in  this  solid  tumor  setting  without  inducing  on-target  off-tumor  toxicity.  We  are  developing  a  next-generation
product  candidate,  which  we  refer  to  as  AUTO6NG,  which  builds  on  this  approach  utilizing  the  same  GD2  CAR  alongside
additional  programming  modules  to  enhance  the  activity  and  persistence.  In  June  2020,  we  presented  preclinical  data  of
AUTO6NG,  including  data  from  a  tumor  model  in  small  cell  lung  cancer  indicating  that  GD2  is  an  attractive  target  for
programmed T cell therapies in that indication. We expect to initiate a Phase 1/2 clinical trial of AUTO6NG in 2021.

Our Pipeline

Our  product  pipeline  is  built  on  our  core  principles  of  modular  innovation  with  protein-based  cell  programming  focused  on  advanced  targeting,
pharmacological control and enhancement of activity. After identifying a cancer target, we select the suite of programming modules that we believe is best
suited to target that particular cancer based on the latest clinical data and the results of our cancer research. The particular modules selected may vary, and
not every product candidate, including our current product candidates, contain all categories of modules. A viral vector is used to introduce combinations of
these  modules  into  the  DNA  of  the  T  cells,  as  depicted  in  the  graphic  below.  The  diagram  below  shows  how  our  programming  modules  relate  to  our
product candidates.

Our programs have been highly tailored and specifically engineered via our proprietary modules, and have the potential to be truly differentiated

assets that could address limitations of current treatments and provide innovative options for patients.

AUTO1 has an optimized engagement of the CD19 target designed to enhance the persistence of AUTO1. We believe that these properties may
enable AUTO1 to be a suitable candidate for the treatment of adult patients with ALL, who tend to be less tolerant of severe toxicity than children with
ALL. There are currently no CAR T cell therapies approved for the treatment of adult ALL. AUTO1/22 builds on the AUTO1 approach utilizing the same
CD19 CAR alongside a novel CD22 CAR designed to reduce antigen negative relapse of disease.

AUTO3 is designed to address a key escape route used by hematological cancers in response to T cell therapies. Cancer cells often mutate and
cease to express the antigen that current therapies were designed to recognize. This loss of the target antigen can lead to antigen negative relapse of disease.
Consequently,  we  have  developed  AUTO3  to  employ  a  dual-targeting  mechanism  because  we  believe  it  may  improve  response  rate  and  durability  of
treatment response and reduce the frequency of cancer relapse when compared to other currently approved single-targeting T cell therapies, including other
CAR T cell therapies.

AUTO4, which we are developing for the treatment of peripheral T-cell lymphoma, employs a novel and differentiated treatment approach. AUTO4
is  designed  to  selectively  kill  cancerous  T  cells  in  a  manner  that  we  believe  will  preserve  a  portion  of  the  patient’s  normal,  healthy  T  cells  to  maintain
immunity. It targets an antigen, TRBC1 found on approximately 40% of T cell lymphomas. Since our AUTO4 approach is a novel mechanism to target T
cells,  we  have  also  programmed  the  product  candidate  with  a  “safety  switch”  in  order  to  allow  physicians  to  manage  toxicity  by  eliminating  the
programmed T cells if a patient experiences severe adverse side effects from the treatment. AUTO5 is a preclinical TRBC2 programmed T cell product
candidate for the treatment of peripheral T-cell lymphoma. TRBC2 is found on approximately 60% of T cell lymphomas. We plan to progress AUTO5 into
the clinic in the second half of 2021, subject to clinical data from the AUTO4 program.

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We are developing AUTO6NG, which builds upon AUTO6 data by incorporating additional programming modules intended to enhance efficacy by

aiming to extend persistence, and address the layers of defense that cancer cells deploy to evade T cell killing.

AUTO7,  our  anti-PSMA  humanized  CAR  T  candidate,  is  being  developed  for  prostate  cancer  indications  and  is  expected  to  start  clinical
development in 2022. AUTO7 is designed to tackle the complex solid tumor environment, through the addition of modules, introduced in the cells via a
gamma retro-viral vector, for potentially improved persistence and resistance in prostate cancer.

AUTO8 is our next-generation product candidate for multiple myeloma which targets both BCMA and a second, undisclosed target. We also plan
on  incorporating  in  this  investigational  CAR  T  cell  therapy  two  programming  modules,  a  dominant  negative  TGFßRII  protein  and  a  truncated  SHP2
protein, designed to block inhibitory signals from the tumor microenvironment. We believe that the final design of AUTO8 has the potential to induce deep
and durable responses and extend the durability of effect over other BCMA CARs currently in development. We expect to initiate a Phase 1 clinical trial in
multiple myeloma in the first half of 2021.

We  expect  to  continue  in  2021  to  expand  our  suite  of  cell  programming  technologies  to  include  programming  modules  designed  for  allogeneic

applications. A first novel allogeneic program is expected to enter the clinic in the first half of 2021.

The manufacture and delivery of programmed T cell therapies to patients involves complex, integrated processes, including harvesting T cells from
patients, programming the T cells ex vivo, or outside the body, multiplying the programmed T cells to obtain the desired dose, and ultimately infusing the
programmed T cells back into a patient’s body. Providing T cell therapies to patients in a commercially successful manner requires a manufacturing process
that  is  reliable,  scalable  and  economical.  We  are  using  a  semi-automated,  fully  enclosed  system  for  cell  manufacturing,  which  is  designed  to  provide  a
common platform suitable for manufacturing all of our product candidates and to allow for rapid development of our product candidates through clinical
trial phases and the regulatory approval processes. In addition, this platform allows for parallel processing and the ability to scale for commercial supply in
a controlled environment and at an economical cost. We intend to continue building internal manufacturing and supply capabilities as well as to utilize the
expertise  of  collaborators  on  some  of  the  aspects  of  product  delivery,  logistics  and  capacity  expansion.  We  believe  having  established  manufacturing
processes  suitable  for  commercialization  early  in  the  development  of  our  T  cell  therapies  will  allow  us  to  focus  on  expanding  manufacturing  capacity
during our clinical trials.

We anticipate that the market for T cell therapies will be characterized by rapid cycling of product improvements. We believe our modular approach
to T cell programming and the common manufacturing platform used across all our T cell therapies will position us to more quickly develop follow-on, or
next- generation, product candidates with enhanced characteristics, such as pharmacological control, insensitivity to checkpoint inhibition or other desirable
features.

Our Management Team

Our management team has a strong track record of accomplishments in the fields of redirected T cell therapies, gene therapy, transplantation and
oncology.  Their  collective  experience  spans  key  areas  of  expertise  required  of  a  fully  integrated  company  delivering  advanced  programmed  T  cell
therapies, including fundamental innovation in therapeutic design, translational medicine and clinical development, process sciences, manufacturing and
commercialization.  We  are  led  by  Dr.  Christian  Itin,  our  Chairman  and  Chief  Executive  Officer.  His  prior  experience  includes  serving  as  the  Chief
Executive Officer of Micromet, Inc., a public biotechnology company acquired by Amgen Inc. in 2012 for $1.2 billion, where he led the development of
blinatumomab, which in 2014 became the first redirected T cell therapy approved by the U.S. Food and Drug Administration, or FDA. Our proprietary and
modular T cell programming technologies were invented by Dr. Martin Pulé, our scientific founder and Senior Vice President and Chief Scientific Officer.
Dr. Pulé has been an innovator in the field of genetic engineering of T cells for cancer treatment for almost 20 years.

Background on T Cells and Cancer Treatment Approaches

    Cancers originate from individual cells that have developed mutations in essential cellular programs, driving increased cell division and growth. A key
control mechanism to detect and eliminate such cells is the patient’s own T cells. T cells are a type of white blood cells used by the human immune system
to defend the body against infectious pathogens and cancerous cells. Using their T cell receptor like a molecular scanner, T cells are able to discriminate
between normal human cells and ones that contain a mutation that alters their function. If the T cell recognizes an altered cell, it becomes activated and kills
that particular cell. For a cancer to grow to the detriment of the patient, cancer cells evolve mechanisms to evade recognition by, or establish other defenses
against, T cells.

T Cell Activation- and Redirection-Based Therapies

    Cancer immunotherapy treatment requires the activation and expansion of cancer-specific T cells, which kill cancer cells by recognizing antigen targets
expressed  on  cancer  cells.  Studies  have  shown  that  tumors  develop  escape  mechanisms  that  prevent  T  cell-mediated  destruction  through  immune
checkpoint proteins, which shut down anti-tumor immunity. Clinical trials have shown that treatment with immune checkpoint inhibitors can restore T cell
activity and results in durable clinical responses. These observations

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have led to the FDA approval of several checkpoint inhibitors including ipilimumab (anti-CTLA-4), nivolumab (anti-PD-1), pembrolizumab (anti-PD-1),
durvalumab  (anti-PD-L1)  and  atezolizumab  (anti-PD-L1).  Treatment  with  checkpoint  inhibitors  has  shown  the  ability  to  activate  CD8+  T  cells,  shrink
tumors, and improve patient survival. While these approaches collectively represented major advances in cancer treatment, they all lack active redirection
of the patient’s T cells to the cancer, eventually limiting clinical activity.

    More recently, redirected T cell therapies that are designed to give the patient’s T cells a new specificity to recognize cancer cells have been developed.
The first approved product of this class is a bi-specific T cell engager called blinatumomab (Blincyto ) from Amgen Inc. Blinatumomab targets the CD19
antigen  on  the  surface  of  B  cells  and  cancers  derived  from  B  cells.  Blinatumomab  received  an  accelerated  approval  for  the  treatment  of  patients  with
relapsed or refractory B cell acute lymphoblastic leukemia, or B-ALL, in 2014, followed by a full approval for all age groups in B-ALL in 2017. In 2017,
the first two genetically programmed redirected T cell therapies were approved, both also targeting CD19, CAR-T therapy Kymriah  by Novartis AG for
pediatric B-ALL and Yescarta  by Kite Pharma, Inc. (acquired by Gilead Sciences, Inc.) for DLBCL. All three of these therapies received breakthrough
therapy designation and showed high response rates and, in a subset of patients, prolonged treatment effects. For those patients experiencing a relapse, the
common causes for relapse are insufficient survival of the programmed T cells, loss of the CD19 target on the cancer cells and upregulation of checkpoint
inhibitor PD-L1 on the cancer cells.

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In view of the limitations of current therapies, there remains a critical unmet medical need for improved T cell therapies. We believe that improving
efficacy and durability over the products currently on the market or in development for the treatment of cancers requires addressing target antigen loss,
countering  checkpoint  inhibition  and  adding  novel  targets  to  expand  the  range  of  indications  amenable  to  programmed  T  cell  therapy.  We  believe  our
clinical-stage product candidates and our approach to T cell programming have the potential to address these limitations.

Programmed T Cell Therapies

Process of T Cell Programming

Existing programmed T cell therapies for oncology have focused on engineering CAR T cells. CARs are membrane-bound proteins, combining the
tumor-recognition properties of an antibody with the naturally occurring T cell activation mechanism. CARs are designed such that a portion on the outside
of the T cell binds to a structure on the surface of a cancer cell and a portion on the inside of a T cell transmits an activating signal and leads the T cell to
attack the cancer cell. The actual steps to create CAR T cells start with leukapheresis, a process in which white blood cells are collected from the patient
and separated from the blood. The sample is then enriched by stimulating the T cells, which causes them to replicate. During that process, a viral gene
vector is used to introduce the genetic information encoding the CAR into the DNA of the T cells. T cells then read this information and produce CARs on
their cell surface. The programmed T cells are then infused back into the patient intravenously following a short course of chemotherapy to condition the
bone marrow to accept the programmed T cells.

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Limitations of Current T Cell Immunotherapies

Although existing T cell immunotherapies, including CAR T therapies, have shown significant efficacy in hematological malignancies, the extent
and duration of the treatment effects and disease remission remain unknown. Optimizing the targeting module of a programmed T cell may enhance its
effect and safety. Also, in response to targeted therapies, cancer cells often mutate and cease to express the antigen the therapy was designed to recognize.
This loss of target antigen leads to patient relapse. Additionally, numerous challenges, including lack of T cell persistence and upregulation of checkpoint
inhibitors,  represent  significant  hurdles  that  need  to  be  addressed  by  new  therapies.  T  cell  immunotherapies  also  have  the  capacity  to  elicit  toxicities
including  CRS,  neurologic  toxicity  and  the  elimination  of  normal  cells  via  on-target  off  tumor  recognition.  Further,  manufacturing  T  cells  can  be
prohibitively  costly  if  the  manufacturing  process  is  not  appropriately  designed  to  support  parallel  processing  and  automation.  Finally,  realization  of  the
potential of this approach across a broad range of solid tumor types will require multiple technology solutions in order to address limitations of the current
generation of therapies. Our broad array of proprietary and modular T cell programming technologies are designed to address these limitations.

Our Solution: Advanced T Cell Programming Using a Modular Approach

We are applying our broad array of T cell programming technologies and capabilities to engineer precisely targeted, controlled and highly active T
cell therapies that are designed to better recognize cancer cells, break down their defense mechanisms and attack and kill these cells. The breadth of our
technology platform allows us to select from a range of programming modules, and our modular approach is designed to enable us to tailor our therapies to
address the specific cancer we are targeting, or to improve an already established therapy, such as by making it suitable for outpatient use. We believe this
capability represents a competitive advantage in the field and will allow us to position our product candidates to have the potential to be best-in-class. 

After identifying a cancer target, we select the suite of programming modules that we believe is best suited to target that particular cancer based on
our latest clinical data and the results of our cancer research. The particular modules selected may vary, and not every product candidate, including our
current product candidates, contain all categories of modules. A viral vector is used to introduce combinations of these modules into the DNA of the T
cells,  as  depicted  in  the  graphic  below.  With  the  exception  of  AUTO1,  all  of  our  product  candidates  contain  two  or  more  programming  modules.  The
diagram in this section following the table summarizing our clinical programs shows how our programming modules relate to our product candidates.

Advanced Targeting Technologies Used in our Modular Approach

We have developed advanced antigen targeting modules to improve the ability of our programmed T cell therapies to selectively identify, target and
destroy  cancer  cells  and  overcome  shortcomings  of  the  current  generation  of  T  cell  therapies.  These  targeting  technologies  include  innovative  binders,
novel targets, dual-targeting and pattern recognition.

Innovative Binders and Novel Targets

Binding domains allow for selective targeting of cancer cells, and the properties of binders are crucial to the performance of T cell therapies. The

binders of each of our programs have been optimized, are novel binders, or bind to novel targets.

The T cells of other CD19 CAR T cell therapies that have been approved or that are in clinical development are engineered to express high affinity
binders that can engage their targets for an extended period of time. This can lead to excessive T cell activation and toxicities caused by cytokine release, as
well as exhaustion of the CAR T cell. The programmed T cells of AUTO1 express a CD19 binder with a fast off-rate, which refers to the rate at which a T
cell disengages from a target antigen. This is similar to the off-rate of naturally occurring T cells. AUTO1, with this enhanced kinetic profile, appears to
result in reduced CRS and in increased T cell engraftment compared to data reported for other CAR T cell product candidates in clinical development for
ALL that use high affinity binders.

AUTO3  includes  an  optimized  CD22  binder  that  increases  the  avidity  for  the  CD22  target.  It  is  challenging  to  target  CD22  for  immunotherapy
because  of  its  large  size  and  extensive  post-translational  modifications.  Our  optimized  CD22  binder  combines  five  CAR  binding  domains  to  allow  for
suitable orientation and efficient target engagement compared to a traditional CAR.

Each of the TRBC1 binder used in AUTO4 and TRBC2 binder used in AUTO5 are highly selective for one of two highly related variants of the
constant domain in T cell receptor beta chains. These binders allow AUTO4 to target TRBC1-positive T cell lymphoma cells without affecting healthy
TRBC2-positive T cells and allow AUTO5 to target TRBC2-positive T cell lymphoma cells without affecting healthy TRBC1-positive T cells

AUTO6  is  designed  to  target  GD2  with  an  optimized  anti-GD2  binder  which  uses  a  humanized  targeting  domain.  Initial  clinical  data  from  an

ongoing Phase 1 clinical trial sponsored by CRUK indicates early signs of clinical activity in the absence of neurotoxicity.

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Dual-Targeting Technology

Escape from T cell recognition by losing the antigen, the very structure the programmed T cell is designed to recognize, is a fundamental defense
mechanism of hematological cancers. All clinical programs targeting CD19, CD22 or BCMA in a single-target approach have reported patients relapsing
with cells that no longer have detectable levels of the target antigen. A notable example of this defense mechanism of cancer cells was reported for children
relapsing under CD19-targeting Kymriah treatment, with more than half the children at time of relapse showing a loss of the CD19 antigen on the recurring
cancer cells.

We  believe  that  directly  targeting  two  antigens  on  a  cancer  cell  will  reduce  the  chances  for  relapse  and  may  also  improve  a  response  in  those
patients  with  low  levels  of  expression  of  a  target  antigen  on  their  cancer  cells.  AUTO8,  our  programmed  T  cell  therapy  for  the  treatment  of  multiple
myeloma, binds to two receptors, BCMA and a second undisclosed target. AUTO3, the first dual-targeting programmed T cell therapy for the treatment of
pediatric ALL and DLBCL, targets both the CD19 and CD22 antigens, both of which are B-cell antigens with similar patterns of expression.

Pattern Recognition Technology

Programmed T cells are very powerful and must be highly selective for the cancer cells in order to avoid unwanted side effects. Particularly for the
treatment  of  solid  tumors,  which  have  greater  complexity,  achieving  a  sufficient  level  of  selectivity  based  on  a  single  target  to  avoid  toxicity  can  be
challenging. For such cancers, we have developed a programming module designed to make a kill decision based on the presence of two or more targets on
the cancer cell. This technology is designed to allow us to program T cells to eliminate tumor cells only if two different targets are both present on the
surface of the cell, thereby sparing healthy cells that express only one of these targets in isolation. We are also developing technology that we believe will
allow us to program T cells to eliminate a tumor if only the tumor target, but not a target only found on healthy cells, is present on the cancer cell.

Pharmacological Control of T Cell Activity

Management of toxicity is a critical step in the successful application of programmed T cell therapies. We have developed multiple technologies

designed to pharmacologically control T cell activity. These technologies fall into two distinct categories: safety switches and tunable T cells.

Safety Switches

Also  referred  to  as  “off  switches”  or  “suicide  switches,”  safety  switches  selectively  eliminate  the  programmed  T  cells  and  are  intended  to  be
triggered in the event a patient suffers certain serious adverse events related to the T cell therapy, such as CRS or neurotoxicity. We incorporate the RQR8
safety  switch  into  some  of  our  programmed  T  cell  product  candidates,  which  allows  us  to  selectively  eliminate  the  programmed  T  cells  by  the
administration of the commercially available monoclonal antibody rituximab, or Rituxan , which binds to the surface of the T cell and thereby triggers cell
death. We use the RQR8 safety switch in our AUTO4 and AUTO6 programs. The next generation of our safety switches, which we plan to incorporate in
our  solid  tumor  programs,  utilizes  rapamycin  activated  Caspase  9  (rapaCasp9),  a  cell  therapy  safety  switch  that  allows  for  selective  elimination  of
programmed T cells using a single therapeutic dose of the commercially available product rapamycin, such as sirolimus or Rapamune . Rapamycin is a
small  molecule  drug,  which  we  expect  will  have  the  benefit  of  better  tissue  penetration  and  may  require  less  time  to  take  effect  as  compared  to  a
monoclonal antibody-activated safety switch.

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Tunable T Cells

Eliminating programmed T cells with a safety switch like RQR8 has the potential to allow the patient to recover from treatment-related side effects
but  also  to  preclude  the  anti-tumor  activity  following  elimination  of  the  programmed  T  cells,  which  could  lead  to  relapse.  To  avoid  this  undesirable
consequence of the safety switch, we are developing several programming modules that are designed to allow tunable programmed T cell responses by
reducing programmed T cell activity if a patient experiences severe toxicity, while also allowing for the subsequent reactivation of programmed T cells,
thereby allowing for the possibility of persistence and sustained anti-tumor activity. One such system we have developed is designed to reversibly dampen
the activity of the programmed T cells by temporarily dislocating the signaling domain on the inside of the T cell from the cancer cell recognition domain
with two commercially available antibiotics, tetracycline and minocycline.

Enhanced T Cell Activity Technologies

We  have  also  developed  a  wide  range  of  technologies  designed  to  inhibit  the  immunosuppressive  effects  of  the  tumor  microenvironment  and

enhance T cell persistence.

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Evading Hostile Tumor Microenvironments Including Checkpoint Inhibition

Proteins  expressed  on  tumor  cells  can  trigger  inhibitory  receptors  on  T  cells  to  block  their  ability  to  eliminate  the  tumor,  such  as  PD-L1/PD-
1 immune checkpoints. These inhibitory receptors act through a common signaling pathway inside the T cell that prevents normal T cell activation. We
have developed a programming module designed to cause T cells to express a modified version of an adaptor protein, SHP2, that in preclinical studies has
been shown to efficiently counteracts the inhibition of T cells resulting from the PD-L1/PD-1checkpoint interaction. Unlike methods that rely on blocking
one inhibitory receptor using antibodies that are separately administered to the patient and are known to have significant side effects on their own, we have
designed this programming module to be engineered into the T cells and not to require the administration of a separate pharmaceutical agent. In addition, it
is designed to simultaneously disarm multiple inhibitory receptors on the cancer cell.

Enhanced T Cell Persistence

Programmed  T  cell  therapies  that  target  hematological  malignancies  are  regularly  stimulated  by  engaging  tumor  and  normal  cells  in  the  bone
marrow and lymph tissue. This continued stimulation helps the programmed T cell survive and persist, allowing them to attack the tumor for an extended
period of time. One of the challenges of targeting some solid tumors is the lack of such easily accessible stimulation for programmed T cells, leading to
poor  persistence  and  a  weak  anti-tumor  activity.  Programmed  T  cell  therapies  have  been  co-administered  with  cytokines  that  boost  T  cell  activity  and
persistence in an attempt to enhance their effect on solid tumors. However, systemic or local administration of cytokines can be toxic. Therefore, we have
developed a technology that is designed to deliver a cytokine signal directly inside our programmed T cells without administration of cytokines themselves.
Depending on the tumor microenvironment, the cytokine persistence signal may be further enhanced by antigens secreted by the tumor. We believe our
approach will be more potent and will have the potential to be less toxic, when compared to approaches that rely on systemic or local delivery of cytokines.

Advanced T Cell Programming is Key for Solid Tumor Programs

Achieving  a  meaningful  and  durable  response  with  programmed  T  cell  therapies  in  the  treatment  of  solid  tumors  is  more  challenging  than  in
hematological  cancers  for  a  variety  of  reasons.  Solid  tumors  have  fewer  suitably  selective,  single  antigen  targets  that  can  be  used  as  a  basis  for  tumor
recognition, and solid tumors employ multiple sophisticated lines of defense to evade T cell killing.

Consequently, in order to be able to tackle the more complex biology of solid tumors, we anticipate that programmed T cell products will need to
employ  multiple  modules  of  technology  to  overcome  these  challenges.  With  our  broad  array  of  proprietary  programming  modules  and  our  ability  to
incorporate  multiple  elements  into  our  programmed  T  cell  product  candidates,  we  believe  we  are  well  positioned  to  design  these  types  of  product
candidates and expand our pipeline into solid tumor indications, including with our development of AUTO6NG and AUTO7.

Advanced T Cell Programming for Allogeneic Programs

We expect to continue to expand our suite of cell programming technologies to include programming modules designed for allogeneic applications
in 2021. There are two key challenges for allogenic T cell therapies. The first is to address Graft vs Host Disease, which is mediated by the T cell Receptor
(TCR)  of  the  donor  cells  used  to  make  the  product  candidate  attacking  the  patient’s  tissues.  This  can  be  life  threatening  for  the  patient.  The  second
challenge  is  to  address  immune  rejection  where  the  patient  T  cells  recognize  and  reject  the  engineered  T  cell  as  foreign  tissue,  thus  preventing  it  from
persisting and potentially limiting its anti-tumor activity. We have developed novel approaches using protein-based programming that integrate with our
existing  T  cell  programming  and  manufacturing  platform.  TCR  expression  is  disrupted  by  intracellular  retention  and  degradation  using  a  single
programming  module  and  programming  modules  are  also  in  development  to  protect  the  donor  cells  from  immune  rejection.  These  approaches  avoid
technical and intellectual property complexities of gene editing. A first novel allogeneic application is expected to enter the clinic in the first half of 2021.

Our Product Candidates for the Treatment of Hematological Cancers

Our three clinical-stage product candidates targeting hematological cancers are AUTO1, AUTO3 and AUTO4. We have additional hematological

product candidates AUTO1NG, AUTO3NG, AUTO5, and AUTO8 in preclinical development.

AUTO1: Our Programmed T Cell Therapy for the Treatment of ALL

Introduction to AUTO1

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AUTO1 is a gene therapy product consisting of autologous enriched T cells that are transduced with a lentiviral vector to express a novel anti-CD19
Chimeric Antigen Receptor (CD19 (CAT) CAR). The transduced T cells express second-generation CARs in which the CD19 CAR construct uses 41BB-ζ
and CD3- ζ endodomains.

CD19 is an ideal target for a CAR T cell therapy as it is a cell surface marker for lymphocytes that is present on most B cell malignancies. CD19 CAR T
cell  therapies  have  proven  effective  in  treating  leukemia  and  lymphoma,  with  efficacy  dependent  on  engraftment  and  expansion  of  the  CAR  T  cells.
However,  rapid  activation  and  expansion  of  CAR  T  cells  can  result  in  CRS  &/or  ICANS,  which  in  some  cases  can  be  life-threatening,  particularly  for
elderly patients and patients with higher tumor burden that have a poor tolerance for toxicity. Furthermore, excessive activation of CAR T cells can lead to
cell exhaustion and limit their persistence, which may impact the durability of therapeutic effect. AUTO1 is an investigational therapy in which a patient’s
T cells are genetically modified to express a novel CD19-specific binder designed to reduce side effects observed with this class of therapeutics.

AUTO1, currently the subject of an ongoing Phase 1/2 trial (FELIX) in adult ALL, has been designed to recognize CD19 and interact with the
target with a fast off-rate enabled by the novel CAT scFv binding domain. This property allows the AUTO1 cells to efficiently recognize cancer cells, inject
cytotoxic proteins to initiate the natural self-destruction process present in all human cells and then rapidly disengage from them in order to engage the next
cancer cell, a process also known as serial killing. Rapid disengagement from the target antigen is expected to minimize excessive activation of the
programmed T cells, reduce toxicity and may also reduce T cell exhaustion.

Our  academic  partner,  UCL,  has  conducted  two  separate  Phase  1  clinical  trials  evaluating  the  safety  and  efficacy  of  AUTO1.  The  first  Phase  1
clinical trial in pediatric ALL patients is named the CARPALL trial the second Phase 1 clinical trial in adult ALL patients is named the ALLCAR19 trial,
latter being the foundation for a Phase 1b/2 clinical trial noted above, which we intend will be a registrational trial.

Clinical Development of AUTO1 in Adult ALL

Background of Adult ALL

AUTO1 is currently being tested in a Phase 1b/2 clinical trial for the treatment of adult ALL, which according to the American Cancer Society is predicted
to  affect  approximately  5,960  adults  in  the  United  States  in  2018.  Combination  chemotherapy  enables  90%  of  adult  patients  to  experience  complete
remission, or CR. Despite this, and in contrast to pediatric ALL, the prognosis of adult ALL is still poor and has not changed significantly during the last
two  to  three  decades,  with  long-term  remission  rates  limited  to  30-40%.  Approximately  50%  of  all  adult  ALL  patients  will  relapse,  and  data  from  the
Medical  Research  Council’s  UKALL12/ECOG  2993  study,  published  in  2007,  found  that  five-year  overall  survival,  or  OS,  rate  in  adults  who  relapse
following standard multi-agent chemotherapy is 7%. The only curative option for relapsed or refractory ALL consists of achieving a second CR by salvage
therapy followed by an allogeneic hematopoietic stem cell transplant, or allo-HSCT. Without allo-HSCT, a subsequent relapse occurs in nearly all patients.
However, less than half of patients achieve a second CR, and therefore only a subset will be eligible for this procedure. Even then, less than one-third of
patients  receiving  the  transplant  are  expected  to  sustain  long-term  disease-free  survival.  Further,  allo-HSCT  is  associated  with  severe  morbidity  and
significant  mortality.  Many  patients  with  relapsed  or  refractory  ALL  will  have  been  maximally  treated  with  chemotherapy,  and  often  do  not  achieve  a
second CR with standard-of-care chemotherapy in order to be eligible for allo-HSCT.

Two targeted immuno therapies have recently been approved for the treatment of adult ALL: blinatumomab and inotuzumab ozogamicin. Both of
these therapies achieve high complete response rates, but durability is limited. In a randomized Phase 3 clinical trial of blinatumomab in heavily pretreated
B-cell precursor ALL, the blinatumomab arm achieved a complete response rate of 44%, of which 76% also achieved MRD-negative CR, and the median
duration  of  remission  was  7.3  months.  The  median  OS  in  those  patients,  though  significantly  improved  compared  to  chemotherapy,  was  still  only  7.7
months. Similarly, in a Phase 3 clinical trial of inotuzumab ozogamicin, a higher percentage of patients achieved MRD-negative CR when treated with
inotuzumab compared to standard-of-care chemotherapy, but the median duration of remission was 4.6 months and median OS was 7.7 months.

AUTO1 Phase 1 Clinical Trial in Adult ALL (ALLCAR19 Trial)

In  the  first  quarter  of  2018,  our  academic  partner  University  College  London,  or  UCL,  initiated  a  single-arm,  open  label,  multi-center  Phase  1
clinical trial of AUTO1, named the ALLCAR19 trial, in patients aged 16 to 65 years with high-risk, relapsed or refractory CD19 positive B-lineage ALL.
The clinical trial was conducted at sites in the United Kingdom. The trial enrolled patients with a high tumor burden; 45% of treated patients had 50% or
greater bone marrow blasts. As of the most recent data cut-off date of November 12, 2020, 20 patients received AUTO1; product for 14 of those patients
was manufactured using a semi-automated, fully-enclosed process, which we anticipate using for commercial supply, if approved.

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As  of  the  most  recent  data  recent  cut-off  date  of  November  12,  2020,  20  patients  in  the  ALLCAR19  trial  had  received  AUTO1.  Based  on  this
preliminary data, we have observed a manageable safety profile for AUTO1, with no patients experiencing Grade 3 or higher CRS (using the Lee grading
criteria, which is the standard currently adopted by the medical community for grading CRS and enables comparison of clinical trial safety data across
clinical  studies)  and  three  patients  (15%)  experiencing  Grade  3  neurotoxicity  that  resolved  with  steroids.  Of  those  20  patients,  three  patients  (15%)
experienced Grade 3 neurotoxicity that resolved swiftly with steroids. We also observed Grade 3 or higher neutropenias in 7 of 20 patients reporting such
neutropenias prior to treatment with AUTO1 and 8 of 17 patients reporting Grade 3 or higher neutropenias 28 days after treatment with AUTO1, with most
resolving by month two or three post-dosing. Of the 20 patients, seven patients died while enrolled in the ALLCAR19 clinical trial, of which two deaths
were determined to be due to progression of the leukemia, one death occurred post progression following infective complication post-transplant and the
four remaining deaths were determined to be due to infectious complications (a common complication of advanced ALL). Of the 19 patients evaluable for
efficacy  (patients  with  at  least  four  weeks  follow-up  as  of  the  data  cut-off  date  of  November  12,2020),  16  patients  (84%)  achieved  minimal  residual
disease, or MRD, negative complete responses following AUTO1 infusion At 12 months post-treatment, we observed that the event free survival was 52%
and overall survival was 63%. In the sub-group of patients that were dosed with AUTO1 manufactured using our semi-automated, fully enclosed system for
manufacturing, 13 were evaluable for efficacy. In this sub-group, as of the data cut-off date of November 12,2020, 12 of the 13 evaluable patients (92%)
had achieved MRD-negative complete response. The EFS and OS data are preliminary considering the small number of patients.

In October 2019, we announced that the U.S. Food and Drug Administration, or FDA, granted orphan drug designation for AUTO1 for the treatment

of ALL. We also received IND clearance for our Phase 1b/2 clinical trial in April 2020.

The ALLCAR19 clinical trial has also been expanded to include three additional cohorts with 10 treated patients in each cohort:
Regimen B: r/r DLBCL (including transformed follicular lymphoma, or FL, but not Richter’s transformation
Regimen C: relapsed or refractory B-cell chronic lymphocytic leukemia / small lymphocytic leukemia
Regimen D: relapsed or refractory indolent B-NHL (either FL, mantle cell lymphoma or marginal zone lymphoma)

•
•
•

UCL has also initiated a Phase 1 exploratory trial of AUTO1 in patients with relapsed or refractory Primary CNS Lymphoma, or PCNSL. This trial, named
CAROUSEL  (NCT04443829),  is  expected  to  initiate  enrolment  in  the  first  quarter  of  2021  and  will  evaluate  the  feasibility  of  generating  AUTO1  and
safety of administration in this patient population

Development Strategy for Adult ALL

In 2020, we initiated a multicenter, single-arm Phase 1b/2 trial of AUTO1 in adult patients with relapsed or refractory ALL, or r/r ALL. We refer to
this trial as the FELIX trial. We are planning to recruit patients in the United Kingdom, the United States and Spain, and clinical trial sites are open for
enrollment in the United Kingdom and the United States. Subject to confirmation by regulatory authorities, we hope that this trial may be a registrational
trial in r/r adult ALL population.

The Phase 1b component of the trial is currently enrolling patients. We anticipate progressing to the Phase 2 portion of the trial in the second half of
2021. The Phase 2 cohort will include approximately 90 patients with morphological disease (>5% blasts in the bone marrow at screening), with ORR as
the primary endpoint; and EFS, DOR and MRD negative CR as key secondary endpoints. An exploratory cohort is also planned in the Phase 2 part of this
trial, in which patients with MRD positive disease in morphological remission as well as patients with isolated extramedullary disease will be treated with
AUTO1.

Background of Pediatric ALL

Pediatric  B-cell  ALL  is  a  type  of  blood  cancer  in  which  the  bone  marrow  makes  too  many  immature  lymphocytes  (named  as  lymphoid  blasts),
which are a type of white blood cell. According to the American Cancer Society, B-cell ALL is most common in childhood, peaking between two and four
years of age. As per the National Cancer Institute Surveillance, Epidemiology and End Results statistics database, there are approximately 3,400 new cases
of pediatric ALL diagnosed in the United States each year.

The current standard of care for both pediatric and adult B-cell ALL patients is a standard regimen of combination chemotherapy. Pediatric patients
typically respond well to the complex first-line chemotherapy treatment. According to the American Cancer Society, the five-year survival rate for children
with  B-cell  ALL  is  more  than  85%  overall.  However,  10  to  20%  of  pediatric  B-cell ALL  patients  relapse  with  chemotherapy-resistant  disease.  These
patients are re-treated with intensive chemotherapy, and those that respond may proceed to receive an allogenic stem cell transplant, or SCT. However, SCT
can be associated with significant long-term morbidity due to the risk of developing graft-versus-host disease, or GVHD, and treatment-related mortality,
although the risk of death have declined with better post-transplant management.

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Patients  with  high-risk  clinical  or  genetic  features  including  gene  abnormalities,  as  well  as  those  who  have  an  inadequate  response  to  initial
chemotherapy, may not respond well with the current available treatments for B-cell ALL (including SCT), some of these patients will have a five-year OS
rate of approximately 15%. Additionally, long-term survival rates are only approximately 10 to 20% among patients receiving a second SCT and negligible
in those unable to proceed to a second transplant.

There is still a significant unmet medical need in pediatric patients with high-risk relapsed or refractory B-cell ALL. CD19 CAR T cell therapies
have been developed for these patients. The approved CD19 CAR T therapy showed approximately 80% of complete molecular response rate. However, at
six months after treatment, approximately 40% of the patients relapsed and the majority of the relapses were CD19 negative disease. In fact, in one study of
CD19-targeting Kymriah treatment, approximately two-thirds of relapses were determined to have been due to loss of CD19 on the target cells.

CD19  CAR  T  cell  therapies  have  been  tested  in  pediatric  ALL  patients  and  have  shown  sustained  responses  without  allo-HSCT.  In  adult  ALL,
however, one of the major challenges has been severe toxicity, including death due to CAR T cell-mediated toxicity observed in the clinical trials of these
products. AUTO1 has been designed to reduce toxicity but still sustain durable CRs, and we believe it has the potential to become a standalone therapy for
adult ALL.

AUTO1 Phase 1 Clinical Trial in Pediatric ALL (CARPALL Trial)

The CARPALL trial was initiated by UCL in the second quarter of 2016 and is a single-arm, open label, multi-center Phase 1 trial enrolling patients
aged 24 years or younger with high-risk relapsed or refractory CD19 positive B-lineage ALL. The main objective of the trial is to evaluate the safety and
efficacy of AUTO1 when administered at a single dose of 1 million cells/kg. The trial has completed enrollment with AUTO-1. However, the extension
arm is now open, and treating pediatric ALL patients with AUTO 1/22 (previously designated AUTO1NG).

As  of  the  data  cut-off  date  of  November  22,  2019,  the  trial  has  enrolled  a  total  of  25  patients,  in  two  cohorts;  one  cohort  utilized  a  manual
manufacturing process (cohort 1) and one cohort utilized a semi-automated fully enclosed manufacturing process (cohort 2). Product was generated for 14
of 17 patients in cohort 1 and the median follow-up for the 14 treated patients was 23 months. Seven patients were treated in cohort 2. The aim of cohort 2
was to increase feasibility of manufacture at scale; one patient died before infusion and product was generated for 100% of patients. Median follow-up for
patients in cohort 2 was seven months.

None of the patients experienced Grade 3 or higher CRS and one patient out of 21 (5%) experienced Grade 4 neurotoxicity, which was deemed
more consistent with fludarabine than CAR-associated neurotoxicity. Two patients experienced Grade 5 sepsis and death, one in the context of progressive
disease  and  the  second  was  considered  related  to  AUTO1.  This  patient  was  in  MRD-negative  CR  and  had  ongoing  Grade  4  cytopenia  associated  with
resistant  HSV  encephalitis.  13  patients  experienced  Grade  4  cytopenias  that  were  ongoing  at  day  28.  Nineteen  of  21  treated  patients  (90%)  achieved
molecular complete remission at post-infusion. Consistent with preclinical results, CAR T cell expansion and persistence was excellent and CARs were
detectable by flow for up to 36 months in four of the patients in cohort 1 who had ongoing responses beyond 12 months. Persistence was noted in 15 of 21
patients at last follow-up, up to 36 months. All of the patients in cohort 2 achieved molecular complete remission at one month post-infusion.

For  cohort  1,  with  a  median  follow-up  of  23  months,  the  overall  survival  at  six  and  12  months  was  86%  and  71%,  respectively,  and  event-free
survival at six and 12 months was 71% and 54%, respectively. In cohort 2, at a median follow-up of 7 months, five patients remain in complete molecular
remission and two patients relapsed. Five of eight evaluable relapses in cohort 1 and cohort 2 combined were due to CD19 negative escape.

AUTO1/22

We commenced a Phase 1 clinical trial in pediatric patients with relapsed or refractory ALL with our next-generation product candidate, AUTO1/22
(previously designated AUTO1NG) in the fourth quarter of 2020. AUTO1/22 is a dual-targeting CAR-T which builds on the AUTO1 approach utilizing the
same CD19 CAR, alongside a novel CD22 CAR designed to reduce antigen negative relapse of disease. We expect to report initial data from this trial in the
fourth quarter of 2021.
AUTO3: Our Programmed T Cell Therapy for the Treatment of Pediatric ALL and Adult DLBCL

Introduction to AUTO3

AUTO3,  the  first  dual-targeting  programmed  T  cell  product  candidate  that  targets  B  cell  antigens  CD19  and  CD22,  is  being  explored  for  the

treatment of pediatric patients with relapsed or refractory B-cell ALL, as well as the treatment of adult patients with DLBCL.

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By simultaneously targeting both B cell antigens, we believe the novel molecular design of AUTO3 addresses a major limitation of current CAR T
cell products that target only CD19 or CD22; the loss or downregulation of the target antigen on the surface of the cancer cell, which is the main cause of
disease relapse after single targeting CAR T-cell therapy.

AUTO3 Phase 1/2 Clinical Trial in Pediatric ALL (AMELIA Trial)

We  initiated  a  single-arm,  open  label,  multi-center  Phase  1/2  clinical  trial  of  AUTO3  in  patients  up  to  24  years  of  age  with  high-risk  relapsed  or
refractory  B-lineage  ALL  in  the  third  quarter  of  2017.  We  refer  to  this  trial  as  the  AMELIA  Trial.  We  presented  final  results  from  this  clinical  trial  in
January 2021, with a data cut-off date of July 29, 2020.

6

Twenty-three patients were screened in this trial, and product was generated in 19 of 20 (95%) patients who underwent leukapheresis and a total of 15
patients received infusion of AUTO3. AUTO3 has been observed to have a manageable safety profile, with no patients experiencing Grade 3 or higher
CRS. One patient treated in the low dose (0.3 x10 /kg) had Grade 3 encephalopathy which was considered likely related to prior intrathecal methotrexate.
The majority of Grade 3 or higher adverse events were determined to be hematological in nature. No patient deaths have been determined to be related to
AUTO3.  The  rate  of  complete  response/complete  response  with  incomplete  bone  marrow  recovery  (CR/CRi)  and  Minimal  Residual  Disease  (MRD)-
Negative Response in the Bone Marrow (PCR)) was 87% (13/15) at month 1 after AUTO3 infusion. Two patients did not respond to the treatment. As of
the data cut-off (29-Jul-2020), 9 patients ultimately had morphological relapse. The majority of the relapses (6 of 9 patients) were CD19-positive/CD22-
positive disease associated with a lack of long-term CAR T cell persistence. The median morphological EFS was 4.9 months (95% CI: 1.64, 12.42). The
most common cause of relapse was due to the absence of CAR T cell persistence. Three of the 9 disease relapses were CD19-negative (including 1 patient
with CD22-positive disease relapse, 1 patient with low expression of CD22, and 1 patient with both antigens lost: CD19-negative/CD22-negative disease
relapse). The 1 case with CD22-negative disease relapse and the 1 disease relapse with downregulation of CD22 expression confirms the activity of the
CD22 CAR in the AUTO3 construct. Based on this small number of 15 patients, 1-year OS is 60%, in the pediatric and young adult relapsed/refractory B-
cell ALL setting.

Development Strategy for Pediatric ALL

In  September  2019,  we  reported  that  we  would  not  be  continuing  development  of  AUTO3  in  pediatric  B-cell  ALL  but  instead  will  continue
development in a pediatric setting with AUTO1 in the form a Pediatric Investigational Program, required by the European Medicines Agency, or EMA, and
with  AUTO1/22,  a  next  generation  version  of  AUTO1,  which  incorporates  the  CD19  CAR  of  AUTO1  and  a  novel  CD22  CAR.  We  initiated  clinical
development of AUTO1/22 in 2020.

Background of DLBCL

Non-Hodgkin lymphoma, or NHL, consists of a diverse group of malignant neoplasms. According to the American Cancer Society, DLBCL is the
most common subtype of NHL, accounting for approximately one-third of the approximately 72,000 adult NHL patients diagnosed in 2017 in the United
States.  DLBCL  arises  from  a  mature  B  cell  that  generally  express  CD19  and  CD22  antigens  on  the  surface.  DLBCL  is  classified  as  an  aggressive
lymphoma, in which survival is measured in months rather than years.

First-line  therapy  usually  consists  of  a  chemotherapy  regimen  known  as  R-CHOP,  which  combines  the  monoclonal  antibody  rituximab  with  the
drugs cyclophosphamide, doxorubicin, vincristine and prednisone. Approximately 50% to 60% of DLBCL patients are cured with first-line therapy and do
not have recurrence of their lymphoma.

For  patients  who  relapse  or  are  refractory  to  first-line  therapy,  the  current  standard  of  care  for  second-line  therapy  consists  of  a  platinum-based
chemotherapy regimen with rituximab. These second-line chemotherapy regimens are either R-ICE, consisting of rituximab, ifosfamide, carboplatin and
etoposide, or R-DHAP, consisting of rituximab, dexamethasone, cytarabine and cisplatin. Patients who respond to second-line therapy may go on to receive
autologous  hematopoietic  stem  cell  transplantation,  or  HSCT.  Patients  who  are  not  candidates  for  HSCT  or  those  who  do  not  respond  to  second-line
therapy or who relapse after HSCT are typically treated with a third-line salvage chemotherapy. These patients have a poor prognosis, and treatment is
generally palliative to try to prevent further cancer growth without the intent to cure.

Indolent lymphomas account for 40% of all NHL cases. The subtypes of indolent lymphoma, including follicular lymphoma and others, initially
respond well to chemotherapy or antibody therapy, or a combination of both. However, in patients with progressive disease or relapse after CR, there is no
defined  standard  of  care,  and  such  patients  are  generally  encouraged  to  participate  in  clinical  trials  whenever  possible.  Relapsed  patients  who  are
symptomatic or need treatment are usually treated with chemotherapy, which is unfortunately not curative. Additionally, a minority of these patients are
eligible to receive HSCT, which provides long-term disease free survival in some cases.

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AUTO3 Phase 1/2 Clinical Trial in Adult DLBCL (ALEXANDER Trial)

We  have  designed  AUTO3  to  address  limitations  of  current  therapies  for  DLBCL.  Simultaneous  targeting  of  both  CD19  and  CD22  antigens  is
designed to reduce CD19 antigen negative disease relapses observed in one-third of the patients relapsing after treatment with Yescarta. Our clinical trial
design also includes the administration of three doses of an anti-checkpoint inhibitor, designed to address tumor relapse due to upregulation of checkpoints
in DLBCL patients treated with CAR T cell therapy. We are initially developing AUTO3 as a third-line therapy for DLBCL.

In  September  2017,  we  initiated  a  single-arm,  open  label,  multi-center  Phase  1/2  clinical  trial  of  AUTO3,  which  included  a  cohort  of  patients
receiving  AUTO3  followed  by  limited  duration  of  consolidation  with  the  anti-PD-1  antibody  pembrolizumab  in  adult  DLBCL  patients  who  have
chemotherapy-refractory  disease  or  with  relapsed  disease  after  two  lines  of  prior  therapy.  We  refer  to  this  trial  as  the  ALEXANDER  Trial.  We  have
completed the Phase 1 dose escalation portion of the trial and have declared a RP2D along with feasibility of use in the outpatient setting.

As of the October 30, 2020 data cut-off date, 49 patients in the ALEXANDER trial have been treated and were evaluable for safety. Across all
dose levels, 43 patients were evaluable for efficacy, with an objective response rate (ORR) of 65% and a CR rate of 51%. Of the 29 evaluable patients
receiving the recommended Phase 2 dose (a dose of ≥ 150 x 10  cells) and pre-conditioning with pembrolizumab at Day -1, the ORR was 66% and the CR
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rate was 55%. A subsequent analysis of these data suggested a superior response rate at higher dose levels, with 15 evaluable patients treated at 450 x 10
cells  achieving  an  ORR  of  87%  and  a  CRR  of  73%.  Across  all  cohorts  in  the  trial,  73%  (16/22)  of  patients  achieving  a  CR  were  without  disease
progression at a median follow up of 4 months (1 – 24 months). AUTO3 was observed to be well tolerated, with low rates of cytokine release syndrome
(CRS) and neurotoxicity (NT). Across all 49 patients, there was only one case of Grade 3 CRS with primary infusion, and only three cases of NT were
reported, with two being ≥ Grade 3. None of the patients achieving a complete response (CR) experienced any NT and all cases of NT were observed in a
setting of disease progression and with confounding factors and minimal or undetectable CAR T cells in peripheral blood. No prophylactic measures of any
kind have been used to manage CRS or NT in this study. Two patients had death possibly related to AUTO3. One in the setting of disease progression and
multiorgan failure and other due to infection in a patient with secondary hemophagocytic lymphohistiocytosis.

6

The  majority  of  patients  receiving  AUTO3  in  the  outpatient  setting  did  not  require  hospital  admission.  Those  patients  admitted  were  managed,
without requiring ICU care. Combined with the overall favorable tolerability data across the Phase 1 trial, the profile of AUTO3 supports administration in
an outpatient setting.

Development Strategy for Adult DLBCL

We  have  completed  enrollment  of  patients  in  the  Phase  1  part  of  this  trial  and  plan  to  seek  a  partner  for  the  before  committing  to  further

development of this program.

AUTO4: Our Programmed T-Cell Lymphoma Program

Introduction to AUTO4

We are developing a programmed T cell product candidate, AUTO4, as a potential treatment for T-cell lymphomas. We are developing this product
candidate with a unique targeting approach that is designed to avoid the severe immunosuppression typically associated with the current investigational
CAR T-cell therapies which uses a pan t-cell antigen. for this disease.

T cells have one of two functionally identical genes, known as TRBC1 and TRBC2. A normal/healthy T cell population contains a mix of cells
expressing  either  TRBC1  or  TRBC2.  Both  forms  are  active  and  provide  the  body  with  natural  immunity,  including  antiviral  immunity.  Because  T-cell
lymphomas are clonal tumors that develop from a single T cell, they are either entirely TRBC1-positive or entirely TRBC2-positive. Currently available
products for the treatment of T-cell lymphoma indiscriminately target all T cells, leading to the severe immunosuppression associated with these treatments.

We have designed AUTO4 as a programmed T cell to specifically target and deplete cells expressing TRBC1, while preserving healthy T cells that
express TRBC2. A normal T cell population consists of varying amounts of TRBC1-positive and TRBC2-positive T cells. Based on the typical distribution
of  TRBC1-positive  and  TRBC2-positive  T  cells,  we  believe  that  patients  treated  with  AUTO4  should  be  left  with  a  population  of  healthy,  functional
polyclonal  T  cells,  which  provides  the  immune  system  of  these  patients  the  ability  to  respond to  bacterial  and  viral  infections  and  other  pathogens.  In
addition, this product candidate will have a built-in safety switch designed to eliminate the programmed CAR T cells in the event a patient suffers certain
serious adverse events related to the CAR T cell therapy, such as CRS or neurotoxicity.

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Companion Diagnostic for AUTO4

We  are  developing  a  proprietary  diagnostic  test  to  distinguish  the  T-cell  lymphoma  which  express  the  clonal  TRBC1-positive  T  cells  versus  the
clonal TRBC2-positive T cells. When a patient presents with T-cell lymphoma, this diagnostic is designed to test the patient’s tumor to assess whether the
tumor is TRBC1-positive or TRBC2-positive, which will determine whether the patient is potentially a candidate to receive AUTO4.

Background of T-Cell Lymphoma

Mature  T  cell  lymphomas  are  aggressive,  treatment  resistant  malignancies  that  are  associated  with  poor  prognosis.  Clinical  application  of
immunotherapeutic approaches has been limited by a lack of target antigens that discriminate malignant from healthy/polyclonal T cells. T-cell lymphoma
is a rare and heterogeneous form of NHL, representing approximately 10 to 20% of NHL cases and 3 to 4% of all hematological malignancies. Most T-cell
lymphomas  are  peripheral  T-cell  lymphomas,  (PTCL),  the  initial  indication  for  which  we  are  developing  AUTO4.  We  estimate  that  PTCL  affects
approximately 2,900 patients in the United States each year. PTCL generally involves high-grade tumors and occurs at a similar age as aggressive B cell
lymphomas, with a relatively high proportion of patients becoming rapidly unwell. For the majority the PTCL subtypes, the five-year survival rate may
range from 18% to 24%. The three most common subtypes of PTCL are peripheral T-cell lymphoma not otherwise specified, or PTCL-NOS, anaplastic
large-cell  lymphoma,  or  ALCL,  and  angioimmunoblastic  T-cell  lymphoma,  or  AITL,  together  accounting  for  approximately  70%  of  all  PTCLs  in  the
United States.

The first-line treatment for PTCL consists of the combination chemotherapy (e.g CHOP, consisting of cyclophosphamide, vincristine, doxorubicin
and  prednisolone).  However,  CHOP  chemotherapy,  complete  response  rates  are  low  and  disease  relapse  is  common.  In  many  treatment  centers,  CHOP
chemotherapy may be consolidated with autologous or allogenic stem cell transplantation in selected patients.

Little  is  understood  in  terms  of  treatment  guidance  for  the  other  PTCL  subtypes  and  these  lymphomas  lack  clear  treatment  guidelines.  A  large
proportion  of  T-cell  lymphoma  patients  are  refractory  to  or  relapse  following  treatment  with  standard  therapies  and  there  remains  a  need  to  develop  an
effective therapy for this currently unmet medical need.

Unlike B cell lymphomas, T-cell lymphomas have not benefited from advances in immunotherapeutic approaches. This is mainly due to the lack of
therapeutic development in T-cell lymphomas to identify suitable target antigens to distinguish malignant T cells from normal/polyclonal T cells. While a
similar problem exists with B cell lymphomas, targeting a pan B cell antigen is an acceptable strategy, as the concomitant depletion of the normal B cell
compartment is well tolerated, and some targeted approaches may be ameliorated by the administration of immunoglobulin. In contrast, targeting a pan T
cell antigen would result in severe immunosuppression, where there is currently no available rescue medication. Some competitors that are pursuing this
approach are planning to use CAR T-cells therapy as a bridging to SCT. However, this approach would only benefit the transplant eligible patients who
may  not  be  the  majority  of  the  T-cell  lymphoma  patients.  There  is  currently  no  programmed  T  cell  therapy  that  is  being  developed  as  a  standalone
treatment.

Clinical Development of AUTO4

Because  AUTO4  represent  a  novel  approach  to  treating  T-cell  lymphomas,  our  development  strategy  for  these  product  candidates  is  based  on

initially commencing a Phase 1/2 clinical trial of AUTO4 for the treatment of TRBC1-positive T-cell lymphoma.

Phase 1/2 Clinical Trial of AUTO4

In the fourth quarter of 2018, we began enrolling patients in a single-arm, open label, multi-center Phase 1/2 clinical trial, Libra T1, in patients with
TRBC1  positive  PTCL-NOS,  AITL  and  ALCL,  the  three  most  common  subtypes  of  PTCL,  for  which  patients  have  failed,  or  have  relapsed  disease
following, at least one prior therapy. We have received approval from the Medicines and Healthcare products Regulatory Agency (2018) and Spanish health
authority (2020), to begin enrollment and we are in the process of enrolling patients. We refer to this trial as the LibrA-T1 trial, which will initially be
conducted at sites in the United Kingdom and Spain. Patients are screened for TRBC status of tumor cells using a CE-marked next-generation sequencing
(NGS) method prior to full enrollment in the trial. Provided that data from the initial patients in the UK and Spain in the trial is satisfactory, we intend to
submit an IND and initiate additional sites for this trial in the United States.

The main objective of the Phase 1 portion of the trial is to evaluate the safety of AUTO4 and to determine a recommended dose for the Phase 2
portion of the trial. The main objective of the Phase 2 portion will be to further evaluate the safety of the treatment and evaluate efficacy endpoints, such as
overall response rate and complete response rate.

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We have designed the trial to evaluate up to three dose levels of AUTO4, beginning with a low dose of 25 million AUTO4 cells in cohorts of three
to six patients. Assuming that we do not observe any DLT, the dose escalation phase of the trial will continue to higher doses of 75 million AUTO4 cells
and 225 million AUTO4 cells/. We expect that we will enroll a total of up to 25 patients in the Phase 1 portion of the trial. Once a recommended phase 2
dose has been identified in the Phase 1 portion of the trial, we intend to treat up to 30 patients in the Phase 2 portion of the trial. In parallel with clinical
development, the companion diagnostic will be developed to enable use within a potential pivotal trial and commercialization.

Development Strategy for AUTO4

Based on our expected enrollment rates for the trial, we anticipate generating clinical proof of concept data in 2021. If the preliminary efficacy data
is positive, we intend to convert the Phase 2 portion to a potential registrational trial, subject to discussions with regulatory authorities. The final number of
patients  to  be  enrolled  in  the  trial,  specific  endpoints  and  other  aspects  of  the  design  of  the  trial  will  be  determined  based  on  feedback  from  regulatory
authorities. If the safety and efficacy data from the Phase 2 portion of the trial are positive, we plan to submit a BLA forAUTO4 as a second-line therapy
for TRBC1-positive T-cell lymphoma patients.

Our Solid Tumor Programs

Solid  tumors  present  a  particular  challenge  to  CAR  T  cell  therapies,  since  solid  tumors  tend  to  fend  off  T  cells  with  upregulation  of  checkpoint
inhibition  and  a  hostile  microenvironment.  In  addition,  contrary  to  hematological  cancer  cells  that  are  readily  accessible  to  programmed  T  cells  in  the
circulating blood of a patient, solid tumors are more difficult for programmed T cells to track down in sufficient numbers to impact the disease. In addition,
the persistence of programmed T cells tends to be limited, which also leads to a reduced effect on solid tumor cells. In addition to the programs we are
currently pursuing described below, we intend to continue to evaluate other possible solid tumor indications.

AUTO6: Our Neuroblastoma Program

Introduction to AUTO6 and AUTO6NG

We have been granted an exclusive, worldwide license under our license agreement with UCLB to AUTO6 (1RG-CART), a programmed T cell
product candidate targeting the glycosphingolipid GD2. CRUK has completed an exploratory Phase 1 clinical trial of AUTO6 in pediatric patients with
neuroblastoma. We are developing a next-generation product candidate, which we refer to as AUTO6NG, incorporating additional programming modules
designed to improve efficacy, safety and persistence of AUTO6. We expect to initiate two Phase 1/2 clinical trials of AUTO6NG, with the first clinical trial
expected to commence in 2021.

Background of Neuroblastoma

Neuroblastoma is a cancer that develops from immature nerve cells found in several areas of the body, and most commonly arises in and around the
adrenal glands, which have similar origins to nerve cells and sit atop the kidneys. However, neuroblastoma can also develop in other areas of the abdomen
and in the chest, neck and near the spine, where groups of nerve cells exist. Neuroblastoma most commonly affects children age five or younger, though it
may rarely occur in older children. According to the American Cancer Society, there are approximately 700 new cases of neuroblastoma each year in the
United States.

Preclinical Studies of AUTO6

In  preclinical  in vitro  studies,  AUTO6  selectively,  effectively  and  efficiently  killed  GD2-expressing  tumor  cells  while  sparing  cells  that  did  not
express GD2. In addition, the RQR8 safety switch activation by rituximab was tested in vitro, where the addition of rituximab was shown to activate the
safety switch and eliminate the programmed T cells from the culture, and residual cells did not possess any intrinsic anti-GD2 activity. This safety switch
activation was also observed in vivo in a mouse model, where the murine analogue of rituximab was able to deplete the GD2-targeting programmed T cell
product candidate from the bone marrow, blood, lymph node and spleen of animals that had previously been engrafted with programmed T cells.

In November 2019, we reported preclinical data of AUTO6NG. AUTO6 had previously shown clinical responses in two of our patients treated at
the  highest  dose  without  inducing  neurotoxicity  in  pediatric  patients  with  relapsed/refractory  neuroblastoma.  Building  on  AUTO6,  in  AUTO6NG  we
introduced  additional  programming  modules  in  order  to  help  the  programmed  T  cells  persist  in  and  withstand  the  hostile  tumor  microenvironment.
AUTO6NG is a programmed T cell therapy incorporating the GD2-targeted CAR T and RQR8 safety switch from AUTO6 but also incorporating three
additional  programming  modules:  (i)  an  IL7  chimeric  cytokine  receptor  designed  to  increase  persistence,  (ii)  a  dominant  negative  TGFbRII  protein
designed  to  block  inhibitor  signals  from  TGFb  and  (iii)  a  truncated  SHP2  protein  designed  to  block  inhibitor  signals  from  PD1.  These  modules  are
delivered, or transduced, into the T cells via two viral vectors. Both single- and dual-transduced CAR T cells were evaluated in vitro for anti-tumor

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activity,  cytokine  secretion,  T  cell  proliferation,  survival,  and  resistance  to  immunosuppressive  pathways.  The  addition  of  these  three  modules  in  the
AUTO6NG product candidate significantly augmented its function by extending T cell persistence and rendering modified T cells resistant to TGFb- and
PD1/PDL1-driven immune inhibition when compared to AUTO6 in vitro. Additionally, intravenous delivery of AUTO6NG in mice with established tumor
burden exhibited potent anti-tumor activity and extended survival, whereas AUTO6 showed no activity in that model.

We presented new preclinical data for AUTO6NG in June 2020 at the American Association for Cancer Research (AACR) Virtual Annual Meeting.
GD2  was  evaluated  as  a  therapeutic  CAR-T  target  antigen  in  Small  Cell  Lung  Cancer  (SCLC).  AUTO6  alone  has  demonstrated  efficacy  in  an  in vitro
SCLC model, but successful tumor targeting alone was not sufficient to drive meaningful in vivo efficacy in the same SCLC model. Autolus has presented
new preclinical data demonstrating the ability to target GD2 in SCLC cell line models in vitro, and the requirement for enhancing modules, designed to
overcome TME suppressive mechanisms, to drive superior in vivo efficacy in a SCLC mouse model. The data suggests that AUTO6NG can overcome the
immune suppressive mechanisms in the TME

We believe these data support the continued development of AUTO6NG, and we expect to initiate the first of two planned Phase 1/2 clinical trials

of AUTO6NG in the first half of 2021.

Phase 1 Dose Escalation Trial of AUTO6 by CRUK in Relapsed or Refractory Neuroblastoma

In the first quarter of 2016, CRUK initiated a single-arm Phase 1 dose escalation trial of AUTO6 in relapsed or refractory neuroblastoma at two
pediatric cancer centers in the United Kingdom. The trial evaluated the safety profile of AUTO6. The Phase 1 trial is also evaluating escalating intensity of
the pre-conditioning regimen along with AUTO6 dose escalation. CRUK plans to enroll 15 to 27 patients in this trial.

As presented at the Annual Meeting of the American Association for Cancer Research in April 2018, twelve patients with relapsed or refractory
neuroblastoma with measurable disease in bone (n=11), bone marrow (n=7) or soft tissue sites (n=9) have been enrolled in the trial. Ten patients have been
treated, with the first six patients receiving a dose of 10 million AUTO6 cells/m , four without preconditioning, one with cyclophosphamide alone and one
with a combination of cyclophosphamide and fludarabine, or cy/flu. A further three patients were treated with a dose of 100 million AUTO6 cells/m  with
cy/flu preconditioning. In the next cohort, a further patient was treated with a dose of one billion AUTO6 cells/m  with cy/flu preconditioning. The trial is
2
continuing to enroll patients at the dose of one billion AUTO6 cells/m .

2

2

2

Updated findings published in Science Translational Medicine suggest that AUTO6 can induce rapid regression of bulky disease in a solid tumor
setting without inducing on-target off-tumor toxicity, despite dose dependent CAR T expansion. CAR T cell expansion has been observed in all six patients
treated at the higher cell dose cohorts in this Phase 1 study. Three of these six patients demonstrated evidence of transient CAR T cell activity, including
cytokine release syndrome, and regression of soft tissue and bone marrow disease activity. The GD2 binder used in AUTO6 has been designed to minimize
on-target  off-tumor  neurotoxicity  associated  with  GD2  expression  at  low  levels  in  pain  fibers  and  the  brain.  Despite  the  presence  of  clear  CAR  T  cell
activity, no neurotoxicity was observed. The publication also suggests that, whilst AUTO6 is a valid and safe strategy for targeting neuroblastoma, further
modifications are required to promote CAR T cell persistence and induce deeper and more durable responses for these patients.

Clinical Development Strategy of AUTO6NG

    Based on preliminary data from proof-of-concept in CRUK’s ongoing Phase 1 clinical trial, we believe it is possible to safely target GD2-expressing
cancers or tumors with a CAR. We are currently developing a next-generation T cell product candidate, which we refer to as AUTO6NG, which builds on
AUTO6 by incorporating additional programming modules intended to enhance the efficacy, safety and persistence of AUTO6.

Because  GD2  is  expressed  in  numerous  pediatric  and  adult  tumors  including  neuroblastoma,  osteosarcoma,  soft  tissue  sarcoma,  melanoma,
astrocytoma and small cell lung cancer, or SCLC, our clinical development strategy is to develop AUTO6NG in parallel in neuroblastoma and in additional
indications. To that end, we are planning to initiate two Phase 1/2 clinical trials of AUTO6NG. The first trial will be in adult patients and is expected to
commence in the first half of 2021, and the second trial will be in pediatric patients and is expected to commence in the second half of 2021.

In  the  first  planned  Phase  1/2  clinical  trial,  we  plan  to  enroll  pediatric  patients  with  relapsed  or  refractory  neuroblastoma  and  osteosarcoma.
Osteosarcoma is the most common type of bone cancer in children and teens, with approximately 800 to 900 new cases diagnosed each year in the United
States, the majority of which will be GD2 positive. Following evaluation of safety and selection of the recommended Phase 2 dose, we plan to initiate the
three-arm Phase 2 portion of the clinical trial, which will enroll patients with neuroblastoma, osteosarcoma and other GD2-positive tumors, respectively, in
each individual arm of the trial. If the preliminary efficacy data from the Phase 2 portion of the trial based on appropriate criteria for individual tumor types
is positive in one or more arms, we intend to discuss with the FDA the possibility of converting the Phase 2 portion into a registrational trial, with separate
arms

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for  each  indication.  The  final  number  of  patients  to  be  enrolled  in  the  trial  and  endpoints  for  each  individual  indication  will  be  determined  based  on
feedback from regulatory authorities.

In our second planned Phase 1/2 trial, we intend to enroll adult patients with metastatic melanoma, SCLC and other GD2-positive malignancies and
who have received at least one prior therapy. The timing of this trial will be staggered with the first Phase 1/2 clinical trial in order to incorporate learnings
from the early dose cohorts of pediatric patients. We anticipate that prior to initiation of this second Phase 1/2 clinical trial, a diagnostic assay for GD2
assessment may be needed. Melanoma is one of the most common types of cancer, with approximately 90,000 new cases diagnosed each year in the United
States. SCLC accounts for about 10-15% of all lung cancer cases, with 30,000 new cases diagnosed each year in the United States. It has been reported that
approximately half of the patients are positive for the GD2 antigen. Following selection of the recommended Phase 2 dose, we plan to initiate the three-arm
Phase 2 portion of the clinical trial, which will enroll patients with melanoma, SCLC and other GD2-positive tumors, respectively, in each individual arm
of the trial. If the preliminary efficacy data such as overall response rate from the Phase 2 portion of the trial is promising in one or more arms, we intend to
discuss with the FDA the possibility of converting the Phase 2 portion into a registrational trial, with separate arms for each indication. The final number of
patients to be enrolled in the trial and endpoints for each individual indication will be determined based on feedback from regulatory authorities.

AUTO7—Our Prostate Cancer Program

We are in preclinical development of AUTO7, a programmed T cell product candidate designed to target and treat prostate cancer. According to the
American Cancer Society, other than skin cancer, prostate cancer is the most common cancer in American men, with approximately 165,000 new cases
diagnosed each year. This program incorporates enhanced safety modules including our small molecule mediated safety switch and enhanced T cell activity
modules that we are developing to overcome the immuno-suppressive effects of the tumor micro-environment and enhance T cell persistence. We have
incorporated a technology in AUTO7 that is designed to deliver a cytokine signal directly inside our programmed T cells. This cytokine persistence signal
is further enhanced by engagement with antigens secreted by the tumor.

We presented new preclinical data for AUTO7 in June 2020 at the American Association for Cancer Research (AACR) Virtual Annual Meeting.
The preclinical data demonstrate that AUTO7 is highly potent in cytotoxicity assays against cells expressing PSMA, even at low levels, and demonstrate
the feasibility of this multi-modular cell programming approach in overcoming the immunotherapeutic challenges presented by advanced prostate cancer,
which is typically otherwise an immunologically cold tumor.

We intend to initiate our clinical development of AUTO7 in H1 2022. We anticipate starting a Phase 1/2 trial in patients with metastatic castration-
resistant prostate cancer to evaluate the safety and identify the optimum Phase 2 dose of AUTO7 in the Phase 1 part of the trial and preliminary efficacy in
the Phase 2 portion of the trial.

Partnerable Coronavirus Disease (COVID-19) Project

Our research team has developed a potentially universal SARS-CoV2 decoy receptor with virus neutralizing activity against SARS-CoV2 and its

variants, and also active against SARS-CoV1.

Manufacture and Delivery of Programmed T Cell Therapies to Patients

We  are  devoting  significant  resources  to  process  development  and  manufacturing  in  order  to  optimize  the  safety  and  efficacy  of  our  product
candidates, as well as to reduce our per unit manufacturing costs and time to market if we obtain regulatory approval for any of our programmed T cell
product candidates.

The manufacture and delivery of programmed T cell therapies to patients involves complex, integrated processes, including harvesting T cells from
patients, manufacturing viral vectors with nucleic acid content encoded with our programming modules, manufacturing programmed T cells using the viral
vectors ex vivo, multiplying the T cells to obtain the desired dose, and ultimately infusing the T cells back into a patient’s body.

Commercial  success  in  T  cell  therapies  requires  a  manufacturing  process  that  is  reliable,  scalable  and  economical.  We  have  established  a
manufacturing process that is scalable and serves as a manufacturing platform designed to support rapid development of our programmed T cell therapy
product  candidates  through  clinical  trial  phases  and  regulatory  approval  processes.  We  are  using  a  semi-automated,  fully  enclosed  system  for  cell
manufacturing, which is designed to provide a common platform suitable for manufacturing all of our product candidates. This platform allows for parallel
processing  having  the  ability  to  scale  for  commercial  supply  in  a  controlled  environment  at  an  economical  cost.  We  have  improved  our  viral  vector
production and viral transduction process to reduce process variability.

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Our manufacturing and logistics process is designed to ensure that product integrity is maintained during shipment along with accurate tracking and
tracing of shipments. We are expanding internal manufacturing and supply capabilities as well as the use of expert service providers on maturing our vein-
to-vein logistics and our gradual capacity expansion.

Our manufacturing and commercialization strategy requires a fully integrated vein-to-vein product delivery cycle. We believe having established
manufacturing processes suitable for commercialization early in the development of our T cell therapies will allow us to focus on expanding manufacturing
capacity during our clinical trials and early commercial launch needs. Over time, we expect to establish regional manufacturing hubs to meet projected
product  requirements  for  commercialization.  We  believe  that  anticipated  future  commercial  requirements  can  be  met.  Our  plan  is  to  establish  our
manufacturing infrastructure in a manner that would ameliorate logistics complexities and costs for all regions on initial commercial launch.

We believe our scalable closed-system manufacturing process, along with our proprietary and modular T cell programming technologies, would be

challenging and costly for potential competitors to replicate.
Manufacturing Agreements

We have manufacturing agreements with King’s College London for early phase vector manufacturing. Autolus also has an internal capability to
produce vector for early and late-stage trials. Additionally, we have also established an agreement with AGC Bio for late stage clinical and commercial
supply  of  vector.  All  vector  manufacturing  is  done  in  accordance  with  Current  Good  Manufacturing  Practice,  or  cGMP,  in  compliant  manufacturing
facilities.  The  manufacturing  agreements  governing  the  external  supply  arrangements  also  provide  for  access  to  services  including  quality  management
systems, qualified persons for product release, office space, frozen storage and warehousing services.

We have established our cell and vector manufacturing capacity at the Cell and Gene Therapy Catapult in Stevenage, United Kingdom. We have 2
cell manufacturing suites capable of supporting clinical and early stage commercial launch. We also have a vector production suite capable of supplying
clinical supplies. The Cell and Gene Therapy Catapult provides for access to architecturally and operationally segregated manufacturing suites. This facility
approach has been reviewed and approved by the MHRA. This operational approach has also been reviewed with the FDA.

In  March  2018,  we  entered  into  a  strategic,  long-term  supply  agreement  with  Miltenyi  Biotec  GmbH,  or  Miltenyi,  for  the  supply  of  Miltenyi’s
CliniMACS Prodigy instruments, reagents and disposables for the manufacture of our programmed T cell therapies for preclinical and clinical use and, if
approved, for commercial use, as well as support services. The supply agreement sets forth procedures to ensure continuity of supply to us of Miltenyi’s
products, both during the clinical phase and any future commercial phase of our product candidates. After the initial ten-year term of the agreement, we
have two separate options to renew the agreement, each for an additional five-year term. The supply agreement contains customary termination provisions,
allowing for termination by a party upon the other party’s uncured material breach, upon the other party’s bankruptcy or insolvency or upon the other party
being subject to an extended period of force majeure events. We may also terminate the supply agreement upon advance written notice, if we decide to
suspend or discontinue the development or commercialization of our product candidates. The supply agreement is governed under the laws of Germany.

Commercialization

Given  our  stage  of  development,  we  have  not  yet  established  a  commercial  infrastructure  or  distribution  capabilities.  We  are  developing  our
clinical-stage programs for the treatment of patients with late-stage or rare hematological cancers and solid tumors, most of whom are treated in specialized
treatment centers or hospitals. With our experience in gene therapy, transplantation and oncology, we aim to provide high levels of service and scientific
engagement at these treatment centers, and to pilot and establish systems necessary for product delivery by the time of launch. We believe this approach
will require less investment in commercial infrastructure compared to the current standard of care. By focusing on these centers, we can begin to build our
commercialization capabilities with limited resources.

We have retained worldwide commercial rights for our product candidates. We currently plan to build our global commercialization capabilities
internally over time such that we are able to commercialize any product candidate for which we may obtain regulatory approval. We may selectively pursue
strategic collaborations with third parties in order to maximize the commercial potential of our product candidates. We generally expect to launch any of
our  products  that  receive  regulatory  approval  in  the  United  States  first,  followed  by  the  European  Union  and  subsequently  in  other  major  markets.  For
AUTO1, we may look to commercialize first in markets outside the United States, if any, where we receive regulatory approvals, with a launch following
regulatory approval in the United States occurring after the earlier of either the expected expiration of any applicable third-party patents covering AUTO1
that expire between 2023 and 2025, the invalidation of such patents or the receipt of a license to such patents on commercially reasonable terms. See “Risk
Factors–Risks Related to our Intellectual Property–Third parties may initiate legal proceedings alleging that we are infringing their intellectual property
rights, the outcome of which would be uncertain and could significantly harm our business.”

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Intellectual Property

Intellectual  property  is  of  vital  importance  in  our  field  and  in  biotechnology  generally.  We  seek  to  protect  and  enhance  proprietary  technology,
inventions  and  improvements  that  are  commercially  important  to  the  development  of  our  business  by  seeking,  maintaining  and  defending  patent  rights,
whether developed internally or licensed from third parties. We will also seek to rely on regulatory protection afforded through orphan drug designations,
data exclusivity, market exclusivity and patent term extensions where available.

Our  intellectual  property  estate,  which  includes  in-licensed  intellectual  property  and  intellectual  property  that  we  own,  is  designed  to  provide
multiple  layers  of  protection.  For  example,  we  are  pursuing  patent  protection  for  core  constructs  used  in  our  product  candidates,  various  methods  of
treatment for particular therapeutic indications using our approach, specific product candidates, innovative manufacturing processes, and constructs that
may be used in future product candidates to improve the ability of our programmed T cells to better recognize and kill cancer cells. A portion of our patent
portfolio  is  directed  to  certain  current  product  candidates  or  technologies  deployed  in  certain  product  candidates,  and  the  remainder  of  the  portfolio  is
directed to alternative approaches, technologies or modules that are not currently deployed in our current product candidates.

As  of  December  31,  2020,  our  patent  portfolio  is  comprised  of  96  patent  families,  of  which  25  patent  families  originated  from  UCLB,  the  technology-
transfer company of UCL, 3 patent families are in-licensed from Noile-Immune Biotech, Inc., and 68 patent families we own and have originated from our
own research. Of the 25 patent families that were originally in-licensed from UCLB, 24 have now been assigned to Autolus under a Deed of Assignment
dated 15 October 2020. We believe that our current patent portfolio, together with our ongoing efforts to develop and patent new technologies, will provide
us  with  substantial  intellectual  property  protection  for  our  product  candidates  and  other  technologies  that  are  not  currently  deployed  in  our  product
candidates.

Commercially or strategically important non-U.S. jurisdictions in which certain patent applications that we have in-licensed are currently pending
include: Europe, Australia, Canada, Japan, China, Brazil, Chile, Israel, India, Republic of Korea, Hong Kong, Mexico, New Zealand, Russian Federation,
Singapore, South Africa, Colombia, Peru, Cuba, Indonesia, Malaysia and Philippines.

Our  strategy  is  to  develop  and  obtain  additional  intellectual  property  covering  innovative  manufacturing  processes  and  methods  for  genetically
engineering  T  cells  expressing  new  constructs  with  properties  that  are  designed  to  improve  the  ability  of  our  programmed  T  cells  to  recognize  and  kill
cancer cells. To support this effort, we have established expertise and development capabilities focused in the areas of T cell programming, preclinical and
clinical  research  and  development,  and  manufacturing  and  manufacturing  process  scale-up,  and  we  expect  that  our  ongoing  research  and  development
activities will yield additional patentable inventions and patent applications that will expand our intellectual property portfolio.

The term of individual patents depends upon the legal term of the patents in the countries in which they are obtained. In most countries in which we
file, the patent term is 20 years from the date of filing of the first non-provisional application to which priority is claimed. In the United States, a patent’s
term may be lengthened by patent term adjustment, which compensates a patentee for administrative delays by the U.S. Patent and Trademark Office in
granting a patent or may be shortened if a patent is terminally disclaimed over an earlier-filed patent. The term of a patent that covers an FDA-approved
drug may also be eligible for a patent term restoration of up to five years under the Hatch-Waxman Act, which is designed to compensate for the patent
term lost during the FDA regulatory review process. The length of the patent term restoration is calculated based on the length of time the drug is under
regulatory review. A patent term restoration under the Hatch-Waxman Act cannot extend the remaining term of a patent beyond a total of 14 years from the
date of product approval and only one patent applicable to an approved drug may be restored. Moreover, a patent can only be restored once, and thus, if a
single patent is applicable to multiple products, it can only be extended based on one product. Similar provisions are available in Europe and certain other
foreign jurisdictions to extend the term of a patent that covers an approved drug. If and when possible, we expect to apply for patent term extensions for
patents covering our product candidates or their methods of use.

Our  commercial  success  may  depend  in  part  on  our  ability  to  obtain  and  maintain  patent  and  other  proprietary  protection  for  commercially
important technology, inventions and know-how related to our business, defend and enforce our patents, preserve the confidentiality of our trade secrets,
and operate without infringing the valid enforceable patents and proprietary rights of third parties. Our ability to stop third parties from making, using,
selling, offering to sell or importing our products may depend on the extent to which we have rights under valid and enforceable patents or trade secrets
that  cover  these  activities.  With  respect  to  both  licensed  and  company-owned  intellectual  property,  we  cannot  be  sure  that  patents  will  be  granted  with
respect to any of our pending patent applications or with respect to any patent applications filed by us in the future, nor can we be sure that any patents, if
granted, will be commercially useful in protecting our commercial products and methods of manufacturing the same. Development and commercialization
of products can be subject to substantial delays and it is possible that, at the time of commercialization, any patent covering the product has expired or will
be in force for only a short period of time following commercialization. Numerous third-party

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U.S. and non-U.S. issued patents exist in the area of programmed T cell therapies, including patents held by our competitors. We cannot predict with any
certainty if any third-party U.S. or foreign patent rights, or other proprietary rights, will be deemed infringed by the use of our technology. Nor can we
predict with certainty which, if any, of these rights will or may be asserted against us by third parties. Should we need to defend ourselves against any such
claims, substantial costs may be incurred. Furthermore, parties making such claims may be able to obtain injunctive or other equitable relief, which could
effectively block our ability to develop or commercialize some or all our products in the United States, European Union and other major markets.

We  may  rely,  in  some  circumstances,  on  trade  secrets  to  protect  our  technology.  However,  trade  secrets  can  be  difficult  to  protect.  We  seek  to
protect our proprietary technology and processes, in part, by entering into confidentiality agreements with our employees, consultants, scientific advisors
and contractors. We also seek to preserve the integrity and confidentiality of our data and trade secrets by maintaining physical security of our premises and
physical  and  electronic  security  of  our  information  technology  systems.  While  we  have  confidence  in  these  individuals,  organizations  and  systems,
agreements or security measures may be breached, and we may not have adequate remedies for any breach. In addition, our trade secrets may otherwise
become  known  or  be  independently  discovered  by  competitors.  To  the  extent  that  our  consultants,  contractors  or  collaborators  use  intellectual  property
owned by others in their work for us, disputes may arise as to the rights in related or resulting know-how and inventions.

Our License Agreement with UCL Business Ltd.

In September 2014, we entered into an exclusive license agreement with UCLB, the technology-transfer company of UCL, for the development and
commercialization rights to certain T cell programming modules. The license agreement was amended and restated in March 2016 to also include certain
development  and  commercialization  rights  to  improvements  and  new  T  cell  programming  modules.  The  license  agreement  was  further  amended  and
restated in March 2018 to include a license to AUTO1, for which UCL is conducting Phase 1 clinical trials in pediatric and adult ALL patients. The license
agreement was further amended and restated in October 2020 to reflect our election to have various patent rights assigned to us, and to include a license to
new technology and further licenses to AUTO1 for which UCL is conducting Phase 1 clinical trials in primary CNS Lymphoma patients. Under the license
agreement, subject to certain limitations, exceptions and retained rights of UCLB, we received an exclusive license of certain patent rights and know-how
owned by UCLB covering T cell programming modules. The licensed rights cover our AUTO1, AUTO3, AUTO4/5 and AUTO6 targeting modules, as well
as additional T cell programming modules and technologies, including dual-targeting technology, pattern recognition technology, safety switches (including
RQR8),  tunable  T  cells,  manufacturing  processes  as  well  as  certain  technology  for  evading  tumor  micro-environments.  We  also  have  option  rights  and
rights of first negotiation to obtain an exclusive license for development and commercialization rights to certain new T cell programming modules.

In  exchange  for  the  rights  under  the  original  license  agreement,  we  granted  UCLB  4,769,994  B  ordinary  shares  of  Autolus  Limited,  which,  in
connection with our corporate reorganization in June 2018, were converted to 1,497,643 ordinary shares of Autolus Therapeutics plc. We also agreed to pay
a management fee, milestone payments and royalties upon future net sales of any products that use the in-licensed rights. The management fee of £120,000
was payable in equal installments on the first four anniversaries of our entry into the original license agreement. In exchange for the additional rights we
received in March 2016 when the license agreement was amended, we issued UCLB an additional 1,000,000 B ordinary shares, which, in connection with
our  corporate  reorganization  in  June  2018,  were  converted  to  313,971  ordinary  shares  of  Autolus  Therapeutics  plc,  and  made  a  one-time  payment  of
£150,000. In exchange for the additional rights we received in March 2018 when the license agreement was further amended, we made an initial payment
of £1.5 million and paid the additional £0.35 million in connection with UCLB's transfer of clinical data to the Company in December 2020.

Under the license agreement, as amended, we are obligated to pay UCLB milestone payments upon the initiation of certain clinical activities in an
aggregate amount of £0.18 million, the receipt of specified regulatory approvals in an aggregate amount of £37.5 million, the start of commercialization in
an aggregate amount of £18 million, and the achievement of net sales levels in an aggregate amount of £51 million. On a per-product basis, these milestone
payments range from £1 million to £18.5 million, depending on which T cell programming modules are used in the product achieving the milestone. Under
the terms of the license, we have the right to grant sub-licenses to third parties, subject to certain restrictions. If we receive any income in connection with
such sublicenses, we must pay UCLB a percentage of the income allocable to the value of the sublicensed intellectual property rights ranging from low
twenties to mid-single digits, decreasing based on the development expenses incurred by us and the passage of time. UCLB has retained the right to use the
licensed T cell programming modules for academic research purposes at UCL and with other academic institutions, subject to certain restrictions.

Upon commercialization of any of our products that use the in-licensed patent rights, we are obligated to pay UCLB a flat royalty for each licensed
product ranging from the low- to mid-single digits, depending on which technologies are deployed in the licensed product, based on worldwide annual net
sales  of  each  licensed  product,  subject  to  certain  reductions,  including  for  the  market  entry  of  competing  products  and  for  loss  of  patent  coverage  of
licensed products. We may deduct from the royalties payable to UCLB half of any payments made to a third party to obtain a license to such third party’s
intellectual property that is necessary to exploit any licensed products. Once net sales of a licensed product have reached a certain specified threshold, we
may exercise an option to buy

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out UCLB’s rights to the remaining milestone payments, royalty payments, and sublicensing revenue payments for such licensed product, on terms to be
negotiated at the time.

As mentioned above, we acquired ownership of the majority of the licensed patent rights under the license agreement (with the exception of the
RQR8  patent  rights)  by  virtue  of  a  Deed  of  Assignment  from  UCLB  which  was  executed  on  15  October  2020.  Our  payment  and  diligence  obligations
remain unaffected by the assignment of the licensed patent rights to us.

Under  the  license  agreement,  we  are  solely  responsible,  at  our  expense,  for  developing  the  products  that  use  the  in-licensed  patent  rights  and
obtaining  all  regulatory  approvals  for  such  products  worldwide.  We  are  also  solely  responsible,  at  our  expense,  for  commercializing  the  products
worldwide after receiving regulatory approval. Further, we are obligated to use commercially reasonable efforts to develop certain products using the patent
rights  pertaining  to  the  T  cell  programming  modules  we  have  licensed  from  UCLB.  Failure  to  achieve  diligence  obligations  may  result  in  loss  of
exclusivity or termination of the license on a program-by-program basis.

The license agreement expires on a product-by-product and country-by-country basis upon the expiration of the royalty term with respect to each
product in each country. We may unilaterally terminate the license agreement for any reason upon advance notice to UCLB. Either party may terminate the
license agreement for the uncured material breach by the other party or for the insolvency of the other party. If UCLB terminates the license agreement
following our insolvency or our material breach of the agreement, or if we terminate the agreement unilaterally, all rights and licenses granted to us will
terminate,  and  all  patent  rights  and  know-how  transferred,  licensed  or  assigned  to  us  pursuant  to  the  agreement  will  revert  back  to  UCLB.  In  addition,
UCLB has the right to negotiate with us for the grant of an exclusive license to our improvements to the T cell programming modules we have licensed on
terms to be agreed upon at the time.

Competition

Presently, the biotechnology and pharmaceutical industries put significant resources in developing novel and proprietary therapies for the treatment
of  cancer.  While  we  believe  that  our  differentiated  product  candidates  and  scientific  expertise  in  the  field  of  cellular  immunotherapy  provide  us  with
competitive advantages, we face potential competition from various sources, including larger and better-funded pharmaceutical, specialty pharmaceutical
and biotechnology companies, as well as from academic institutions, governmental agencies and public and private research institutions. We anticipate that
we will face intense and increasing competition as new drugs and therapies enter the market and advanced technologies become available. Due to their
promising  clinical  therapeutic  effect  in  clinical  exploratory  trials,  advanced  T  cell  therapies,  redirected  T  cell  therapies  in  general  and  antibody-drug
conjugates  are  being  pursued  by  multiple  biotechnology  and  pharmaceutical  companies,  including  Novartis  AG,  or  Novartis,  Gilead  Sciences,  Inc.,  or
Gilead, Bristol-Myers Squibb, or BMS, Janssen Biotech Inc., bluebird bio, Inc., or bluebird bio, Roche Holding AG, Seattle Genetics, and Amgen Inc. Our
competitors may succeed in developing, acquiring or licensing technologies and products that are more effective, more effectively marketed and sold or
less costly than any product candidates that we may develop, which could render our product candidates non-competitive and obsolete.

Currently there are now four approved anti-CD19 CAR T cell therapies, including Tecartus from Kite/Gilead and Breynazi from BMS, which were
approved in February 2021.These companies and products will compete directly with AUTO3, our dual-targeting CD19/CD22 programmed T cell product
candidate and AUTO1, our CD19 targeting programmed T cell product candidate in some indications.

The BCMA space continues to be highly competitive with the first CAR T cell therapy in this indication anticipated to come to market in the near
future. Bluebird bio, Inc., in collaboration with BMS, is developing a BCMA CAR T cell therapy for the treatment of multiple myeloma. Nanjing Legend
Biotech  and  Janssen  Biotech,  Inc.,  a  subsidiary  of  Johnson  &  Johnson,  are  collaborating  on  the  development  of  a  similar  therapy.  In  addition,  some
companies, such as Gilead, BMS and Poseida Therapeutics Inc. are also developing BCMA CAR T cell therapies for the treatment of multiple myeloma.
The BCMA landscape continues to evolve, with BMS deciding to abandon a second BCMA CAR-T candidate in February 2021. Some companies like
Amgen, BMS and Genentech, Inc., a member of the Roche Group, are developing BCMA-targeting T cell engagers for the treatment of multiple myeloma,
which are expected to compete directly with CAR-T approaches. All of these therapies will compete directly with our AUTO8 product candidate.

While we believe that other known types of immunotherapies may potentially be used in conjunction with CAR T cell therapies, such as checkpoint
inhibitors, to enhance efficacy, we do not currently expect substantial direct competition from these other types of immunotherapies. However, we cannot
predict whether other types of immunotherapies may be enhanced and show greater efficacy, and we may have direct and substantial competition from such
immunotherapies in the future.

In  addition,  more  effective  small  molecules,  cancer  vaccines  and  other  approaches  may  be  developed  and  used  as  first  line  or  second  line

treatments, which would reduce the market opportunity for our programmed T cell therapies.

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Many of our competitors, either alone or with their strategic collaborators, have substantially greater financial, technical and human resources than
we do. Accordingly, our competitors may be more successful than we are in obtaining approval for treatments and achieving widespread market acceptance
and  may  render  our  treatments  obsolete  or  non-competitive.  Mergers  and  acquisitions  in  the  biotechnology  and  pharmaceutical  industries  may  result  in
even more resources being concentrated among a smaller number of our competitors. These competitors also compete with us in recruiting and retaining
qualified  scientific  and  management  personnel  and  establishing  clinical  study  sites  and  patient  registration  for  clinical  studies,  as  well  as  in  acquiring
technologies complementary to, or necessary for, our programs. Smaller or early-stage companies may also prove to be significant competitors, particularly
through collaborative arrangements with large and established companies.

We  anticipate  that  we  will  face  intense  and  increasing  competition  as  new  products  and  therapies  enter  the  market  and  advanced  technologies
become available. We expect any treatments that we develop and commercialize to compete on the basis of, among other things, efficacy, safety, delivery,
price and the availability of reimbursement from government and other third-party payors.

Our commercial opportunity could be reduced or eliminated if our competitors develop and commercialize products that are safer, more effective,
have fewer or less severe side effects, are more convenient or are less expensive or better reimbursed than any products that we may commercialize. Our
competitors also may obtain FDA, EMA or other regulatory approval for their products more rapidly than we may obtain approval for ours, which could
result in our competitors establishing a strong market position for either the product or a specific indication before we are able to enter the market.

Government Regulation and Product Approval

As a biopharmaceutical company, we are subject to extensive regulation. Our programmed T cell product candidates, if approved, will be regulated
as  biological  medicines.  With  this  classification,  commercial  production  of  our  products  will  need  to  occur  in  registered  and  licensed  facilities  in
compliance with current Good Manufacturing Practices, or cGMPs, for biologics.

Human immunotherapy products are a new category of therapeutics. The FDA categorizes human cell- or tissue-based products as either minimally
manipulated or more than minimally manipulated and has determined that more than minimally manipulated products require clinical trials to demonstrate
product safety and efficacy and the submission of a Biologics License Application, or BLA, for marketing authorization.

Government authorities in the United States (at the federal, state and local level) and in other countries and jurisdictions, including the European
Union,  extensively  regulate,  among  other  things,  the  research,  development,  preclinical  and  clinical  testing,  manufacturing,  quality  control,  labeling,
packaging, storage, record-keeping, promotion, advertising, sale, distribution, post-approval monitoring and reporting, marketing and export and import of
biopharmaceutical products such as those we are developing. Our product candidates must be approved by the FDA before they may be legally marketed in
the United States and by the appropriate foreign regulatory agency before they may be legally marketed in foreign countries. Generally, our activities in
other  countries  will  be  subject  to  regulation  that  is  similar  in  nature  and  scope  as  that  imposed  in  the  United  States,  although  there  can  be  important
differences.  Additionally,  some  significant  aspects  of  regulation  in  Europe  are  addressed  in  a  centralized  way,  but  country-specific  regulation  remains
essential in many respects. The process for obtaining regulatory marketing approvals and the subsequent compliance with applicable federal, state, local
and foreign statutes and regulations require the expenditure of substantial time and financial resources.

U.S. Product Development Process

In  the  United  States,  the  FDA  regulates  biological  products  under  the  Public  Health  Service  Act,  or  PHSA,  and  the  Federal  Food,  Drug  and
Cosmetic Act, or FDCA, and implementing regulations. Products are also subject to other federal, state and local statutes and regulations. The process of
obtaining  regulatory  approvals  and  the  subsequent  compliance  with  appropriate  federal,  state,  local  and  foreign  statutes  and  regulations  require  the
expenditure  of  substantial  time  and  financial  resources.  Failure  to  comply  with  the  applicable  U.S.  requirements  at  any  time  during  the  product
development process, approval process or after approval, may subject an applicant to administrative or judicial sanctions. FDA sanctions could include,
among other actions, refusal to approve pending applications, withdrawal of an approval, a clinical hold, warning letters and similar public notice of alleged
non-compliance with laws, product recalls or withdrawals from the market, product seizures, total or partial suspension of production or distribution, fines,
refusals of government contracts, restitution, disgorgement of profits, or civil or criminal penalties. Any agency or judicial enforcement action could have a
material adverse effect on us. The process required by the FDA before a biological product may be approved for marketing in the United States generally
involves the following:

•

completion of preclinical laboratory tests and animal studies according to Good Laboratory Practices, or GLPs, and applicable requirements for
the humane use of laboratory animals or other applicable regulations;

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•

•

•

•
•
•

submission to the FDA of an Investigational New Drug Application, or IND, which must become effective before human clinical trials may begin;
performance  of  adequate  and  well-controlled  human  clinical  trials  according  to  the  FDA’s  regulations  commonly  referred  to  as  Good  Clinical
Practices, or GCPs, and any additional requirements for the protection of human research subjects and their health information, to establish the
safety and efficacy of the proposed biological product for its intended use;
preparation and submission to the FDA of a Biologics License Application, or BLA, for marketing approval that includes substantive evidence of
safety, purity, and potency from results of nonclinical testing and clinical trials;
satisfactory  completion  of  one  or  more  FDA  inspections  of  the  manufacturing  facility  or  facilities  where  the  biological  product  is  produced  to
assess  compliance  with  cGMP  to  assure  that  the  facilities,  methods  and  controls  used  in  product  manufacture  are  adequate  to  preserve  the
biological  product’s  identity,  strength,  quality  and  purity  and,  if  applicable,  the  FDA’s  current  Good  Tissue  Practices,  or  GTPs,  for  the  use  of
human cellular and tissue products;
potential FDA audit of the nonclinical study and clinical trial sites that generated the data in support of the BLA;
payment of user fees for FDA review of the BLA; and
FDA  acceptance,  review  and  approval,  or  licensure,  of  the  BLA,  which  might  include  review  by  an  advisory  committee,  a  panel  typically
consisting of independent clinicians and other experts who provide recommendations as to whether the application should be approved and under
what conditions.

Before  testing  any  biological  product  candidate,  including  our  product  candidates,  in  humans,  the  product  candidate  must  undergo  rigorous  the
preclinical testing. Preclinical tests, also referred to as nonclinical studies, include laboratory evaluations as well as in vitro and animal studies to assess the
potential  safety  and  efficacy  of  the  product  candidate.  After  sufficient  preclinical  testing  has  been  conducted,  the  conduct  of  the  preclinical  tests  must
comply with federal regulations and requirements including GLPs. The clinical trial sponsor must submit an IND to the FDA before clinical testing can
begin in the United States. An IND must contain the results of the preclinical tests, manufacturing information, analytical data, any available clinical data
or literature, a proposed clinical protocol, an investigator’s brochure, a sample informed consent form, and other materials. Clinical trial protocols detail,
among other things, the objectives of the clinical trial, dosing procedures, subject selection and exclusion criteria, and the parameters to be used to monitor
subject safety, including stopping rules that assure a clinical trial will be stopped if certain adverse events should occur. Each protocol and any amendments
to the protocol must be submitted to the FDA as part of the IND. Some preclinical testing, such as toxicity studies, may continue even after the IND is
submitted.

The IND automatically becomes effective 30 days after receipt by the FDA, unless the FDA raises concerns or questions regarding the proposed
clinical  trials  or  places  the  trial  on  a  clinical  hold  within  that  30-day  time  period.  In  such  a  case,  the  IND  sponsor  and  the  FDA  must  resolve  any
outstanding concerns before the clinical trial can begin. The FDA may also impose clinical holds on a biological product candidate at any time before or
during clinical trials due to safety concerns or non-compliance. If the FDA imposes a clinical hold, trials may not recommence without FDA authorization
and then only under terms authorized by the FDA. Accordingly, we cannot be sure that submission of an IND will result in the FDA allowing clinical trials
to begin, or that, once begun, issues will not arise that suspend or terminate such trials.

Further, each clinical trial must be reviewed and approved by an independent institutional review board, or IRB, at or servicing each institution at
which the clinical trial will be conducted. An IRB is charged with protecting the welfare and rights of trial participants and considers such items as whether
the risks to individuals participating in the clinical trials are minimized and are reasonable in relation to anticipated benefits. The IRB also approves the
form and content of the informed consent that must be signed by each clinical trial subject or his or her legal representative and must monitor the clinical
trial  until  completed.  Clinical  trials  involving  recombinant  or  synthetic  nucleic  acid  molecules  also  must  be  reviewed  by  an  institutional  biosafety
committee, or IBC, a local institutional committee that reviews and oversees basic and clinical research conducted at that institution. The IBC assesses the
safety of the research and identifies any potential risk to public health or the environment.

Clinical  trials  involve  the  administration  of  the  biological  product  candidate  to  healthy  volunteers  or  patients  under  the  supervision  of  qualified
investigators, generally physicians not employed by or under the trial sponsor’s control. Clinical trials must be conducted and monitored in accordance with
the FDA’s regulations comprising the GCP requirements, including the requirement that all research patients provide informed consent.

Human clinical trials are typically conducted in three sequential phases that may overlap or be combined:

•

Phase 1. The biological product is initially introduced into healthy human subjects and tested for safety. In the case of some products for severe or
life-threatening diseases, especially when the product may be too inherently toxic to ethically administer to healthy volunteers, the initial human
testing is often conducted in patients with the target disease or condition.

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•

Phase 2. The biological product is evaluated in a limited patient population to identify possible adverse effects and safety risks, to preliminarily
evaluate the efficacy of the product for specific targeted diseases and to determine dosage tolerance, optimal dosage and dosing schedule.
Phase  3.  Clinical  trials  are  undertaken  to  further  evaluate  dosage,  clinical  efficacy,  potency,  and  safety  in  an  expanded  patient  population,
generally  at  geographically  dispersed  clinical  trial  sites.  These  clinical  trials  are  intended  to  generate  enough  data  to  statistically  evaluate  the
efficacy and safety of the product for approval, to establish the overall risk to benefit profile of the product and to provide an adequate basis for
product labeling.

Phase 1, Phase 2 and Phase 3 clinical trials may not be completed successfully within any specified period, if at all.

Post-approval clinical trials, sometimes referred to as Phase 4 clinical trials, may be conducted after initial marketing approval. These clinical trials

are used to gain additional experience from the treatment of patients in the intended therapeutic indication, particularly for long-term safety follow-up.

During all phases of clinical development, regulatory agencies require extensive monitoring and auditing of all clinical activities, clinical data, and
clinical trial investigators. Annual progress reports detailing the results of the clinical trials must be submitted to the FDA. Written IND safety reports must
be promptly submitted to the FDA, and the investigators for serious and unexpected adverse events, any findings from other studies, tests in laboratory
animals or in vitro testing that suggest a significant risk for human patients, or any clinically important increase in the rate of a serious suspected adverse
reaction over that listed in the protocol or investigator brochure. The sponsor must submit an IND safety report within 15 calendar days after the sponsor
determines that the information qualifies for reporting. The sponsor also must notify the FDA of any unexpected fatal or life-threatening suspected adverse
reaction within seven calendar days after the sponsor’s initial receipt of the information. The FDA or the sponsor or its data safety monitoring board, an
independent  group  of  experts  that  evaluates  study  data  for  safety  and  makes  recommendations  concerning  continuation,  modification,  or  termination  of
clinical trials, may suspend or terminate a clinical trial at any time on various grounds, including a finding that the research patients are being exposed to an
unacceptable  health  risk,  including  risks  inferred  from  other  unrelated  immunotherapy  trials.  Similarly,  an  IRB  can  suspend  or  terminate  approval  of  a
clinical trial at its institution if the clinical trial is not being conducted in accordance with the IRB’s requirements or if the biological product has been
associated with unexpected serious harm to patients.

Because this is a relatively new and expanding area of novel therapeutic interventions, there can be no assurance as to the length of the trial period,
the  number  of  patients  the  FDA  will  require  to  be  enrolled  in  the  trials  in  order  to  establish  the  safety,  efficacy,  purity  and  potency  of  immunotherapy
products, or that the data generated in these trials will be acceptable to the FDA to support marketing approval.

Concurrently with clinical trials, companies usually complete additional nonclinical studies and must also develop additional information about the
physical characteristics of the biological product as well as finalize a process for manufacturing the product in commercial quantities in accordance with
cGMP  requirements.  To  help  reduce  the  risk  of  the  introduction  of  adventitious  agents  with  use  of  biological  products,  the  PHSA  emphasizes  the
importance of manufacturing control for products whose attributes cannot be precisely defined. The manufacturing process must be capable of consistently
producing quality batches of the product candidate and, among other things, the sponsor must develop methods for testing the identity, strength, quality,
potency and purity of the final biological product. Additionally, appropriate packaging must be selected and tested and stability studies must be conducted
to demonstrate that the biological product candidate does not undergo unacceptable deterioration over its shelf life.

U.S. Review and Approval Processes

After  the  completion  of  clinical  trials  of  a  biological  product,  FDA  approval  of  a  BLA  must  be  obtained  before  commercial  marketing  of  the
biological product. The BLA must include results of product development, laboratory and animal studies, human trials, information on the manufacture and
composition of the product, proposed labeling and other relevant information. The testing and approval processes require substantial time and effort and
there can be no assurance that the FDA will accept the BLA for filing and, even if filed, that any approval will be granted on a timely basis, if at all as the
FDA has significant discretion to approve or reject the BLA and to require additional preclinical or clinical studies.

Under the Prescription Drug User Fee Act, or PDUFA, as amended, each BLA must be accompanied by a significant user fee. The FDA adjusts the
PDUFA user fees on an annual basis. PDUFA also imposes an annual program fee for approved biological products. Fee waivers or reductions are available
in certain circumstances, including a waiver of the application fee for the first application filed by a small business. Additionally, no user fees are assessed
on BLAs for products designated as orphan drugs, unless the product also includes a non-orphan indication.

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Within 60 days following submission of the application, the FDA reviews a BLA submitted to determine if it is substantially complete before the
agency accepts it for filing. The FDA may refuse to file any BLA that it deems incomplete or not properly reviewable at the time of submission and may
request additional information. In this event, the BLA must be resubmitted with the additional information. The resubmitted application also is subject to
review before the FDA accepts it for filing. Once the submission is accepted for filing, the FDA begins an in-depth substantive review of the BLA. The
FDA reviews the BLA to determine, among other things, whether the proposed product is safe, potent, and/or effective for its intended use, and has an
acceptable purity profile, and whether the product is being manufactured in accordance with cGMP to assure and preserve the product’s identity, safety,
strength, quality, potency and purity. The FDA may refer applications for novel biological products or biological products that present difficult questions of
safety or efficacy to an advisory committee, typically a panel that includes clinicians and other experts, for review, evaluation and a recommendation as to
whether the application should be approved and under what conditions. The FDA is not bound by the recommendations of an advisory committee, but it
considers such recommendations carefully when making decisions. During the biological product approval process, the FDA also will determine whether a
Risk Evaluation and Mitigation Strategy, or REMS, is necessary to ensure that the benefits of the product outweigh its risks and to assure the safe use of the
biological product, which could include medication guides, physician communication plans, or elements to assure safe use, such as restricted distribution
methods, patient registries and other risk minimization tools. FDA determines the requirement for a REMS, as well as the specific REMS provisions, on a
case-by-case basis. If the FDA concludes a REMS is needed, the sponsor of the BLA must submit a proposed REMS. The FDA will not approve a BLA
without a REMS, if required.

Before approving a BLA, the FDA will inspect the facilities at which the product is manufactured. The FDA will not approve the product unless it
determines that the manufacturing processes and facilities are in compliance with cGMP requirements and adequate to assure consistent production of the
product within required specifications. For immunotherapy products, the FDA also will not approve the product if the manufacturer is not in compliance
with the GTPs, to the extent applicable. These are FDA regulations and guidance documents that govern the methods used in, and the facilities and controls
used  for,  the  manufacture  of  human  cells,  tissues,  and  cellular  and  tissue  based  products,  or  HCT/Ps,  which  are  human  cells  or  tissue  intended  for
implantation, transplant, infusion, or transfer into a human recipient. The primary intent of the GTP requirements is to ensure that cell and tissue based
products are manufactured in a manner designed to prevent the introduction, transmission and spread of communicable disease. FDA GTP regulations also
require  tissue  establishments  to  register  and  list  their  HCT/Ps  with  the  FDA  and,  when  applicable,  to  evaluate  donors  through  screening  and  testing.
Additionally,  before  approving  a  BLA,  the  FDA  will  typically  inspect  one  or  more  clinical  sites  to  assure  that  the  clinical  trials  were  conducted  in
compliance with IND trial requirements and GCP requirements.

To  assure  cGMP,  GTP  and  GCP  compliance,  an  applicant  must  incur  significant  expenditure  of  time,  money  and  effort  in  the  areas  of  training,

recordkeeping, production, and quality control.

Notwithstanding  the  submission  of  relevant  data  and  information,  the  FDA  may  ultimately  decide  that  the  BLA  does  not  satisfy  its  regulatory
criteria for approval and deny approval. If the agency decides not to approve the BLA in its present form, the FDA will issue a Complete Response Letter,
which  generally  outlines  the  specific  deficiencies  in  the  BLA  identified  by  the  FDA  and  may  require  additional  clinical  or  other  data  or  impose  other
conditions that must be met in order to secure final approval of the application. The deficiencies identified may be minor, for example, requiring labeling
changes, or major, for example, requiring additional clinical trials. Even with the submission of additional information, the FDA may ultimately decide that
the application does not satisfy the regulatory criteria for approval. If a Complete Response Letter is issued, the applicant may either resubmit the BLA,
addressing all of the deficiencies identified in the letter, or withdraw the application.

If  a  product  receives  regulatory  approval,  the  approval  is  limited  to  the  conditions  of  use  (e.g.,  patient  population,  indication)  described  in  the

application.

Further,  the  FDA  may  require  that  certain  contraindications,  warnings  or  precautions  be  included  in  the  product  labeling,  or  otherwise  limit  the
scope of any approval. In addition, the FDA may require post marketing clinical trials, sometimes referred to as Phase 4 clinical trials, designed to further
assess  a  biological  product’s  safety  and  effectiveness,  and  testing  and  surveillance  programs  to  monitor  the  safety  of  approved  products  that  have  been
commercialized. After approval, many types of changes to the approved product, such as adding new indications, manufacturing changes and additional
labeling claims, are subject to further testing requirements and FDA review and approval.

In  addition,  under  the  Pediatric  Research  Equity  Act,  or  PREA,  a  BLA  or  supplement  to  a  BLA  must  contain  data  to  assess  the  safety  and
effectiveness of the product for the claimed indications in all relevant pediatric subpopulations and to support dosing and administration for each pediatric
subpopulation for which the product is safe and effective. The FDA may grant deferrals for submission of data or full or partial waivers.

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Post-Approval Requirements

Any products for which we receive FDA approvals are subject to continuing regulation by the FDA, including, among other things, record-keeping
requirements, reporting of adverse experiences with the product, providing the FDA with updated safety and efficacy information, product sampling and
distribution  requirements,  and  complying  with  FDA  promotion  and  advertising  requirements,  which  include,  among  others,  standards  for  direct-to-
consumer advertising, restrictions on promoting products for uses or in patient populations that are not described in the product’s approved uses (known as
“off-label  use”),  limitations  on  industry-sponsored  scientific  and  educational  activities,  and  requirements  that  important  safety  information  and  material
facts  related  to  the  product  be  disclosed.  Although  physicians  may  prescribe  legally  available  products  for  off-label  uses,  if  the  physicians  deem  to  be
appropriate in their professional medical judgment, manufacturers may not market or promote such off-label uses. The FDA and other agencies actively
enforce the laws and regulations prohibiting the promotion of off-label uses, and a company that is found to have improperly promoted off-label uses may
be subject to significant civil, criminal and administrative liability.

In  addition,  quality  control  and  manufacturing  procedures  must  continue  to  conform  to  applicable  manufacturing  requirements  after  approval  to
ensure  the  long-term  stability  of  the  product.  We  rely,  and  expect  to  continue  to  rely,  on  third  parties  for  the  production  of  clinical  and  commercial
quantities of our products in accordance with cGMP regulations. cGMP regulations require among other things, quality control and quality assurance as
well  as  the  corresponding  maintenance  of  records  and  documentation  and  the  obligation  to  investigate  and  correct  any  deviations  from  cGMP.
Manufacturers and other entities involved in the manufacture and distribution of approved products are required to register their establishments with the
FDA and certain state agencies, and are subject to periodic unannounced inspections by the FDA and certain state agencies for compliance with cGMP and
other laws. Accordingly, manufacturers must continue to expend time, money, and effort in the area of production and quality control to maintain cGMP
compliance. Discovery of problems with a product after approval may result in restrictions on a product, manufacturer, or holder of an approved BLA,
including, among other things, recall or withdrawal of the product from the market.

The  FDA  also  may  require  post-marketing  testing,  known  as  Phase  4  testing,  and  surveillance  to  monitor  the  effects  of  an  approved  product.
Discovery of previously unknown problems with a product, including adverse events of unanticipated severity or frequency, with manufacturing processes,
or  the  failure  to  comply  with  applicable  FDA  requirements  can  have  negative  consequences,  including  adverse  publicity,  judicial  or  administrative
enforcement,  complete  withdrawal  from  the  market,  product  recalls,  warning  letters  from  the  FDA,  mandated  corrective  advertising  or  communications
with doctors, product seizure or detention, injunctions, and civil or criminal penalties, among others. Newly discovered or developed safety or effectiveness
data  may  require  changes  to  a  product’s  approved  labeling,  including  the  addition  of  new  warnings  and  contraindications,  and  also  may  require  the
implementation  of  other  risk  management  measures.  Also,  new  government  requirements,  including  those  resulting  from  new  legislation,  may  be
established, or the FDA’s policies may change, which could delay or prevent regulatory approval of our products under development.

U.S. Marketing Exclusivity

The Biologics Price Competition and Innovation Act amended the PHSA to authorize the FDA to approve similar versions of innovative biologics,
commonly  known  as  biosimilars.  Biosimilars  are  approved  pursuant  to  an  abbreviated  pathway  whereby  applicants  need  not  submit  the  full  slate  of
preclinical and clinical data, and approval is based in part on the FDA’s findings of safety, purity, and potency for the original biologic (i.e., the reference
product).  Original  BLAs  are  eligible  to  receive  12  years  of  exclusivity  from  the  time  of  first  licensure  of  the  product,  which  prevents  the  FDA  from
approving any biosimilars to the reference product through the abbreviated pathway, but does not prevent approval of BLAs that are accompanied by a full
data  package  and  that  do  not  rely  on  the  reference  product.  A  biosimilar  may  be  approved  if  the  product  is  highly  similar  to  the  reference  product
notwithstanding minor differences in clinically inactive components and there are no clinically meaningful differences with the reference product in terms
of the safety, purity, and potency.

Pediatric  exclusivity  is  another  type  of  regulatory  market  exclusivity  in  the  United  States.  Pediatric  exclusivity,  if  granted,  adds  six  months  to
existing exclusivity periods and patent terms. This six-month exclusivity, which runs from the end of other exclusivity protection or patent term, may be
granted based on the voluntary completion of a pediatric trial in accordance with an FDA-issued “Written Request” for such a trial.

Coverage, Pricing and Reimbursement

Significant uncertainty exists as to the coverage and reimbursement status of any product candidates for which we obtain regulatory approval. In
the  United  States  and  markets  in  other  countries,  sales  of  any  products  for  which  we  receive  regulatory  approval  for  commercial  sale  will  depend,  in
significant part, on the extent to which third-party payors provide coverage, and establish adequate reimbursement levels for such products. In the United
States, third-party payers include federal and state healthcare programs, private managed care organizations, health insurers and other organizations. The
process  for  determining  whether  a  third-party  payer  will  provide  coverage  for  a  product  may  be  separate  from  the  process  of  establishing  the
reimbursement rate that such a

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payor will pay for the product. Third-party payors may limit coverage to specific products on an approved list, also known as a formulary, which might not
include  all  of  the  FDA-approved  products  for  a  particular  indication.  Third-party  payors  are  increasingly  challenging  the  price,  examining  the  medical
necessity of and reviewing the cost-effectiveness of medical products, therapies and services, in addition to questioning their safety and efficacy.

Reimbursement may impact the demand for, and/or the price of, any product candidate which obtains marketing approval. Even if coverage and
reimbursement is obtained for a given product candidate by a third-party payor, the resulting reimbursement payment rates may not be adequate or may
require  co-payments  that  patients  find  unacceptably  high.  Patients  who  are  prescribed  medications  for  the  treatment  of  their  conditions,  and  their
prescribing physicians, generally rely on third-party payors to reimburse all or part of the costs associated with those medications. Patients are unlikely to
use  a  product,  and  physicians  may  be  less  likely  to  prescribe  a  product,  unless  coverage  is  provided  and  reimbursement  is  adequate  to  cover  all  or  a
significant portion of the cost of the product. Therefore, coverage and adequate reimbursement is critical to new drug product acceptance.

Additionally, we are developing a proprietary diagnostic test for use with certain of our product candidates. The diagnostic test will require separate
regulatory  approval  in  addition  to  the  regulatory  approval  of  AUTO4  and  AUTO5.  Failure  to  obtain  marketing  approval  for  the  diagnostic  test  could
prevent us from commercializing either AUTO4 or AUTO5 unless another similar diagnostic test for distinguishing TRBC1-positive and TRBC2-positive
T cell lymphomas is commercially available.

Different  pricing  and  reimbursement  schemes  exist  in  other  countries.  In  the  EU,  governments  influence  the  price  of  pharmaceutical  products
through their pricing and reimbursement rules and control of national healthcare systems that fund a large part of the cost of those products to consumers.
Some jurisdictions operate positive and negative list systems under which products may only be marketed once a reimbursement price has been agreed. To
obtain  reimbursement  or  pricing  approval,  some  of  these  countries  may  require  the  completion  of  additional  clinical  trials  that  compare  the  cost-
effectiveness of a particular product candidate to currently available therapies. Other member states allow companies to fix their own prices for medicines,
but monitor and control company profits. In addition, in some countries, cross-border imports from low-priced markets exert a commercial pressure on
pricing within a country.

The downward pressure on healthcare costs in general, particularly prescription drugs and biologics, has become very intense. Governments have
shown  significant  interest  in  implementing  cost-containment  programs,  including  price  controls,  restrictions  on  reimbursement  and  requirements  for
substitution of generic products. As a result, increasingly high barriers are being erected to the entry of new products. The marketability of any product
candidates for which we receive regulatory approval for commercial sale may suffer if the government and third-party payors fail to provide coverage and
adequate reimbursement. In addition, emphasis on managed care in the United States has increased and we expect will continue to increase the pressure on
healthcare pricing. Coverage policies and third-party reimbursement rates may change at any time. Even if favorable coverage and reimbursement status is
attained for one or more products for which we receive regulatory approval, less favorable coverage policies and reimbursement rates may be implemented
in the future.

Healthcare Laws Governing Interactions with Healthcare Providers

In  addition  to  FDA  restrictions  on  marketing  of  pharmaceutical  products,  several  other  types  of  state  and  federal  laws  restrict  our  business
activities, including certain marketing practices. These laws include, without limitation, anti-kickback laws, false claims laws, data privacy and security
laws, as well as transparency laws regarding payments or other items of value provided to healthcare providers.

The  U.S.  federal  Anti-Kickback  Statute  prohibits  any  person  or  entity  from,  among  other  things,  knowingly  and  willfully  offering,  paying,
soliciting or receiving remuneration, directly or indirectly, in cash or in kind, to induce or reward, or in return for, purchasing, leasing, ordering or arranging
for the purchase, lease or order of any healthcare item, good, facility or service reimbursable, in whole or in part, under Medicare, Medicaid or other federal
healthcare  programs.  The  term  ‘‘remuneration’’  has  been  broadly  interpreted  to  include  anything  of  value.  This  statute  has  been  interpreted  to  apply  to
arrangements  between  pharmaceutical  manufacturers  on  the  one  hand  and  prescribers,  purchasers  and  formulary  managers  on  the  other  hand.  Although
there are a number of statutory exceptions and regulatory safe harbors protecting certain common activities from prosecution or other regulatory sanctions,
the exceptions and safe harbors are drawn narrowly, and practices that involve remuneration that are alleged to be intended to induce prescribing, purchases
or recommendations may be subject to scrutiny if they do not qualify for an exception or safe harbor. Failure to meet all of the requirements of a particular
applicable statutory exception or regulatory safe harbor does not make the conduct per se illegal under the U.S. federal Anti-Kickback Statute. Instead, the
legality of the arrangement will be evaluated on a case-by-case basis based on a cumulative review of all its facts and circumstances. Several courts have
interpreted  the  statute’s  intent  requirement  to  mean  that  if  any  one  purpose  of  an  arrangement  involving  remuneration  is  to  induce  referrals  of  federal
healthcare  covered  business,  the  U.S.  federal  Anti-Kickback  Statute  has  been  violated.  Additionally,  the  intent  standard  under  the  U.S.  federal  Anti-
Kickback Statute was amended by the Patient Protection and Affordable Care Act of 2010, as amended by the Health Care and Education Reconciliation
Act of 2010, collectively the Affordable Care Act, or ACA, to a stricter standard such that a person or entity no longer

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needs to have actual knowledge of the statute or specific intent to violate it in order to have committed a violation. In addition, the ACA codified case law
that  a  claim  including  items  or  services  resulting  from  a  violation  of  the  U.S.  federal  Anti-Kickback  Statute  constitutes  a  false  or  fraudulent  claim  for
purposes of the U.S. federal False Claims Act.

Federal civil and criminal false claims laws and civil monetary penalties laws, including the U.S. federal False Claims Act, , which can be enforced
through civil whistleblower or qui tam actions, prohibit any person or entity from, among other things, knowingly presenting, or causing to be presented, a
false  claim  for  payment  to  the  federal  government,  or  knowingly  making,  or  causing  to  be  made,  a  false  statement  to  have  a  false  claim  paid.
Pharmaceutical and other healthcare companies have been prosecuted under these laws for, among other things, allegedly inflating drug prices they report
to pricing services, which in turn were used by the government to set Medicare and Medicaid reimbursement rates, and for allegedly providing free product
to customers with the expectation that the customers would bill federal programs for the product. In addition, certain marketing practices, including off-
label promotion, may also violate false claims laws. Further, pharmaceutical manufacturers can be held liable under the U.S. federal False Claims Act even
when they do not submit claims directly to government payors if they are deemed to “cause” the submission of false or fraudulent claims.

The U.S. federal Health Insurance Portability and Accountability Act of 1996, or HIPAA, created new federal criminal statutes that prohibit among
other actions, knowingly and willfully executing, or attempting to execute, a scheme to defraud any healthcare benefit program, including private third-
party  payors,  knowingly  and  willfully  embezzling  or  stealing  from  a  healthcare  benefit  program,  willfully  obstructing  a  criminal  investigation  of  a
healthcare offense, and knowingly and willfully falsifying, concealing or covering up a material fact or making any materially false, fictitious or fraudulent
statement in connection with the delivery of or payment for healthcare benefits, items or services. Like the U.S. federal Anti-Kickback Statute, the ACA
amended the intent standard for certain healthcare fraud under HIPAA such that a person or entity no longer needs to have actual knowledge of the statute
or specific intent to violate it in order to have committed a violation.

HIPAA, as amended by the Health Information Technology for Economic and Clinical Health Act, or HITECH, and their implementing regulations,
impose  certain  requirements  on  “covered  entities,”  including  certain  healthcare  providers,  health  plans  and  healthcare  clearinghouses,  as  well  as  their
respective “business associates” that create, receive, maintain or transmit individually identifiable health information for or on behalf of a covered entity,
and their covered subcontractors, relating to the privacy, security, transmission and breach of individually identifiable health information. Further, HITECH
also created four new tiers of civil monetary penalties, amended HIPAA to make civil and criminal penalties directly applicable to business associates, and
gave state attorneys general new authority to file civil actions for damages or injunctions in U.S. federal courts to enforce HIPAA and seek attorneys’ fees
and costs associated with pursuing federal civil actions.

Additionally,  the  federal  Physician  Payments  Sunshine  Act,  created  under  the  ACA,  and  its  implementing  regulations,  require  certain
manufacturers  of  drugs,  devices,  biologics  and  medical  supplies  for  which  payment  is  available  under  Medicare,  Medicaid  or  the  Children’s  Health
Insurance Program (with certain exceptions) to annually report to the Centers for Medicare and Medicaid Services, or CMS, information related to certain
payments or other transfers of value provided to physicians (defined to include doctors, dentists, optometrists, podiatrists and chiropractors) and teaching
hospitals, or to entities or individuals at the request of, or designated on behalf of, the physicians and teaching hospitals as well as certain ownership and
investment interests held by physicians and their immediate family members. Beginning in 2022, applicable manufacturers also will be required to report
information  regarding  payments  and  transfers  of  value  provided  to  physician  assistants,  nurse  practitioners,  clinical  nurse  specialists,  certified  nurse
anesthetists, anesthesiologist assistants, and certified nurse-midwives.

Additionally,  similar  healthcare  laws  and  regulations  in  the  European  Union  and  other  jurisdictions,  including  reporting  requirements  detailing
interactions with and payments to healthcare providers and laws governing the privacy and security of certain protected information, such as GDPR, which
imposes obligations and restrictions on the collection and use of personal data relating to individuals located in the European Union (including health data).

Finally, the majority of states also have statutes or regulations similar to the aforementioned federal laws, some of which are broader in scope and
apply  to  items  and  services  reimbursed  under  Medicaid  and  other  state  programs,  or,  in  several  states,  apply  regardless  of  the  payor.  Some  state  laws
require  pharmaceutical  companies  to  comply  with  the  pharmaceutical  industry’s  voluntary  compliance  guidelines  and  the  relevant  compliance  guidance
promulgated  by  the  federal  government  in  addition  to  requiring  drug  manufacturers  to  report  information  related  to  payments  to  clinicians  and  other
healthcare providers or marketing expenditures. Some states and local jurisdictions require the registration of pharmaceutical sales representatives. State
and foreign laws also govern the privacy and security of health information in some circumstances, many of which differ from each other in significant
ways and often are not preempted by HIPAA, thus complicating compliance efforts.

Because of the breadth of these laws and the narrowness of their exceptions and safe harbors, it is possible that business activities can be subject to
challenge  under  one  or  more  of  such  laws.  The  scope  and  enforcement  of  each  of  these  laws  is  uncertain  and  subject  to  rapid  change  in  the  current
environment of healthcare reform, especially in light of the lack of applicable precedent and

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regulations.  Federal  and  state  enforcement  bodies  have  recently  increased  their  scrutiny  of  interactions  between  healthcare  companies  and  healthcare
providers, which has led to a number of investigations, prosecutions, convictions and settlements in the healthcare industry.

Ensuring  that  business  arrangements  with  third  parties  comply  with  applicable  healthcare  laws  and  regulations  is  costly  and  time  consuming.  If
business  operations  are  found  to  be  in  violation  of  any  of  the  laws  described  above  or  any  other  applicable  governmental  regulations  a  pharmaceutical
manufacturer may be subject to penalties, including civil, criminal and administrative penalties, damages, fines, disgorgement, individual imprisonment,
exclusion from governmental funded healthcare programs, such as Medicare and Medicaid, contractual damages, reputational harm, diminished profits and
future earnings, additional reporting obligations and oversight if subject to a corporate integrity agreement or other agreement to resolve allegations of non-
compliance with these laws, and curtailment or restructuring of operations, any of which could adversely affect a pharmaceutical manufacturer’s ability to
operate its business and the results of its operations.

Healthcare Reform Efforts

A primary trend in the United States healthcare industry and elsewhere is cost containment. Over the last several years, there have been federal and
state proposals and legislation enacted regarding the pricing of pharmaceutical and biopharmaceutical products, limiting coverage and reimbursement for
drugs and other medical products, and making changes to healthcare financing and the delivery of care in the United States. We do not expect this trend to
dissipate with the advent of a new administration; but may take an initial back seat to its work on COVID 19 vaccination and prevention efforts.

During his 2020 campaign, President Biden stressed his desire to expand and strengthen the Affordable Care Act, in which he played a large role to
help pass through Congress in 2010. Specifically, this includes lowering the age of Medicare eligibility to 60, expanding coverage to those who are caught
in the gap between Medicaid eligibility and qualifying for ACA subsidies, and potentially creating a “public option” to cover those who have systemically
been  uninsured.  We  also  expect  the  Biden  administration  will  overturn  several  of  the  last  minute  regulations  and  waivers  approved  by  the  Trump
administration including Medicaid work requirements. In fact, the administration on Day 1 issued a regulatory freeze to all federal agencies and requested
agencies delay the effective date of new regulations until the new appointed agency head has reviewed those regulations.

We expect the Biden Administration may ramp up its efforts on drug pricing as it explores ways to pay for its various efforts to combat COVID 19,
health  equity,  insurance  coverage,  and  other  priorities.  One  area  we  anticipate  increased  interest  is  direct  Medicare  price  negotiation  with  drug
manufacturers, however, it will remain challenging for Democrats to pass partisan legislation despite having control of the White House and Congress. The
current 50-50 split in the Senate (with the Vice President breaking any tie votes) means that Democrats will need to either pursue legislation that will get
some Republican votes or use the budget reconciliation mechanism to pass a more limited set of proposals with only 51 votes. We also anticipate the new
administration will continue to use executive orders and Medicare demonstration authority to control drug costs. While the courts have essentially deemed
the Trump administration’s Most Favored Nations model—which would have benchmarked Medicare payment rates on international reference prices for
drugs—dead in the water, some of the proposals within the model may resurface under a new name. Other areas of reform like copay accumulators and Part
D reform may also have impact on drug pricing.

Further, there remains heightened Congressional scrutiny in the United States of pharmaceutical pricing practices designed to, among other things,
bring more transparency in product pricing, review the relationship between pricing and manufacturer patient programs, and reform government program
reimbursement methodologies for products. We expect both the Biden Administration’s first budget proposal and Congressional actors to begin hearings in
earnest once relief measures for COVID 19 and other “first 100 day” issues like immigration, rejoining various International accords, and staffing at federal
agencies,  has  taken  place.  US  states  are  also  not  immune  to  such  discussions.  At  the  state  level,  legislatures  have  increasingly  enacted  legislation  and
implemented  regulations  designed  to  control  pharmaceutical  and  biological  product  pricing,  including  price  or  patient  reimbursement  constraints,
discounts,  restrictions  on  certain  product  access  and  marketing  cost  disclosure  and  transparency  measures,  and,  in  some  cases,  designed  to  encourage
importation from other countries and bulk purchasing. In addition, regional healthcare authorities and individual hospitals are increasingly using bidding
procedures to determine what pharmaceutical products and which suppliers will be included in their prescription drug and other healthcare programs. It is
also possible that additional governmental action is taken in response to the COVID-19 pandemic.

In addition, other federal health reform measures have been proposed and adopted in the United States that could impact cell therapy. Most notably,
the previous administration supported and promulgated a rule related to value based payment alternatives in the Medicaid program. Medicaid is a jointly
run federal and state program that provides health benefits coverage for low-income residents and children. In exchange for broad coverage in Medicaid,
drug manufacturers are required to sign a Medicare Drug Rebate agreement which requires them to offer Medicaid programs the “best price” available for a
particular product. This “best price” takes into consideration any rebates or concessions manufacturers offer, with some exceptions. The final rule would
exempt value-based or outcomes-based payment arrangements from the definition of “best price” which provides manufacturers more flexibility to work
with

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commercial payers and states on innovate payment mechanisms for high-cost cell and gene therapies. While Medicaid is not a significant driver of cell
therapy sales it is a bellwether program and one we watch closely.

FCPA, the Bribery Act and Other Laws

The FCPA, prohibits any U.S. individual or business from paying, offering, or authorizing payment or offering of anything of value, directly or
indirectly, to any foreign official, political party or candidate for the purpose of influencing any act or decision of the foreign entity in order to assist the
individual or business in obtaining or retaining business. The FCPA also obligates companies whose securities are listed in the United States to comply
with  accounting  provisions  requiring  the  company  to  maintain  books  and  records  that  accurately  and  fairly  reflect  all  transactions  of  the  corporation,
including international subsidiaries, and to devise and maintain an adequate system of internal accounting controls for international operations. Activities
that violate the FCPA, even if they occur wholly outside the United States, can result in criminal and civil fines, imprisonment, disgorgement, oversight,
and debarment from government contracts.

Our operations are also subject to non-U.S. anti-corruption laws such as the Bribery Act. As with the FCPA, these laws generally prohibit us and
our employees and intermediaries from authorizing, promising, offering, or providing, directly or indirectly, improper or prohibited payments, or anything
else of value, to government officials or other persons to obtain or retain business or gain some other business advantage. Under the Bribery Act, we may
also be liable for failing to prevent a person associated with us from committing a bribery offense.

We are also subject to other laws and regulations governing our international operations, including regulations administered by the governments of
the United Kingdom and the United States and authorities in the European Union, including applicable export control regulations, economic sanctions and
embargoes on certain countries and persons, anti-money laundering laws, import and customs requirements and currency exchange regulations, collectively
referred to as trade control laws.

Failure  to  comply  with  the  Bribery  Act,  the  FCPA  and  other  anti-corruption  laws  and  trade  control  laws  could  subject  us  to  criminal  and  civil

penalties, disgorgement and other sanctions and remedial measures, and legal expenses.

Review and Approval of New Drug Products in the European Union

In  the  European  Union,  medicinal  products,  including  advanced  therapy  medicinal  products,  or  ATMPs,  are  subject  to  extensive  pre-  and  post-
market regulation by regulatory authorities at both the European Union and national levels. ATMPs comprise gene therapy products, somatic-cell therapy
products and tissue engineered products, which are cells or tissues that have undergone substantial manipulation and that are administered to human beings
in order to regenerate, repair or replace a human tissue. We anticipate that our T cell therapy products will be regulated as ATMPs in the European Union.
There  is  legislation  at  a  European  Union  level  relating  to  the  standards  of  quality  and  safety  for  the  collection  and  testing  of  human  blood  and  blood
components  for  use  in  cell-based  therapies,  which  could  apply  to  our  products.  Additionally,  there  may  be  local  legislation  in  various  European  Union
Member States, which may be more restrictive than the European Union legislation, and we would need to comply with such legislation to the extent it
applies.

Clinical Trials

Clinical trials of medicinal products in the European Union must be conducted in accordance with European Union and national regulations and the
International  Conference  on  Harmonization,  or  ICH,  guidelines  on  Good  Clinical  Practices,  or  GCP.  Additional  GCP  guidelines  from  the  European
Commission, focusing in particular on traceability, apply to clinical trials of ATMPs. The sponsor must take out a clinical trial insurance policy, and in most
European Union countries, the sponsor is liable to provide “no fault” compensation to any study subject injured in the clinical trial.

Prior to commencing a clinical trial, the sponsor must obtain a clinical trial authorization from the competent authority, and a positive opinion from
an independent ethics committee. The application for a clinical trial authorization must include, among other things, a copy of the trial protocol and an
investigational medicinal product dossier containing information about the manufacture and quality of the medicinal product under investigation. Currently,
clinical trial authorization applications must be submitted to the competent authority in each EU Member State in which the trial will be conducted. Under
the  new  Regulation  on  Clinical  Trials,  which  is  expected  to  take  effect  in  December  2021,  there  will  be  a  centralized  application  procedure  where  one
national authority takes the lead in reviewing the application and the other national authorities have only a limited involvement. Any substantial changes to
the trial protocol or other information submitted with the clinical trial applications must be notified to or approved by the relevant competent authorities
and  ethics  committees.  Medicines  used  in  clinical  trials  must  be  manufactured  in  accordance  with  cGMP.  Other  national  and  European  Union-wide
regulatory requirements also apply.

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During the development of a medicinal product, the EMA and national medicines regulators within the European Union provide the opportunity for
dialogue and guidance on the development program. At the EMA level, this is usually done in the form of scientific advice, which is given by the Scientific
Advice Working Party of the Committee for Medicinal Products for Human Use, or CHMP. A fee is incurred with each scientific advice procedure. Advice
from  the  EMA  is  typically  provided  based  on  questions  concerning,  for  example,  quality  (chemistry,  manufacturing  and  controls  testing),  nonclinical
testing and clinical studies, and pharmacovigilance plans and risk-management programs. In accordance with the EMA’s policy, scientific advice will not
be legally binding with regard to any future marketing authorization application of the product concerned.

Marketing Authorizations

In order to market a new medicinal product in the European Union, a company must submit and obtain approval from regulators of a marketing

authorization application, or MAA. The process for doing this depends, among other things, on the nature of the medicinal product.

The centralized procedure results in a single marketing authorization, or MA, granted by the European Commission that is valid across the EEA
(i.e., the European Union as well as Iceland, Liechtenstein and Norway). The centralized procedure is compulsory for human drugs that are: (i) derived
from  biotechnology  processes,  such  as  genetic  engineering,  (ii)  contain  a  new  active  substance  indicated  for  the  treatment  of  certain  diseases,  such  as
HIV/AIDS, cancer, diabetes, neurodegenerative diseases, autoimmune and other immune dysfunctions and viral diseases, (iii) officially designated orphan
medicines and (iv) advanced-therapy medicines, such as gene therapy, somatic cell therapy or tissue-engineered medicines. The centralized procedure may
at  the  request  of  the  applicant  also  be  used  in  certain  other  cases.  Therefore,  the  centralized  procedure  would  be  mandatory  for  the  products  we  are
developing.

The Committee for Advanced Therapies, or CAT, is responsible in conjunction with the CHMP for the evaluation of ATMPs. The CAT is primarily
responsible for the scientific evaluation of ATMPs and prepares a draft opinion on the quality, safety and efficacy of each ATMP for which a marketing
authorization application is submitted. The CAT’s opinion is then taken into account by the CHMP when giving its final recommendation regarding the
authorization of a product in view of the balance of benefits and risks identified. Although the CAT’s draft opinion is submitted to the CHMP for final
approval,  the  CHMP  may  depart  from  the  draft  opinion,  if  it  provides  detailed  scientific  justification.  The  CHMP  and  CAT  are  also  responsible  for
providing  guidelines  on  ATMPs  and  have  published  numerous  guidelines,  including  specific  guidelines  on  gene  therapies  and  cell  therapies.  These
guidelines provide additional guidance on the factors that the EMA will consider in relation to the development and evaluation of ATMPs and include,
among  other  things,  the  preclinical  studies  required  to  characterize  ATMPs;  the  manufacturing  and  control  information  that  should  be  submitted  in  a
marketing  authorization  application;  and  post-approval  measures  required  to  monitor  patients  and  evaluate  the  long  term  efficacy  and  potential  adverse
reactions of ATMPs. Although these guidelines are not legally binding, we believe that our compliance with them is likely necessary to gain and maintain
approval for any of our product candidates.

Under the centralized procedure in the European Union, the maximum timeframe for the evaluation of an MAA by the EMA is 210 days. This
excludes so-called clock stops, during which additional written or oral information is to be provided by the applicant in response to questions asked by the
CHMP. At the end of the review period, the CHMP provides an opinion to the European Commission. If this is opinion favorable, the Commission may
then adopt a decision to grant an MA. In exceptional cases, the CHMP might perform an accelerated review of an MAA in no more than 150 days. This is
usually when the product is of major interest from the point of view of public health and, in particular, from the viewpoint of therapeutic innovation.

The European Commission may grant a so-called “marketing authorization under exceptional circumstances”. Such authorization is intended for
products for which the applicant can demonstrate that it is unable to provide comprehensive data on the efficacy and safety under normal conditions of use,
because the indications for which the product in question is intended are encountered so rarely that the applicant cannot reasonably be expected to provide
comprehensive  evidence,  or  in  the  present  state  of  scientific  knowledge,  comprehensive  information  cannot  be  provided,  or  it  would  be  contrary  to
generally accepted principles of medical ethics to collect such information. Consequently, marketing authorization under exceptional circumstances may be
granted subject to certain specific obligations, which may include the following:

•

•

•

the applicant must complete an identified program of studies within a time period specified by the competent authority, the results of which form
the basis of a reassessment of the benefit/risk profile;
the medicinal product in question may be supplied on medical prescription only and may in certain cases be administered only under strict medical
supervision, possibly in a hospital and in the case of a radiopharmaceutical, by an authorized person; and
the  package  leaflet  and  any  medical  information  must  draw  the  attention  of  the  medical  practitioner  to  the  fact  that  the  particulars  available
concerning the medicinal product in question are as yet inadequate in certain specified respects.

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A  marketing  authorization  under  exceptional  circumstances  is  subject  to  annual  review  to  reassess  the  risk-benefit  balance  in  an  annual
reassessment procedure. Continuation of the authorization is linked to the annual reassessment and a negative assessment could potentially result in the
marketing authorization being suspended or revoked. The renewal of a marketing authorization of a medicinal product under exceptional circumstances,
however, follows the same rules as a “normal” marketing authorization. Thus, a marketing authorization under exceptional circumstances is granted for an
initial five years, after which the authorization will become valid indefinitely, unless the EMA decides that safety grounds merit one additional five-year
renewal.

The  European  Commission  may  also  grant  a  so-called  “conditional  marketing  authorization”  prior  to  obtaining  the  comprehensive  clinical  data
required for an application for a full marketing authorization. Such conditional marketing authorizations may be granted for product candidates (including
medicines designated as orphan medicinal products), if (i) the risk-benefit balance of the product candidate is positive, (ii) it is likely that the applicant will
be in a position to provide the required comprehensive clinical trial data, (iii) the product fulfills an unmet medical need and (iv) the benefit to public health
of the immediate availability on the market of the medicinal product concerned outweighs the risk inherent in the fact that additional data are still required.
A  conditional  marketing  authorization  may  contain  specific  obligations  to  be  fulfilled  by  the  marketing  authorization  holder,  including  obligations  with
respect to the completion of ongoing or new studies, and with respect to the collection of pharmacovigilance data. Conditional marketing authorizations are
valid for one year, and may be renewed annually, if the risk-benefit balance remains positive, and after an assessment of the need for additional or modified
conditions and/or specific obligations. The timelines for the centralized procedure described above also apply with respect to the review by the CHMP of
applications for a conditional marketing authorization.

The European Union medicines rules expressly permit the EU Member States to adopt national legislation prohibiting or restricting the sale, supply
or use of any medicinal product containing, consisting of or derived from a specific type of human or animal cell, such as embryonic stem cells. While the
products we have in development do not make use of embryonic stem cells, it is possible that the national laws in certain EU Member States may prohibit
or restrict us from commercializing our products, even if they have been granted an EU marketing authorization.

Data Exclusivity

Marketing authorization applications for generic medicinal products do not need to include the results of preclinical and clinical trials, but instead
can  refer  to  the  data  included  in  the  marketing  authorization  of  a  reference  product  for  which  regulatory  data  exclusivity  has  expired.  If  a  marketing
authorization is granted for a medicinal product containing a new active substance, that product benefits from eight years of data exclusivity, during which
generic marketing authorization applications referring to the data of that product may not be accepted by the regulatory authorities, and a further two years
of market exclusivity, during which such generic products may not be placed on the market. The two-year period may be extended to three years if during
the first eight years a new therapeutic indication with significant clinical benefit over existing therapies is approved.

There is a special regime for biosimilars, or biological medicinal products that are similar to a reference medicinal product but that do not meet the
definition of a generic medicinal product, for example, because of differences in raw materials or manufacturing processes. For such products, the results of
appropriate preclinical or clinical trials must be provided, and guidelines from the EMA detail the type of quantity of supplementary data to be provided for
different types of biological product. There are no such guidelines for complex biological products, such as gene or cell therapy medicinal products, and so
it is unlikely that biosimilars of those products will currently be approved in the European Union. However, guidance from the EMA states that they will be
considered in the future in light of the scientific knowledge and regulatory experience gained at the time.

Pediatric Development

In the European Union, companies developing a new medicinal product must agree to a Pediatric Investigation Plan, or PIP, with the EMA and
must conduct pediatric clinical trials in accordance with that PIP, unless a deferral or waiver applies, (e.g., because the relevant disease or condition occurs
only in adults). The marketing authorization application for the product must include the results of pediatric clinical trials conducted in accordance with the
PIP, unless a waiver applies, or a deferral has been granted, in which case the pediatric clinical trials must be completed at a later date. Products that are
granted a marketing authorization on the basis of the pediatric clinical trials conducted in accordance with the PIP are eligible for a six month extension of
the protection under a supplementary protection certificate (if any is in effect at the time of approval) or, in the case of orphan medicinal products, a two
year  extension  of  the  orphan  market  exclusivity.  This  pediatric  reward  is  subject  to  specific  conditions  and  is  not  automatically  available  when  data  in
compliance with the PIP are developed and submitted.

Post-Approval Controls

The holder of a marketing authorization must establish and maintain a pharmacovigilance system and appoint an individual qualified person for
pharmacovigilance, or QPPV, who is responsible for oversight of that system. Key obligations include expedited reporting of suspected serious adverse
reactions and submission of periodic safety update reports, or PSURs.

All  new  marketing  authorization  applications  must  include  a  risk  management  plan,  or  RMP,  describing  the  risk  management  system  that  the

company will put in place and documenting measures to prevent or minimize the risks associated with the product. The

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regulatory  authorities  may  also  impose  specific  obligations  as  a  condition  of  the  marketing  authorization.  Such  risk-minimization  measures  or  post-
authorization obligations may include additional safety monitoring, more frequent submission of PSURs, or the conduct of additional clinical trials or post-
authorization safety studies. RMPs and PSURs are routinely available to third parties requesting access, subject to limited redactions. All advertising and
promotional activities for the product must be consistent with the approved summary of product characteristics, and therefore all off-label promotion is
prohibited.  Direct-to-consumer  advertising  of  prescription  medicines  is  also  prohibited  in  the  European  Union.  Although  general  requirements  for
advertising and promotion of medicinal products are established under EU directives, the details are governed by regulations in each EU Member State and
can differ from one country to another.

Pricing and Reimbursement in the European Union

Governments  influence  the  price  of  medicinal  products  in  the  European  Union  through  their  pricing  and  reimbursement  rules  and  control  of
national healthcare systems that fund a large part of the cost of those products to consumers. Some jurisdictions operate positive and negative list systems
under which products may only be marketed once a reimbursement price has been agreed. To obtain reimbursement or pricing approval, some of these
countries may require the completion of clinical trials that compare the cost-effectiveness of a particular product candidate to currently available therapies.
Other  EU  Member  States  allow  companies  to  fix  their  own  prices  for  medicines,  but  monitor  and  control  company  profits.  The  downward  pressure  on
healthcare costs in general, particularly prescription medicines, has become very intense. As a result, increasingly high barriers are being erected to the
entry of new products.

Brexit and the Regulatory Framework in the United Kingdom

On June 23, 2016, the electorate in the United Kingdom voted in favor of leaving the European Union (commonly referred to as ‘‘Brexit’’) and the
United Kingdom officially withdrew from the European Union on January 31, 2020. Pursuant to the formal withdrawal arrangements agreed between the
United Kingdom and the European Union, the United Kingdom was subject to a transition period until December 31, 2020, or the Transition Period, during
which  European  Union  rules  continued  to  apply.  A  trade  and  cooperation  agreement,  or  the  Trade  and  Cooperation  Agreement,  that  outlines  the  future
trading relationship between the United Kingdom and the European Union was agreed in December 2020.

Brexit may influence the attractiveness of the United Kingdom as a place to conduct clinical trials. The European Union’s regulatory environment
for clinical trials is being harmonized as part of the Clinical Trial Regulations, which are due to enter into full effect at the end of 2021, but it is currently
unclear as to what extent the United Kingdom will seek to align its regulations with the European Union. Failure of the United Kingdom to closely align its
regulations with the EU may have an effect on the cost of conducting clinical trials in the United Kingdom as opposed to other countries and/or make it
harder to seek a marketing authorization for our product candidates on the basis of clinical trials conducted in the United Kingdom.

In  the  short  term  there  will  be  few  changes  to  clinical  trials  that  only  have  sites  in  the  United  Kingdom.  The  MHRA  have  confirmed  that  the
sponsor of a clinical trial can be based in the EEA for an initial period following Brexit. Further investigational medicinal products can be supplied directly
from  the  EU/EEA  to  a  trial  site  in  Great  Britain  without  further  oversight  until  1  January  2022,  and  to  Northern  Ireland  beyond  such  date.  The  United
Kingdom is now a “third country” for the purpose of clinical trials that have sites in the EEA. For such trials the sponsor/legal representative must be based
in the EEA, and the trial must be registered on the EU Clinical Trials Register (including data on sites outside of the EEA).

The  data  exclusivity  periods  in  the  United  Kingdom  are  currently  in  line  with  those  in  the  European  Union,  but  the  Trade  and  Cooperation
Agreement provides that the periods for both data and market exclusivity are to be determined by domestic law, and so there could be divergence in the
future. It is currently unclear whether the MHRA in the United Kingdom is sufficiently prepared to handle the increased volume of marketing authorization
applications that it is likely to receive.

Orphan designation in Great Britain following Brexit is based on the prevalence of the condition in Great Britain as opposed to the current position
where prevalence in the European Union is the determinant. It is therefore possible that conditions that are currently designated as orphan conditions in
Great Britain will no longer be and that conditions that are not currently designated as orphan conditions in the European Union will be designated as such
in Great Britain.

Business Update: COVID-19 Response, Program Prioritization and Corporate Adjustments

With  the  global  spread  of  the  ongoing  coronavirus  2019,  or  COVID-19,  pandemic,  we  established  a  cross-functional  task  force  and  have
implemented business continuity plans designed to address and mitigate its impact on our employees and business. While we have not experienced any
significant  financial  impact  to  date,  the  overall  disruption  caused  by  the  COVID-19  pandemic  on  global  healthcare  systems,  and  the  other  risks  and
uncertainties  associated  with  the  pandemic,  could  cause  our  business,  financial  condition,  results  of  operations  and  growth  prospects  to  be  materially
adversely affected.

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In  March  2020,  our  global  workforce  transitioned  to  working  remotely  with  the  exception  of  clinical  trial  related  activities  that  required
laboratory-based activity or manufacturing. We implemented protocols and procedures to ensure the safety of our employees working on site, including
requirement  to  wear  personal  protective  equipment,  temperature  checks  at  entry  and  offered  COVID-19  testing  for  any  employee  with  symptoms  or  at
suspected  risk  of  exposure  to  virus.  In  June  2020,  we  began  the  implementation  of  our  workplace  re-entry  plan,  based  on  a  phased  approach  that  is
principles-based and local in design, with a focus on continuity of patient treatment and working to bring its workforce back on-site safely. We have also
implemented policies to control and limit office and lab access in line with social distancing guidelines and for contact tracing if needed.

We continue to track COVID-19 developments in Europe and the United States closely for their potential impact on our clinical trial sites, logistics
and supply chain to ensure we can continue to maintain clinical trial conduct and data integrity. As the patients in our clinical trials are severely immune
suppressed as a consequence of their underlying disease and the treatment they receive in the trials, we are also monitoring other transmissible infectious
diseases, including influenza.

In  January  2021,  we  announced  the  prioritization  of  the  AUTO1  program  and  our  intention  to  seek  a  partner  for  the  AUTO3  program  before
progressing AUTO3 into its next phase of development. We also announced an adjustment of our workforce and infrastructure footprint during the first
quarter of 2021, which will involve an overall reduction in headcount of approximately 20%.

C. Organizational structure.

The following diagram illustrates our corporate structure:

In  December  2020,  Autolus  Limited  transferred,  by  way  of  an  interim  distribution  in  kind  to  its  immediate  parent,  Autolus  Holdings  (UK)
Limited, the entire issued share capital of its wholly owned subsidiary, Autolus Inc., our U.S. subsidiary which was incorporated under the laws of the State
of Delaware in October 2017. In anticipation of Brexit, Autolus Limited transferred all of its German assets to Autolus GmbH in December 2020.

D. Property, plant and equipment.

    Our corporate headquarters are located at 58 Wood Lane, White City, London W12 7RZ, United Kingdom, where we lease approximately 14,908 square
feet of office space. The lease is non-cancellable and is scheduled to terminate in September 2025. We have a one-time right, upon six months' notice, to
terminate the lease effective in September 2020. In addition, from September 2020 onward, the landlord has the option to terminate the lease on 12 months'
notice. The landlord exercised its option to give notice in September 2020 to terminate the Forest House lease and pay the Company a break-lease payment
fee in September 2021.

We  also  sublease  a  manufacturing  suite,  consisting  of  approximately  8,750  square  feet  of  manufacturing  space,  at  the  Cell  and  Gene  Therapy
Catapult  manufacturing  center  in  Stevenage,  United  Kingdom.  The  lease  is  non-cancellable  and  is  scheduled  to  terminate  in  September  2023,  with  the
option to renew or terminate the lease in May 2021.

In September 2018, we entered into a binding arrangement for a lease for a manufacturing facility, consisting of approximately 39,558 square feet,

in Enfield, United Kingdom. The lease term is 15 years, commenced in February 2019, with an

89

option  to  terminate  the  lease  in  February  2029.  We  initially  planned  on  initiating  manufacturing  activities  at  this  facility  in  2020;  however,  following  a
strategic review of our manufacturing plan, we chose to discontinue the fit-out of manufacturing capability at the Enfield facility in December 2019.

    In October 2018, we entered into a sublease for 27,502 square feet of office space in Rockville, Maryland. On February 27, 2020, we terminated the
sublease of this office space and concurrently entered into a direct lease with the building owner for the same premises. The lease is non-cancellable and is
scheduled to terminate in March 2025.

In November 2018, we entered into a lease for approximately 32,673 square feet of office space in White City Place, London, due to our increased

headcount. This new space will serve as our new corporate headquarters; we took occupancy in January 2019. The lease term expires in 2026.

In January 2019 we entered into a lease for 84,264 square feet of office and manufacturing space in Rockville, Maryland, under which the lease
term commenced in August 2020 and expires in June 2036.The lease agreement required us to enter into a lease provided that the landlord completed the
required leasehold improvements described in the agreement; the improvements were completed in August 2020.

In May 2020, we executed an arrangement with Catapult Limited to lease a manufacturing suite at the Cell and Gene Therapy Catapult

manufacturing center in Stevenage, United Kingdom for a term through April 2024.

We anticipate leasing additional office and manufacturing space as we add employees, and we believe that suitable additional or substitute space

will be available as needed to accommodate any such expansion of our operations.

Item 4A. Unresolved Staff Comments.

Not applicable.

Item 5. Operating and Financial Review and Prospects.

The  following  discussion  and  analysis  of  our  financial  condition  and  results  of  operations  should  be  read  together  with  our  consolidated  financial
statements and related notes appearing elsewhere in this Annual Report. Some of the information contained in this discussion and analysis or set forth
elsewhere in this Annual Report, including information with respect to our plans and strategy for our business and related financing, includes forward-
looking statements that involve risks and uncertainties. As a result of many factors, including those set forth in the Item 3.D. “Risk Factors” section of this
Annual Report, our actual results could differ materially from the results described in or implied by these forward-looking statements. Please also see the
section titled “Cautionary Statement Regarding Forward-Looking Statements.”

We maintain our books and records in pounds sterling, our results are subsequently converted to U.S. dollars and we prepare our consolidated financial
statements in accordance with U.S. GAAP. All references in this Annual Report to “$” are to U.S. dollars and all references to “£” are to pounds sterling.
Our consolidated balance sheets as of December 31, 2020 and 2019 have been translated from pounds sterling into U.S. dollars at the rate of £1.00 to
$1.3663  and  £1.00  to  $1.3268. Our consolidated statements  of  operations  and  cash  flows  for  the  years  ended  December  31,  2020  and  2019,  the  three
months ended December 31, 2018, and the year ended September 30, 2018 have been translated from pounds sterling to U.S. dollars at the rate of £1.00 to
$1.2862, £1.00 to $1.2738, £1.00 to $1.2870, and £1.00 to $1.3459. These translations should not be considered representations that any such amounts
have been, could have been or could be converted into U.S. dollars at that or any other exchange rate as of that or any other date.

We have historically conducted our business through Autolus Limited, and therefore our historical consolidated financial statements previously presented
the consolidated results of operations of Autolus Limited. Following the completion of our IPO in June 2018, our consolidated financial statements present
the consolidated results of operations of Autolus Therapeutics plc.

We have adopted the amendments to Item 5.A, effective as of February 10, 2021 for the year ended December 31, 2020.

A.

 Operating results.

Overview

90

    
We are a biopharmaceutical company developing next-generation programmed T cell therapies for the treatment of cancer. Using our broad suite of
proprietary  and  modular  T  cell  programming  technologies,  we  are  engineering  precisely  targeted,  controlled  and  highly  active  T  cell  therapies  that  are
designed to better recognize cancer cells, break down their defense mechanisms and attack and kill these cells. We believe our programmed T cell therapies
have  the  potential  to  be  best-in-class  and  offer  cancer  patients  substantial  benefits  over  the  existing  standard  of  care,  including  the  potential  for  cure  in
some patients.

Since our inception in July 2014, we have devoted substantially all of our resources to conducting preclinical studies and clinical trials, organizing
and staffing our company, business planning, raising capital and establishing our intellectual property portfolio. We do not have any products approved for
sale and have not generated any revenue from product sales. We have funded our operations to date primarily with sales of our equity securities, including
American Depositary Shares, or ADSs. As of December 31, 2020, we had received net proceeds of $516.1 million from sales of our equity securities. In
September 2020, we entered into an Open Market Sale Agreement
, or the Sales Agreement, with Jefferies LLC, or Jefferies, where we may offer and sell
ADSs having an aggregate offering price of up to $100 million from time to time through Jefferies, acting as sales agent. As of the date of this Annual
Report,  we  have  sold  an  aggregate  of  1.7  million  of  our  ADSs  resulting  in  net  proceeds  of  $15.3  million.  Further,  in  February  2021,  we  closed  an
underwritten  public  offering  of  16,428,572  ADSs,  including  the  exercise  in  full  by  the  underwriters  of  their  option  to  purchase  an  additional  2,142,857
ADSs, at a public offering price of $7.00 per ADS. We received net proceeds of $108.1 million after underwriting discounts. We do not expect to generate
significant revenue unless and until we obtain marketing approval for and commercialize one of our product candidates.

SM

Since  our  inception,  we  have  incurred  significant operating  losses.  For  the  years  ended  December  31,  2020  and  2019,  the  three  months  ended
December 31, 2018, and the year ended September 30, 2018, we incurred net losses of $142.1 million, $123.8 million, $20.6 million, and $44.8 million,
respectively. As of December 31, 2020, we had an accumulated deficit of $379.2 million.

We expect to continue to incur significant expenses for the foreseeable future as we advance our product candidates through preclinical and clinical
development, seek regulatory approval and pursue commercialization of any approved product candidates. In addition, if we obtain marketing approval for
any  of  our  product  candidates,  we  expect  to  incur  significant  commercialization  expenses  related  to  product  manufacturing,  marketing,  sales  and
distribution. In addition, we may incur expenses in connection with the in-license or acquisition of additional product candidates. Furthermore, we have
incurred and expect to continue to incur, additional costs associated with operating as a public company, including significant legal, accounting, investor
relations and other expenses that we did not incur as a private company.

As a result, we will need substantial additional funding to support our continuing operations and pursue our growth strategy. Until such time as we
can generate significant revenue from product sales, if ever, we expect to finance our operations through the sale of equity, debt financings or other capital
sources, including potential collaborations with other companies or other strategic transactions. We may be unable to raise additional funds or enter into
such other agreements or arrangements when needed on favorable terms, or at all. If we fail to raise capital or enter into such agreements as, and when,
needed, we may have to significantly delay, scale back or discontinue the development and commercialization of one or more of our drug candidates or
delay our pursuit of potential in-licenses or acquisitions.

Based on our current clinical development plans, we believe our existing cash of $153.3 million at December 31, 2020, along with the additional

net proceeds of $123.4 million from sale of our ADSs under our at-the market facility program in January 2021 and our follow-on capital raise in February
2021 will be able to fund our current and planned operating expenses and capital expenditure requirements through at least the next 12 months from the
date of this Annual Report. The forecast of cash resources is forward-looking information that involves risks and uncertainties, and the actual amount of
our expenses could vary materially and adversely as a result of a number of factors. We have based these estimates on assumptions that may prove to be
wrong, and our expenses could prove to be significantly higher than we currently anticipate. Management does not know whether additional financing will
be on terms favorable or acceptable to us when needed, if at all. If adequate additional funds are not available when required, or if we are unsuccessful in
entering into partnership agreements for further development of our product candidates, management may need to curtail its development efforts and
planned operations.

We announced an adjustment of our workforce and infrastructure footprint during the first quarter of 2021, which will involve an overall reduction
in  headcount  of  approximately  20%.  We  expect  to  realize  cash  savings,  on  an  annualized  basis,  of  approximately  $15  million  per  annum  once  the
operational changes are fully implemented. In the short-term, we expect there to be an increase in the first half of 2021 to both research and development
expenses and selling, general, and administrative expenses due to restructuring charges of approximately $2.5 million, combined.

Components of Our Results of Operations

Grant Income

Grant income consists of proceeds from government research grants used to perform specific research and development activities. We recognize

grant income over the period in which we recognize the related costs covered under the terms and conditions

91

of the grant. We have received grants from the U.K. government, which are repayable under certain circumstances, including breach or noncompliance with
the terms of the grant. For grants with refund provisions, we review the grant to determine the likelihood of repayment. If the likelihood of repayment of
the grant is determined to be remote, then the grant is recognized as grant income.

License Revenue

We account for our revenue pursuant to the provisions of Accounting Standards Codification, or ASC, Topic 606, Revenue from Contracts with

Customers, or ASC Topic 606.

We have no products approved for commercial sale and have not generated any revenue from commercial product sales to date. The total revenue
to date has been generated from a license agreement with an investee company of Syncona, our principal shareholder. The terms of the agreement includes
a non-refundable license fee, payments based upon achievement of clinical development and regulatory objectives, and royalties on product sales.

In determining the appropriate amount of revenue to be recognized as we fulfill our obligations under our agreements, we perform the following
steps: (i) identification of the promised goods or services in the contract; (ii) determination of whether the promised goods or services are performance
obligations, including whether they are distinct in the context of the contract; (iii) measurement of the transaction price, including the constraint on variable
consideration; (iv) allocation of the transaction price to the performance obligations based on estimated selling prices; and (v) recognition of revenue when
(or as) the we satisfy each performance obligation.

License Fees and Multiple Element Arrangements

If  a  license  to  our  intellectual  property  is  determined  to  be  distinct  from  the  other  performance  obligations  identified  in  the  arrangement,  we
recognize revenues from non-refundable, upfront fees allocated to the license at such time as the license is transferred to the licensee and the licensee is
able  to  use,  and  benefit  from,  the  license.  For  licenses  that  are  bundled  with  other  promises,  we  utilize  judgment  to  assess  the  nature  of  the  combined
performance  obligations  to  determine  whether  the  combined  performance  obligations  are  satisfied  over  time  or  at  a  point  in  time  and,  if  over  time,  the
appropriate method of measuring progress for purposes of recognizing revenue from non-refundable, upfront fees. We evaluate the measure of progress
each reporting period and, if necessary, adjusts the measure of performance and related revenue recognition.

Appropriate  methods  of  measuring  progress  include  output  methods  and  input  methods.  In  determining  the  appropriate  method  for  measuring
progress, we consider the nature of service that we promise to transfer to the customer. When we decide on a method of measurement, we will apply that
single method of measuring progress for each performance obligation satisfied over time and will apply that method consistently to similar performance
obligations and in similar circumstances.

Contingent Research Milestone Payments

ASC Topic 606 constrains the amount of variable consideration included in the transaction price in that either all, or a portion, of an amount of
variable  consideration  should  be  included  in  the  transaction  price.  The  variable  consideration  amount  should  be  included  only  to  the  extent  that  it  is
probable  that  a  significant  reversal  in  the  amount  of  cumulative  revenue  recognized  will  not  occur  when  the  uncertainty  associated  with  the  variable
consideration is subsequently resolved. The assessment of whether variable consideration should be constrained is largely a qualitative one that has two
elements: the likelihood of a change in estimate, and the magnitude thereof. Variable consideration is not constrained if the potential reversal of cumulative
revenue recognized is not significant, for example.

If the consideration in a contract includes a variable amount, we will estimate the amount of consideration in exchange for transfer of promised
goods or services. The consideration also can vary if our entitlement to the consideration is contingent on the occurrence or non-occurrence of a future
event.  We  consider  contingent  research  milestone  payments  to  fall  under  the  scope  of  variable  consideration,  which  should  be  estimated  for  revenue
recognition purposes at the inception of the contract and reassessed ongoing at the end of each reporting period.

We assess whether contingent research milestones should be considered variable consideration that should be constrained and thus not part of the
transaction price. This includes an assessment of the probability that all or some of the milestone revenue could be reversed when the uncertainty around
whether or not the achievement of each milestone is resolved, and the amount of reversal could be significant.

GAAP provides factors to consider when assessing whether variable consideration should be constrained. All of the factors should be considered,

and no factor is determinate. We consider all relevant factors.

92

Royalty Revenue

For arrangements that include sales-based royalties, including milestone payments based on the level of sales, and the license is deemed to be the
predominant item to which the royalties relate, we recognize revenue at the later of (i) when the related sales occur, or (ii) when the performance obligation
to which some or all of the royalty has been allocated has been satisfied (or partially satisfied).

Operating Expenses

Research and Development Expenses

Research and development expenses consist of costs incurred in connection with the research and development of our product candidates, which
are partially offset by research and development expenditure tax credits provided by Her Majesty’s Revenue & Customs, or HMRC. We expense research
and development costs as incurred. These expenses include:

•

expenses incurred under agreements with contract research organizations, or CROs, as well as investigative sites and consultants that conduct our
clinical trials, preclinical studies and other scientific development services;

• manufacturing scale-up expenses and the cost of acquiring and manufacturing preclinical and clinical trial materials;

•

•

•

•

•

employee-related expenses, including salaries, related benefits, travel and share-based compensation expense for employees engaged in research
and development functions;

expenses incurred for outsourced professional scientific development services;

costs for laboratory materials and supplies used to support our research activities;

allocated facilities costs, depreciation and other expenses, which include rent and utilities; and

upfront, milestone and management fees for maintaining licenses under our third-party licensing agreements.

We recognize external development costs based on an evaluation of the progress to completion of specific tasks using information provided to us by

our service providers.

Our  direct  research  and  development  expenses  are  tracked  on  a  program-by-program  basis  for  our  product  candidates  and  consist  primarily  of
external costs, such as fees paid to outside consultants and CROs in connection with our preclinical development, manufacturing and clinical development
activities. Our direct research and development expenses by program also include fees incurred under license agreements. We do not allocate employee
costs or facility expenses, including depreciation or other indirect costs, to specific programs because these costs are deployed across multiple programs
and, as such, are not separately classified. We use internal resources primarily to oversee research and development as well as for managing our preclinical
development, process development, manufacturing and clinical development activities.

The following table summarizes our research and development expenses incurred by program:

Direct research and development expenses

B cell malignancies (AUTO1 & AUTO3)

Other projects (AUTO 2, 4, 5, 6, 7, & 8)

Total direct research and development expense

Research and discovery expense and unallocated costs:

Personnel related (including share-based compensation)

Indirect research and development expense

Total research and development expenses

Year Ended December 31,

2020-2019

2019-2018

2020

2019

(unaudited)
2018
(in thousands)

Change

Change

$

29,335  $

15,346  $

5,436  $

13,989  $

3,366 

32,701 

4,368 

19,714 

3,998 

9,434 

(1,002)

12,987 

58,171 

44,016 

54,187 

31,517 

21,453 

17,412 

3,984 

12,499 

$

134,888  $

105,418  $

48,299  $

29,470  $

9,910 

370 

10,280 

32,734 

14,105 

57,119 

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    Research and development activities are central to our business model. Product candidates in later stages of clinical development generally have higher
development costs than those in earlier stages of clinical development, primarily due to the increased size and duration of later-stage clinical trials. As a
result, we expect that our research and development expenses will increase substantially over the next few years as we increase personnel costs, initiate and
conduct additional clinical trials and prepare regulatory filings related to our product candidates. We also expect to incur additional expenses related to
milestone, royalty payments and maintenance fees payable to third parties with whom we have entered into license agreements to acquire the rights related
to our product candidates.

The successful development and commercialization of our product candidates is highly uncertain. At this time, we cannot reasonably estimate or
know the nature, timing and costs of the efforts that will be necessary to complete the clinical development of any of our product candidates or when, if
ever, material net cash inflows may commence from sales of any of our product candidates. This uncertainty is due to the numerous risks and uncertainties
associated with development and commercialization activities, including the uncertainty of:

•

•

•

•

•

•

•

•

the scope, progress, outcome and costs of our clinical trials and other research and development activities, including establishing an appropriate
safety profile with IND-directed studies;

successful patient enrollment in, and the initiation and completion of, clinical trials;

the timing, receipt and terms of any marketing approvals from applicable regulatory authorities;

establishing commercial manufacturing capabilities or making arrangements with third-party manufacturers;

development and timely delivery of commercial-grade drug formulations that can be used in our clinical trials and for commercial manufacturing;

obtaining, maintaining, defending and enforcing patent claims and other intellectual property rights;

significant and changing government regulation;

launching commercial sales of our product candidates, if and when approved, whether alone or in collaboration with others;

• maintaining a continued acceptable safety profile of the product candidates following approval; and

•

significant competition and rapidly changing technologies within the biopharmaceutical industry.

We may never succeed in achieving regulatory approval for any of our product candidates. We may obtain unexpected results from our clinical
trials. We may elect to discontinue, delay or modify clinical trials of some product candidates or focus on others. Any changes in the outcome of any of
these variables with respect to the development of our product candidates in clinical development could mean a significant change in the costs and timing
associated with the development of these product candidates. For example, if the EMA, the FDA, or another regulatory authority were to delay our planned
start of clinical trials or require us to conduct clinical trials or other testing beyond those that we currently expect or if we experience significant delays in
enrollment in any of our planned clinical trials, we could be required to expend significant additional financial resources and time on the completion of
clinical development of that product candidate. Commercialization of our product candidates will take several years and millions of dollars in development
costs.

General and Administrative Expenses

General and administrative expenses consist primarily of salaries, related benefits, travel and share-based compensation expense for personnel in
executive, finance, legal and administrative functions. General and administrative expenses also include allocated facility-related costs, patent filing and
prosecution costs and professional fees for marketing, insurance, legal, consulting, accounting and audit services.

We  anticipate  that  our  general  and  administrative  expenses  will  increase  in  the  future  as  we  increase  our  headcount  to  support  the  planned
development  of  our  product  candidates.  Additionally,  if  we  believe  a  regulatory  approval  of  one  of  our  product  candidates  appears  likely,  we  would
anticipate an increase in payroll and expense as a result of our preparation for commercial operations, especially as it relates to the sales and marketing of
our product candidate.

We have experienced, and expect to continue to experience, increased expense with being a public company, including increased accounting, audit,
legal,  regulatory  and  compliance  costs  associated  with  maintaining  compliance  with  Nasdaq  listing  rules  and  SEC  requirements,  director  and  officer
insurance premiums, as well as higher investor and public relations costs.

94

Other Income (Expense)

Other income consists primarily of interest income earned on our cash balances held at a commercial bank and foreign currency transaction gains.

Other expense consists primarily of foreign currency transaction losses.

Income Tax Benefit

We are subject to corporate taxation in the United Kingdom and in the United States. Due to the nature of our business, we have generated losses
since inception. Our income tax credit recognized represents the sum of the research and development tax credits recoverable in the United Kingdom and
income tax payable in the United States.

As a company that carries out extensive research and development activities, we benefit from the U.K. research and development tax credit regime
under the scheme for small or medium-sized enterprises, or SMEs, and also claim a Research and Development Expenditure Credit, or RDEC, to the extent
that our projects are grant funded. Under the SME regime, we are able to surrender some of our trading losses that arise from our qualifying research and
development activities for a cash rebate ranging from 21.67% to 33.35% of such qualifying research and development expenditure. The net tax benefit of
the RDEC is expected to be 10.5% in 2021 and future periods. We meet the conditions of the SME regime, but also can make claims under the RDEC
regime  to  the  extent  that  our  projects  are  grant  funded.  Qualifying  expenditures  largely  comprise  employment  costs  for  research  staff,  consumables,
outsourced CRO costs and utilities costs incurred as part of research projects. Certain subcontracted qualifying research and development expenditures are
eligible for a cash rebate of up to 21.67%. A large portion of costs relating to our research and development, clinical trials and manufacturing activities are
eligible for inclusion within these tax credit cash rebate claims.

We  may  not  be  able  to  continue  to  claim  research  and  development  tax  credits  under  the  SME regime in the future because we  may  no  longer

qualify as a small or medium-sized company. However, we should continue to be able to make claims under the RDEC regime.

Un-surrendered U.K. losses may be carried forward indefinitely to be offset against future taxable profits, subject to numerous utilization criteria
and restrictions. The amount that can be offset each year is limited to £5.0 million plus an incremental 50% of U.K. taxable profits. After accounting for tax
credits receivable, there were accumulated tax losses for carry forward in the United Kingdom of $211.7 million as of December 31, 2020. We currently do
not recognize a deferred tax asset from our accumulated losses and record a full valuation allowance against the net deferred tax asset as the recoverability
due to future taxable profits is unknown.

In the event we generate revenues in the future, we may benefit from the U.K. “patent box” regime that allows profits attributable to revenues from

patents or patented products to be taxed at effective rate of 10%.

Value Added Tax, or VAT, is broadly charged on all taxable supplies of goods and services by VAT-registered businesses. Under current rates, an
amount of 20% of the value, as determined for VAT purposes, of the goods or services supplied is added to all sales invoices and is payable to HMRC.
Similarly, VAT paid on purchase invoices is generally reclaimable from HMRC.

95

Results of Operations

Comparison of Years Ended December 31, 2020 and 2019

The following table summarizes our results of operations for the years ended December 31, 2020 and 2019.

Grant income
License revenue
Operating expenses:

Research and development
General and administrative

          Loss on impairment of leasehold improvements
Total operating expenses, net
Other income (expense):
          Interest income
          Other income (expense)
Total other income, net
Net loss before income tax

Income tax benefit

Net loss attributable to ordinary shareholders

Grant Income

Year Ended December 31,

2020

2019

Change

$

$

$

1,473 
242 

2,908 
— 

$

(134,888)
(34,972)
— 
(168,145)

536 
1,352 
1,888 
(166,257)
24,163 
(142,094)

$

(105,418)
(39,452)
(4,102)
(146,064)

2,542 
4,514 
7,056 
(139,008)
15,159 
(123,849)

$

(1,435)
242 

(29,470)
4,480 
4,102 
(22,081)

(2,006)
(3,162)
(5,168)
(27,249)
9,004 
(18,245)

Grant  income  decreased  to  $1.5  million  for  the  year  ended  December  31,  2020  from  $2.9  million  for  the  year  ended  December  31,  2019.  The
decrease in grant income of $1.4 million was related to a one-time grant completed in the year ended December 31, 2019 submitted to the U.K. government
as part of the reimbursement terms of government research grants used to perform specific research and development activities.

License Revenue

The $0.2 million of license revenue relates to the grant of a license to an investee company of Syncona, our principal shareholder, in the year ended

December 31, 2020.

Research and Development Expenses

Research and development expenses increased to $134.9 million for the year ended December 31, 2020 from $105.4 million for the year ended
December 31, 2019. Cash costs, which exclude depreciation and amortization as well as share-based compensation, increased to $116.9 million from $83.4
million. The increase in research and development cash costs of $33.5 million consisted primarily of (i) an increase of $8.8 million in compensation and
employment related costs, net of lower travel costs, due to an increase in employee headcount to support the advancement of our product candidates in
clinical development, (ii) an increase of $14.4 million in project expenses as a consequence of the advancement of our clinical portfolio which includes
research and process development and manufacturing activities necessary to prepare, activate, and monitor clinical trial programs, (iii) an increase of $6.0
million  in  facilities  costs  related  to  the  commencement  of  a  lease  for  an  additional  manufacturing  suite  and  the  continued  scaling  of  manufacturing
operations,  (iv)  an  increase  of  $4.0  million  in  IT  infrastructure  and  support  for  information  systems  related  to  the  conduct  of  clinical  trials  and
manufacturing operations, (v) an increase of $0.5 million related to legal fees, and (vi) an increase of $1.7 million related to cell logistics, which is offset by
a reduction in materials purchases of $0.7 million and license fees of $1.1 million.

Non-cash costs decreased to $18.1 million for the year ended December 31, 2020 from $22.0 million for the year ended December 31, 2019. The
$3.9 million decrease is related to a decrease of $4.8 million share-based compensation expense as a result of a lower fair value of options recognized in the
period, offset by a $0.9 million increase in depreciation.

96

General and Administrative Expenses

General  and  administrative  expenses  decreased  to  $35.0  million  for  the  year  ended  December  31,  2020  from  $39.5  million  for  the  year  ended
December 31, 2019. Cash  costs,  which  exclude  depreciation  as  well  as  share-based  compensation  increased  to  $27.4  million  from  $26.6  million.  There
were increases of $1.3 million of costs related to D&O insurance and intellectual property costs and $0.1 million of facilities cost, offset by decreases of
$0.5 million of compensation and other employment related costs and $0.1 million in general office expense.

Non-cash costs decreased to $7.6 million for the year ended December 31, 2020 from $12.9 million for the year ended December 31, 2019. The
decrease of $5.3 million is mainly attributed to lower share-based compensation expenses as a result of the lower fair value of stock options recognized
during the period.

Loss on Impairment of Leasehold Improvements

There was no loss on impairment of leasehold improvements for the year ended December 31, 2020 as compared to $4.1 million for the year ended

December 31, 2019 which related to the discontinuation of the fit-out of our manufacturing capacity at the Enfield, U.K. facility.

Interest Income

Interest income decreased to $0.5 million for the year ended December 31, 2020 from $2.5 million for the year ended December 31, 2019. This

decrease is due to the lower cash balances held during the year combined with lower interest rates for cash held on deposit.

Other Income (Expense)

Other income decreased to $1.4 million for the year ended December 31, 2020 from $4.5 million for the year ended December 31, 2019 primarily due to a
weakening of the U.S. dollar exchange rate relative to the pound sterling. The decrease of $4.6 million in the year ended December 31, 2020 was offset by
lease termination gains of $1.5 million.

    Income Tax Benefits

Income tax benefit increased to $24.2 million for the year ended December 31, 2020 from $15.2 million for the year ended December 31, 2019 due
to  additional  U.K.  research  and  development  tax  credits  receivable  from  HMRC.  Research  and  development  credits  are  obtained  at  a  maximum  rate  of
33.35% of our qualifying research and development expenses, and the increase in the net credit was primarily attributable to an increase in our eligible
research and development expenses.

Comparison of Years Ended December 31, 2019 and 2018

The following table summarizes our results of operations for the years ended December 31, 2019 and 2018. The unaudited information for the year
ended  December  31,  2018  has  been  calculated  by  adding  (i)  our  results  for  the  nine  months  ended  September  30,  2018,  derived  from  our  audited
consolidated financial statements for the fiscal year ended September 30, 2018 included in our Annual Report on Form 20-F with the SEC on November
23, 2018 and (ii) our results for the three month transition period ended December 31, 2018, derived from the audited consolidated financial statements
included in our Transition Report on Form 20-F with the SEC on February 25, 2019.

97

Grant income
Operating expenses:

Research and development
General and administrative

          Loss on impairment of leasehold improvements
Total operating expenses, net
Other income (expense):
          Interest income
          Other income (expense)
Total other income, net
Net loss before income tax

Income tax benefit

Net loss attributable to ordinary shareholders

$

Grant Income

Year Ended December 31,

2019

(unaudited)
2018

Change

$

2,908 

$

1,472 

$

(105,418)
(39,452)
(4,102)
(146,064)

2,542 
4,514 
7,056 
(139,008)
15,159 
(123,849)

$

(48,299)
(27,299)
— 
(74,126)

2,011 
5,752 
7,763 
(66,363)
8,488 
(57,875)

$

1,436 

(57,119)
(12,153)
(4,102)
(71,938)

531 
(1,238)
(707)
(72,645)
6,671 
(65,974)

Grant  income  increased  to  $2.9  million  for  the  year  ended  December  31,  2019  from  $1.5  million  for  the  year  ended  December  31,  2018.  The

increase in grant income of $1.4 million was related to an increase in reimbursable expenditures submitted in 2019 to the UK government.

Research and Development Expenses

Research  and  development  expenses  increased  to  $105.4  million  for  the  year  ended  December  31,  2019  from  $48.3  million  for  the  year  ended
December  31,  2018.  Cash  costs,  which  exclude  depreciation  as  well  as  share-based  compensation,  increased  to  $83.4  million  from  $41.5  million.  The
increase  in  research  and  development  cash  costs  of  $41.9  million  consisted  primarily  of  an  increase  in  compensation-related  costs  of  $20.0  million
primarily due to an increase in headcount to support the advancement of our product candidates in clinical development and investment in manufacturing
facilities and equipment, an increase of $4.1 million in research and manufacturing consumables, in part due to the migration and expansion of our research
and  process  development  laboratories  from  Forest  House  to  our  new  location  in  the  Media  Works  facility,  preparations  in  advance  of  any  potential
disruption to supply arrangements that may occur due to Brexit, as well as validation and training costs as part of the start-up at the Catapult facility, an
increase of $10.2 million in facility costs primarily related to the increased leased facilities, an increase of $3.8 million in project expenses related to the
activities  necessary  to  prepare,  activate,  and  monitor  clinical  trial  programs,  an  increase  of  $1.5  million  in  legal  and  professional  fees  that  includes  a
decrease in milestone payments of $0.5 million consisting of a milestone payment payable to UCL Business plc in 2018 and a milestone payment payable
to Noile-Immune Biotech Inc. in 2019, and an increase in IT and general office expenses of $2.3 million.

Non-cash costs increased to $22.0 million for the year ended December 31, 2019 from $6.7 million for the year ended December 31, 2018. The
$15.3 million increase is related to an increase of $12.7 million share-based compensation expense as a result of an increase in headcount, and consequent
increase in the number of options granted, and an increase in the fair value of stock for options expensed during the year ended December 31, 2019, and an
increase  of  $2.6  million  of  depreciation  related  to  the  purchase  of  equipment  to  support  our  clinical  trials  and  research  activities  and  leasehold
improvements.

General and Administrative Expenses

General  and  administrative  expenses  increased  to  $39.5  million  for  the  year  ended  December  31,  2019  from  $27.3  million  for  the  year  ended
December  31,  2018.  Cash  costs,  which  exclude  depreciation  as  well  as  share-based  compensation  increased  to  $26.6  million  from  $21.4  million.  The
increase of $5.2 million consisted primarily of an increase in compensation-related costs of $2.6 million due to an overall increase in headcount, an increase
of $1.9 million in commercial costs, an increase in legal and professional fees of $1.0 million, and an increase of $0.7 million in facility costs related to
lease and maintenance costs, offset in part by a decrease of $1.0 million in IT charges, project expenses, and other office expenses.

Non-cash costs increased to $12.9 million for the year ended December 31, 2019 from $5.9 million for the year ended December 31, 2018. The

increase is primarily due to an increase of $7.3 million share-based compensation expense as a result of an

98

increase in headcount, and consequent increase in the number of options granted, and an increase in the fair value of stock for options expensed during the
period, an increase of $0.1 million of depreciation, offset in part by impairment expense of $0.4 million related to discontinued software that occurred in
2018.

Loss on Impairment of Leasehold Improvements

Loss  on  impairment  of  leasehold  improvements  of  $4.1  million  for  the  year  ended  December  31,  2019  related  to  the  impairment  of  leasehold
improvements as we discontinued the fit-out of our manufacturing capacity at the Enfield, U.K. facility. No such loss of leasehold improvements occurred
in 2018.

Interest Income

Interest income increased to $2.5 million for the year ended December 31, 2019 from $2.0 million for the year ended December 31, 2018 primarily

due to a higher cash balance related to our capital raise activity in 2018 and 2019.

Other Income (Expense)

Other income decreased to $4.5 million for the year ended December 31, 2019 from $5.8 million for the year ended December 31, 2018 primarily due to
foreign currency gains related to the U.S dollar strength relative to the British pound during 2019 as compared to 2018.

    Income Tax Benefits

Income tax benefits increased to $15.2 million for the year ended December 31, 2019 from $8.5 million for the year ended December 31, 2018 due
to  additional  U.K.  research  and  development  tax  credits  receivable  from  HMRC.  Research  and  development  credits  are  obtained  at  a  maximum  rate  of
33.35% of our qualifying research and development expenses, and the increase in the net credit was primarily attributable to an increase in our eligible
research and development expenses.

B. Liquidity and capital resources.

Since our inception, we have not generated any product revenue and have incurred operating losses and negative cash flows from our operations. We
expect to incur significant expenses and operating losses for the foreseeable future as we advance our product candidates through preclinical and clinical
development, seek regulatory approval and pursue commercialization of any approved product candidates. We expect that our research and development
and general and administrative costs will increase in connection with our planned research activities. As a result, we will need additional capital to fund our
operations until such time as we can generate significant revenue from product sales.

We do not currently have any approved products and have never generated any revenue from product sales. We have funded our operations to date
primarily with proceeds from government grants and sales of our preferred and ordinary shares. Through December 31, 2020, we have received aggregate
net cash proceeds of $516.1 million from sales of our equity securities. As of December 31, 2020, we had cash of $153.3 million. We raised additional net
proceeds of $123.4 million from the sale of ADSs under our at-the-market facility in January 2021 and our follow-on capital raise in February 2021.

We currently have no ongoing material financing commitments, such as lines of credit or guarantees, that are expected to affect our liquidity over the
next five years, other than our lease obligations and supplier purchase commitments described below. We also do not currently have any off-balance sheet
arrangements.

ATM Facility

In September 2020, we entered into an Open Market Sale Agreement

, or the Sales Agreement, with Jefferies LLC, or Jefferies, under which we
may, at our option, offer and sell ADSs having an aggregate offering price of up to $100.0 million from time to time through Jefferies, acting as sales agent.
Any such sales, made through our sales agent can be made by any method that is deemed an “at-the-market offering” as defined in Rule 415 promulgated
under the Securities Act, or in other transactions pursuant to an effective shelf registration statement on Form F-3. We agreed to pay Jefferies a commission
of 3.0% of the gross proceeds of any sales of ADSs sold pursuant to the Sales Agreement. During the year ended December 31, 2020, we did not sell any
ADSs under the Sales Agreement.

SM

As of the date of this Annual Report, we have sold an aggregate of 1,718,506 ADSs under the Sales Agreement, for net proceeds of $15.3 million.

99

Cash Flows

The following table summarizes our cash flows for each of the periods presented:

Year Ended December 31,

(in thousands)

2020

$(117,758)
(14,681)
74,415
679
$(57,345)

2019

$(101,484)
(18,668)
108,863
5,164
$(6,125)

(unaudited)
2018

$(42,525)
(13,189)
156,487
(12,202)
$88,571

Net cash used in operating activities
Net cash used in investing activities
Net cash provided by financing activities
Effect of exchange rate changes on cash
Net (decrease) increase in cash

Net Cash Used in Operating Activities

During the year ended December 31, 2020, operating activities used $117.8 million of cash, resulting from our net loss of $142.1 million, offset by
net cash used resulting from changes in our operating assets and liabilities of $1.4 million and by non-cash charges of $22.9 million. The non-cash charges
primarily related to share based compensation of $20.1 million and depreciation of $5.7 million, offset by a gain on lease incentive of $1.3 million and a
gain on lease termination of $0.2 million. Net cash used resulting from changes in our operating assets and liabilities for the year ended December  31,
2020 consisted primarily of a $5.3 million increase in prepaid expenses and other assets, current and non-current and $0.5 million increase in long term
deposits, offset in part by a $7.0 million increase in accounts payable and accrued expenses and other liabilities.

During the year ended December 31, 2019, operating activities used $101.5 million of cash, resulting from our net loss of $123.8 million, and net
cash used resulting from changes in our operating assets and liabilities of $16.2 million, partially offset by non-cash charges of $38.6 million, primarily
related to share based compensation of $30.2 million and leasehold improvement impairment of $4.1 million. Net cash used resulting from changes in our
operating assets and liabilities for the year ended December 31, 2019 consisted primarily of a $21.0 million increase in prepaid expenses and other assets,
current and non-current offset in part by a $3.5 million increase in accounts payable and accrued expenses and other liabilities.

During the year ended December 31, 2018, operating activities used $42.5 million of cash, resulting from our net loss of $57.9 million, offset in
part by net cash provided by changes in our operating assets and liabilities of $3.7 million and non-cash charges of $11.7 million. Net cash used in changes
in our operating assets and liabilities for the year ended December 31, 2018 consisted primarily of a $9.3 million increase in prepaid expenses and other
assets, current and non-current and a $1.3 million increase in long-term deposits, partially offset by a $12.9 million increase in accrued expenses and other
liabilities and a $1.3 million increase in accounts payable.

Net Cash Used in Investing Activities

During the years ended December 31, 2020 and 2019, we used $14.7 million and $18.7 million, respectively, of cash in investing activities, which

consisted primarily of purchases of property and equipment.

During the years ended December 31, 2019 and 2018, we used $18.7 million and $13.2 million, respectively, of cash in investing activities, all of

which consisted primarily of purchases of property and equipment.

Net Cash Provided by Financing Activities

        During  the  years  ended  December  31,  2020  and  2019,  net  cash  provided  by  financing  activities  was  $74.4 million  and  $108.9  million,  respectively,
consisting of net cash proceeds from our January 2020 and April 2019 follow-on capital raises.

During  the  year  ended  December  31,  2019  and  2018,  net  cash  provided  by  financing  activities  was  $108.9  million  and  $156.5  million,

respectively, consisting of net cash proceeds from our April 2019 follow-on capital raise and June 2018 IPO.

100

        
Cash Denomination

The following table reflects cash denominations in U.S. dollars and pound sterling as of:

December 31,

2020

2019

(in thousands)

Total cash held

$

153,299 

$

210,643 

U.S. dollars
British sterling*
* British sterling amount disclosed include immaterial amounts of Swiss franc and Euro account
balances.

62,200 
66,700 

29,800 
136,900 

$
£

$
£

Funding Requirements

We expect our expenses to increase substantially in connection with our ongoing activities, particularly as we advance the preclinical activities and

clinical trials of our product candidates. Our expenses will increase as we:

• seek regulatory approvals for any product candidates that successfully complete preclinical and clinical trials;

• establish  a  sales,  marketing  and  distribution  infrastructure  in  anticipation  of  commercializing  of  any  product  candidates  for  which  we  may  obtain

marketing approval and intend to commercialize on our own or jointly;

• hire additional clinical, medical, and development personnel;

• expand our infrastructure and facilities to accommodate our growing employee base; and

• maintain, expand and protect our intellectual property portfolio.

Our  primary  uses  of  capital  are,  and  we  expect  will  continue  to  be,  compensation  and  related  expenses,  clinical  costs,  external  research  and
development  services,  laboratory  and  related  supplies,  legal  and  other  regulatory  expenses,  and  administrative  and  overhead  costs.  Our  future  funding
requirements will be heavily determined by the resources needed to support development of our product candidates.

Based on our current clinical development plans, we believe our existing cash of $153.3 million at December 31, 2020, along with the net proceeds
from sale of our ADSs in January and February 2021 of $123.4 million, will enable us to fund our current and planned operating expenses and capital
expenditure requirements for at least the next 12 months. We have based these estimates on assumptions that may prove to be wrong, and we could utilize
our available capital resources sooner than we expect. If we receive regulatory approval for our other product candidates, we expect to incur significant
commercialization expenses related to product manufacturing, sales, marketing and distribution, depending on where we choose to commercialize. We may
also require additional capital to pursue in-licenses or acquisitions of other product candidates.

Because  of  the  numerous  risks  and  uncertainties  associated  with  research,  development  and  commercialization  of  pharmaceutical  product
candidates, we are unable to estimate the exact amount of our working capital requirements. Our future funding requirements will depend on and could
increase significantly as a result of many factors, including:

• the scope, progress, outcome and costs of our clinical trials and other research and development activities;

• the costs, timing, receipt and terms of any marketing approvals from applicable regulatory authorities;

• the costs of future activities, including product sales, medical affairs, marketing, manufacturing and distribution, for any of our product candidates for

which we receive marketing approval;

• the revenue, if any, received from commercial sale of our products, should any of our product candidates receive marketing approval;

• the costs and timing of hiring new employees to support our continued growth;

• the costs of preparing, filing and prosecuting patent applications, maintaining and enforcing our intellectual property rights and defending intellectual

property-related claims; and

• the extent to which we in-license or acquire additional product candidates or technologies.

101

Until such time, if ever, that we can generate product revenue sufficient to achieve profitability, we expect to finance our cash needs through equity
offerings.  To  the  extent  that  we  raise  additional  capital  through  the  sale  of  equity,  your  ownership  interest  will  be  diluted.  If  we  raise  additional  funds
through other third-party funding, collaborations agreements, strategic alliances, licensing arrangements or marketing and distribution arrangements, we
may have to relinquish valuable rights to our technologies, future revenue streams, research programs or product candidates or grant licenses on terms that
may not be favorable to us. If we are unable to raise additional funds through equity financings when needed, we may be required to delay, limit, reduce or
terminate our product development or future commercialization efforts or grant rights to develop and market products or product candidates that we would
otherwise prefer to develop and market ourselves.

Lease, Purchase, and Other Obligations
We have operating lease obligations related to our property, plant and equipment. The details of these leases are disclosed in Item 4D. "Property,
Plant and Equipment." within this annual report. The obligations related to the leases arrangements both short- and long-term is given in the Notes to the
Consolidated Financial Statements, specifically, Note 11, "Commitment and Contingencies."

We enter into contracts in the normal course of business with CROs and other third parties for clinical trials and preclinical research studies and
testing. These contracts are generally cancellable by us upon prior notice. Payments due upon cancellation consist only of payments for services provided
or expenses incurred, including noncancellable obligations of our service providers, up to the date of cancellation.

We have contingent payment obligations that we may incur upon achievement of clinical, regulatory and commercial milestones, as applicable, or
royalty payments that we may be required to make under our license agreements with UCLB and Noile-Immune Biotech; however, the amount, timing and
likelihood of such payments are not known as of December 31, 2020.

Critical Accounting Policies and Significant Judgments and Estimates

Our financial statements are prepared in accordance with U.S. GAAP. The preparation of our financial statements and related disclosures requires
us to make estimates and judgments that affect the reported amounts of assets, liabilities, costs and expenses, and the disclosure of contingent assets and
liabilities in our financial statements. We base our estimates on historical experience, known trends and events and various other factors that we believe are
reasonable under the circumstances, the results of which form the basis for making judgments about the carrying values of assets and liabilities that are not
readily apparent from other sources. We evaluate our estimates and assumptions on an ongoing basis. Our actual results may differ from these estimates
under different assumptions or conditions.

While our significant accounting policies are described in more detail in Note 2 to our financial statements appearing at the end of this Annual
Report, we believe that the following accounting policies are those most critical to the judgments and estimates used in the preparation of our financial
statements.

Share-Based Compensation

We issue ordinary shares as well as options and other securities exercisable for or convertible into ordinary shares or ADSs, as incentives to our
employees and directors, and employees and consultants of our subsidiaries. To the extent such incentives are in the form of share options, the options are
granted pursuant to the terms of our 2017 Share Option Plan, or the 2017 Plan, or pursuant to the terms of our 2018 Equity Incentive Plan, or the 2018 Plan.
Options granted under the 2017 Plan and 2018 Plan, as well as shares granted as employee incentives, typically vest over a four-year service period with
25% of the award vesting on the first anniversary of the commencement date and the balance vesting monthly over the remaining three years, unless the
awards contain specific performance vesting provisions. For equity awards issued that have both a performance vesting condition and a services condition,
or performance awards, once the performance criteria is achieved, the performance awards are then subject to a four-year service vesting with 25% of the
performance award vesting on the first anniversary of the performance condition being achieved, with the balance vesting monthly over the remaining three
years. For certain members of senior management and directors, the board has approved an alternative vesting schedule for the equity awards. The options
granted under the 2017 Plan and 2018 Plan generally expire ten years from the date of grant. We expect our share-based compensation expense for awards
granted to employees, directors and other service providers to increase in future periods due to the planned increases in our headcount, particularly as we
move toward commercialization of our product candidates.

We  recognize  compensation  expense  for  equity  awards  based  on  the  grant  date  fair  value  of  the  award.  For  equity  awards  that  vest  based  on  a
service  condition,  the  share-based  compensation  expense  is  recognized  on  a  straight-line  basis  over  the  requisite  service  period.  For  equity  awards  that
contain  both  performance  and  service  conditions,  we  recognize  share-based  compensation  expense  ratably  over  the  requisite  service  period  when  the
achievement of a performance-based milestone is probable based on the

102

relative  satisfaction  of  the  performance  condition  as  of  the  reporting  date.  We  use  the  fair  value  of  our  ordinary  shares  to  determine  the  fair  value  of
restricted share awards.

Share-based  compensation  is  recognized  as  an  expense  in  the  financial  statements  based  on  the  grant  date  fair  value  over  the  requisite  service
period. For awards granted to our employees and directors that vest based on service conditions, we use the accelerated method to allocate compensation
expense to reporting periods. We do not adjust share-based compensation for estimated forfeitures and account for forfeitures when they occur.

We use the Black-Scholes option pricing model to estimate the fair value of share options. This option-pricing model requires the input of various

subjective assumptions, including the option’s expected life and the price volatility of the security.

The fair value of each share option grant is estimated on the date of grant using the Black-Scholes option pricing model and applying assumptions
used in connection with option grants made during the periods covered by these financial statements. Assumptions used in the option pricing model include
the following:

Expected volatility.  We lack company-specific historical and implied volatility information for our ADSs for expected terms greater than 2.5 years.
Therefore, we use a combination of the historical volatility of our ADSs and also the expected share volatility based on the historical volatility of
publicly traded peer companies and expect to continue to do so until such time as we have adequate historical data regarding the volatility of our
own traded security price.

Expected term.    The  expected  term  of  options  granted  represents  the  weighted  average  period  of  time  that  options  granted  are  expected  to  be
outstanding  giving  consideration  to  vesting  schedules  and  our  historical  exercise  patterns.  The  expected  term  of  our  share  options  has  been
determined utilizing the “simplified” method for awards that qualify as “plain-vanilla” options.

Risk-free interest rate.  The risk-free interest rate is determined by reference to the U.S. Treasury yield curve in effect at the time of grant of the
award for time periods that are approximately equal to the expected term of the award.

Expected dividend.  Expected dividend yield of zero is based on the fact that we have never paid cash dividends on ordinary shares and do not
expect to pay any cash dividends in the foreseeable future.

Fair value of ordinary shares.  Options granted after our IPO are issued at the fair market value of our ADSs at the date the grant is approved by the
Board.

Prior to the IPO, we calculated the fair value of our ordinary shares in accordance with the guidelines in the American Institute of Certified Public
Accountants’  Accounting  and  Valuation  Guide,  Valuation  of  Privately-Held-Company  Equity  Securities  Issued  as  Compensation.  The  valuations  of
ordinary shares were prepared using a market approach, based on precedent transactions in the shares, to estimate our total equity value using the option-
pricing method, or OPM, which used a combination of market approaches and an income approach to estimate our enterprise value.

The OPM derives an equity value such that the value indicated is consistent with the investment price, and it provides an allocation of this equity
value to each class of our securities. The OPM treats the various classes of shares as call options on the total equity value of a company, with exercise
prices based on the value thresholds at which the allocation among the various holders of a company’s securities changes. Under this method, each class of
shares has value only if the funds available for distribution to shareholders exceed the value of the share liquidation preferences of the class or classes of
shares with senior preferences at the time of the liquidity event. Key inputs and assumptions used in the OPM calculation include the following:

Expected volatility.    We applied re-levered equity volatility based on the historical unlevered and re-levered equity volatility of publicly traded
peer companies.

Expected dividend.    Expected dividend yield of zero is based on the fact that the Company has never paid cash dividends on ordinary shares and
does not expect to pay any cash dividends in the foreseeable future.

Expected term.    The expected term of the option or the estimated time until a liquidation event.

Risk-free interest rate.    The risk-free interest rate is determined by reference to the U.S. Treasury yield curve for the period commensurate with the
expected of the exit event.

When considering the fair value of options granted in the period prior to the IPO, management considered probability-weighted scenarios based on
the relative likelihoods of completing the IPO and remaining a privately-held company. In the IPO scenarios, the fair value was calculated by dividing our
total estimated equity value by the number of fully diluted ordinary shares outstanding, and then discounting the implied per-share value at a rate intended
to approximate our cost of equity between the share option grant date and the expected IPO date. The stay-private scenario utilized an OPM "Backsolve"
calculation to estimate our equity value implied by the purchase price of the series A preference shares in September 2017.

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Accrued Research and Development Expenses

As  part  of  the  process  of  preparing  consolidated  financial  statements,  the  Company  is  required  to  estimate  accruals  for  research  and  development
expenses. This process involves reviewing and identifying services which have been performed by third parties on the Company’s behalf and determining
the  value  of  these  services.  In  addition,  the  Company  makes  estimates  of  costs  incurred  to  date  but  not  yet  invoiced,  in  relation  to  external  Clinical
Research  Organizations  and  clinical  site  costs.  The  Company  analyzes  the  progress  of  clinical  trials,  including  levels  of  patient  enrollment;  invoices
received and contracted costs, when evaluating the adequacy of the accrued liabilities for research and development. The Company makes judgments and
estimates in determining the accrued balance in any accounting period.

Income Taxes

We  account  for  income  taxes  under  the  asset  and  liability  method  which  includes  the  recognition  of  deferred  tax  assets  and  liabilities  for  the
expected future tax consequences of events that have been included in our financial statements. Under this approach, deferred taxes are recorded for the
future  tax  consequences  expected  to  occur  when  the  reported  amounts  of  assets  and  liabilities  are  recovered  or  paid.  The  provision  for  income  taxes
represents income taxes paid or payable for the current year plus deferred taxes. Deferred taxes result from differences between the financial statements and
tax bases of our assets and liabilities, and are adjusted for changes in tax rates and tax laws when changes are enacted. The effects of future changes in
income tax laws or rates are not anticipated.

We are subject to corporation taxes in the United Kingdom and the United States. The calculation of our tax provision involves the application of

U.K. tax law and requires judgment and estimates.

We evaluate the realizability of our deferred tax assets at each reporting date. The ultimate realization of deferred tax assets is dependent upon the
generation of future taxable income of the same character and in the same jurisdiction. We consider all available positive and negative evidence in making
this assessment, including, but not limited to, the scheduled reversal of deferred tax liabilities, projected future taxable income, and tax planning strategies.
In circumstances where there is sufficient negative evidence indicating that our deferred tax assets are not more likely than not realizable, we establish a
valuation allowance.

We use a two-step approach to recognizing and measuring uncertain tax positions. The first step is to evaluate tax positions taken or expected to be
taken in a tax return by assessing whether they are more likely than not sustainable, based solely on their technical merits, upon examination, and including
resolution of any related appeals or litigation process. The second step is to measure the associated tax benefit of each position as the largest amount that
we believe is more likely than not realizable. Differences between the amount of tax benefits taken or expected to be taken in our income tax returns and
the amount of tax benefits recognized in our financial statements represent our unrecognized income tax benefits, which we either record as a liability or as
a reduction of deferred tax assets.

Deferred Tax and Current Tax Credits

Tax on the profit or loss for the year comprises current and deferred tax. Tax is recognized in the statement of operations, except to the extent that it
relates to items recognized directly in equity, in which case it is recognized in equity. Current tax is the expected tax payable on the taxable income or loss
for the year, using tax rates enacted or substantively enacted at the balance sheet date, and any adjustment to tax payable in respect of previous years. Tax
credits are accrued for the year based on calculations that conform to the U.K. research and development tax credit regime, under both the SME and large
company regimes. We meet the conditions of the SME regime, but also can make claims under the RDEC regime to the extent that our projects are grant
funded.

We  may  not  be  able  to  continue  to  claim  research  and  development  tax  credits  under  the  SME  regime  in  the  future  because  we  may  no  longer

qualify as a small or medium-sized company. However, we should continue to be able to make claims under the RDEC regime.

Deferred tax is recognized on temporary differences between the carrying amounts of assets and liabilities for financial reporting purposes and the
amounts used for taxation purposes. The amount of deferred tax is based on the expected manner of realization or settlement of the carrying amount of
assets and liabilities, using tax rates enacted or substantively enacted at the balance sheet date. A deferred tax asset is recognized only to the extent that it is
probable that future taxable profits will be available against which the asset can be utilized. No deferred tax assets are recognized on our losses carried
forward and other attributes because there is currently no indication that we will make sufficient profits to utilize these attributes.

JOBS Act

On April 5, 2012, the Jumpstart Our Business Startups Act, or the JOBS Act, was enacted. The JOBS Act provides that, among other things, an
emerging growth company can take advantage of an extended transition period for complying with new or revised accounting standards. As an emerging
growth company, we have irrevocably elected not to take advantage of the extended

104

 
transition period afforded by the JOBS Act for implementation of new or revised accounting standards and, as a result, we will comply with new or revised
accounting standards on the relevant dates on which adoption of such standards is required for non-emerging growth public companies.

In addition, we intend to rely on the other exemptions and reduced reporting requirements provided by the JOBS Act. Subject to certain conditions
set forth in the JOBS Act, we are entitled to rely on certain exemptions as an “emerging growth company,” we are not required to, among other things,
(i)  provide  an  auditor’s  attestation  report  on  our  system  of  internal  controls  over  financial  reporting  pursuant  to  Section  404(b),  (ii)  provide  all  of  the
compensation  disclosure  that  may  be  required  of  non-emerging  growth  public  companies  under  the  Dodd-Frank  Wall  Street  Reform  and  Consumer
Protection Act, (iii) comply with any requirement that may be adopted by the Public Company Accounting Oversight Board regarding mandatory audit
firm rotation or a supplement to the auditor’s report providing additional information about the audit and the financial statements (auditor discussion and
analysis),  and  (iv)  disclose  certain  executive  compensation-related  items  such  as  the  correlation  between  executive  compensation  and  performance  and
comparisons  of  the  chief  executive  officer’s  compensation  to  median  employee  compensation.  These  exemptions  will  apply  for  a  period  of  five  years
following the completion of our IPO or until we no longer meet the requirements of being an emerging growth company, whichever is earlier.

Recently Issued Accounting Pronouncements

A description of recently issued accounting pronouncements that may potentially impact our financial position and results of operations is

disclosed in Note 2, “Summary of Significant Accounting Policies,” to our consolidated financial statements included in this Annual Report.

C. Research and development, patents and licenses, etc.

Full details of our research and development activities and expenditures are given in Item 4.B. "Information on the Company – Business Overview”
and Item 5.A. "Operating Results” within this Annual Report.

D. Trend information.

See Item 5.A. "Operating Results” and Item 5.B. "Liquidity and Capital Resources” within this Annual Report.

E. Off-balance sheet arrangements.

We have elected to voluntarily comply with Item 5.E of the Instructions to Form 20-F, as effective February 10, 2021 and are omitting this disclosure
in reliance thereon.

F. Tabular disclosure of contractual obligations.

We have elected to voluntarily comply with Item 5.F of the Instructions to Form 20-F, as effective February 10, 2021 and are omitting this disclosure
in reliance thereon. These disclosures are now incorporated into Item 5.B. "Liquidity and Capital Resources” within this Annual Report.

G. Safe harbor.

This Annual Report contains forward-looking statements within the meaning of Section 27A of the Securities Act and Section 21E of the Exchange

Act and as defined in the Private Securities Litigation Reform Act of 1995. See the section titled “Cautionary Statement Regarding Forward-Looking
Statements” at the beginning of this Annual Report.

Item 6. Directors, Senior Management and Employees

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A. Directors and senior management.

The following table sets forth information regarding members of our senior management and our directors, including their ages as of March 1, 2021.

NAME

Senior Management:
Christian Itin, Ph.D.
Andrew Oakley
Martin Pulé, MBBS
Muhammad Al-Hajj, Ph.D.
Christopher Vann
Matthias Alder
    David Brochu

Non-Executive Directors:
Joseph Anderson, Ph.D.
Jay Backstrom, M.D., MPH
Linda Bain
John Berriman
Cynthia Butitta
Kapil Dhingra, M.D.
Martin Murphy, Ph.D.

Senior Management

AGE

POSITION(S)

56
58
48
50
56
56
65

61
66
50
72
66
61
52

Chief Executive Officer and Chairman of the Board of Directors
Senior Vice President, Chief Financial Officer
Senior Vice President, Founder, Chief Scientific Officer
Senior Vice President, Translational Sciences
Senior Vice President, Chief Operating Officer
Senior Vice President, Chief Business Officer and Company Secretary

    Senior Vice President, Chief Technical Officer

Director
Director
Director
Director
Director
Director
Director

    Christian Itin, Ph.D. has served as our Chief Executive Officer since March 2016 and as Chairman of our board of directors since October 2014. Prior to
joining us, Dr. Itin served as chief executive officer and chairman of the board of directors at Cytos Biotechnology Ltd, a biotechnology company, from
November 2012 until it merged with Kuros Biosurgery Holding Ltd in January 2016. From January 2016 until June 2018, he served as chairman, and from
June 2018 to May 2019 as non-executive director, of Kuros Biosciences Ltd. Prior to that, Dr. Itin served as president, chief executive officer and director
of Micromet, Inc., a biopharmaceutical company, from 2006 until it was acquired by Amgen Inc. in 2012. From 1999 until 2006, he served in a number of
capacities with Micromet, Inc.’s subsidiary, Micromet AG, including head of IP and licensing, vice president of business and corporate development, chief
business officer and ultimately as its chief executive officer. Before joining Micromet, Dr. Itin was a co-founder of Zyomyx, a protein chip company. Dr.
Itin also serves as a non-executive director of Kymab Ltd., a privately held biopharmaceutical company. Dr. Itin received a Diploma in Biology and a Ph.D.
in Cell Biology from the University of Basel, Switzerland. In addition, he also performed post-doctoral research at the Biocenter of University of Basel and
at the Stanford University School of Medicine. We believe that Dr. Itin is qualified to serve on our board of directors because of his deep knowledge of our
company and his extensive experience serving in executive and non-executive leadership positions at other public and private biotechnology companies.

    Andrew Oakley has served as our Chief Financial Officer since June 2018. Prior to joining Autolus, Mr. Oakley served as the chief financial officer for
Sosei Group from February 2017 until June 2018 and as its executive vice president from August 2017 to June 2018. From January 2015 until June 2016,
Mr. Oakley served as chief financial officer, company secretary and executive board member of Vectura group plc. From January 2003 until August 2013,
Mr. Oakley served as the chief financial officer for Actelion Ltd. He also previously served in a senior finance capacity for the global holding companies of
Accenture,  held  executive  positions  in  major  multinational  building  material  companies  and  spent  several  years  as  an  equity  analyst  with  banks  in
Australia, the United Kingdom and the United States. Mr. Oakley holds a Bachelor of Economics Degree from Macquarie University and an M.B.A. from
London Business School. Mr. Oakley is a member of the Australian Institute of Chartered Accountants.

Martin Pulé, MBBS has served as our Senior Vice President, Founder and Chief Scientific Officer since August 2014. He also served as a member
of our board of directors from August 2014 to June 2018. Dr. Pulé has served as a clinical senior lecturer in the Department of Haematology at University
College London Cancer Institute since 2010 and as an Honorary Consultant in Haematology at University College London Hospital since 2010. He entered
the T cell engineering field in 2001 as a travelling

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Fulbright Scholar at the Center for Cell and Gene Therapy at Baylor College of Medicine, Houston, Texas. Dr. Pulé holds a Bachelor of Medicine and
Bachelor of Surgery (MBBS) from University College Dublin and is a Fellow of the Royal College of Pathologists.

Muhammad Al-Hajj, Ph.D. has served as our Senior Vice President, Translational Sciences since July 2017. Prior to joining us, he served as vice
president,  discovery  and  translational  medicine  at  Sanford  Burnham  Medical  Institute  from  July  2015  to  July  2017.  Prior  to  that,  he  served  as  senior
director, biology and translational medicine in oncology research and development at GlaxoSmithKline plc from 2009 to June 2015. His other experience at
large pharmaceutical companies includes serving as group leader in oncology research and development at AstraZeneca AB from 2007 to 2009 and as lab
head and group leader in oncology research and development at Novartis from 2003 to 2007. Dr. Al-Hajj earned his B.S. in mathematics and biology from
the American University of Beirut. He holds a Ph.D. in molecular genetics from the Wayne State University and completed a postdoctoral fellowship in
cancer and stem cell biology at the University of Michigan Medical School.

Christopher  Vann  has  served  as  our  Senior  Vice  President,  Chief  Operating  Officer  since  October  2016.  Prior  to  joining  us,  he  worked
at  Hoffmann-La  Roche’s  Swiss  headquarters  from  February  1994  to  September  2016,  most  recently  serving  as  its  commercial  director  from  December
2011 to September 2016 where he was primarily responsible for leading the lung cancer commercial team and general management of the Tarceva brand.
Mr. Vann has significant experience of global lifecycle management of oncology products as well as implementing marketing strategy at a regional and
national  level.  This  includes  launching  several  oncology,  immunology  and  transplant  products  in  the  United  States,  United  Kingdom,  Romania,  Russia,
South Africa and countries in Asia, including Japan. Mr. Vann holds a B.S. in Toxicology and Pharmacology from the School of Pharmacy, University of
London.

Matthias  Alder  has  served  as  our  Senior  Vice  President,  Chief  Business  Officer  and  Company  Secretary  since  July  2017,  and  as  our  General
Counsel  between  July  2017  and  August  2018.  Prior  to  joining  us,  he  served  as  executive  vice  president  for  business  development  and  licensing  from
October  2014  to  March  2017  and  as  general  counsel  and  corporate  secretary  from  May  2015  to  July  2017  at  Sucampo  Pharmaceuticals,  Inc.,  a
biopharmaceutical company which was subsequently acquired by Mallinckrodt Pharmaceuticals. Prior to this, Mr. Alder served as executive vice president
of corporate development and legal affairs and corporate secretary at Cytos Biotechnology AG, a biopharmaceutical company focused on the development
of  targeted  immunotherapies,  from  2013  to  October  2014.  From  2006  to  2012,  Mr. Alder  held  various  executive  management  roles  at  Micromet,  Inc.,
serving as senior vice president for administration, general counsel and secretary at the time of the acquisition of Micromet by Amgen Inc. in 2012. He was
also  a  partner  in  the  Life  Sciences  Transactions  Practice  at  Cooley  LLP  from  1997  to  2006,  where  he  represented  biotech  companies  in  strategic
transactions with pharmaceutical companies. Earlier in his career, Mr. Alder was in-house counsel at Ciba-Geigy and Novartis. Mr. Alder holds law degrees
from the University of Basel and the University of Miami and is qualified to practice law in Switzerland and the United States.

David Brochu has served as our Senior Vice President, Chief Technical Officer since January 2021, our Senior Vice President, Head of Product
Delivery between October 2019 and his promotion, and our Vice President of Technical Operations between March and October 2019. David previously
served  at  Kedrion  USA  as  vice  president  of  technical  operations  and  program  head  for  Kedrion  SpA’s  next  generation  IVIG  development  and
industrialization  effort.  In  this  position,  David  also  established  various  operations  functions  for  the  company,  including  manufacturing,  supply  chain,
engineering,  quality,  regulatory,  program  management,  and  development  in  support  of  this  effort.  Prior  to  this,  he  was  the  vice  president  of  plasma
collection operations for Talecris Biotherapeutics (formerly Bayer HealthCare LLC), where he led the operations buildout in the Western United States.
Earlier in his career, David was the senior director of process development and technology at Talecris leading the preparation and implementation of the
technology, process development, and facilities investment strategy for the plasma products business. In this role he led teams of scientists, engineers and
technicians  in  the  development  of  new  protein  therapeutics  from  research  to  commercial  launch.  Prior  to  Talecris,  he  held  engineering  and  technical
operations  leadership  roles  at  Bayer  and  Warner  Lambert  in  the  US,  EU  and  South  America.  David  has  over  30  years  of  operational  and  development
experience. He received a B.S. in chemical engineering from Northeastern University.

Non-Executive Directors

Joseph Anderson, Ph.D. has  served  on  our  board  of  directors  since  February  2016.  He  is  a  Partner  in  the  Sofinnova  Crossover  Fund,  which  he
joined in October 2020. Previously, he was the chief executive officer and a member of the board of directors of Arix Bioscience plc, a global life sciences
company,  where  he  held  similar  positions  since  January  2016.  He  has  founded  and  managed  public  equity  funds  and  been  a  member  of  the  following
boards  of  directors:  Algeta  ASA  (acquired  by  Bayer  AG)  from  2009  to  2013,  Amarin  plc  from  October  2009  to  2013,  Cytos  Biotechnology  Ltd,  a
biotechnology company, from 2012 until it merged with Kuros Biosurgery Holding Ltd in January 2016 and Epigenomics AG from 2012 to 2014. He was a
partner at Abingworth LLP, an international investment group dedicated to the life sciences and healthcare sectors, from January 2004 through December
2015. From October 1999 through December 2003, Dr. Anderson was previously at First State Investments in London, part of the Commonwealth Bank of
Australia,  where  he  was  head  of  global  healthcare  equities  and  portfolio  manager.  Prior  to  this,  he  was  a  pharmaceuticals  analyst  at  investment  bank,
Dresdner Kleinwort Benson from June 1998 through October 1999. From 1990 to 1998, Dr. Anderson

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established  and  was  head  of  the  strategy  unit  at  The  Wellcome  Trust,  one  of  the  world’s  largest  medical  foundations.  Dr.  Anderson  holds  a  Doctor  of
Philosophy  in  Biochemistry  from  the  University  of  Aston  and  a  Bachelor  of  Science  in  Biological  Science  from  Queen  Mary  College,  University  of
London.  He  was  originally  nominated  to  our  board  of  directors  by  Arix  Bioscience  Holdings  Limited  pursuant  to  our  March  2016  Subscription  and
Shareholders’  Agreement,  which  granted  Arix  the  right  to  appoint  one  individual  as  a  director,  and  re-elected  to  a  three-year  term  at  our  2019  Annual
General Meeting of Shareholders. We believe that Dr. Anderson is qualified to serve on our board of directors because of his extensive experience serving
on boards of directors of life science companies.

Jay  T.  Backstrom,  MD,  MPH  has  served  on  our  board  of  directors  since  August  2020.  Dr.  Backstrom  has  served  as  Executive  Vice  President,
Research and Development, at Acceleron Pharma since December 2019. He previously served as the Chief Medical Officer for the Celgene Corporation
since April 2016. Dr. Backstrom joined Celgene in March 2008 as vice president of clinical research and development for the MDS and AML therapeutic
area. He subsequently served as senior vice president of clinical research and development for hematology and oncology and the head of global regulatory
affairs  before  being  appointed  the  Chief  Medical  Officer,  a  position  he  held  up  through  the  completion  of  the  Bristol  Myers  Squibb-Celgene  merger  in
December 2019. After earning his medical degree at the Lewis Katz School of Medicine at Temple University in Philadelphia, he completed his medical
training  including  serving  as  Chief  Medical  Resident  in  the  Department  of  Medicine  at  Temple  University  Hospital.  Dr,  Backstrom  received  a  master’s
degree in public health from Saint Louis University. We believe that Dr. Backstrom is qualified to serve on our board of directors because of his extensive
experience as a pharmaceutical company executive and in the clinical development and regulation of pharmaceuticals, including CAR T therapies.

Linda Bain has served on our board of directors since June 2018. She currently serves as the chief financial officer of Codiak BioSciences, Inc., a
position she has held since December 2015. She has also served as a non-executive director of Arvinas, Inc. since June 2020. Prior to joining Codiak, Ms.
Bain served as the chief financial officer and treasurer of Avalanche Biotechnologies, Inc. from April 2014 until November 2015. Previously, Ms. Bain
served at bluebird bio, Inc., a gene therapy biotechnology company, as vice president of finance and business operations from October 2011 to March 2014,
and chief accounting officer and treasurer from June 2013 to March 2014. From September 2008 to September 2011, Ms. Bain served as vice president of
finance at Genzyme Corporation. From September 2007 to September 2008, she served as vice president at Fidelity Investments, and from May 2000 to
September  2007,  she  held  a  number  of  positions  at  AstraZeneca  plc.  She  received  her  B.S.  degree  in  Accounting  and  Business  Administration  and  an
Honors Degree in Accounting and Business Administration from the University of the Free State in South Africa. Ms. Bain is a certified public accountant.
We believe that Ms. Bain is qualified to serve on our board of directors because of her extensive experience in our industry, her background in accounting
and finance and her leadership skills.

John Berriman has served on our board of directors since August 2014. He has served as chairman of the boards of directors of Confo Therapeutics
NV since December 2016, Depixus SAS since December 2015, and Autifony Therapeutics Ltd since 2011. He previously served as chairman of the board
of directors of ReNeuron Group plc between April 2015 and September 2020; as chairman of the board of directors of Heptares Therapeutics Ltd from
2007 until it was sold to Sosei Group in February 2015; as chairman of the board of directors of Algeta ASA from 2004 through its listing on the Oslo
Stock Exchange in 2007 and subsequently served as deputy chairman from 2008 until it was sold to Bayer AG in 2014; and as a director of Micromet, Inc.
from May 2006 until it was sold to Amgen Inc. in 2012. Prior to this, from 1997 to 2004, he was a director of Abingworth Management, an international
healthcare venture capital firm, where he was involved in founding, financing and serving as a director of several biotechnology companies in Europe and
the United States, many of which obtained listings on public stock exchanges. Prior to that, Mr. Berriman spent 14 years with Celltech Group plc and was a
member of its board when it listed on the London Stock Exchange in 1994. He holds a Master’s degree in Chemical Engineering from the University of
Cambridge and an M.B.A. from the London Business School. We believe that Mr. Berriman is qualified to serve on our board of directors because of his
extensive experience in our industry, including his strategic management and operational experience, his experience serving on public company boards and
his experience with public offerings, private investments and mergers.

Cynthia Butitta has served on our board of directors since March 2018. Ms. Butitta served as the executive vice president and chief financial officer
of Kite Pharma Inc., a biopharmaceutical company, from January 2014 to May 2016 and as its chief operating officer from March 2014 to September 2017.
From May 2011 to December 2012, she was senior vice president and chief financial officer at NextWave Pharmaceuticals, Inc., a specialty pharmaceutical
company. Prior to that, Ms. Butitta served as chief operating officer of Telik, Inc., a biopharmaceutical company, from March 2001 to December 2010 and
as its chief financial officer from August 1998 to December 2010. Ms. Butitta also served as principal accounting officer of Telik, Inc. until December
2010. She has served as a director of publicly held biopharmaceutical companies UroGen Pharma Ltd. since October 2017, and Olema Pharmaceuticals
Inc. since August 2020. Ms. Butitta holds a B.S. degree with honors in Business and Accounting from Edgewood College in Madison, Wisconsin and an
M.B.A. in Finance from the University of Wisconsin, Madison. We believe that Ms. Butitta is qualified to serve on our board of directors because of her
extensive financial and operational experience within the biotechnology and high-technology industries, as well as her leadership skills.

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Kapil Dhingra, M.D. has served on our board of directors since August 2014. Dr. Dhingra currently serves as the managing member of KAPital
Consulting, LLC, a healthcare consulting firm that he founded in June 2008. Dr. Dhingra has over 25 years of experience in oncology clinical research and
drug development. From 1999 to 2008, Dr. Dhingra worked at Hoffmann-La Roche,where he served in roles of increasing responsibility, most recently as
vice president, head of the oncology disease biology leadership team and head of oncology clinical development. From 2000 to 2008, he held a clinical
affiliate appointment at Memorial Sloan Kettering Cancer Center. From 1996 to 1999, Dr. Dhingra worked at Eli Lilly and Company where he served in
roles of increasing responsibility, most recently as senior clinical research physician. Dr. Dhingra also served as a clinical associate professor of medicine at
the  Indiana  University  School  of  Medicine  from  1997  to  1999.  Prior  to  Eli  Lilly  and  Company,  Dr.  Dhingra  was  a  member  of  the  faculty  of  the  MD
Anderson  Cancer  Center  of  the  University  of  Texas  from  1989  to  1996.  Dr.  Dhingra  has  served  on  the  boards  of  directors  of  Replimune  Limited,  a
biotechnology company, since July 2017, Median Technologies, a medical imaging software company, since June 2017, Five Prime Therapeutics, Inc., a
biotechnology company, since December 2015, and Black Diamond Therapeutics, Inc. since January 2021. Dr. Dhingra previously served as a member of
the boards of directors of BioVex from 2009 until its acquisition by Amgen Inc. in 2011, Micromet, Inc. from February 2009 until its acquisition by Amgen
Inc. in 2012, YM Biosciences Inc. from 2012 until its acquisition by Gilead Sciences, Inc. in February 2013, Algeta ASA from 2010 until its acquisition by
Bayer in March 2014 and EpiTherapeutics ApS from January 2014 until its acquisition by Gilead in May 2015, Advanced Accelerator Applications S.A., a
pharmaceutical company, from April 2014 until its acquisition by Novartis in January 2018, and Exosome Diagnostics from 2012 until its acquisition by
Bio-Techne  Corporation  in  August  2018.  Dr.  Dhingra  holds  an  M.D.  from  the  All  India  Institute  of  Medical  Services  in  New  Delhi,  India  and  has
performed postgraduate work at the All India Institute of Medical Services, the Lincoln Medical and Mental Health Center of New York Medical College
and Emory University School of Medicine. We believe that Dr. Dhingra is qualified to serve on our board of directors because of his extensive experience
in executive positions with several pharmaceutical companies and in the clinical development of pharmaceuticals in several therapeutic areas, including in
oncology, and his experience serving on the boards of several publicly traded life science companies.

Martin  Murphy,  Ph.D.  has  served  on  our  board  of  directors  since  September  2014.  He  has  served  as  the  chief  executive  officer  of  Syncona
Investment Management Limited, part of the global life science company Syncona Ltd, since December 2016 and previously founded Syncona Partners
LLP and served as its chief executive officer from May 2012 to December 2016. Previously, he was a partner at MVM Life Science Partners LLP, a venture
capital  company  focused  on  life  science  and  healthcare  investments,  from  2003  to  2012.  During  his  time  at  MVM,  Dr.  Murphy  was  a  member  of  the
management  and  investment  committees  and  led  MVM’s  European  operations.  Before  MVM,  Dr.  Murphy  worked  at  3i  Group  plc  and  McKinsey  &
Company. He has a Ph.D. in Biochemistry from the University of Cambridge. Dr. Murphy was originally nominated to our board of directors by Syncona
Portfolio Limited pursuant to our September 2014 Subscription and Shareholders’ Agreement, which granted Syncona the right to appoint two individuals
as directors, and was re-elected to a three-year term at our 2019 Annual General Meeting of Shareholders. We believe that Dr. Murphy is qualified to serve
on our board of directors because of his extensive experience as an investor, particularly in the life sciences industry.

Family Relationships

There are no family relationships among any of our senior management or our directors.

B. Compensation.

The following discussion provides the amount of compensation paid, and benefits in-kind granted, by us and our subsidiaries to our directors, members of
our senior management and non-employee directors for services in all capacities to us and our subsidiaries for the year ended December 31, 2020, as well
as the amount contributed by us or our subsidiaries into money purchase plans for the year ended December 31, 2020 to provide pension, retirement or
similar benefits to, our directors, members of our senior management and non-employee directors.

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Director Compensation

For the year ended December 31, 2020, the table below sets forth the compensation paid to our directors. In the case of Dr. Itin, our chief executive
officer, the table below sets forth the compensation paid to him for services as a member of our senior management. He does not receive any compensation
for serving as an executive director. All such amounts are established and paid in pounds sterling.

Name

Christian Itin, Ph.D.

Executive Director

Joseph Anderson, Ph.D.

Non-Executive Director

Jay Backstrom, M.D, MPH

Non-Executive Director

Linda Bain

Non-Executive Director

John Berriman

Non-Executive Director

Cynthia Butitta

Non-Executive Director

Kapil Dhingra, M.D.

Non-Executive Director

Martin Murphy, Ph.D.

Non-Executive Director

Salary/Fees

Annual Bonus

Benefit

Compensation

Total

Pension

All Other

£

£

£

£

£

£

£

£

381,615  £

191,344  £

—  £

2,543,717  £

3,116,676 

37,050  £

—  £

—  £

86,766  £

123,816 

16,500  £

—  £

—  £

73,139  £

89,639 

42,750  £

—  £

—  £

133,006  £

175,756 

48,450  £

—  £

—  £

104,653  £

153,103 

38,475  £

—  £

—  £

117,873  £

156,348 

45,600  £

—  £

—  £

104,653  £

150,253 

32,775  £

—  £

—  £

86,766  £

119,541 

Non-Executive Letters of Appointment

Non-executive directors are engaged on letters of appointment that set out their duties and responsibilities. The non-executive directors do not receive

benefits upon termination or resignation from their respective positions as directors.

Non-Executive Director Compensation Policy

In January 2020, following market research and advice from its compensation consultant, our board of directors amended our non-executive director

compensation policy.

Under this policy, we pay each of our non-executive directors a cash retainer for service on our board of directors and committees of our board of
directors.  Our  lead  independent  director  also  receives  an  additional  cash  retainer.  These  retainers  are  payable  in  arrears  in  twelve  equal  monthly
installments at the end of each calendar month, provided that the amount of such payment will be prorated for any portion of such month that the director is
not serving on our board. Non-executive directors residing outside the United Kingdom will be paid the applicable amounts converted from pounds sterling
into a currency of their request at the time of payment. We will also reimburse our directors for their reasonable out-of-pocket expenses in connection with
attending board and committee meetings.

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Non-executive directors are eligible to receive cash compensation as follows:

Annual retainer
Additional retainer for lead independent director
Additional retainer for audit committee chair
Additional retainer for audit committee member
Additional retainer for compensation committee chair
Additional retainer for compensation committee member
Additional retainer for nominating and governance committee chair
Additional retainer for nominating and governance committee member
Additional retainer for research and development committee chair
Additional retainer for research and development committee member

Equity Compensation

Annual Cash Retainer
(£)

30,000 
12,000 
12,000 
6,000 
9,000 
4,500 
6,000 
3,000 
12,000 
6,000 

In addition to cash compensation, each non-executive director is eligible to receive share options under our equity incentive plans. Any share options
granted under this policy shall have a term of ten years from the date of grant, subject to earlier termination in connection with a termination of service.
Vesting schedules for equity awards are subject to the non-executive director’s continuous service on each applicable vesting date.

Notwithstanding  any  vesting  schedule,  for  each  non-executive  director  who  remains  in  continuous  service  with  us  until  immediately  prior  to  the
closing of a change in control (as such term is defined in our 2018 Plan), the shares subject to his or her then-outstanding initial or annual equity awards
that were granted pursuant to this policy will become fully vested immediately prior to the closing of such change in control.

Upon the termination of the membership of the non-executive director on the board for any reason, his or her options granted under this policy shall
remain exercisable for three months following his or her date of termination (or such longer period as the board may determine in its discretion on or after
the date of grant of such options).

Initial Award

Each new non-executive director elected to our board of directors will be granted an initial, one-time equity award of options to purchase 25,000 of our
ADSs  or  ordinary  shares  on  the  date  of  such  director’s  initial  election  or  appointment  to  the  board  of  directors,  which  will  vest  in  equal  monthly
installments through the third anniversary of the grant date.

Annual Awards

On the date of each of our annual meeting of shareholders, each non-executive director that continues to serve will be granted an option to purchase

12,500 of our ADSs or ordinary shares, which will vest in equal monthly installments through the first anniversary of the grant date.

Senior Management Compensation

The compensation for each member of our executive management comprises the following elements: base salary, annual bonus, personal benefits,
pension or 401(k) plan and long-term incentives. For the year ended December 31, 2020, the aggregate compensation accrued or paid to the members of
our  senior  management  for  services,  whether  or  not  a  director,  in  all  capacities  was  $10.6  million.  The  amount  set  aside  or  accrued  by  us  to  provide
pension, retirement or similar benefits to members of senior management amounted to a total of $8,100 in the year ended December 31, 2020.

Bonus Plan

Management Incentive Compensation Plan

On May 17, 2016, the board of directors adopted the Management Incentive Compensation Plan. The Management Incentive Compensation Plan is

designed to offer annual incentive compensation to our members of senior management and managers by

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rewarding  the  achievement  of  corporate  goals  and  specifically  measured  personal  goals  that  are  consistent  with  and  support  the  achievement  of  the
corporate goals. The key terms of the Management Incentive Compensation Plan are summarized below.

Administration and Eligibility

Our  Chief  Executive  Officer  is  responsible  for  the  administration  of  the  Management  Incentive  Compensation  Plan;  however,  the  compensation
committee  of  the  board  of  directors  is  responsible  for  approving  any  incentive  awards  to  our  Chief  Executive  Officer  and  other  members  of  our  senior
management.

In order to be eligible to receive an incentive award under the Management Incentive Compensation Plan, an individual must have been employed
with us for at least three consecutive months during a plan year, which runs from January 1 to December 31, and must achieve a rating of at least 75% of
his or her personal goal.

Form and Determination of Incentive Awards

Incentive award payments may be made in cash, or, at the discretion of the compensation committee and subject to the approval of our board of

directors, through the issuance of equity.

An  individual’s  potential  incentive  award  is  calculated  by  multiplying  his  or  her  base  salary  as  of  the  end  of  the  plan  year  by  the  participant’s
“target award multiplier,” which is a percentage ranging from 10% to 50%. The resulting amount is then divided between a corporate component and an
individual component based on the weighting assigned for the individual’s management level. After the end of the plan year, the actual achievement of the
corporate and individual goals is determined, each expressed as a percentage of complete achievement, resulting in the calculation of the individual’s total
incentive award.

Annual performance reviews for participants in the Management Incentive Compensation Plan are completed following the end of the applicable

plan year, with payment of incentive awards made as soon as practicable thereafter.

Termination of Employment

If a participant in the Management Incentive Compensation Plan gives or receives notice of termination or his or her employment is terminated
prior to the payment of an incentive award under the Management Incentive Compensation Plan, our board of directors has discretion as to whether or not
to pay an incentive award and whether to pay the full amount of the incentive award or a portion thereof.

Amendment

Our  board  of  directors  may  abolish  or  alter  the  Management  Incentive  Compensation  Plan  at  any  time  before,  during  or  after  a  plan  year  is

completed.

Senior Management Employment Arrangements

We have entered into arrangements with members of our senior management to grant restricted shares that are subject to vesting and a repurchase

right in favor of us in the event the individual terminates his or her employment prior to the vesting date. 

In order to align the interests of our executive management with our shareholders, members of our executive management are eligible to receive
share-based awards pursuant to our equity incentive plans. The amount of the awards will generally be subject to the discretion of our board of directors
and our compensation committee.

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Outstanding Equity Awards, Grants and Option Exercise

The following table summarizes the options that we granted to members of our board of directors and senior management pursuant to the terms of

the 2017 Plan or 2018 Plan during the year ended December 31, 2020.

Name

Senior Management
   Nushmia Khokhar, M.D.*
Non-Executive Directors
   Joseph Anderson, Ph.D.
   Jay Backstrom, M.D, MPH
   Linda Bain
   John Berriman
   Cynthia Butitta
   Kapil Dhingra, M.D.
   Martin Murphy, Ph.D.

Ordinary Share Underlying

Option

Exercise Price

Grant

Date

Expiration

Date

20,000 

12,500 
25,000 
12,500 
12,500 
12,500 
12,500 
12,500 

13.97 

8/3/2020

8/3/2030

13.25 
13.97 
13.25 
13.25 
13.25 
13.25 
13.25 

6/18/2020
8/3/2020
6/18/2020
6/18/2020
6/18/2020
6/18/2020
6/18/2020

6/18/2030
8/3/2030
6/18/2030
6/18/2030
6/18/2030
6/18/2030
6/18/2030

* Dr. Khokhar, our Senior Vice President, Clinical Development since August 2020, has resigned from the Company effective mid-March 2021.

As of December 31, 2020, members of our board of directors and senior management held options to purchase an aggregate of 3,247,041 ordinary
shares and restricted share awards covering an aggregate of 1,610,946 ordinary shares. No options were exercised by any members of our board of directors
and senior management during the year ended December 31, 2020.

Equity Incentive Plans

We have granted equity securities under a share option plan and an equity incentive plan, which are summarized in this section.

2017 Share Option Plan

        In  2017,  our  board  of  directors  and  shareholders  approved  the  2017  Plan  to  provide  equity  incentives  to  certain  eligible  employees  and  directors,
consultants and advisors. The 2017 Plan provided for the grant of potentially tax-favored Enterprise Management Incentives, or EMI, options to our U.K.
employees and for the grant of options to our U.S. employees. The 2017 Plan terminated in connection with our IPO; accordingly, as of September 30,
2018, there were no shares available for future grants under the 2017 Plan. Options previously granted pursuant to the 2017 Plan and that are currently
outstanding remain subject to the terms of the 2017 Plan.

2018 Equity Incentive Plan

The 2018 Plan was approved by our board of directors and shareholders in June 2018 and became effective as of our IPO. The 2018 Plan allows for
the  grant  of  equity-based  incentive  awards  to  our  employees  and  directors,  including  directors  who  are  also  our  employees.  Except  where  the  context
indicates otherwise, references hereunder to our ordinary shares shall be deemed to include a number of ADSs equal to the number of ordinary shares. The
material terms of the 2018 Plan are summarized below:

Eligibility and Administration

Our  employees  and  directors,  and  employees  and  consultants  of  our  subsidiaries,  referred  to  as  service  providers  are  eligible  to  receive  awards
under the 2018 Plan. The 2018 Plan is administered by our board of directors, which may delegate its duties and responsibilities to one or more committees
of  our  directors  and/or  officers  (referred  to  as  the  plan  administrator  below),  subject  to  certain  limitations  imposed  under  the  2018  Plan,  and  other
applicable  laws  and  stock  exchange  rules.  Our  board  of  directors  has  delegated  concurrent  authority  to  administer  the  2018  Plan  to  the  compensation
committee. The plan administrator has the authority to take all actions and make all determinations under the 2018 Plan, to interpret the 2018 Plan and
award agreements and to adopt, amend and repeal rules for the administration of the 2018 Plan as it deems advisable. The plan administrator also has the
authority  to  determine  which  eligible  service  providers  receive  awards,  grant  awards,  set  the  terms  and  conditions  of  all  awards  under  the  2018  Plan,
including  any  vesting  and  vesting  acceleration  provisions,  and  designate  whether  such  awards  will  cover  our  ordinary  shares  or  ADSs,  subject  to  the
conditions and limitations in the 2018 Plan.

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Shares Available for Awards

    The maximum number of ordinary shares that may be issued under our 2018 Plan was initially 3,281,622 shares, which consists of 3,025,548 ordinary
shares under the 2018 Plan at the time of its adoption and 256,074 ordinary shares that remained available for future grants under the 2017 Plan at the time
of its termination. Additionally, the number of ordinary shares reserved for issuance under the 2018 Plan will automatically increase on October 1st of each
year, for a period of not more than ten years, commencing on October 1, 2018 and ending on (and including) October 1, 2027, by an amount equal to the
lesser of (i) 4% of the total number of ordinary shares outstanding on September 30 of the same calendar year or (ii) such fewer number of ordinary shares
as the board of directors may designate prior to the applicable October 1st date. As of December 31, 2020, an updated maximum of 8,778,719 ordinary
shares may be issued under the 2018 Plan, of which 4,071,011 were available for future grant as of that date. No more than 14,000,000 shares may be
issued under the 2018 Plan upon the exercise of incentive share options.

If an award under the 2018 Plan, or any prior equity incentive plan, expires, lapses or is terminated, exchanged for cash, surrendered, repurchased,
canceled without having been fully exercised or forfeited, any unused shares subject to the award will, as applicable, become or again be available for new
grants under the 2018 Plan. Awards granted under the 2018 Plan in substitution for any options or other equity or equity-based awards granted by an entity
before the entity’s merger or consolidation with us or our acquisition of the entity’s property or stock will not reduce the shares available for grant under the
2018 Plan, but will count against the maximum number of shares that may be issued upon the exercise of incentive options.

Awards

The 2018 Plan provides for the grant of options, share appreciation rights, or SARs, restricted shares, dividend equivalents, restricted share units, or
RSUs, and other share-based awards. All awards under the 2018 Plan will be set forth in award agreements, which will detail the terms and conditions of
awards, including any applicable vesting and payment terms, change of control provisions and post-termination exercise limitations. A brief description of
each award type follows.

Options and SARs. Options provide for the purchase of our ordinary shares in the future at an exercise price set on the grant date. SARs entitle their
holder, upon exercise, to receive from us an amount equal to the appreciation of the shares subject to the award between the grant date and the exercise
date. The plan administrator will determine the number of shares covered by each option and SAR, the exercise price of each option and SAR and the
conditions and limitations applicable to the exercise of each option and SAR.

Restricted Shares and RSUs.  Restricted  shares  are  an  award  of  nontransferable  ordinary  shares  that  remain  forfeitable  unless  and  until  specified
conditions are met and which may be subject to a purchase price. RSUs are contractual promises to deliver our ordinary shares in the future, which may
also remain forfeitable unless and until specified conditions are met and may be accompanied by the right to receive the equivalent value of dividends paid
on our ordinary shares prior to the delivery of the underlying shares. The plan administrator may provide that the delivery of the shares underlying RSUs
will  be  deferred  on  a  mandatory  basis  or  at  the  election  of  the  participant.  The  terms  and  conditions  applicable  to  restricted  shares  and  RSUs  will  be
determined by the plan administrator, subject to the conditions and limitations contained in the 2018 Plan.

Other Share-Based Awards. Other share-based awards are awards of fully vested ordinary shares and other awards valued wholly or partially by
referring  to,  or  otherwise  based  on,  our  ordinary  shares  or  other  property.  Other  share-based  awards  may  be  granted  to  participants  and  may  also  be
available as a payment form in the settlement of other awards, as standalone payments and as payment in lieu of compensation to which a participant is
otherwise  entitled.  The  plan  administrator  will  determine  the  terms  and  conditions  of  other  share-based  awards,  which  may  include  any  purchase  price,
performance goal, transfer restrictions and vesting conditions.

Performance Criteria

The plan administrator may select performance criteria for an award to establish performance goals for a performance period.

Certain Transactions

In connection with certain corporate transactions and events affecting our ordinary shares, including a change of control, another similar corporate
transaction or event, another unusual or nonrecurring transaction or event affecting us or our financial statements or a change in any applicable laws or
accounting principles, the plan administrator has broad discretion to take action under the 2018 Plan to prevent the dilution or enlargement of intended
benefits, facilitate the transaction or event or give effect to the change in applicable laws or accounting principles. This includes canceling awards for cash
or property, accelerating the vesting of awards, providing for the assumption or substitution of awards by a successor entity, adjusting the number and type
of shares subject to outstanding awards and/or with respect to which awards may be granted under the 2018 Plan and replacing or terminating awards

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under  the  2018  Plan.  In  addition,  in  the  event  of  certain  non-reciprocal  transactions  with  our  shareholders,  the  plan  administrator  will  make  equitable
adjustments to the 2018 Plan and outstanding awards as it deems appropriate to reflect the transaction.

Plan Amendment and Termination

Our board of directors may amend or terminate the 2018 Plan at any time; however, no amendment, other than an amendment that increases the
number of shares available under the 2018 Plan, may materially and adversely affect an award outstanding under the 2018 Plan without the consent of the
affected participant and shareholder approval will be obtained for any amendment to the extent necessary to comply with applicable laws. Further, the plan
administrator  cannot,  without  the  approval  of  our  shareholders,  amend  any  outstanding  option  or  SAR  to  reduce  its  price  per  share  or  cancel  any
outstanding option or SAR in exchange for cash or another award under the 2018 Plan with an exercise price per share that is less than the exercise price
per share of the original option or SAR. The 2018 Plan will remain in effect until the tenth anniversary of its effective date unless earlier terminated by our
board of directors. No awards may be granted under the 2018 Plan after its termination.

Transferability and Participant Payments

Except  as  the  plan  administrator  may  determine  or  provide  in  an  award  agreement,  awards  under  the  2018  Plan  are  generally  non-
transferrable, except by will or the laws of descent and distribution, or, subject to the plan administrator’s consent, pursuant to a domestic relations order,
and are generally exercisable only by the participant. With regard to tax withholding obligations arising in connection with awards under the 2018 Plan, and
exercise price obligations arising in connection with the exercise of options under the 2018 Plan, the plan administrator may, in its discretion, accept cash,
wire transfer or cheque, our ordinary shares that meet specified conditions, a promissory note, a “market sell order,” such other consideration as the plan
administrator deems suitable or any combination of the foregoing.

Non-U.S. Participants

The  plan  administrator  may  modify  awards  granted  to  participants  who  are  non-U.S.  nationals  or  employed  outside  the  United  States  or

establish sub-plans or procedures to address differences in laws, rules, regulations or customs of such foreign jurisdictions.

U.S. Taxpayers

Awards may be granted under the 2018 Plan to U.S. taxpayers.

2018 Non-Employee Sub Plan

    The 2018 Non-Employee Sub Plan will govern equity awards granted to our non-executive directors and our service providers. The 2018 Non-Employee
Sub Plan was adopted under the 2018 Plan and provides for equity- and cash-based awards to be made on identical terms to awards made under our 2018
Plan.  If  all  or  any  part  of  an  award  granted  under  the  2018  Non-Employee  Sub  Plan  expires,  lapses  or  is  terminated,  exchanged  for  cash,  surrendered,
repurchased, canceled without having been fully exercised or forfeited, any unused shares covered by the award will become or again be available for new
grants under the 2018 Non-Employee Sub Plan.

C. Board practices.

Composition of Our Board of Directors

Our board of directors presently has eight members. As a foreign private issuer, under the listing requirements and rules of Nasdaq, we are not required to
have  independent  directors  on  our  board  of  directors,  except  that  our  audit  committee  is  required  to  consist  fully  of  independent  directors,  subject  to
certain phase-in schedules. However, our board of directors has determined that Drs. Anderson, Backstrom, Dhingra and Murphy, Mses. Butitta and Bain
and Mr. Berriman, representing seven of our eight directors, do not have a relationship that would interfere with the exercise of independent judgment in
carrying out the responsibilities of director and that each of these directors is “independent” as that term is defined under Nasdaq rules.

In accordance with our Articles of Association, our board of directors are divided into three classes with staggered three-year terms. At each annual
general  meeting  of  shareholders,  the  directors  whose  terms  expire  will  retire  and  are  eligible  for  re-appointment  by  ordinary  resolution  at  such  annual
general meeting. At each annual general meeting, the successors to directors whose terms then expire or the directors who have been re-appointed will be
elected to serve from the time of election and qualification until the third annual meeting following election. Our directors are divided among the three
classes as follows:

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• Class I, which consists of Joseph Anderson, Jay Backstrom and Martin Murphy, whose terms will expire at our 2022 annual general meeting;

• Class II, which consists of John Berriman and Kapil Dhingra, whose terms will expire at our 2023 annual general meeting; and

• Class III, which consists of Christian Itin, Cynthia Butitta and Linda Bain, whose terms will expire at our 2021 annual general meeting.

Each director shall serve until his or her successor is duly elected and qualified or until his or her earlier death, resignation or removal.

Committees of Our Board of Directors

Our  board  of  directors  has  four  standing  committees:  an  audit  committee,  a  compensation  committee,  a  nominating  and  corporate  governance
committee,  and  a  research  and  development  committee.  The  board  has  adopted  a  written  charter  for  each  of  the  committees  below  that  is  available  to
shareholders on our website at http://www.autolus.com/investor-relations/corporate-governance/documents-charters.

Audit Committee

The audit committee is composed of Ms. Bain (chair), Dr. Anderson and Ms. Butitta, and assists the board of directors in overseeing our accounting
and financial reporting processes. The audit committee consists exclusively of members of our board who are financially literate, and our board of directors
has determined that Ms. Bain is an “audit committee financial expert” as defined by applicable SEC rules and has the requisite financial sophistication as
defined  under  the  applicable  Nasdaq  rules  and  regulations.  Our  board  of  directors  has  determined  that  each  member  of  the  audit  committee  is  an
independent director under Nasdaq listing rules and under Rule 10A-3 under the Exchange Act. Our audit committee meets at least four times per year and
oversees  and  reviews  our  internal  controls,  accounting  policies  and  financial  reporting,  and  provides  a  forum  through  which  our  independent  registered
public accounting firm reports. Our audit committee meets regularly with our independent registered public accounting firm without management present.

The primary functions of the audit committee include:

• recommending the appointment of the independent auditor to shareholders for approval at the general meeting of shareholders;

• the appointment, compensation, retention and oversight of any accounting firm engaged for the purpose of preparing or issuing an audit report

or performing other audit services;

• pre-approving the audit services and non-audit services to be provided by our independent auditor before the auditor is engaged to render such

services;

• evaluating the independent auditor’s qualifications, performance and independence, and presenting its conclusions to the full board of directors

on at least an annual basis;

• reviewing and discussing with management and our independent registered public accounting firm our financial statements and our financial

reporting process; and

• reviewing, approving or ratifying any related party transactions.

Compensation Committee

The  compensation  committee  is  composed  of  Mr.  Berriman  (chairman),  Ms.  Butitta  and  Dr.  Murphy.  Under  SEC  and  Nasdaq  rules,  there  are
heightened independence standards for members of the compensation committee, including a prohibition against the receipt of any compensation from us
other  than  standard  board  member  fees.  Although  foreign  private  issuers  are  not  required  to  meet  this  heightened  standard,  all  of  our  compensation
committee members meet this heightened standard.

The primary functions of the compensation committee include:

• identifying, reviewing, overseeing and proposing policies relevant to the compensation and benefits of our directors and senior management;

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• evaluating the performance of senior management in light of such policies and reporting to the board; and

• overseeing and administering our share option plan, equity incentive plan and other benefit plans in operation from time to time.

Nominating and Corporate Governance Committee

The nominating and corporate governance committee is composed of Dr. Dhingra (chairman), Dr. Anderson and Ms. Bain.

The primary functions of the nominating and corporate governance committee include:

• drawing up selection criteria and appointment procedures for directors;

• recommending nominees for appointment to our board of directors and its corresponding committees; and

• assessing the functioning of individual members of our board of directors and management and reporting the results of such assessment to the

full board of directors.

Research and Development Committee

    The research and development committee is composed of Drs. Dhingra (chairman), Backstrom, Itin and Murphy.

    The primary functions of the research and development committee include:

• overseeing the Company's scientific, technical, research and development strategy, and the implementation thereof;

• advising  our  board  of  directors  and  management  regarding  program  prioritization,  clinical  development  strategy,  regulatory  strategy  and

interactions, intellectual property, product manufacture and supply, and related matters; and

• reviewing and assessing business development opportunities related to research collaborations, licensing or strategic transactions.

D. Employees.

As of December 31, 2020, we had 376 full-time employees, 88 of whom hold Ph.D. or M.D. degrees. Of these 376 employees, 317 are engaged in research
and  development  activities  and  59  are  engaged  in  business  development,  finance,  information  systems,  facilities,  human  resources  or  administrative
support. None of our employees is subject to a collective bargaining agreement. We consider our relationship with our employees to be good.

Function:

Administrative
Research and development

Total
Geography:

United Kingdom
Germany and Switzerland
United States

E. Share ownership.

2020

At December 31,
2019

2018

59
317
376

340
2
34

43
247
290

251
6
33

38
162
200

177
3
20

For information regarding the share ownership of our directors and members senior management, see Item 6.B—"Compensation” and Item 7.A—"Major
Shareholders.”

Item 7. Major Shareholders and Related Party Transactions

A.

Major shareholders.

117

    The following table sets forth information with respect to the beneficial ownership of our ordinary shares as of February 12, 2021 by:

•
•
•

each beneficial owner of 5% or more of our outstanding ordinary shares;
each of our current directors and each member of our senior management; and
all of our directors and senior management as a group.

    Beneficial ownership is determined in accordance with the rules of the SEC. These rules generally attribute beneficial ownership of securities to persons
who possess sole or shared voting power or investment power with respect to those securities and include ordinary shares that can be acquired within 60
days  of  February  12,  2021.  These  ordinary  shares,  however,  are  not  included  in  the  computation  of  the  percentage  ownership  of  any  other
person.    Percentage  ownership  calculations  are  based  on  70,507,309  ordinary  shares  outstanding  (including  ordinary  shares  in  the  form  of  ADSs)  as  of
February 12, 2021.

    Except as otherwise indicated, all of the shares reflected in the table are ordinary shares and all persons listed below have sole voting and investment
power with respect to the shares beneficially owned by them, subject to applicable community property laws. The information is not necessarily indicative
of beneficial ownership for any other purpose.

    Except as otherwise indicated, the addresses of the persons listed in the table is c/o Autolus Therapeutics plc, 58 Wood Lane, White City, London W12
7RZ, United Kingdom.

NAME OF BENEFICIAL OWNER

Number of Ordinary Shares
Beneficially Owned (#)

Percent of Ordinary Shares
Beneficially Owned (%)

5% or Greater Shareholders:

Syncona Portfolio Limited (1)
    PPF Capital Partners Fund B.V. (2)

Senior Management and Directors:
Christian Itin, Ph.D. (3)
Andrew Oakley (4)
Martin Pulé, MBBS (5)
Muhammad Al-Hajj, Ph.D. (6)
Christopher Vann (7)
Matthias Alder (8)
David Brochu (9)
Nushmia Khokhar (10)
Joseph Anderson, Ph.D. (11)
Jay Backstrom, M.D., MPH (12)
Linda Bain (13)
John Berriman (14)
Cynthia Butitta (15)
Kapil Dhingra, M.D. (16)
Martin Murphy, Ph.D. (17)

All directors and senior management as a group (15 persons) (18)

19,527,162
14,251,020

1,450,403
256,657 
808,332
178,481
290,549
297,260
71,664 
94,054
34,374
5,555
55,959 
182,805
80,676
120,011
19,561,536

23,488,316

27.7 %
20.2 %

2.0 %
*
1.1 %
*
*
*
*
*
*
*
*
*
*
*
27.7 %

32.5 %

* Represents beneficial ownership of less than one percent.
(1)The information shown is based, in part, upon disclosures filed on a Schedule 13G/A on February 12, 2020 by Syncona Portfolio Limited. The number reported consists of (i) 12,180,333 ordinary
shares and (ii) 3,775,401 ADSs. In addition, 3,571,428 ADS were purchased from the Company's February 2021 offering. Syncona Portfolio Limited is a wholly owned subsidiary of Syncona Holdings
Limited, which, in turn, is a wholly controlled subsidiary of Syncona Limited, a publicly-listed company. Each of Syncona Holdings Limited and Syncona Limited may be deemed to have voting and
dispositive power over the securities held by Syncona Portfolio Limited. Investment and voting decisions with respect to these securities are made by Syncona Portfolio Limited acting upon the
recommendation of an investment committee of Syncona Investment Management Limited, also a subsidiary of Syncona Holdings Limited. The members of this investment committee consist of Nigel
Keen, Martin Murphy and Chris Hollowood. The address for Syncona Portfolio Limited is Arnold House, St Julian's Avenue, St Peter Port, Guernsey GY1 3RD, Channel Islands.
(2) The information shown is based, in part, upon disclosures filed on a Schedule 13D/A on August 13, 2020 by PPF Capital Partners Fund B.V., PPF Group N.V. and Petr Kellner. The number reported
consists of 14,251,020 ADSs. The principal shareholder of PPF Capital Partners Fund B.V. is PPF Group N.V., which is ultimately beneficially owned by Petr Kellner. The address of the principal office
of each of PPF Group and PPF Capital is Strawinskylaan 933, 1077XX Amsterdam, The Netherlands. The address of the principal office of Petr Kellner is c/o PPF a.s., Evropská 2690/17, P.O. Box 177,
160 41 Prague 6, Czech Republic.
(3) Consists of (i) 1,066,009 ordinary shares issuable upon conversion of restricted ordinary shares and (ii) 384,394 ordinary shares underlying options that are vested and exercisable within 60 days of
February 12, 2021.
(4) Consists of ordinary shares underlying options that are vested and exercisable within 60 days of February 12, 2021.
(5) Consists of (i) 538,677 ordinary shares, (ii) 160,064 ordinary shares issuable upon conversion of restricted ordinary shares, and (iii) 109,591 ordinary shares underlying options that are vested and
exercisable within 60 days of February 12, 2021.

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(6) Consists of ordinary shares underlying options that are vested and exercisable within 60 days of February 12, 2021.
(7) Consists of (i) 112,211 ordinary shares issuable upon conversion of restricted ordinary shares and (ii) 178,338 ordinary shares underlying options that are vested and exercisable within 60 days of
February 12, 2021.
(8) Consists of (i) 125,588 ordinary shares issuable upon conversion of restricted ordinary shares and (ii) 171,672 ordinary shares underlying options that are vested and exercisable within 60 days of
February 12, 2021.
(9) Consists of ordinary shares underlying options that are vested and exercisable within 60 days of February 12, 2021.
(10) Consists of ordinary shares underlying options that are vested and exercisable within 60 days of February 12, 2021.
(11) Consists of ordinary shares underlying options that are vested and exercisable within 60 days of February 12, 2021.
(12) Consists of ordinary shares underlying options that are vested and exercisable within 60 days of February 12, 2021.
(13) Consists of ordinary shares underlying options that are vested and exercisable within 60 days of February 12, 2021.
(14) Consists of (i) 62,794 ordinary shares and (ii) 73,537 ordinary shares issuable upon conversion of restricted ordinary shares, and (iii) 46,474 ordinary shares underlying options that are vested and exercisable
within 60 days of February 12, 2021.
(15) Consists of (i) 10,000 ADSs and (ii) 70,676 ordinary shares underlying options that are vested and exercisable within 60 days of February 12, 2021
(16) Consists of (i) 73,537 ordinary shares issuable upon conversion of restricted ordinary shares and (ii) 46,474 ordinary shares underlying options that are vested and exercisable within 60 days of February 12, 2021.
(17) Consists of (i) the shares set forth in footnote (1) above. Dr. Murphy is the chief executive officer of Syncona Investment Management Limited (both Syncona Investment Management Limited and Syncona
Portfolio Limited are subsidiaries of Syncona Limited) and (ii) and 34,374 ordinary shares underlying options that are vested and exercisable within 60 days of February 12, 2021.
(18) Consists of (i) 20,138,633 ordinary shares, (ii) 1,610,946 ordinary shares issuable upon conversion of restricted ordinary shares and (ii) 1,738,737 ordinary shares underlying options that are vested and exercisable
within 60 days of February 12, 2021.

    The significant changes in the beneficial ownership percentage held by our major shareholders during the past three years result from our June 2018
initial public offering of ADSs, and our April 2019, January 2020, and February 2021 follow-on offerings of ADSs and the dilution resulting from these
offerings.

    As of February 12, 2021, assuming that all of our ordinary shares represented by ADSs are held by residents of the United States other than ADSs held
by the entities set forth in the table above and certain other holders that we know to be non-residents of the United States, we estimate that approximately
27.5% of our outstanding ordinary shares (including ordinary shares underlying ADSs) were held in the United States by 82 holders of record. The actual
number of holders is greater than these numbers of record holders, and includes beneficial owners whose ordinary shares are held in street name by brokers
and other nominees. This number of holders of record also does not include holders whose shares may be held in trust by other entities.

B. Related party transactions.

Policies and Procedures for Related Person Transactions

We  have  adopted  a  related  person  transaction  policy  that  sets  forth  our  procedures  for  the  identification,  review,  consideration  and  approval  or
ratification of related person transactions. For purposes of our policy only, a related person transaction is a transaction, arrangement or relationship, or any
series of similar transactions, arrangements or relationships, in which we or any of our subsidiaries and any related person are, were or will be participants
in which the amount involved exceeds $120,000 or which is unusual in its nature or conditions. Transactions involving compensation for services provided
to us as an employee or director are not covered by this policy. A related person is any executive officer, director or beneficial owner of more than 5% of
any class of our voting securities, including any of their immediate family members and any entity owned or controlled by such persons.

    For so long as we qualify as a foreign private issuer, a related person will be any:

•
•
•

•

•

enterprise that directly or indirectly controls or is controlled by or is under common control with us;
enterprise over which we have a significant influence or which has significant influence over us;
individual owning, directly or indirectly, an interest in our voting power that gives them significant influence over us, and close members
of any such individual’s family;
persons having authority or responsibility for planning, directing or controlling our activities, including directors and senior management
and close members of such individuals’ families; or
enterprise in which a substantial interest in our voting power is owned, directly or indirectly, by any person described above or over
which such a person is able to exercise significant influence, including enterprises owned by our directors or major shareholders and
enterprises that have a member of key management in common with us.

    If we cease to be a foreign private issuer, then, under our policy, a related person will be any:

•

•
•
•

person who is, or at any time since the beginning of our last fiscal year was, a director or member of senior management of us or a
nominee to become a director of us;
security holder known by us to be the beneficial owner of more than 5% of any class of our voting securities;
immediate family member of any of the foregoing; and
firm, corporation or other entity in which any of the foregoing persons is an executive, partner or principal or similar control position or
in which such person has a 5% or greater beneficial ownership interest.

119

    
    Under the policy, if a transaction has been identified as a related person transaction, including any transaction that was not a related person transaction
when originally consummated or any transaction that was not initially identified as a related person transaction prior to consummation, our management
must present information regarding the related person transaction to our audit committee, or, if audit committee approval would be inappropriate, to another
independent body of our board of directors for review, consideration and approval or ratification. The presentation must include a description of, among
other things, the material facts, the interests, direct and indirect, of the related persons, the benefits to us of the transaction and whether the transaction is on
terms  that  are  comparable  to  the  terms  available  to  or  from,  as  the  case  may  be,  an  unrelated  third  party  or  to  or  from  employees  generally.  Under  the
policy,  we  will  collect  information  that  we  deem  reasonably  necessary  from  each  director,  member  of  senior  management  and,  to  the  extent  feasible,
significant shareholder to enable us to identify any existing or potential related person transactions and to effectuate the terms of the policy. In addition,
under  our  Code  of  Business  Conduct  and  Ethics,  our  employees,  members  of  senior  management  and  directors  have  an  affirmative  responsibility  to
disclose any transaction or relationship that reasonably could be expected to give rise to a conflict of interest.

Transactions with Our Principal Shareholders, Directors and Members of our Senior Management

The following is a description of related party transactions we have entered into since January 1, 2018 with our directors, members of our senior
management and holders of more than 5% of our outstanding voting securities and their affiliates, whom we refer to as our related persons, in which the
amount  involved  exceeds  $120,000  and  that  are  material  to  us,  other  than  the  compensation  arrangements  we  describe  in  Item  6.B.  “Management  -
Compensation of Senior Management and Directors.”

Participation in April 2019 Public Offering

In  our  April  2019  public  offering,  Syncona  Portfolio  Limited  and  Arix  Bioscience  Holdings  Limited  purchased  1,000,000  ADSs  and  208,333

ADSs, respectively. This purchase was made through the underwriters at the public offering price.

Participation in January 2020 Public Offering

In our January 2020 public offering, Syncona Portfolio Limited purchased 1,363,636 ADSs. This purchase was made through the underwriters at

the public offering price.

Participation in February 2021 Public Offering

    In our February 2021 public offering, Syncona Portfolio Limited purchased 3,571,428 ADSs. This purchase was made through the underwriters at the
public offering price.

Entities affiliated with Syncona

In September 2020, we entered into a license agreement with an investee company of Syncona Portfolio Limited, a holder of more than 5% of our
share capital. This agreement generated $242,000 of license revenue which is recognized in the Consolidated Statement of Operations for the year ended
December 31, 2020.

Agreements with Our Senior Management and Directors

       We  have  entered  into  service  agreements  with  the  members  of  our  senior  management  and  non-executive  directors.  See  Item  6.B.  “Management—
Compensation of Senior Management and Directors.” These agreements contain customary provisions and representations, including confidentiality, non-
competition, non-solicitation and inventions assignment undertakings by the members of our senior management. However, the enforceability of the non-
competition provisions may be limited under applicable law.

Indemnification Agreements

        We  entered  into  a  deed  of  indemnity  with  each  of  our  directors  and  members  of  our  senior  management.  These  agreements  and  our  Articles  of
Association require us to indemnify our directors and senior management to the fullest extent permitted by law.

Registration Rights Agreement

    We have entered into a registration rights agreement with certain of our existing shareholders pursuant to which we have granted them customary
registration rights for the resale of the ordinary shares held by certain of our existing shareholders.

C. Interests of experts and counsel.

120

Not applicable.

Item 8. Financial Information.

A.

Consolidated Statements and Other Financial Information.

    Our consolidated financial statements are appended at the end of this Annual Report, starting on page F-1.

Dividend Policy

    We have never declared or paid a dividend, and we do not anticipate declaring or paying dividends in the foreseeable future. We intend to retain all
available funds and any future earnings to fund the development and expansion of our business.

    Under English law, among other things, we may only pay dividends if we have sufficient distributable reserves (on a non-consolidated basis), which are
our accumulated realized profits that have not been previously distributed or capitalized less our accumulated realized losses, so far as such losses have not
been previously written off in a reduction or reorganization of capital.

Legal Proceedings

    We are not currently a party to any material legal proceedings. From time to time, we may be a party to litigation or subject to claims incident to the
ordinary course of business. Although the results of litigation and claims cannot be predicted with certainty, we currently believe that the final outcome of
these ordinary course matters will not have a significant effect on our financial position or profitability. Regardless of the outcome, litigation can have an
adverse impact on us because of defense and settlement costs, diversion of management resources and other factors.

B. Significant Changes.

Not applicable.

Item 9. The Offer and Listing.

A. Offer and listing details.

    Our ADS began trading on the Nasdaq Global Select Market under the symbol “AUTL” since June 22, 2018. Prior to that date, there was no public
trading market for ADSs or our ordinary shares.  

B. Plan of distribution.

    Not applicable.

C. Markets.

    Our ADSs have been listed on the Nasdaq Global Select Market under the symbol “AUTL” since June 22, 2018. Our ordinary shares are not listed.

D. Selling shareholders.

    Not applicable.

E. Dilution.

    Not applicable.

F. Expenses of the issue.

    Not applicable.

Item 10. Additional Information.

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A. Share capital.

Not applicable.

B. Memorandum and articles of association.

General

We are a public limited company, originally incorporated pursuant to the laws of England and Wales in February 2018 as a private company with
limited  liability  called  Autolus  Therapeutics  Limited.  Autolus  Limited  was  originally  incorporated  under  the  laws  of  England  and  Wales  in  July  2014.
Pursuant to the terms of our corporate reorganization, the shareholders of Autolus Limited exchanged each of the shares held by them in Autolus Limited
for the same number and class of newly issued shares of Autolus Therapeutics Limited and, as a result, Autolus Limited became a wholly owned subsidiary
of Autolus Therapeutics Limited. On June 18, 2018, Autolus Therapeutics Limited re-registered as a public limited company and was renamed Autolus
Therapeutics plc. On June 22, 2018, our outstanding preferred and ordinary shares were converted into a single class of ordinary shares and various classes
of deferred shares, and we completed our initial public offering of ADSs on the Nasdaq Global Select Market.

We are registered with the Registrar of Companies in England and Wales under number 11185179, and our registered office is at Forest House, 58

Wood Lane, White City, London W12 7RZ, United Kingdom.

Certain  resolutions  were  passed  by  our  shareholders  in  connection  with  our  initial  public  offering,  including  a  special  resolution  approving  the
adoption of new articles of association that became effective upon the admission of our ADSs to trading on Nasdaq. Our articles of association authorize
our directors, for the purposes of section 551 of the U.K. Companies Act 2006, or the Companies Act, to issue shares in the company and grant rights to
subscribe for or convert any securities into shares in the company up to a maximum aggregate nominal amount of $8,400 for a period of five years. See
“Key Provisions of Our Articles of Association” below for further information.

Issued Share Capital

Effective from June 26, 2018, the board of directors has the authority to allot new ordinary shares or to grant rights to subscribe for or to convert any
security into ordinary shares in the Company up to a maximum aggregate nominal amount of $8,400. This authority runs for five years and will expire on
June  26,  2023.  Effective  from  June  26,  2018,  the  board  also  has  the  authority  to  allot  ordinary  shares  for  cash  or  to  grant  rights  to  subscribe  for  or  to
convert any security into ordinary shares in the Company without first offering them to existing shareholders in proportion to their existing holdings up to
an aggregate maximum nominal amount of $8,400. This authority runs for five years and will expire on June 26, 2023.

As of February 12. 2021, our issued capital share consisted of 70,507,309 ordinary shares, with a nominal value of $0.000042 per share, (ii) 34,425
deferred shares, with a nominal value of £0.00001 per share, (iii) 88,893,548 B deferred shares, with a nominal value of £0.00099 per share and (iv) 1 C
deferred share, with a nominal value of £0.000008. Each issued share has been fully paid.

Ordinary Shares

Our ordinary shares have the rights and restrictions described in “Key Provisions of Our Articles of Association” below. The following summarizes

the rights of holders of our ordinary shares:

•

•

each holder of our ordinary shares is entitled to one vote per ordinary share on all matters to be voted on by shareholders generally;

the holders of the ordinary shares shall be entitled to receive notice of, attend, speak and vote at our general meetings; and

holders  of  our  ordinary  shares  are  entitled  to  receive  such  dividends  as  are  recommended  by  our  directors  and  declared  by  our

•
shareholders.

We are required by the Companies Act to keep a register of our shareholders. Under English law, the ordinary shares are deemed to be issued when
the  name  of  the  shareholder  is  entered  in  our  register  of  members.  The  register  of  members  therefore  is  prima  facie  evidence  of  the  identity  of  our
shareholders, and the shares that they hold. The register of members generally provides

Registered Shares

122

limited,  or  no,  information  regarding  the  ultimate  beneficial  owners  of  our  ordinary  shares.  Our  register  of  members  is  maintained  by  our  registrar,
Computershare Investor Services plc.

Holders of our ADSs will not be treated as our shareholders and their names will therefore not be entered in our share register. The Depositary, the
custodian or their nominees will be the holder of the ordinary shares underlying our ADSs. Holders of our ADSs have a right to receive the ordinary shares
underlying their ADSs. For discussion on our ADSs and ADS holder rights, see “Description of American Depositary Shares” in this Annual Report.

Under  the  Companies  Act,  we  must  enter  an  allotment  of  shares  in  our  register  of  members  as  soon  as  practicable  and  in  any  event  within  two
months of the allotment. We also are required by the Companies Act to register a transfer of shares (or give the transferee notice of and reasons for refusal
as the transferee may reasonably request) as soon as practicable and in any event within two months of receiving notice of the transfer.

We, any of our shareholders or any other affected person may apply to the court for rectification of the register of members if:

•

the name of any person, without sufficient cause, is wrongly entered in or omitted from our register of members; or

there is a default or unnecessary delay in entering on the register the fact of any person having ceased to be a member or on which we

•
have a lien, provided that such delay does not prevent dealings in the shares taking place on an open and proper basis.

Registration Rights

We and the holders of certain of our ordinary shares are parties to a registration rights agreement that provides the following registration rights:

    Demand Registration on Form F-1 - each holder shall be entitled to demand registrations on Form F-1, provided that these demand registration
rights may only be exercised by holders who hold, in the aggregate, not less than 25% of the aggregate number of shares then outstanding and
held  by  all  holders  who  are  party  to  the  agreement,  and  provided  further  that  the  we  shall  not  be  required  to  effect  a  demand  registration
statement after we have effected two demand registration statements, and such registration statements have been declared or ordered effective.

    Demand Registration on Form F-3 - each holder shall be entitled to unlimited demand registrations on Form F-3, if we are eligible to register
shares on Form F-3, provided that these demand registration rights may only be exercised by holders who hold, in the aggregate, not less than
10%  of  the  aggregate  number  of  shares  then  outstanding  and  held  by  all  holders  who  are  party  to  the  agreement.  These  demand  registration
rights may not be exercised more than twice in any twelve-month period.

        Piggyback  Registration  -  each  holder  shall  be  entitled  to  piggyback  registration  rights,  subject,  in  the  case  of  an  underwritten  offering,  to
customary reductions by the underwriter, provided that the aggregate number of securities of the holders included in the registration may not be
reduced to less than 30% of the total number of securities registered.

        Expenses  -  We  will  pay  all  registration  expenses  relating  to  the  exercise  of  the  registration  rights  above,  including  the  reasonable  fees  and

expenses of legal counsel to the participating holders up to a maximum of $50,000 in the aggregate per registration.

Preemptive Rights

English law generally provides shareholders with statutory preemptive rights when new shares are issued for cash; however, it is possible for the
articles  of  association,  or  shareholders  by  way  of  a  special  resolution  at  a  general  meeting,  to  disapply  preemptive  rights.  Such  a  disapplication  of
preemptive rights may be for a maximum period of up to five years from the date of adoption of the articles of association, if the disapplication is contained
in the articles of association, or from the date of the shareholder special resolution, if the disapplication is by shareholder special resolution. In either case,
this disapplication would need to be renewed by our shareholders upon its expiration (i.e., at least every five years). Our articles of association disapply
preemptive  rights  for  a  period  of  five  years  from  the  date  of  adoption,  which  was  June  26,  2018.  This  disapplication  will  need  to  be  renewed  upon
expiration (i.e., at least every five years) to remain effective, but may be sought more frequently for additional five-year terms (or any shorter period). Such
preemption rights have been disapplied by our articles of association that became effective upon the completion of our initial public offering in June 2018
and which remain in force at the date of this Annual Report.

123

 
 
Purchase of Own Shares

English law permits a public limited company to purchase its own shares out of the distributable profits of the company or the proceeds of a fresh
issue of shares made for the purpose of financing the purchase, subject to complying with procedural requirements under the Companies Act and provided
that its articles of association do not prohibit it from doing so. Our articles of association, a summary of which is provided below, do not prohibit us from
purchasing our own shares. A public limited company must not purchase its own shares if, as a result of the purchase, there would no longer be any issued
shares of the company other than redeemable shares or shares held as treasury shares.

Any such purchase will be either a “market purchase” or “off market purchase,” each as defined in the Companies Act. A “market purchase” is a
purchase made on a “recognized investment exchange” (other than an overseas exchange) as defined in the UK Financial Services and Markets Act 2000,
or  FSMA.  An  “off  market  purchase”  is  a  purchase  that  is  not  made  on  a  “recognized  investment  exchange.”  Both  “market  purchases”  and  “off  market
purchases” require prior shareholder approval by way of an ordinary resolution. In the case of an “off market purchase,” a company’s shareholders, other
than the shareholders from whom the company is purchasing shares, must approve the terms of the contract to purchase shares and in the case of a “market
purchase,” the shareholders must approve the maximum number of shares that can be purchased and the maximum and minimum prices to be paid by the
company.

The Nasdaq Global Select Market is an “overseas exchange” for the purposes of the Companies Act and does not fall within the definition of a
“recognized  investment  exchange”  for  the  purposes  of  FSMA  and  any  purchase  made  by  us  would  accordingly  need  to  comply  with  the  procedural
requirements under the Companies Act that regulate “off market purchases.”

A share buy-back by a company of its shares will give rise to U.K. stamp duty reserve tax and stamp duty at the rate of 0.5% of the amount or value
of the consideration payable by the company (rounded up to the next £5.00), and such stamp duty reserve tax or stamp duty will be paid by the company.
The  charge  to  stamp  duty  reserve  tax  will  be  cancelled  or,  if  already  paid,  repaid  (generally  with  interest),  where  a  transfer  instrument  for  stamp  duty
purposes  has  been  duly  stamped  within  six  years  of  the  charge  arising  (either  by  paying  the  stamp  duty  or  by  claiming  an  appropriate  relief)  or  if  the
instrument is otherwise exempt from stamp duty.

Distributions and Dividends

Under the Companies Act, before a company can lawfully make a distribution or dividend, it must ensure that it has sufficient distributable reserves,
as determined on a non-consolidated basis. The basic rule is that a company’s profits available for the purpose of making a distribution are its accumulated,
realized profits, so far as not previously utilized by distribution or capitalization, less its accumulated, realized losses, so far as not previously written off in
a reduction or reorganization of capital duly made. The requirement to have sufficient distributable reserves before a distribution or dividend can be paid
applies to us and to each of our subsidiaries that has been incorporated under English law.

As a public company, it will not be sufficient that we have made a distributable profit for the purpose of making a distribution. An additional capital
maintenance requirement will be imposed on us to ensure that the net worth of the company is at least equal to the amount of its capital. A public company
can only make a distribution:

    if, at the time that the distribution is made, the amount of its net assets (that is, the total excess of assets over liabilities) is not less than the total

of its called up share capital and undistributable reserves; and

    if, and to the extent that, the distribution itself, at the time that it is made, does not reduce the amount of its net assets to less than that total.

Disclosure of Interest in Shares

Pursuant to Part 22 of the Companies Act, we are empowered to give by notice in writing to any person whom we know or have reasonable cause to
believe  to  be  interested  in  our  shares,  or  at  any  time  during  the  three  years  immediately  preceding  the  date  on  which  the  notice  is  issued  has  been  so
interested, within a reasonable time to disclose to us particulars of that person’s interest and, so far as is within his or her knowledge, particulars of any
other interest that subsists or subsisted in those shares.

124

 
Under our articles of association, if a person defaults in supplying us with the required particulars in relation to the shares in question, or default

shares within the prescribed period, our board of directors may by notice direct that:

    the relevant shareholder shall not be entitled in respect of the default shares to be present or vote, either in person or by proxy, at any general
meeting or separate meeting of the holders of a class of shares or upon any poll or to exercise any other right conferred by the membership in
relation to any such meeting;

    where the default shares represent at least 0.25% of their class, (a) any dividend or other money payable in respect of the default shares shall be
retained by us without liability to pay interest, and/or (b) no transfers by the relevant shareholder of any default shares may be registered, unless
the shareholder himself or herself is not in default and the shareholder proves to the satisfaction of the board of directors that no person in default
as regards to supplying such information is interested in any of the default shares; and/or

    any shares held by the relevant shareholder in uncertificated form shall be converted into certificated form.

Key Provisions of Our Articles of Association

The following is a summary of certain key provisions of our articles of association, which were adopted by a special resolution of our shareholders
passed in June 2018. Please note that this is only a summary and is not intended to be exhaustive. For further information please refer to the full version of
our articles of association, which is included as an exhibit to this Annual Report

The  articles  of  association  contain  no  specific  restrictions  on  our  purpose  and  therefore,  by  virtue  of  section  31(1)  of  the  Companies  Act,  our

purpose is unrestricted.

The articles of association contain, among other things, provisions to the following effect:

Share Capital

Our share capital currently consists of ordinary shares, deferred shares, B deferred shares and C deferred shares. We may issue shares with such
rights or restrictions as may be determined by ordinary resolution, including shares which are to be redeemed, or are liable to be redeemed at our option or
the option of the holder of such shares.

Voting

Holders  of  ordinary  shares  have  the  right  to  receive  notice  of,  and  to  vote  at,  our  general  meetings.  Any  resolution  put  to  the  vote  of  a  general
meeting must be decided exclusively on a poll. Each shareholder who is present in person (or, being a corporation, by representative) or by proxy has one
vote in respect of every share held by him.

Variation of Rights

Whenever our share capital is divided into different classes of shares, the special rights attached to any class may be varied or abrogated either (i)
with the consent in writing of the holders of three-quarters in nominal value of the issued shares of that class, (ii) with the authority of a special resolution
passed at a separate meeting of the holders of the shares of that class or (iii) in any other way as expressly provided for in relation to such rights, and may
be so varied and abrogated while the company is a going concern.

Dividends

We may, subject to the provisions of the Companies Act and our articles of association, by ordinary resolution from time to time declare dividends
to  be  paid  to  shareholders  not  exceeding  the  amount  recommended  by  our  board  of  directors.  Subject  to  the  provisions  of  the  Companies  Act,  in  the
discretion of board of directors, on the basis that our profits justify such payments, the board of directors may pay interim dividends on any class of our
shares.

Any  dividend  unclaimed  after  a  period  of  12  years  from  the  date  such  dividend  was  declared  or  became  payable  shall,  if  the  board  of  directors
resolve, be forfeited, cease to remain owing and shall revert to us. No dividend or other moneys payable on or in respect of a share shall bear interest as
against us.

Transfer of Ordinary Shares

Each member may transfer all or any of his shares which are in certificated form by means of an instrument of transfer in writing in any usual form

or in any other form which the board of directors may approve.

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The board of directors may, in its absolute discretion, refuse to register a transfer of certificated shares unless:

(i)    it is for a share which is fully paid up;

(ii)    it is for a share upon which the company has no lien;

(iii)    it is only for one class of share;

(iv)    it is in favor of a single transferee or no more than four joint transferees;

(v)    it is duly stamped or is duly certificated or otherwise shown to the satisfaction of the board of directors to be exempt from stamp duty; and

(vi)        it  is  delivered  for  registration  to  the  registered  office  of  the  company  (or  such  other  place  as  the  board  of  directors  may  determine),
accompanied (except in the case of a transfer by a person to whom the company is not required by law to issue a certificate and to whom a
certificate has not been issued or in the case of a renunciation) by the certificate for the shares to which it relates and such other evidence as
the board of directors may reasonably require to prove the title of the transferor (or person renouncing) and the due execution of the transfer
or renunciation by him or, if the transfer or renunciation is executed by some other person on his behalf, the authority of that person to do
so.

Allotment of Shares and Preemption Rights

Subject  to  the  Companies  Act  and  to  any  rights  attached  to  existing  shares,  any  share  may  be  issued  with  or  have  attached  to  it  such  rights  and
restrictions as the company may by ordinary resolution determine, or if no ordinary resolution has been passed or so far as the resolution does not make
specific provision, as the board of directors may determine (including shares which are to be redeemed, or are liable to be redeemed at the option of the
company or the holder of such shares).

In accordance with section 551 of the Companies Act, the board of directors may be generally and unconditionally authorized to exercise for each
prescribed  period  all  the  powers  of  the  company  to  allot  shares  up  to  an  aggregate  nominal  amount  equal  to  the  amount  stated  in  the  relevant  ordinary
resolution  authorizing  such  allotment.  The  authorities  referred  to  above  are  included  in  our  articles  of  association  that  became  effective  upon  the
completion of our initial public offering in June 2018 and which remain in force at the date of this Annual Report.

The  provisions  of  section  561  of  the  Companies  Act  (which  confer  on  shareholders  rights  of  preemption  in  respect  of  the  allotment  of  equity
securities which are paid up in cash) apply to the company except to the extent disapplied by special resolution of the shareholders of the company, or in
the company’s articles of association. Such preemption rights have been disapplied by our articles of association that became effective upon the completion
of our initial public offering in June 2018 and which remain in force at the date of this Annual Report.  

Alteration of Share Capital

In accordance with the Companies Act, the company may by ordinary resolution consolidate its share capital into shares of larger nominal value
than its existing shares, or sub-divide its shares into shares of a smaller amount than the existing shares, and may in each case determine that the shares
resulting from such sub-division or share consolidation may have a preference or advantage or be subject to a particular restriction.

The company may, in accordance with the Companies Act, reduce or cancel its share capital or any capital redemption reserve or share premium

account in any manner and with and subject to any conditions, authorities and consents required by law.

Board of Directors

Unless otherwise determined by the company by ordinary resolution, the number of directors (other than any alternate directors) shall not be less

than two and not more than 15.

Subject to the articles of association and the Companies Act, the company may by ordinary resolution appoint a person who is willing to act as a
director  and  the  board  of  directors  shall  have  power  at  any  time  to  appoint  any  person  who  is  willing  to  act  as  a  director,  in  both  cases  either  to  fill  a
vacancy or as an addition to the existing board of directors, provided the total number of directors shall not exceed the maximum number of 15.

Our articles of association provide that our board of directors is divided into three classes, each of which will consist, as nearly as possible, of one-
third of the total number of directors constituting our entire board and which will serve staggered three-year terms. At each annual general meeting, the
successors to directors whose terms then expire will be elected to serve from the time of election and qualification until the third annual meeting following
election.

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At every subsequent annual general meeting, any director who either (i) has been appointed by the board of directors since the last annual general
meeting or (ii) was not appointed or reappointed at one of the preceding two annual general meetings, must retire from office and may offer themselves for
reappointment by the shareholders by ordinary resolution.

Subject to the provisions of the articles of association, the board of directors may regulate their proceedings as they deem appropriate. A director

may, and the secretary at the request of a director shall, call a meeting of the directors.

The quorum for a meeting of the board of directors may be determined by the board and until otherwise determined, it is set at two directors.

Questions  and  matters  requiring  resolution  arising  at  a  meeting  shall  be  decided  by  a  majority  of  votes  of  the  participating  directors,  with  each
director having one vote. In the case of an equality of votes, the chairman will have a casting vote or second vote, unless he or she is not entitled to vote on
the resolution in question.

Directors shall be entitled to receive such compensation as the board shall determine for their services to the company as directors, and for any other
service which they undertake for the company provided that the aggregate fees payable to the directors must not exceed $2,500,000 per annum or such
higher amount as may from time to time be decided by ordinary resolution. The directors shall also be entitled to be paid all reasonable expenses properly
incurred by them in connection with their attendance at meetings of shareholders or class meetings, board of director or committee meetings or otherwise in
connection with the exercise of their powers and the discharge of their responsibilities in relation to the company.

     The board of directors may, in accordance with the requirements in the articles of association, authorize any matter proposed to them by any director
which would, if not authorized, involve a director breaching their duty under the Companies Act, to avoid conflicts of interests.

A director seeking authorization in respect of such conflict shall declare to the board of directors the nature and extent of his interest in a conflict as
soon  as  is  reasonably  practicable.  The  director  shall  provide  the  board  with  such  details  of  the  matter  as  are  necessary  for  the  board  to  decide  how  to
address the conflict together with such additional information as may be requested by the board.

Any authorization by the board of directors will be effective only if:

(i)    to the extent permitted by the Companies Act, the matter in question shall have been proposed by any director for consideration in the same

way that any other matter may be proposed to the directors under the provisions of the articles of association;

(ii)        any  requirement  as  to  the  quorum  for  consideration  of  the  relevant  matter  is  met  without  counting  the  conflicted  director  and  any  other

conflicted director; and

(iii)        the  matter  is  agreed  to  without  the  conflicted  director  voting  or  would  be  agreed  to  if  the  conflicted  director’s  and  any  other  interested

director’s vote is not counted.

Subject to the provisions of the Companies Act, every director, secretary or other officer of the company (other than an auditor) is entitled to be

indemnified against all losses and liabilities incurred in connection with his or her duties and powers.

General Meetings

The company must convene and hold annual general meetings once a year in accordance with the Companies Act. Under the Companies Act, an

annual general meeting must be called by notice of at least 21 days.

No  business  shall  be  transacted  at  any  general  meeting  unless  a  quorum  is  present  when  the  meeting  proceeds  to  business,  but  the  absence  of  a
quorum shall not preclude the choice or appointment of a chairman of the meeting which shall not be treated as part of the business of the meeting. Unless
otherwise provided by the articles of association, two shareholders present in person or by proxy and entitled to vote shall be a quorum for all purposes.

Borrowing Powers

Subject to the articles of association and the Companies Act, the board of directors may exercise all of the powers of the company to:

(a)    borrow money;

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(b)    indemnify and guarantee;

(c)    mortgage or charge the assets of the company;

(d)    create and issue debentures and other securities; and

(e)    give security either outright or as collateral security for any debt, liability or obligation of the company or of any third party.

Capitalization of Profits

The directors may, if they are so authorized by an ordinary resolution of the shareholders, decide to capitalize any undivided profits of the company
not required for paying any preferential dividend (whether or not they are available for distribution), or any sum standing to the credit of the company’s
share premium account, capital redemption reserve or any other reserve or fund of the company which is available for distribution. The directors may also,
subject to the aforementioned ordinary resolution, appropriate any sum which they so decide to capitalize to the persons who would have been entitled to it
if it were distributed by way of dividend and in the same proportions.

Uncertificated Shares

Subject to the Companies Act, the board of directors may permit title to shares of any class to be issued or held otherwise than by a certificate and to

be transferred by means of a “relevant system” (e.g., DTC) without a certificate.

The board of directors may take such steps as it sees fit in relation to the evidencing of and transfer of title to uncertificated shares, any records

relating to the holding of uncertificated shares and the conversion of uncertificated shares to certificated shares, or vice versa.

The company may by notice in writing to the holder of an uncertificated share, require that share to be converted into certificated form.

The board of directors may take such other action that the board considers appropriate to achieve the sale, transfer, disposal, forfeiture, re-allotment

or surrender of an uncertified share or otherwise to enforce a lien in respect of it.

Choice of Forum

Our articles of association will provide that the U.S. federal district courts will be the exclusive forum for resolving any complaint asserting a cause

of action arising under the Securities Act.

Other Relevant United Kingdom Laws and Regulations

Mandatory Bid

(i)    The Takeover Code does not currently apply to the company. However if the company were to become subject to the Takeover Code in the

future, the following provisions will apply. Under Rule 9 of the Takeover Code, where:

a.    any person, together with persons acting in concert with him, acquires, whether by a series of transactions over a period of time or not,
an interest in shares which (taken together with shares in which he is already interested, and in which persons acting in concert with
him are interested) carry 30% or more of the voting rights of a company; or

b.    any person who, together with persons acting in concert with him, is interested in shares which in the aggregate carry not less than 30%
of the voting rights of a company but does not hold shares carrying more than 50% of such voting rights and such person, or any
person acting in concert with him, acquires an interest in any other shares which increases the percentage of shares carrying voting
rights in which he is interested; such person shall, except in limited circumstances, be obliged to extend offers, on the basis set out in
Rules 9.3, 9.4 and 9.5 of the Takeover Code, to the holders of any class of equity share capital, whether voting or non-voting, and
also  to  the  holders  of  any  other  class  of  transferable  securities  carrying  voting  rights.  Offers  for  different  classes  of  equity  share
capital must be comparable; the Takeover Panel should be consulted in advance in such cases.

(ii)    An offer under Rule 9 of the Takeover Code must be in cash and at the highest price paid for any interest in the shares by the person required

to make an offer or any person acting in concert with him during the 12 months prior to the announcement of the offer.

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(iii)    Under the Takeover Code, a “concert party” arises where persons acting together pursuant to an agreement or understanding (whether formal
or  informal  and  whether  or  not  in  writing)  actively  cooperate,  through  the  acquisition  by  them  of  an  interest  in  shares  in  a  company,  to
obtain or consolidate control of the company. “Control” means holding, or aggregate holdings, of an interest in shares carrying 30% or more
of the voting rights of the company, irrespective of whether the holding or holdings give de facto control.

Squeeze-out

(i)    Under sections 979 to 982 of the Companies Act, if an offeror were to acquire, or unconditionally contract to acquire, not less than 90% in
value  of  the  ordinary  shares  of  the  company  and  90%  of  the  voting  rights  carried  by  the  ordinary  shares  of  the  company,  it  could  then
compulsorily  acquire  the  remaining  10%.  It  would  do  so  by  sending  a  notice  to  outstanding  shareholders  telling  them  that  it  will
compulsorily acquire their shares, provided that no such notice may be served after the end of: (a) the period of three months beginning with
the  day  after  the  last  day  on  which  the  offer  can  be  accepted;  or  (b)  if  earlier,  and  the  offer  is  not  one  to  which  section  943(1)  of  the
Companies Act applies, the period of six months beginning with the date of the offer.

(ii)    Six weeks following service of the notice, the offeror must send a copy of it to the company together with the consideration for the ordinary
shares to which the notice relates, and an instrument of transfer executed on behalf of the outstanding shareholder(s) by a person appointed
by the offeror.

(iii)    The company will hold the consideration on trust for the outstanding shareholders.

Sell-out

(i)    Sections 983 to 985 of the Companies Act also give minority shareholders in the company a right to be bought out in certain circumstances by
an offeror who has made a takeover offer. If a takeover offer relating to all the ordinary shares of the company is made at any time before
the  end  of  the  period  within  which  the  offer  could  be  accepted  and  the  offeror  held  or  had  agreed  to  acquire  not  less  than  90%  of  the
ordinary shares, any holder of shares to which the offer related who had not accepted the offer could by a written communication to the
offeror  require  it  to  acquire  those  shares.  The  offeror  is  required  to  give  any  shareholder  notice  of  his  right  to  be  bought  out  within  one
month of that right arising. The offeror may impose a time limit on the rights of minority shareholders to be bought out, but that period
cannot end less than three months after the end of the acceptance period, or, if longer a period of three months from the date of the notice.

(ii)    If a shareholder exercises his rights, the offeror is bound to acquire those shares on the terms of the offer or on such other terms as may be

agreed.

Differences in Corporate Law

The  applicable  provisions  of  the  Companies  Act  differ  from  laws  applicable  to  U.S.  corporations  and  their  shareholders.  Set  forth  below  is  a
summary of certain differences between the provisions of the Companies Act applicable to us and the General Corporation Law of the State of Delaware
relating to shareholders’ rights and protections. This summary is not intended to be a complete discussion of the respective rights and it is qualified in its
entirety by reference to Delaware law and English law.

Number of Directors

ENGLAND

the  Companies  Act,  a
Under 
public 
limited  company  must
have at least two directors and the
number of directors may be fixed
by or in the manner provided in a
company’s articles of association.

DELAWARE
Under  Delaware  law,  a  corporation
must have at least one director and the
number of directors shall be fixed by or
in the manner provided in the bylaws.

129

 
Removal of Directors

Vacancies of Board of Directors

ENGLAND

(which 

remove 

the  Companies  Act,
Under 
shareholders  may 
a
director  without  cause  by  an
ordinary 
is
resolution 
passed  by  a  simple  majority  of
those  voting  in  person  or  by
proxy  at  a  general  meeting)
irrespective  of  any  provisions  of
any  service  contract  the  director
has  with  the  company,  provided
28  clear  days’  notice  of 
the
resolution  has  been  given  to  the
company and its shareholders. On
receipt  of  notice  of  an  intended
resolution  to  remove  a  director,
the company must forthwith send
a copy of the notice to the director
other
concerned. 
procedural requirements under the
Companies  Act  must  also  be
followed,  such  as  allowing  the
director  to  make  representations
against  his  or  her  removal  either
at the meeting or in writing.

Certain 

except 

(i)  unless 

DELAWARE
Under  Delaware  law,  any  director  or
the  entire  board  of  directors  may  be
removed, with or without cause, by the
holders of a majority of the shares then
entitled  to  vote  at  an  election  of
directors, 
the
certificate  of  incorporation  provides
otherwise,  in  the  case  of  a  corporation
whose  board  of  directors  is  classified,
stockholders  may  effect  such  removal
only  for  cause,  or  (ii)  in  the  case  of  a
corporation  having  cumulative  voting,
if less than the entire board of directors
is  to  be  removed,  no  director  may  be
removed without cause if the votes cast
against his removal would be sufficient
to elect him if then cumulatively voted
at  an  election  of  the  entire  board  of
directors,  or,  if  there  are  classes  of
directors,  at  an  election  of  the  class  of
directors of which he is a part.

ENGLAND
Under English law, the procedure
by which directors, other than a
company’s initial directors, are
appointed is generally set out in a
company’s articles of association,
provided that where two or more
persons are appointed as directors
of a public limited company by
resolution of the shareholders,
resolutions appointing each
director must be voted on
individually.

DELAWARE

Under Delaware law, vacancies and
newly created directorships may be
filled by a majority of the directors then
in office (even though less than a
quorum) or by a sole remaining director
unless (i) otherwise provided in the
certificate of incorporation or bylaws of
the corporation or (ii) the certificate of
incorporation directs that a particular
class of stock is to elect such director,
in which case a majority of the other
directors elected by such class, or a sole
remaining director elected by such
class, will fill such vacancy.

130

Annual General Meeting

General Meeting

ENGLAND

DELAWARE

Under the Companies Act, a
public limited company must hold
an annual general meeting in each
six-month period following the
company’s annual accounting
reference date.

Under Delaware law, the annual
meeting of stockholders shall be held at
such place, on such date and at such
time as may be designated from time to
time by the board of directors or as
provided in the certificate of
incorporation or by the bylaws.

DELAWARE
Under Delaware law, special meetings
of the stockholders may be called by
the board of directors or by such person
or persons as may be authorized by the
certificate of incorporation or by the
bylaws.

ENGLAND

Under the Companies Act, a
general meeting of the
shareholders of a public
limited company may be called
by the directors.

Shareholders holding at least
5% of the paid-up capital of
the company carrying voting
rights at general meetings
(excluding any paid up capital
held as treasury shares) can
require the directors to call a
general meeting and, if the
directors fail to do so within a
certain period, may themselves
convene a general meeting.

131

 
Notice of General Meeting

Proxy

ENGLAND
Under the Companies Act, at least
21 days’ notice must be given for
an annual general meeting and
any resolutions to be proposed at
the meeting. Subject to a
company’s articles of association
providing for a longer period, at
least 14 days’ notice is required
for any other general meeting of a
public limited company. In
addition, certain matters, such as
the removal of directors or
auditors, require special notice,
which is 28 days’ notice. The
shareholders of a company may in
all cases consent to a shorter
notice period, the proportion of
shareholders’ consent required
being 100% of those entitled to
attend and vote in the case of an
annual general meeting and, in
the case of any other general
meeting, a majority in number of
the members having a right to
attend and vote at the meeting,
being a majority who together
hold not less than 95% in nominal
value of the shares giving a right
to attend and vote at the meeting.

ENGLAND
Under the Companies Act, at any
meeting of shareholders, a
shareholder may designate
another person to attend, speak
and vote at the meeting on their
behalf by proxy.

132

DELAWARE
Under Delaware law, unless otherwise
provided in the certificate of
incorporation or bylaws, written notice
of any meeting of the stockholders
must be given to each stockholder
entitled to vote at the meeting not less
than ten nor more than 60 days before
the date of the meeting and shall
specify the place, date, hour and
purpose or purposes of the meeting.

DELAWARE
Under Delaware law, at any meeting of
stockholders, a stockholder may
designate another person to act for such
stockholder by proxy, but no such
proxy shall be voted or acted upon after
three years from its date, unless the
proxy provides for a longer period. A
director of a Delaware corporation may
not issue a proxy representing the
director’s voting rights as a director.

Preemptive Rights

Authority to Allot

ENGLAND

Under the Companies Act,
“equity securities,” being
(i) shares in the company other
than shares that, with respect to
dividends and capital, carry a
right to participate only up to a
specified amount in a distribution,
referred to as “ordinary shares,”
or (ii) rights to subscribe for, or to
convert securities into, ordinary
shares, proposed to be allotted for
cash must be offered first to the
existing equity shareholders in the
company in proportion to the
respective nominal value of their
holdings, unless an exception
applies or a special resolution to
the contrary has been passed by
shareholders in a general meeting
or the articles of association
provide otherwise in each case in
accordance with the provisions of
the Companies Act.

ENGLAND

Under the Companies Act, the
directors of a company must not
allot shares or grant rights to
subscribe for or convert any
security into shares unless an
exception applies or an ordinary
resolution to the contrary has
been passed by shareholders in a
general meeting or the articles of
association provide otherwise, in
each case in accordance with the
provisions of the Companies Act.

133

DELAWARE
Under Delaware law, shareholders have
no preemptive rights to subscribe to
additional issues of stock or to any
security convertible into such stock
unless, and except to the extent that,
such rights are expressly provided for
in the certificate of incorporation.

DELAWARE
Under Delaware law, if the
corporation’s charter or certificate of
incorporation so provides, the board of
directors has the power to authorize the
issuance of stock. The board may
authorize capital stock to be issued for
consideration consisting of cash, any
tangible or intangible property or any
benefit to the corporation or any
combination thereof. It may determine
the amount of such consideration by
approving a formula. In the absence of
actual fraud in the transaction, the
judgment of the directors as to the
value of such consideration is
conclusive.

Liability of Directors and Officers

DELAWARE
Under Delaware law, a corporation’s
certificate of incorporation may
include a provision eliminating or
limiting the personal liability of a
director to the corporation and its
stockholders for damages arising
from a breach of fiduciary duty as a
director. However, no provision can
limit the liability of a director for:

any breach of the director’s duty
of loyalty to the corporation or its
stockholders;

acts or omissions not in good faith
or that involve intentional
misconduct or a knowing violation
of law;

intentional or negligent payment
of unlawful

ENGLAND

Under the Companies Act, any
provision, whether contained in a
company’s articles of association
or any contract or otherwise, that
purports to exempt a director of a
company, to any extent, from any
liability that would otherwise
attach to him in connection with
any negligence, default, breach of
duty or breach of trust in relation
to the company, is void. Any
provision by which a company
directly or indirectly provides an
indemnity, to any extent, for a
director of the company or of an
associated company against any
liability attaching to him in
connection with any negligence,
default, breach of duty or breach
of trust in relation to the company
of which he is a director is also
void except as permitted by the
Companies Act, which provides
exceptions for the company to
(i) purchase and maintain
insurance against such liability;
(ii) provide a “qualifying third
party indemnity,” or an indemnity
against liability incurred by the
director to a person other than the
company or an associated
company or criminal proceedings
in which he is convicted; and
(iii) provide a “qualifying pension
scheme indemnity,” or an
indemnity against liability
incurred in connection with the
company’s activities as trustee of
an occupational pension plan.

ENGLAND

DELAWARE

134

 
 
 
Voting Rights

Delaware law provides that, unless
otherwise provided in the certificate of
incorporation, each stockholder is
entitled to one vote for each share of
capital stock held by such stockholder.

Under English law, unless a poll
is demanded by the shareholders
of a company or is required by the
chairman of the meeting or the
company’s articles of association,
shareholders shall vote on all
resolutions on a show of hands.
Under the Companies Act, a poll
may be demanded by (i) not
fewer than five shareholders
having the right to vote on the
resolution; (ii) any shareholder(s)
representing not less than 10% of
the total voting rights of all the
shareholders having the right to
vote on the resolution (excluding
any voting rights attaching to
treasury shares); or (iii) any
shareholder(s) holding shares in
the company conferring a right to
vote on the resolution (excluding
any voting rights attaching to
treasury shares) being shares on
which an aggregate sum has been
paid up equal to not less than 10%
of the total sum paid up on all the
shares conferring that right. A
company’s articles of association
may provide more extensive
rights for shareholders to call a
poll.

Under English law, an ordinary
resolution is passed on a show
of hands if it is approved by a
simple majority (more than
50%) of the votes cast by
shareholders present (in person
or by proxy) and entitled to
vote. If a poll is demanded, an
ordinary resolution is passed if
it is approved by holders
representing a simple majority
of the total voting rights of
shareholders present, in person
or by proxy, who, being
entitled to vote, vote on the
resolution. Special resolutions
require the affirmative vote of
not less than 75% of the votes
cast by shareholders present, in
person or by proxy, at the
meeting.

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Shareholder Vote on Certain Transactions

ENGLAND
The Companies Act provides for
schemes of arrangement, which
are arrangements or compromises
between a company and any class
of shareholders or creditors and
used in certain types of
reconstructions, amalgamations,
capital reorganizations or
takeovers. These arrangements
require:

the approval at a shareholders’ or
creditors’ meeting convened by
order of the court, of a majority
in number of shareholders or
creditors representing 75% in
value of the capital held by, or
debt owed to, the class of
shareholders or creditors, or class
thereof present and voting, either
in person or by proxy; and

the approval of the court.

DELAWARE
Generally, under Delaware law, unless
the certificate of incorporation
provides for the vote of a larger
portion of the stock, completion of a
merger, consolidation, sale, lease or
exchange of all or substantially all of a
corporation’s assets or dissolution
requires:

the approval of the board of directors;
and

the approval by the vote of the holders
of a majority of the outstanding stock
or, if the certificate of incorporation
provides for more or less than one vote
per share, a majority of the votes of the
outstanding stock of the corporation
entitled to vote on the matter.

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Standard of Conduct for Directors

ENGLAND

Under English law, a director
owes various statutory and
fiduciary duties to the company,
including:

to act in the way he considers, in
good faith, would be most likely
to promote the success of the
company for the benefit of its
members as a whole;

to avoid a situation in which he
has, or can have, a direct or
indirect interest that conflicts, or
possibly conflicts, with the
interests of the company;

to act in accordance with the
company’s constitution and only
exercise his powers for the
purposes for which they are
conferred;

to exercise independent judgment;

to exercise reasonable care, skill
and diligence;

not to accept benefits from a third
party conferred by reason of his
being a director or doing, or not
doing, anything as a director; and

to declare any interest that he has,
whether directly or indirectly, in a
proposed or existing transaction
or arrangement with the company.

DELAWARE
Delaware law does not contain specific
provisions setting forth the standard of
conduct of a director. The scope of the
fiduciary duties of directors is generally
determined by the courts of the State of
Delaware. In general, directors have a
duty to act without self-interest, on a
well-informed basis and in a manner
they reasonably believe to be in the
best interest of the stockholders.

Directors of a Delaware corporation
owe fiduciary duties of care and loyalty
to the corporation and to its
shareholders. The duty of care
generally requires that a director acts in
good faith, with the care that an
ordinarily prudent person would
exercise under similar circumstances.
Under this duty, a director must inform
himself of all material information
reasonably available regarding a
significant transaction. The duty of
loyalty requires that a director act in a
manner he reasonably believes to be in
the best interests of the corporation. He
must not use his corporate position for
personal gain or advantage. In general,
but subject to certain exceptions,
actions of a director are presumed to
have been made on an informed basis,
in good faith and in the honest belief
that the action taken was in the best
interests of the corporation. However,
this presumption may be rebutted by
evidence of a breach of one of the
fiduciary duties. Delaware courts have
also imposed a heightened standard of
conduct upon directors of a Delaware
corporation who take any action
designed to defeat a threatened change
in control of the corporation.

In addition, under Delaware law, when
the board of directors of a Delaware
corporation approves the sale or break-
up of a corporation, the board of
directors may, in certain circumstances,
have a duty to obtain the highest value
reasonably available to the
shareholders.

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Shareholder Litigation

ENGLAND
Under English law, generally, the
company, rather than its
shareholders, is the proper
claimant in an action in respect of
a wrong done to the company or
where there is an irregularity in
the company’s internal
management. Notwithstanding
this general position, the
Companies Act provides that (i) a
court may allow a shareholder to
bring a derivative claim (that is,
an action in respect of and on
behalf of the company) in respect
of a cause of action arising from a
director’s negligence, default,
breach of duty or breach of trust
and (ii) a shareholder may bring a
claim for a court order where the
company’s affairs have been or
are being conducted in a manner
that is unfairly prejudicial to some
of its shareholders.

DELAWARE
Under Delaware law, a stockholder
may initiate a derivative action to
enforce a right of a corporation if the
corporation fails to enforce the right
itself. The complaint must:

state that the plaintiff was a
stockholder at the time of the
transaction of which the plaintiff
complains or that the plaintiff’s
shares thereafter devolved on the
plaintiff by operation of law; and

allege with particularity the efforts
made by the plaintiff to obtain the
action the plaintiff desires from the
directors and the reasons for the
plaintiff’s failure to obtain the
action; or

state the reasons for not making
the effort.

Additionally, the plaintiff must
remain a stockholder through the
duration of the derivative suit. The
action will not be dismissed or
compromised without the approval of
the Delaware Court of Chancery.

C. Material contracts.

    In addition to the contracts described elsewhere in this Annual Report, the following are summaries of each material contract to which we are a party for
the two years preceding the date of this Annual Report.

Underwriting Agreements

    We entered into an underwriting agreement with Goldman Sachs & Co. LLC and Jefferies LLC as representatives of the underwriters, on June 21, 2018,
with respect to the ADSs sold in our IPO. We have agreed to indemnify the underwriters against certain liabilities, including liabilities under the Securities
Act, and to contribute to payments the underwriters may be required to make in respect of such liabilities.

We entered into an underwriting agreement with Goldman Sachs & Co. LLC and Jefferies LLC as representatives of the underwriters, on April 10,
2019, with respect to the ADSs sold in our April 2019 public offering. We have agreed to indemnify the underwriters against certain liabilities, including
liabilities under the Securities Act, and to contribute to payments the underwriters may be required to make in respect of such liabilities.

We entered into an underwriting agreement with J.P. Morgan Securities LLC and Jefferies LLC as representatives of the underwriters, on January
22,  2020,  with  respect  to  the  ADSs  sold  in  our  January  2020  public  offering.  We  have  agreed  to  indemnify  the  underwriters  against  certain  liabilities,
including liabilities under the Securities Act, and to contribute to payments the underwriters may be required to make in respect of such liabilities.

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We  entered  into  an  underwriting  agreement  with  J.P.  Morgan  Securities  LLC  and  Wells  Fargo  Securities,  LLC  as  representatives  of  the
underwriters,  on  February  9,  2021,  with  respect  to  the  ADSs  sold  in  our  February  2021  public  offering.  We  have  agreed  to  indemnify  the  underwriters
against certain liabilities, including liabilities under the Securities Act, and to contribute to payments the underwriters may be required to make in respect
of such liabilities.

Sales Agreement

On September 18, 2020, we entered into an Open Market Sale Agreement

 with Jefferies LLC, or the Sales Agreement, pursuant to which we
may issue and sell our ADSs having aggregate offering sales proceeds of up to $100,000,000, from time to time, in an at-the-market offering pursuant to
which  Jefferies  will  act  as  sales  agent.  The  Sales  Agreement  contains  customary  representations  and  warranties  of  the  parties  and  indemnification  and
contribution  provisions  under  which  we  and  Jefferies  have  agreed  to  indemnify  each  other  against  certain  liabilities,  including  liabilities  under  the
Securities Act. Each party has right, by giving written notice as specified in the Sales Agreement, to terminate the sales agreement.

SM

University College of London Business Ltd. (UCLB) License

In September 2014, we entered into an exclusive license agreement, or the License, with UCL Business Ltd., or UCLB, the technology transfer
company of University College London, or UCL, to obtain licenses to certain technology rights in the field of cancer therapy and diagnosis. In March 2016
and March 2018, the License was amended to include additional rights. In October 2020, the License was again amended and restated to reflect our election
to have various UCLB patent rights assigned to us, as well as to provide us with access to additional licensed technologies. The License contains customary
indemnification and other risk allocation terms with respect to the licensed technologies and our uses thereof.

    For additional information on our material contracts, please see “Item 4. Information on the Company,” “Item 6. Directors, Senior Management and
Employees,” and “Item 7.B. Related Party Transactions” of this Annual Report.

D. Exchange controls.

    There are no governmental laws, decrees, regulations or other legislation in the United Kingdom that may affect the import or export of capital, including
the availability of cash and cash equivalents for use by us, or that may affect the remittance of dividends, interest, or other payments by us to non-resident
holders of our ordinary shares or ADSs, other than withholding tax requirements. There is no limitation imposed by the laws of England and Wales or our
articles of association on the right of non-residents to hold or vote shares.

E. Taxation.

    The following summary contains a description of material U.K. and U.S. federal income tax consequences of the acquisition, ownership and disposition
of our ADSs. This summary should not be considered a comprehensive description of all the tax considerations that may be relevant to beneficial owners of
ADSs.

Material U.S. Federal Income Tax Considerations for U.S. Holders

    The following is a description of the material U.S. federal income tax consequences to the U.S. Holders described below of owning and disposing of our
ADSs.  It  is  not  a  comprehensive  description  of  all  tax  considerations  that  may  be  relevant  to  a  particular  person’s  decision  to  acquire  securities.  This
discussion applies only to a U.S. Holder that holds our ADSs as a capital asset for tax purposes (generally, property held for investment). In addition, it
does  not  describe  all  of  the  tax  consequences  that  may  be  relevant  in  light  of  a  U.S.  Holder’s  particular  circumstances,  including  state  and  local  tax
consequences,  estate  tax  consequences,  alternative  minimum  tax  consequences,  the  potential  application  of  the  Medicare  contribution  tax,  and  tax
consequences applicable to U.S. Holders subject to special rules, such as:

•

•

•

•

banks, insurance companies, and certain other financial institutions;

U.S. expatriates and certain former citizens or long-term residents of the United States;

dealers or traders in securities who use a mark-to-market method of tax accounting;

persons holding ADSs as part of a hedging transaction, “straddle,” wash sale, conversion transaction or integrated transaction or persons
entering into a constructive sale with respect to ADSs;

139

•

•

•

•

•

•

•

•

persons whose “functional currency” for U.S. federal income tax purposes is not the U.S. dollar;

brokers, dealers or traders in securities, commodities or currencies;

tax-exempt entities or government organizations;

S  corporations,  partnerships,  or  other  entities  or  arrangements  classified  as  partnerships  for  U.S.  federal  income  tax  purposes  (and
investors therein);

regulated investment companies or real estate investment trusts;

persons who acquired ADSs pursuant to the exercise of any employee share option or otherwise as compensation;

persons that own or are deemed to own 10 percent or more of our shares including shares represented by ADSs (by vote or value); and

persons holding our ADSs in connection with a trade or business, permanent establishment, or fixed base outside the United States.

    If an entity that is classified as a partnership for U.S. federal income tax purposes holds ADSs, the U.S. federal income tax treatment of a partner will
generally  depend  on  the  status  of  the  partner  and  the  activities  of  the  partnership.  Partnerships  holding  ADSs  and  partners  in  such  partnerships  are
encouraged to consult their tax advisers as to the particular U.S. federal income tax consequences of holding and disposing of ADSs.

        U.S.  Holders  that  own  (directly,  indirectly,  or  constructively  through  the  application  of  attribution  rules)  10%  or  more  of  our  total  combined  voting
power or value could be subject to adverse U.S. federal income tax consequences pursuant to the controlled foreign corporation rules due to our ownership
of a U.S. subsidiary. Such prospective Holders should consult with their tax advisors as to the tax consequences of acquiring, owning and disposing of our
ADSs.

        The  discussion  is  based  on  the  Internal  Revenue  Code  of  1986,  as  amended,  or  the  Code,  administrative  pronouncements,  judicial  decisions,  final,
temporary and proposed Treasury Regulations, and the income tax treaty between the United Kingdom and the United States, or the Treaty, all as of the
date hereof, changes to any of which may affect the tax consequences described herein—possibly with retroactive effect.

    A “U.S. Holder” is a holder who, for U.S. federal income tax purposes, is a beneficial owner of ADSs who is eligible for the benefits of the Treaty and is:

(i)

(ii)

a citizen or individual resident of the United States;

a corporation, or other entity taxable as a corporation, created or organized in or under the laws of the United States, any state
therein or the District of Columbia;

(iii)

an estate the income of which is subject to U.S. federal income taxation regardless of its source; or

(iv)

a  trust  if  (1)  a  U.S.  court  is  able  to  exercise  primary  supervision  over  the  administration  of  the  trust  and  one  or  more  U.S.
persons have authority to control all substantial decisions of the trust or (2) the trust has a valid election to be treated as a U.S.
person under applicable U.S. Treasury Regulations.

    U.S. Holders are encouraged to consult their tax advisers concerning the U.S. federal, state, local and foreign tax consequences of owning and disposing
of ADSs in their particular circumstances.

    THESE PARAGRAPHS ARE A SUMMARY OF CERTAIN U.S. TAX CONSIDERATIONS AND ARE INTENDED AS A GENERAL GUIDE
ONLY. IT IS RECOMMENDED THAT ALL HOLDERS OF ADSs OBTAIN ADVICE AS TO THE CONSEQUENCES OF THE ACQUISITION,
OWNERSHIP AND DISPOSAL OF THE ADSs IN THEIR OWN SPECIFIC CIRCUMSTANCES FROM THEIR OWN TAX ADVISORS.

    The discussion below assumes that the representations contained in the deposit agreement are true and that the obligations in the deposit agreement and
any related agreement will be complied with in accordance with their terms. A U.S. Holder of an ADS will generally be treated for U.S. federal income tax
purposes as the beneficial owner of the ordinary shares represented by the ADS, and, accordingly, no gain or loss will be recognized upon an exchange of
ADSs for ordinary shares.

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Passive Foreign Investment Company Rules

    If we are classified as a passive foreign investment company, or a PFIC in any taxable year, a U.S. Holder will be subject to special rules generally
intended  to  reduce  or  eliminate  any  benefits  from  the  deferral  of  U.S.  federal  income  tax  that  a  U.S.  Holder  could  derive  from  investing  in  a  non-U.S.
company that does not distribute all of its earnings on a current basis.

A non-U.S. corporation will be classified as a PFIC for any taxable year in which, after applying certain look-through rules, either:

•

•

at least 75% of its gross income is passive income (such as interest income); or

at least 50% of its gross assets (determined on the basis of a quarterly average) is attributable to assets that produce passive income or are
held for the production of passive income.

    We will be treated as owning our proportionate share of the assets and earning our proportionate share of the income of any other corporation, the equity
of which we own, directly or indirectly, 25% or more (by value).

    Based on our current estimates of expected gross assets and income for the current taxable year, we do not believe we will be a PFIC for the year ending
December 31, 2020. However, the application of the PFIC rules is subject to uncertainty in several respects, and therefore, no assurances can be provided
with respect to our PFIC status for our taxable year ending December 31, 2020 or with regard to our PFIC status in the past or in the future.

    A separate determination must be made after the close of each taxable year as to whether we are a PFIC for that year. As a result, our PFIC status may
change from year to year. The total value of our assets for purposes of the asset test generally will be calculated using the market price of the ADSs, which
may fluctuate considerably. Fluctuations in the market price of the ADSs may result in our being a PFIC for any taxable year.

    If we are classified as a PFIC in any year with respect to which a U.S. Holder owns ADSs, we will continue to be treated as a PFIC with respect to such
U.S. Holder in all succeeding years during which the U.S. Holder owns the ADSs, regardless of whether we continue to meet the tests described above
unless (i) we cease to be a PFIC and the U.S. Holder has made a “deemed sale” election under the PFIC rules, (ii) we cease to be a PFIC and the U.S.
Holder  has  a  valid  mark-to-  market  election  in  effect  (as  described  below)  or  (iii)  the  U.S.  Holder  makes  a  Qualified  Electing  Fund  Election,  or  QEF
Election, with respect to all taxable years during such U.S. Holders holding period in which we are a PFIC. However, a U.S. Holder may make a QEF
Election with respect to our ADSs only if we annually provide such U.S. Holder with certain tax information, and we currently do not intend to prepare or
provide such information. As a result, the QEF Election is not expected to be available to a U.S. Holder and the remainder of this discussion assumes that
such election will not be available. If the “deemed sale” election is made, a U.S. Holder will be deemed to have sold the ADSs the U.S. Holder holds at
their fair market value and any gain from such deemed sale would be subject to the rules described below. After the deemed sale election, so long as we do
not become a PFIC in a subsequent taxable year, the U.S. Holder’s ADSs with respect to which such election was made will not be treated as shares in a
PFIC and the U.S. Holder will not be subject to the rules described below with respect to any “excess distribution” the U.S. Holder receives from us or any
gain from an actual sale or other disposition of the ADSs. U.S. Holders should consult their tax advisors as to the possibility and consequences of making a
deemed sale election if we cease to be a PFIC and such election becomes available.

    For each taxable year we are treated as a PFIC with respect to U.S. Holders, U.S. Holders will be subject to special tax rules with respect to any “excess
distribution” such U.S. Holder receives and any gain such U.S. Holder recognizes from a sale or other disposition (including a pledge) of ADSs, unless (i)
such U.S. Holder makes a QEF Election with respect to all taxable years of a U.S. Holder’s holding period during which we are a PFIC or makes a purging
election to cause a deemed sale of the ADSs at their fair market value in conjunction with a QEF election (however, as discussed above, such elections are
expected and assumed not to be available) or (ii) our ADSs constitute “marketable“ securities, and such U.S. Holder makes a mark-to-market election as
discussed  below.  Distributions  a  U.S.  Holder  receives  in  a  taxable  year  that  are  greater  than  125%  of  the  average  annual  distributions  a  U.S.  Holder
received during the shorter of the three preceding taxable years or the U.S. Holder’s holding period for the ADSs will be treated as an excess distribution.
Under these special tax rules:

•

•

the excess distribution or gain will be allocated ratably over a U.S. Holder’s holding period for the ADSs;

the amount allocated to the current taxable year, and any taxable year prior to the first taxable year in which we became a PFIC, will be
treated as ordinary income; and

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•

the  amount  allocated  to  each  other  year  will  be  subject  to  the  highest  tax  rate  in  effect  for  that  year  and  the  interest  charge  generally
applicable to underpayments of tax will be imposed on the resulting tax attributable to each such year.

        The  tax  liability  for  amounts  allocated  to  years  prior  to  the  year  of  disposition  or  the  year  of  an  “excess  distribution”  cannot  be  offset  by  any  net
operating losses for such years, and gains (but not losses) realized on the sale of the ADSs cannot be treated as capital, even if a U.S. Holder holds the
ADSs as capital assets.

    If we are a PFIC, a U.S. Holder will generally be subject to similar rules with respect to distributions we receive from, and our dispositions of the stock
of, any of our direct or indirect subsidiaries that also are PFICs, as if such distributions were indirectly received by, and/or dispositions were indirectly
carried out by, such U.S. Holder. U.S. Holders should consult their tax advisors regarding the application of the PFIC rules to our subsidiaries.

    U.S. Holders can avoid the interest charge on excess distributions or gain relating to the ADSs by making a mark-to-market election with respect to the
ordinary shares, provided that the ADSs are “marketable.” ADSs will be marketable if they are “regularly traded” on certain U.S. stock exchanges or on a
foreign  stock  exchange  that  meets  certain  conditions.  For  these  purposes,  the  ordinary  shares  or  ADSs  will  be  considered  regularly  traded  during  any
calendar year during which they are traded, other than in de minimis quantities, on at least 15 days during each calendar quarter. Any trades that have as
their principal purpose meeting this requirement will be disregarded. Our ADSs are listed on Nasdaq, which is a qualified exchange for these purposes.
Consequently, if our ADSs remain listed on Nasdaq and are regularly traded, and you are a holder of ADSs, we expect the mark-to-market election would
be available to U.S. Holders if we are a PFIC. Each U.S. Holder should consult its tax advisor as to the whether a mark-to-market election is available or
advisable with respect to the ADSs.

    A U.S. Holder that makes a mark-to-market election must include as ordinary income for each year an amount equal to the excess, if any, of the fair
market value of the ADSs at the close of the taxable year over the U.S. Holder’s adjusted tax basis in the ADSs. Accordingly, such mark-to-market election
may accelerate the recognition of income without a corresponding receipt of cash. An electing holder may also claim an ordinary loss deduction for the
excess, if any, of the U.S. Holder’s adjusted basis in the ADSs over the fair market value of the ADSs at the close of the taxable year, but this deduction is
allowable only to the extent of any net mark-to-market gains for prior years. Gains from an actual sale or other disposition of the ADSs will be treated as
ordinary income, and any losses incurred on a sale or other disposition of the ADSs will be treated as an ordinary loss to the extent of any net mark-to-
market gains for prior years. Once made, the election cannot be revoked without the consent of the Internal Revenue Service, or the IRS, unless the ADSs
cease to be marketable.

    However, a mark-to-market election generally cannot be made for equity interests in any lower- tier PFICs that we own, unless shares of such lower-tier
PFIC are themselves “marketable.” As a result, even if a U.S. Holder validly makes a mark-to-market election with respect to our ADSs, the U.S. Holder
may continue to be subject to the PFIC rules (described above) with respect to its indirect interest in any of our investments that are treated as an equity
interest in a PFIC for U.S. federal income tax purposes. U.S. Holders should consult their tax advisors as to the availability and desirability of a mark-to-
market election, as well as the impact of such election on interests in any lower-tier PFICs.

    Unless otherwise provided by the U.S. Treasury, each U.S. shareholder of a PFIC is required to file an annual report containing such information as the
U.S. Treasury may require. A U.S. Holder’s failure to file the annual report will cause the statute of limitations for such U.S. Holder’s U.S. federal income
tax return to remain open with regard to the items required to be included in such report until three years after the U.S. Holder files the annual report, and,
unless such failure is due to reasonable cause and not willful neglect, the statute of limitations for the U.S. Holder’s entire U.S. federal income tax return
will remain open during such period. U.S. Holders should consult their tax advisors regarding the requirements of filing such information returns under
these rules.

ADSs

    A U.S. Holder of ADSs will generally be treated for U.S. federal income tax purposes as the owner of the underlying ordinary shares that such ADSs
represent. Accordingly, no gain or loss will be recognized if a U.S. Holder exchanges ADSs for the underlying shares represented by those ADSs.

    The U.S. Treasury has expressed concern that parties to whom ADSs are released before shares are delivered to the Depositary or intermediaries in the
chain of ownership between holders and the issuer of the security underlying the ADSs, may be taking actions that are inconsistent with the claiming of
foreign  tax  credits  by  U.S.  Holders  of  ADSs.  These  actions  would  also  be  inconsistent  with  the  claiming  of  the  reduced  rate  of  tax,  described  below,
applicable  to  dividends  received  by  certain  non-corporate  U.S.  Holders.  Accordingly,  the  creditability  of  non-U.S.  withholding  taxes  (if  any),  and  the
availability of the reduced tax rate for

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dividends received by certain non-corporate U.S. Holders, each described below, could be affected by actions taken by such parties or intermediaries.

Taxation of Distributions

    Subject to the discussion above under “Passive Foreign Investment Company Rules,” distributions paid on ADSs, other than certain pro rata distributions
of ADSs, will generally be treated as dividends to the extent paid out of our current or accumulated earnings and profits (as determined under U.S. federal
income  tax  principles).  Because  we  may  not  calculate  our  earnings  and  profits  under  U.S.  federal  income  tax  principles,  we  expect  that  distributions
generally  will  be  reported  to  U.S.  Holders  as  dividends.  Subject  to  applicable  limitations,  dividends  paid  to  certain  non-corporate  U.S.  Holders  may  be
taxable at preferential rates applicable to “qualified dividend income.” However, the qualified dividend income treatment will not apply if we are treated as
a PFIC with respect to the U.S. Holder for the taxable year in which a dividend is paid or the preceding year. The amount of the dividend will be treated as
foreign-source dividend income to U.S. Holders and will not be eligible for the dividends-received deduction generally available to U.S. corporations under
the Code. Dividends will generally be included in a U.S. Holder’s income on the date of the U.S. Holder’s receipt of the dividend. The amount of any
dividend  income  paid  in  foreign  currency  will  be  the  U.S.  dollar  amount  calculated  by  reference  to  the  exchange  rate  in  effect  on  the  date  of  actual  or
constructive receipt, regardless of whether the payment is in fact converted into U.S. dollars. If the dividend is converted into U.S. dollars on the date of
receipt, a U.S. Holder should not be required to recognize foreign currency gain or loss in respect of the dividend income. A U.S. Holder may have foreign
currency gain or loss if the dividend is converted into U.S. dollars after the date of receipt. Such gain or loss would generally be treated as U.S.-source
ordinary  income  or  loss.  The  amount  of  any  distribution  of  property  other  than  cash  (and  other  than  certain  pro  rata  distributions  of  ADSs  or  rights  to
acquire ADSs) will be the fair market value of such property on the date of distribution.

    For foreign tax credit limitation purposes, our dividends will generally be treated as passive category income. Because no U.K. income taxes will be
withheld from dividends on ADSs, there will be no creditable foreign taxes associated with any dividends that a U.S. Holder will receive.

Sale or Other Taxable Disposition of ADSs

    Subject to the discussion above under “Passive Foreign Investment Company Rules,” gain or loss realized on the sale or other taxable disposition of
ADSs will be capital gain or loss, and will be a long-term capital gain or loss if the U.S. Holder held the ADSs for more than one year. The amount of the
gain or loss will equal the difference between the U.S. Holder’s tax basis in the ADSs disposed of and the amount realized on the disposition, in each case
as determined in U.S. dollars. This gain or loss will generally be U.S.-source gain or loss for foreign tax credit purposes. The deductibility of capital losses
is subject to limitations.

    If the consideration received by a U.S. Holder is not paid in U.S. dollars, the amount realized will be the U.S. dollar value of the payment received
determined  by  reference  to  the  spot  rate  of  exchange  on  the  date  of  the  sale  or  other  disposition.  However,  if  the  ADSs  are  treated  as  traded  on  an
“established securities market” and a U.S. Holder is either a cash basis taxpayer or an accrual basis taxpayer that has made a special election (which must
be applied consistently from year to year and cannot be changed without the consent of the IRS), such U.S. Holder will determine the U.S. dollar value of
the amount realized in a non-U.S. dollar currency by translating the amount received at the spot rate of exchange on the settlement date of the sale. If a U.S.
Holder is an accrual basis taxpayer that is not eligible to or does not elect to determine the amount realized using the spot rate on the settlement date, such
U.S.  Holder  will  recognize  foreign  currency  gain  or  loss  to  the  extent  of  any  difference  between  the  U.S.  dollar  amount  realized  on  the  date  of  sale  or
disposition and the U.S. dollar value of the currency received at the spot rate on the settlement date.

        WE  STRONGLY  URGE  YOU  TO  CONSULT  YOUR  TAX  ADVISOR  REGARDING  THE  IMPACT  OF  OUR  PFIC  STATUS  ON  YOUR
INVESTMENT IN THE ADSs AS WELL AS THE APPLICATION OF THE PFIC RULES TO YOUR INVESTMENT IN THE ADSs.

Information Reporting and Backup Withholding

    Payments of dividends and sales proceeds that are made within the United States or through certain U.S.-related financial intermediaries generally are
subject to information reporting, and may be subject to backup withholding, unless (i) the U.S. Holder is a corporation or other exempt recipient or (ii) in
the case of backup withholding, the U.S. Holder provides a correct taxpayer identification number and certifies that it is not subject to backup withholding
(generally, by providing an IRS Form W-9).

    Backup withholding is not an additional tax. The amount of any backup withholding from a payment to a U.S. Holder will be allowed as a credit against
the holder’s U.S. federal income tax liability and may entitle it to a refund, provided that the required information is timely furnished to the IRS.

143

Information with Respect to Foreign Financial Assets

    Certain U.S. Holders who are individuals (and, under regulations, certain entities) may be required to report information relating to the ordinary shares or
ADSs, subject to certain exceptions (including an exception for ordinary shares or ADSs held in accounts maintained by certain U.S. financial institutions).
Such U.S. Holders who fail to timely furnish the required information may be subject to a penalty. Additionally, if a U.S. Holder does not file the required
information, the statute of limitations with respect to tax returns of the U.S. Holder to which the information relates may not close until three years after
such information is filed. U.S. Holders should consult their tax advisers regarding their reporting obligations with respect to their ownership and disposition
of the ADSs.

U.K. Taxation

    The following is intended as a general guide to current U.K. tax law and HM Revenue & Customs, or HMRC, published practice applying as at the date
of this Annual Report (both of which are subject to change at any time, possibly with retrospective effect) relating to the holding of ADSs. It does not
constitute legal or tax advice and does not purport to be a complete analysis of all U.K. tax considerations relating to the holding of ADSs, or all of the
circumstances in which holders of ADSs may benefit from an exemption or relief from U.K. taxation. It is written on the basis that we do not (and will not)
directly or indirectly derive 75% or more of our qualifying asset value from U.K. land, and that we are and remain solely resident in the United Kingdom
for tax purposes and will therefore be subject to the U.K. tax regime and not the U.S. tax regime save as set out above under “Material U.S. Federal Income
Tax Considerations for U.S. Holders.”

    Except to the extent that the position of non-U.K. resident persons is expressly referred to, this guide relates only to persons who are resident (and, in the
case of individuals, domiciled or deemed domiciled) for tax purposes solely in the United Kingdom and do not have a permanent establishment, branch,
agency  (or  equivalent)  or  fixed  base  in  any  other  jurisdiction  with  which  the  holding  of  the  ADSs  is  connected,  or  U.K.  Holders,  who  are  absolute
beneficial owners of the ADSs (where the ADSs are not held through an Individual Savings Account or a Self-Invested Personal Pension) and who hold the
ADSs as investments.

    This guide may not relate to certain classes of U.K. Holders, such as (but not limited to):

•

•

•

•

•

•

•

•

•

persons who are connected with the company;

financial institutions;

insurance companies;

charities or tax-exempt organizations;

collective investment schemes;

pension schemes;

market makers, intermediaries, brokers or dealers in securities;

persons who have (or are deemed to have) acquired their ADSs by virtue of an office or employment or who are or have been officers or
employees of the company or any of its affiliates; and

individuals who are subject to U.K. taxation on a remittance basis.

    The decision of the First-tier Tribunal (Tax Chamber) in HSBC Holdings PLC and The Bank of New York Mellon Corporation v HMRC (2012) cast
some doubt on whether a holder of a depositary receipt is the beneficial owner of the underlying shares. However, based on published HMRC guidance we
would expect that HMRC will regard a holder of ADSs as holding the beneficial interest in the underlying shares and therefore these paragraphs assume
that a holder of ADSs is the beneficial owner of the underlying ordinary shares and any dividends paid in respect of the underlying ordinary shares (where
the dividends are regarded for U.K. purposes as that person’s own income) for U.K. direct tax purposes.

    THESE PARAGRAPHS ARE A SUMMARY OF CERTAIN U.K. TAX CONSIDERATIONS AND ARE INTENDED AS A GENERAL GUIDE
ONLY.  IT  IS  RECOMMENDED  THAT  ALL  HOLDERS  OF  ADSs  OBTAIN  ADVICE  AS  TO  THE  CONSEQUENCES  OF  THE
ACQUISITION, OWNERSHIP AND DISPOSAL OF THE ADSs IN THEIR OWN

144

    
SPECIFIC  CIRCUMSTANCES  FROM  THEIR  OWN  TAX  ADVISORS.  IN  PARTICULAR,  NON-U.K.  RESIDENT  OR  DOMICILED
PERSONS ARE ADVISED TO CONSIDER THE POTENTIAL IMPACT OF ANY RELEVANT DOUBLE TAXATION AGREEMENTS.

Dividends

Withholding Tax

    Dividends paid by us will not be subject to any withholding or deduction for or on account of U.K. tax.

Income Tax

    An individual U.K. Holder may, depending on his or her particular circumstances, be subject to U.K. tax on dividends received from us. An individual
holder of ADSs who is not resident for tax purposes in the United Kingdom should not be chargeable to U.K. income tax on dividends received from us
unless he or she carries on (whether solely or in partnership) a trade, profession or vocation in the United Kingdom through a branch or agency to which
the  ADSs  are  attributable.  There  are  certain  exceptions  for  trading  in  the  United  Kingdom  through  independent  agents,  such  as  some  brokers  and
investment managers.

    All dividends received by an individual U.K. Holder from us or from other sources will form part of that U.K. Holder’s total income for income tax
purposes and will constitute the top slice of that income. A nil rate of income tax will apply to the first £2,000 of taxable dividend income received by the
individual U.K. Holder in a tax year. Income within the nil rate band will be taken into account in determining whether income in excess of the £2,000 tax-
free allowance falls within the basic rate, higher rate or additional rate tax bands. Dividend income in excess of the tax-free allowance will (subject to the
availability of any income tax personal allowance) be taxed at 7.5% to the extent that the excess amount falls within the basic rate tax band, 32.5% to the
extent that the excess amount falls within the higher rate tax band and 38.1% to the extent that the excess amount falls within the additional rate tax band.

Corporation Tax

    A corporate holder of ADSs who is not resident for tax purposes in the United Kingdom should not be chargeable to U.K. corporation tax on dividends
received from us unless it carries on (whether solely or in partnership) a trade in the United Kingdom through a permanent establishment to which the
ADSs are attributable.

    Corporate U.K. Holders should not be subject to U.K. corporation tax on any dividend received from us so long as the dividends qualify for exemption,
which should be the case, although certain conditions must be met. If the conditions for the exemption are not satisfied, or such U.K. Holder elects for an
otherwise exempt dividend to be taxable, U.K. corporation tax will be chargeable on the amount of any dividends (at the current rate of 19%).

Chargeable Gains

    A disposal or deemed disposal of ADSs by a U.K. Holder may, depending on the U.K. Holder’s circumstances and subject to any available exemptions
or reliefs (such as the annual exemption), give rise to a chargeable gain or an allowable loss for the purposes of U.K. capital gains tax and corporation tax
on chargeable gains.

    If an individual U.K. Holder who is subject to U.K. income tax at either the higher or the additional rate is liable to U.K. capital gains tax on the disposal
of ADSs, the current applicable rate will be 20%. For an individual U.K. Holder who is subject to U.K. income tax at the basic rate and liable to U.K.
capital  gains  tax  on  such  disposal,  the  current  applicable  rate  would  be  10%,  save  to  the  extent  that  any  capital  gains,  when  aggregated  with  the  U.K.
Holder’s other taxable income and gains in the relevant tax year, exceed the unused basic rate tax band. In that case, the rate currently applicable to the
excess would be 20%

    If a corporate U.K. Holder becomes liable to U.K. corporation tax on the disposal (or deemed disposal) of ADSs, the main rate of U.K. corporation tax
(currently 19%) would apply.

    A holder of ADSs which is not resident for tax purposes in the United Kingdom should not normally be liable to U.K. capital gains tax or corporation tax
on chargeable gains on a disposal (or deemed disposal) of ADSs unless the person is carrying on (whether solely or in partnership) a trade, profession or
vocation in the United Kingdom through a branch or agency (or, in the case of a corporate holder of ADSs, through a permanent establishment) to which
the ADSs are attributable. However, an individual holder of ADSs who has ceased to be resident for tax purposes in the United Kingdom for a period of
less than five years and who disposes of

145

ADSs  during  that  period  may  be  liable  on  his  or  her  return  to  the  United  Kingdom  to  U.K.  tax  on  any  capital  gain  realized  (subject  to  any  available
exemption or relief).

Stamp Duty and Stamp Duty Reserve Tax

    The discussion below relates to the holders of our ordinary shares or ADSs wherever resident, however it should be noted that special rules may apply to
certain persons such as market makers, brokers, dealers or intermediaries.

Issue of Ordinary Shares

No U.K. stamp duty or stamp duty reserve tax, or SDRT, is payable on the issue of the underlying ordinary shares in the company.

Transfers of Ordinary Shares

    An unconditional agreement to transfer ordinary shares in certificated form will normally give rise to a charge to SDRT at the rate of 0.5% of the amount
or value of the consideration payable for the transfer. The purchaser of the shares is liable for the SDRT. Transfers of ordinary shares in certificated form
are generally also subject to stamp duty at the rate of 0.5% of the amount or value of the consideration given for the transfer (rounded up to the next £5.00).
Stamp duty is normally paid by the purchaser. The charge to SDRT will be canceled or, if already paid, repaid (generally with interest), where a transfer
instrument has been duly stamped within six years of the charge arising (either by paying the stamp duty or by claiming an appropriate relief) or if the
instrument is otherwise exempt from stamp duty.

    An unconditional agreement to transfer ordinary shares to, or to a nominee or agent for, a person whose business is or includes the issue of depositary
receipts or the provision of clearance services will generally be subject to SDRT (or, where the transfer is effected by a written instrument, stamp duty) at a
higher rate of 1.5% of the amount or value of the consideration given for the transfer unless the clearance service has made and maintained an election
under section 97A of the U.K. Finance Act 1986, or a section 97A election. It is understood that HMRC regards the facilities of DTC as a clearance service
for these purposes and we are not aware of any section 97A election having been made by DTC.

    However, no SDRT is generally payable where the transfer of ordinary shares to a clearance service or depositary receipt system is an integral part of an
issue of share capital.

    Any stamp duty or SDRT payable on a transfer of ordinary shares to a depositary receipt system or clearance service will in practice generally be paid by
the participants in the clearance service or depositary receipt system.

Transfers of ADSs

No SDRT should be required to be paid on a paperless transfer of ADSs through the clearance service facilities of DTC, provided that no section

97A election has been made by DTC, and such ADSs are held through DTC at the time of any agreement for their transfer.

No U.K. stamp duty will in practice be payable on a written instrument transferring an ADS provided that the instrument of transfer is executed
and  remains  at  all  times  outside  the  United  Kingdom.  Where  these  conditions  are  not  met,  the  transfer  of,  or  agreement  to  transfer,  an  ADS  could,
depending on the circumstances, attract a charge to U.K. stamp duty at the rate of 0.5% of the amount or value of the consideration. If it is necessary to pay
stamp duty, it may also be necessary to pay interest and penalties.

F. Dividends and paying agents.

Not applicable.

G. Statement by experts.

Not applicable.

H. Documents on display.

        We  are  subject  to  the  information  reporting  requirements  of  the  Exchange  Act  applicable  to  foreign  private  issuers  and  file  reports  under  those
requirements with the SEC. Those reports may be inspected without charge at the locations described below. As a

146

foreign private issuer, we are exempt from the rules under the Exchange Act related to the furnishing and content of proxy statements, and our officers,
directors and principal shareholders are exempt from the reporting and short-swing profit recovery provisions contained in Section 16 of the Exchange Act.
In addition, we are not required under the Exchange Act to file periodic reports and financial statements with the SEC as frequently or as promptly as U.S.
companies whose securities are registered under the Exchange Act.

    We maintain a corporate website at www.autolus.com. Information contained in, or that can be accessed through, our website is not a part of, and shall
not  be  incorporated  by  reference  into,  this  Annual  Report.  We  have  included  our  website  address  in  this  Annual  Report  solely  as  an  inactive  textual
reference.

    The SEC maintains a website (www.sec.gov) that contains reports, proxy and information statements and other information regarding registrants, such as
us, that file electronically with the SEC.

    With respect to references made in this Annual Report to any contract or other document of our company, such references are not necessarily complete
and you should refer to the exhibits attached or incorporated by reference to this Annual Report for copies of the actual contract or document.

I. Subsidiary Information.

Not applicable.

Item 11.     Quantitative and Qualitative Disclosures About Market Risk.

    We are exposed to market risks in the ordinary course of our business, which are principally limited to interest rate fluctuations and foreign currency
exchange rate fluctuations. We maintain significant amounts of cash and cash equivalents that are in excess of federally insured limits in various currencies,
placed with one or more financial institutions for varying periods according to expected liquidity requirements.

Interest Rate Risk

    As of December 31, 2020 and 2019, we had cash of $153.3 million and $210.6 million, respectively. Our exposure to interest rate sensitivity is impacted
by changes in the underlying U.S. and U.K. bank interest rates. Our surplus cash and cash equivalents have been invested in interest-bearing savings and
money market accounts from time to time. We have not entered into investments for trading or speculative purposes. Due to the conservative nature of our
investment portfolio, which is predicated on capital preservation of investments with short-term maturities, we do not believe an immediate one percentage
point change in interest rates would have a material effect on the fair market value of our portfolio, and we, therefore, do not expect our operating results or
cash flows to be significantly affected by changes in market interest rates.

Foreign Currency Exchange Risk

    We maintain our consolidated financial statements in our functional currency, which is pounds sterling. Monetary assets and liabilities denominated in
currencies other than the functional currency are translated into the functional currency at rates of exchange prevailing at the balance sheet dates. Non-
monetary assets and liabilities denominated in foreign currencies are translated into the functional currency at the exchange rates prevailing at the date of
the transaction. Exchange gains or losses arising from foreign currency transactions are included in the determination of net income (loss) for the respective
periods. We recorded a foreign exchange loss of $0.2 million for the year ended December 31, 2020, a foreign exchange gain of $4.6 million for the year
ended  December  31,  2019,  $1.1  million  for  the  three  months  ended  December  31,  2018,  and  a  $4.0  million  foreign  exchange  loss  for  the  year  ended
December 31, 2018, respectively, which are included in other income in the statements of operations and comprehensive loss.

    For financial reporting purposes, our consolidated financial statements are prepared using the functional currency and translated into the U.S. dollar.
Assets and liabilities are translated at the exchange rates at the balance sheet dates and revenue and expenses are translated at the average exchange rates
and shareholders’ equity is translated based on historical exchange rates. Translation adjustments are not included in determining net income (loss) but are
included in foreign exchange adjustment to accumulated other comprehensive income (loss), a component of shareholders’ equity.

    We do not currently engage in currency hedging activities in order to reduce our currency exposure, but we may begin to do so in the future. Instruments
that may be used to hedge future risks may include foreign currency forward and swap contracts. These

147

instruments  may  be  used  to  selectively  manage  risks,  but  there  can  be  no  assurance  that  we  will  be  fully  protected  against  material  foreign  currency
fluctuations.

Item 12. Description of Securities Other than Equity Securities.

A. Debt Securities

Not applicable.

B. Warrants and Rights.

Not applicable.

C. Other Securities.

Not applicable.

D. American Depositary Shares.

    Citibank, N.A., as Depositary bank, registers and delivers our American Depositary Shares, also referred to as ADSs. Each ADS represents one ordinary
share (or a right to receive one ordinary share) deposited with Citibank, N.A., London Branch, or any successor, as custodian for the Depositary. Each ADS
will also represent any other securities, cash or other property which may be held by the Depositary in respect of the Depositary facility. The Depositary’s
corporate office at which the ADSs are administered is located at 388 Greenwich Street, New York, New York 10013.  A deposit agreement among us, the
Depositary and the ADS holders sets out ADS holder rights as well as the rights and obligations of the Depositary. A form of the deposit agreement is
incorporated by reference as an exhibit to this Annual Report.

Fees and Charges

    The following table shows the fees and charges that a holder of our ADSs may have to pay, either directly or indirectly. The majority of these costs are
set by the Depositary bank and are subject to change:

SERVICE
Issuance of ADSs (e.g., an issuance of ADS upon a deposit of ordinary shares or upon a change in the ADS(s)-to-

ordinary shares ratio), excluding ADS issuances as a result of distributions of ordinary shares

Cancellation of ADSs (e.g., a cancellation of ADSs for delivery of deposited property or upon a change in the

ADS(s)-to-ordinary shares ratio, or for any other reason)

Distribution of cash dividends or other cash distributions (e.g., upon a sale of rights and other entitlements)
Distribution of ADSs pursuant to (i) share dividends or other free share distributions, or (ii) exercise of rights to

purchase additional ADSs

Distribution of securities other than ADSs or rights to purchase additional ADSs (e.g., upon a spin-off)
ADS Services

FEE

Up to $0.05 per ADS issued

Up to $0.05 per ADS canceled
Up to $0.05 per ADS held

Up to $0.05 per ADS held
Up to $0.05 per ADS held
Up to $0.05 per ADS held on 
the applicable record date(s) 
established by the Depositary

As an ADS holder, you will also be responsible to pay certain charges such as:

•

•

taxes (including applicable interest and penalties) and other governmental charges;

the registration fees as may from time to time be in effect for the registration of ordinary shares on the share register and applicable to transfers
of ordinary shares to or from the name of the custodian, the Depositary or any nominees upon the making of deposits and withdrawals,
respectively;

• certain cable, telex and facsimile transmission and delivery expenses;

•

the expenses and charges incurred by the Depositary in the conversion of foreign currency;

148

 
•

•

the fees and expenses incurred by the Depositary in connection with compliance with exchange control regulations and other regulatory
requirements applicable to ordinary shares, ADSs and ADRs; and

the fees and expenses incurred by the Depositary, the custodian or any nominee in connection with the servicing or delivery of deposited
property.

149

Item 13. Defaults, Dividend Arrearages and Delinquencies.

Not applicable.

PART II

Item 14. Material Modifications to the Rights of Security Holders and Use of Proceeds.

A.     Not applicable.

B.     Not applicable.

C.     Not applicable.

D.     Not applicable.

E.     Use of Proceeds.

    Not applicable.

Item 15. Controls and Procedures.

A. Disclosure Controls and Procedures.

    We maintain “disclosure controls and procedures,” as defined in Rules 13a-15(e) and 15d-15(e) under the Securities Exchange Act of 1934, as amended,
or the Exchange Act, that are designed to ensure that information required to be disclosed by us in the reports that we file or submit under the Exchange
Act  is  recorded,  processed,  summarized  and  reported,  within  the  time  periods  specified  in  the  SEC’s  rules  and  forms,  and  is  accumulated  and
communicated to our management, including our Chief Executive Officer and Chief Financial Officer, as appropriate to allow timely decisions regarding
required disclosure. Our management, with the participation of our Chief Executive Officer and Chief Financial Officer, has evaluated the effectiveness of
our disclosure controls and procedures (as defined in Exchange Act Rules 13a-15(e) and 15d-15(e)) as of December 31, 2020. Based on such evaluation,
our  Chief  Executive  Officer  and  Chief  Financial  Officer  have  concluded  that,  as  of  December  31,  2020,  our  disclosure  controls  and  procedures  were
effective.

B. Management’s Annual Report on Internal Control over Financial Reporting.

    Our management is responsible for establishing and maintaining adequate internal control over our financial reporting, as such term is defined in Rule
13a-15(f) under the Exchange Act.

    Our management, under the supervision and with the participation of our Chief Executive Officer and Chief Financial Officer, conducted an assessment
of the evaluation of the effectiveness of our internal control over financial reporting as of December 31, 2020 based on the criteria set forth in “Internal
Control - Integrated Framework (2013)” issued by the Committee of Sponsoring Organizations of the Treadway Commission.

    Based on this assessment, our management concluded that our internal control over financial reporting was effective as of December 31, 2020.

C. Attestation Report of the Registered Public Accounting Firm.

    This Annual Report does not include an attestation report of our registered public accounting firm due to a transition period established by rules of the
SEC for newly public companies. For so long as we qualify as an "emerging growth company" as defined under the JOBS Act, our independent registered
accounting firm is not required to issue an attestation report on our internal control over financial reporting.

150

    
D. Changes in Internal Control Over Financial Reporting.

    There were no changes in our internal control over financial reporting (as defined in Rule 13a-15(f) of the Exchange Act) that occurred during the period
covered by this Annual Report that has materially affected, or is reasonably likely to materially affect, our internal control over financial reporting.

Item 15T. Controls and Procedures.

Not applicable.

Item 16. [Reserved]

Item 16A. Audit Committee Financial Expert.

    Our Board has determined that Ms. Bain is an “audit committee financial expert” as defined by SEC rules and has the requisite financial sophistication
under the applicable rules and regulations of the Nasdaq Stock Market. Ms. Bain is independent as such term is defined in Rule 10A-3 under the Exchange
Act and under the listing standards of the Nasdaq Stock Market.

Item 16B. Code of Ethics.

    We have adopted a Code of Business Conduct and Ethics, or the Code of Ethics, that is applicable to all of our employees, officers and directors and is
available on our website at https://www.autolus.com/investor-relations/corporate-governance/documents-charters. Information contained on, or that can be
accessed through, our website does not constitute a part of this report and is not incorporated by reference herein. If we make any amendment to the Code
of Ethics or grant any waivers, including any implicit waiver, from a provision of the Code of Ethics, we will disclose the nature of such amendment or
waiver on our website to the extent required by the rules and regulations of the SEC. Under Item 16B of Form 20-F, if a waiver or amendment of the Code
of Ethics applies to our principal executive officer, principal financial officer, principal accounting officer or controller and relates to standards promoting
any of the values described in Item 16B(b) of Form 20-F, we are required to disclose such waiver or amendment on our website in accordance with the
requirements of Instruction 4 to such Item 16B.

Item 16C. Principal Accountant Fees and Services.

        Ernst  &  Young  LLP  has  served  as  our  independent  registered  public  accountant  since  September  2017  and  has  audited  our  consolidated  financial
statements for the years ended December 31, 2020 and 2019, the three months ended December 31, 2018 and the years ended September 30, 2018, 2017
and 2016.

    The following table shows the aggregate fees for services rendered by Ernst & Young LLP to us and our subsidiaries for the year ended December 31,
2020 and 2019.

Audit fees
Audit-related fees
Total

$

$

2020

2019*

Year Ended December 31,

(in thousands)

697 
247 
944 

$

$

649 
114 
763 

*Fees for 2019 have been reclassified for comparability due to a change in the definitions for audit fees and audit-related fees in the current year

Audit fees.  Audit fees consisted of fees for the audit of our annual financial statements and other professional services provided in connection with the
statutory and regulatory filings or engagements, including fees for the review of our interim financial information.

151

Audit-related fees. Audit related fees include fees for assurance reporting on our current and historical financial information included in our SEC
registration statements in connection with our follow-on capital raises and our at-the-market facility program, including services that generally only the
independent accountant can reasonably provide such as comfort letters.

Audit Committee Pre-Approval Policies and Procedures

Our audit committee reviews and pre-approves the scope and the cost of audit services related to us and permissible non-audit services performed by the
independent auditors, other than those for de minimis services which are approved by the audit committee prior to the completion of the audit. All of the
services related to us provided by Ernst & Young LLP during the year ended December 31, 2020 have been pre-approved by the audit committee.

Item 16D. Exemptions from the Listing Standards for Audit Committees.

Not applicable

Item 16E Purchases of Equity Securities by the Issuer and Affiliated Purchasers.

Not applicable.

Item 16F. Change in Registrant’s Certifying Accountant.

Not applicable

Item 16G. Corporate Governance.

    We are a “foreign private issuer,” as defined by the SEC. As a result, in accordance with Nasdaq listing requirements, we may rely on home country
governance  requirements  and  certain  exemptions  thereunder  rather  than  complying  with  Nasdaq  corporate  governance  standards.  While  we  voluntarily
follow most Nasdaq corporate governance rules, we may choose to take advantage of the following limited exemptions:

▪

▪

▪

▪

Exemption  from  filing  quarterly  reports  on  Form  10-Q  containing  unaudited  financial  and  other  specified  information  or  current  reports  on
Form 8-K upon the occurrence of specified significant events.

Exemption from Section 16 rules requiring insiders to file public reports of their share ownership and trading activities and liability for insiders
who profit from trades in a short period of time, which will provide less data in this regard than shareholders of U.S. companies that are subject to
the Exchange Act.

Exemption from the requirement that our board have a compensation committee that is composed entirely of independent directors with a written
charter addressing the committee’s purpose and responsibilities.

Exemption from the requirement to have independent director oversight of director nominations.

We intend to follow U.K. corporate governance practices in lieu of Nasdaq corporate governance requirements as follows:

▪ We  do  not  intend  to  follow  Nasdaq  Rule  5620(c)  regarding  quorum  requirements  applicable  to  meetings  of  shareholders.  Such  quorum
requirements  are  not  required  under  English  law.  In  accordance  with  generally  accepted  business  practice,  our  Articles  of  Association  provide
alternative quorum requirements that are generally applicable to meetings of shareholders.

▪ We do not intend to follow Nasdaq Rule 5605(b)(2), which requires that independent directors regularly meet in executive sessions where only

independent directors are present. Our independent directors may choose to meet in executive sessions at their discretion.

Although  we  may  rely  on  certain  home  country  corporate  governance  practices,  we  must  comply  with  Nasdaq’s  Notification  of  Noncompliance
requirement (Nasdaq Rule 5625) and the Voting Rights requirement (Nasdaq Rule 5640). Further, we must have an audit committee that satisfies Nasdaq
Rule 5605(c)(3), which addresses audit committee responsibilities and authority and requires that the audit committee consist of members who meet the
independence requirements of Nasdaq Rule 5605(c)(2)(A)(ii).

152

We intend to take all actions necessary for us to maintain compliance as a foreign private issuer under the applicable corporate governance requirements of
the Sarbanes-Oxley Act, the rules adopted by the SEC and Nasdaq listing rules. Accordingly, our shareholders will not have the same protections afforded
to shareholders of companies that are subject to all of the corporate governance requirements of Nasdaq.

Item 16H. Mine Safety Disclosure

Not applicable

153

Item 17. Financial Statements.

See the financial statements beginning on page F-1 of this Annual Report.

PART III

Item 18. Financial Statements.

Not applicable.

Item 19. Exhibits.

EXHIBIT
NUMBER

1.1

2.1

2.2

2.3
2.4

4.1#

4.2#

4.3+

4.4+
4.5+

DESCRIPTION OF EXHIBIT

Articles of Association of Autolus
Therapeutics plc.
Deposit Agreement by and among the
registrant, Citibank, N.A., as the Depositary
bank and the holders and beneficial owners
of American Depositary Shares issued
thereunder.
Form of American Depositary Receipt
(included in exhibit 2.1).

Autolus Therapeutics plc, Registration
Rights Agreement, dated as June 26, 2018
Description of Securities
License Agreement, dated as of September
25, 2014 by and between the registrant and
UCL Business Ltd., as amended on March
2, 2016 and March 28, 2018.
Supply Agreement, dated as of March 23,
2018, by and between the registrant and
Miltenyi Biotec GmbH.
Autolus Therapeutics plc 2018 Equity
Incentive Plan.
Non-employee Sub Plan to the Autolus
Therapeutics plc 2018 Equity Incentive
Plan.
Management Incentive Compensation Plan.

INCORPORATED BY REFERENCE

SCHEDULE/
FORM

FILE NUMBER

EXHIBIT

FILE
DATE

Form F-1/A

333-224720

3.1

6/19/18

Form F-6/A

333-224837

99.(a)

6/19/18

Form F-6/A

333-224837

99.(a)

6/19/18

Form 20-F

001-38547

2.4

3/3/20

Form F-1/A

333-224720

10.1

5/10/18

Form F-1/A

333-224720

Form F-1/A

333-224720

Form F-1/A
Form F-1/A

333-224720
333-224720

10.2

10.3

10.4
10.5

6/8/18

6/19/18

6/19/18
6/8/18

154

Form F-1/A

333-224720

10.6

6/8/18

Form 20-F

001-38547

8.1 3/3/20

4.6+

4.7*†
8.1

12.1*

 12.2*

13.1**

15.1*
101.INS*
101.SCH*

101.CAL*

101.DEF*

101.LAB*

101.PRE*

Form of Deed of Indemnity between the
registrant and each of its members of senior
management and directors.
Amendment to License Agreement, dated
as of September 25, 2014 by and between
the registrant and UCL Business Ltd., dated
as of October 15, 2020.
Subsidiaries of the registrant.

Certification of Principal Executive Officer
Pursuant to Rules 13a-14(a) and 15d-14(a)
under the Securities Exchange Act of 1934,
as Adopted Pursuant to Section 302 of the
Sarbanes-Oxley Act of 2002.
Certification of Principal Financial Officer
Pursuant to Rules 13a-14(a) and 15d-14(a)
under the Securities Exchange Act of 1934,
as Adopted Pursuant to Section 302 of the
Sarbanes-Oxley Act of 2002.
Certification of Principal Executive Officer
and Principal Financial Officer Pursuant to
18 U.S.C. Section 1350, as Adopted
Pursuant to Section 906 of the Sarbanes-
Oxley Act of 2002.
Consent of Ernst & Young LLP.
XBRL Instance Document
XBRL Taxonomy Extension Schema
Document
XBRL Taxonomy Extension Calculation
Linkbase Document
XBRL Taxonomy Extension Definition
Linkbase Document
XBRL Taxonomy Extension Label
Linkbase Document
XBRL Taxonomy Extension Presentation
Linkbase Document

+    Indicates management contract or compensatory plan.
#    Confidential treatment has been granted as to portions of the exhibit (indicated by asterisks). Confidential materials omitted and filed separately with

the Securities and Exchange Commission.

*    Filed herewith.
**    Furnished herewith.
†    Certain portions of this exhibit (indicated by asterisks) have been omitted because they are not material and would likely cause competitive harm to

Autolus Therapeutics plc if publicly disclosed.

155

The registrant hereby certifies that it meets all of the requirements for filing on Form 20-F and that it has duly caused and authorized the undersigned to
sign this annual report on its behalf.

SIGNATURES

Date: March 4, 2021

AUTOLUS THERAPEUTICS PLC

By:

/s/ Christian Itin
Christian Itin 
Chief Executive Officer

156

 
INDEX TO CONSOLIDATED FINANCIAL STATEMENTS

Report of Independent Registered Public Accounting Firm
Consolidated Balance Sheets as of December 31, 2020 and 2019
Consolidated Statements of Operations and Comprehensive Loss for the years ended December 31, 2020 and 2019, the three months ended
December 31, 2018, and the year ended September 30, 2018
Consolidated Statements of Shareholders’ Equity for the years ended December 31, 2020 and 2019, the three months ended December 31,
2018, and the year ended September 30, 2018
Consolidated Statements of Cash Flows for the years ended December 31, 2020 and 2019, the three months ended December 31, 2018, and the
year ended September 30, 2018
Notes to Consolidated Financial Statements

F-2
F-3

F-4

F-5

F-6

F-7

F-1

Report of Independent Registered Public Accounting Firm

To the Shareholders and the Board of Directors of Autolus Therapeutics plc

Opinion on the Financial Statements

We have audited the accompanying consolidated balance sheets of Autolus Therapeutics plc (the Company) as of December 31, 2020 and 2019, the related
consolidated statements of operations and comprehensive loss, shareholders' equity and cash flows for the years ended December 31, 2020 and 2019, three-
month period ended December 31, 2018, and the year ended September 30, 2018, and the related notes (collectively referred to as the “consolidated
financial statements”). In our opinion, the consolidated financial statements present fairly, in all material respects, the financial position of the Company at
December 31, 2020 and 2019, and the results of its operations and its cash flows for the years ended December 31, 2020 and 2019, the three-month period
ended December 31, 2018 and the year ended September 30, 2018, in conformity with U.S. generally accepted accounting principles.

Adoption of ASU No. 2016-02

As discussed in Note 2 to the consolidated financial statements, the Company changed its method of accounting for leases in 2019 due to the adoption of
Accounting Standard Update (ASU) No. 2016-02, Leases (Topic 842) and the related amendments.

Basis for Opinion

These financial statements are the responsibility of the Company's management. Our responsibility is to express an opinion on the Company’s financial
statements based on our audits. We are a public accounting firm registered with the Public Company Accounting Oversight Board (United States)
(PCAOB) and are required to be independent with respect to the Company in accordance with the U.S. federal securities laws and the applicable rules and
regulations of the Securities and Exchange Commission and the PCAOB.

We conducted our audits in accordance with the standards of the PCAOB. Those standards require that we plan and perform the audit to obtain reasonable
assurance about whether the financial statements are free of material misstatement, whether due to error or fraud. The Company is not required to have, nor
were we engaged to perform, an audit of its internal control over financial reporting. As part of our audits we are required to obtain an understanding of
internal control over financial reporting but not for the purpose of expressing an opinion on the effectiveness of the Company's internal control over
financial reporting. Accordingly, we express no such opinion.

Our audits included performing procedures to assess the risks of material misstatement of the financial statements, whether due to error or fraud, and
performing procedures that respond to those risks. Such procedures included examining, on a test basis, evidence regarding the amounts and disclosures in
the financial statements. Our audits also included evaluating the accounting principles used and significant estimates made by management, as well as
evaluating the overall presentation of the financial statements. We believe that our audits provide a reasonable basis for our opinion.

/s/ Ernst & Young LLP

We have served as the Company’s auditor since 2017.

Reading, United Kingdom

March 4, 2021

F-2

AUTOLUS THERAPEUTICS PLC

Consolidated Balance Sheets
(In thousands, except share and per share amounts)

Assets
Current assets:
Cash
Restricted cash
Prepaid expenses and other current assets
Total current assets

Non-current assets:
Property and equipment, net
Prepaid expenses and other non-current assets
Right of use asset, net
Long-term deposits
Deferred tax asset
Intangible assets, net

Total assets
Liabilities and shareholders' equity
Current liabilities:

Accounts payable
Accrued expenses and other liabilities
Lease liability
Total current liabilities

Non-current liabilities:
Lease liability
Total liabilities

Shareholders' equity:
Ordinary shares, $0.000042 par value; 200,000,000 shares authorized at December 31, 2020 and 2019,
52,346,231 and 44,983,006 shares issued and outstanding at December 31, 2020 and 2019
Deferred shares,£0.00001 par value; 34,425 shares authorized, issued and outstanding at December 31, 2020
and 2019
Deferred B shares, £0.00099 par value; 88,893,548 shares authorized, issued and outstanding at December 31,
2020 and 2019
Deferred C shares, £0.000008 par value; 1 share authorized, issued and outstanding at December 31, 2020
and 2019
Additional paid-in capital
Accumulated other comprehensive loss
Accumulated deficit
Total shareholders' equity

Total liabilities and shareholders' equity

December 31,

2020

2019

153,299  $
786 
42,899 
196,984 

38,046 
3,033 
51,637 
2,625 
1,754 
158 
294,237  $

2,263 
27,781 
3,590 
33,634 

50,571 
84,205 

3 

— 

118 

— 
595,016 
(5,861)
(379,244)
210,032 
294,237  $

210,643 
787 
37,826 
249,256 

28,164 
— 
23,409 
2,040 
410 
254 
303,533 

1,075 
21,398 
2,511 
24,984 

23,710 
48,694 

2 

— 

118 

— 
500,560 
(8,691)
(237,150)
254,839 
303,533 

$

$

$

The accompanying notes are an integral part of these consolidated financial statements.

F-3

AUTOLUS THERAPEUTICS PLC

Consolidated Statements of Operations and Comprehensive Loss
(In thousands, except share and per share amounts)

For the Year Ended
December 31,

2020

2019

For the
three-
months
ended
December
31,
2018

For the
Year
Ended
September
30,

2018

$

1,473  $
242 

2,908  $
— 

296  $
— 

1,407 
— 

Grant income
License revenue
Operating expenses:

Research and development
General and administrative

(134,888)
(34,972)

(105,418)
(39,452)

       Loss on impairment of
leasehold improvements
Total operating expenses, net
Other income (expense):
Interest income
Other income (expense)

Total other income, net
Net loss before income tax
Income tax benefit
Net loss attributable to ordinary
shareholders
Other comprehensive (loss)
income:
Foreign currency exchange
translation adjustment

Total comprehensive loss

— 
(168,145)

(4,102)
(146,064)

536 
1,352 
1,888 
(166,257)
24,163 

2,542 
4,514 
7,056 
(139,008)
15,159 

(17,713)
(7,593)

— 
(25,010)

660 
1,097 
1,757 
(23,253)
2,605 

(36,150)
(22,790)

— 
(57,533)

1,532 
3,970 
5,502 
(52,031)
7,280 

(142,094)

(123,849)

(20,648)

(44,751)

2,830 
(139,264)

6,797 
(117,052)

(5,568)
(26,216)

(6,071)
(50,822)

Basic and diluted net loss per
ordinary share
Weighted-average basic and diluted
ordinary shares

$

(2.76) $

(2.88) $

(0.52) $

(1.42)

51,558,075  43,065,542 

39,366,634 31,557,034 

The accompanying notes are an integral part of these consolidated financial statements.

F-4

AUTOLUS THERAPEUTICS PLC

Consolidated Statements of Shareholders’ Equity
(In thousands, except share amounts)

Ordinary shares

Deferred Shares

Deferred B shares

Deferred C Shares

Shares

29,962,742  $

Amount
1 

Shares Amount
—  $ — 

Shares

Amount
—  $ — 

Shares
— 

Balance at September 30, 2017
Issuance of ordinary shares,
net of issuance costs
Issuance of deferred shares
Share-based compensation
expense
Unrealized loss on foreign
currency translation
Net loss

Balance at September 30, 2018
Share-based compensation
expense
Restricted shares - forfeited
Unrealized gain on foreign
currency translation
Net loss

Balance at December 31, 2018
Issuance of ordinary shares,
net of issuance costs
Share-based compensation
expense
Restricted shares - forfeited
Exercise of stock options
Unrealized gain on foreign
currency translation
Net loss

Balance at December 31, 2019
Issuance of ordinary shares,
net of issuance costs
Share-based compensation
expense
Restricted shares - forfeited
Exercise of stock options
Unrealized gain on foreign
currency translation
Net loss

10,183,440 

— 

— 
— 

40,146,182  $

— 
(565)

— 
— 

40,145,617  $

4,830,000 

— 
(4,335)
11,724 

— 
— 

44,983,006  $

7,250,000 

— 
(1,969)
115,194 

— 

Balance at December 31, 2020

52,346,231  $

1 
— 

— 

— 
— 
2 

— 
— 

— 
— 
2 

— 

— 
— 
— 

— 
— 
2 

1 

— 
— 
— 

— 
— 
3 

— 
34,425 

— 

— 
— 

— 

— 
— 

— 
— 
34,425  $ — 

— 
— 

— 
— 

— 
— 

— 
— 
34,425  $ — 

— 

— 
— 
— 

— 
— 
34,425 

— 

— 
— 
— 

— 
— 
34,425 

— 

— 
— 
— 

— 
— 
— 

— 

— 
— 
— 

— 
— 
— 

— 
88,893,548 

118 

— 

— 
— 

88,893,548  $

118 

— 
— 

— 
— 

88,893,548  $

— 

— 
— 
— 

— 
— 

88,893,548  $

— 

— 
— 
— 

— 
— 

88,893,548  $

— 
— 

— 
— 
118 

— 

— 
— 
— 

— 
— 
118  $

— 

— 
— 
— 

— 
— 
118 

— 
1 

— 

— 
— 
1 

— 
— 

— 
— 
1 

— 

— 
— 
— 

— 
— 
1 

— 

— 
— 
— 

— 
— 
1 

Additional
Paid in
Capital
—  $ 194,351  $

Amount
$

Accumulated
other
comprehensive
loss

Accumulated
deficit

(3,849) $

(47,902) $

— 
— 

— 

156,802 
— 

6,765 

— 

— 

— 
— 

— 

Total
142,601 

156,803 
118 

6,765 

— 
— 
—  $ 357,918  $

— 
— 

(6,071)
— 
(9,920) $

— 
(44,751) $
(92,653) $

(6,071)
(44,751)
255,465 

— 
— 

3,393 
— 

— 
— 

— 
— 

3,393 
— 

— 
— 
—  $ 361,311  $

— 
— 

(5,568)
— 
(15,488) $

— 
(20,648) $
(113,301) $

(5,568)
(20,648)
232,642 

$

$

— 

— 
— 
— 

108,815 

30,386 
— 
48 

— 

— 
— 
— 

— 

— 
— 
— 

108,815 

30,386 
— 
48 

— 
— 
—  $ 500,560  $

— 
— 

6,797 
— 
(8,691) $

— 
(123,849)
(237,150) $

6,797 
(123,849)
254,839 

— 

— 
— 
— 

73,952 

20,021 
— 
483 

— 

— 
— 
— 

— 

— 
— 
— 

73,953 

20,021 
— 
483 

— 
— 
—  $ 595,016  $

— 
— 

2,830 
— 
(5,861) $

— 
(142,094)
(379,244) $

2,830 
(142,094)
210,032 

The accompanying notes are an integral part of these consolidated financial statements.

F-5

AUTOLUS THERAPEUTICS PLC

Consolidated Statements of Cash Flows
(In thousands)

Cash flows from operating activities:

Net loss

Adjustments to reconcile net loss to net cash used in operating activities:

Depreciation
Loss on disposal of fixed assets and intangible assets
Non-cash share-based compensation (net of amount capitalized)
Gain on lease incentive and reassessment
Loss on impairment of leasehold improvements
Gain on termination of operating lease
Deferred income tax

Changes in operating assets and liabilities
Prepaid expenses and other current assets

Prepaid expenses and other non-current assets
Long-term deposits
Accounts payable
Right of use assets, net
Accrued expenses and other liabilities
 Lease liabilities

Net cash used in operating activities
Cash flows from investing activities:
Purchases of property and equipment
Purchase of intangible assets

Net cash used in investing activities
Cash flows from financing activities:

Proceeds of issuance of ordinary shares, net of issuance costs

Net cash provided by financing activities
Effect of exchange rate changes on cash

Net (decrease) increase in cash

Cash and restricted cash, beginning of period
Cash and restricted cash, end of period

Supplemental cash flow information

Cash paid for taxes

Supplemental non-cash flow information

Property and equipment purchases included in accounts payable or accrued
Leased assets terminated and obtained in exchange for operating lease liabilities, net
Reduction in right of use asset
Leased assets obtained in exchange for operating lease liabilities
Capitalized share-based compensation, net of forfeitures
Capitalized implementation costs included in accrued expenses

Reconciliation of cash and restricted cash reported within the consolidated balance sheets

Cash
Short-term restricted cash
Total cash and restricted cash

Year Ended December 31,
2019
2020

Three Months
Ended December
31,
2018

Year Ended
September 30,
2018

$

(142,094) $

(123,849)

$

(20,648)

$

(44,751)

5,658 
— 
20,107 
(1,335)
— 
(160)
(1,344)

(2,660)

(2,637)
(508)
1,439 
3,670 
5,608 
(3,502)
(117,758)

(14,681)
— 
(14,681)

74,415 
74,415 
679 
(57,345)

211,430 
154,085 

4,611 
43 
30,212 
— 
4,102 
— 
(399)

(21,008)
— 
(702)
(1,451)
2,551 
4,958 
(552)
(101,484)

(18,341)
(327)
(18,668)

108,863 
108,863 
5,164 
(6,125)

217,555 
211,430 

645 
394 
3,393 
— 
— 
— 
— 

(3,521)
— 
(1,285)
(723)
— 
1,960 
— 
(19,785)

(4,422)
— 
(4,422)

— 
— 
(5,327)
(29,534)

247,089 
217,555 

1,841 

1,535 

— 

2,499 
2,487 
— 
26,956 
(86)

144 

153,299 
786 
154,085 

2,818 
— 
1,919 
— 
174 

— 

210,643 
787 
211,430 

3,343 
— 
— 
— 
— 

— 

217,450 
105 
217,555 

1,709 
8 
6,765 
— 
— 
— 
— 

(7,132)
— 
— 
838 
— 
11,026 
— 
(31,537)

(9,119)
(412)
(9,531)

156,920 
156,920 
(5,833)
110,019 

137,070 
247,089 

— 

328 
— 
— 
— 
— 

— 

246,984 
105 
247,089 

The accompanying notes are an integral part of these consolidated financial statements.

F-6

AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements

Note 1. Nature of the Business

    Autolus Therapeutics plc (the “Company”) is a biopharmaceutical company developing next-generation programmed T cell therapies for the treatment of
cancer. Using its broad suite of proprietary and modular T cell programming technologies, the Company is engineering precisely targeted, controlled and
highly active T cell therapies that are designed to better recognize cancer cells, break down their defense mechanisms and attack and kill these cells. The
Company  believes  its  programmed  T  cell  therapies  have  the  potential  to  be  best-in-class  and  offer  cancer  patients  substantial  benefits  over  the  existing
standard of care, including the potential for cure in some patients.

    The Company is a public limited company incorporated in England and Wales. On June 22, 2018, the Company completed its initial public offering
("IPO")  of  American  Depositary  Shares  (“ADSs”).  In  the  IPO,  the  Company  sold  an  aggregate  of  10,147,059 ADSs  representing  the  same  number  of
ordinary shares, including 1,323,529 ADSs pursuant to the underwriters’ option to purchase additional ADSs, at a public offering price of $17.00 per ADS.
Net proceeds were $156.5 million, after deducting underwriting discounts and commissions and offering expenses paid by the Company.

        On  April  15,  2019,  the  Company  completed  an  underwritten  public  offering  of  4,830,000 ADSs  representing  4,830,000  ordinary  shares,  at  a  public
offering  price  of  $24.00  per  ADS,  which  includes  an  additional  630,000 ADSs  issued  upon  the  exercise  in  full  of  the  underwriters’  option  to  purchase
additional ADSs.  Aggregate net proceeds to the Company, after underwriting discounts and offering expenses, were $108.8 million.

    On January 27, 2020, the Company completed an underwritten public offering of 7,250,000 ADSs representing 7,250,000 ordinary shares, at a public
offering price of $11.00 per ADS.  Aggregate net proceeds to the Company, after underwriting discounts, were $75.0 million.

The  Company  is  a  continuation  of  Autolus  Limited  and  its  subsidiaries.  In  connection  with  the  IPO,  the  Company  completed  a  corporate
reorganization,  which  has  been  accounted  for  as  a  combination  of  entities  under  common  control.  The  corporate  reorganization  has  been  given
retrospective effect in these financial statements and such financial statements represent the financial statements of Autolus Therapeutics plc. In connection
with the corporate reorganization, outstanding restricted share awards and option grants of Autolus Limited were exchanged for share awards and option
grants of Autolus Therapeutics plc with identical restrictions.

        The  Company  is  subject  to  risks  and  uncertainties  common  to  early-stage  companies  in  the  biotechnology  industry,  including,  but  not  limited  to,
development  by  competitors  of  new  technological  innovations,  dependence  on  key  personnel,  protection  of  proprietary  technology,  compliance  with
government  regulations  and  the  ability  to  secure  additional  capital  to  fund  operations.  Product  candidates  currently  under  development  will  require
significant additional research and development efforts, including preclinical and clinical testing and regulatory approval, prior to commercialization. These
efforts require significant amounts of capital, adequate personnel and infrastructure and extensive compliance-reporting capabilities. Even if the Company’s
product development efforts are successful, it is uncertain when, if ever, the Company will realize revenue from its product sales.
    The Company has funded its operations primarily with proceeds from the sale of its equity securities. The Company has incurred recurring losses since
its inception, including net losses of $142.1 million and $123.8 million for the years ended December 31, 2020 and 2019, $20.6 million for the three
months ended December 31, 2018, and $44.8 million for the year ended September 30, 2018, respectively. In addition, as of December 31, 2020 and 2019,
the Company had an accumulated deficit of $379.2 million and $237.2 million, respectively. The Company expects to continue to generate operating losses
for the foreseeable future. The future viability of the Company beyond that point is dependent on its ability to raise additional capital to finance its
operations. The Company’s inability to raise additional capital as and when needed could have a negative impact on its financial condition and ability to
pursue its business strategies. There can be no assurances, however, that the current operating plan will be achieved or that additional funding will be
available on terms acceptable to the Company, or at all. The Company believes the cash on hand at December 31, 2020 of $153.3 million, with the
additional net proceeds from sale of ADSs under the Company’s at-the market facility program in January 2021 and its follow-on capital raise in February
2021 of $123.4 million, will be sufficient to fund the Company’s operations for at least 12 months from the issuance date of these financial statements.

F-7

AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

Note 2. Summary of Significant Accounting Policies

Basis of Presentation

    The accompanying consolidated financial statements include those of the Company, Autolus Limited, and its U.S. subsidiary, Autolus Inc., and have
been prepared in conformity with accounting principles generally accepted in the United States of America (“U.S. GAAP”). All intercompany accounts and
transactions have been eliminated upon consolidation.

Use of Estimates

    The preparation of consolidated financial statements in conformity with U.S. GAAP requires management to make estimates and assumptions that affect
the reported amounts of assets and liabilities, the disclosure of contingent assets and liabilities at the date of the consolidated financial statements and the
reported  amounts  of  income  and  expenses  during  the  reporting  periods.  Significant  estimates  and  assumptions  reflected  in  these  consolidated  financial
statements include, but are not limited to, the accrual for research and development expenses, the fair value of ordinary shares, share-based compensation
and income taxes. Estimates are periodically reviewed in light of changes in circumstances, facts and experience. Changes in estimates are recorded in the
period in which they become known. Actual results could differ materially from those estimates.

Cash and Cash Equivalents

    The Company considers cash and cash equivalents in the consolidated financial statements to include cash at banks with a maturity of less than three
months, which is subject to an insignificant risk of changes in value.

Restricted Cash

    The Company entered into a lease that requires a letter of credit supported by $0.6 million deposit held by the Company's bank for the duration of the
lease and a credit card arrangement that requires a security deposit of $0.2 million. The Company includes the restricted cash balance in cash and cash
equivalents when reconciling beginning-of-period and end-of-period total amounts shown on the Company's consolidated statements of cash flows.

Fair Value Measurements

The carrying amounts reported in the balance sheets for cash, prepaid expenses and other assets, accounts payable and accrued expenses and other

liabilities approximate their fair value because of the short-term nature of these instruments.

Concentration of Credit Risk

        Financial  instruments  that  subject  the  Company  to  credit  risk  consist  primarily  of  cash  and  cash  equivalents.  The  Company  places  cash  and  cash
equivalents  in  established  financial  institutions.  The  Company  has  no  significant  off-balance-sheet  risk  or  concentration  of  credit  risk,  such  as  foreign
exchange contracts, options contracts, or other foreign hedging arrangements.

Implementation Costs in a Cloud Computing Arrangement

The Company’s cloud computing arrangements primarily comprise hosting arrangements which are service contracts, whereby the Company gains
remote  access  to  use  enterprise  software  hosted  by  the  vendor  or  another  third  party  on  an  as-needed  basis  for  a  period  of  time  in  exchange  for  a
subscription fee. Implementation costs for cloud computing arrangements are capitalized if certain criteria are met and consist of internal and external costs
directly attributable to developing and configuring cloud computing software for its intended use. These capitalized implementation costs are presented in
the condensed consolidated balance sheet in prepaid expenses and other assets, current and non-current, and are generally amortized over the fixed, non-
cancellable term of the associated hosting arrangement on a straight-line basis.

Property and Equipment

Property  and  equipment  are  recorded  at  cost  and  depreciated  or  amortized  using  the  straight-line  method  over  the  estimated  useful  lives  of  the
respective assets. As of December 31, 2020 and 2019, the Company’s property and equipment consisted of office equipment, lab equipment, furniture and
fixtures, and leasehold improvements. The office equipment has an estimated useful life of three years, lab equipment has an estimated useful life of five or
ten years, and furniture and fixtures have an estimated useful life of five years. Leasehold improvements are depreciated over the shorter of the lease term
or the estimated useful life of the asset. Assets under construction consist of costs incurred with leasehold improvements and, once placed into service, will
be depreciated over the

F-8

AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

shorter  of  the  lease  term  or  the  estimated  useful  life  of  the  asset.  Upon  retirement  or  sale,  the  cost  of  assets  disposed  of,  and  the  related  accumulated
depreciation, are removed from the accounts and any resulting gain or loss is included in the statement of operations and other comprehensive loss. Repairs
and  maintenance  expenditures,  which  are  not  considered  improvements  and  do  not  extend  the  useful  life  of  property  and  equipment,  are  expensed  as
incurred.

The Company evaluates an asset for potential impairment when events or changes in circumstances indicate the carrying value of the asset may not
be recoverable. Recoverability is measured by comparing the book value of the asset to the expected future net undiscounted cash flows that the asset is
expected to generate. If such asset is considered to be impaired, the impairment to be recognized is measured by the amount by which the book value of the
asset exceeds the fair value. The Company did not recognize an impairment nor a disposal for the year ended December 31, 2020, but did recognize a loss
on impairment of $4.1 million for the year ended December 31, 2019 related to leasehold improvements, as the Company chose to discontinue the fit-out of
the  Company's  manufacturing  capacity  at  the  Enfield,  U.K.  facility.  There  were  no  impairments  or  disposals  for  the  three  months  ended  December  31,
2018, and an asset disposal of less than $10,000 for the year ended September 30, 2018.
    The Company routinely evaluates the useful life attributed to its assets. During the second quarter ended June 30, 2019, the Company determined that the
useful lives of certain lab equipment should be increased from five-years to ten-years based on expectation of future usability. The Company accounted for
this as a change in estimate that was applied prospectively, effective April 1, 2019. This change in useful life resulted in a reduction of depreciation expense
of $0.3 million, and an increase in basic and diluted earnings per share of $0.01, for the year ended December 31, 2019.

Leases

        Effective  January  1,  2019,  the  Company  adopted  Accounting  Standards  Codification  (“ASC”),  Topic  842,  Leases  (“ASC  842”),  using  the  required
modified retrospective approach and utilizing the effective date as its date of initial application, for which prior periods are presented in accordance with
the previous guidance in ASC 840, Leases.

    At the inception of an arrangement, the Company determines whether the arrangement is or contains a lease based on the unique facts and circumstances
present. Most leases with a term greater than one year are recognized on the balance sheet as right-of-use assets, lease liabilities and, if applicable, long-
term lease liabilities. The Company has elected not to recognize on the balance sheet leases with terms of one year or less. Operating lease liabilities and
their corresponding right-of-use assets are recorded based on the present value of lease payments over the expected remaining lease term. However, certain
adjustments to the right-of-use asset may be required for items such as incentives received, initial direct costs, or prepayments. The interest rate implicit in
lease  contracts  is  typically  not  readily  determinable.  As  a  result,  the  Company  utilizes  its  incremental  borrowing  rates,  which  are  the  rates  incurred  to
borrow on a collateralized basis over a similar term an amount equal to the lease payments in a similar economic environment.

    In accordance with the guidance in ASC 842, components of a lease should be split into three categories: lease components (e.g., land, building, etc.),
non-lease components (e.g., common area maintenance, consumables, etc.), and non-components (e.g., property taxes, insurance, etc.). Then the fixed and
in-substance fixed contract consideration (including any related to non-components) must be allocated based on the respective relative fair values to the
lease components and non-lease components.

    Although separation of lease and non-lease components is required, certain practical expedients are available. Entities may elect the practical expedient
to not separate lease and non-lease components. Rather, they would account for each lease component and the related non-lease component together as a
single component. For new and amended leases beginning in 2019, the Company elected the practical expedients to account for the lease and non-lease
components  for  leases  for  classes  of  all  underlying  assets  and  allocate  all  of  the  contract  consideration  to  the  lease  component  only.  The  Company
determined the underlying lease to be the predominant component, and therefore, the entire agreement was accounted for under ASC 842.

Intangible Assets Subject to Amortization

    The Company’s intangible assets have been related to acquired software licenses with finite lives are amortized over their useful lives and reviewed for
impairment  whenever  events  or  changes  in  circumstances  indicate  that  the  carrying  amount  of  the  asset  may  not  be  recoverable.  If  any  indicators  were
present,  the  Company  would  test  for  recoverability  by  comparing  the  carrying  amount  of  the  asset  to  the  net  undiscounted  cash  flows  expected  to  be
generated from the asset. If those net undiscounted cash flows do not exceed the carrying amount (i.e., the asset is not recoverable), the Company would
perform the next step, which is to determine the fair value of the asset and record an impairment loss, if any. The Company evaluates the useful lives for
these  intangible  assets  each  reporting  period  to  determine  whether  events  and  circumstances  warrant  a  revision  in  their  remaining  useful  lives.  The
Company did not recognize an impairment loss in the years ended December 31, 2020 and 2019. The Company recognized an impairment loss of $0.4
million for the three months ended December 31, 2018 related to software which the Company elected to discontinue. The Company did not recognize an
impairment loss in the year ended September 30, 2018.

F-9

AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

Segment Information

        Operating  segments  are  defined  as  components  of  an  enterprise  about  which  separate  discrete  information  is  available  for  evaluation  by  the  chief
operating decision maker in deciding how to allocate resources and assess performance. The Company and the Company’s chief operating decision maker,
the Company’s Chief Executive Officer, view the Company’s operations and manages its business as a single operating segment, which is the business of
developing and commercializing gene therapies. The Company operates in two geographic regions: the United Kingdom and the United States. A majority
of the Company’s assets are held in the United Kingdom.

Deferred Rent and Lease Incentives

    Prior to the adoption of ASC 842, rent expense and lease incentives from operating leases were recognized on a straight-line basis over the lease term. 
The Company has operating leases that include rent escalation payment terms and a rent-free period. Deferred rent represents the difference between actual
operating lease payments and straight-line rent expense over the term of the lease. Upon adoption of ASC 842, the Company no longer records or presents
deferred rent.

Research and Development Costs

    Research and development costs are expensed as incurred. Research and development expenses consist of costs incurred in performing research and
development activities, including salaries, share-based compensation and benefits, depreciation expense, third-party license fees, external costs of outside
vendors engaged to conduct clinical development activities, clinical trials, costs to manufacture clinical trial materials and certain tax credits associated
with research and development activities. The Company recorded the U.K.'s research and development expenditure credit ("RDEC") of $0.1 million and
$0.2 million for the years ended December 31, 2020 and 2019, $37,000 for the three months ended December 31, 2018, and $0.2 million for the year ended
September 30, 2018, respectively, as reductions of research and development expenses within the Company’s statement of operations and comprehensive
loss.

Accrued Research and Development Expenses

 As  part  of  the  process  of  preparing  consolidated  financial  statements,  the  Company  is  required  to  estimate  accruals  for  research  and  development
expenses. This process involves reviewing and identifying services which have been performed by third parties on the Company’s behalf and determining
the value of these services. In addition, the Company makes estimates of costs incurred to date but not yet invoiced, in relation to external clinical research
organizations  and  clinical  site  costs.  The  Company  analyzes  the  progress  of  clinical  trials,  including  levels  of  patient  enrollment;  invoices  received  and
contracted costs, when evaluating the adequacy of the accrued liabilities for research and development. The Company makes judgments and estimates in
determining the accrued balance in any accounting period.

Share-Based Compensation

    The Company recognizes compensation expense for equity awards based on the grant date fair value of the award. The Company recognizes share-based
compensation expense for awards granted to employees that have a graded vesting schedule based on a service condition only on a straight-line basis over
the requisite service period for each separately vesting portion of the award as if the award was, in substance, multiple awards (the “graded-vesting
attribution method”), based on the estimated grant date fair value for each separately vesting tranche. For equity awards with a graded vesting schedule and
a combination of service and performance conditions, the Company recognizes share-based compensation expense using a graded-vesting attribution
method over the requisite service period when the achievement of a performance-based milestone is probable, based on the relative satisfaction of the
performance condition as of the reporting date. The Company accounts for forfeitures as they occur.

    For share-based awards granted to consultants and non-employees, compensation expense is recognized using the graded-vesting attribution method over
the period during which services are rendered by such consultants and non-employees until completed. The measurement date for employee awards is the
date of grant, and share-based compensation costs are recognized as expense over the employees’ requisite service period, which is the vesting period, on
an accelerated basis. In the year ended December 31, 2019 the Company adopted Accounting Standards Update (“ASU”) No. 2018-07, “Compensation —
Stock  Compensation  (Topic  718):  Improvements  to  Non-employee  Share-Based  Payment  Accounting”  (“ASU  No.  2018-07”),  prior  to  which  the
measurement  date  for  non-employee  awards  was  generally  the  date  the  services  were  completed,  resulting  in  financial  reporting  period  adjustments  to
share-based compensation during the vesting terms for changes in the fair value of the awards. After the adoption of

F-10

AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

ASU  No.  2018-07,  the  measurement  date  for  non-employee  awards  is  the  later  of  the  adoption  date  of  ASU  No.  2018-07  or  the  date  of  grant,  without
changes in the fair value of the award.

    The fair value of each share option grant is estimated on the date of grant using the Black-Scholes option pricing model. See Note 8 for the Company’s
assumptions used in connection with option grants made during the periods covered by these consolidated financial statements. Assumptions used in the
option pricing model include the following:

Expected volatility.  We lack company-specific historical and implied volatility information for our ADSs for expected terms greater than 2.5 years.
Therefore, we use a combination of the historical volatility of our ADSs and also the expected share volatility based on the historical volatility of
publicly traded peer companies and expect to continue to do so until such time as we have adequate historical data regarding the volatility of our
own traded security price

Expected term.    The expected term of the Company’s share options has been determined utilizing the “simplified” method for awards that qualify
as “plain-vanilla” options.

Risk-free interest rate.    The risk-free interest rate is determined by reference to the U.S. Treasury yield curve in effect at the time of grant of the
award for time periods that are approximately equal to the expected term of the award.

Expected dividend.    Expected dividend yield of zero is based on the fact that the Company has never paid cash dividends on ordinary shares and
does not expect to pay any cash dividends in the foreseeable future.

Fair value of ordinary shares.    Options granted after the Company’s IPO are issued at the fair market value of the Company’s ADS at the date the
grant is approved by the Board.

Prior to the IPO, the Company calculated the fair value of its ordinary shares in accordance with the guidelines in the American Institute of Certified Public
Accountants’  Accounting  and  Valuation  Guide,  Valuation  of  Privately-Held-Company  Equity  Securities  Issued  as  Compensation.  The  Company’s
valuations of ordinary shares were prepared using a market approach, based on precedent transactions in the shares, to estimate the Company’s total equity
value  using  the  option-pricing  method  (“OPM”),  which  used  a  combination  of  market  approaches  and  an  income  approach  to  estimate  the  Company’s
enterprise value.

    The OPM derives an equity value such that the value indicated is consistent with the investment price, and it provides an allocation of this equity value
to each class of the Company’s securities. The OPM treats the various classes of shares as call options on the total equity value of a company, with exercise
prices based on the value thresholds at which the allocation among the various holders of a company’s securities changes. Under this method, each class of
shares has value only if the funds available for distribution to shareholders exceed the value of the share liquidation preferences of the class or classes of
shares with senior preferences at the time of the liquidity event. Key inputs and assumptions used in the OPM calculation include the following:

Expected volatility.    The Company applied re-levered equity volatility based on the historical unlevered and re-levered equity volatility of publicly
traded peer companies.

Expected dividend.    Expected dividend yield of zero is based on the fact that the Company has never paid cash dividends on ordinary shares and
does not expect to pay any cash dividends in the foreseeable future.

Expected term.    The expected term of the option or the estimated time until a liquidation event.

Risk-free interest rate.    The risk-free interest rate is determined by reference to the U.S. Treasury yield curve for the period commensurate with the
expected of the exit event.

    When considering the fair value of options granted in the period prior to the IPO, the Company considered probability-weighted scenarios based on the
relative likelihoods of completing the IPO and remaining a privately-held company. In the IPO scenarios, the fair value was calculated by dividing the total
estimated equity value by the number of fully diluted ordinary shares outstanding, and then discounting the implied per-share value at a rate intended to
approximate  the  Company's  cost  of  equity  between  share  option  grant  date  and  the  expected  IPO  date.  The  stay-private  scenario  utilized  an  OPM
"Backsolve" calculation to estimate its equity value implied by the purchase price of the series A preference shares in September 2017. In March and May
2018, the Company issued share option grants to employees that applied a 50% and 80% probability weighting of an IPO, respectively, to the fair value of
the underlying ordinary share utilized in the Black-Scholes option pricing model.

Foreign Currency Translation

The Company maintains its financial statements in its functional currency, which is the pounds sterling. Monetary assets and liabilities denominated
in currencies other than the functional currency are translated into the functional currency at rates of exchange prevailing at the balance sheet dates. Non-
monetary assets and liabilities denominated in foreign currencies are translated into the functional currency at the exchange rates prevailing at the date of
the transaction. Exchange gains or losses arising from foreign currency transactions are included in the determination of net income (loss) for the respective
periods. The Company recorded foreign

F-11

  
AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

exchange loss of $0.2 million for the year ended December 31, 2020, foreign exchange gains of $4.6 million, $1.1 million, and $4.0 million for the year
ended December 31, 2019, the three months ended December 31, 2018, and the year ended September 30, 2018, respectively, which are included in other
income in the statements of operations and comprehensive loss.

For financial reporting purposes, the financial statements of the Company have been translated into U.S. dollars. Assets and liabilities have been
translated at the exchange rates at the balance sheet dates, while revenue and expenses are translated at the average exchange rates over the reporting period
and  shareholders’  equity  amounts  are  translated  based  on  historical  exchange  rates  as  of  the  date  of  each  transaction.  Translation  adjustments  are  not
included  in  determining  net  income  (loss)  but  are  included  in  foreign  exchange  adjustment  to  other  comprehensive  loss,  a  component  of  shareholders’
equity.

Patent Costs

The  Company  expenses  patent  prosecution  and  related  legal  costs  as  they  are  incurred  and  classifies  such  costs  as  general  and  administrative
expenses in the accompanying statements of operations and comprehensive loss. The Company recorded patent expenses of $2.1 million and $2.0 million
for  the  years  ended  December  31,  2020  and  2019,  $0.2  million  for  the  three  months  ended  December  31,  2018,  and  $1.0  million  for  the  year  ended
September 30, 2018, respectively.

Grant Income

The Company has received research grants under which it is reimbursed for specific research and development activities. Payments received are
recognized as income in the statements of operations and comprehensive loss over the period in which the Company recognizes the related costs. At the
time the Company recognizes grant income, it has complied with the conditions attached to it and the receipt of the reimbursement is reasonably assured.
The Company has received grants from the U.K. government, which are repayable under certain circumstances, including breach or noncompliance. For
grants  with  refund  provisions,  the  Company  reviews  the  grant  to  determine  the  likelihood  of  repayment.  If  the  likelihood  of  repayment  of  the  grant  is
determined to be remote, then the grant is recognized as grant income. The Company has determined that the likelihood of any repayment events included
in its current grants is remote.

License Revenue

The  Company  accounts  for  its  revenues  pursuant  to  the  provisions  of  Accounting  Standards  Codification  (“ASC”)  Topic  606,  Revenue  from

Contracts with Customers (“ASC Topic 606”).

The Company has no products approved for commercial sale and has not generated any revenue from commercial product sales. The total revenue
to date has been generated from a license agreement with an investee company of one our affiliates. The terms of the agreement include a non-refundable
license fee, payments based upon achievement of clinical development and regulatory objectives, and royalties on product sales.

In determining the appropriate amount of revenue to be recognized as the Company fulfills its obligations under its agreements, the Company

performs the following steps: (i) identification of the promised goods or services in the contract; (ii) determination of whether the promised goods or
services are performance obligations, including whether they are distinct in the context of the contract; (iii) measurement of the transaction price, including
the constraint on variable consideration; (iv) allocation of the transaction price to the performance obligations based on estimated selling prices; and (v)
recognition of revenue when (or as) the Company satisfies each performance obligation.

License Fees and Multiple Element Arrangements

If  a  license  to  the  Company’s  intellectual  property  is  determined  to  be  distinct  from  the  other  performance  obligations  identified  in  the
arrangement, the Company recognizes revenues from non-refundable, upfront fees allocated to the license at such time as the license is transferred to the
licensee and the licensee is able to use, and benefit from, the license. For licenses that are bundled with other promises, the Company utilizes judgment to
assess the nature of the combined performance obligations to determine whether the combined performance obligations are satisfied over time or at a point
in  time  and,  if  over  time,  the  appropriate  method  of  measuring  progress  for  purposes  of  recognizing  revenue  from  non-refundable,  upfront  fees.  The
Company evaluates the measure of progress each reporting period and, if necessary, adjusts the measure of performance and related revenue recognition.

Appropriate  methods  of  measuring  progress  include  output  methods  and  input  methods.  In  determining  the  appropriate  method  for  measuring
progress, the Company considers the nature of service that the Company promises to transfer to the customer. When the Company decides on a method of
measurement, the Company will apply that single method of measuring progress for each performance obligation satisfied over time and will apply that
method consistently to similar performance obligations and in similar circumstances.

F-12

AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

Contingent Research Milestone Payments

ASC Topic 606 constrains the amount of variable consideration included in the transaction price in that either all, or a portion, of an amount of
variable  consideration  should  be  included  in  the  transaction  price.  The  variable  consideration  amount  should  be  included  only  to  the  extent  that  it  is
probable  that  a  significant  reversal  in  the  amount  of  cumulative  revenue  recognized  will  not  occur  when  the  uncertainty  associated  with  the  variable
consideration is subsequently resolved. The assessment of whether variable consideration should be constrained is largely a qualitative one that has two
elements: the likelihood of a change in estimate, and the magnitude thereof. Variable consideration is not constrained if the potential reversal of cumulative
revenue recognized is not significant, for example.

If the consideration in a contract includes a variable amount, the Company will estimate the amount of consideration in exchange for transfer of
promised  goods  or  services.  The  consideration  also  can  vary  if  the  Company’s  entitlement  to  the  consideration  is  contingent  on  the  occurrence  or  non-
occurrence  of  a  future  event.  The  Company  considers  contingent  research  milestone  payments  to  fall  under  the  scope  of  variable  consideration,  which
should be estimated for revenue recognition purposes at the inception of the contract and reassessed ongoing at the end of each reporting period.

The Company assesses whether contingent research milestones should be considered variable consideration that should be constrained and thus
not  part  of  the  transaction  price.  This  includes  an  assessment  of  the  probability  that  all  or  some  of  the  milestone  revenue  could  be  reversed  when  the
uncertainty around whether or not the achievement of each milestone is resolved, and the amount of reversal could be significant.

GAAP provides factors to consider when assessing whether variable consideration should be constrained. All of the factors should be considered,

and no factor is determinate. The Company considers all relevant factors.

For the year ended December 31, 2020, the Company has not recognized any variable consideration with regards to the development milestones
which are included in the license agreement that was executed in the third quarter of the year. This is due to the fact that those development milestones have
not yet been met and the recognition of the related revenue is not yet probable.

Income Taxes

The Company accounts for income taxes under the asset and liability method which includes the recognition of deferred tax assets and liabilities for
the  expected  future  tax  consequences  of  events  that  have  been  included  in  the  Company’s  financial  statements.  Under  this  approach,  deferred  taxes  are
recorded for the future tax consequences expected to occur when the reported amounts of assets and liabilities are recovered or paid. The provision for
income taxes represents income taxes paid or payable for the current year plus deferred taxes. Deferred taxes result from differences between the financial
statements and tax bases of the Company’s assets and liabilities, and are adjusted for changes in tax rates and tax law when changes are enacted. The effects
of future changes in income tax laws or rates are not anticipated.

The Company is subject to income taxes in the United Kingdom and the United States. The calculation of the Company’s tax provision involves the

application of United Kingdom tax law and requires judgement and estimates.

The  Company  evaluates  the  realizability  of  its  deferred  tax  assets  at  each  reporting  date,  and  establishes  a  valuation  allowance  when  it  is  more

likely than not that all or a portion of its deferred tax assets will not be realized.

The ultimate realization of deferred tax assets is dependent upon the generation of future taxable income of the same character and in the same
jurisdiction. The Company considers all available positive and negative evidence in making this assessment, including, but not limited to, the scheduled
reversal of deferred tax liabilities, projected future taxable income, and tax planning strategies. In circumstances where there is sufficient negative evidence
indicating that the Company’s deferred tax assets are not more likely than not realizable, the Company establishes a valuation allowance.

The Company uses a two-step approach for recognizing and measuring uncertain tax positions. The first step is to evaluate tax positions taken or
expected  to  be  taken  in  a  tax  return  by  assessing  whether  they  are  more  likely  than  not  sustainable,  based  solely  on  their  technical  merits,  upon
examination, and including resolution of any related appeals or litigation process. The second step is to measure the associated tax benefit or each position
as the largest amount that the Company believes is more likely than not realizable. Differences between the amount of tax benefits taken or expected to be
taken in the Company’s income tax returns and the amount of tax benefits recognized in the its financial statements represent the Company’s unrecognized
income tax benefits, which it either records as a liability or reduction of deferred tax assets.

F-13

AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

Income Tax Credit

The  Company  benefits  from  the  U.K.  research  and  development  tax  credit  regime  under  both  the  small  and  medium  sized  enterprise,  or  SME,
scheme and by claiming an RDEC in respect of grant funded projects. Under the SME regime, a portion of the Company’s losses can be surrendered for a
cash rebate of up to 33.35% of eligible expenditures. Such credits are accounted for within the tax provision in the year in which the expenditures were
incurred.

Comprehensive Loss

The Company follows the provisions of the Financial Accounting Standards Board (“FASB”) Accounting Standards Codification (“ASC”) Topic
220, Comprehensive Income, which establishes standards for the reporting and display of comprehensive income and its components. Comprehensive gain
or  loss  is  defined  to  include  all  changes  in  equity  during  a  period  except  those  resulting  from  investments  by  owners  and  distributions  to  owners.  The
Company recorded a gain of $2.8 million and $6.8 million for the years ended December 31, 2020 and 2019, a loss of $5.6 million for the three months
ended December 31, 2018, and a loss $6.1 million for the year ended September 30, 2018, respectively to foreign currency translation.

Net Loss per Share

Basic and diluted net loss per ordinary share is determined by dividing net loss by the weighted average number of ordinary shares outstanding
during the period. For all periods presented, the preferred shares and outstanding but unvested restricted shares and share options have been excluded from
the calculation, because their effects would be anti-dilutive. Therefore, the weighted average shares outstanding used to calculate both basic and diluted
loss per share are the same for all periods presented.

The following potentially dilutive securities have been excluded from the calculation of diluted net loss per share due to their anti-dilutive effect:

Unvested restricted shares and units
Incentive share options

Total

Ordinary Share Conversion

Year Ended December 31,

2020

2019

505,383 
5,611,429 
6,116,812 

814,744 
5,963,239 
6,777,983 

Three Months Ended
December 31,
2018

Years Ended
September 30,
2018

708,834 
3,711,274 
4,420,108 

815,632 
2,065,481 
2,881,113 

    On the date of the IPO, the Company converted its outstanding preferred and ordinary shares as discussed in Note 7.  All share and per share information
has been retroactively adjusted to reflect the share conversion.

Emerging Growth Company Status

The Company is an “emerging growth company,” as defined in the Jumpstart Our Business Startups Act (“JOBS Act”) and may take advantage of
certain  exemptions  from  various  reporting  requirements  that  are  applicable  to  other  public  companies  that  are  not  emerging  growth  companies.  The
Company may take advantage of these exemptions until the Company is no longer an emerging growth company. Section 107 of the JOBS Act provides
that an emerging growth company can take advantage of the extended transition period afforded by the JOBS Act for the implementation of new or revised
accounting  standards.  The  Company  has  elected  not  to  use  the  extended  transition  period  for  complying  with  any  new  or  revised  financial  accounting
standards provided pursuant to Section 7(a)(2)(B) of the Securities Act.

These exemptions provided by the JOBS Act will apply up until the last day of the fiscal year following the fifth anniversary of the IPO or such
earlier time that the Company is no longer meets the requirements of being an emerging growth company. The Company would cease to be an emerging
growth company if it has more than $1.07 billion in annual revenue, has more than $700 million in market value of its securities held by non-affiliates (and
it has been a public company for at least 12 months, and has filed one annual report on Form 20-F), or it issues more than $1 billion of non-convertible debt
securities over a three-year period.

F-14

 
AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

JOBS Act

    On April 5, 2012, the Jumpstart Our Business Startups Act, or the JOBS Act, was enacted. The JOBS Act provides that, among other things, an emerging
growth  company  can  take  advantage  of  an  extended  transition  period  for  complying  with  new  or  revised  accounting  standards.  As  an  emerging  growth
company, the Company has irrevocably elected not to take advantage of the extended transition period afforded by the JOBS Act for the implementation of
new or revised accounting standards and, as a result, the Company will comply with new or revised accounting standards on the relevant dates on which
adoption of such standards is required for non-emerging growth public companies.

    In addition, the Company intends to rely on the other exemptions and reduced reporting requirements provided by the JOBS Act. Subject to certain
conditions set forth in the JOBS Act, the Company is entitled to rely on certain exemptions as an “emerging growth company.” As an emerging growth
company, the Company is not required to, among other things, (i) provide an auditor’s attestation report on the Company’s system of internal controls over
financial  reporting  pursuant  to  Section  404(b),  (ii)  provide  all  of  the  compensation  disclosure  that  may  be  required  of  non-emerging  growth  public
companies under the Dodd-Frank Wall Street Reform and Consumer Protection Act, (iii) comply with any requirement that may be adopted by the Public
Company Accounting Oversight Board regarding mandatory audit firm rotation or a supplement to the auditor’s report providing additional information
about the audit and the financial statements (auditor discussion and analysis), and (iv) disclose certain executive compensation-related items such as the
correlation  between  executive  compensation  and  performance  and  comparisons  of  the  chief  executive  officer’s  compensation  to  median  employee
compensation.  These  exemptions  will  apply  for  a  period  of  five  years  following  the  completion  of  the  IPO  or  until  the  Company  no  longer  meets  the
requirements of being an emerging growth company, whichever is earlier.

Recently issued accounting pronouncements adopted

    In August 2018, the FASB issued ASU 2018-15, Customer’s Accounting for Implementation Costs Incurred in a Cloud Computing Arrangement That Is
a  Service  Contract,  a  new  standard  on  a  customer's  accounting  for  implementation,  set-up,  and  other  upfront  costs  incurred  in  a  cloud  computing
arrangement  ("CCA")  that  aligns  the  requirements  for  capitalizing  implementation  costs  in  a  CCA  service  contract  with  existing  internal-use  software
guidance. The standard is effective for interim and annual periods beginning after December 15, 2019, with early adoption permitted, and can be adopted
prospectively  or  retrospectively.  The  Company  adopted  the  new  standard  on  January  1,  2020  on  a  prospective  basis.  The Company’s CCAs are service
contracts for the hosting of software primarily related to recording and tracking information related to its clinical trials, including but not limited to patient
data and clinical manufacturing. The capitalized implementation costs are presented in the condensed consolidated balance sheet in prepaid expenses and
other  assets,  current  and  non-current.  The  deferred  implementation  costs  will be  expensed  over  the  term  of  the  hosting  arrangement,  which  is  the  non-
cancelable term of the arrangement plus any reasonably certain renewal periods. As of December 31, 2020, $1.0 million was recorded in prepaid expenses
and other assets, current and $1.1 million was recorded to prepaid expenses and other assets, non-current as deferred implementation costs. During the year
ended December 31, 2020, $0.3 million of deferred implementation costs were expensed.

Recently issued accounting pronouncements not adopted

Other accounting standards that have been issued by the FASB or other standards-setting bodies that do not require adoption until a future date are

not expected to have a material impact on the Company's financial statements upon adoption.

F-15

AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

Note 3. Prepaid Expenses and Other Current Assets

    Prepaid expenses and other current assets consisted of the following (in thousands):

Research and development claims receivable
Prepayments
VAT receivable
Lease incentive receivable
Other asset
Grant income receivable
Other receivable
Deferred cost

Total prepaid expenses and other current assets

December 31,

2020

2019

$

$

24,711  $
10,206 
3,124 
1,237 
— 
414 
199
3,008 
42,899  $

27,567 
7,023 
1,928 
— 
279 
547 
482
— 
37,826 

    The decrease to research and development claims receivable is due the fact that as of December 31, 2020 the research and development claim receivable
is solely for the year ended December 31, 2020; however, the research and development claims receivable as of December 31, 2019 were related to a claim
receivable for both the 15-month period ended December 31, 2019 and the year ended September 30, 2018.

The increase in prepayments in the year ended December 31, 2020 is primarily related to $1.9 million of prepayments for facilities and manufacturing and
$1.4 million for prepayments of clinical trial costs.

The  increase  in  deferred  cost  in  the  year  ended  December  31,  2020  is  related  to  $1.0  million  in  capitalized  implementation  costs  related  to  hosting
arrangements and $2.0 million of non-refundable upfront costs related to payments for future manufacturing slots related to our clinical programs.

Note 4. Property and Equipment, Net

    Property and equipment, net consisted of the following (in thousands):

Lab equipment
Office equipment
Furniture and fixtures
Leasehold improvements
Assets under construction
Less: accumulated depreciation

Total property and equipment, net

December 31,

2020

2019

$

$

23,491  $
2,928 
1,340 
10,629 
14,321 
(14,663)
38,046  $

18,214 
2,211 
1,301 
10,316 
4,687 
(8,565)
28,164 

    Depreciation expense recorded for the years ended December 31, 2020 and 2019 was $5.7 million and $4.6 million, the three months ended December
31, 2018 was $0.6 million and, for the year ended September 30, 2018 was $1.7 million, respectively. The increase in assets under construction and lab
equipment relates to the build out of manufacturing facilities as we continue scaling our manufacturing operations.

Note 5. Intangible Asset

    The following table summarizes the carrying amount of the Company's intangible assets, net of accumulated amortization (in thousands):

F-16

AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

Software license

Estimated Life 
(years)
3

December 31, 2020

Cost

Accumulated
Amortization

Net

$

291  $

(133)

$

158 

    Amortization expense for the years ended December 31, 2020 and 2019 was $91,000 and $47,000, respectively. Assuming no changes in the gross cost
basis of intangible assets, the total estimated amortization expense for finite-lived intangible assets is approximately $92,000 and $66,000 for each of the
years ending December 31, 2021 through December 31, 2022.

Note 6. Accrued Expenses and Other Liabilities

    Accrued expenses and other liabilities consisted of the following (in thousands):

Compensation and benefits
Research and development costs
UCLB milestone
Professional fees
Corporate tax
Other liabilities

Total accrued expenses and other liabilities

$

December 31,

2020

2019

8,732  $

15,343 
205 
3,005 
— 
496 
27,781  $

6,568 
10,449 
663 
2,611 
391 
716 
21,398 

Other liabilities primarily consisted of the business rates for UK property in the amount of $0.4 million and $0.2 million as of December 31, 2020

and 2019, respectively. As of December 31, 2019, other liabilities also consisted of the current portion of the lease incentive liability of $0.3 million.

Note 7. Shareholders’ Equity

Ordinary Shares

    Each holder of ordinary shares is entitled to one vote per ordinary share and to receive dividends when and if such dividends are recommended by the
board of directors and declared by the shareholders.  As of December 31, 2020, the Company has not declared any dividends.

Effective from June 26, 2018, the board of directors has the authority to allot new ordinary shares or to grant rights to subscribe for or to convert any
security into ordinary shares in the Company up to a maximum aggregate nominal amount of $8,400. This authority runs for five years and will expire on
June  26,  2023.  Effective  from  June  26,  2018,  the  board  also  has  the  authority  to  allot  ordinary  shares  for  cash  or  to  grant  rights  to  subscribe  for  or  to
convert any security into ordinary shares in the Company without first offering them to existing shareholders in proportion to their existing holdings up to
an aggregate maximum nominal amount of $8,400. This authority runs for five years and will expire on June 26, 2023.

    As of December 31, 2020, the Company’s issued capital share consisted of 52,346,231 ordinary shares, with a nominal value of $0.000042 per share, (ii)
34,425 deferred shares, with a nominal value of £0.00001 per share, (iii) 88,893,548 B deferred shares, with a nominal value of £0.00099 per share and (iv)
1 C deferred share, with a nominal value of £0.000008. Each issued share has been fully paid.

Initial Public Offering and Impact of Corporate Reorganization

        On  June  18,  2018,  Autolus  Therapeutics  Limited  re-registered  as  a  public  limited  company  and  its  name  was  changed  from  Autolus  Therapeutics
Limited to Autolus Therapeutics plc (see Note 1).

F-17

    
AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

    On June 26, 2018, the Company closed its IPO.  In the IPO, the Company sold an aggregate of 10,147,059 ADSs representing the same number of
ordinary shares at a public offering price of $17.00 per ADS, which included the full exercise by the underwriters of their option to purchase additional
ADSs.  Net  proceeds  were  approximately  $156.5  million,  after  deducting  underwriting  discounts,  and  commissions  and  offering  expenses  paid  by  the
Company of $16.0 million. Upon the closing of the IPO, each separate class of ordinary shares of Autolus Therapeutics plc was converted into a single
class of ordinary shares of Autolus Therapeutics plc as described further below.

    Prior to the Company’s June 2018 reorganization and IPO, the Company had issued series A preferred shares, ordinary B shares, and ordinary C shares
to fund its operations and upon the completion of the IPO, the different classes of shares were converted into a single class of ordinary shares on a 3.185-
for-1 basis and created various classes of deferred shares.

    The following deferred share classes were created:

        Deferred Shares  -  The  34,425  deferred  shares,  aggregate  nominal  value  less  than  $1.00,  existed  in  Autolus  Limited  and  were  re-created  in  Autolus
Therapeutics  plc  as  part  of  the  share  exchange  to  place  Autolus  Therapeutics  as  the  ultimate  parent  entity.  The  Company  was  required  to  replicate  the
shares  to  ensure  the  existing  share  has  the  correct  nominal  value  to  ensure  stamp  duty  mirroring  relief  is  available  on  the  subsequent  share  for  share
exchange. These deferred shares have no voting rights.

Deferred B Shares - The deferred shares were the product of the reorganization of the series A preferred shares and ordinary B shares into ordinary
shares. The nominal residual value was utilized by management as the required £50,000 of share capital to re-register Autolus Therapeutics Limited as
Autolus Therapeutics plc. The resulting 88,893,548 deferred shares, aggregate nominal value of $118,000, is presented as a separate class of equity on the
balance sheet and statement of shareholder’s equity. These deferred B shares have no voting rights.

Deferred C Share - The deferred share, nominal value less than $1.00, was created when the shares in the Company were redenominated from
pounds sterling to U.S. Dollars as part of the capital reduction to deal with rounding issues that would otherwise have unbalanced the company’s nominal
share capital. This deferred C share has no voting rights.

Open Market Sale Agreement

In September 2020, the Company entered into an Open Market Sale Agreement, or the Sales Agreement, with Jefferies LLC, or Jefferies, under
which the Company may, at its option, offer and sell ADSs having an aggregate offering price of up to $100.0 million from time to time through Jefferies,
acting as sales agent. Any such sales made through Jefferies can be made by any method that is deemed an “at-the-market offering” as defined in Rule 415
promulgated under the Securities Act, or in other transactions pursuant to an effective shelf registration statement on Form F-3. The Company agreed to
pay Jefferies a commission of 3.0% of the gross proceeds of any sales of ADSs sold pursuant to the Sales Agreement. During the year ended December 31,
2020, the Company did not sell any ADSs under the Sales Agreement.

Note 8. Share Based Compensation

        In  February  2017,  the  Company’s  board  of  directors  adopted  the  2017  Share  Option  Plan,  or  the  2017  Plan.  The  2017  Plan  was  set  to  expire  on
February 21, 2027. The 2017 Plan provided for the grant of potentially tax-favored Enterprise Management Incentives, or EMI, options to the Company's
U.K. employees and for the grant of options to its U.S. employees.

    In June 2018, as part of the Company's reorganization and IPO, the Company’s board of directors and shareholders approved the 2018 Equity Incentive
Plan, or the 2018 Plan. The initial maximum number of ordinary shares that may be issued under the 2018 Plan was 3,281,622. This number consists of
3,025,548  new  ordinary  shares  and  256,074  ordinary  shares  that  would  have  otherwise  remained  available  for  future  grants  under  the  2017  Plan.  The
number of ordinary shares reserved for issuance under the 2018 Plan will automatically increase on October 1st of each year, for a period of not more than
ten years, commencing on October 1, 2018 and ending on (and including) October 1, 2027, by an amount equal to the lesser of (i) 4% of the total number
of ordinary shares outstanding on September 30th of the same calendar year or (ii) such fewer number of ordinary shares as the board of directors may
designate prior to the applicable October 1st date. The updated maximum number of ordinary shares that may be issued under the 2018 Plan is 8,778,719 as
of December 31, 2020. The total Shares issued under the 2018 Plan may be authorized but unissued shares, shares purchased on the open market, treasury
shares or ADSs. No more than 14,000,000 shares may be issued under the 2018 Plan upon the exercise of incentive share options.

    Options granted under the 2018 Plan and 2017 Plan, as well as restricted shares granted as employee incentives, typically vest over a four-year service
period with 25% of the award vesting on the first anniversary of the commencement date and the balance

F-18

AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

vesting monthly over the remaining three years, unless the award contains specific performance vesting provisions. For equity awards issued that have both
a  performance  vesting  condition  and  a  services  condition,  once  the  performance  criteria  is  achieved,  the  awards  are  then  subject  to  a  four-year  service
vesting  with  25%  of  the  award  vesting  on  the  first  anniversary  of  the  performance  condition  being  achieved  and  the  balance  vesting  monthly  over  the
remaining three years. Options granted under the 2018 Plan and 2017 Plan generally expire 10 years from the date of grant. For certain senior members of
management and directors, the board of directors has approved an alternative vesting schedule. Restricted stock units awarded in December 2019 vest over
a 3-year service period with 50% of the award vesting one-and-half years from commencement date and the remaining 50% of the award vesting at the end
of the third year. No restricted stock units were awarded in the year ended December 31, 2020.

Share Option Valuation

The assumptions (see Note 2) used in the Black-Scholes option pricing model to determine the fair value of the share options granted to employees

and directors during the years ended December 31, 2020 and 2019, the three months ended December 31, 2018, and the year ended September 30, 2018
were as follows:

Expected option life (years)
Risk-free interest rate
Expected volatility
Expected dividend yield

Year Ended December 31,

2020
5.27 to 6.08
0.31% to 1.66%
76.38% to 81.45%
—%

2019
5.27 to 6.08
1.39% to 2.66%
72.30% to 76.22%
—%

Three Months Ended
December 31,
2018
6.08
2.70% to 3.09%
69.52% to 71.26%
—%

Year Ended
September 30,
2018
6.08
2.61% to 3.00%
68.15% to 72.99%
—%

Share Options

The table below summarizes activity for the years ended December 31, 2020 and 2019.

Outstanding as of December 31, 2018

Granted
Exercised
Forfeited

Outstanding as of December 31, 2019

Granted
Exercised
Forfeited

Outstanding as of December 31, 2020
Exercisable as of December 31, 2020
Vested and expected to vest as of December 31, 2020

Number of 
Options

Weighted- 
Average 
Exercise 
Price

Weighted- 
Average 
Remaining 
Contractual 
Term 
(Years)

Aggregate 
Intrinsic 
Value

3,711,274  $
2,438,145 
(11,724)
(201,456)
5,936,239  $
474,225 
(115,194)
(683,841)
5,611,429 
2,664,362  $
5,611,429  $

19.25 
15.22 
3.94 
19.72 
17.71 
11.61
4.07
20.06
17.19
17.36 
17.19 

9.47 $
— 
—  $
— 
9.02 $
— 
— 
— 
7.96 $
7.36 $
7.96 $

51,464 
— 
198 
— 
11,873 
— 
— 
— 
4,262 
3,432 
4,262 

The aggregate intrinsic value of share options is calculated as the difference between the exercise price of the share options and the fair value of the

Company’s restricted ordinary shares for those share options that had exercise prices lower than the fair value of the Company’s restricted ordinary shares.

F-19

AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

The  weighted  average  grant-date  fair  value  of  share  options  granted  was  $7.82,  $10.07,  $20.04,  and  $8.55  per  share  for  the  years  ended

December 31, 2020 and 2019, for the three months ended December 31, 2018, and for the year ended September 30, 2018, respectively.

    As of December 31, 2020, the total unrecognized compensation expense related to unvested options was $13.2 million, which the Company expects to
recognize over a weighted average vesting period of 2.57 years.

Restricted Ordinary Shares

The assumptions (Note 2) used in the OPM to determine the fair value of the ordinary shares for the following dates are as follows:

Expected term
Risk-free interest rate
Expected volatility
Expected dividend yield

March 2, 
2016

2.8 years
1.0 %
73.2 %
0.0 %

April 26, 
2017
1.2 years
1.0 %
76.6 %
0.0 %

September 25, 
2017

0.8 years
1.3 %
71.0 %
0.0 %

March 31,
2018
1.8 years
2.1 %
71 %
0.0 %

May 31, 2018
1.8 years
2.1 %
71 %
0.0 %

    A summary of the changes in the Company’s restricted ordinary shares during the years ended December 31, 2020 and 2019 are as follows and reflect
the conversion of ordinary shares in the current and previous years.

Unvested and outstanding at December 31, 2018

Granted
Vested
Forfeited

Unvested and outstanding at December 31, 2019

Granted
Vested
Forfeited

Unvested and outstanding at December 31, 2020

Number of 
restricted 
shares

Weighted average 
grant date 
fair value

708,834  $

0 
(389,755)
(4,335)
314,744 
— 
(222,392)
(1,969)
90,383 

4.10 
0.00 
4.31 
4.43 
4.22 
0
4.24
4.44
4.17

As of December 31, 2020, there was unrecognized compensation of $0.04 million, which will be recognized over 0.59 years.

Restricted Stock Units

    A restricted stock unit (“RSU”) represents the right to receive one of the Company’s ADSs upon vesting of the RSU. The fair value of each RSU is
based on the closing price of the Company’s ADSs on the date of grant. The Company grants RSUs with service conditions that vest over 3-year service
period with 50% of the award vesting one-and-half years from commencement date and the remaining 50% of the award vesting at the end of the third year.

F-20

    
AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

During the year ended December 31, 2020, the Company did not grant RSUs under the 2018 Plan. The following is a summary of RSU activity for the
2018 Plan for the years ended December 31, 2020 and 2019, respectively:

Unvested and outstanding at December 31, 2018

Granted
Vested
Forfeited

Unvested and outstanding at December 31, 2019

Granted
Vested
Forfeited

Unvested and outstanding at December 31, 2020

Number of 
restricted 
units

Weighted average 
grant date 
fair value

— 
500,000 
— 
— 
500,000 
— 
— 
(85,000)
415,000 

$

$

— 
12.09
— 
— 
12.09
— 
— 
12.09 
12.09 

    As of December 31, 2020, there was $2.4 million of unrecognized compensation costs related to unvested RSUs, which are expected to be recognized
over a weighted average period of 1.96 years.

Share-based Compensation Expense

    Share-based compensation expense recorded as research and development and general and administrative expenses is as follows (in thousands):

Year Ended December 31,

2020

2019

Three Months December 31,
2018

Years Ended September
30,
2018

Research and development
General and administrative
Capitalized to fixed assets

Total share-based compensation

$
$
$
$

12,992 
7,115 
(86)
20,021 

$

$

17,761  $
12,451 
174 
30,386  $

1,906  $
1,487 
— 
3,393  $

3,116 
3,649 
— 
6,765 

Note 9. License Agreements

University College of London Business Ltd. (UCLB) License

In September 2014, the Company entered into an exclusive license agreement (the “License”) with UCL Business Ltd. (“UCLB”), the technology
transfer company of University College London (“UCL”), to obtain licenses to certain technology rights in the field of cancer therapy and diagnosis. In
March 2016, the License was amended to include additional rights.

As  part  of  the  consideration  for  the  License  in  September  2014,  the  Company  issued  1,497,643  ordinary  shares  to  UCLB.  The  Company  paid

upfront fees of $0.3 million and issued an additional 313,971 ordinary shares to UCLB when the License was amended in March 2016.

In March 2018, the License was further amended and restated to include a license to the Company's product candidate, AUTO1, for which UCL is
conducting Phase 1 clinical trials of AUTO1 in pediatric and adult ALL patients. The Company paid an upfront fee of £1.5 million for consideration for the
amended and restated License and paid the additional £0.35 million in connection with UCLB's transfer of clinical data to the Company in December 2020.
No equity was issued as part of the upfront fee consideration.

F-21

    
 
AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

Additionally, the Company may be obligated to make payments to UCLB under the amended and restated License upon the initiation of certain
clinical activities in an aggregate amount of £0.18 million, the receipt of specified regulatory approvals in an aggregate amount of £37.5 million, the start of
commercialization in an aggregate amount of £18 million, and the achievement of net sales levels in an aggregate amount of £51 million, as well as royalty
payments based on possible future sales resulting from the utilization of the licensed technologies. On a per-product basis, these milestone payments range
from £1 million to £18.5 million, depending on which T cell programming modules are used in the product achieving the milestone.

Upon commercialization of any of the Company’s products that use the in-licensed patent rights, the Company will be obligated to pay UCLB a
flat royalty for each licensed product ranging from the low- to mid-single digits, depending on which technologies are deployed in the licensed product,
based on worldwide annual net sales of each licensed product, subject to certain reductions, including for the market entry of competing products and for
loss of patent coverage of licensed products. The Company may deduct from the royalties payable to UCLB one-half of any payments made to a third party
to obtain a license to such third party’s intellectual property that is necessary to exploit any licensed products. Once net sales of a licensed product have
reached  a  certain  specified  threshold,  the  Company  may  exercise  an  option  to  buy  out  UCLB’s  rights  to  the  remaining  milestone  payments,  royalty
payments, and sublicensing revenue payments for such licensed product, on terms to be negotiated at the time.

The License expires on a product-by-product and country-by-country basis upon the expiration of the royalty term with respect to each product in
each country. The Company may unilaterally terminate the license agreement for any reason upon advance notice to UCLB. Either party may terminate the
License  for  the  uncured  material  breach  by  the  other  party  or  for  the  insolvency  of  the  other  party.  If  UCLB  terminates  the  License  following  the
Company’s  insolvency  or  the  Company’s  material  breach  of  the  License,  or  if  the  Company  terminates  the  License  unilaterally,  all  rights  and  licenses
granted to the Company will terminate, and all patent rights and know-how transferred to the Company pursuant to the License will revert back to UCLB,
unless  and  to  the  extent  the  Company  has  exercised  its  option  to  acquire  ownership  of  the  licensed  patent  rights.  In  addition,  UCLB  has  the  right  to
negotiate with the Company for the grant of an exclusive license to the Company’s improvements to the T cell programming modules the Company has
licensed on terms to be agreed upon at the time.

Noile-Immune Biotech Inc.

In  November  2019,  the  Company  entered  into  an  exclusive  license  agreement  with  Noile-Immune  Biotech  Inc.  ("Noile")  under  which  the
Company will have the right to develop CAR T cell therapies incorporating Noile’s PRIME (proliferation-inducing and migration-enhancing) technology.
The PRIME technology is designed to improve proliferation and trafficking into solid tumors of both engineered CAR T cells as well as the patient’s own T
cells.

The Company paid an upfront fee and may be obligated to make additional payments to Noile upon the achievement of development milestones
and receipt of regulatory approvals product sale milestones, as well as royalty payments based on possible future sales resulting from the utilization of the
licensed technology.

Note 10. Income Taxes

The Company recorded an income tax benefit of $24.2 million, $15.2 million, $2.6 million, and $7.3 million for the years ended December 31,

2020 and 2019, the three months ended December 31, 2018, and the year ended September 30, 2018, respectively.

F-22

 
AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

A reconciliation of income tax expense (benefit) at the statutory corporate income tax rate to the income tax expense (benefit) at the Company’s

effective income tax rates is as follows (in thousands): 

Net loss before taxes
U.K. statutory tax rate
Income tax benefit at U.K. statutory tax rate
Tax incentives / credits
Non-deductible expenses
Adjustments in respect of prior years
Operating losses
Tax  on  property,  plant,  equipment  and
intangibles
Other, net
Foreign rate differential

Total income tax benefit
Current income tax benefit
Deferred income tax benefit

$

Year ended December 31,

2020
(166,257)

$

2019
(139,008)

$

Three months ended
December 31,
2018
(23,253)

$

19.0 %

19.0 %

19.0 %

(31,589)
(23,076)
797 
(1,088)
28,672 

547 
1,552 
22 
(24,163)
(22,819)
(1,344)

$
$

(26,411)
(16,312)
4,048 
96 
21,643 

267 
1,510 
— 
(15,159)
(14,749)
(410)

$

(4,423)
(3,234)
127 
265 
3,605 

140 
915 
— 
(2,605)
(2,605)
— 

Year ended September 30,
2018

$

$
$

(52,031)

19.0 %

(9,886)
(7,296)
1,553 
(13)
7,317 

233 
812 
— 
(7,280)
(7,280)
— 

Effective rate of income tax

14.5 %

10.9 %

11.2 %

14.0 %

The effective tax rates in the above table for the years ended December 31, 2020 and 2019, the three months ended December 31, 2018, and the
year ended September 30, 2018 is lower than the main rate of U.K. tax primarily due to administration of the U.K. research and development tax credit,
which is included within the tax incentive/credits line in the table above.

Deferred tax assets and liabilities consisted of the following at December 31, 2020 and 2019 (in thousands): 

Deferred tax assets:
Other differences
Tax losses
Fixed assets
Total deferred tax assets
Valuation allowances
Net deferred tax asset (liability)

December 31,

2020

2019

$

$

14,135  $
40,221 
3,934 
58,290 
(56,536)

1,754  $

1,522 
19,624 
896 
22,042 
(21,632)
410 

Deferred tax assets resulting from loss carry forwards, fixed assets and retirement benefits, with total deferred tax assets increasing by $1.3 million
in  2020.  The  Company  has  recorded  a  valuation  allowance  against  the  net  deferred  tax  asset  where  the  recoverability  due  to  future  taxable  profits  is
unknown. The $1.8 million deferred tax asset balance is related to the Company's U.S. entity.

At December 31, 2020, the Company had U.K. trading losses carry forward of $211.7 million. These losses are carried forward indefinitely under

local law, but are subject to numerous utilization criteria and restrictions.

As required by the authoritative guidance on accounting for income taxes, the Company evaluates the realizability of deferred tax assets at each
reporting date. Accounting for income taxes guidance requires that a valuation allowance be established when it is more likely than not that all or a portion
of the deferred tax assets will not be realized. In circumstances where this is sufficient negative evidence indicating that the deferred tax assets are not more
likely than not realizable, the Company establishes a valuation allowance. The Company recorded valuation allowances in the amounts of $56.5 million
and $21.6 million at December 31, 2020 and 2019, respectively.

F-23

AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

Note 11. Commitments and Contingencies

License Agreement

The  Company  has  entered  into  an  exclusive  license  agreement,  as  amended,  with  UCLB  (see  Note  9).  In  connection  with  the  UCLB  license
agreement,  the  Company  is  required  to  make  annual  license  payments  and  may  be  required  to  make  payments  upon  the  achievement  of  specified
milestones. The Company has estimated the probability of the Company achieving each potential milestone in accordance with ASC 450, Contingencies.
The Company concluded that, as of December 31, 2020, there were no other milestones for which the likelihood of achievement was probable.

Legal Proceedings

From time to time, the Company may be a party to litigation or subject to claims incident to the ordinary course of business. Regardless of the
outcome, litigation can have an adverse impact on the Company because of defense and settlement costs, diversion of management resources and other
factors. The Company was not a party to any litigation and did not have contingency reserves established for any liabilities as of December 31, 2020.

Leases

    The Company leases certain office space, laboratory space, and equipment. At the inception of an arrangement, the Company determines whether the
arrangement  is  or  contains  a  lease  based  on  the  unique  facts  and  circumstances  present.  The  Company  does  not  recognize  right-of-use  assets  or  lease
liabilities  for  leases  determined  to  have  a  term  of  12  months  or  less.  Many  of  the  Company's  leases  contain  variable  non-lease  components  such  as
maintenance, taxes, insurance, and similar costs for the spaces it occupies. For new and amended leases beginning in 2019, the Company has elected the
practical expedient not to separate these non-lease components of leases for classes of all underlying assets and instead account for them as a single lease
component  for  all  leases.  The  Company  recognizes  on  a  straight-lines  basis  the  net  fixed  payments  of  operating  leases  over  the  lease  term.  Variable
executory costs, as it relates to net leases, are excluded from the calculation of the lease liability and the Company expenses the variable lease payments in
the period in which it incurs the obligation to pay such variable amounts and will be included in variable lease costs in the leases footnote disclosure.

    These variable lease payments are not included in the Company's calculation of its right-of-use assets or lease liabilities.

        In  adopting  ASC  842,  in  the  year  ended  December  31,  2019,  the  Company  elected  the  package  of  practical  expedients  which,  among  other  things,
allowed it to retain the classification of its leases in place at the effective date of ASC 842.

The Company’s corporate headquarters are located in London, United Kingdom. As of December 31, 2020, and 2019, the Company leased space at
this location from Imperial (Forest House) Limited under a ten year lease, the term of which commenced in September 2015. The lease included an option
for the Company to lease additional space within a 15-month period, which the Company exercised in October 2016. The exercise of the option resulted in
a  separate  new  lease  with  a  concurrent  term  through  August  2025.  The  Company  and  the  landlord  has  the  option  to  early  terminate  both  leases  in
September 2020 and the landlord has the option to give notice to terminate the lease from September 2020 onward. The Company has measured its right-
of-use assets and lease liabilities based on lease terms ending in September 2025. The landlord exercised its option to give notice in September 2020 to
terminate the Forest House lease and pay the Company a break-lease payment fee in September 2021. The Company recorded a $1.3 million gain upon the
notice of lease termination and lease incentive receivable of $1.2 million in the third quarter of 2020. In addition to base rent, the Company is obligated to
pay its proportionate share of building operating expenses and real estate taxes in excess of base year amounts. These costs are considered to be variable
lease payments and are not included in the determination of the lease’s right-of-use asset or lease liability.

Prior to the lease commencement date of the Forest House leases, the Company, in conjunction with the landlord, made improvements to the leased
space. The total cost of these improvements was funded by the landlord, a portion of the cost will be reimbursed by the Company over the term of the
leases. The total cost of the improvements was capitalized as leasehold improvements on the Company’s balance sheet, with an offset to long-term lease
incentive obligation for the portion funded by the landlord and other long-term payables for the portion to be repaid to the landlord. The lease related to this
facility is classified as an operating lease.

In  September  2017,  the  Company  executed  an  arrangement  with  Catapult  Limited  to  lease  a  manufacturing  suite  at  the  Cell  and  Gene  Therapy

Catapult manufacturing center in Stevenage, United Kingdom for a term through May 2021, at which time the

F-24

    
AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

Company has the option to renew or terminate the lease. The lease related to this facility is classified as an operating lease. The lease has a six-month rent-
free period. In addition to base rent, the Company is obligated to pay its proportionate share of building operating expenses and real estate taxes in excess
of base year amounts. These costs are considered to be variable lease payments and are not included in the determination of the lease’s right-of-use asset or
lease liability. In December 2018, the Company executed an additional lease arrangement for additional manufacturing space for a term through September
2023, at which time the Company has the option to renew or terminate the lease.

    In June 2018, the Company signed a binding letter of intent to enter into a lease for office and laboratory space in White City, London. The letter of
intent  required  the  Company  to  enter  into  a  ten-year  lease  provided  that  the  landlord  completed  the  required  leasehold  improvements  described  in  the
agreement. The leasehold improvements were completed and the lease commenced in January 2019. The Company has the option to terminate the lease in
November 2026. In addition to base rent, the Company is obligated to pay its proportionate share of building operating expenses and real estate taxes in
excess of base year amounts. These costs are considered to be variable lease payments and are not included in the determination of the lease’s right-of-use
asset or lease liability. The lease agreement includes an option to lease additional space. As of December 31, 2020, the Company capitalized $6.7 million as
leasehold improvements.

    In September 2018, the Company signed a binding letter of intent to enter into a lease for manufacturing space in Enfield, United Kingdom. The letter of
intent  required  the  Company  to  enter  into  a  15-year  lease  provided  that  the  landlord  completed  the  required  leasehold  improvements  described  in  the
agreement. The Company executed lease agreements for three manufacturing space units, each for 15-year lease terms. The leases commenced in February
2019  with  option  to  terminate  the  lease  in  February  2029.  In  addition  to  base  rent,  the  Company  is  obligated  to  pay  its  proportionate  share  of  building
operating expenses and real estate taxes in excess of base year amounts. These costs are considered to be variable lease payments and are not included in
the  determination  of  the  lease’s  right-of-use  asset  or  lease  liability.  The  Company  expensed  $4.1  million  of  leasehold  improvements  from  assets  under
construction as of December 31, 2019 as a result of discontinuing the fit-out of the manufacturing facility. The Company reduced the right-of-use asset and
lease liability based on the contractual option termination date. The Company is actively seeking to sub-lease or assign the lease arrangements to a third
party. The Company completed an asset impairment analysis of the right-of-use lease concluding the undiscounted cash flows exceeded the carrying value
as of December 31, 2020.

    In October 2018, the Company executed an agreement to sublease office space in Rockville, Maryland for a term through October 2021. The Company
then  terminated  the  sublease  in  February  2020  and  immediately  entered  into  a  five-year  lease  for  the  same  space  with  the  landlord.  As  a  result  of  the
sublease termination, the Company recognized a $0.2 million gain in other income (expense) in the first quarter of 2020. The lease related to this facility is
classified as an operating lease. The Company is obligated to pay its proportionate share of building operating expenses and real estate taxes in excess of
base year amounts. These costs are considered to be variable lease payments and are not included in the determination of the lease’s right-of-use asset or
lease liability.

        In  January  2019,  the  Company  executed  a  lease  agreement  to  lease  additional  office  and  manufacturing  space  in  Rockville,  Maryland.  The  lease
agreement  required  the  Company  to  enter  into  a  lease  provided  that  the  landlord  completes  the  required  leasehold  improvements  described  in  the
agreement. The lease commenced in August 2020 for a term through June 2036. The lease related to this facility is classified as an operating lease. The
Company has capitalized $2.4 million in leasehold improvements as assets under construction as of December 31, 2020.

In May 2020, the Company executed an arrangement with Catapult Limited to lease a manufacturing suite at the Cell and Gene Therapy Catapult
manufacturing center in Stevenage, United Kingdom for a term through April 2024. The lease related to this facility is classified as an operating lease. In
addition to base rent, the Company is obligated to pay its proportionate share of building operating expenses and real estate taxes in excess of base year
amounts.  These  costs  are  considered  to  be  variable  lease  payments  and  are  not  included  in  the  determination  of  the  lease’s  right-of-use  asset  or  lease
liability.    

       The  Company  identified  and  assessed  the  following  significant  assumptions  in  recognizing  its  right-of-use  assets  and  corresponding  lease  liabilities
during the adoption of ASC 842, in the year ended December 31, 2019:

• As the Company's leases do not provide an implicit rate, it estimated the incremental borrowing rate for each lease based on a yield curve analysis,
utilizing the interest rate derived from the fair value analysis of its existing leases and adjusting it for factors that appropriately reflect the profile
of secured borrowing over the lease term. For leases existing as of the adoption date, the Company has utilized its incremental borrowing rate
based  on  the  remaining  lease  term  as  of  the  adoption  date.  For  leases  that  commenced  after  the  adoption  date,  the  Company  determined  the
incremental borrowing rate based on the lease term as determined at the commencement date of the lease.

F-25

AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

•

•

The expected lease terms include both contractual lease periods and, when applicable, cancellable option periods where failure to exercise such
options would result in an economic penalty.

Since the Company elected to account for each lease component and its associated non-lease components as a single combined lease component,
all contract consideration was allocated to the combined lease component.

Leases (in thousands)

Balance Sheet Classification

Assets
   Operating lease assets
Liabilities
Current
   Operating lease liabilities
Noncurrent
   Operating lease liabilities

Total lease liabilities

Operating right-of-use assets

Current liabilities: Operating lease liabilities

Non-current liabilities: Operating lease liabilities

Lease cost (in thousands)
Operating lease cost
Variable lease cost

Statement of Operations Classification
Operating expenses: research and development
Operating expenses: research and development

Operating lease cost
Variable lease cost

Operating expenses: general and administrative
Operating expenses: general and administrative

Short term lease costs

Total lease cost

As of December 31,

2020

2019

51,637  $

23,409 

3,590  $

50,571  $
54,161  $

Year Ended December 31,
2019
2020

$

5,266 
1,212 

1,380 
270 

282 
8,410 

$

2,511 

23,710 
26,221 

3,156 
1,222 

1,241 
250 

270 
6,139 

$

$

$
$

$

$

Other information (in thousands)
Cash paid for amounts included in the measurement of lease liabilities
   Operating cash flows from operating leases
Right-of-use assets obtained in exchange for new operating lease liabilities
Weighted-average remaining lease term — operating leases (years)
Weighted-average discount rate — operating leases

December 31,

2020

2019

$
$

4,896 
30,786 

$
$

11.2
9.0 %

2,415 
26,124 

7.7
6.8 %

F-26

AUTOLUS THERAPEUTICS PLC
Notes to Consolidated Financial Statements — Continued

    Future fixed payments for non-cancellable operating leases in effect as of December 31, 2020 are payable as follows:

Maturity of lease liabilities for the years ending December 31,
2021
2022
2023
2024
2025
Thereafter
Total lease payments
Less: imputed interest

Present value of lease liabilities

Operating Leases
(in thousands)

7,026 
9,112 
8,644 
8,150 
7,044 
51,259 
91,235 
(37,074)
54,161 

$

$

    The Company recognized rent expense on a straight-line basis over the respective lease period and has recorded deferred rent for rent expense incurred
but not yet paid prior to the adoption of ASC 842.

The  Company  recorded  rent  expense  totaling  $0.5  million  for  the  three  months  ended  December  31,  2018,  and  $0.9  million for  the  year  ended

September 30, 2018, respectively.

Note 12. Employee Benefit Plans

        In  the  United  Kingdom,  the  Company  makes  contributions  to  private  defined  benefit  pension  schemes  on  behalf  of  its  employees.  The  Company
expensed $1.3 million, $1.0 million, $0.2 million, and $0.5 million in contributions in the years ended December 31, 2020 and 2019, three months ended
December 2018, and the year ended September 30, 2018, respectively.

    In the United States, the Company established a defined contribution savings plan under Section 401(k) of the Internal Revenue Code subsequent to
September 30, 2018. The plan covers substantially all U.S. employees who meet minimum age and service requirements and allows participants to defer a
portion of their annual compensation on a pre-tax basis. The Company matches employee contributions up to four percent of the employee’s annual salary.
The  Company  expensed  $0.3  million,  $0.2  million  and  $18,000  in  contributions  in  the  years  ended  December  31,  2020  and  2019  and  the  three  months
ended  December  31,  2018,  respectively.  No  matching  contributions  were  recorded  for  the  year  ended  September  30,  2018.  The  Company  pays  all
administrative fees related to the Plan.

Note 13. Subsequent Events

    The Company evaluated subsequent events through March 4, 2021 the date on which these financial statements were issued.

In January 2021, the Company announced the prioritization of the AUTO1 program and its intention to seek a partner for the AUTO3 program
before progressing AUTO3 into the next phase of development. The Company also announced an adjustment of its workforce and infrastructure footprint
during the first quarter of 2021, which will involve an overall reduction in headcount of approximately 20%. The company expects to realize cash savings,
on an annualized basis, of approximately $15 million per annum once the operational changes are fully implemented. In the short-term, we expect there to
be an increase in the first half of 2021 to both research and development expenses and selling, general, and administrative expenses due to restructuring
charges of approximately $2.5 million, combined.

In January 2021, the Company sold an aggregate of 1.7 million ADSs under the Sales Agreement, resulting in net proceeds of $15.3 million.

On  February  12,  2021,  the  Company  completed  an  underwritten  public  offering  of  14,285,715  ADSs,  which  includes  the  full  exercise  by  the
underwriters  to  purchase  an  additional  2,142,857  ADSs,  at  a  public  offering  price  of  $7.00  per  ADS.  Aggregate  net  proceeds  to  the  Company,  after
underwriting discounts, were $108.1 million. .

F-27

    
    
Exhibit 2.4

DESCRIPTION OF THE REGISTRANT'S SECURITIES
REGISTERED PURSUANT TO SECTION 12 OF THE
SECURITIES EXCHANGE ACT OF 1934

The following description sets forth certain material terms and provisions of the securities of Autolus Therapeutics plc (“Autolus,” the “Company,” “we,”
“us,” and “our”) that are registered under Section 12 of the Securities Exchange Act of 1934, as amended (the “Exchange Act”). This description also
summarizes relevant provisions of English law, including the U.K. Companies Act 2006 (the “Companies Act”). The following summary does not purport
to be complete and is subject to, and is qualified in its entirety by reference to, the applicable provisions of English law and our articles of association, a
copy of which is incorporated by reference as an exhibit to the Annual Report on 20-F of which this Exhibit is a part. We encourage you to read our articles
of association and the applicable provisions of English law for additional information.

General

Our  securities  include  (a)  our  ordinary  shares,  nominal  value  $0.000042  per  share,  and  (b)  our  American  Depositary  Shares  (the  “ADSs”),  each
representing one ordinary share, nominal value $0.000042 per share. Our ordinary shares are registered under the Exchange Act not for trading, but only in
connection with the listing of the ADSs on The Nasdaq Global Select Market.

Our ADSs are listed on The Nasdaq Global Select Market under the trading symbol “AUTL.”

The following is a description of the rights of (i) the holders of ordinary shares and (ii) ADS holders. Ordinary shares underlying the outstanding ADSs are
held by Citibank N.A., as depositary.

Ordinary Shares

The following is a summary of the rights of our holders of our ordinary shares as specified in our articles of association which was adopted by a special
resolution of our shareholders passed in June 2018.

Type and Class of Securities

Each ordinary share has a nominal value of $0.000042.

Preemptive Rights

English law generally provides shareholders with statutory preemptive rights when new shares are issued for cash; however, it is possible for the articles of
association, or shareholders by way of a special resolution at a general meeting, to disapply preemptive rights. Such a disapplication of preemptive rights
may be for a maximum period of up to five years from the date of adoption of the articles of association, if the disapplication is contained in the articles of
association,  or  from  the  date  of  the  shareholder  special  resolution,  if  the  disapplication  is  by  shareholder  special  resolution.  In  either  case,  this
disapplication  would  need  to  be  renewed  by  our  shareholders  upon  its  expiration  (i.e.,  at  least  every  five  years).  Our  articles  of  association  disapply
preemptive  rights  for  a  period  of  five  years  from  the  date  of  adoption,  which  was  June  26,  2018.  This  disapplication  will  need  to  be  renewed  upon
expiration (i.e., at least every five years) to remain effective, but may be sought more frequently for additional five-year terms (or any shorter period).

Limits of Qualifications

Not applicable.

Registration Rights

We and the holders of certain of our ordinary shares are parties to a registration rights agreement that provides the following registration rights:

• Demand Registration on Form F-1 - each holder is entitled to demand registrations on Form F-1, provided that these demand registration rights
may only be exercised by holders who hold, in the aggregate, not less than 25% of the aggregate number of shares then outstanding and held by
all holders who are party to the agreement, and provided further that the we shall not be required to effect a demand registration statement after
we have effected two demand registration statements, and such registration statements have been declared or ordered effective.

Exhibit 2.4

• Demand Registration on Form F-3 - each holder is entitled to unlimited demand registrations on Form F-3, if we are eligible to register shares on
Form F-3, provided that these demand registration rights may only be exercised by holders who hold, in the aggregate, not less than 10% of the
aggregate number of shares then outstanding and held by all holders who are party to the agreement. These demand registration rights may not be
exercised more than twice in any twelve-month period.
Piggyback Registration - each holder is entitled to piggyback registration rights, subject, in the case of an underwritten offering, to customary
reductions by the underwriter, provided that the aggregate number of securities of the holders included in the registration may not be reduced to
less than 30% of the total number of securities registered.
Expenses  -  We  will  pay  all  registration  expenses  relating  to  the  exercise  of  the  registration  rights  above,  including  the  reasonable  fees  and
expenses of legal counsel to the participating holders up to a maximum of $50,000 in the aggregate per registration.

•

•

Memorandum and Articles of Association

Our ordinary shares have the rights and restrictions described in “Key Provisions of Our Articles of Association” below. The following summarizes the
rights of holders of our ordinary shares:

•
•
•

each holder of our ordinary shares is entitled to one vote per ordinary share on all matters to be voted on by shareholders generally;
the holders of the ordinary shares shall be entitled to receive notice of, attend, speak and vote at our general meetings; and
holders of our ordinary shares are entitled to receive such dividends as are recommended by our directors and declared by our shareholders.

Key Provisions of Our Articles of Association

The following is a summary of certain key provisions of our articles of association. Please note that this is only a summary and is not intended to be

exhaustive.

The articles of association contain no specific restrictions on our purpose and therefore, by virtue of section 31(1) of the Companies Act, our purpose is
unrestricted. The articles of association contain, among other things, provisions to the following effect:

Share Capital

Our share capital currently consists of ordinary shares, deferred shares, B deferred shares and C deferred shares. We may issue shares with such rights or
restrictions  as  may  be  determined  by  ordinary  resolution,  including  shares  which  are  to  be  redeemed,  or  are  liable  to  be  redeemed  at  our  option  or  the
option of the holder of such shares.

Voting

Holders of ordinary shares have the right to receive notice of, and to vote at, our general meetings. Any resolution put to the vote of a general meeting must
be decided exclusively on a poll. Each shareholder who is present in person (or, being a corporation, by representative) or by proxy has one vote in respect
of every share held by such holder.

Variation of Rights

Exhibit 2.4

Whenever our share capital is divided into different classes of shares, the special rights attached to any class may be varied or abrogated either (i) with the
consent in writing of the holders of three-quarters in nominal value of the issued shares of that class, (ii) with the authority of a special resolution passed at
a separate meeting of the holders of the shares of that class or (iii) in any other way as expressly provided for in relation to such rights, and may be so
varied and abrogated while the company is a going concern.

Dividends

We may, subject to the provisions of the Companies Act and our articles of association, by ordinary resolution from time to time declare dividends to be
paid to shareholders not exceeding the amount recommended by our board of directors. Subject to the provisions of the Companies Act, in the discretion of
board of directors, on the basis that our profits justify such payments, the board of directors may pay interim dividends on any class of our shares.

Any dividend unclaimed after a period of 12 years from the date such dividend was declared or became payable shall, if the board of directors resolve, be
forfeited, cease to remain owing and shall revert to us. No dividend or other moneys payable on or in respect of a share shall bear interest as against us.

Transfer of Ordinary Shares

Each member may transfer all or any of his shares which are in certificated form by means of an instrument of transfer in writing in any usual form or in
any other form which the board of directors may approve.

The board of directors may, in its absolute discretion, refuse to register a transfer of certificated shares unless:

(i)

(ii)

(iii)

(iv)

(v)

(vi)

it is for a share which is fully paid up;

it is for a share upon which the company has no lien;

it is only for one class of share;

it is in favor of a single transferee or no more than four joint transferees;

it is duly stamped or is duly certificated or otherwise shown to the satisfaction of the board of directors to be exempt from
stamp duty; and

it  is  delivered  for  registration  to  the  registered  office  of  the  company  (or  such  other  place  as  the  board  of  directors  may
determine), accompanied (except in the case of a transfer by a person to whom the company is not required by law to issue a
certificate and to whom a certificate has not been issued or in the case of a renunciation) by the certificate for the shares to
which it relates and such other evidence as the board of directors may reasonably require to prove the title of the transferor
(or  person  renouncing)  and  the  due  execution  of  the  transfer  or  renunciation  by  him  or,  if  the  transfer  or  renunciation  is
executed by some other person on his behalf, the authority of that person to do so.

Allotment of Shares and Preemption Rights

Subject to the Companies Act and to any rights attached to existing shares, any share may be issued with or have attached to it such rights and restrictions
as  the  company  may  by  ordinary  resolution  determine,  or  if  no  ordinary  resolution  has  been  passed  or  so  far  as  the  resolution  does  not  make  specific
provision, as the board of directors may determine (including shares which are to be redeemed, or are liable to be redeemed at the option of the company or
the holder of such shares).

In  accordance  with  section  551  of  the  Companies  Act,  the  board  of  directors  may  be  generally  and  unconditionally  authorized  to  exercise  for  each
prescribed  period  all  the  powers  of  the  company  to  allot  shares  up  to  an  aggregate  nominal  amount  equal  to  the  amount  stated  in  the  relevant  ordinary
resolution authorizing such allotment. The authorities referred to above are

 
Exhibit 2.4

included in our articles of association that became effective upon the completion of our initial public offering in June 2018 and which remain currently in
force as of December 31, 2019.

The provisions of section 561 of the Companies Act (which confer on shareholders rights of preemption in respect of the allotment of equity securities
which  are  paid  up  in  cash)  apply  to  the  company  except  to  the  extent  disapplied  by  special  resolution  of  the  shareholders  of  the  company,  or  in  the
company’s articles of association. Our articles of association disapply preemptive rights for a period of five years from the date of adoption, which was
June 26, 2018. This disapplication will need to be renewed upon expiration (i.e., at least every five years) to remain effective, but may be sought more
frequently for additional five-year terms (or any shorter period).

Alteration of Share Capital

In accordance with the Companies Act, the company may by ordinary resolution consolidate its share capital into shares of larger nominal value than its
existing shares, or sub-divide its shares into shares of a smaller amount than the existing shares, and may in each case determine that the shares resulting
from such sub-division or share consolidation may have a preference or advantage or be subject to a particular restriction.

The company may, in accordance with the Companies Act, reduce or cancel its share capital or any capital redemption reserve or share premium account in
any manner and with and subject to any conditions, authorities and consents required by law.

Uncertificated Shares

Subject to the Companies Act, the board of directors may permit title to shares of any class to be issued or held otherwise than by a certificate and to be
transferred by means of a “relevant system” (e.g., DTC) without a certificate.  

The board of directors may take such steps as it sees fit in relation to the evidencing of and transfer of title to uncertificated shares, any records relating to
the holding of uncertificated shares and the conversion of uncertificated shares to certificated shares, or vice versa.

The company may by notice in writing to the holder of an uncertificated share, require that share to be converted into certificated form.

The  board  of  directors  may  take  such  other  action  that  the  board  considers  appropriate  to  achieve  the  sale,  transfer,  disposal,  forfeiture,  re-allotment  or
surrender of an uncertified share or otherwise to enforce a lien in respect of it.

Choice of Forum

Our articles of association provide that the U.S. federal district courts will be the exclusive forum for resolving any complaint asserting a cause of action
arising under the Securities Act.

Board of Directors

Our articles of association provide that our board of directors is divided into three classes, each of which will consist, as nearly as possible, of one-third of
the total number of directors constituting our entire board and which will serve staggered three-year terms. At each annual general meeting, the successors
to directors whose terms then expire will be elected to serve from the time of election and qualification until the third annual meeting following election.

At every subsequent annual general meeting, any director who either (i) has been appointed by the board of directors since the last annual general meeting
or (ii) was not appointed or reappointed at one of the preceding two annual general meetings, must retire from office and may offer themselves for
reappointment by the shareholders by ordinary resolution.

American Depositary Shares

Depositary

Exhibit 2.4

We have appointed Citibank, N.A. (“Citibank”) as the depositary for the ADSs pursuant to a deposit agreement (the “Deposit Agreement”). Citibank’s
depositary offices are located at 388 Greenwich Street, New York, New York 10013.

Provisions

ADSs represent ownership interests in securities that are on deposit with the depositary. ADSs may be represented by certificates that are commonly known
as American Depositary Receipts (“ADRs”). The depositary typically appoints a custodian to safekeep the securities on deposit. In this case, the custodian
is Citibank, N.A., London Branch located at 25 Canada Square, Canary Wharf, London, E14 5LB, United Kingdom.

The following is a summary of the material provisions of the Deposit Agreement. For more complete information, you should read the Deposit Agreement
and Form of ADR. The Deposit Agreement has been filed with the SEC as an exhibit to the Form F-6/A filed on June 19, 2018. A copy of the Deposit
Agreement may be obtained from the SEC’s website (www.sec.gov). Please refer to File No. 333-224837.

Each ADS represents the right to receive, and to exercise the beneficial ownership interests in, one ordinary share that is on deposit with the depositary
and/or custodian. An ADS also represents the right to receive, and to exercise the beneficial interests in, any other property received by the depositary or
the custodian on behalf of the owner of the ADS but that has not been distributed to the owners of ADSs because of legal restrictions or practical
considerations. We and the depositary may agree to change the ADS-to-ordinary share ratio by amending the Deposit Agreement. This amendment may
give rise to, or change, the depositary fees payable by ADS owners. The custodian, the depositary and their respective nominees will hold all deposited
property for the benefit of the holders and beneficial owners of ADSs. The deposited property does not constitute the proprietary assets of the depositary,
the custodian or their nominees. Beneficial ownership in the deposited property will under the terms of the Deposit Agreement be vested in the beneficial
owners of the ADSs. The depositary, the custodian and their respective nominees are the record holders of the deposited property represented by the ADSs
for the benefit of the holders and beneficial owners of the corresponding ADSs. A beneficial owner of ADSs may or may not be the holder of ADSs.
Beneficial owners of ADSs are able to receive, and to exercise beneficial ownership interests in, the deposited property only through the registered holders
of the ADSs, the registered holders of the ADSs (on behalf of the applicable ADS owners) only through the depositary, and the depositary (on behalf of the
owners of the corresponding ADSs) directly, or indirectly, through the custodian or their respective nominees, in each case upon the terms of the Deposit
Agreement.

Beneficial owners of ADSs, or holders, are parties to the Deposit Agreement and therefore are bound to its terms and to the terms of any ADR that
represents the ADSs. The Deposit Agreement and the ADR specify our rights and obligations as well as holders’ rights and obligations as owner of ADSs
and those of the depositary. Holders appoint the depositary to act on their behalf in certain circumstances. The Deposit Agreement and the ADRs are
governed by New York law. However, our obligations to the holders of ordinary shares will continue to be governed by the laws of England and Wales,
which may be different from the laws in the United States.

In addition, applicable laws and regulations may require holders to satisfy reporting requirements and obtain regulatory approvals in certain circumstances.
Neither the depositary, the custodian, us or any of their or our respective agents or affiliates shall be required to take any actions whatsoever on behalf of
the holders to satisfy such reporting requirements or obtain such regulatory approvals under applicable laws and regulations.

Holders will not be treated as one of our shareholders and will not have direct shareholder rights. The depositary will hold on the behalf of holders the
shareholder rights attached to the ordinary shares underlying the ADSs. Holders are able to exercise the shareholders rights for the ordinary shares
represented by their ADSs through the depositary only to the extent contemplated in the Deposit Agreement. To exercise any shareholder rights not
contemplated in the Deposit Agreement holders will need to arrange for the cancellation of their ADSs and become a direct shareholder. The manner in
which ADSs are owned (e.g., in brokerage account vs. as registered holder, or as holder of certificated vs. uncertificated) may affect the holder’s rights and
obligations, and the manner in which the depositary’s services are made available.

Holders may hold ADSs either by means of an ADR registered in their name, through a brokerage or safekeeping account, or through an account
established by the depositary in their name reflecting the registration of uncertificated ADSs directly on the books of the depositary (commonly referred to
as the direct registration system or DRS). The direct registration system reflects the uncertificated (book-entry) registration of ownership of ADSs by the
depositary. Under the direct registration system, ownership of ADSs is evidenced by periodic statements issued by the depositary to the holders. The direct
registration system includes automated transfers between the depositary and The Depository Trust Company, or DTC, the

Exhibit 2.4

central book-entry clearing and settlement system for equity securities in the United States. If a holder decides to hold their ADSs through a brokerage or
safekeeping account, they must rely on the procedures of their broker or bank to assert their rights as ADS owner. Banks and brokers typically hold
securities such as the ADSs through clearing and settlement systems such as DTC. The procedures of such clearing and settlement systems may limit the
holder’s ability to exercise their rights as an owner of ADSs. All ADSs held through DTC are registered in the name of a nominee of DTC. This summary
description assumes the holder has opted to own the ADSs directly by means of an ADS registered in their name at the relevant time.

The registration of the ordinary shares in the name of the depositary or the custodian shall, to the maximum extent permitted by applicable law, vest in the
depositary or the custodian the record ownership in the applicable ordinary shares with the beneficial ownership rights and interests in such ordinary shares
being at all times vested with the beneficial owners of the ADSs representing the ordinary shares. The depositary or the custodian shall at all times be
entitled to exercise the beneficial ownership rights in all deposited property, in each case only on behalf of the holders and beneficial owners of the ADSs
representing the deposited property.

Dividends and Other Distributions

Holders generally have the right to receive the distributions we make on the securities deposited with the custodian. Receipt of these distributions may be
limited, however, by practical considerations and legal limitations. Holders will receive such distributions under the terms of the Deposit Agreement in
proportion to the number of ADSs held as of the specified record date, after deduction the applicable fees, taxes and expenses.

Distributions of Cash

Whenever we make a cash distribution for the securities on deposit with the custodian, we will deposit the funds with the custodian. Upon receipt of
confirmation of the deposit of the requisite funds, the depositary will arrange for the funds received in a currency other than U.S. dollars to be converted
into U.S. dollars and for the distribution of the U.S. dollars to the holders, subject to the laws and regulations of England and Wales.

The conversion into U.S. dollars will take place only if practicable and if the U.S. dollars are transferable to the United States. The depositary will apply the
same method for distributing the proceeds of the sale of any property (such as undistributed rights) held by the custodian in respect of securities on deposit.

The distribution of cash will be made net of the fees, expenses, taxes and governmental charges payable by holders under the terms of the Deposit
Agreement. The depositary will hold any cash amounts it is unable to distribute in a non-interest bearing account for the benefit of the applicable holders
and beneficial owners of ADSs until the distribution can be effected or the funds that the depositary holds must be escheated as unclaimed property in
accordance with the laws of the relevant states of the United States.

Distributions of Shares

Whenever we make a free distribution of ordinary shares for the securities on deposit with the custodian, we will deposit the applicable number of ordinary
shares with the custodian. Upon receipt of confirmation of such deposit, the depositary will either distribute to holders new ADSs representing the ordinary
shares deposited or modify the ADS-to-ordinary shares ratio, in which case each ADS held will represent rights and interests in the additional ordinary
shares so deposited. Only whole new ADSs will be distributed. Fractional entitlements will be sold and the proceeds of such sale will be distributed as in
the case of a cash distribution.

The distribution of new ADSs or the modification of the ADS-to-ordinary share ratio upon a distribution of ordinary shares will be made net of the fees,
expenses, taxes and governmental charges payable by holders under the terms of the Deposit Agreement. In order to pay such taxes or governmental
charges, the depositary may sell all or a portion of the new ordinary shares so distributed.

No such distribution of new ADSs will be made if it would violate a law (e.g., the U.S. securities laws) or if it is not operationally practicable. If the
depositary does not distribute new ADSs as described above, it may sell the ordinary shares received upon the terms described in the Deposit Agreement
and will distribute the proceeds of the sale as in the case of a distribution of cash.

Exhibit 2.4

Distributions of Rights

Whenever we intend to distribute rights to purchase additional ordinary shares, we will give prior notice to the depositary and we will assist the depositary
in determining whether it is lawful and reasonably practicable to distribute rights to purchase additional ADSs to holders.

The depositary will establish procedures to distribute rights to purchase additional ADSs to holders and to enable such holders to exercise such rights if it is
lawful and reasonably practicable to make the rights available to holders of ADSs, and if we provide all of the documentation contemplated in the Deposit
Agreement (such as opinions to address the lawfulness of the transaction). Holders may have to pay fees, expenses, taxes and other governmental charges
to subscribe for the new ADSs upon the exercise of rights. The depositary is not obligated to establish procedures to facilitate the distribution and exercise
by holders of rights to purchase new ordinary shares other than in the form of ADSs.

The depositary will not distribute the rights if:

•
•
•

we do not timely request that the rights be distributed to the holders or we request that the rights not be distributed to the holders;
we fail to deliver satisfactory documents to the depositary; or
it is not reasonably practicable to distribute the rights.

The depositary will sell the rights that are not exercised or not distributed if such sale is lawful and reasonably practicable. The proceeds of such sale will
be distributed to holders as in the case of a cash distribution. If the depositary is unable to sell the rights, it will allow the rights to lapse.

Elective Distributions

Whenever we intend to distribute a dividend payable at the election of shareholders either in cash or in additional shares, we will give prior notice thereof
to the depositary and will indicate whether we wish the elective distribution to be made available to the holders. In such case, we will assist the depositary
in determining whether such distribution is lawful and reasonably practicable.

The depositary will make the election available to the holders only if it is reasonably practicable and if we have provided all of the documentation
contemplated in the Deposit Agreement. In such case, the depositary will establish procedures to enable the holders to elect to receive either cash or
additional ADSs, in each case as described in the Deposit Agreement.

If the election is not made available to the holders, they will receive either cash or additional ADSs, depending on what a shareholder in England and Wales
would receive upon failing to make an election, as more fully described in the Deposit Agreement.

Other Distributions

Whenever we intend to distribute property other than cash, ordinary shares or rights to purchase additional ordinary shares, we will notify the depositary in
advance and will indicate whether we wish such distribution to be made to the holders. If so, we will assist the depositary in determining whether such
distribution to holders is lawful and reasonably practicable.

If it is reasonably practicable to distribute such property to the holders and if we provide all of the documentation contemplated in the Deposit Agreement,
the depositary will distribute the property to the holders in a manner it deems practicable.

The distribution will be made net of fees, expenses, taxes and governmental charges payable by holders under the terms of the Deposit Agreement. In order
to pay such taxes and governmental charges, the depositary may sell all or a portion of the property received.

The depositary will not distribute the property and will sell the property if:

Exhibit 2.4

•
•
•

we do not request that the property be distributed to the holders or if we ask that the property not be distributed to the holders;
we do not deliver satisfactory documents to the depositary; or
the depositary determines that all or a portion of the distribution is not reasonably practicable.

The proceeds of such a sale will be distributed to holders as in the case of a cash distribution.

Redemption

Whenever we decide to redeem any of the securities on deposit with the custodian, we will notify the depositary in advance. If it is practicable and if we
provide all of the documentation contemplated in the Deposit Agreement, the depositary will provide notice of the redemption to the holders.

The custodian will be instructed to surrender the shares being redeemed against payment of the applicable redemption price. The depositary will convert
the redemption funds received into U.S. dollars upon the terms of the Deposit Agreement and will establish procedures to enable holders to receive the net
proceeds from the redemption upon surrender of their ADSs to the depositary. Holders may have to pay fees, expenses, taxes and other governmental
charges upon the redemption of ADSs. If less than all ADSs are being redeemed, the ADSs to be retired will be selected by lot or on a pro rata basis, as the
depositary may determine.

Changes Affecting Ordinary Shares

The ordinary shares held on deposit for the holders’ ADSs may change from time to time. For example, there may be a change in nominal or par value,
split-up, cancellation, consolidation or any other reclassification of such ordinary shares or a recapitalization, reorganization, merger, consolidation or sale
of assets of the company.

If any such change were to occur, the holder’s ADSs would, to the extent permitted by law and the Deposit Agreement, represent the right to receive the
property received or exchanged in respect of the ordinary shares held on deposit. The depositary may in such circumstances deliver new ADSs to the
holders, amend the Deposit Agreement, the ADRs and the applicable Registration Statement(s) on Form F-6, call for the exchange of existing ADSs for
new ADSs and take any other actions that are appropriate to reflect as to the ADSs the change affecting the ordinary shares. If the depositary may not
lawfully distribute such property to the holders, the depositary may sell such property and distribute the net proceeds to the holders as in the case of a cash
distribution.

Transfer, Combination and Split Up of ADRs

ADR holders are entitled to transfer, combine or split up their ADRs and the ADSs evidenced thereby. For transfers of ADRs, holders will have to
surrender the ADRs to be transferred to the depositary and also must:

•
•

•

•

ensure that the surrendered ADR is properly endorsed or otherwise in proper form for transfer;
provide such proof of identity and genuineness of signatures as the depositary deems appropriate;

provide any transfer stamps required by the State of New York or the United States; and

pay all applicable fees, charges, expenses, taxes and other government charges payable by ADR holders pursuant to the terms of the
Deposit Agreement, upon the transfer of ADRs.

To have ADRs either combined or split up, holders must surrender the ADRs in question to the depositary with their request to have them combined or split
up, and must pay all applicable fees, charges and expenses payable by ADR holders, pursuant to the terms of the Deposit Agreement, upon a combination
or split up of ADRs.

Withdrawal of Ordinary Shares Upon Cancellation of ADSs

Holders are entitled to present their ADSs to the depositary for cancellation and then receive the corresponding number of underlying ordinary shares at the
custodian’s offices. The ability to withdraw the ordinary shares held in respect of the ADSs may be limited by the legal considerations in the United States
and in England and Wales applicable at the time of withdrawal. In order to withdraw the ordinary shares represented by the ADSs, holders will be required
to pay to the depositary the fees for cancellation of ADSs and any charges and taxes payable upon the transfer of the ordinary shares.

Exhibit 2.4

Holders assume the risk for delivery of all funds and securities upon withdrawal. Once canceled, the ADSs will not have any rights under the Deposit
Agreement.

If a holder’s ADSs are registered in the holder’s name, the depositary may ask the holder to provide proof of identity and genuineness of any signature and
such other documents as the depositary may deem appropriate before it will cancel ADSs. The withdrawal of the ordinary shares represented by the
holder’s ADSs may be delayed until the depositary receives satisfactory evidence of compliance with all applicable laws and regulations. Please keep in
mind that the depositary will only accept ADSs for cancellation that represent a whole number of securities on deposit.

Holders have the right to withdraw the securities represented by their ADSs at any time except as a result of:

•

•

•
•

temporary delays that may arise because (i) the transfer books for the ordinary shares or ADSs are closed, or (ii) ordinary shares are
immobilized on account of a shareholders’ meeting or a payment of dividends;

obligations to pay fees, taxes and similar charges;
restrictions imposed because of laws or regulations applicable to ADSs or the withdrawal of securities on deposit; and/or

other circumstances specifically contemplated by Section I.A.(l) of the General Instructions to Form F-6 (as such General Instructions may
be amended from time to time).

The Deposit Agreement may not be modified to impair a holder’s right to withdraw the securities represented by their ADSs except to comply with
mandatory provisions of law.

Voting Rights

Holders generally have the right under the Deposit Agreement to instruct the depositary to exercise the voting rights for the ordinary shares represented by
their ADSs.

At our request, the depositary will distribute to the holders any notice of shareholders’ meeting received from us together with information explaining how
to instruct the depositary to exercise the voting rights of the securities represented by ADSs. In lieu of distributing such materials, the depositary may
distribute to holders of ADSs instructions on how to retrieve such materials upon request.

If the depositary timely receives voting instructions from a holder of ADSs, it will endeavor to vote (or cause the custodian to vote) the securities (in person
or by proxy) represented by the holder’s ADSs as follows: the depositary will vote (or cause the custodian to vote) the securities represented by ADSs in
accordance with the voting instructions received from the holders of ADSs. If the depositary does not receive voting instructions from a holder of ADSs as
of the applicable ADS record date on or before the date established by the depositary for such purpose, such holder will be deemed, and the depositary will
deem such holder, to have instructed the depositary to give a discretionary proxy to a person designated by us to vote the securities represented by ADSs;
provided, however, that no such discretionary proxy will be given by the depositary with respect to any matter to be voted upon as to which we inform the
depositary that (a) we do not wish such proxy to be given, (b) substantial opposition exists or (c) the rights of holders of securities represented by ADSs
may be adversely affected.

Securities for which no voting instructions have been received will not be voted (except as otherwise contemplated in the Deposit Agreement). Please note
that the ability of the depositary to carry out voting instructions may be limited by practical and legal limitations and the terms of the securities on deposit.
We cannot assure holders that they will receive voting materials in time to enable holders to return voting instructions to the depositary in a timely manner.

Amendments and Termination

We may agree with the depositary to modify the Deposit Agreement at any time without the holders’ consent. We undertake to give holders 30 days’ prior
notice of any modifications that would materially prejudice any of their substantial rights under the Deposit Agreement. We do not consider to be
materially prejudicial to the holders’ substantial rights any modifications or supplements that are reasonably necessary for the ADSs to be registered under
the Securities Act or to be eligible for book-entry settlement, in each case without imposing or increasing the fees and charges holders are required to pay.
In addition, we may not be able to provide holders with prior notice of any modifications or supplements that are required to accommodate compliance
with applicable provisions of law.

Exhibit 2.4

Holders are bound by the modifications to the Deposit Agreement if they continue to hold their ADSs after the modifications to the Deposit Agreement
become effective. The Deposit Agreement cannot be amended to prevent holders from withdrawing the ordinary shares represented by their ADSs (except
as permitted by law).

We have the right to direct the depositary to terminate the Deposit Agreement. Similarly, the depositary may in certain circumstances on its own initiative
terminate the Deposit Agreement. In either case, the depositary must give notice to the holders at least 30 days before termination. Until termination,
holders’ rights under the Deposit Agreement will be unaffected.

Termination

After termination, the depositary will continue to collect distributions received (but will not distribute any such property until holders request the
cancellation of their ADSs) and may sell the securities held on deposit. After the sale, the depositary will hold the proceeds from such sale and any other
funds then held for the holders of ADSs in a non-interest bearing account. At that point, the depositary will have no further obligations to holders other than
to account for the funds then held for the holders of ADSs still outstanding (after deduction of applicable fees, taxes and expenses).

Books of Depositary

The depositary will maintain ADS holder records at its depositary office. Holders may inspect such records at such office during regular business hours but
solely for the purpose of communicating with other holders in the interest of business matters relating to the ADSs and the Deposit Agreement.

The depositary will maintain in New York facilities to record and process the issuance, cancellation, combination, split-up and transfer of ADSs. These
facilities may be closed from time to time, to the extent not prohibited by law.

Transmission of Notices, Reports and Proxy Soliciting Material

The depositary makes available for holders’ inspection at its office all communications that it receives from us as a holder of deposited securities that we
make generally available to holders of deposited securities. Subject to the terms of the Deposit Agreement, the depositary will send holders copies of those
communications or otherwise make those communications available to the holders if we ask it to.

Limitations on Obligations and Liabilities

The Deposit Agreement limits our obligations and the depositary’s obligations to holders. Please note the following:

Exhibit 2.4

•

•

•

•

•

•

•

•

•

•

•

We and the depositary are obligated only to take the actions specifically stated in the Deposit Agreement without negligence or bad
faith.
The depositary disclaims any liability for any failure to carry out voting instructions, for any manner in which a vote is cast or for the
effect of any vote, provided it acts in good faith and without negligence and in accordance with the terms of the Deposit Agreement.

The depositary disclaims any liability for any failure to determine the lawfulness or practicality of any action, for the content of any
document forwarded to holders on our behalf or for the accuracy of any translation of such a document, for the investment risks
associated with investing in ordinary shares, for the validity or worth of the ordinary shares, for any tax consequences that result from
the ownership of ADSs, for the credit-worthiness of any third party, for allowing any rights to lapse under the terms of the Deposit
Agreement, for the timeliness of any of our notices or for our failure to give notice.

We and the depositary will not be obligated to perform any act that is inconsistent with the terms of the Deposit Agreement.

We and the depositary disclaim any liability if we or the depositary are prevented or forbidden from or subject to any civil or criminal
penalty or restraint on account of, or delayed in, doing or performing any act or thing required by the terms of the Deposit Agreement,
by reason of any provision, present or future of any law or regulation, or by reason of present or future provision of any provision of
our articles of association, or any provision of or governing the securities on deposit, or by reason of any act of God or war or other
circumstances beyond our control.

We and the depositary disclaim any liability by reason of any exercise of, or failure to exercise, any discretion provided for in the
Deposit Agreement or in our articles of association or in any provisions of or governing the securities on deposit.

We and the depositary further disclaim any liability for any action or inaction in reliance on the advice or information received from
legal counsel, accountants, any person presenting ordinary shares for deposit, any holder of ADSs or authorized representatives thereof,
or any other person believed by either of us in good faith to be competent to give such advice or information.

We and the depositary also disclaim liability for the inability by a holder to benefit from any distribution, offering, right or other
benefit that is made available to holders of ordinary shares but is not, under the terms of the Deposit Agreement, made available to
any applicable holder.
We and the depositary may rely without any liability upon any written notice, request or other document believed to be genuine and
to have been signed or presented by the proper parties.

We and the depositary also disclaim liability for any consequential or punitive damages for any breach of the terms of the Deposit
Agreement.

No disclaimer of any Securities Act liability is intended by any provision of the Deposit Agreement.

Nothing in the Deposit Agreement gives rise to a partnership or joint venture, or establishes a fiduciary relationship, among us, the depositary bank and
ADS holders.

Nothing in the Deposit Agreement precludes Citibank (or its affiliates) from engaging in transactions in which parties adverse to us or the ADS owners
have interests, and nothing in the Deposit Agreement obligates Citibank to disclose those transactions, or any information obtained in the course of those
transactions, to us or to the ADS owners, or to account for any payment received as part of those transactions.

Pre-Release Transactions

Subject to the terms and conditions of the Deposit Agreement, the depositary may issue to broker/dealers ADSs before receiving a deposit of ordinary
shares or release ordinary shares to broker/dealers before receiving ADSs for cancellation. These transactions are commonly referred to as “pre-release
transactions,” and are entered into between the depositary and the applicable broker/dealer. The Deposit Agreement limits the aggregate size of pre-release
transactions (not to exceed 30% of the ordinary shares on deposit in the aggregate, but such limit may be changed or disregarded from time to time as the
depositary deems appropriate) and imposes a number of conditions on such transactions (e.g., the need to receive collateral, the type of collateral required,
the representations required from brokers, etc.). The depositary may retain the compensation received from the pre-release transactions.

Taxes

Holders are responsible for the taxes and other governmental charges payable on the ADSs and the securities represented by the ADSs. We, the depositary
and the custodian may deduct from any distribution the taxes and governmental charges

Exhibit 2.4

payable by holders and may sell any and all property on deposit to pay the taxes and governmental charges payable by holders.

The depositary may refuse to issue ADSs, to deliver, transfer, split and combine ADRs or to release securities on deposit until all taxes and charges are paid
by the applicable holder. The depositary and the custodian may take reasonable administrative actions to obtain tax refunds and reduced tax withholding for
any distributions on behalf of the holders. However, holders may be required to provide to the depositary and to the custodian proof of taxpayer status and
residence and such other information as the depositary and the custodian may require to fulfill legal obligations. Holders are required to indemnify us, the
depositary and the custodian for any claims with respect to taxes based on any tax benefit obtained for holders.

Foreign Currency Conversion

The depositary will arrange for the conversion of all foreign currency received into U.S. dollars if such conversion is practical, and it will distribute the
U.S. dollars in accordance with the terms of the Deposit Agreement. Holders may have to pay fees and expenses incurred in converting foreign currency,
such as fees and expenses incurred in complying with currency exchange controls and other governmental requirements.

If the conversion of foreign currency is not practical or lawful, or if any required approvals are denied or not obtainable at a reasonable cost or within a
reasonable period, the depositary may take the following actions in its discretion:

•

•
•

Convert the foreign currency to the extent practical and lawful and distribute the U.S. dollars to the holders for whom the conversion and
distribution is lawful and practical.
Distribute the foreign currency to holders for whom the distribution is lawful and practical.
Hold the foreign currency (without liability for interest) for the applicable holders.

Governing Law/Waiver of Jury Trial

The Deposit Agreement and the ADRs will be interpreted in accordance with the laws of the State of New York. The rights of holders of ordinary shares
(including ordinary shares represented by ADSs) are governed by the laws of England and Wales.

The Deposit Agreement provides that, to the fullest extent permitted by law, ADS holders waive the right to a jury trial in any legal proceeding against us
or the depositary arising out of or relating to the ADSs, the Deposit Agreement and any transactions contemplated therein. If we or the depositary opposed
a jury trial demand based on the waiver, the court would determine whether the waiver was enforceable in the facts and circumstances of that case in
accordance with applicable case law.

CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

LICENCE Agreement
As amended on 15 October 2020

(1)    AUTOLUS LIMITED

(2)    UCL BUSINESS LTD

CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

Contents

Clause    Page

1.    DEFINITIONS AND INTERPRETATION    2

2.    LICENCE GRANT    22

3.    SUB-LICENSING    25

4.    RETAINED RIGHTS, ACADEMIC RESEARCH & RESTRICTIONS    27

5.    ACCESS TO IMPROVEMENTS AND NEW INVENTIONS    33

6.    ACCESS TO [***] PROGRAM    35

7.    ACCESS TO [***]    37

8.    INFORMATION AND ACCESS TO MARTIN PULE’S OTHER PROGRAMS & OTHER RESTRICTIONS    38

9.    MATERIALS TRANSFER AND ENABLEMENT OF THE LICENSED RIGHTS    40

10.    OPTION TO ACQUIRE PROGRAM IP    41

11.    DILIGENCE OBLIGATIONS    42

12.    UCLB MANAGEMENT FEE AND SHARES    46

13.    MILESTONE PAYMENTS    47

14.    ROYALTIES    51

15.    SUB-LICENSEE PAYMENTS    55

16.    REPORTING AND PAYMENT PROVISIONS    58

17.    BUY-OUT OPTION    61

18.    INTELLECTUAL PROPERTY PROSECUTION AND MAINTENANCE    61

19.    INTELLECTUAL PROPERTY ENFORCEMENT    64

20.    CONFIDENTIALITY    65

21.    WARRANTIES AND COVENANTS    67

22.    LIMITATION OF LIABILITY    67

23.    INDEMNITY AND INSURANCE    68

24.    TERMINATION    69

25.    CONSEQUENCES OF TERMINATION    70

26.    FORCE MAJEURE    72

    1

CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

27.    FURTHER ASSURANCE    72

28.    PUBLICITY    72

29.    ASSIGNMENT    73

30.    NOTICES    73

31.    MISCELLANEOUS PROVISIONS    74

32.    COUNTERPARTS    75

33.    DISPUTE RESOLUTION, GOVERNING LAW AND JURISDICTION    75

Schedule 1: The Programs    77

Schedule 2: Program IP    87

Schedule 3: UCL Background    172

Part A : UCL Know-How    172

Part B : UCL Background Materials    178

Schedule 4: Manufacturing Know-How    183

Schedule 5: Disclosure Process    185

Schedule 6    186

Part A : Existing Licences    186

Part B : Commercial Agreements    188

Schedule 7: Permitted Studies    189

Schedule 8    191

Part A : Net Sales Definition    191

Part B : Net Receipts Definition    193

Part C : Expert Procedure    194

Schedule 9: Warranties and Covenants    195

Part A : Mutual Warranties & Representations    195

Part B : UCLB Warranties & Covenants at the Effective Date    195

Part C : UCLB Warranties & Covenants at the Amendment Date    196

PART d : uclb warranties & covenants at the second amendment date    197

PART E : uclb warranties & covenants at the THIRD amendment date    198

Schedule 10: Royalty and Net Receipts Statements    200

Schedule 11: Release of Patient Clinical Data    201

    2

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

Schedule 12: CD19 SEQUENCE    203

Schedule 13    204

Schedule 14: ASSIGNED PATENTS    205

    3

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

THIS AGREEMENT  was  made  as  of  25  September  2014  (the  “Effective Date”),  was  amended  as  of  2  March  2016  (the  “Amendment
Date”), was amended and restated as of 28 March 2018 (the “Second Amendment Date”), and is hereby amended and restated as of 15
October 2020 (the “Third Amendment Date”)
BY AND BETWEEN:
(1)

AUTOLUS Limited, a company duly organised and validly existing under the laws of England (company number 09115837) with its
registered office at Forest House, 58 Wood Lane, London, W12 7RZ (“Autolus”); and
UCL  BUSINESS  LTD,  a  public  company  duly  organised  and  validly  existing  under  the  laws  of  England  (company  number
02776963),  previously  named  UCL  Business  Plc,  with  its  registered  office  at  The  Network  Building,  97  Tottenham  Court  Road,
London, W1T 4TP (“UCLB”).

(2)

WHEREAS:
(A)

Autolus  has  been  established  for  the  purposes  of  exploitation  of  certain  of  the  Technology  (as  defined  hereunder)  in  the  field  of
cancer therapy and diagnosis;
UCLB is a public company established by UCL to assist in the commercialisation of technology arising from UCL’s faculties and is
responsible for technology development and commercialisation transactions for UCL;
Dr  Martin  Pule  is  a  leading  academic  and  expert  in  the  development  of  T-cell  therapies  for  the  treatment  of  cancer  and  is  an
employee  and  academic  at  UCL  and  leads  and  supervises  the  MP  Laboratory  (as  defined  below),  and  is  also  the  Senior  Vice
President and Chief Scientific Officer of Autolus;
Through his own research and the research undertaken at the MP Laboratory, certain inventions, discoveries and know-how have
been developed concerning the modification and utilisation of T-cells for cancer therapy, in particular focusing on the identification
and development of certain chimeric antigen receptors that are engineered into human T-cells;
Pursuant  to  his  employment  conditions,  and  the  conditions  of  employment  or  studentship  existing  amongst  those  who  work  or
collaborate  in  the  MP  Laboratory  and  UCL’s  governance,  all  Intellectual  Property  generated  by  MP  or  by,  at  or  within  the  MP
Laboratory  (irrespective  of  the  individual  or  their  status  within  the  MP  Laboratory)  are  initially  owned  by  UCL  and  UCLB  has  the
automatic exclusive right to assign and/or license all such Intellectual Property by virtue of its arrangements with UCL;
MP has assigned all right to the technology to be licensed and/or assigned hereunder to UCLB;
Autolus was granted a licence to the Original Programs with effect from the Effective Date and now with effect from the Amendment
Date the Additional Programs (except for the TRBC2 Dx Program) are to be added to this Agreement;
Autolus is granted rights to the Technology, including an option to acquire certain of the Technology, in each case upon the terms of
this Agreement, and UCLB wishes to grant such rights to Autolus and does so with the consent and support of UCL and MP;
Autolus  obtained  rights  and  licenses  to  certain  additional  Technology  relating  to  the  CAT19  Program  with  effect  from  the  Second
Amendment Date pursuant to the terms of this Agreement;
Autolus is further interested in obtaining rights and licenses to certain additional Technology relating to the TRBC2 Dx Program with
effect from the Third Amendment Date pursuant to the terms of this Agreement; and
Autolus is further interested in obtaining rights and licenses to certain additional Technology relating to the CAROUSEL Study with
effect from the Third Amendment Date pursuant to the terms of this Agreement.

(B)

(C)

(D)

(E)

(F)
(G)

(H)

(I)

(J)

(K)

NOW,  THEREFORE,  the  Parties,  in  consideration  of  the  mutual  covenants  and  undertakings  herein  and  for  other  good  and  valuable
consideration, intending to be legally bound, HEREBY AGREE as follows:
1.
a.

In  this  Agreement,  each  of  the  capitalised  words  and  expressions  set  out  below  shall  have  the  meanings  set  forth  against  that

DEFINITIONS AND INTERPRETATION

capitalised word or expression, unless expressly provided otherwise:
“Academic Information” has the meaning set out in Clause 4.5;

    <#>

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

“Academic Collaborator(s)”  means  any  Academic  Organisation  which  is  actively  collaborating  with  UCL  or  CRUK  on  Academic
Research permitted pursuant to Clause 4;
“Academic Organisation” means an organisation engaged in the conduct of academic research or the non-commercial funding of
academic  research,  comprising  academic  institutions,  charities,  non-for-profit  organisations  and  government  bodies  including  the
national health service and equivalent organisations anywhere in the world;
“Academic Research”  means  academic  research  which  is  undertaken  by  UCL  alone  or  in  collaboration  with  another  Academic
Organisation and, without limiting the foregoing, excluding any Commercial Research;
“Academic Reports” has the meaning set out in Clause 4.5.2;
“Academic Rights” has the meaning set out in Clause 4.1;
“Additional LG Milestone Payment” has the meaning set out in Clause 13.3;
“Additional Licensed Patents” means the ccCAR Patent Rights, iCAR Patent Rights, Epitope Tag Patent Rights, Retrostim Patent
Rights, RapaiCASP9 Patents Rights, TetCAR Patent Rights and ZAP-CAR Patent Rights;
“Additional Milestone”  means  the  achievement  of  a  milestone  triggering  an  Additional  LG  Milestone  Payment  or  Additional  ZC
Milestone Payment in accordance with Clause 13.3 or Clause 13.4 respectively;

“Additional  Program”  means  one  of  the  ccCAR  Program,  iCAR  Program,  Epitope  Tag  Program,  Retrostim  Program,
RapaiCASP9 Program, TetCAR Program, the ZAP-CAR Program, or TRBC2 Dx Program, and “Additional Programs” means any
combination of two or more of the foregoing, as the context requires;

“Additional  Program  IP”  means  the  ccCAR  Program  IP,  iCAR  Program  IP.  Epitope  Tag  Program  IP,  Retrostim  Program  IP,

RapaiCASP9 Program IP, TetCAR Program IP, the ZAP-CAR Program IP and TRBC2 Dx Program IP;

“Additional  Program  Licence”  means  any  one  of  the  ccCAR  Licence,  iCAR  Licence,  Epitope  Tag  Licence,  Retrostim  Licence,
RapaiCASP9 Licence, TetCAR Licence, the ZAP-CAR Licence, and the TRBC2 Dx Licence and “Additional Program Licences” means any
two or more of the foregoing;

“Additional  Royalty  Product”  means  any  one  of  the  ccCAR  Product,  iCAR  Product,  Epitope  Tag  Product,  Retrostim  Product,
RapaiCASP9 Product, TetCAR Product, or ZAP-CAR Product, and the TRBC2 Dx Product and “Additional Royalty Products” means any
two or more of the foregoing;

“Additional ZC Milestone Payment” has the meaning set out in Clause 13.4;
“Affiliate” means any entity that directly or indirectly controls, is controlled by, or is under common control with a Party, for so long
as such control exists. For the purposes of this definition of Affiliate and the definition of Tobacco Party, “control” and “controlled”
means either (a) with respect to any person or entity, ownership directly or indirectly of more than fifty (50%) per cent of the shares of
stock entitled to vote for the election of directors, in the case of a company or corporation, or more than fifty (50%) per cent of equity
interest in the case of any other type of legal entity, status as a general partner in any partnership, or any other arrangement whereby
a person controls or has the right to control the board of directors or equivalent governing body of the relevant entity, or the ability
generally to cause the direction of the management or policies of an entity. In the case of certain entities organised under the laws of
certain countries, where the maximum percentage ownership permitted by law for a foreign investor is less than fifty (50%) per cent,
in such case such lower percentage shall be substituted in the preceding sentence provided that such foreign investor has the power
to  direct  the  management  and  policies  of  such  entity.  For  the  purposes  of  this  Agreement  (i)  UCL  shall  be  deemed  an  Affiliate  of
UCLB and vice versa; and (ii) Autolus’ Affiliates shall be limited to its subsidiaries (as defined in section 1159 of the Companies Act
2006) from time to time;
“Agreement” means this agreement together with its schedules, each as may be amended from time to time in accordance with the
terms of this Agreement;

    1

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

“ALLCAR19  Study”  means  the  Permitted  Study  described  in  Schedule  1  under  the  heading  CAT19  Program  and  conducted  in
accordance with the [***] Contract under UCL’s sponsorship;
“Amendment Date” has the meaning set out in the initial paragraph of this Agreement;
“Antigen” means any protein or other molecular structure including for the avoidance of doubt a cell surface sugar or part thereof
(including  any  polypeptide  or  fragment  thereof)  whether  glycosylated  or  otherwise  and  is  capable  of  or  demonstrates  any  binding
affinity with any antibody, antigen-binding domain or other engager protein or any fragment of any of the foregoing;
“Assigned Patents” has the meaning set out in Clause 10.1;
“Assignee Entity” means any third party to which Autolus assigns an Assigned Patent together with any successor in title to such
Assigned Patent and any licensee or sub-licensee of such third party or its successors under such Assigned Patent;
“Autolus Improvement” means to the extent Controlled by Autolus (free from any restriction or encumbrance) (i) any improvement
to the applicable Licensed Patent(s) in respect of which a Licence has been terminated by UCLB, where use or deployment of such
improvement in a product would infringe the applicable Licensed Patent(s) for which such Licence has been terminated; and, (ii) the
Test and Regulatory Data which has been generated by or on behalf of Autolus in its Exploitation of the applicable Licensed Patents
for which the Licence has been terminated by UCLB;
“Background Licence” has the meaning set out in Clause 2.1;
“Background Materials” means (i) those materials listed in Part B of Schedule 3; and (ii) any materials Controlled by UCLB (free
from any restriction or encumbrance) which are not Program Materials and which (a) have been used in connection with any of the
Programs  prior  to  the  Effective  Date;  and/or  (b)  are  used  in  connection  with  the  [***]  Program,  NSG  Program  and/or  the  ZipCAR
Program  after  the  Effective  Date;  and/or  (c)  have  been  used  in  connection  with  any  of  the  Additional  Programs  prior  to  the
Amendment  Date  or  the  CAT19  Program  prior  to  the  Second  Amendment  Date  or  the  TRBC2  Dx  Program  on  or  after  the
Amendment Date but prior to the Third Amendment Date;
“BCMA”  means  the  specific  Antigen  (or  part  thereof)  coded  for  by  reference  to  the  sequence  defined  in  Schedule  1  under  the
BCMA-CAR Program;
“[***]” means [***] in which one of the [***] intended to [***] on the [***] is capable of binding to [***];
“[***] Option” means Autolus’s rights under Clause 7;
“BCMA Field” means all uses without restriction except for use in [***];
“BCMA Licence” has the meaning set out in Clause 2.2.2;
“BCMA  Product”  means  any  product  or  therapy  which  within  the  BCMA  Field  (i)  targets  BCMA  and  is  covered  by,  uses,
incorporates or has been developed using any of the BCMA Program IP; or (ii) targets BCMA and has been developed using any of
the  UCL  Background  IP  or  Manufacturing  Know-How;  or  (iii)  were  it  not  for  the  license  to  BCMA  Program  IP  hereunder,  would
otherwise infringe any Patent Right licensed hereunder under the BCMA Program IP or, were it not for the assignment of the BCMA-
CAR  Patent  Rights  to  Autolus  (or  subsequent  assignment  or  licence  grant  to  an  Assignee  Entity)  would  otherwise  infringe  such
assigned BCMA-CAR Patent Rights;
“[***]  Existing  Patent”  means  patent  application  [***]  and  all  Patent  Rights  derived  therefrom  including  International  Patent
Application No. [***];
“[***] Period” has the meaning set out in Clause 7.1;
“[***] Program” means the research conducted with respect to [***] by MP and/or the MP Laboratory, as of the Effective Date and
thereafter during the [***] Period;
“[***]  Program  IP”  means,  excluding  Program  IP,  UCL  Background  IP  and  Manufacturing  Know-How,  (i)  all  Know-How  and
inventions generated, created or developed pursuant to the [***] Program excluding CGK as of the licence grant date (in respect of
the [***] Program IP); and (ii) the [***] Existing Patent and all Patent Rights filed in respect of the research undertaken pursuant to the
[***] Program from time to time (“[***] Patent Rights”);
“BCMA-CAR Program” means the program of research defined in Schedule 1 under the title BCMA-CAR Program as conducted
prior to the Effective Date;

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

“BCMA Program IP” means, excluding Manufacturing Know-How, (i) those patent applications listed in Schedule 2 under the title
BCMA-CAR  Patents  and  all  Patent  Rights  derived  therefrom  (“BCMA-CAR  Patent  Rights”);  (ii)  the  Know-How  described  in
Schedule  2  under  the  title  BCMA-CAR  Know-How;  (iii)  the  materials  listed  in  Schedule  2  under  the  title  BCMA-CAR  Materials
together with all IP in the same; and (iv) all Know-How and materials (together with all IP in the same) developed by MP and/or the
MP  Laboratory  pursuant  specifically  to  the  BCMA-CAR  Program  prior  to  the  Effective  Date,  including  the  technology  claimed  or
disclosed  in  the  BCMA-CAR  Patent  Rights  as  of  the  Effective  Date,  other  than  that  which  is  CGK  as  of  the  Effective  Date  or
disclosed in the sections of the BCMA-CAR Patent Rights that refer to, describe or disclose prior art including CGK existing as of the
Effective Date;
“BiTE” means a bi-specific T-cell engager protein comprising two or more antigen-binding domains which bind a T-cell and a target
cell thereby bringing the two cells into close proximity intended to modulate or trigger an immune response against the target cell,
and for the avoidance of doubt is not a CAR;
“Buy-Out Option” has the meaning set out in Clause 17.1;
“CAROUSEL Clinical Study Agreement” means the [***] entered into between UCL and [***];
“CAROUSEL  Clinical  Study  Results”  means  all  data,  information,  know-how  and  other  intellectual  property  and  results  arising
from the CAROUSEL Study including the protocol for the CAROUSEL Study and the Test and Regulatory Data resulting from the
CAROUSEL Study;
“CAROUSEL Patient Clinical Data”  means  patient  clinical  data  generated  in  the  performance  of  the  CAROUSEL  Study  after  the
Third Amendment Date;
“CAROUSEL Program IP” means, excluding Manufacturing Know-How, (i) the Know-How described in Schedule 2 under the title
CAROUSEL Know-How, including CAROUSEL Patient Clinical Data generated prior to the Third Amendment Date once the same is
disclosed to Autolus in accordance with Clause 9.9; and (ii) any Know-How generated in the performance of the CAROUSEL Study
after the Third Amendment Date, which shall include CAROUSEL Patient Clinical Data once disclosed to Autolus in accordance with
Clause 9.9;
“CAROUSEL Study” means the Permitted Study described in Schedule 7 under the heading CAT19 Studies with the study acronym
“CAROUSEL”;
“CARPALL and ALLCAR19 Patient Clinical Data” means  patient  clinical  data  generated  from  either  the  CARPALL  Study  or  the
ALLCAR19  Study  and  (i)  described  in  Schedule  2  under  the  title  CAT19  Know-How,  with  the  heading  Patient  Clinical  Data  or  (ii)
which is otherwise generated after the Second Amendment Date but excluding [***];
“CARPALL Study” means the Permitted Study described in Schedule 1 under the heading CAT19 Program and conducted under
UCL’s  sponsorship  with  funding  from  (i)  Children  with  Cancer  UK,  (ii)  Great  Ormond  St  Hospital  Children’s  Charity  and  (iii)  J  P
Moulton Charitable Foundation;
“CAT19 1 Gen Product”  means  the  CAT19  Product  developed  in  the  CAT19  Program  (the  “Original  CAT19  Product”),  and  any
other  CAT19  Product  as  may  be  modified  by  Autolus  or  any  Sub-Licensee  after  the  Second  Amendment  Date  that  is  developed
using  any  of  the  CARPALL  and  ALLCAR19  Patient  Clinical  Data  either  in  a  Phase  2  study  following  directly  a  clinical  study
conducted under the CAT19 Program, or pursuant to a bridging study or similar adjunctive study intended to demonstrate that the
modified CAT19 Product is equivalent to the Original CAT19 Product, but excluding any CAT19 CNS Product;
“CAT19 Binder Product” means a CAT19 Product that is not a CAT19 1  Gen Product or a CAT19 CNS Product;
“CAT19 CNS Licence” has the meaning set out in Clause 2.2.18;
“CAT19  CNS  Product”  means  a  CAT19  Product  for  the  treatment  of  Primary  CNS  Lymphoma  which  is  covered  by,  uses,
incorporates or has been developed using any of the CAROUSEL Program IP;
“CAT19 Licence” has the meaning set out in Clause 2.2.16;
“CAT19 Product” means any product or therapy which (i) targets the CD19 Antigen and is covered by, uses, incorporates or has
been developed using any of the CAT19 Program IP; or (ii) were it not for the license to CAT19 Program IP hereunder, or the

st 

st

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

assignment  to  Autolus  (or  subsequent  assignment  or  licence  grant  to  an  Assignee  Entity)  of  the  CAT19  Patent  Rights,  would
otherwise infringe any Patent Right licensed under the CAT19 Program IP;
“CAT19 Program” means the program of research defined in Schedule 1 under the title CAT19 Program as conducted prior to the
Second Amendment Date;
“CAT19 Program IP” means excluding Manufacturing Know-How, (i) those patent applications listed in Schedule 2 under the title
CAT19 Patents and all Patent Rights derived therefrom (“CAT19 Patent Rights”); (ii) the Know-How described in Schedule 2 under
the  title  CAT19  Know-How,  including  CARPALL  and  ALLCAR19  Patient  Clinical  Data  generated  prior  to  the  Second  Amendment
Date  once  the  same  is  disclosed  to  Autolus  in  accordance  with  Clause  9.8;  (iii)  the  materials  listed  in  Schedule  2  under  the  title
CAT19  Materials  together  with  all  IP  in  the  same;  and  (iv)  any  Know-How  generated  in  the  performance  of  the  ALLCAR19  and
CARPALL  Studies  after  the  Second  Amendment  Date,  which  shall  include  CARPALL  and  ALLCAR19  Patient  Clinical  Data  once
disclosed to Autolus in accordance with Clause 9.8, but excluding any such Know-How in which Autolus has rights pursuant to any
agreement  entered  into  in  accordance  with  Clause  4.12  and  excluding  any  Know-How  owned  by  [***]  pursuant  to  the  [***]
Agreements;
“CAR”  means  a  chimeric  antigen  receptor  (also  known  as  chimeric  immunoreceptors  or  artificial  T-Cell  receptors)  comprising  an
extra-cellular  Antigen  binding  or  recognition  domain  and  one  or  more  intra-cellular  signalling  domains,  wherein  the  extra-cellular
Antigen binding or recognition domain has binding specificity for one particular Antigen, and for the avoidance of doubt is not a BiTE;
“ccCAR Licence” has the meaning set out in Clause 2.2.7;
“ccCAR Product” means any product or therapy which, were it not for the licence to ccCAR Patent Rights granted hereunder or the
assignement  to  Autolus  (or  subsequent  assignment  or  licence  grant  to  an  Assignee  Entity)  of  the  ccCAR  Patent  Rights,  would
otherwise infringe the ccCAR Patent Rights in the relevant country of sale;
“ccCAR Program” means the program of research defined in Schedule 1 under the title ccCAR Program as conducted prior to the
Amendment Date;
“ccCAR Program IP” means excluding Manufacturing Know-How, (i) those patent applications listed in Schedule 2 under the title
ccCAR Patents and all Patent Rights derived therefrom (“ccCAR Patent Rights”); (ii) the Know-How described in Schedule 2 under
the title ccCAR Know-How; and (iii) the materials listed in Schedule 2 under the title ccCAR Materials together with all IP in the same;
“CDA” the confidentiality agreement executed between [***];
“CD19 Antigen” means the specific Antigen (or part thereof) coded for by reference to the sequence set out in Schedule 12;
“CD19 Clinical Study Results” means all data, information, know-how and other intellectual property and results arising from the
ALLCAR19 Study or the CARPALL Study including the protocol for the ALLCAR19 Study and for the CARPALL Study and the Test
and Regulatory Data resulting from the ALLCAR19 Study and from the CARPALL Study;
“CD19 Clinical Study NHS Foundation Trusts” means UCL Hospitals NHS Foundation Trust, The Christie NHS Foundation Trust,
Oxford University Hospitals NHS Foundation Trust, Great Ormond Street Hospital for Children NHS Foundation Trust, Manchester
University NHS Foundation Trust and such other NHS Trusts as may act as study sites in respect of the ALLCAR19 Study and/or the
CARPALL Study from time to time;
“CD19  Clinical  Study  Agreements”  means  (i)  the  [***]  Contract,  (ii)  the  [***]  Grant,  (iii)  the  [***]  Agreements  and  (iv)  the
agreements  entered  into  between  UCL  and  [***]  in  respect  of  their  conduct  of  the  CARPALL  Study  or  the  ALLCAR19  Study,  as
applicable;
“CD19 Field” means all uses without restriction;
“[***]” means the specific [***] defined in Schedule 1 under the [***] Program;
“[***] Exercise Period” has the meaning set out in Clause 2.4;
“[***] Licence” has the meaning set out in Clause 2.5;
“[***] Licence Notice” has the meaning set out in Clause 2.4;
“[***] Option” means Autolus’s rights under Clauses 2.4 and 2.5;
“[***] Product” means any product or therapy which (i) targets [***] and is covered by, uses, incorporates or has been developed
using any of the [***] Program IP; or (ii) targets [***] and has been developed using any of the UCL Background IP or Manufacturing

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

Know-How;  or  (iii)  were  it  not  for  the  license  to  [***]  Program  IP  hereunder,  would  otherwise  infringe  any  Patent  Right  licensed
hereunder under the [***] Program IP;
“[***] Program” means the program of research defined in Schedule 1 under the title [***] Program, conducted prior to the Effective
Date together with the program of research conducted after the Effective Date as described in Schedule 1 under the title (B): [***]
Project Plan;
“[***] Program IP” means, excluding Manufacturing Know-How, (i) the Know-How described in Schedule 2 under the title [***] Know-
How;  (ii)  the  materials  listed  in  Schedule  2  under  the  title  [***]  Materials  together  with  all  IP  in  the  same;  (iii)  all  Know-How  and
materials (together with all IP in the same) developed by MP and/or the MP Laboratory pursuant specifically to the [***] Program prior
to the Effective Date other than that which is CGK as of the date the [***] Option is exercised; (iv) any Patent Rights filed by or on
behalf of UCLB from time to time in respect of any work undertaken by MP and/or the MP Laboratory prior to, on or after the Effective
Date under or pursuant to the agreed project plan for the [***] Program as set out in Schedule 1 (“[***] Patent Rights”); and (v) all
Know-How  and  materials  developed  after  the  Effective  Date  by  MP  and/or  the  MP  Laboratory  pursuant  specifically  to  the  agreed
project plan for the [***] Program as set out in Schedule 1, including the technology claimed or disclosed in the [***] Patent Rights (if
any) as of the date the [***] Option is exercised, other than that which is CGK as of the date the [***] Option is exercised, or disclosed
in  the  sections  of  the  [***]  Patent  Rights  that  refer  to,  describe  or  disclose  prior  art  including  CGK  existing  as  of  the  date  the  [***]
Option is exercised; references to CGK in this definition shall exclude CGK that became CGK through disclosure after the Effective
Date  by  publication  of  the  [***]  Patent  Rights  or  by  MP  making  a  publication  in  accordance  with  this  Agreement  or  by  Autolus’s
publication;
“CGK”  or  “Common  General  Knowledge”  means  all  technical  and  other  information,  knowledge,  ideas,  inventions,  concepts,
discoveries, data, designs, know-how, formulae, methods, sequences, models, procedures, designs, protocols, trials, tests, methods,
processes, specifications and techniques which are not confidential and are publicly available at the applicable time;
“Collaborative Research Project” has the meaning set out in Clause 4.7;
“Combination Product” has the meaning set out in paragraph 4 of Part A of Schedule 8;
“Commercial Licence” has the meaning set out in Clause 3.3;
“Commercial Research” means any research (i) that is, in whole or part, funded by a person or entity that is not a Funder; or (ii)
that is undertaken at the request of or for the benefit of any entity that is not an Academic Organisation involved in such research; or
(iii) that is undertaken (as opposed to funded) in collaboration with any entity which is not an Academic Organisation; or (iv) under
which a Third Party, which is not an Academic Organisation participating in such research, will acquire any rights in (including by way
of assignment or licence) or control over the results of such research;
“Competitive  Product”  means  any  product  or  therapy  that  is  competitive  to  or  equivalent  (whether  structurally,  functionally  or
through mechanism of action) to any Royalty Product, including any product or therapy that may be considered a generic, biosimilar
and/or a “me-too” product or therapy or otherwise infringes any of the Licensed Patents or makes use of any of the Technology;
“Complementary  Diagnostic  Product”  means  a  Royalty  Product  that  is  not  directly  of  therapeutic  application  and  is  used  for
diagnosis, assessment or detection;
“Competing Entrant” has the meaning set out in Clause 14.11;
“Confidential Information” has the meaning set out in Clause 20.1;
“Control” or “Controlled” in respect of a Party, that Intellectual Property (or those materials) which such Party (which in the case of
UCLB means UCLB or UCL) (i) owns and is able, without breaching any obligation owed by it or (by rule of law) having to obtain the
consent of any co-owner, to license (or in the case of materials supply) the same to a third party (including a Party to this Agreement)
for the applicable Royalty Product or use; or (ii) is licensed to use (other than by the other Party to this Agreement) and is entitled
under the terms of the licence to which they are a party to sub-license (or in the case of materials supply) the same to a third party
(including a Party to this Agreement) for the applicable Royalty Product or use without having to obtain consent from such third party;

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

“Core Countries” means [***] and [***] and “Core Country” means any one of them;
“Cover”, “Covering” or “Covered” means (for the purposes of Clauses 13.2.1, 13.2.2, 14.4 and paragraph 1.8 of Part B Schedule
9, paragraph 1.7 of Part C Schedule 9 and paragraph 1.7 of Part D Schedule 9) in the case of a product or a therapy that (i) such
product  or  therapy  would,  were  it  not  for  the  applicable  licence  granted  and  subsisting  hereunder,  infringe  the  applicable  Patent
Rights  so  licensed  hereunder;  or  (ii)  in  the  case  of  the  Assigned  Patents  such  product  or  therapy  would,  were  it  not  for  the
assignment  to  Autolus  (or  subsequent  assignment  or  licence  grant  to  an  Assignee  Entity)  of  such  Assigned  Patent,  infringe  the
applicable Patent Rights so assigned;
“CRUK” means Cancer Research UK (registered as a company limited by guarantee in England and Wales under number 4325234)
and registered charity (number 1089464 in England and Wales and registered in Scotland under number SC041666 and in the Isle of
Man under number 1103);
“CRUK Funders” means Cancer Research Technology Limited, with company number 01626049 (“CRT”), CRUK, Leukaemia and
Lymphoma  Research  (registered  as  a  company  limited  by  guarantee  in  England  and  Wales  under  number  738089  and  registered
charity  number  216032  in  England  and  Wales  and  registered  in  Scotland  under  number  SC037529)  (“LLR”)  and  Kay  Kendall
Leukaemia Fund (a registered charity in England and Wales with number 290772) (“KKLF”);
“CRUK Indemnified Parties” has the meaning set out in Clause 23.1.1;
“CRUK Study” has the meaning set out in Clause 11.1.1(ii);
“Defaulting Party” has the meaning set out in Clause 24.4;
“Developing Country” or “Developing Countries” means those countries that are: (a) eligible for support from [***]; and (b) to the
extent not included in (a), defined as at the Second Amendment Date by the [***], as such definitions may be amended from time to
time by the World Bank:
“Direct Licence” has the meaning set out in Clause 29.6;
“Disclosing Party” has the meaning set out in Clause 20.1;
“Disclosure Notification” has the meaning in Clause 5.2;
“Enforcement Action” has the meaning set out in Clause 19.2;
“Epitope Tag Field” means all uses and applications without restriction;
“Epitope Tag Licence” has the meaning set out in Clause 2.2.12;
“Epitope  Tag  Product”  means  any  product  or  therapy  within  the  Epitope  Tag  Field:  (i)  which,  were  it  not  for  the  licence  granted
hereunder or the assignment to Autolus (or subsequent assignment or licence grant to an Assignee Entity) of the Epitope Tag Patent
Rights,  would  infringe  the  Epitope  Tag  Patent  Rights;  and/or  (ii)  the  manufacture  of  which  product  or  therapy  were  it  not  for  the
licence  granted  hereunder  or  the  assignment  to  Autolus  (or  subsequent  assignment  or  licence  grant  to  an  Assignee  Entity)  of  the
Epitope Tag Patent Rights, would infringe the Epitope Tag Patent Rights; and/or (iii) where the manufacture of any materials used in
connection  with  the  manufacture  of  such  product  or  therapy,  were  it  not  for  the  licence  granted  hereunder  or  the  assignment  to
Autolus  (or  subsequent  assignment  or  licence  grant  to  an  Assignee  Entity)  of  the  Epitope  Tag  Patent  Rights,  would  infringe  the
Epitope Tag Patent Rights (including without limitation the manufacture of the viral vector ultimately used for the manufacture of such
product or therapy);
“Epitope Tag Program” means the program of research defined in Schedule 1 under the title Epitope Tag Program as conducted
prior to the Amendment Date;
“Epitope Tag Program IP” means excluding Manufacturing Know-How, (i) those patent applications listed in Schedule 2 under the
title  Epitope  Tag  Patents  and  all  Patent  Rights  derived  therefrom  (“Epitope Tag Patent Rights”);  (ii)  the  Know-How  described  in
Schedule 2 under the title Epitope Tag Know-How; and (iii) the materials listed in Schedule 2 under the title Epitope Tag Materials
together with all IP in the same;
“Existing Licences” has the meaning set out in Clause 4.9;
“Exploit” and “Exploiting” means to make, have made, import, export, use, sell or offer for sale, including to research, experiment,
develop, commercialise, obtain and maintain Regulatory Approvals, manufacture, to have manufactured, hold or keep (whether for
disposal  or  otherwise),  have  used,  export,  transport,  distribute,  promote,  market  or  have  sold  or  otherwise  dispose  of,  and
“Exploitation” shall mean the act of Exploiting;
“Extended Period” has the meaning set out in Clause 7.1;

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

“Field” means all uses and applications without restriction;
“Final Written Report” means with respect to the applicable Program, the final written report approved by MP and written following
completion of the applicable Program in accordance with the applicable project plan, and which shall provide a complete report of the
results, findings and activities arising from that Program;
“First Commercial Sale” means the first arm’s length commercial sale of the applicable Royalty Product in a country by Autolus or
by a Sub-Licensee or by an Assignee Entity pursuant to this Agreement, in each case following the grant of a Marketing Approval for
the applicable Royalty Product in such country;
“Funder”  means  an  academic,  charitable  or  other  not-for-profit  organisation  including  academic  institutions,  charities,  and
government bodies;
“GD2” means the specific Antigen (or part thereof) coded for by reference to the sequence defined in Schedule 1 under the GD2
Program;
“GD2 Clinical Study Results”  means  all  data,  information,  know-how  and  other  intellectual  property  and  results  arising  from  the
CRUK Study including the protocol for the CRUK Study and the Test and Regulatory Data resulting from the CRUK Study;
“GD2 Diligence Obligation” has the meaning set out in Clause 11.1.1;
“GD2 Licence” has the meaning set out in Clause 2.2.3;
“GD2 Product” means any product or therapy which (i) targets GD2 and is covered by, uses, incorporates or has been developed
using any of the GD2 Program IP; or (ii) targets GD2 and has been developed using any of the UCL Background IP or Manufacturing
Know-How; or (iii) were it not for the license to GD2 Program IP hereunder, would otherwise infringe any Patent Right licensed under
the GD2 Program IP or, were it not for the assignment to Autolus (or subsequent assignment or licence grant to an Assignee Entity)
of the GD2 Patent Rights, would otherwise infringe such assigned GD2 Patent Rights;
“GD2  Program”  means  the  program  of  research  defined  in  Schedule  1  under  the  title  GD2  Program  as  conducted  prior  to  the
Effective Date;
“[***] Period” has the meaning set out in Clause 6.1;
“[***] Program” means the program of research concerning [***] being undertaken by [***] using the [***] as is used in the [***] and
any  other  research  undertaken  by  [***]  with  respect  to  [***]  in  collaboration  with  [***]  in  each  case  up  to  the  Effective  Date  and
thereafter during the [***] Period;
“[***]  Program  IP”  means,  excluding  Program  IP,  UCL  Background  IP  and  Manufacturing  Know-How,  (i)  all  Know-How  and
inventions  generated,  created  or  developed  pursuant  to  the  [***]  Program  excluding  CGK  as  of  the  licence  grant  date,  and  (ii)  all
Patent Rights filed in respect of the research undertaken pursuant to the [***] Program;
“GD2 Program IP” means, excluding Manufacturing Know-How, (i) those patent applications listed in Schedule 2 under the title GD2
Patents and all Patent Rights derived therefrom (“GD2 Patent Rights”); (ii) the Know-How described in Schedule 2 under the title
GD2  Know-How;  (iii)  the  materials  listed  in  Schedule  2  under  the  title  GD2  Materials  together  with  all  IP  in  the  same;  and  (iv)  all
Know-How and materials (together with all IP in the same) developed by MP and/or the MP Laboratory pursuant specifically to the
GD2 Program prior to the Effective Date including the technology claimed or disclosed in the GD2 Patent Rights as of the Effective
Date,  other  than  that  which  is  CGK  as  of  the  Effective  Date  or  disclosed  in  the  sections  of  the  GD2  Patent  Rights  that  refer  to,
describe or disclose prior art including CGK existing as of the Effective Date;
“[***] Grant” means the grant from [***], and associated grant terms and conditions, dated [***];
“iCAR Licence” has the meaning set out in Clause 2.2.8;
“iCAR Product” means any product or therapy which, were it not for the licence to iCAR Patent Rights granted hereunder or the
assignment to Autolus (or subsequent assignment or licence grant to an Assignee Entity) of the iCAR Patent Rights, would otherwise
infringe the iCAR Patent Rights in the relevant country of sale;
“iCAR Program”  means  the  program  of  research  defined  in  Schedule  1  under  the  title  iCAR  Program  as  conducted  prior  to  the
Amendment Date;
“iCAR  Program  IP”  means  excluding  Manufacturing  Know-How,  (i)  those  patent  applications  listed  in  Schedule  2  under  the  title
iCAR Patents and all Patent Rights derived therefrom (“iCAR Patent Rights”); (ii) the Know-How described in Schedule 2

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

under the title iCAR Know-How; and (iii) the materials listed in Schedule 2 under the title iCAR Materials together with all IP in the
same;
“Improvement” means (a) any improvement, enhancement, development or advancement (together with all Intellectual Property in
the same) over any of the Original Program IP in each case as created, generated or developed by MP (in the course of his activities
or  role  at  or  for  UCL  or  UCLB)  and/or  the  MP  Laboratory,  and  which,  if  used,  deployed  or  incorporated  in  the  Exploitation  of  any
product, process or service would result in such product, process or service being within the scope of or otherwise dependent upon
(that is, in each case, in the absence of any licence, would otherwise infringe) any one or more claims of any of the Original Licensed
Patents; and (b) any CAR developed or generated by MP (in the course of his activities or role at or for UCL or UCLB) or the MP
Laboratory that targets any of GD2, BCMA, TRBC1, TRBC2, the CD19 Antigen and/or [***];
“Improvement  Negotiation  Period”  means  in  respect  of  each  Improvement  or  New  Invention  (as  applicable),  the  period
commencing with the date the applicable Disclosure Notification (complying with Clause 5.2.2) is deemed to have been received by
Autolus and expiring [***] after the date such Disclosure Notification is deemed to have been so received;
“Improvement Period” has the meaning set out in Clause 5.1;
“Indemnity Claim” has the meaning set out in Clause 23.1;
“Indemnified Party” has the meaning set out in Clause 23.1;
“Insolvency Event” means the occurrence of any of the following events or circumstances (or any analogous event or circumstance
in a jurisdiction other than England and Wales) in relation to the relevant entity: (i) being deemed unable to pay its debts as defined
in section 123 Insolvency Act 1986 without any requirement to prove any matter stated in that section to a court, (ii) entering into a
voluntary arrangement or any other composition, scheme or arrangement with (or assignment for the benefit of) its creditors; (iii) the
appointment of a receiver, administrator or insolvency manager over the whole or the majority of its business or assets, and which
appointment is not appealed or set aside within twenty (20) days; (iv) an order is made or a resolution is passed for its winding up
(except  for  the  purposes  of  a  bona  fide  solvent  reorganisation);  (v)  an  order  for  bankruptcy  or  dissolution  or  the  making  of  an
administration order is made which is not appealed or set aside within thirty (30) days of it being made; or (vi) ceasing to carry on
business for any period in excess of thirty (30) days or claiming the benefit of any statutory moratorium;
“Intellectual Property” or “IPR” or “IP” means all Patent Rights, claims in or rights to Patent Rights, rights in designs (including
design patents, registered designs and unregistered designs), copyright, rights in software, database rights, rights in data, inventions,
rights  in  Know-How,  trade  secrets  and  confidential  information,  and  any  and  all  other  similar  or  equivalent  rights  to  any  of  the
foregoing situated in any country in the world, in each case for their full term and any extensions, together with applications for any of
the foregoing, the right to apply for any of the foregoing in any part of the world and the right to claim priority in respect of any of the
foregoing;
“[***] Materials” means the [***] (as listed in Schedule 2 under the title [***] Materials – [***] Materials), complementarity determining
regions (“CDRs”) from such [***], and any derivatives, fragments, modifications, enhancements of such [***] or the CDRs from [***];
“[***]” means [***];
“[***] Agreements” means the agreements entered into between UCL and [***] (i) dated [***] for [***] to manufacture the viral vector
used in the CARPALL Study, (ii) dated [***] for [***] to manufacture the viral vector used in the CARPALL Study, (iii) dated [***] for
[***] to manufacture the viral vector used in the ALLCAR19 Study, and (iv) dated [***] for [***] to manufacture the viral vector used in
the ALLCAR19 Study;
“Know-How” means all technical and other information, knowledge, ideas, concepts, discoveries, data, designs, know-how, trade
secrets,  inventions  (which  at  the  relevant  time  are  not  the  subject  of  a  Patent  Right)  formulae,  methods,  software  sequences,
models,  procedures,  designs  for  experiments,  trials  and  tests  and  results  of  the  same,  testing  methods,  processes,  specifications
and techniques, clinical data and manufacturing data including all Test and Regulatory Data;
“Know-How Period” has the meaning set out in Clause 14.8;

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

“Laboratory Notebooks” means all laboratory notebooks in UCL’s possession, custody or control which emanate from laboratories
at UCL, in so far as they concern or contain information relating to any of the Programs, Program IP, Materials or UCL Background
IP;
“Licence” means one of the BCMA Licence, [***] Licence (if Autolus exercises its rights under Clause 2.4), GD2 Licence, Logic Gate
Licence,  NSG  Licence,  RQR8  Licence,  TRBC1/2  Licence,  ZipCAR  Licence,  ccCAR  Licence,  iCAR  Licence,  Epitope  Tag  Licence,
Retrostim  Licence,  RapaiCASP9  Licence,  TetCAR  Licence,  ZAP-CAR  Licence,  CAT19  Licence,  TRBC2  Dx  Licence,  CAT19  CNS
Licence, Manufacturing Licence, the Background Licence and the licences granted under Clause 2.3;
“Licensed  Patents”  means  the  Original  Licensed  Patents,  the  Additional  Licensed  Patents,  the  CAT19  Patent  Rights  and  the
TRBC2  Dx  Patent  Rights.  For  the  avoidance  of  doubt,  notwithstanding  their  assignment  to  Autolus  pursuant  to  the  Patent
Assignment, or subsequent assignment to an Assignee Entity, the Assigned Patents shall continue to be included within the definition
of Licensed Patents;
“Limb (i)(b) Notice” has the meaning set out in Clause 11.1.1(ii);
“Logic Gate Licence” has the meaning set out in Clause 2.2.4;
“Logic Gate Product” means any product or therapy which is covered by, uses or incorporates any of the Logic Gate Program IP;
“Logic Gate Program” means the program of research defined in Schedule 1 under the title Logic Gate Program as conducted prior
to the Effective Date;
“Logic Gate Program IP” means, excluding Manufacturing Know-How, (i) those patent applications listed in Schedule 2 under the
title  Logic  Gate  Patents  and  all  Patent  Rights  derived  therefrom  (“Logic  Gate  Patent  Rights”);  (ii)  the  Know-How  described  in
Schedule 2 under the title Logic Gate Know-How; (iii) the materials listed in Schedule 2 under the title Logic Gate Materials together
with  all  IP  in  the  same;  and  (iv)  all  Know-How  and  materials  (together  with  all  IP  in  the  same)  developed  by  MP  and/or  the  MP
Laboratory pursuant specifically to the Logic Gate Program prior to the Effective Date including the technology claimed or disclosed
in the Logic Gate Patent Rights filed as of the Effective Date (or the Amendment Date for those Logic Gate Patents Rights filed as of
the Amendment Date), other than that which is CGK as of the Effective Date (or, in respect of those Logic Gate Patent Rights filed as
of the Amendment Date, the Amendment Date) or disclosed in the sections of the Logic Gate Patent Rights that refer to, describe or
disclose prior art including CGK existing as of the Effective Date or Amendment Date, as applicable, provided that to the extent that
any materials in (iii) or (iv) incorporate any of the following binders, CD5, CD19, CD22, CD23, CD33 and/or EGFRvIII, such binders
together  with  IP  in  the  same  are  excluded.  References  to  CGK  in  this  definition  shall  exclude  CGK  that  became  CGK  through
disclosure after the Effective Date by publication of the Logic Gate Patent Rights (filed as of the Effective Date), or by MP making a
publication in accordance with this Agreement or by Autolus’ publication;
“Manufacturing Licence” has the meaning set out in Clause 2.3;
“Manufacturing Know-How” means that Know-How generated by each of [***] prior to the Effective Date, irrespective of its use or
application in any of the Programs, which is required for or beneficial to practice any of the Program IP licensed hereunder, including
that listed in Schedule 4;
“Marketing Approval Application” or “MAA” means an application for a Marketing Approval;
“Marketing Approval”  or  “MA”  means  those  Regulatory  Approval(s)  required  by  applicable  laws  and  regulations  in  a  particular
country or territory in order to sell or commercially supply a medicinal product and/or device in that country or territory;
“Martin Pule” or “MP” means Dr Martin Pule of [***] for so long as he is employed by, consults for or otherwise holds any position at
or undertakes or supervises any research at UCL or UCLB;
“Match Period” has the meaning set out in Clause 5.7;
“Materials” means Program Materials and Background Materials;
“Milestone” means a Success Milestone, an Additional Milestone or a Sales Milestone;
“Milestone  Payment”  means  a  Success  Milestone  Payment,  an  Additional  Milestone  Payment,  a  Sales  Milestone  Payment,  an
Additional LG Milestone Payment or an Additional ZC Milestone Payment;

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

“MP Laboratory” means from time to time prior to, on and following the Effective Date, those members of the research group(s) at
UCL lead by MP, who at the relevant time were or are under the supervision or direction of MP;
“Net Receipts” has the meaning in Part B of Schedule 8;
“Net Sales” has the meaning in Part A of Schedule 8;
“New Invention” means any discovery, invention (whether patentable or not), Know-How or improvement concerning any CAR or
CARs  (including  methods  for  manufacture  of  any  CAR  or  CARs)  which  is  not  an  Improvement  and  which  has  been  discovered,
generated, identified or invented by MP and/or by any member(s) of the MP Laboratory, and which is Controlled by UCLB either at
the time of creation, development or generation or upon UCL or UCLB acquiring or being granted a licence to the same from the
inventors or individuals responsible for the creation, development or generation of the same;
“[***]  Contract”  means  the  [***]  contract  entered  into  between  (i)  [***]  and  (ii)  [***]  on  [***],  together  with  the  associated  letter
agreement between (i) [***] and (ii) [***] dated [***];
“Non-Defaulting Party” has the meaning set out in Clause 24.4;
“Notice Period” has the meaning set out in Clause 24.4.1;
“NSG Licence” has the meaning set out in Clause 2.2.14;
“NSG Product” means any product or therapy which is covered by, uses or incorporates any of the NSG Program IP;
“NSG Program” means the program of research defined in Schedule 1 under the title New Suicide Gene Program conducted prior
to the Effective Date together with the program of research conducted after the Effective Date and prior to the Amendment Date as
described in Schedule 1 under the title (B) New Suicide Gene Project Plan;
“NSG  Program  IP”  means,  excluding  Manufacturing  Know-How,  (i)  the  Know-How  described  in  Schedule  2  under  the  title  NSG
Know-How; (ii) the materials listed in Schedule 2 under the title NSG Materials together with all IP in the same; (iii) all Know-How and
materials (including all IP in the same) developed by MP and/or the MP Laboratory pursuant specifically to the NSG Program prior to
the Effective Date, other than that which is CGK as of the Amendment Date; (iv) any Patent Rights filed by or on behalf of UCLB from
time to time and in respect of any work undertaken by MP and/or the MP Laboratory for the NSG Program prior to the Amendment
Date and all Patent Rights resulting therefrom, including those as set out in Schedule 1 (“NSG Patent Rights”); and (v) all Know-
How and materials (including all IP in the same) developed after the Effective Date but prior to the Amendment Date by MP and/or
the  MP  Laboratory  pursuant  specifically  to  the  agreed  project  plan  for  the  NSG  Program  as  set  out  in  Schedule  1  including  the
technology claimed or disclosed in the NSG Patent Rights (if any) as of the Amendment Date, other than that which is CGK as of the
Amendment Date, or disclosed in the sections of the NSG Patent Rights that refer to, describe or disclose prior art including CGK
existing  as  of  the  Amendment  Date;  references  to  CGK  in  this  definition  shall  exclude  CGK  that  became  CGK  through  disclosure
after the Effective Date by publication of the NSG Patent Rights or by MP making a publication in accordance with this Agreement or
by Autolus’s publication;
“Option” means the [***] Option;
“Original  Licensed  Patents”  means  the  BCMA-CAR  Patent  Rights,  the  [***]  Patent  Rights  (if  Autolus  exercises  its  rights  under
Clause 2.4), the GD2 Patent Rights, the TRBC1/2 Patent Rights, the RQR8 Patent Rights, the NSG Patent Rights, the Logic Gate
Patent Rights and the ZipCAR Patent Rights;
“Original Program” means one of the GD2 Program, the BCMA-CAR Program, the TRBC1/2 Program, the [***] Program (if Autolus
exercises its rights under Clause 2.5), the RQR8 Program, the NSG Program, the Logic Gate Program or the ZipCAR Program, and
“Original Programs” means any combination of two or more of the foregoing, as the context requires;
“Original Program IP” means the GD2 Program IP, the BCMA Program IP, the TRBC1/2 Program IP, the RQR8 Program IP, the
Logic Gate Program IP, the NSG Program IP and the ZipCAR Program IP; and (ii) where the [***] Option has been exercised, the
[***] Program IP;
“Original Program Licence” means any one of the BCMA Licence, GD2 Licence, Logic Gate Licence, RQR8 Licence, TRBC1/2
Licence, NSG Licence, ZipCAR Licence, and

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

where  the  [***]  Option  has  been  exercised,  the  [***]  Licence,  and  “Original  Program  Licences”  means  any  two  or  more  of  the
foregoing;
“Original  Royalty  Product”  means  any  one  of  the  BCMA  Product,  GD2  Product,  Logic  Gate  Product,  RQR8  Product,  TRBC1/2
Product,  NSG  Product,  ZipCAR  Product,  and  where  the  [***]  Option  has  been  exercised,  [***]  Product,  and  “Original  Royalty
Products” means any two or more of the foregoing;
“Other Technology” has the meaning set out in Clause 8.3;
“Party”  or  “Parties”  means  Autolus  or  UCLB,  or  both  Autolus  and  UCLB,  as  the  context  requires,  including  their  respective
successors in title, permitted assignees and transferees from time to time (if any);
“Patent Assignment” means the deed of assignment between the Parties in respect of the Assigned Patents dated on the Third
Amendment Date;
“Patent Prosecution Costs” means those professional service fees and costs reasonably charged by a Third Party for the provision
of patent filing, prosecution (including defending oppositions and interferences), maintenance and renewal services with respect to
the applicable Licensed Patent, including all official fees, charges and surcharges properly incurred in the provision of such services;
“Patent Rights” means all patent rights, claims in any patent right, applications for patents and the right to apply for patent rights in
any part of the world including all divisionals, reissues, extensions, substitutions, confirmations, registrations, revalidations, additions,
continuations in-part and any supplementary protection certificates or patent term extensions and where referred to in the context of
a schedule hereto shall include all patent rights from time to time derived from, claiming priority from, issued or granted from those
patent rights listed in such schedule;
“Patent Sale” has the meaning set out in Clause 15.1;
“Permitted Studies” means those clinical studies identified and briefly described in Schedule 7;
“Primary CNS Lymphoma” means lymphoma of CNS cells that has originated within the central nervous system as the primary site
of the lymphoma, as opposed to a secondary lymphoma of the CNS;
“Program” means any one of the Original Programs, the Additional Programs or the CAT19 Program, and “Programs” means any
combination of two or more of the Original Programs, Additional Programs and the CAT19 Program, as the context requires;
“Program IP” means the Original Program IP, the Additional Program IP, the CAT19 Program IP and the CAROUSEL Program IP;
“Program Licence” means one of the BCMA Licence, [***] Licence (if Autolus exercises its rights under Clause 2.5), GD2 Licence,
Logic Gate Licence, NSG Licence, RQR8 Licence, TRBC1/2 Licence, ZipCAR Licence, ccCAR Licence, Epitope Tag Licence, iCAR
Licence,  RapaiCASP9  Licence,  Retrostim  Licence,  TetCAR  Licence,  ZAP-CAR  Licence,  CAT19  Licence,  TRBC2  Dx  Licence  and
CAT19 CNS Licence;
“Program Materials” means those materials forming part of the Program IP;
“Purple Book Reference” has the meaning set out in Clause 18.8;
“Quarterly” or “Quarter” means a period of three calendar months each commencing on 31 March, 30 June, 30 September or 31
December;

“RapaiCASP9 Licence” has the meaning set out in Clause 2.2.9;
“RapaiCASP9 Product”  means  any  product  or  therapy  which,  were  it  not  for  the  licence  to  RapaiCASP9  Patent  Rights  granted
hereunder  or  the  assignment  to  Autolus  (or  subsequent  assignment  or  licence  grant  to  an  Assignee  Entity)  of  the  RapaiCASP9
Patent Rights, would otherwise infringe the RapaiCASP9 Patent Rights in the relevant country of sale;
“RapaiCASP9 Program” means the program of research defined in Schedule 1 under the title RapaiCASP9 Program as conducted
prior to the Amendment Date;
“RapaiCASP9 Program IP” means excluding Manufacturing Know-How, (i) those patent applications listed in Schedule 2 under the
title RapaiCASP9 Patents and all Patent Rights derived therefrom (“RapaiCASP9 Patent Rights”); (ii) the Know-How described in
Schedule 2 under the title RapaiCASP9 Know-How; and (iii) the materials listed in Schedule 2 under the title RapaiCASP9 Materials
together with all IP in the same;
“Recipient Party” has the meaning set out in Clause 20.1;

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

“Referral Notice” has the meaning set out in paragraph 1 of Part C Schedule 8;
“Regulatory Approval” means any and all approvals (including any applicable supplements, amendments, pre and post approvals,
and  approvals  of  applications  for  regulatory  exclusivity),  licenses,  registrations,  or  authorisations  of  any  federal,  national,
multinational,  state,  provincial  or  local  regulatory  agency,  department,  bureau,  commission,  council  or  other  governmental  entity
necessary for the manufacture, distribution, use, testing, development, storage, import, export, transport, promotion, marketing and
sale of a medicinal product in a country or countries;
“Regulatory  Authority”  means  any  governmental  or  regulatory  authority  responsible  for  assessing  and/or  granting  Regulatory
Approvals (including any ethics committees) and “Regulatory Authorities” shall mean more than one such authority;
“Regulatory Exclusivity” means on a country by country basis that a Third Party under applicable laws concerning the application
for  and  grant  of  Marketing  Approvals  having  jurisdiction  over  that  country  at  the  relevant  time  (i)  is  precluded  from  submitting  a
Marketing Approval Application for any Competitive Product; or (ii) in respect of any Competitive Product, is prohibited from relying
upon  Autolus’s,  or  its  Affiliates’  or  Sub-Licensee’s  or  any  Assignee  Entity’s  product  dossier  or  Regulatory  Submission  (from  which
such party obtained a Marketing Approval) concerning any Royalty Product, under any applicable abridged or streamlined procedure
to obtain a Marketing Approval for such Competitive Product;
“Regulatory Submission”  means  in  respect  of  a  Royalty  Product,  the  package  or  packages  of  data,  pre-clinical  and  clinical  trial
data and materials, information, results, materials and samples (including any Test and Regulatory Data and/or the drug master file
or part thereof) submitted to a Regulatory Authority in support of a Marketing Approval Application or any other Regulatory Approval;
“Retained Patents” means the RQR8 Patent Rights and the [***] Existing Patent;
“Retrostim Field” means all uses and applications without restriction;
“Retrostim Licence” has the meaning set out in Clause 2.2.13;
“Retrostim  Product”  means  any  product  or  therapy  within  the  Retrostim  Field:  (i)  which,  were  it  not  for  the  licence  granted
hereunder or the assignment to Autolus (or subsequent assignment or licence grant to an Assignee Entity) of the Retrostim Patent
Rights, would infringe the Retrostim Patent Rights; and/or (ii) the manufacture of which product or therapy were it not for the licence
granted hereunder or the assignment to Autolus (or subsequent assignment or licence grant to an Assignee Entity) of the Retrostim
Patent Rights, would infringe the Retrostim Patent Rights, and/or (iii) where the manufacture of any materials used in connection with
the  manufacture  of  such  product  or  therapy,  were  it  not  for  the  licence  granted  hereunder  or  the  assignment  to  Autolus  (or
subsequent assignment or licence grant to an Assignee Entity) of the Retrostim Patent Rights, would infringe the Retrostim Patent
Rights;
“Retrostim Program” means the program of research defined in Schedule 1 under the title Retrostim Program as conducted prior to
the Amendment Date;
“Retrostim Program IP” means excluding Manufacturing Know-How, (i) those patent applications listed in Schedule 2 under the title
Retrostim Patents and all Patent Rights derived therefrom (“Retrostim Patent Rights”); (ii) the Know-How described in Schedule 2
under the title Retrostim Know-How; and (iii) the materials listed in Schedule 2 under the title Retrostim Materials together with all IP
in the same;
“Royalty” or “Royalties” has the meaning set out in Clause 14.1;
“Royalty Expiry” has the meaning set out in Clause 14.6;
“Royalty Product” means any one of the Original Royalty Products, Additional Royalty Products or CAT19 Products, and “Royalty
Products”  shall  be  constructed  to  mean  any  combination  of  two  or  more  of  the  Original  Royalty  Products,  Additional  Royalty
Products and CAT19 Products, as the context requires;
“RQR8 Field” means all uses and applications without restriction except for any uses or activities relating to any allogeneic T-cell
therapy  (being  the  therapeutic  transplantation  of  genetically  engineered  T-cell  lymphocyte  to  a  recipient  patient  from  a  genetically
non-identical  donor  of  the  same  species),  including  any  activities  of  research,  development  or  commercialisation  of  a  therapeutic
relating to any allogeneic T-cell therapy;
“RQR8 Licence” has the meaning set out in Clause 2.2.5;

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

“RQR8  Permitted  Studies”  means  those  studies  identified  and  briefly  described  in  Schedule  7  under  the  title  RQR8  Permitted
Studies;
“RQR8 Product” means any product or therapy within the RQR8 Field which is covered by, uses or incorporates any of the RQR8
Program IP;
“RQR8  Program”  means  the  program  of  research  defined  in  Schedule  1  under  the  title  RQR8  Program  conducted  prior  to  the
Effective Date;
“RQR8 Program IP” means, excluding Manufacturing Know-How, (i) those patent applications listed in Schedule 2 under the title
RQR8 Patents at all Patent Rights derived therefrom (“RQR8 Patent Rights”); (ii) the Know-How described in Schedule 2 under the
title RQR8 Know-How; (iii) the materials listed in Schedule 2 under the title RQR8 Materials together with all IP in the same; and (iv)
all  Know-How  developed  by  MP  and/or  the  MP  Laboratory  pursuant  specifically  to  the  RQR8  Program  prior  to  the  Effective  Date
including the technology claimed or disclosed in the RQR8 Patent Rights as of the Effective Date, other than that which is CGK as of
the Effective Date or disclosed in the sections of the RQR8 Patent Rights that refer to, describe or disclose prior art including CGK
existing as of the Effective Date;
“Sales Milestone” has the meaning set out in Clause 13.4;
“Sales Milestone Payment” has the meaning set out in Clause 13.4;
“Second Amendment Date” has the meaning set out in the first paragraph of this Agreement;
“SPC” has the meaning set out in Clause 18.8;
“SSA” means the Subscription and Shareholders’ Agreement between Autolus, UCLB, Dr Martin Pule, John Berriman and Syncona
LLP dated as of the Effective Date;
“Sublicence” has the meaning set out in Clause 15.1;
“Sub-Licensee” means any person (including an Affiliate) to whom Autolus: (i) sub-licenses all or part of the Intellectual Property
licensed to it under this Agreement in respect of Royalty Products; or (ii) licences any or all of the Assigned Patents in respect of
Royalty Products;
“Sublicence Or Patent Sale Payment” has the meaning set out in Clause 15.1;
“Success Milestone” has the meaning set out in Clause 13.1.1;
“Success Milestone Payment” has the meaning set out in Clause 13.1.1;
“Surrender” or “Surrendered” means in respect of any Patent Rights, any of (i) ceasing to maintain (by payment of renewal fees or
otherwise)  the  applicable  Patent  Rights;  (ii)  withdrawing,  surrendering,  dedicating  to  the  public  or  allowing  the  applicable  Patent
Rights  to  lapse;  (iii)  in  the  case  of  a  pending  application  de-designating,  or  not  validating  or  ratifying  in,  a  country  covered  by  the
application or not entering into the national or regional phase in a country designated in the international or convention application or
(iv) consenting to or ceasing to defend an application, action or litigation for revocation;
“Technology” collectively means all Program IP, Manufacturing Know-How, UCL Background IP and the Materials;
“Term” has the meaning in Clause 24.1;
“Territory” means all countries throughout the World;
“Test and Regulatory Data” means any and all pre-clinical and clinical test data, test designs and protocols, results and data from
pre-clinical and clinical studies, information contained in or submitted in government licenses and applications therefor, government
certifications  and  findings,  and  related  materials,  data  from  pre-clinical  and  clinical  experiments  demonstrating  or  supporting
bioavailability, bioequivalence, data concerning any adverse drug reactions;
“TetCAR Field” means all uses and applications without restriction;
“TetCAR Licence” has the meaning set out in Clause 2.2.10;
“TetCAR Product” means any product or therapy within the TetCAR Field which, were it not for the licence to TetCAR Patent Rights
granted hereunder or the assignment to Autolus (or subsequent assignment or licence grant to an Assignee Entity) of the TetCAR
Patent Rights, would otherwise infringe the TetCAR Patent Rights in the relevant country of sale;
“TetCAR Program” means the program of research defined in Schedule 1 under the title TetCAR Program as conducted prior to the
Amendment Date;
“TetCAR Program IP” means excluding Manufacturing Know-How, (i) those patent applications listed in Schedule 2 under the title
TetCAR Patents and all Patent Rights

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

derived therefrom (“TetCAR Patent Rights”); (ii) the Know-How described in Schedule 2 under the title TetCAR Know-How; and (iii)
the materials listed in Schedule 2 under the title TetCAR Materials together with all IP in the same;
“Third Party” means any person other than the Parties or their respective Affiliates or MP;
“Third Party Access Rights” has the meaning set out in Clause 14.9;
“Tobacco Party” means: (i) any entity which develops, sells or manufactures tobacco products; and/or (ii) any entity which makes
the majority of its profits from the importation, marketing, sale or disposal of tobacco products. Furthermore, Tobacco Party shall include any
entity that is controlled by or controls any entity referred to in (i) or (ii); controlled and controls having the meaning defined in the definition of
Affiliate;

“TP Fees” has the meaning set out in Clause 14.9;
“TRBC1”  means  the  specific  Antigen  (or  part  thereof)  coded  for  by  reference  to  the  sequence  defined  in  Schedule  1  under  the
TRBC1/2 Program;
“TRBC1/2 Licence” has the meaning set out in Clause 2.2.6;
“TRBC1/2  Product”  means  any  product  therapy  which  (i)  targets  TRBC1/2  and  is  covered  by,  uses,  incorporates  or  has  been
developed  using  any  of  the  TRBC1/2  Program  IP;  or  (ii)  targets  TRBC1  and/or  TRBC2  and  has  been  developed  using  any  of  the
UCL Background IP or Manufacturing Know-How; or (iii) were it not for the license to TRBC1/2 Program IP hereunder, would infringe
any Patent Right licensed under the TRBC1/2 Program IP or, were it not for the assignment of the TRBC1/2 Patent Rights to Autolus
(or subsequent assignment or licence grant to an Assignee Entity), would infringe any TRBC1/2 Patent Rights;
“TRBC1/2 Program” means the program of research defined in Schedule 1 under the title TRBC1/2 Program conducted prior to the
Effective Date;
“TRBC1/2 Program IP” means, excluding Manufacturing Know-How, (i) those patent applications listed in Schedule 2 under the title
TRBC1/2-CAR  Patents  and  all  Patent  Rights  derived  therefrom  (“TRBC1/2  Patent  Rights”);  (ii)  the  Know-How  described  in
Schedule 2 under the title TRBC1/2 Know-How; and (iii) the materials listed in Schedule 2 under the titles “TRBC1/2 Materials” and
“TRBC1/2 Materials – [***]” together with all IP in the same; and (iv) all Know-How and materials (together with all IP in the same)
developed by MP and/or the MP Laboratory pursuant specifically to the TRBC1/2 Program prior to the Effective Date including the
technology  claimed  or  disclosed  in  the  TRBC1/2  Patent  Rights  as  of  the  Effective  Date,  other  than  that  which  is  CGK  as  of  the
Effective Date or disclosed in the sections of the TRBC1/2 Patent Rights that refer to, describe or disclose prior art including CGK
existing as of the Effective Date;
“TRBC2”  means  the  Antigen  (or  part  thereof)  coded  for  by  reference  to  the  sequence  defined  in  Schedule  1  under  the  TRBC1/2
Program;
“TRBC2 Dx Licence” has the meaning set out in Clause 2.2.17;
“TRBC2 Dx Product” means any product or therapy: (i) which, were it not for Autolus’ ownership of the TRBC2 Dx Patent Rights (or
subsequent assignment or licence of the TRBC2 Dx Patent Rights to an Assignee Entity), would otherwise infringe any TRBC2 Dx
Patent  Rights;  and/or  (ii)  the  manufacture  of  which  product  or  therapy  were  it  not  for  Autolus’  ownership  of  the  TRBC2  Dx  Patent
Rights (or subsequent assignment or licence of the TRBC2 Dx Patent Rights to an Assignee Entity), would infringe any TRBC2 Dx
Patent  Rights;  and/or  (iii)  where  the  manufacture  of  any  materials  used  in  connection  with  the  manufacture  of  such  product  or
therapy, were it not for Autolus’ ownership of the TRBC2 Dx Patent Rights (or subsequent assignment or licence of the TRBC2 Dx
Patent Rights to an Assignee Entity), would infringe any TRBC2 Dx Patent Rights;
“TRBC2 Dx Program” means the program of research defined in Schedule 1 under the title TRBC2 Dx Program as conducted prior
to the Third Amendment Date;
“TRBC2 Dx Program IP” means excluding Manufacturing Know-How, (i) those patent applications listed in Schedule 2 under the
title  TRBC2  Dx  Patents  and  all  Patent  Rights  derived  therefrom  (“TRBC2  Dx  Patent  Rights”);  and  (ii)  the  materials  listed  in
Schedule 2 under the title TRBC2 Dx Materials together with all IP in the same;
“UCL” means University College London, an institution incorporated in the United Kingdom by Royal Charter and having its address
at Gower Street, London, WC1E 6BT;

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

“UCL Background IP” means excluding Program IP and Manufacturing Know-How, (i) all Know-How listed in Part A of Schedule 3;
(ii) excluding that listed in Part A of Schedule 3, all other Know-How (excluding binder domains) developed by MP and/or the MP
Laboratory  pursuant  to  or  in  conjunction  with  any  of  the  Original  Programs  (or  any  combination  of,  or  across  any  of,  the  Original
Programs) prior to the Effective Date (or in the case of the New Suicide Gene Program, or the ZipCAR Program, respectively prior to
the Amendment Date, or in the case of the [***] Program prior to the date of exercise of the [***] Option) to the extent that it is not
CGK as of the Effective Date and UCLB Controls the same; (iii) excluding that listed in Part A of Schedule 3, all other Know-How
(excluding  binder  domains)  developed  by  MP  and/or  the  MP  Laboratory  pursuant  to  or  in  conjunction  with  any  of  the  Additional
Programs (or any combination of, or across any of, the Additional Programs) prior to the Amendment Date to the extent that it is not
CGK as of the Amendment Date and UCLB Controls the same; (iv) excluding that listed in Part A of Schedule 3, all other Know-How
(excluding binder domains) developed by MP and/or the MP Laboratory pursuant to or in conjunction with any of the CAT19 Program
prior to the Second Amendment Date to the extent that it is not CGK as of the Second Amendment Date and UCLB Controls the
same; (v) excluding that listed in Part A of Schedule 3, all other Know-How (excluding binder domains) developed by MP and/or the
MP Laboratory pursuant to or in conjunction with the CAROUSEL STUDY prior to the Third Amendment Date to the extent that it is
not CGK as of the Third Amendment Date and UCLB Controls the same (vi) excluding that listed in Part A of Schedule 3, all other
Know-How (excluding binder domains) developed by MP and/or the MP Laboratory pursuant to or in conjunction with the TRBC2 Dx
Program  on  or  after  the  Amendment  Date  but  prior  to  the  Third  Amendment  Date  to  the  extent  that  it  is  not  CGK  as  of  the  Third
Amendment Date and UCLB Controls the same;
“UCLB Indemnified Parties” has the meaning set out in Clause 23.1.2;
“Unresolved Matter” has the meaning set out in Clause 33.2;
“Valid Claim”  means  a  claim  within  (i)  an  issued  or  granted  and  unexpired  Patent  Right,  including  any  Licensed  Patent;  or  (ii)  a
pending application for a Patent Right including an application with respect to any Licensed Patents, which has not been pending for
more than eight (8) years from the date of the priority filing from which such pending application originates, and in each case of (i)
and  (ii)  above,  which  has  not  been  held  unenforceable,  unpatentable  or  invalid  by  a  decision  of  a  court  or  government  body  of
competent  jurisdiction,  or  where  appealed  within  the  time  allowed  for  appeal  has  not  been  held  unenforceable,  unpatentable  or
invalid  by  an  appellate  body  of  competent  jurisdiction  (including  by  the  highest  appellate  court  in  the  relevant  jurisdiction  where
appealed  to  that  court),  or,  which  has  not  been  withdrawn,  cancelled,  revoked,  disclaimed,  or  rendered  unenforceable  through
disclaimer  or  otherwise,  or  which  has  not  been  donated  or  dedicated  to  the  public,  Surrendered  or  which  has  not  been  deemed
invalid by an interference or opposition panel or court as part of any interference or opposition proceeding;
“Year” means a period of twelve (12) months commencing on 1 January;
“ZAP-CAR Licence” has the meaning set out in Clause 2.2.11;
“ZAP-CAR Product” means any product or therapy which, were it not for the licence to ZAP-CAR Patent Rights granted hereunder
or  the  assignment  to  Autolus  (or  subsequent  assignment  or  licence  grant  to  an  Assignee  Entity)  of  the  ZAP-CAR  Patent  Rights,
would otherwise infringe the ZAP-CAR Patent Rights in the relevant country of sale;
“ZAP-CAR Program” means the program of research defined in Schedule 1 under the title ZAP-CAR Program as conducted prior to
the Amendment Date;
“ZAP-CAR Program IP” means excluding Manufacturing Know-How, (i) those patent applications listed in Schedule 2 under the title
ZAP-CAR Patents and all Patent Rights derived therefrom (“ZAP-CAR Patent Rights”); (ii) the Know-How described in Schedule 2 under
the title ZAP-CAR Know-How; and (iii) the materials listed in Schedule 2 under the title ZAP-CAR Materials together with all IP in the same;

“ZipCAR Licence” has the meaning set out in Clause 2.2.15;
“ZipCAR Product” means any product or therapy which is covered by, uses or incorporates any of the ZipCAR Program IP;

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

“ZipCAR Program” means the program of research defined in Schedule 1 under the title ZipCAR Program conducted prior to the
Effective  Date  together  with  the  program  of  research  conducted  after  the  Effective  Date  and  prior  to  the  Amendment  Date  as
described in Schedule 1 under the title (B): ZipCAR Project Plan;
“ZipCAR  Program  IP”  means,  excluding  Manufacturing  Know-How  (i)  the  Know-How  described  in  Schedule  2  under  the  title
ZipCAR Know-How; (ii) the materials listed in Schedule 2 under the ZipCAR Materials together with all IP in the same; (iii) all Know-
How  and  materials  (including  all  IP  in  the  same)  developed  by  MP  and/or  the  MP  Laboratory  pursuant  specifically  to  the  ZipCAR
Program prior to the Effective Date other than that which is CGK as of the Amendment Date; (iv) any Patent Rights filed by or on
behalf of UCLB from time to time in respect of any work undertaken by MP and/or the MP Laboratory for the ZipCAR Program prior
to the Amendment Date and all Patent Rights therefrom, including those as set out in Schedule 1 (“ZipCAR Patent Rights”); and (v)
all Know-How and materials (including IP in the same) developed after the Effective Date but prior to the Amendment Date by MP
and/or the MP Laboratory pursuant specifically to the agreed project plan for the ZipCAR Program as set out in Schedule 1, including
the technology claimed or disclosed in the ZipCAR Patent Rights (if any) as of the Amendment Date, other than that which is CGK as
of the Amendment Date, or disclosed in the sections of the ZipCAR Patent Rights that refer to, describe or disclose prior art including
CGK existing as of the Amendment Date, provided that to the extent that any materials in (ii), (iii) or (v) incorporate the binder CD33,
such  binder  together  with  IP  in  the  same  is  excluded;  references  to  CGK  in  this  definition  shall  exclude  CGK  that  became  CGK
through disclosure after the Effective Date by publication of the ZipCAR Patent Rights or by MP making a publication in accordance
with this Agreement or by Autolus’s publication;

In this Agreement, unless the context requires otherwise:
use of the singular includes the plural and vice versa and use of any gender includes the other genders;
any reference to “this Agreement” is a reference to this Agreement as from time to time amended, varied or extended in any way;

and,

“undertaking” shall have the meaning given by section 1161 Companies Act 2006 save that for the purposes of this Agreement and

for the avoidance of doubt, an undertaking shall include a limited liability partnership.

In this Agreement unless otherwise specified:
any  reference  to  a  recital,  clause,  paragraph  or  schedule  is  to  the  relevant  recital,  clause,  paragraph  or  schedule  of  or  to  this
Agreement, and any reference in a schedule to a part or a paragraph (as opposed to a clause) is to a part or a paragraph of that
schedule;

any  reference  to  a  “person”  includes  an  individual,  firm,  partnership,  body  corporate,  corporation,  association,  organisation,

government, state, foundation and trust, in each case whether or not having separate legal personality;

“parent undertaking” and “subsidiary undertaking” shall have the respective meanings given by section 1162 Companies Act 2006
save  that  for  the  purposes  of  this  Agreement,  an  undertaking  shall  be  treated  as  a  member  of  another  undertaking  if  any  of  the
shares in that other undertaking are registered in the name of another person (or its nominee) as security (or in connection with the
taking of security) from the first undertaking or any of that first undertaking's subsidiary undertakings;

any reference to a statute, statutory provision or subordinate legislation (“legislation”) shall be construed as referring to that legislation
as  amended  and  in  force  from  time  to  time  and  to  any  legislation  which  re-enacts,  re-writes  or  consolidates  (with  or  without
modification) any such legislation;

any reference to an English legal term or concept or any court, official, governmental or administrative authority or agency in England
includes, in respect of any jurisdiction other than England, a reference to whatever most closely approximates in that jurisdiction to
the relevant English legal term;

b.
c.
d.

e.

f.
g.

h.

i.

j.

k.

l.
m.

any reference to an agreement includes any form of arrangement, whether or not in writing and whether or not legally binding;
“writing”  shall  include  any  modes  of  reproducing  words  in  a  legible  and  non-transitory  form  excluding  (unless  expressly  stated  to

include) email, SMS and other temporary

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

n.
o.

p.

q.
r.

2.
a.

i..

ii..

transient electronic messaging systems and “written” shall be construed accordingly; and,

a period of time being specified which dates from a given day or the day of an act or event, it shall be calculated exclusive of that day.
In this Agreement, the words “other”, “including”, “includes”, “include”, “in particular” and any similar words, shall not limit the general

effect of words that precede or follow them and accordingly, the ejusdem generis rule shall not apply.

Any undertaking by or obligation on a Party not to do any act or thing includes an undertaking not to allow, cause or assist the doing of
that act or thing and to exercise all rights of control over the affairs of any other person which that Party is able to exercise (directly or
indirectly) in order to secure performance of that undertaking, which in the case of UCLB shall include UCL as a person over whom
UCLB can exercise control.

The index to and the headings in this Agreement are for information only and are to be ignored in construing the same.
Where  this  Agreement  refers  to  a  Person  being  “free”  to  do  something,  this  shall  be  construed  as  that  Person  not  being  prevented,

whether by law, equity or contract, from doing that thing.
LICENCE GRANT

UCL Background Licence

Subject to Clause 4, UCLB hereby grants to Autolus for the full duration of the Term and throughout the Territory a licence to the UCL
Background IP and Background Materials (including the right to sub-license through multiple tiers) within the Field to use the same:

for any and all acts of Exploitation with respect to any and all of the Royalty Products, which licence shall be exclusive to Autolus

(to the exclusion of UCLB, UCL and any Third Party); and

without prejudice to Clause 2.1.1, for any other purpose or act of Exploitation without restriction (including in respect of Royalty
Products  if  the  licence  under  Clause  2.1.1  terminates),  which  licence  shall  be  non-exclusive,  royalty-free,  irrevocable  and
perpetual;

(collectively the “Background Licence”). It is acknowledged that the Background Licence granted under Clause 2.1.1 shall include
Exploitation of the [***] Product upon election of the [***] Option, and shall include Exploitation of the NSG Product and shall include
Exploitation  of  the  ZipCAR  Product  and  that  UCLB  shall  not  (and  shall  procure  that  UCL  shall  not)  grant  any  rights  under  UCL
Background IP and Background Materials with respect to any and all acts of Exploitation with respect to the [***] Product until such
time (if any) as Autolus’s right to exercise the [***] Option has expired. The Background Licence shall be assignable as part of any
assignment  of  this  Agreement  in  accordance  with  Clause  29.  Autolus  shall  not  be  entitled  to  assign  the  Background  Licence
independently of the other Licences.

b.

Program Licences

Subject to Clause 4, UCLB hereby grants to Autolus for the full duration of the Term and throughout the Territory an exclusive (save
as  provided  in  Clause  2.2.3  and  Clause  2.2.6)  licence  (to  the  exclusion  of  UCLB,  UCL  and  any  Third  Party)  which  shall  be  sub-
licensable through multiple tiers:

with effect from the Effective Date to the BCMA Program IP within the BCMA Field to use the same for any act of Exploitation

concerning any product, therapy, service or process without restriction;

with effect from the Effective Date to the [***] Existing Patent within the BCMA Field to use the same for any act of Exploitation
concerning  any  product,  therapy,  service  or  process  without  restriction  (collectively  the  licences  under  Clauses  2.2.1  and
2.2.2 being the “BCMA Licence”);

with effect from the Effective Date to the GD2 Program IP within the Field to use the same for any act of Exploitation concerning
any  product,  therapy,  service  or  process  without  restriction,  save  that  where  the  provisions  of  Clause  9.7  apply  and  GD2
Clinical  Study  Results  are  deemed  to  be  part  of  the  GD2  Program  IP,  Autolus’s  licence  pursuant  to  this  Clause  2.2.3  in
respect  of  the  GD2  Clinical  Study  Results  only  shall  be  exclusive  to  those  aspects  of  the  GD2  Clinical  Study  Results  that
relate directly to and only to the GD2 Product and non-exclusive to

i..

ii..

iii..

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

all other aspects of the GD2 Clinical Study Results for use in connection with CARs (“GD2 Licence”);

with  effect  from  the  Effective  Date  to  the  Logic  Gate  Program  IP  within  the  Field  to  use  the  same  for  any  act  of  Exploitation

concerning any product, therapy, service or process without restriction (“Logic Gate Licence”);

with  effect  from  the  Effective  Date  to  the  RQR8  Program  IP  within  the  RQR8  Field  to  use  the  same  for  any  act  of  Exploitation

concerning any product, therapy, service or process without restriction (the “RQR8 Licence”);

with  effect  from  the  Effective  Date  to  the  TRBC1/2  Program  IP  within  the  Field  to  use  the  same  for  any  act  of  Exploitation
concerning  any  product,  therapy,  service  or  process  without  restriction  save  that  this  licence  pursuant  to  Clause  2.2.6  in
relation to the [***] only shall be limited to the [***] (i) for the purpose of Exploiting a TRBC1/2 Product in humans (including
for  the  avoidance  of  doubt,  use  of  [***]  for  pre-clinical  studies  in  animals  or  otherwise  for  such  Exploitation  of  a  TRBC1/2
Product) and (ii) for Exploiting companion diagnostics for use with a TRBC1/2 Product; (“TRBC1/2 Licence”);

with  effect  from  the  Amendment  Date  to  the  ccCAR  Program  IP  within  the  Field  to  use  the  same  for  any  act  of  Exploitation

concerning any product, therapy, service or process without restriction (“ccCAR Licence”);

with  effect  from  the  Amendment  Date  to  the  iCAR  Program  IP  within  the  Field  to  use  the  same  for  any  act  of  Exploitation

concerning any product, therapy, service or process without restriction (“iCAR Licence”);

with effect from the Amendment Date to the RapaiCASP9 Program IP within the Field to use the same for any act of Exploitation

concerning any product, therapy, service or process without restriction (“RapaiCASP9 Licence”);

with  effect  from  the  Amendment  Date  to  the  TetCAR  Program  IP  within  the  TetCAR  Field  to  use  the  same  for  any  act  of

Exploitation concerning any product, therapy, service or process without restriction (“TetCAR Licence”);

with effect from the Amendment Date to the ZAP-CAR Program IP within the Field to use the same for any act of Exploitation

concerning any product, therapy, service or process without restriction (“ZAP-CAR Licence”);

with effect from the Amendment Date to the Epitope Tag Program IP within the Epitope Tag Field to use the same for any act of

Exploitation concerning any product, therapy, service or process without restriction (the “Epitope Tag Licence”);

with  effect  from  the  Amendment  Date  to  the  Retrostim  Program  IP  within  the  Retrostim  Field  to  use  the  same  for  any  act  of

Exploitation concerning any product, therapy, service or process without restriction (the “Retrostim Licence”);

with  effect  from  the  Amendment  Date  to  the  NSG  Program  IP  within  the  Field  to  use  the  same  for  any  act  of  Exploitation

concerning any product, therapy, service or process without restriction (“NSG Licence”);

with  effect  from  the  Amendment  Date  to  the  ZipCAR  Program  IP  within  the  Field  to  use  the  same  for  any  act  of  Exploitation

concerning any product, therapy, service or process without restriction (“ZipCAR Licence”);

with  effect  from  the  Second  Amendment  Date  to  the  CAT19  Program  IP  within  the  CD19  Field  to  use  the  same  for  any  act  of

Exploitation concerning any product, therapy, service or process without restriction (“CAT19 Licence”);

with  effect  from  the  Third  Amendment  Date  to  the  TRBC2  Dx  Program  IP  within  the  Field  to  use  the  same  for  any  act  of

Exploitation concerning any product, therapy, service or process without restriction (“TRBC2 Dx Licence”); and

with effect from the Third Amendment Date to the CAROUSEL Program IP within the CD19 Field to use the same for any act of

Exploitation concerning any product, therapy, service or process without restriction (“CAT19 CNS Licence”),

it  being  acknowledged  that  each  of  the  foregoing  Licences  granted  under  this  Clause  2.2(i)  may  only  be  revoked  or  terminated
pursuant  to  a  right  of  termination  under  Clause  11.1.5,  11.2.3,  11.3.2  or  Clause  24  (as  applicable)  in  respect  of  the  applicable
Licence(s); and (ii) may all be assigned together (but not individually) as part of any assignment of this Agreement in accordance with
Clause 29.

iv..

v..

vi..

vii..

viii..

ix..

x..

xi..

xii..

xiii..

xiv..

xv..

xvi..

xvii..

xviii..

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

c.

Manufacturing Know-How

i..
ii..

Subject  to  Clause  4,  UCLB  hereby  grants  to  Autolus  for  the  full  duration  of  the  Term  and  throughout  the  Territory  a  non-exclusive
licence under the Manufacturing Know-How (including the right to sub-license through multiple tiers) within the Field to use the same:

for any act of Exploitation of the Royalty Products in the Territory; and
without prejudice to Clause 2.3.1 for any other act of Exploitation or purpose without restriction (including in respect of a Royalty
Product  if  the  licence  under  Clause  2.1.1  terminates),  in  which  case  such  licence  shall  be  royalty-free,  irrevocable  and
perpetual;

the (“Manufacturing Licence”). The Manufacturing Licence may be assigned together (but not alone) as part of any assignment of
this Agreement in accordance with Clause 29.

d.

Option to a Licence under the [***] Program IP

i..

ii..

iii..

iv..

v..

e.

At any time up until [***] following Autolus’s receipt of the Final Written Report concerning the [***] Program in accordance with the
[***]  project  plan  set  out  in  Schedule  1  (“[***]  Exercise  Period”),  Autolus  shall  have  the  right  upon  written  notice  to  UCLB,  to
exercise its option for the grant of a licence on the terms of this Agreement to the [***] Program IP (“[***] Licence Notice”). Prior to
service of a [***] Licence Notice:

UCLB shall keep Autolus informed, on a reasonably frequent basis (or otherwise upon request by Autolus), of (i) the status and
performance of the [***] Program against the [***] project plan set out in Schedule 1; (ii) developments and advancements in
the  [***]  Program  since  the  last  report;  (iii)  identification  and  disclosure  of  any  [***]  Program  IP  not  previously  disclosed  to
Autolus; (iv) disclosure at least 30 days in advance of filing of any draft patent filing for any [***] Patent Rights intended to be
filed; and (v) details of all [***] Patent Rights that have been filed and their current prosecution status;

Autolus shall have the right (prior to filing and thereafter) to participate with UCLB and input into any draft patent filing for any [***]
Patent  Rights  that  are  filed  or  are  intended  to  be  filed  and  UCLB  shall  consider,  take  into  account  and  implement  any
reasonable recommendations made by Autolus with respect to such filings or proposed filings;

UCLB  shall  use  reasonable  endeavours  to  seek  appropriate  IP  protection  for  any  results  and  inventions  arising  from  the  [***]

Program;

UCL  and  UCLB  shall  ensure  that  all  Intellectual  Property  generated  by  MP  or  the  MP  Laboratory  in  connection  with  the  [***]

Program shall be owned by UCLB and shall be held subject to the terms of this Agreement; and,

UCLB  shall  not,  and  shall  procure  that  UCL  shall  not,  assign,  grant  (whether  conditional,  optional  or  otherwise)  any  rights  to,
encumber, charge, mortgage, license, sell, Exploit, or waive any rights over any of the [***] Program IP, nor enable another to
do so.

Automatically upon service of a [***] Licence Notice by Autolus, UCLB shall have granted to Autolus for the remaining duration of the
Term (from the date of such notice) and throughout the Territory, an exclusive (to the exclusion of UCLB, UCL and any Third Party)
licence  to  the  [***]  Program  IP  within  the  Field  which  shall  be  sub-licensable  through  multiple  tiers  to  use  the  same  for  any  act  of
Exploitation  concerning  any  product,  therapy,  service  or  process  without  restriction  (“[***]  Licence”).  Upon  the  grant  of  the  [***]
Licence, the licences to UCL Background IP, Background Materials and Manufacturing Know-How shall automatically be extended
such  that  references  to  “Royalty Products”  in  Clauses  2.1.1  and  2.3.1  shall  automatically  include  [***]  Products  and  the  relevant
schedules in respect of Patent Rights shall be updated. The [***] Licence (i) may only be revoked or terminated pursuant to a right of
termination  under  Clause  11.1.5(ii)  in  respect  of  the  [***]  Licence  or  Clause  24;  and  (ii)  may  be  assigned  but  only  as  part  of  the
assignment of this Agreement in accordance with Clause 29.

f.

If  Autolus  does  not  serve  a  [***]  Licence  Notice  on  UCLB  prior  to  expiry  of  the  [***]  Exercise  Period,  thereafter  without  otherwise
affecting any other Licences, including in each case the exclusivity granted in respect of the same, Autolus shall cease to have any
right or interest in or to the [***] Program IP and UCLB shall be free to develop and exploit the [***] Program IP as it thinks fit, without
any reference to Autolus.

g.

Option to a Licence under the NSG Program IP

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

UCLB  acknowledges  that  Autolus  has  exercised  its  option  for  the  grant  of  a  licence  on  the  terms  of  this  Agreement  to  the  NSG
Program IP with effect from the Amendment Date.

h.

Option to a Licence under the ZipCAR Program IP

UCLB acknowledges that Autolus has exercised its option for the grant of a licence on the terms of this Agreement to the ZipCAR
Program IP with effect from the Amendment Date.

i.

i..

ii..

3.
a.

i..

ii..

Schedule Updates

The Parties have agreed a list of Know-How and materials generated during the period commencing after the Effective Date and
ending  as  of  the  Amendment  Date  which  are  included  in  the  NSG  Licence  and  ZipCAR  Licence,  but  which  are  not
necessarily  exhaustive  of  all  Know-How  and  materials  licensed  under  the  NSG  Licence  and  ZipCAR  Licence.  The  Parties
further agree that such list, to the extent incomplete, may be updated from time to time pursuant to the provisions of Clause
9.1 to include Know-How and materials under the NSG Program and ZipCAR Program.

Upon exercise of the [***] Option the Parties shall, purely for convenience and future reference purposes, co-operate to agree a
list of Know-How and materials generated after the Effective Date that are included in the [***] Licence. The grant of the [***]
Licence  is  not  dependent  on  agreeing  such  list,  and  such  list  shall  not  fetter,  vary  or  otherwise  limit  the  terms  of  this
Agreement or the scope or inclusion of Know-How or materials under the [***] Licence unless the Parties expressly agree in
writing that it shall do.

SUB-LICENSING

Autolus shall be entitled to sub-license any of the Technology licensed to it hereunder or licence any of the Assigned Patents through
multiple tiers and without restriction save that (i) no sub-licence or licence may be granted to a Tobacco Party and (ii) UCLB shall
have  a  right  to  object  to  the  grant  of  a  sub-license  or  licence  by  Autolus  to  other  Third  Parties  solely  in  the  following  specific
circumstances:

UCLB may only object in respect of a proposed sub-licensee or licensee if, due to the nature of that proposed sub-licensee’s or
licensee’s  business,  the  grant  of  the  sub-license  or  licence  to  that  entity  will,  in  the  reasonable  and  measured  opinion  of
UCLB, have a material detrimental impact on the reputation of UCL by its association; and

if  the  circumstances  in  Clause  3.1.1  apply,  UCLB  shall  only  have  the  right  to  object  provided  that  it  serves  written  notice  of  its
objection setting out the grounds for its objection within [***] of written notice from Autolus of the identity of the proposed sub-
licensee.

b.

If UCLB has objected to the grant of a sub-license or licence in accordance with Clause 3.1, Autolus may either accept that objection

and not grant (or terminate) the sub-license or licence or if it disputes the objection the following shall apply:

UCLB shall procure that representatives from UCLB and UCL shall meet with Autolus within [***] of the objection to enable the
Parties and UCL to discuss the proposed sub-license or licence and the reasons for the perceived risk that an association
will have a material detrimental impact on UCL’s reputation and, in good faith, seek ways in which to overcome or mitigate
such risk to a pragmatic and reasonably acceptable position;

if UCLB agrees that the risk is acceptable or UCLB and Autolus agree on any conditions to include in a sub-license or licence to
avert  or  mitigate  the  risk  then  Autolus  shall  be  entitled  to  grant  (or  maintain)  the  sub-license  or  licence  subject  to  any
agreement reached between UCLB and Autolus;

if  within  [***]  of  the  objection,  (i)  UCLB  and  Autolus  are  unable  to  reach  an  agreement  and  Autolus  still  wishes  to  grant  (or
maintain) a sub-license or licence or (ii) representatives of UCLB and UCL do not or are unable to meet with Autolus; then
Autolus  shall  be  entitled  to  refer  the  objection  to  a  person  nominated  by  the  chairman  of  the  Wellcome  Trust  to  the
determination identified below (the “Appointed Expert”). The nomination shall be subject to the Appointed Expert agreeing
to be so appointed and the terms of that appointment set by the Wellcome Trust. The costs of the Appointed Expert shall be
borne by the Parties equally. The Appointed Expert shall be entitled to consider any information presented to the Appointed
Expert by UCLB or Autolus (provided that

i..

ii..

iii..

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

each Party shall copy to the other Party all information provided to the Appointed Expert at the same time) and any other
information that the Appointed Expert may consider relevant. The Appointed Expert shall make his or her decision as expert
and not as arbiter, and the decision of the Appointed Expert shall be final and binding save in the case of manifest error. If, in
the  Appointed  Expert’s  opinion  the  Appointed  Expert  considers  the  grant  of  a  sub-licence  or  licence  to  the  Third  Party
objected to by UCLB will, by virtue of the nature of the business of that Third Party entity, be materially detrimental to the
reputation of UCL, then Autolus shall not grant (or shall terminate) such sub-licence or licence. In all other circumstances,
irrespective  of  any  objection  by  UCLB  or  UCL,  Autolus  shall  be  entitled  to  grant  (or  maintain)  the  sub-licence  or  licence
without  restriction  or  condition.  UCLB  and  Autolus  hereby  irrevocably  agree,  accept  and  acknowledge  that  neither  the
Wellcome Trust, the Chairman of the Wellcome Trust nor the Appointed Expert shall have any liability to UCLB or Autolus (or
any Third Party or any other person) by virtue of the provisions of this Clause or exercise of decisions pursuant this Clause,
and UCLB and Autolus hereby undertake not to make or bring any claim against any of the Wellcome Trust, the Chairman of
the Wellcome Trust or the Appointed Expert with respect to performance in connection with the foregoing or this Agreement.

c.

i..

ii..

iii..

In so far as Autolus grants a sub-licence of rights under the Technology to a sub-licensee or a licence of the Assigned Patents to a
licensee (other than in respect of a material transfer agreement, contract research agreement or manufacturing agreement, provided
they do not include rights to sell products or services incorporating the Technology) (“Commercial Licence”), Autolus shall in such
circumstances enter into a written agreement with each sub-licensee or licensee (provided that this obligation to enter into a written
agreement shall not apply where, and for so long as, the sub-licensee or licensee is an Affiliate of Autolus). Additionally,

Autolus  shall  ensure  that  the  provisions  of  the  sub-licence  or  licence  agreement  do  not  grant  rights  in  the  Technology  beyond
those  granted  hereunder  and  impose  obligations  and  restrictions  on  the  sub-licensee  or  licensee  consistent  with  the
obligations and restrictions imposed on Autolus hereunder under (i) Clauses 3, 16 and 20 and (ii) Clauses 23.1.1 and 23.2 to
23.5 should any CRUK Rights be sub-licensed;

Autolus  shall  ensure  that  the  sub-licence  or  licence  agreement  imposes  obligations  of  confidentiality  on  the  sub-licensee  or

licensee which are no less onerous than those set out in Clause 20;

the  sub-licence  or  licence  agreement  shall,  if  required  by  Autolus,  be  novated  to  UCLB  (which  novation  UCLB  will  accept
provided that, in the case of the Assigned Patents, such Assigned Patents have been assigned to UCLB) on termination of
this Agreement, provided that (i) the sub-licensee or licensee is willing to accept the novation of any sub-licence or licence
agreement  upon  such  termination  and  make  payment  of  sums  otherwise  payable  under  this  Agreement  for  the  sub-
licensee’s (and its sub-sublicensees') Exploitation of the Technology or licensee’s (and its sub-licensees’) Exploitation of the
Assigned Patents directly to UCLB; (ii) at the time of novation the sub-licensee or licensee is not in breach of its obligations
to  Autolus  under  the  sub-licence  agreement;  (iii)  the  sub-licence  or  licence  agreement  does  not  impose  on  UCLB  any
obligations or commitments beyond those included in this Agreement; and (iv) the sub-licence or licence agreement includes
terms (at a minimum) consistent with those in Clauses 3, 11, 13, 14, 15, 16, 20 and 23, 24.4, 24.5, 24.6, 24.7 and 24.8 of this
Agreement failing which the sub-licence or licence agreement shall automatically terminate.

d.

e.

Autolus shall be liable to UCLB for all acts and omissions of its sub-licensees and licensees (other than those whose sub-license or
licence has novated to UCLB) that, if committed by Autolus, would constitute a breach of any of the provisions of this Agreement.
Autolus shall provide UCLB with written notice of any Commercial Licence and, to the extent it is able to do so, provide UCLB with a

copy of any Commercial Licence (with confidential and financial information redacted) promptly following its execution.

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

f.

4.
a.

i..

ii..

b.

i..

Notwithstanding the above, where the sub-licence relates only to UCL Background IP or Manufacturing Know-How the provisions of

Clauses 3.3.1, 3.3.3, and 3.5 shall not apply and such sub-licences shall not terminate on termination of this Agreement.
RETAINED RIGHTS, ACADEMIC RESEARCH & RESTRICTIONS, COLLABORATIVE PROGRAMS

The licences to Program IP and the Background Licence, to the extent each of them are exclusive (including the [***] Program IP), are
subject to: (a) UCLB reserving for and having the right to grant the limited non-assignable, worldwide, perpetual and irrevocable right,
and (b) Autolus hereby granting to UCLB a limited, non-exclusive, non-assignable, worldwide, royalty free, registrable, perpetual and
irrevocable licence under the Assigned Patents with the right to grant sub-licences, in each case:

to UCL to enable UCL to use the (i) licensed Program IP; and/or (ii) each of the UCL Background IP and Background Materials
forming part of the exclusive licence pursuant to Clause 2.1.1; and/or (iii) Assigned Patents, solely to teach and undertake
Academic Research at UCL and/or (subject to Clause 4.7) in collaboration with other Academic Organisations; and

to CRUK to enable CRUK, and those scientists employed at Academic Organisations funded by CRUK and/or those scientists
employed  by  CRUK  and  other  Funders  who  have  provided  funding  for  the  CRUK  Study  to  use  the  GD2  Clinical  Study
Results  to  undertake  academic  research  that  is  not  Commercial  Research  alone  or  in  collaboration  with  other  Academic
Organisations;

but for no other purposes (collectively the “Academic Rights”). For the avoidance of doubt, UCLB and/or UCL (including MP and
MP Laboratory) are not precluded under this Clause 4, from using and/or disclosing for any purpose without any restriction, the UCL
Background  IP  and  the  Background  Materials  that  are  subject  to  the  non-exclusive  licence  granted  to  Autolus  pursuant  to  Clause
2.1.2 where such use and/or disclosure does not relate to the use of any of the Program IP. Also, for the avoidance of doubt, this
Clause 4 shall not apply to the exercise of rights by UCLB, UCL or any Third Party under the RQR8 Program IP outside the RQR8
Program Field, the TetCAR Program IP outside the TetCAR Field or the CAT19 Program IP outside the CD19 Field.

The Academic Rights shall be subject to the following:

UCLB and UCL shall be entitled to apply for, obtain and use Third Party funding from a Funder for any of the Academic Research,

provided that:
(1) during  the  Improvement  Period  only  and  where  the  Academic  Research  relates  to  or  is  likely  to  give  rise  to
Improvements,  UCLB  and  UCL  shall  be  entitled  to  accept  such  Third  Party  funding  if  there  are  no  restrictions  or
conditions on UCLB and UCL by the Third Party Funder with respect to Exploitation of the funded Improvements other
than the requirement that UCLB and/or UCL seek the consent of the Funder prior to UCLB’s grant of any commercial
licence to exploit any such Improvement;

(2) during  the  Improvement  Period  only  and  where  the  Academic  Research  relates  to  or  is  likely  to  give  rise  to
Improvements,  UCLB  and  UCL  shall  not  without  the  prior  written  consent  of  Autolus  accept  any  such  funding  if  such
Third Party Funder imposes restrictions or conditions on the Exploitation of Improvements funded in whole or in part by
that Third Party in addition to the requirement in (i) above;

(3) during  the  Improvement  Period  only  and  where  the  Academic  Research  relates  to  or  is  likely  to  give  rise  to  CARs
developed by MP and/or the MP Laboratory (but such research does not relate to Improvements), UCLB shall use its
reasonable  endeavours  to  obtain,  or  not  accept  a  restriction  over,  commercialisation  rights  from  such  Third  Party
Funder,  in  which  case,  the  provisions  of  Clause  4.2.1(i)  and  (ii)  above  shall  apply  (and  the  Parties  acknowledge  that
where UCLB is unable to secure commercialisation rights from any Third Party Funder such Intellectual Property shall
not fall within the definition of New Inventions);

(4) during the [***] Period only and where the Academic Research relates to the [***] Program IP, UCLB and UCL shall be
entitled to accept such Third Party funding if there are no restrictions or conditions on UCLB and UCL by the Funder
with respect to Exploitation of the [***] Program IP, other than the

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

requirement  by  the  Third  Party  Funder  that  UCLB  and/or  UCL  shall  seek  the  consent  of  that  Funder  prior  to  UCLB’s
grant of any commercial licence to exploit any such [***] Program IP; and

(5) during the [***] Period only and where the Academic Research relates to the [***] Program IP, UCLB and UCL shall not
without the prior written consent of Autolus accept any such funding if such Third Party Funder imposes restrictions or
conditions on the commercial exploitation of [***] Program IP funded in whole or in part by that Third Party in addition to
the requirement in Clause 4.2.1(iv) above;

UCLB  shall  and  shall  procure  that  UCL  shall  maintain  the  confidentiality  of  the  Program  IP  (with  the  exception  of  information
disclosed in the Licensed Patents once published) and shall impose the same restrictions on its Academic Collaborators (and
enforce the same), save that UCL and its Academic Collaborators shall be entitled to publish the results of their Academic
Research under these Academic Rights subject to and in accordance with the provisions of Clause 4.5;

with  the  exception  of  the  CRUK  Study  and  the  Permitted  Studies,  or  as  otherwise  permitted  under  Clause  4.2.5,  no  studies
intended  to  be  (or  required  by  applicable  laws,  ethical  requirements  or  standards  or  otherwise  to  be)  conducted  in
accordance  with  the  standards  of  GLP  (good  laboratory  practice),  GMP  (good  manufacturing  practice)  and/or  GCP  (good
clinical practice) may be conducted under the Academic Rights without the prior written consent of Autolus, and UCLB shall
procure that an equivalent restriction is imposed on and complied with by all Academic Collaborators and shall ensure that
the Program Materials and Background Materials (that are subject to the exclusive licence granted pursuant to Clause 2.1.1)
are not provided to any Third Party for the purposes of any such study that has not been authorised by Autolus;

with the exception of the CRUK Study and the Permitted Studies, no clinical studies or treatment of patients may be conducted
under  the  Academic  Rights  except  for  academic  clinical  studies  conducted  with  the  involvement  of  MP  and/or  MP’s
Laboratory,  which  studies  shall  only  be  conducted  with  the  prior  written  consent  of  Autolus  (such  consent  not  to  be
unreasonably withheld or delayed), and UCLB shall procure that an equivalent restriction is imposed on and complied with by
all Academic Collaborators and shall ensure that the Program Materials and Background Materials (that are subject to the
exclusive licence granted pursuant to Clause 2.1.1) are not provided to any Third Party for the purposes of any such study
that has not been authorised by Autolus;

CRUK shall be permitted to use the GD2 Clinical Study Results for the purposes of clinical research with the prior written consent

of Autolus, such consent not to be unreasonably withheld or delayed;

UCLB  shall  procure  the  lawful  disclosure  to  Autolus  (free  of  any  restriction)  of  the  Regulatory  Submissions  for  each  of  the
CARPALL  Study,the  ALLCAR19  Study  and  the  CAROUSEL  Study  and,  subject  to  the  requirements  of  any  Regulatory
Authority,  shall  notify  Autolus  if  there  are  any  changes  to  the  Regulatory  Submissions  for  either  study.  Autolus  shall  be
entitled  to  comment  on  the  content  of  any  draft  Regulatory  Submission  and/or  any  proposed  changes  to  any  Regulatory
Submission for the CARPALL Study, the ALLCAR19 Study, or the CAROUSEL Study in advance of their submission to any
Regulatory  Authority.  UCLB  shall  procure  that  UCL  shall  consider  any  such  Autolus  comments  but  Autolus  acknowledges
that  UCL  shall  be  under  no  obligation  to  take  any  such  comments  into  account  and  the  contents  of  any  Regulatory
Submissions (and any amendments) shall be determined by UCL as sponsor of the CARPALL Study, the ALLCAR19 Study
and the CAROUSEL Study in its absolute discretion;

subject  to  the  requirements  of  any  Regulatory  Authority,  UCLB  shall  procure  that  UCL  shall  promptly  provide  to  Autolus
information regarding any Suspected Unexpected Serious Adverse Reactions or other adverse events resulting from either
the  CARPALL  Study,  the  ALLCAR19  Study  or  the  CAROUSEL  Study  and  which  are  reported  to  any  Regulatory  Authority;
and

ii..

iii..

iv..

v..

vi..

vii..

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

viii..

Clause 4.2.6 and 4.2.7 may be subject to additional terms agreed in any collaboration agreement entered into in accordance with

Clause 4.12;

save  that  where  Program  IP  is  in  or  comes  into  the  public  domain  (other  than  as  a  result  of  a  breach  of  any  obligation  of
confidentiality  owed  by  UCLB,  UCL  or  an  Academic  Collaborator),  the  provisions  of  this  Clause  4  shall  not  apply  in  respect  of
Academic  Research  relating  to  such  Program  IP,  except  where  the  Academic  Research  is  conducted  by  MP  and/or  the  MP
Laboratory in which case these provisions shall continue to apply.

The Parties acknowledge that UCLB, UCL and/or MP have entered or have agreed to enter into agreements governing the Permitted
Studies and that UCLB, UCL and/or MP are free to continue in those studies to the extent of the scope of such studies as outlined in
Schedule  7  and  the  provisions  of  this  Clause  4  shall  not  (with  the  exception  of  the  CRUK  Study  and  subject  to  Clause  4.11,  the
ALLCAR19 Study, the CARPALL Study and the CAROUSEL Study) apply to those Permitted Studies in that regard.

UCLB shall procure the lawful disclosure to Autolus (free of any restriction) of the protocol (once finalised) for the CRUK Study and,
subject to the requirements of any Regulatory Authority, shall notify Autolus if there are any changes to the protocol after it has been
finalised. The Parties acknowledge that RQR8 Permitted Studies listed with the identifiers DCAR19 and UCART19 in Schedule 7 are
being  conducted  with  a  commercial  Third  Party  and,  as  such,  Autolus  shall  have  no  access  to  the  protocol  or  results  from  those
RQR8 Permitted Studies.

If  UCL  or  any  Academic  Collaborator  (which  for  the  purposes  of  this  Clause  4.5  shall  include  any  of  their  respective  academics,
employees or students), wish to publish (including by way of publication of any thesis) (i) any of the Program IP; or (ii) with respect to
each Improvement or New Invention for the duration of the period prior to and during the applicable Improvement Negotiation Period,
any results of Academic Research that amount to Improvements or New Inventions; or (iii) for the duration of the [***] Period, any
results  of  Academic  Research  that  relate  to  [***]  Program  IP  (each  of  (i),  (ii)  and  (iii)  being  “Academic Information”)  then  UCLB
shall procure that:

UCL and the Academic Collaborator shall refrain from making any publication (or submitting for approval any publication) of any
of the information in the Academic Information (or the Academic Information in its entirety) pending conclusion of all steps
required under this Clause 4.5;

UCL  or  the  Academic  Collaborator(s)  must  first,  via  UCLB,  give  to  Autolus,  in  advance,  a  copy  of  the  proposed  publication
containing the Academic Information and which is intended to be disclosed or published, or submitted for publication at least
[***] before its presentation or intended submission for publication (“Academic Reports”);

upon  receipt  of  the  Academic  Reports,  Autolus  shall  within  [***]  of  receipt  either  approve  or  (where  Autolus  has  legitimate
commercial concerns including wanting to seek patent protection of the relevant Academic Information) refuse the request for
publication, and failing receipt of Autolus’s notice within the [***] time period, the request for publication shall be deemed to
be approved in the form in which the Academic Reports were provided to Autolus pursuant to Clause 4.5.2;

where the request for publication is refused, the refusal shall be communicated to UCLB in writing, following which UCLB shall
procure that UCL and the Academic Collaborator(s) shall refrain from making any publication of the Academic Reports (and
the Academic Information contained in such Academic Reports) for a period no less than [***] (or no less than [***] if agreed
by UCLB) from the date of notification from Autolus refusing the request for publication;

if consent is given to the request for publication, or where refused the period of [***] (or such longer period as agreed) pursuant to
Clause  4.5.4  has  expired,  UCL  and/or  the  Academic  Collaborator(s)  may  proceed  to  publish  the  Academic  Reports  in  the
form in which they were provided to Autolus pursuant to Clause 4.5.2;

notwithstanding  the  above,  this  Clause  4.5  shall  not  prevent  or  hinder  any  student  from  submitting  for  degrees  of  UCL  or  any
Academic  Collaborator  any  thesis  containing  Academic  Information  or  from  following  the  procedures  of  UCL  or  any
Academic Collaborator for examination and for admission to postgraduate degree

c.

d.

e.

i..

ii..

iii..

iv..

v..

vi..

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

f.

g.

i..

ii..

iii..

iv..

v..

vi..

h.

i.

status provided that such procedures require the thesis to be placed on restricted access within the library of UCL for at least
[***] from the date of placement of the thesis at the library of UCL.

Where  Autolus  has  exercised  its  right  to  refuse  the  request  for  publication  of  any  Academic  Reports,  then  Autolus  and  UCLB  shall
promptly and collaboratively work together in order to assess, and where appropriate, file and seek appropriate patent protection for
the Academic Information. Where UCLB files for patent protection, the costs for seeking such protection shall be borne by UCLB.
UCL shall have the limited right to sub-license the Academic Rights through one tier only, to Academic Collaborators for named staff to
work on with respect to a collaborative Academic Research project in conjunction with UCL (a “Collaborative Research Project”).
Such sub-licence shall:

only be granted provided that all results and Intellectual Property arising from a Collaborative Research Project shall be subject to
Autolus’s  right  to  exercise  its  rights  over  Improvements  in  accordance  with  the  provisions  of  Clause  5  (and  the  Parties
acknowledge that this obligation shall not apply in respect of results or Intellectual Property which are not Improvements);
be in writing and the Intellectual Property licensed, and Materials provided, shall be limited for use solely by the named individuals

at the Academic Collaborator and for that Collaborative Research Project only;

be subject to all the conditions and restrictions set out in this Clause 4, which UCL and UCLB shall ensure are binding on such

Academic Collaborator, with UCLB procuring compliance;

automatically terminate on conclusion of the Collaborative Research Project, and require the return of any Materials provided by

UCL (or its academics or laboratories) under or pursuant to the Collaborative Research Project;

not permit, and positively restrict, the Academic Collaborator from undertaking any of the studies referred to in Clauses 4.2.3 and

4.2.4 without Autolus’s prior written consent; and,

be promptly notified to Autolus, within [***] of grant of such sub-licence and, to the extent it is able to do so, provide Autolus with a
copy  of  the  agreement  with  respect  to  the  Collaborative  Research  Project  (with  confidential  and  financial  information
redacted) promptly following its execution.

UCLB shall procure the compliance of any Academic Collaborator(s) with the provisions of this Clause 4 by including these terms in an
agreement with the Academic Collaborator(s). UCLB shall be responsible to Autolus for any act or omission of a sub-licensee who is
granted a sub-license by UCL under Clause 4.7 where such act or omission if committed by UCL or UCLB would be a breach of this
Agreement.

UCLB warrants and represents to Autolus that the agreements disclosed to Autolus and identified in Part A of Schedule 6 comprise all
licences, consents, waivers and/or permissions (whether express or implied) granted by UCL and/or UCLB in respect of any of the
Program IP or UCL Background IP (that is subject to the exclusive licence pursuant to Clause 2.1.1) to any Third Party (“Existing
Licences”), save that Existing Licences shall not include arrangements relating to the use of the RQR8 Program IP outside of the
RQR8 Field or arrangements relating to the use of the UCL Background IP for purposes which are unrelated to any of the Program
IP or Royalty Products. Notwithstanding the foregoing, UCLB shall not be deemed in breach of the requirements or restrictions under
this Clause 4 with respect to those Existing Licences provided that such Existing Licences shall not be amended after the Effective
Date without the prior written consent of Autolus (not to be unreasonably withheld or delayed) nor shall their duration be extended or
consent granted under them to allow materials to be provided to a Third Party. The Parties confirm that UCL has agreed to provide to
[***] CAT19 Materials for use in the research collaboration between [***] and UCL referred to in Part A of Schedule 6 and that the
Existing Licence relating to such research collaboration shall apply.

j.

Save for the limited right granted to UCL under Clause 4.1 to undertake Academic Research, UCLB shall retain no other rights that
deviate from or otherwise encumber, limit or affect the licences (including their scope, termination and duration) granted to Autolus
hereunder.

k.

CARPALL, ALLCAR19 and CAROUSEL Studies

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

The Parties acknowledge that:

in  respect  of  the  ALLCAR19  Study,  UCL  has  various  reporting  commitments  to  [***]  and  [***]  is  entitled  to  use,  publish  and
disclose CD19 Clinical Study Results relating to the ALLCAR19 Study in accordance with the terms of the [***] Contract;
in respect of the CARPALL Study, UCL has various reporting commitments to [***], and [***] is entitled to use CD19 Clinical Study

Results relating to the CARPALL Study in accordance with the terms of the [***] Grant;

in  respect  of  the  CAROUSEL  Study,  UCL  has  various  reporting  commitments  to  [***]  in  accordance  with  the  terms  of  the

CAROUSEL Clinical Study Agreement;

in  respect  of  the  ALLCAR19  Study,  [***]  is  entitled  to  use  CD19  Clinical  Study  Results  relating  to  the  ALLCAR19  Study  in

accordance with the terms of the [***] Contract;

the  [***]  are  permitted  to  use,  publish  and  disclose  the  CD19  Clinical  Study  Results  in  accordance  with  the  terms  of  the

agreements entered into between [***] and UCL that govern the conduct of the relevant clinical study;

[***] is entitled to use, publish and disclose certain CD19 Clinical Study Results in accordance with the [***] Agreements;
where there is any conflict or inconsistency between the provisions of the CD19 Clinical Study Agreements and the provisions of
this Clause 4 in respect of the use, publication or disclosure of the CD19 Clinical Study Results, the provisions of this Clause
4  shall  not  apply  to  the  use,  publication  or  disclosure  of  the  CD19  Clinical  Study  Results  by  [***],  [***],  [***],  or  [***]  (as
applicable)  and  the  provisions  of  the  relevant  CD19  Clinical  Study  Agreements  shall  prevail,  (provided  that  any  such  use,
publication or disclosure may be subject to further terms agreed in any collaboration agreement entered into in accordance
with Clause 4.12);

where [***], [***] or [***] (as applicable) propose to make any publication or presentation which includes any of the CD19 Clinical
Study  Results  and  submits  the  same  to  UCL  for  review,  UCLB  shall  procure  that  UCL  shall  provide  a  copy  of  the  draft
publication  to  Autolus  for  review  as  soon  as  reasonably  practicable,  but  Autolus  acknowledges  that  it  may  receive  the
proposed publication or presentation less than [***] before its presentation or intended submission for publication;

subject  to  Clause  4.11.8,  Autolus  shall  be  entitled  to  review  any  draft  publications  relating  to  the  CD19  Clinical  Study  Results
submitted to UCL by [***], [***] or [***] (as applicable) in accordance with the principles set out in Clause 4.5, but subject to
Clause 4.11.10 and to the extent consistent with UCL’s rights under the relevant CD19 Clinical Study Agreements;

in respect of any draft publications or presentations relating to the CD19 Clinical Study Results and/or the CAROUSEL Clinical
Study Results, Autolus confirms that under Clause 4.5.3, Autolus’ grounds to refuse publication shall be for the purposes of a
delay to seek patent protection in respect of any CD19 Clinical Study Results and/or CAROUSEL Clinical Study Results (as
applicable) in accordance with Clause 4.6 or to prevent the publication of any of Autolus’ Confidential Information; and

where  there  is  any  conflict  or  inconsistency  between  the  provisions  of  the  CD19  Clinical  Study  Agreements  and/or  the
CAROUSEL Clinical Study Agreement and the provisions of this Clause 4, UCLB shall not be under any obligation to amend
the terms of any CD19 Clinical Study Agreement and/or the CAROUSEL Clinical Study Agreement (or otherwise impose on
the  parties  to  the  CD19  Clinical  Study  Agreements  and/or  CAROUSEL  Clinical  Study  Agreement  any  obligation  required
under  this  Clause  4  which  is  not  included  in  any  CD19  Clinical  Study  Agreement  and/or  the  CAROUSEL  Clinical  Study
Agreement)  and  UCLB  shall  not  be  in  breach  of  the  provisions  in  this  Clause  4  to  the  extent  of  any  such  conflict  or
inconsistency.  The  provisions  of  Clauses  4.7  and  4.8  shall  not  apply  to  the  CD19  Clinical  Study  Agreements  or  the
CAROUSEL  Clinical  Study  Agreement  and  the  conduct  of  the  CARPALL  Study  and  ALLCAR19  Study  shall  be  under  the
terms of the CD19 Clinical Study Agreements and the conduct of the CAROUSEL Study shall be under the terms of the or
the CAROUSEL Clinical Study Agreement.

i..

ii..

iii..

iv..

v..

vi..
vii..

viii..

ix..

x..

xi..

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

UCLB confirms that any site agreements in respect of the ALLCAR19 Study entered into with any CD19 Clinical Study NHS
Foundation  Trusts  after  the  Second  Amendment  Date  shall  be  on  substantially  the  same  terms  as  those  study  site
agreements entered into prior to the Second Amendment Date.

Autolus  and  UCL  propose  to  collaborate  on  the  ALLCAR19  Study  and  CARPALL  Study.  Autolus  shall  use  commercially  reasonable
efforts to enter into collaboration and funding agreements with UCL with respect to i) the ALLCAR19 Study as soon as practicable
after the Second Amendment Date and ii) any extension of the CARPALL Study as soon as practicable after UCLB has procured the
release of all the CARPALL and ALLCAR19 Patient Clinical Data generated before the Second Amendment Date. UCLB shall use
commercially reasonable efforts to facilitate such discussions between UCL and Autolus with a view to two separate collaboration
agreements being entered into within this timescale. Such agreements will include terms typical for a clinical research collaboration,
including  a  joint  governance  structure  to  lead  this  program,  and  a  contribution  of  expertise  and  funding  to  be  agreed  in  order  to
establish  a  manufacturing  process  suitable  for  commercial  scale  and  lay  the  foundation  for  a  registration  study.  The  total  value  of
such commitment by Autolus shall be mutually agreed by Autolus and UCL in the applicable collaboration and funding agreement.
In respect of the ALLCAR19 Study, notwithstanding the provisions of Clause 4.12, Autolus confirms that, as a minimum, it shall pay to
UCL  the  sum  of  £875,000  (eight  hundred  and  seventy-five  thousand  pounds),  being  [***]  (the “Autolus  Funding  Commitment”).
The  Parties  acknowledge  that  the  consent  given  [***]  to  permit  the  licensing  of  the  Test  and  Regulatory  Data  arising  from  the
ALLCAR19 Study to Autolus under this Agreement is dependent on the Autolus Funding Commitment.

The parties intend that the collaboration agreement relating to the ALLCAR19 Study referred to in Clause 4.12 shall include details of
Autolus’ total funding commitment in respect of the ALLCAR19 Study (which may be higher than the Autolus Funding Commitment,
but not lower), together with payment terms. The Autolus Funding Commitment is binding on Autolus, irrespective of whether or not
the  parties  enter  into  any  such  collaboration  agreement.  Autolus  shall  pay  the  Autolus  Funding  Commitment  to  UCL  in  quarterly
instalments. UCL shall be entitled to invoice Autolus for the Autolus Funding Commitment in quarterly instalments in such amounts
as  reflect  the  instalments  payable  by  [***]  under  the  [***]  Contract,  as  amended  from  time,  and  the  parties  acknowledge  that  the
quarterly instalments will vary in amount from quarter to quarter. UCL shall invoice Autolus for the first quarterly instalment in April
2018. Autolus shall pay all invoices within [***] of the date of receipt of the invoice from UCL.

ACCESS TO IMPROVEMENTS AND NEW INVENTIONS

The provisions of this Clause 5 shall apply to each and every Improvement and New Invention that is generated, reduced to practice or
otherwise  discovered  or  identified  at  any  time  up  until  and  including  the  [***]  anniversary  of  the  Effective  Date  (“Improvement
Period”). For  clarity,  in  respect  of  CD19  CARs  or  binders,  the  Improvement  Period  shall  commence  on  the  Second  Amendment
Date. UCLB shall procure that UCL, MP and the MP Laboratory shall comply with the terms of this Clause 5 and notify UCLB of all
Improvements and New Inventions in order that UCLB shall be able to comply with this Clause 5.

From the Effective Date until expiry of the Improvement Period, UCLB shall notify Autolus of each and every Improvement and New
Invention generated, developed or arising in such period within [***] of UCLB’s receipt of an invention disclosure form describing the
Improvement  or  New  Invention  (in  respect  of  each  Improvement  and  each  New  Invention,  each  notification  to  Autolus  being  a
“Disclosure Notification”). In respect of Improvements and New Inventions which UCLB is obliged to notify to Autolus:

UCLB shall procure the exclusive right (to the exclusion of UCL and the inventor(s)) to license, assign, exploit or otherwise grant
any rights to that Improvement or New Invention (as applicable) to, any Third Party, and shall ensure that such Improvement
or  New  Invention  (as  applicable)  is  kept  confidential  including  by  its  Academic  Collaborators  (subject  to  the  publications
procedure under Clause 4.5) until Autolus ceases to have any rights (exercisable,

l.

m.

n.

5.
a.

b.

i..

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

ii..

iii..

iv..

v..

c.

d.

i..

ii..

iii..

negotiable or otherwise) to that Improvement or New Invention (as applicable) under the terms of this Clause 5;

UCLB shall procure that the Disclosure Notification shall provide (i) reasonable details of that Improvement or New Invention (as
applicable)  with  a  similar  level  of  detail  as  that  which  UCLB  provided  to  Syncona  Management  LLP  for  the  purpose  of
Syncona  Management  LLP  evaluating  the  Program  IP  and  (ii)  any  draft  or  pending  application  for  registered  patent
protection  in  respect  of  that  Improvement  or  New  Invention  (as  applicable);  in  each  case  disclosure  to  Autolus  shall  be
subject to the confidentiality obligations of this Agreement;

UCLB shall procure that Autolus shall have reasonable access to those individuals at UCL that invented, generated, discovered
or developed that Improvement or New Invention (as applicable) in order to allow a confidential discussion as to the nature
and features of that Improvement or New Invention (as applicable) and its application;

UCLB  shall  not,  until  Autolus  ceases  to  have  any  rights  (exercisable,  negotiable  or  otherwise)  to  that  Improvement  or  New
Invention  (as  applicable)  under  the  terms  of  this  Clause  5,  encumber,  charge,  mortgage,  license,  sell,  assign,  Exploit  or
otherwise grant any other right or enable any Third Party to Exploit the Improvement or New Invention (as applicable);

provided  that  Autolus  acknowledges  that  Improvements  and  New  Inventions  may  be  subject  to  the  rights  of  Funders  in
accordance with Clause 4.2.1 and that Autolus’s ability to exercise its rights under this Clause 5 may be dependent on any
such  Funder  providing  its  consent  to  the  terms  of  Exploitation  as  agreed  by  the  Parties  in  accordance  with  Clause  4.2.1,
which UCLB shall use reasonable endeavours to obtain; In particular the consent of CRUK, KK and LLF may be required in
respect of any licence of Improvements relating to TRBC1/2 and the consent of KK and LLF may be required in respect of
any licence of Improvements relating to BCMA CAR.

Following the date of the Disclosure Notification, Autolus shall have the right, exercisable at any time up until expiry of the Improvement
Negotiation Period (irrespective of whether that is before or after the Improvement Period) to exercise its right of first negotiation in
respect  of  obtaining  a  worldwide,  assignable,  sub-licensable  (through  multiple  tiers)  licence  within  the  Field  of  the  applicable
Improvement  or  New  Invention  that  is  the  subject  of  the  Disclosure  Notification,  such  licence  to  be  exclusive  or  non-exclusive,  as
agreed by the Parties.

Upon Autolus exercising its right of first negotiation in respect of an Improvement or a New Invention (as applicable) by way of serving
a  written  notice  on  UCLB,  the  following  shall  apply  until  expiry  of  the  applicable  Improvement  Negotiation  Period  for  that
Improvement or New Invention (as applicable), unless extended by agreement between the Parties:

unless the Parties agree to terminate negotiations during the Improvement Negotiation Period, Autolus and UCLB shall promptly
and  actively  negotiate  throughout  the  Improvement  Negotiation  Period,  in  good  faith  and  acting  reasonably,  fair  and
reasonable  terms  for  and  a  conclusive  agreement  upon  which  that  Improvement  or  New  Invention  (as  applicable)  may  be
licensed to Autolus;

in so far as UCLB fails to comply with the provisions of Clause 5.4.1, the Improvement Negotiation Period shall be extended by a
period  equal  to,  or  otherwise  fairly  calculated  to,  compensate  for  any  delay  in  or  absence  from  a  negotiation  by  UCLB  in
accordance with the principles under Clause 5.4.1;

in its negotiations around the fair and reasonable financial and other terms for a licence of Improvements (as opposed to New
Inventions), UCLB shall have regard to the existing licensing and financial structure under this Agreement with respect to the
Royalty Product to which such Improvement relates or is applicable.

e.

If Autolus, by written notice, elects not to continue with negotiations over an Improvement or a New Invention, then without prejudice to
the remainder of this Clause 5 or any other Improvements or New Inventions, the Parties shall be released from their then current

    28

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

obligations to negotiate in accordance with Clause 5.4 with respect to that particular Improvement or New Invention.

f.

Subject to UCLB’s compliance with Clause 5.4.1, if the Improvement Negotiation Period has expired with respect to an Improvement or

New Invention (as applicable), and that Improvement or New Invention (as applicable) has not been licensed to Autolus, then:

i..

ii..

in respect of that Improvement, subject to the terms of Clause 5.7, UCLB shall be entitled to (i) instigate negotiations with Third
Parties for the grant of a licence of that Improvement; or (ii) engage in negotiations solicited by Third Parties to agree terms
for the grant of a licence of that Improvement to that Third Party; or

in  respect  of  that  New  Invention,  Autolus  shall  cease  to  have  any  further  rights  to  negotiate  or  match  terms  for  any  licence  of
rights  to  that  New  Invention  under  this  Clause  5,  save  that  the  foregoing  shall  be  without  prejudice  to  any  other  New
Inventions.

g.

In negotiating with any Third Party to grant a licence of an Improvement to that Third Party, for a period of [***] after the expiry of the

Improvement Negotiation Period (“Match Period”) or Autolus having served notice under Clause 5.5:

i..

ii..

h.

i.

6.
a.

UCLB  shall  not  accept  or  agree  any  term  (or  overall  set  or  combination  of  terms)  that  are  equal  to  or  less  preferential  or
favourable  to  UCLB  than  the  most  preferential  or  favourable  matching  or  equivalent  term  (or,  as  applicable,  matching  or
equivalent set or combination of terms) as offered by Autolus to UCLB; and,

where UCLB has reached an agreement (in principle) on all material terms for the licence of the Improvement with the Third Party
(which is in compliance with Clause 5.7.1), but prior to concluding the definitive binding agreement on such terms with the
Third Party, UCLB shall promptly first offer such terms to Autolus for acceptance within the Match Period whereupon:
(1) during such Match Period UCLB shall not execute or enter into such agreement with any Third Party or otherwise look to

license or otherwise Exploit or grant rights to Exploit the Improvement with any Third Party; and,
if such terms are accepted by Autolus, UCLB shall conclude the definitive agreement with Autolus on those same terms
(or, if the Parties agree otherwise, more preferential or favourable terms for Autolus); or,
if rejected by Autolus, UCLB shall conclude the definitive agreement with that Third Party on those same terms.

(2)

(3)

The provisions of Clause 5.7 shall apply each and every time that UCLB instigates in or engages in negotiations with any Third Party
concerning  an  Improvement  during  the  Match  Period,  such  that  if  negotiations  with  a  Third  Party  break  down  and  either  re-
commence with that Third Party or new negotiations begin with a different Third Party to that Improvement, the provisions of Clause
5.7 shall continue to apply again.

Where  during  the  Improvement  Period  MP  and/or  any  member  of  the  MP  Laboratory  generates,  identifies  or  invents  any  discovery,
invention (whether patentable or not), Know-How or improvement concerning any CAR or CARs (including methods for manufacture
of any CAR or CARs) which is not an Improvement and which has been discovered, generated, identified or invented together with
another  Academic  Organisation(s)  such  that  UCLB  does  not  Control  the  same  and  it  does  not  fall  within  the  definition  of  New
Inventions, UCLB shall use reasonable endeavours to agree with the other Academic Organisation(s) that UCLB shall be the lead
commercialisation partner and that the relevant technology will be offered to Autolus in accordance with this Clause 5.
ACCESS TO [***] PROGRAM

The provisions of this Clause 6 shall apply to the [***] Program as it is developed from time to time up until the [***] anniversary of the
Effective Date (“[***] Period”). UCLB shall procure that UCL and [***] shall comply with the terms of this Clause 6 and notify UCLB
of  the  progress,  development  and  generation  of  IP  under  the  [***]  Program  in  order  that  UCLB  shall  be  able  to  comply  with  this
Clause 6. During the [***] Period, UCLB shall keep Autolus appraised on a reasonable basis of the progress and development of the
[***] Program and the inventions arising therefrom.

b.

If at any time during the [***] Period, any bona fide decision is made by UCLB, UCL or [***] to seek a licensee to Exploit any of the [***]

Program IP or any bona fide approach is

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

made  by  a  Third  Party  to  seek  rights  to  or  under  any  of  the  [***]  Program  IP,  and  in  respect  of  which  UCLB  genuinely  intends  to
engage in, UCLB shall provide written notice to Autolus together with (under the confidentiality obligations of this Agreement):

a description of the [***] Program and related [***] Program IP; and
a copy of any application for Patent Rights in respect of any invention arising from the [***] Program that has been filed in respect

of inventions arising from the [***] Program.

Following service of a notice in accordance with Clause 6.2:

at  Autolus’s  request,  UCLB  shall  use  reasonable  endeavours  to  procure  that  Autolus  shall  have  access  to  those  individuals  at
UCL that work on the [***] Program or invented, generated, discovered or developed the [***] Program IP in order to allow a
confidential discussion as to the nature and features of the [***] Program and the [***] Program IP and its application; and,
Autolus shall have the option to negotiate with UCLB to agree terms for the grant of a licence (which may be exclusive or non-

exclusive as the Parties agree) to [***] Program IP.

Upon exercise of its right to negotiate with UCLB, UCLB and Autolus shall negotiate the terms for such licence in good faith and acting
reasonably and seeking fair and reasonable commercial terms and Autolus’s right to negotiate with Autolus shall expire [***] following
Autolus’s receipt of UCLB’s notice of served under Clause 6.2.

If Autolus, by written notice, elects not to continue with negotiations over the [***] Program IP, then the Parties shall be released from
their then current obligations to negotiate in accordance with Clause 6.3.2. If the Parties are unable to agree licence terms within the
[***] period referred to in Clause 6.4 above, then UCLB shall be under no further obligation to Autolus and shall be free to license or
commercialise the same [***] Program IP as it thinks fit.

UCLB confirms that save for having to seek the consent of the Wellcome Trust and the National Institute for Health Research for the
Exploitation of the [***] Program IP, it otherwise Controls that [***] Program IP existing as of the Effective Date. For the avoidance of
doubt,  nothing  in  this  Agreement  shall  impose  any  restrictions  or  controls  on  UCL’s  ability  to  publish  and  to  undertake  Academic
Research  or  obtain  Third  Party  funding  from  a  Funder  for  Academic  Research  or  otherwise  collaborate  with  any  Academic
Organisation in respect of the [***] Program and, as a result, the [***] Program IP may be subject to rights of Third Party Funders and
may  not  be  Controlled  by  UCLB.  UCLB  shall  procure  that  during  the  [***]  Period,  UCL  shall  not  and  [***]  shall  not  undertake  any
Commercial Research with respect to the [***] Program or accept Third Party funding other than from Funders for the [***] Program.
Autolus acknowledges that UCLB or UCL shall not under any circumstances be required to restrict [***] from undertaking any work
(academic or commercial) either independently or in collaboration with any Third Party, on any program (including a [***] program)
that is not a [***] Program.

Upon  the  Parties  concluding  terms  for  a  license  to  the  [***]  Program  IP  as  provided  for  in  this  Clause  6,  UCLB  shall  procure  full

disclosure to Autolus of the [***] Program including the [***] Program IP and relevant UCL Background IP used for the same.

Until the expiry of the [***] Period, UCLB shall not, and shall procure that UCL and [***] shall not, grant any rights to, license, assign,
grant  an  option  to,  Exploit,  or  otherwise  encumber,  mortgage,  charge  or  otherwise  waive  rights  in  respect  of  the  [***]  Program  IP,
save  that  this  shall  not  prevent  UCLB,  UCL  and/or  [***]  from  securing  funding  from  Funders  pursuant  to  Clause  6.6  above  and/or
collaborating with Academic Organisations.
ACCESS TO [***]

The provisions of this Clause 7 shall apply to the [***] Program as it is developed from time to time up until expiry of the [***] Period.
The “[***] Period”  shall  be  for  a  minimum  of  [***]  from  the  Effective  Date,  and  may  be  extended  by  Autolus  (upon  written  notice
served  prior  to  the  [***]  anniversary  of  the  Effective  Date)  for  the  Extended  Period.  The “Extended  Period”  shall  mean  the  day
commencing on the [***] anniversary of the Effective Date and ending on the earlier of (i) Autolus and UCLB concluding a definitive
agreement for a licence to any of the [***] Program IP; (ii) Autolus terminating its rights under this Clause 7 by written notice; or (iii)
the [***] anniversary of the Effective Date.

i..
ii..

c.

i..

ii..

d.

e.

f.

g.

h.

7.
a.

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

b.

c.

d.

i..

ii..

iii..

The  Parties  acknowledge  and  agree  that  Autolus  has  served  notice  to  extend  the  [***]  Period  beyond  its  [***],  and  is  under  the
obligation to reimburse UCLB [***] of those Patent Prosecution Costs incurred by UCLB for activities during the Extended Period in
respect of the [***] Patents that are the subject of the [***] Option. Autolus shall not be responsible for any Patent Prosecution Costs
incurred before or after the Extended Period.

UCLB shall and shall procure that UCL (including MP and the MP Laboratory) shall comply with the terms of this Clause 7 and notify
UCLB of the progress, development and generation of Intellectual Property under the [***] Program in order that UCLB shall be able
to comply with this Clause 7.

From the Effective Date until expiry of the [***] Period, UCLB shall:

from time to time (and upon reasonable request from Autolus):

(1) notify Autolus of the development, progress and advances made in the [***] Program since the last update report was

provided;

(2) notify Autolus of any change in status of MP’s position at UCL (including termination of the same) or the MP Laboratory;
(3) disclose to Autolus (under the confidentiality obligations of this Agreement) any planned application for a Patent Right in
respect  of  any  invention  arising  from  the  [***]  Project  at  least  [***]  prior  to  it  being  filed,  and  thereafter  keep  Autolus
informed of all developments in the prosecution of such Patent Right(s); and,

(4) procure the exclusive right (to the exclusion of UCL and the inventor(s)) to license, assign, exploit or otherwise grant any
rights to the [***] Program IP until Autolus ceases to have any rights (exercisable, negotiable or otherwise) to the [***]
Program under the terms of this Clause 7;

use reasonable endeavours to procure that Autolus shall have access to those individuals at UCL that work on the [***] Program
or invented, generated, discovered or developed any of the [***] Program IP in order to allow a confidential discussion as to
the nature and features of the [***] Program and the [***] Program IP and its application;

not,  until  Autolus  ceases  to  have  any  rights  (exercisable,  negotiable  or  otherwise)  to  the  [***]  Program  under  the  terms  of  this
Clause 7, encumber, charge, mortgage, license, sell, assign, Exploit or otherwise grant any other right or enable any Third
Party to Exploit the [***] Program IP,

provided that Autolus acknowledges that certain [***] Program IP (i) existing as of the Effective Date is subject to those rights of Third
Party  Funders  which  are  disclosed  in  Schedule  6  and  described  as  having  funded  such  [***]  Program  IP;  and  (ii)  arising  after  the
Effective Date may be subject to the rights of Third Party Funders in accordance with Clause 4.2.1; and that in case of (i) and (ii)
Autolus’s ability to exercise its rights under this Clause 7 in respect of such [***] Program IP may be dependent on any such Third
Party Funder providing its consent to the terms of Exploitation as agreed by the Parties in accordance with Clause 4.2.1; In particular
consent of [***] may be required in respect of any licence of [***] Program IP developed prior to or after the Effective Date.

e.

Autolus shall have the right, exercisable at any time up until expiry of the [***] Period to exercise its right of first negotiation in respect of
obtaining a worldwide, assignable, sub-licensable licence (which may be exclusive or non-exclusive as the Parties may agree), within
the Field of the [***] Program IP (or, as the Parties may agree, any part thereof).

f.

Upon  Autolus  exercising  its  right  of  first  negotiation  in  respect  of  the  [***]  Program  by  way  of  serving  a  written  notice  on  UCLB,  the

following shall apply until expiry of the [***] Period (unless extended by agreement between the Parties):

unless the Parties agree to terminate negotiations during the [***] Period, Autolus and UCLB shall promptly and actively negotiate
throughout that period, in good faith and acting reasonably, fair and reasonable terms for a conclusive agreement upon which
the [***] Program IP may be licensed to Autolus;

in  so  far  as  UCLB  fails  to  comply  with  the  provisions  of  Clause  7.6.1,  does  not  actively  and  properly  participate  in  such
negotiations or does not act reasonably or in good faith, the [***] Period shall be extended by a period equal to, or otherwise
fairly calculated to, compensate for any delay in, or absence from a negotiation by UCLB in accordance with the principles
under Clause 7.6.1.

i..

ii..

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

g.

h.

If Autolus, by written notice, elects not to continue with negotiations over the [***] Program IP, then the Parties shall be released from

their then current obligations to negotiate in accordance with Clause 7.6.

Subject to UCLB’s compliance with Clause 7.6.1, upon expiry of the [***] Period and provided that Autolus has not been granted an
exclusive  license  to  the  [***]  Program  IP,  UCLB  shall  then  be  entitled  to  negotiate  with  any  Third  Party  to  grant  a  licence  to  or
otherwise Exploit the [***] Program IP subject always to the BCMA Licence with respect to the [***] Existing Patent.

i.

Upon  the  Parties  concluding  terms  for  a  license  to  the  [***]  Program  IP  as  provided  for  in  this  Clause  7,  UCLB  shall  procure  full

8.
a.

b.

i..

ii..
iii..
iv..

c.

disclosure to Autolus of the [***] Program including the [***] Program IP and relevant UCL Background IP used for the same.
The provisions of this Clause 7 shall be without prejudice to UCLB’s obligations under Clause 18 with respect to the [***] Existing
Patent.
INFORMATION AND ACCESS TO MARTIN PULE’S OTHER PROGRAMS & OTHER RESTRICTIONS

During  the  Improvement  Period,  the  Parties  shall  hold  regular  meetings  to  review  the  activities  of  MP  and  the  MP  Laboratory.  Such

meeting shall be held every month, or at such intervals as the Parties shall otherwise agree.

At each meeting UCLB shall update Autolus (under the confidentiality obligations of this Agreement) in respect of:

research  activities  being  undertaken  by  MP  and  the  MP  Laboratory,  together  with  progress  and  advances  made  under  all

research activities since the last meeting was held;

any change in status of MP’s position at UCL (including termination of the same) or the MP Laboratory;
any collaboration with Academic Collaborators entered into by MP and/or the MP Laboratory; and,
any proposed applications for Patent Rights;

provided  that  the  foregoing:  (i)  shall  not  require  any  disclosure  of  research  activities  being  undertaken  by  MP  on  behalf  of  [***]
pursuant  to  his  [***]  existing  as  of  the  Effective  Date;  and  (ii)  shall  not  require  disclosure  of  any  of  research  activities  being
undertaken  by  MP  and/or  the  MP  Laboratory  where  UCL,  MP  and/or  MP  Laboratory  are  restricted  from  disclosing  any  such
information as a result of obligations of confidentiality owed to a Third Party.

For any new inventions generated, reduced to practice or otherwise developed by or under the supervision of MP (whilst employed at
UCL) and/or the MP Laboratory (including therapeutic immune therapies, engineered T-cells, CARs or BiTEs, and any manufacturing
techniques and/or research tools relating to the foregoing) from time to time up until the [***] anniversary of the Effective Date and in
respect  of  which  Autolus  does  not  have  any  rights  pursuant  to  Clauses  5,  6  or  7  (“Other Technology”),  UCLB  shall  provide  and
procure that Autolus shall have a right of first review according to Clause 8.4.

d. Where Other Technology is Controlled by UCLB or UCL and where UCLB or UCL makes any bona fide decision to seek a licensee to
Exploit  any  Other  Technology,  or  any  bona  fide  approach  is  made  by  a  Third  Party  to  seek  rights  to  or  under  any  of  the  Other
Technology which UCLB or UCL genuinely intends to engage in, UCLB shall not, and shall procure that UCL shall not, disclose the
Other Technology to any Third Party for the purposes of instigating or encouraging any licensing discussions nor offer to license the
Other Technology to any Third Party, in each case prior to disclosing to Autolus at least [***] in advance (and in no less detail than it
will disclose to any Third Party) the same Other Technology.

Commercial Restrictions

The Parties acknowledge that Autolus’s business is primarily based upon the Exploitation of CARs and this licence of Technology
and involvement of MP as an employee to Autolus is crucial and fundamental to its business. Accordingly, in recognition of
the foregoing UCLB hereby agrees and undertakes that for a period of [***] after the Effective Date UCLB shall procure that
for so long as MP is employed by or holds any position or undertakes or supervises any research at UCL during such period,
that with respect to any CARs:

e.

i..

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

(1) MP shall not undertake any Commercial Research himself or through the MP Laboratory;
(2) without Autolus’s prior written consent, neither MP or UCL shall accept or use any funding to enable MP to undertake

any Commercial Research himself or through the MP Laboratory; and

(3) without Autolus’s prior written consent, MP shall not participate in, contribute to or otherwise supervise any Commercial

Research;

save that the foregoing shall not prevent MP (i) undertaking that Commercial Research which is on-going as of the Effective
Date  and  which  is  the  subject  of  the  those  agreements  disclosed  to  Autolus  and  listed  in  Part  B  of  Schedule  6,  (ii)  from
undertaking any Approved Activity as such term is defined in the SSA (effective at the same date as the Effective Date); and
(iii)  from  undertaking  Commercial  Research  on  behalf  of  Autolus.  For  the  avoidance  of  doubt,  the  provisions  of  this
Agreement do not apply to any work (commercial, academic or otherwise) conducted in relation to CARs in any laboratory of
UCL without any assistance from and/or involvement of MP.

Should Autolus be sold (comprising the transfer of all shares in Autolus to a Third Party or the sale of all assets of the business of
Autolus to a Third Party) and MP ceases to be employed or provide consultancy to Autolus (in the case of a share sale) or
the Third Party acquirer (in the case of either a share sale or asset sale) then the restriction under Clause 8.5 shall terminate
on the earlier of (i) [***] after the Effective Date; or (ii) [***] following the completion of the relevant sale.

MATERIALS TRANSFER AND ENABLEMENT OF THE LICENSED RIGHTS

From the Effective Date and thereafter until the [***] anniversary of the Effective Date:

UCLB  shall  procure  the  disclosure  to  Autolus  by  UCL,  MP  and  the  MP  Laboratory  of  all  Technology  licensed  hereunder  in

accordance with the timeline and practical disclosure steps set out in Schedule 5;

After compliance with Clause 9.1.1, UCLB shall thereafter continue to disclose to Autolus, and procure the disclosure to Autolus
by UCL, MP and the MP Laboratory, of any Technology not disclosed under Clause 9.1.1, at Autolus’s request from time to
time;

UCLB  shall  procure  that  each  of  UCL,  MP  and  the  MP  Laboratory  shall  help  facilitate  any  technology  transfer  or  teach-in

(including any demonstrations) concerning any of the Technology; and,

it being understood that such disclosure should be in the English language and should be disclosed in a structured and helpful
manner  to  enable  the  proper  understanding,  benefit  and  access  to  the  technology  in  respect  of  each  Program  and  the
Program IP.

ii..

9.
a.

i..

ii..

iii..

iv..

b. Where the Technology comprises Materials, UCLB shall procure the delivery of a reasonable quantity of such Materials and UCL shall
itself be entitled to retain a reasonable quantity of the Materials for the exercise of its Academic Rights subject to and in accordance
with the provisions of Clause 4.

c. With  respect  to  materials  which  as  of  the  Effective  Date  are  not  Program  Materials,  have  been  used  in  connection  with  any  of  the
Original Programs, but are not Controlled by UCLB and so do not fall within the definition of Background Materials, UCLB shall use
its  reasonable  endeavours  to  obtain  consent  for  the  transfer  of  such  materials  to  Autolus  (and  for  the  licensing  of  associated
Intellectual Property) or otherwise assist Autolus in obtaining access to and the right to use such materials. Where UCLB uses its
commercially  reasonable  efforts  to  obtain  such  consent,  it  shall  not  be  obliged  to  make  any  payment  but,  should  any  payment  be
demanded by the relevant Third Party Autolus may, at its discretion, make such payment to obtain consent. This Clause 9.3 shall not
apply in respect of materials that are “off the shelf”, such as reagents and other commercially available Third Party materials.

UCL shall retain ownership and possession of all Laboratory Notebooks and UCLB shall procure throughout the Term:

physical  access  for  Autolus  (including  the  right  for  Autolus  to  physically  borrow  from  UCL’s  possession  and  copy),  upon
reasonable notice, of any Laboratory Notebooks in so far as they concern any of the Programs, Program IP, Materials or UCL
Background IP;

d.

i..

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

ii..

iii..
e.

f.

that  all  Laboratory  Notebooks  shall  be  kept  reasonably  safe  and  secure  and  protected  from  loss,  damage  or  destruction  in

accordance with standard UCL process; and,

that all Laboratory Notebooks shall not be destroyed without first offering the same to be transferred to Autolus.

UCLB  shall  co-operate  and  collaborate  with  Autolus  to  provide,  and  procure,  guidance,  information  and  know-how  from  time  to  time

from MP and MP Laboratory about Program IP and the Technology.

UCLB shall not and shall procure that UCL (including MP and the MP Laboratory) shall not transfer, lend, supply or otherwise provide
(i) any Laboratory Notebooks to any Third Party and/or (ii) any Program Materials to any Third Party except (a) in the exercise of
UCL’s Academic Rights subject to and in accordance with the provisions of Clause 4; and/or (b) with respect to those specific parts
of the Laboratory Notebooks or specific Program Materials under (1) the RQR8 Program IP in the exercise of UCLB’s rights to the
extent  permitted  outside  of  the  RQR8  Field;  and/or  (2)  the  TetCAR  Program  IP  in  the  exercise  of  UCLB’s  rights  to  the  extent
permitted outside of the TetCAR Field.

g. Without prejudice to Clause 4.4, UCLB shall procure the transfer to Autolus of a copy of the Test and Regulatory Data resulting from
the  CRUK  Study,  once  such  Test  and  Regulatory  Data  has  been  released  by  the  sponsor  and  UCLB  has  authority  from  any
applicable Regulatory Authority to disclose the same to Autolus. Upon the disclosure of such Test and Regulatory Data to Autolus,
the GD2 Licence shall automatically be extended to include a licence to the GD2 Clinical Study Results and the GD2 Program IP
shall be deemed to include GD2 Clinical Study Results.

h.

i.

UCLB  shall  procure  the  transfer  to  Autolus  of  a  copy  of  the  Test  and  Regulatory  Data  resulting  from  the  CARPALL  Study  and
ALLCAR19  Study  which  shall  include  the  CARPALL  and  ALLCAR19  Patient  Clinical  Data  once  such  CARPALL  and  ALLCAR19
Patient Clinical Data has been released by the sponsor and UCLB has authority to disclose the same to Autolus. The frequency and
timing  of  such  transfers  of  the  CARPALL  and  ALLCAR19  Patient  Clinical  Data  is  detailed  in  Schedule  11.  Upon  the  disclosure  of
such  CARPALL  and  ALLCAR19  Patient  Clinical  Data  to  Autolus,  the  CAT19  Licence  shall  automatically  be  extended  to  include  a
licence to the CARPALL and ALLCAR19 Patient Clinical Data released and the CAT19 Program IP shall be deemed to include such
CARPALL and ALLCAR19 Patient Clinical Data.

UCLB shall procure the transfer to Autolus of a copy of the Test and Regulatory Data resulting from the CAROUSEL Study which shall
include the CAROUSEL Patient Clinical Data once such CAROUSEL Patient Clinical Data has been released by the sponsor and
UCLB has authority to disclose the same to Autolus. The frequency and timing of such transfers of the CAROUSEL Patient Clinical
Data is detailed in Schedule 11. Upon the disclosure of such CAROUSEL Patient Clinical Data to Autolus, the CAT19 CNS Licence
shall automatically be extended to include a licence to the CAROUSEL Patient Clinical Data released and the CAROUSEL Program
IP shall be deemed to include such CAROUSEL Patient Clinical Data.

j.

UCLB shall procure that UCL (including MP and those engaged in the MP Laboratory) shall:

keep the Program IP confidential (subject to any disclosure in accordance with patent prosecution of the Licensed Patents);
not disclose the Program IP to any Third Party, other than as expressly permitted in the course of Academic Research pursuant to

Clause 4; and,

not  enable  or  assist  any  Third  Party  to  Exploit  any  of  the  Program  IP  other  than  (i)  as  expressly  permitted  in  the  course  of
Academic Research pursuant to Clause 4, or (ii) in connection with (1) the RQR8 Program IP to the extent permitted outside
of the RQR8 Field; (2) the Epitope Tag Program IP to the extent permitted outside of the Epitope Tag Field; (3) the Retrostim
Program IP to the extent permitted outside of the Retrostim Field; and/or (4) the TetCAR Program IP to the extent permitted
outside of the TetCAR Field.

OPTION TO ACQUIRE PROGRAM IP

UCLB acknowledges that Autolus has exercised its right to acquire ownership of all of the Licensed Patents, but excluding all Retained

Patents, as set forth in Schedule 14 (collectively, the “Assigned Patents”), with effect from the Third Amendment Date.

i..
ii..

iii..

10.
a.

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

b.

Autolus  acknowledges  that  the  assignment  of  the  Assigned  Patents  with  effect  from  the  Third  Amendment  Date  shall  not  extinguish
Autolus’s (or its successors) obligation to pay Royalties for sales of Royalty Product or other financial commitments under Clauses
13, 14 and 15.

c.

Autolus shall not be entitled to assign the Assigned Patents to any third party other than in accordance with the following provisions.

Autolus shall be entitled to assign the Assigned Patents in whole or part to:

i..
ii..

an acquirer of all or substantially all of Autolus’s business provided Autolus complies with Clause 29.2; or
a third party which is not acquiring all or substantially all of Autolus’s business provided that:

i.

ii.

iii.

the provisions of Clause 3.1 and 3.2 shall apply with respect to any proposed assignment as if it were a sub-licence and
Autolus shall procure that such third party and its successors in title to the Assigned Patents agree to, and comply with,
legally binding provisions in substantially the same form as Clause 3 in respect of any licences granted by such Third
Party under the Assigned Patents;
the assignment to such third party and its successors in title to the Assigned Patents shall be made subject to the non-
exclusive licence under the Assigned Patents granted by Autolus to UCLB purusant to Clause 4.1; and
prior to such assignment becoming effective, Autolus and UCLB shall agree in good faith, and enter into, a reasonable
and proportionate legally binding agreement which safeguards UCLB’s interests as regards the third party assignee and
any  successors  to  the  Assigned  Patents  in  the  following:  (a)  the  Royalties  and  Milestones  and  other  financial
commitments under Clauses 13, 14 and 15 in respect of Royalty Products developed and sold by an Assignee Entity;
(b) the maintenance and defence of such Assigned Patents; and (c) the assignment of such Assigned Patents to UCLB
on termination of a Program Licence or this Agreement as a whole pursuant to Clauses 25.1.2, 25.2.2, and/or 25.4.2.

DILIGENCE OBLIGATIONS

Diligence Obligations for Original Royalty Products

With respect to diligence obligations on Autolus concerning the GD2 Product, the Parties agree as follows:

iv.

v.

vi.

The “GD2 Diligence Obligation” shall mean (i) either (a) Autolus has initiated a follow-up clinical study with the same
CAR construct as used in the CRUK Study; or if necessary (b) Autolus has initiated formal pre-clinical development of a
new  GD2  Product  not  used  in  the  CRUK  Study;  and  (ii)  as  soon  as  is  reasonably  practicable  following  marketing
approval of the GD2 Product in each of the relevant territories, Autolus (or its Sub-licensee) has commenced sale of the
GD2 Product in either (a) the [***] (if reimbursement has been granted in the [***]) and one other Core Country or one of
[***] or (b) at least any two of [***] or any of the Core Countries (other than the [***]), in the event reimbursement is not
granted in the [***];
If Autolus fails to achieve limb (i)(a) of the GD2 Diligence Obligation within [***] of completion (being delivery of the final
written  report  to  Autolus  concerning  the  study)  of  the  Cancer  Research  UK  sponsored  GD2-CAR  clinical  trial
(CRUKD/15/001)  (“CRUK  Study”),  UCLB  may  serve  written  notice  on  Autolus  to  terminate  the  GD2  Licence  which
termination shall be effective within [***] of Autolus’s receipt of the same unless Autolus has previously served or within
the [***] period serves notice on UCLB that it has or will initiate pre-clinical development as an alternative under limb (i)
(b) of the GD2 Diligence Obligation (a “Limb (i)(b) Notice”);
If Autolus, having served a Limb (i)(b) Notice, fails to achieve limb (i)(b) of the GD2 Diligence Obligation within [***] of
completion of the CRUK Study, UCLB may serve written notice on Autolus to terminate the GD2 Licence in which case
the provisions of Clauses 11.1.4 and 11.1.5 shall govern the right to terminate the GD2 Licence;

11.
a.

i..

    35

CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

vii.

Autolus  shall  be  obliged  to  use  its  commercially  reasonable  efforts  to  fulfil  limb  (ii)  of  the  GD2  Diligence  Obligation
following  marketing  approval  of  the  GD2  Product  and  the  provisions  of  Clauses  11.1.4  and  11.1.5  shall  govern  any
allegation of breach of such obligation or termination of the GD2 Licence under this Clause 11.

Subject to Clause 11.1.3, and without prejudice to Clause 11.1.1, Autolus shall use its commercially reasonable efforts to develop
Original  Royalty  Products  under  any  of  the  Program  Licences  in  respect  of  the  Original  Program  IP  (other  than  a  GD2
Product)  provided  that  it  is  acknowledged  that  Autolus  shall  not  be  obliged  to  develop  an  Original  Royalty  Product  under
each and every Program Licence in respect of the Original Program IP at all times during the relevant period. It is recognised
that in complying with the above, Autolus shall have the right to determine in its sole discretion the prioritisation, on a purely
commercial basis, of the various Royalty Products for development, and that Autolus’s compliance with the foregoing shall be
assessed on the basis of the whole of its Royalty Product (Original Royalty Product, Additional Royalty Product and CAT19
Product) and other product development portfolio.

Once all of the Tranche 3 Subscription Shares (as such term is defined in the SSA) have been issued by Autolus, the diligence
obligations  set  out  under  Clause  11.1.2  above  shall  cease  to  apply  other  than  for  Autolus  seeking  to  develop  one  BCMA
Product, a [***] Product (if the licence has been exercised) and a TRBC1/2 Product.

Non-compliance  with  Clause  11.1.1  and/or  11.1.2  shall  not  result  in  a  right  to  terminate  this  Agreement  or  any  financial  or
equitable  remedy  (including  any  remedy  in  damages),  but  UCLB’s  sole  remedy  for  non-compliance  shall  be  limited  to  the
right to terminate those specific Original Program Licences granted under this Agreement for which Autolus is in breach in
accordance with Clause 11.1.5. It is acknowledged that notwithstanding any delay in development of one or more Original
Royalty Products, a breach of Clause 11.1.1 and/or 11.1.2 may be remedied by Autolus subsequently undertaking activities
to develop the applicable Original Royalty Product following UCLB’s written notice referred to below and, as such, a delay in
development timeline shall not be an un-remediable breach. Prior to exercising any right of termination UCLB shall first be
obliged  to  provide  Autolus  with  a  written  notice  setting  out  the  basis  for  its  allegation  of  breach  by  Autolus  under  Clause
11.1.1 and/or 11.1.2, which notice shall set out the deficiencies by Autolus and set out a series of reasonable activities UCLB
consider  sufficient  to  remedy  the  breach.  For  the  avoidance  of  doubt,  UCLB's  list  of  suggested  activities  shall  not  be  a
definitive list of what is required to remedy any breach. Upon Autolus’s receipt of such notice, the Parties shall, promptly, in
good  faith  and  acting  reasonably,  (i)  discuss  ways  for  Autolus  to  remedy  or  undertake  activities  in  compliance  with  the
obligations under Clause 11.1.1 and/or 11.1.2 and (ii) agree a reasonable period of time within which Autolus will be required
to undertake such activities. If the Parties fail to agree the period which Autolus has to undertake the activities, Autolus shall
have  [***]  from  the  date  Autolus  or  UCLB  serves  written  notice  stating  in  its  view  that  an  agreement  under  (ii)  cannot  be
reached  to  comply  with  its  obligations  under  Clause  11.1.1  and/or  11.1.2  for  the  Original  Royalty  Product(s)  in  respect  of
which the breach has occurred.

Provided that Clause 11.1.4 has been complied with and the process set out therein followed, and provided that following the [***]
period  (or  such  other  period  agreed  between  the  Parties)  Autolus  is  still  in  breach  of  the  same  obligations  under  Clause
11.1.1  and/or  11.1.2  in  respect  of  the  development  of  one  or  more  Original  Royalty  Products  that  were  the  subject  of  the
original breach notice under Clause 11.1.4, UCLB shall be entitled upon immediate written notice to terminate the Program
Licence(s) granted as follows:

viii.

UCLB shall be entitled to terminate the GD2 Licence where, in breach of its obligations hereunder, Autolus has not met
and failed to remedy the GD2 Diligence Obligation; and

ii..

iii..

iv..

v..

    36

CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

b.

i..

ix.

UCLB  shall  be  entitled  to  terminate,  on  an  Original  Program  Licence  by  Original  Program  Licence  basis,  the  Original
Program Licence applicable to the Original Royalty Product where, in breach of its obligations hereunder in respect of
such Original Royalty Product, Autolus has not met and has failed to remedy its diligence obligation under Clause 11.1.1
and/or 11.1.2 to develop such Royalty Products.

Diligence Obligations for Additional Royalty Products

Autolus  shall  use  its  commercially  reasonable  efforts  to  develop  at  least  one  Additional  Royalty  Product  under  each  of  the
Additional Program Licences (with the exception of the iCAR Program Licence in respect of which no diligence obligations
shall  apply  under  this  Agreement),  it  being  acknowledged  that  (i)  an  individual  Additional  Royalty  Product  may  utilise
technology  under  more  than  one  of  the  Additional  Program  Licences  and  hence  fulfil  the  foregoing  diligence  obligation  for
more  than  one  Additional  Program  Licence  and  (ii)  should  all  the  Patent  Rights  either  licensed  hereunder  or  assigned  to
Autolus under the Patent Assignment that are applicable to an Additional Royalty Product either be Surrendered or cease to
have  any  Valid  Claim  Autolus  shall  be  deemed  to  have  complied  with  its  foregoing  obligation.  The  foregoing  obligation  is
subject to the following:

x.

xi.

xii.

UCLB shall be entitled to assess Autolus’s compliance with the foregoing obligation [***], by assessing the development
work undertaken by or on behalf of Autolus under each of the Additional Program Licences over [***] (each period being
an “Assessment Period”). For this purpose, [***] before the end of any Assessment Period in respect of each Additional
Program Licence, Autolus shall provide UCLB with a written report that is detailed enough for UCLB to assess if Autolus’
activities are in accordance with this Clause 11.2;
Autolus’s diligence obligation under this Clause 11.2.1 shall, subject to the remaining provisions of this Clause 11.2.1,
expire upon the [***] anniversary of: [***];
Autolus will have met its diligence obligations under this Clause 11.2.1 for any Additional Program Licence where, in an
applicable Assessment Period:
(a) Autolus  has  spent  an  amount  equivalent  to  GBP  £[***]  on  development  activities  under  the  Additional  Program
Licence in question. Where development of a Royalty Product utilises technologies licensed under more than one
Additional  Program  Licence,  in  order  to  assess  if  an  amount  equivalent  to  GBP  £[***]  has  been  spent  on
development activities under the Additional Program Licence in question, the total amount of investment made by
Autolus in the applicable Assessment Period shall be apportioned equally (on a numerical basis and not a value
basis)  across  the  applicable  Additional  Program  Licences,  or  where  Autolus  reasonably  believes  that  it  is  more
appropriate for the apportionment to be across the applicable Additional Program Licences in which investment has
been  made  on  a  value  basis  (rather  than  on  an  equal  numerical  basis)  then  at  the  time  of  reporting  pursuant  to
Clause (i), Autolus shall disclose to UCLB the details of the basis according to which Autolus has determined the
apportionment  of  investment  to  be  allocated  for  each  of  the  Additional  Program  Licences  by  reference  to  value.
UCLB shall in good faith consider Autolus’ proposal. If UCLB agrees with Autolus, or does not object to Autolus’
proposal in writing within [***] of Autolus’ notification, the apportionment of the investment for the Royalty Product
across  the  Additional  Program  Licences  shall  be  carried  out  in  accordance  with  the  proposal  put  forward  by
Autolus.  If  UCLB  does  not  agree  with  Autolus’  proposal  and  notifies  Autolus  in  writing  within  [***]  of  Autolus’
notification setting out the reasons for its disagreement, the investment made by Autolus with respect to the Royalty
Product in the applicable Assessment Period shall be apportioned equally across the applicable Additional Program
Licences; or

    37

CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

ii..

iii..

(b) Autolus  has  conducted  research  and  development  activities  over  the  applicable  Assessment  Period  which  in
UCLB’s reasonable opinion are sufficient to demonstrate that Autolus has used commercially reasonable efforts to
develop one or more Additional Royalty Products under the Additional Program Licence in question;
If Autolus (or its Sub-licensee) has filed any IND, CTA or comparable application for undertaking any clinical trial in respect of an
Additional Royalty Product, then notwithstanding the provisions of Clause 11.2.1(iii), Autolus shall be deemed to have fulfilled
its diligence obligations under this Clause 11.2 with respect to such Additional Royalty Product and the Additional Program
Licence(s) applicable to such Additional Royalty Product;

If  Autolus  does  not  meet  (or  is  not  deemed  to  have  met)  its  diligence  obligations  for  any  Additional  Program  Licence  in  any
Assessment  Period,  (i)  UCLB  shall  be  entitled  to  terminate  the  specific  Additional  Program  Licence  in  question  by  serving
written  notice  of  termination  on  Autolus  within  [***]  of  expiry  of  the  applicable  Assessment  Period  in  which  case  the
termination of the Additional Program Licence shall have immediate effect, but failing which the Additional Program Licence
may not be terminated for non-compliance during such Assessment Period; and (ii) UCLB shall not have a right to terminate
this Agreement as a whole or any financial or equitable remedy (including any remedy in damages), but UCLB’s sole remedy
for non-compliance shall be limited to the right to terminate the Additional Program Licence in question.

c.

Diligence Obligations for the CAT19 Product

With respect to diligence obligations on Autolus concerning the CAT19 Product, the Parties agree as follows:

The “CAT19 Diligence Obligation” shall mean (i) either (a) Autolus has initiated a follow-up clinical study with a CAT19 1  Gen
Product,  or  (b)  Autolus  has  initiated  formal  pre-clinical  development  of  a  CAT19  Binder  Product;  and  (ii)  as  soon  as  is
reasonably practicable following marketing approval of the CAT19 Product in each of the relevant territories, Autolus (or its
Sub-licensee) has commenced sale of the CAT19 Product in either (a) the [***] (if reimbursement has been granted in the
[***]) and one other Core Country or one of [***] or (b) at least any two of [***] or any of the Core Countries (other than the
[***]), in the event reimbursement is not granted in the [***].

st

If Autolus fails to achieve the CAT19 Diligence Obligation under limb (i) within [***] of completion of the ALLCAR19 Study (being
delivery of the final written report to Autolus concerning the relevant study) or fails to use its commercially reasonable efforts
to fulfil limb (ii) of the CAT19 Diligence Obligation following Marketing Approval of the CAT19 Product, then UCLB may serve
written  notice  on  Autolus  to  terminate  the  CAT19  Licence  and  the  CAT19  CNS  Licence,  in  which  case  the  provisions  of
Clauses 11.1.4 and 11.1.5 shall govern the right to terminate the CAT19 Licence and the CAT19 CNS Licence (in which case,
references  in  Clauses  11.1.4  and  11.1.5  to  Original  Program  Licences  and  Original  Royalty  Product  shall  be  read  as  the
CAT19 Licence and CAT19 Product and CAT19 CNS Licence and CAT19 CNS Product and references within Clauses 11.1.4
and 11.1.5 to Clause 11.1.1 and Clause 11.1.2 shall be read as references to Clause 11.3.1).

The  Parties  recognise  the  importance  of  making  pharmaceutical  products  available  in  Developing  Countries,  to  the  extent
practicable. However,  the  Parties  acknowledge  the  early  stage  nature  of  the  CAT19  Program  IP  and  acknowledge  that  a
substantial investment would be required to bring CAT19 Products to market in Developing Countries, especially given the
clinical  infrastructure  required  to  support  the  administration  of  a  CAR  T-cell  therapy,  the  unfamiliarity  of  such  therapies  to
local  regulatory  authorities  and  the  uncertainty  associated  with  pricing  and  reimbursement  strategies  in  Developing
Countries.

Diligence Provisions relating to all Programs

It is acknowledged that Exploitation by or on behalf of Autolus, Autolus’s Affiliates and/or Sub-licensees or Assignee Entities of

any Royalty Product shall, for the purposes of this Clause 11, be considered activities of Autolus for assessing its

i..

ii..

iii..

d.

i..

    38

CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

use  of  commercially  reasonable  efforts  and  compliance  with  Clause  11.  Without  prejudice  to  the  provisions  of  Clause
11.2.1(i), with effect from UCLB ceasing to have a director or observer status on the board of Autolus, thereafter by [***] of
each  year,  Autolus  shall  provide  UCLB  with  a  written  report  that  will  include  a  summary  of  its  development  timelines  and
major  development  steps  in  relation  to  the  Royalty  Products  that  were  taken  in  the  previous  twelve  (12)  months  and  will
include  development  timelines,  budget  and  major  development  steps  that  Autolus  anticipates  shall  be  undertaken  with
respect to the Royalty Products for the following twelve (12) months. In addition to the foregoing (but without prejudice to the
provisions of Clause 11.2.1(i)), UCLB shall be entitled, if reasonable, to request details of FTE resource allocation and CRO
costs incurred by Autolus. The foregoing obligation shall cease to apply with effect from the [***] anniversary of the Effective
Date.

Upon  termination  of  any  Program  Licence  pursuant  to  this  Clause  11,  UCLB  shall  have  the  option  to  negotiate  with  Autolus  to
agree terms for the assignment of or grant of an exclusive licence to those Autolus Improvements Controlled by Autolus free
of  restrictions  and  encumbrances  and  which  specifically  relate  to  the  Program  Licence  that  has  been  terminated.  The
foregoing right of UCLB to negotiate an assignment or licence with Autolus for the Autolus Improvements shall expire [***]
following notice of termination served under Clause 11.1.5, Clause 11.2.3 or Clause 11.3.2. This Clause 11.4.2 shall survive
the termination of this Agreement for [***] from the date of termination during which UCLB shall have a right to exercise its
right under Clause 11.4.2.

Autolus  shall,  by  written  notice,  promptly  notify  UCLB  in  the  event  that  its  Board  takes  any  decision  to  permanently  terminate
development  of  any  Royalty  Products  under  a  particular  Program  Licence,  whereupon  Autolus  shall  have  no  further
obligation to develop or Exploit any Royalty Product applicable to that Program Licence and the relevant Program Licence
shall terminate as of the date of Autolus’s written notice and the relevant provisions of Clause 25 shall apply.

UCLB MANAGEMENT FEE AND SHARES

Autolus shall, during the Term of this Agreement, make a maximum of [***] payments to UCLB each of GBP £[***], with each annual
payment being made within [***] of receipt of a VAT invoice addressed to Autolus, the first of which shall be issued no earlier than the
first  anniversary  of  the  Effective  Date,  and  thereafter  for  the  remaining  three  (3)  annual  payments  they  shall  be  issued  on  each
subsequent anniversary of the Effective Date.

In  consideration  of  UCLB  entering  into  the  Agreement  with  effective  date  25  September  2014,  Autolus  has  issued  and  allocated  to

UCLB 4,769,994 B Ordinary Shares (as such terms is defined in the SSA) in Autolus.

In consideration of UCLB granting the Additional Program Licences (except the TRBC2 Dx Licence) to Autolus pursuant to the deed of
variation  of  the  Agreement  with  effective  date  2  March  2016,  Autolus  has  issued  and  allocated  to  UCLB  1,000,000  additional  B
Ordinary  Shares  (as  such  term  is  defined  in  the  SSA)  each  credited  as  fully  paid  up  to  £1.00  per  B  Ordinary  Share.  In  addition,
Autolus  has  paid  UCLB  the  sum  of  £150,000  (one  hundred  and  fifty  thousand  pounds  sterling)  within  14  days  of  the  Amendment
Date.

ii..

iii..

12.
a.

b.

c.

d.

In consideration of UCLB entering into this Amended and Restated Licence Agreement effective as of the Second Amendment Date,

Autolus shall pay UCLB within [***] of the Second Amendment Date:

the sum of £1,000,000 (one million pounds sterling); and
the sum of £500,000 (five hundred thousand pounds sterling) in recognition of the Second Amendment Date being a date no later

than [***].

In  consideration  of  the  transfer  of  a  copy  of  all  the  CARPALL  and  ALLCAR19  Patient  Clinical  Data  generated  as  at  the  Third

Amendment Date, Autolus shall pay UCLB the sum of £[***] within [***] of the Third Amendment Date.

In  consideration  of  UCLB  entering  into  this  Amended  and  Restated  Licence  Agreement  effective  as  of  the  Third  Amendment  Date,

Autolus shall pay UCLB within [***] of the Third Amendment Date the sum of £[***] in recognition of the TRBC2 Dx Licence.
MILESTONE PAYMENTS

One-Off Success Milestone Payments

i..
ii..

e.

f.

13.
a.

    39

CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

i..

During the Term of this Agreement, upon the occurrence of any milestone applicable to the relevant Royalty Product set out in the
table below (each a “Success Milestone”) Autolus shall, in accordance with Clause 16, pay a sum equal to the amount set
against that Success Milestone in the table below (each amount being a “Success Milestone Payment”).

Royalty Product

Success Milestone

Success Milestone
Payment
([***])

GD2 Product

BCMA Product

TRBC1/2 Product

TRBC1/2 Product

[***] Product

NSG Product

Logic Gate Product

Logic Gate Product

ZipCAR Product

ZipCAR Product

ccCAR Product

Epitope Tag Product

iCAR Product

RapaiCASP9 Product

Retrostim Product

TetCAR Product

ZAP-CAR Product

CAT19 Product

CAT19 Product

CAT19 Product

CAT19 Product

CAT19 Product

TRBC2 Dx Product

TRBC2 Dx Product

TRBC2 Dx Product

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

xiv.
xv.

The payment of Success Milestone Payments under Clause 13.1.1 above is subject to the following:
xiii.

each  Success  Milestone  Payment  set  out  above  shall  be  payable  once  only,  irrespective  of  the  number  of  Royalty
Products achieving the applicable Success Milestone;
the aggregate maximum payment under Clause 13.1.1 and 13.1.2 shall never exceed GBP £[***] ([***] pounds sterling);
if a particular Royalty Product triggers two or more Success Milestones (on the basis that by definition it may fall within
more than one definition of a Royalty Product), then:
(c)

if  the  occurrence  of  the  Success  Milestones  are  simultaneous  with  each  other,  then  only  the  highest  value
applicable Success Milestone Payment triggered at that time shall be payable in respect of that Royalty Product,
and the other Success Milestone Payments triggered at the same time shall not be payable; or
if  the  occurrence  of  the  Success  Milestones  are  sequential,  then  with  the  exception  of  the  Success  Milestones
referred to in Clause (C) the Success Milestone Payment attributable to the first Success Milestone applicable to
that  Royalty  Product  shall  be  paid  in  full,  and  the  Success  Milestone  Payment  attributable  to  the  next  sequential
Success

(d)

ii..

    40

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

Milestone  shall  be  payable  at  [***]  of  the  Success  Milestone  value  set  out  in  the  table  above,  and  no  further
Success Milestones shall be payable in respect of that Royalty Product;

(f)

(e) with respect to the TRBC1/2 Product, there shall be no reduction in the Success Milestone Payment identified as
number  (4)(ii),  in  the  above  table  by  virtue  of  such  Success  Milestone  being  a  sequential  Success  Milestone  in
respect of the same Royalty Product (TRBC1/2 Product), unless such Success Milestone is a [***], where upon it
shall be payable at [***] of the Success Milestone value set out at (4(ii)) in the table above, and no further Success
Milestones shall be payable in respect of that TRBC1/2 Product;
in  calculating  Net  Sales  for  the  applicable  Royalty  Product  (i)  the  currency  exchange  mechanism  set  out  in  this
Agreement  to  calculate  the  relevant  Net  Sales  shall  be  applied  and  (ii)  sales  of  any  Complementary  Diagnostic
Product shall be excluded from Net Sales for the purposes of calculating whether a Success Milestone has been
triggered;
in the case of Success Milestones 7 and 8 above concerning Logic Gate Products, the reference to a “first Logic
Gate Product” and a “second Logic Gate Product” shall mean that the first and second Logic Gate Products shall
each [***]; and,
in the case of Success Milestones 9 and 10 above concerning ZipCAR Products, the reference to a “first ZipCAR
Product” and a “second ZipCAR Product” shall mean that the first and second ZipCAR Products shall each [***].

(g)

(h)

Additional Milestone Payments

During the Term, if Success Milestones 7 and 8 have both been achieved (irrespective of whether UCLB has received payments
with respect to either or both of Success Milestones 7 and 8), should any subsequent Logic Gate Product achieve [***] (being
any Logic Gate Product that [***]), Autolus shall, in accordance with Clause 16, pay to UCLB a one-off milestone payment of
GBP £[***] upon the [***] of such subsequent Logic Gate Product (“Additional LG Milestone Payment”) provided that at the
time  of  [***].  A  “Milestone  Logic  Gate  Product”  is  any  Logic  Gate  Product  in  respect  of  which  a  Success  Milestone
Payment  or  Additional  LG  Milestone  Payment  has  been  paid  or  is  payable.  For  the  avoidance  of  doubt,  subject  to  the
requirement of [***], there shall be no limit on the number of Additional LG Milestone Payments that may be payable under
this Clause 13.2.1. Upon there ceasing to be [***], this Clause 13.2.1 shall cease to apply.

During the Term, if Success Milestones 9 and 10 have both been achieved (irrespective of whether UCLB has received payments
with respect to either or both of Success Milestones 9 and 10), should any subsequent ZipCAR Product achieve [***] (being a
ZipCAR Product that [***]), Autolus shall, in accordance with Clause 16, pay to UCLB a one-off milestone payment of GBP £
[***] upon [***] of such subsequent ZipCAR Product (“Additional ZC Milestone Payment”) provided that at the time of [***].
A “Milestone ZipCAR Product” is any ZipCAR Product in respect of which a Success Milestone Payment or Additional ZC
Milestone Payment has been paid or is payable. For the avoidance of doubt, subject to the requirement of [***], there shall be
no  limit  on  the  number  of  Additional  ZC  Milestone  Payments  that  may  be  payable  under  this  Clause  13.2.2.  Upon  there
ceasing to be [***], this Clause 13.2.2 shall cease to apply.

By way of example only to assist interpretation of Clauses 13.1 to 13.2.2 (inclusive) assuming:

a Royalty Product is a [***] Product, a [***] Product and a [***] Product, the first set of Success Milestones to be achieved for this
Royalty Product would, on a simultaneous basis, be the Success Milestones for the [***] (number [***] in above table) and
[***]  Product  (number  [***]  in  the  above  table).  Upon  achievement  of  the  foregoing  simultaneous  Success  Milestones,  the
payment provisions of Clause [13.1.2(iii)(A) ]would apply and so UCLB would be entitled to receive a

b.

i..

ii..

c.

i..

    41

CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

payment  of  £[***]  (the  higher  of  the  simultaneous  Success  Milestones).  The  second  set  of  Success  Milestones  to  be
achieved for this Royalty Product would be the Success Milestone for the [***] Product (number [***] in the above table). As
the Success Milestone for the [***] Product ([***]) in respect of the Royalty Product would be achieved after achievement of
the  Success  Milestones  [***],  the  Success  Milestone  for  the  [***]  would  be  a  sequential  Success  Milestone  that  would  be
subject to the provisions of Clause [13.1.2(iii)(B)]. Accordingly, if the Success Milestone for the [***] is achieved in relation to
the aforementioned Royalty Product, UCLB would be entitled to receive a payment of £[***] ([***] of the Success Milestone
([***]));

a Royalty Product is a [***] Product, a [***] Product and a [***] Product, the only Success Milestone for which a payment is due to
UCLB  would  be  the  Success  Milestone  for  the  [***]  Product  (number  [***]  in  the  above  table).  Accordingly,  if  the  Success
Milestone  for  the  [***]  Product  is  achieved  in  relation  to  the  aforementioned  Royalty  Product,  UCLB  would  be  entitled  to
receive a payment of £[***].

Sales Milestones

During  the  Term  Autolus  shall  pay  to  UCLB  each  of  the  following  one-off  sales-related  milestone  payments  (each  a  “Sales
Milestone Payment”) payable in the Year that aggregate annual global Net Sales of all Royalty Products first exceed the
following thresholds (each a “Sales Milestone”) calculated from the Effective Date:

ii..

d.

i..

Aggregate annual global Net Sales of all Royalty Products

GBP £[***]

GBP £[***]

Sales Milestone Payment
([***])

[***]

[***]

In calculating aggregate annual global Net Sales, (i) the currency exchange mechanism set out in this Agreement to calculate the
relevant  Net  Sales  shall  be  applied;  and  (ii)  sales  of  any  Complementary  Diagnostic  Product  shall  not  be  included  in  Net
Sales for the purposes of calculating whether a Sales Milestone has been triggered. Each of the Sales Milestone Payments
in this Clause 13.4.2 shall be paid once only irrespective of the number of Royalty Products and shall be paid in accordance
with Clause 16.

ROYALTIES

On a Program Licence by Program Licence basis, in partial consideration of the grant of that particular Program Licence, during the
Royalty  Term  Autolus  shall  pay  to  UCLB  a  royalty  on  Net  Sales  of  the  applicable  Royalty  Product  supplied  by  Autolus  or  its  Sub-
Licensees or any Assignee Entity within the applicable field under that Program Licence within the Territory, such royalty calculated
as  the  percentage  value  of  the  Net  Sales  at  the  following  rates  subject  to  the  terms  and  conditions  of  this  Agreement,  and  in
particular the remaining provisions of this Clause 14 (individually per Royalty Product a “Royalty” and collectively the “Royalties”):

ii..

14.
a.

    42

CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

Royalty Product and Field

Royalty Rate

Net Sales of BCMA Products within the BCMA Field

Net Sales of [***] Product within the Field (with effect from grant of the [***] Licence)

Net Sales of GD2 Products within the Field

Net Sales of Logic Gate Products within the Field

Net Sales of NSG Products within the Field

Net Sales of RQR8 Product within the Field

Net Sales of TRBC1/2 Products within the Field

Net Sales of ZipCAR Products within the Field

Net Sales of ccCAR Products within the Field

Net Sales of Epitope Tag Products within the Field

Net Sales of iCAR Products within the Field

Net Sales of RapaiCASP9 Products within the Field

Net Sales of Retrostim Products within the Field

Net Sales of TetCAR Products within the Field

Net Sales of ZAP-CAR Products within the Field

Net Sales of CAT19 1  Gen Product within the Field

st

Net Sales of CAT19 CNS Product within the Field

Net Sales of CAT19 Binder Product within the Field

Net Sales of TRBC2 Dx Products within the Field

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

b.

c.

d.

Subject  to  Clause  14.4,  only  one  Royalty  Rate  shall  be  payable  per  Royalty  Product  and  the  Royalty  payable  on  a  Royalty  Product

shall be calculated only once and payable only once.

The Royalty Rate in respect of a Royalty Product set out above shall be adjusted, as applicable, in accordance with the provisions of
Clause 14.4 to 14.10, and the order of reduction or adjustment in the Royalty Rate or Royalty due shall be applied sequentially in the
order of those remaining clauses.
Multiple Royalty Product Adjustments

No Royalty Product shall trigger more than one Royalty payment, such that if a product or therapy falls within two or more categories
of  Royalty  Product  (such  as  a  GD2  Product  being  Covered  by  RQR8  Program  IP)  then  the  maximum  Royalty  payable  for  that
particular product or therapy shall be calculated as a percentage of the Net Sales for such Royalty Product at a rate being the sum of
[***] payable pursuant to Clause 14.1 for such Royalty Product, plus [***] of the next [***] payable pursuant to Clause 14.1 for such
Royalty Product.

e.

Adjustment to Royalty Rate for Sub-Licensees

Excluding  the  Royalty  Rate  for  the  GD2  Product,  where  Autolus  has  granted  rights  under  the  Technology  to  any  Sub-Licensee  in
respect  of  any  Royalty  Product  and  at  the  effective  date  of  such  sub-licence,  the  Royalty  Product  has  only  been  the  subject  of
preclinical  development  (being  any  development  activities  prior  to  commencement  of  a  phase  I  trial)  conducted  by  or  on  behalf  of
Autolus, then the Royalty due to UCLB on Net Sales of that Royalty Product made by the Sub-Licensee, irrespective of the Royalty
Rate(s) applicable to such Net Sales set out above, shall not exceed [***] of the sums received by Autolus from such Sub-Licensee in
respect of such Net Sales.

f.

Royalty Rate Reductions

In respect of each Royalty Product and on a country by country basis, the Royalty Rate applicable to the Net Sales for such Royalty
Product shall be reduced by the percentages set out in the table below where the applicable circumstance exists or does not exist, as
the  context  requires.  Furthermore,  where  a  product  or  therapy  falls  within  two  or  more  definitions  of  a  Royalty  Product,  then  the
following circumstances shall be assessed on an individual Royalty Product by Royalty Product basis (and hence separate Program
IP by Program IP basis) such that the Royalty Rate in respect of the product or therapy falling within one Royalty Product definition
may be adjusted differently to the Royalty Rate that would be applicable for such same product or therapy also falling within a second
definition for another Royalty Product.

    43

CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

Circumstance in the country of sale in respect of the applicable Royalty Product

(A) the sale of the Royalty Product in the country of sale would were it not for the specific Licence granted
hereunder in respect of Licensed Patents under one category of Program IP specific to that Royalty
Product or the ownership by Autolus or the relevant Assignee Entity of the relevant Licensed Patents, at the
time of sale infringe a Valid Claim of such Licensed Patents included in that Program IP.

(B) where the Royalty Product is either an Epitope Tag Product and/or a Retrostim Product (i) such Royalty
Product would were it not for the specific Licence granted hereunder in respect of the Epitope Tag Patent
Rights and/or Retrostim Patent Rights, as applicable to that Royalty Product, or the ownership by Autolus
or the relevant Assignee Entity of the Epitope Tag Patent Rights and/or Retrostim Patent Rights, at the time
of sale infringe a Valid Claim of the Epitope Tag Patent Rights and/or Retrostim Patent Rights in that
country of sale; (ii) the manufacture of such Royalty Product or the manufacture of any component used in
its manufacture would, were it not for the specific Licence granted hereunder in respect of the Epitope Tag
Patent Rights or the ownership by Autolus or the relevant Assignee Entity of the Epitope Tag Patent Rights,
have infringed a Valid Claim of the Epitope Patent Tag Rights at the time of sale in the country of sale for
such Royalty Product (as if such Royalty Product or any component used in the manufacture of such
Royalty Product had been manufactured in the country of sale) and/or (iii) the manufacture of such Royalty
Product or the manufacture of any component used in its manufacture would, were it not for the specific
Licence granted hereunder in respect of the Retrostim Patent Rights, or the ownership by Autolus or the
relevant Assignee Entity of the Retrostim Patent Rights have infringed a Valid Claim of the Retrostim Patent
Rights at the time of its sale in the country of sale for such Royalty Product (as if such Royalty Product or
any component used in the manufacture of such Royalty Product had been manufactured in the country of
sale).

(C) Circumstance (A) above does not exist, but the Original Royalty Product or the CAT19 1  Gen Product
or the CAT19 CNS Product benefits from Regulatory Exclusivity in that country of sale.

st

(D) Neither of the circumstances (A) or (C) exists, but the sale of the Original Royalty Product or CAT19 1
Gen Product or CAT19 CNS Product is made in the country during the Know-How Period applicable to that
Original Royalty Product or the CAT19 1  Gen Product or CAT19 CNS Product.

st

st

(E) Neither of the circumstances (A) or (C) exists, but the sale of the CAT19 Binder Product is made in the
country during the Know-How Period applicable to the CAT19 Binder Product, provided that the CAT19
Binder Product is not Covered by any Program IP in addition to the CAT19 Program IP at the time of sale.

(F) (i) in the case of an Original Royalty Product or CAT19 Product, [***] for that applicable Original Royalty
Product or CAT19 Product; or (ii) in the case of an Additional Royalty Product (that is not an Epitope Tag
Product or a Retrostim Product), [***] for that applicable Additional Royalty Product; or (iii) in the case of an
Epitope  Tag  Product  or  a  Retrostim  Product,  neither  circumstance  (A)  or  (B)  apply  for  such  Epitope  Tag
Product or Retrostim Product (each being a “Royalty Expiry”).

Percentage reduction to the
Royalty Rate

[***]%

(with respect to the Royalty Rate
applicable only to the use of any
Licensed Patents excluding the
Epitope Tag Patents and
Retrostim Patent Rights)

[***]%

(with respect to the Royalty Rate
applicable only to the use of
Epitope Tag Patents or Retrostim
Patent Rights only)

[***]%

[***]%

[***]%

[***]%

g.

By  way  of  example  only  to  assist  interpretation  of  the  foregoing,  assuming  a  Royalty  Product  is  (i)  both  a  [***]  Product  and  a  [***]
Product, but that in the country of sale there are no Valid Claims remaining under the [***] Patent Rights and one Valid Claim remains
under the [***] Patent Rights which would, were it not for the Licence to the [***] Program

    44

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

IP  or  the  ownership  by  Autolus  or  the  relevant  Assignee  Entity  of  the  [***]  Patent  Rights,  be  infringed  by  the  sale  of  such  Royalty
Product, and there remains Regulatory Exclusivity then the applicable royalty would be [***] being [***] of the [***] Royalty Rate and
[***] of the [***] Royalty Rate; or (ii) both a [***] Product and an [***] Product, but that in the country of sale there are no Valid Claims
remaining under the [***] Patent Rights or [***] Patent Rights, but one Valid Claim remains under the [***] Patent Rights in the country
of  sale  of  the  [***]  Product,  which  Valid  Claim  would,  were  it  not  for  the  Licence  to  the  [***]  IP  or  the  ownership  by  Autolus  or  the
relevant Assignee Entity of the [***] Patent Rights, be infringed at the time of the sale of such [***] Product, by the manufacture of
such  [***]  Product  or  by  the  manufacture  of  materials  subsequently  used  in  the  manufacture  of  the  such  [***]  Product  (as  if  such
manufacture was taking place in the country of sale), and the Royalty Product is sold during the Know-How Period, then the total
applicable royalty would be[***], being [***] of the [***] Royalty Rate [***][***][***] and [***] of the [***] Royalty Rate [***][***][***].

For the purposes of Clause 14.6:

st

the  “Know-How  Period”  means  (a)  with  respect  to  Original  Royalty  Products  the  period  of  time  in  a  particular  country
commencing with the Effective Date and expiring on the earlier of (i) the [***] of the Effective Date or (ii) [***] from the date of
First Commercial Sale in the country in question for the applicable Original Royalty Product, and (b) with respect to a CAT19
1  Gen Product the period of time in a particular country commencing with the Second Amendment Date and expiring on the
earlier  of  (i)  the  [***]  of  the  Second  Amendment  Date  or  (ii)  [***]  from  the  date  of  First  Commercial  Sale  in  the  country  in
question for the applicable CAT19 1  Gen Product and (c) with respect to a CAT19 Binder Product the period of time in a
particular  country  commencing  with  the  Second  Amendment  Date  and  expiring  on  the  earlier  of  (i)  the  [***]  of  the  Second
Amendment  Date  or  (ii)  [***]  from  the  date  of  First  Commercial  Sale  in  the  country  in  question  for  the  applicable  CAT19
Binder Product, and (d) with respect to a CAT19 CNS Product the period of time in a particular country commencing with the
Third Amendment Date and expiring on the earlier of (i) the [***] of the Third Amendment Date or (ii) [***] from the date of
First Commercial Sale in the country in question for the applicable CAT19 CNS Product; and,

st

no Royalty shall be payable on an Additional Royalty Product unless condition (A) or condition (B) exists at the time of sale.

Royalty Stacking

If Autolus, its Affiliates or any Sub-Licensee or any Assignee Entity in-licenses or acquires (a) any rights for Exploitation of the BioVec
cell line referred to in Schedule 13 for the purpose of Exploiting the same GD2 Product that is the subject of the CRUK Study; or (b)
any Patent Rights from any Third Party or, subject to Clause 14.10 from UCLB, and such Patent Rights are required (as reasonably
assessed, based on such rights blocking Exploitation) to Exploit any Royalty Product(s) in any way (“Third Party Access Rights”);
to the extent Autolus, its Affiliates or its Sub-Licensee or Assignee Entity is required (under (a) and/or (b)) to pay any consideration,
royalties, monies, milestones, or other fees under or in connection with the aforementioned use of such cell line and/or Third Party
Access Rights applicable to any Royalty Product(s) (“TP Fees”), such TP Fees shall be deductible from Royalties otherwise due on
those Royalty Product(s) as follows:

the deduction from Royalties of the TP Fees payable in respect of the [***][***][***] shall be limited to [***], and only [***] of the

value of those TP Fees [***] may be deducted;

in the case of TP Fees paid for Third Party Access Rights, a maximum deduction of [***] of the total Royalty that would otherwise

be payable were it not for this Clause;

for  the  purpose  of  this  Clause,  Third  Party  Access  Rights  shall  include  the  in-licensing  or  acquisition  of  [***]  or  equivalent

technology in so far as it relates to [***] but shall not include the in-licensing of any other unpatented technology.

If Autolus, its Affiliates or any Sub-Licensee or any Assignee Entity in-licenses any Patent Right from UCLB that is (i) in the name of
UCLB as of the Effective Date; and/or (ii) is filed by or on behalf of or at the direction of UCLB within [***] after the Effective Date in

h.

i..

ii..
i.

i..

ii..

iii..

j.

    45

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

k.

i..

respect of any invention recorded in an invention disclosure form logged in UCLB’s database and categorised as “Biopharm” and
with the status “being assessed” as of [***] prior to the Effective Date, then where such Patent Right is required to Exploit any Royalty
Product(s)  in  any  way  (as  reasonably  assessed,  based  on  such  rights  blocking  Exploitation),  if  Autolus,  its  Affiliates  or  any  Sub-
Licensee or any Assignee Entity is required under the terms of such licence to pay any consideration, royalties, monies, milestones,
or other fees under or in connection with such rights, the Royalty in respect of such Royalty Product(s) shall be reduced by [***].

Diminished Royalty Product

If  a  Third  Party  (that  is  not  authorised  as  a  Sub-licensee  or  an  Assignee  Entity  to  Exploit  a  particular  Royalty  Product)
commences  Exploitation  of  any  Competitive  Product  in  a  country  within  the  Territory  that  infringes  any  of  the  Intellectual
Property  licensed  hereunder  or  any  Assigned  Patents  (each  an  “Competing  Entrant”),  and  UCLB  and/or  Autolus
commence litigation against such Competing Entrant in respect of such Competitive Product, then in so far as any Royalties
are  due  for  sales  of  Royalty  Product(s)  in  the  country  where  litigation  is  ongoing  and  in  respect  of  which  the  Competitive
Product  is  competitive,  such  Royalties  will  be  paid  into  escrow  by  Autolus  pending  resolution  of  such  litigation.  Upon
conclusion of such litigation, the Royalties due on those Royalty Products sold during the period in which the litigation was
on-going, shall be re-calculated (based on the final outcome of the patent position, status of Regulatory Exclusivity available
and Know-How Period as at the conclusion of the litigation) and the funds held in escrow shall be distributed according to
such re-calculation.

ii..

All  interest  earned  on  the  sums  paid  into  escrow  pursuant  to  this  clause  shall  accrue  to  the  benefit  of  the  escrow  account  for

distribution in accordance with Clause 14.11.

l.

Royalty Term

The Royalty Term shall commence on the Effective Date and on a country by country basis and Royalty Product by Royalty Product
basis, shall expire automatically upon there being a Royalty Expiry in such country for such Royalty Product. Upon such expiry the
rights  and  licences  granted  under  this  Agreement  to  Autolus  in  respect  of  such  Royalty  Product  and  country  (including  any  sub-
licences granted by Autolus in respect thereof) shall become irrevocable, perpetual, royalty free and fully paid up.
SUB-LICENSEE PAYMENTS

If Autolus, upon (i) granting a sub-license of any Technology or a licence of Assigned Patents to a Sub-Licensee for the right to Exploit
one  or  more  Royalty  Products  (each  being  a  “Sublicence”),  or  (ii)  assigning  any  Assigned  Patent  to  a  third  party  (each  being  a
“Patent Sale”) receives in consideration of that grant or sale any Net Receipts, Autolus shall, subject to Clause 15.2 and Clause 16,
make payments to UCLB from time to time calculated by reference to a percentage of Net Receipts received by Autolus under the
Sublicence or Patent Sale in accordance with the applicable percentage set out below (“Sublicence Or Patent Sale Payment”):

where (i) the Sublicence includes a sub-licence under any of the BCMA Licence, the [***] Licence, the GD2 Licence, the Logic
Gate Licence, the NSG Licence, the RQR8 Licence, the TRBC1/2 Licence and/or the ZipCAR Licence or a licence under any
Assigned  Patents  which  were  previously  licensed  under  any  such  Program  Licence  but  not  under  any  Additional  Program
Licences, or (ii) the Patent Sale includes assignment of any of the [***] Patent Rights, the GD2 Patent Rights, the Logic Gate
Patent Rights, the NSG Patent Rights, the TRBC1/2 Patent Rights and/or the ZipCAR Patent Rights: - 

15.
a.

i..

    46

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

Circumstances at the time of grant of the Sublicence or Patent Sale

Percentage of Net Receipts

At the time of grant of the Sublicence or completion of such Patent Sale, the aggregate
investment (by way of cash or debt) in Autolus from its incorporation date by its shareholders or
investors for the development of the Technology is less than GBP £[***]

At the time of grant of the Sublicence or completion of such Patent Sale, the aggregate
investment (by way of cash or debt) in Autolus from its incorporation date by its shareholders or
investors for the development of the Technology is equal to or more than GBP £[***] (unless the
circumstances below apply)

At the time of grant of the Sublicence or completion of such Patent Sale, at least [***] have
passed since the aggregate investment (by way of cash or debt) in Autolus from its incorporation
date by its shareholders or investors for the development of the Technology first equalled or
exceeded GBP £[***] (in which case none of the above provisions shall apply)

[***]%

[***]%

[***]%

ii..

where (i) the Sublicence is in respect of any of the ccCAR Licence, the Epitope Tag Licence, the iCAR Licence, the RapaiCASP9
Licence, the Retrostim Licence, the TetCAR Licence, and/or the ZAP-CAR Licence or a licence under any Assigned Patents
which  were  previously  licensed  under  any  such  Program  Licence  (and  does  not  include  a  sub-licence  under  any  of  the
Original Program Licences or a licence under any Assigned Patents which were previously licensed under any such Program
Licence), or (ii) the Patent Sale is in respect of any of the ccCAR Patent Rights, the Epitope Tag Patent Rights, the iCAR
Patent Rights, the RapaiCASP9 Patent Rights, the Retrostim Patent Rights and/or the ZAP-CAR Patent Rights (and does not
include  a  sub-licence  under  any  of  the  Original  Program  Licences  or  a  licence  under  any  Assigned  Patents  which  were
previously licensed under any such Program Licence):

Circumstances at the time of grant of the Sublicence or Patent Sale

Percentage of Net Receipts

The Sublicence is granted or Patent Sale is completed within [***] following the Amendment
Date

The Sublicence is granted or Patent Sale is completed between [***] and [***] following the
Amendment Date

The Sublicence is granted or Patent Sale is completed anytime after [***] following the
Amendment Date

[***]%

[***]%

[***]%

iii..

where  the  Sublicence  is  in  respect  of  the  CAT19  Licence  and/or  under  the  CAT19  CNS  Licence  (and  does  not  include  a  sub-
licence  under  any  of  the  Original  Program  Licences  or  the  Additional  Program  Licences  or  a  licence  under  any  Assigned
Patents which were previously licensed under any such Program Licence):

Circumstances at the time of grant of the Sublicence

Percentage of Net Receipts

The Sublicence is granted within [***] following the Second Amendment Date

The Sublicence is granted between [***] and [***] following the Second Amendment Date

The Sublicence is granted anytime after [***] following the Second Amendment Date

[***]%

[***]%

[***]%

    47

CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

iv..

where  Autolus  grants  sub-licences  under  any  of  the  Original  Program  Licences,  the  Additional  Program  Licences,  the  CAT19
Licence  and/or  under  the  CAT19  CNS  Licence  or  a  licence  under  any  Assigned  Patents  which  were  previously  licensed
under any such Program Licence or assigns an Assigned Patent which were previously licensed under any such Program
Licence  to  the  same  Sub-Licensee  or  Asignee  Entity  (irrespective  of  whether  such  sub-licences,  licences  or  patent
assignments form part of the same, separate or a connected Sublicence or Patent Sale), then subject first to any reduction of
the value of Net Revenues attributable to Intellectual Property that is not exclusively Technology (pursuant to Clause 15.2),
for  the  purposes  of  calculating  the  Sublicence  Or  Patent  Sale  Payment  due  to  UCLB  in  respect  of  the  Original  Program
Licences, the Additional Program Licences, the CAT19 Licence and/or the CAT19 CNS Licence or Assigned Patents which
were previously licensed under any such Program Licence the following shall apply:

xvi.

xvii.

xviii.

to the extent Clause 15.2.1 applies, an adjustment will be made to the total value (after any adjustment pursuant to the
provisions of Clause 15.2.1) after which the remaining value of the Net Receipts shall first be divided in proportion to the
value fairly and reasonably attributable to the Original Program Licences and/or Assigned Patents which were previously
licensed  under  the  Original  Program  Licences,  the  value  fairly  and  reasonably  attributable  to  the  Additional  Program
Licences  and/or  Assigned  Patents  which  were  previously  licensed  under  the  Additional  Program  Licences,  the  value
fairly  and  reasonably  attributable  to  the  CAT19  Licence  and  the  value  fairly  and  reasonably  attributable  to  the  CAT19
CNS Licence;
at UCLB’s request, Autolus shall provide details to UCLB of the basis of Autolus’ proposed apportionment of the total
value of Net Receipts between the Original Program Licences and/or Assigned Patents which were previously licensed
under the Original Program Licences, the Additional Program Licences and/or Assigned Patents which were previously
licensed under the Additional Program Licences, the CAT19 Licence and the CAT19 CNS Licence pursuant to (i); and,
following  an  agreed  apportionment  of  the  total  value  of  Net  Receipts  between  the  Original  Program  Licences  and/or
Assigned  Patents  which  were  previously  licensed  under  the  Original  Program  Licences,  the  Additional  Program
Licences and/or Assigned Patents which were previously licensed under the Additional Program Licences, the CAT19
Licence and the CAT19 CNS Licence pursuant to (i), the Sublicence Or Patent Sale Payment shall be calculated) using
the applicable percentages set out in the tables under Clauses 15.1.1, 15.1.2, and 15.1.3.

Each Sublicence Or Patent Sale Payment under Clause 15.1 is subject to the following:

where the Sublicence or Patent Sale includes a grant of rights to Intellectual Property which is not exclusively Technology, then
for the purposes of calculating the Sublicence Or Patent Sale Payment, the value of Net Receipts shall first be adjusted to a
value attributable to the Technology sub-licensed, or in the case of the Assigned Patents, licensed or assigned, to the Sub-
Licensee  or  Assignee  Entity  (as  applicable)  which  will  be  calculated  in  direct  proportion  to  the  value  fairly  and  reasonably
attributed to Technology licensed hereunder (or in the case of the Assigned Patents, assigned to Autolus or Assignee Entity)
as  against  all  other  Intellectual  Property  licensed  or  assigned  to  the  Third  Party  under  the  Sublicence  or  Patent  Sale  (as
applicable). At UCLB’s request Autolus shall provide details to UCLB of the basis of any proposed apportionment;

either party may refer any dispute relating to any apportionment of values either under any of Clauses 15.1.4 and/or 15.2.1 to the

Expert in accordance with Part C of Schedule 8;

Sublicence Or Patent Sale Payments in respect of Net Receipts received under a particular Sublicence or Patent Sale shall, on a

country by country basis, cease to be payable:

b.

i..

ii..

iii..

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

xx.

xix.

xxiv.

xxiii.

xxi.
xxii.

under Clause 15.1.1 where Licensed Patents are sub-licensed, or in the case of Assigned Patents licensed or assigned,
to the Sub-Licensee or Assignee Entity (as applicable) in that country, upon the later of (i) the [***]; or (ii) [***] after the
Effective Date;
under Clause 15.1.1 where no Licensed Patents are sub-licensed to the Sub-Licensee in that country or in the case of
Assigned Patents licensed or assigned, [***] after the Effective Date;
under Clause 15.1.2 upon [***] under the Additional Program Licences so sub-licensed or assigned;
under Clause 15.1.3 where Licensed Patents are sub-licensed to the Sub-Licensee in that country, upon the later of (i)
[***] in that country; or (ii) [***] after the Second Amendment Date;
under  Clause  15.1.3  where  no  Licensed  Patents  are  sub-licensed  to  the  Sub-Licensee  in  that  country,  [***]  after  the
Second Amendment Date; and
under  Clause  15.1.4,  according  to  the  relevant  periods  set  out  above  under  this  Clause  15.2.3  applicable  to  the
Technology licensed under the Original Program Licences, the Additional Program Licences and/or the CAT19 Licence;
and/or the relevant Assigned Patents which were previously licensed under any such Program Licence.
In the event that any of the Milestones are achieved by a Sub-Licensee or Assignee Entity (as opposed to by Autolus) in respect of a
particular Royalty Product then Autolus shall be entitled to offset against the corresponding Milestone Payment payable to UCLB the
amount of Sublicence Or Patent Sale Payments payable to UCLB in respect of a Sublicence to that Royalty Product or Patent Sale
of Assigned Patents Covering such Royalty Product on or before the date that the Milestone Payment is triggered. In the event that
any  Milestone  Payment  is  triggered  by  a  second  or  subsequent  Royalty  Product  (derived  from  the  same  Program  IP),  the  right  to
offset Sublicence Or Patent Sale Payments against such Milestone Payment shall exclude any previous Sublicence Or Patent Sale
Payments to the extent that they have already been offset in relation to the previous Milestone Payment provided that any excess of
a  Sublicence  Or  Patent  Sale  Payment  not  offset  shall  be  capable  of  offset  against  future  Milestone  Payments  relating  to  Royalty
Products derived from the same Program IP. Where the amount of the Milestone Payment exceeds the Sublicence Or Patent Sale
Payments that can be offset against it, Autolus shall pay to UCLB the shortfall against that Milestone Payment in accordance with
Clause 13.
REPORTING AND PAYMENT PROVISIONS

Payment Provisions for Milestone Payments and Sublicence Or Patent Sale Payments

Milestone Payments and Sublicence Or Patent Sale Payments shall all be made in accordance with the following procedure:

Autolus  shall,  within  [***],  of  the  occurrence  of  a  Milestone  or  receipt  of  Net  Receipts  triggering  a  Sublicence  Or  Patent  Sale
Payment, notify UCLB of such occurrence, and in the case of a receipt of Net Receipts Autolus shall include in its notification
confirmation  of  what  sum  is  payable  by  way  of  a  Sublicence  Or  Patent  Sale  Payment  and  its  notification  shall  include  the
information  listed  in  Schedule  10  in  so  far  as  relevant  to  the  calculation  of  a  Milestone  or  Sublicence  Or  Patent  Sale
Payment;

UCLB shall send to Autolus a VAT invoice addressed to Autolus in respect of the applicable payment due under either Clause 13

or Clause 15;

Autolus shall pay such invoice within [***] of the date of receipt of the same by Autolus.

Payment Provisions for Royalties

With effect from the First Commercial Sale of the first Royalty Product to be sold and throughout the remainder of the Royalty
Term, Autolus shall provide UCLB with a written report showing the gross selling price of those Royalty Products (triggering
Royalties or Milestones) sold by Autolus and its Sub-Licensees or any Assignee Entity in the preceding Quarter together with
the  calculations  of  Net  Sales,  which  report  shall  include  the  information  listed  in  Schedule  10  to  the  extent  relevant  to  the
calculation of Net Sales.

c.

16.
a.

i..

ii..

iii..
b.

i..

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

ii..

iii..

iv..

Quarterly reports shall be due within [***] of the close of every Quarter. Autolus shall keep accurate records in sufficient detail to

enable the Royalties and payable hereunder to be determined.

After receipt of the Quarterly report referred to in Clause 16.2.2, UCLB shall send to Autolus a VAT invoice addressed to Autolus

in respect of the applicable payment due under Clause 14 as indicated in the royalty report.

Royalties shall be due and payable within [***] of the date such invoice is received by Autolus in accordance with Clause 16.2.3.

Payments of Royalties due in whole or in part may be made in advance of such due date.

c.

Late Payments

Any payment of any amount under this Agreement not received on the due date specified in accordance with this Clause 16 shall
accrue interest thereafter on the sum due and owing from the date payment is due until the date payment is received at an annual
interest rate equal to [***].

d.

Currency Conversion

All amounts payable pursuant to this Agreement shall be payable in Pounds Sterling by bank transfer to a bank account designated
from  time  to  time  in  writing  by  UCLB.  In  calculating  Net  Sales,  Sublicence  Or  Patent  Sale  Payments  and  Royalties  under  this
Agreement, where receipts are received in a currency other than Pounds Sterling, such sums shall be calculated as Pounds Sterling
by converting such sums according to the spot rate for the Pound Sterling against the applicable currency as of midday on the day at
the end of the applicable calendar Quarter, as such rate is advertised by the Financial Times in London.

e. Withholding

All amounts due under the Agreement shall be made after deduction of any withholding taxes, charges or other duties in the country
of payment. Where any amount due to be paid under this Agreement is subject to any withholding or similar other tax, the Parties
shall take reasonable steps to do such reasonable acts and things and sign such deeds and documents as reasonably appropriate to
assist  them  to  take  advantage  of  any  applicable  double  taxation  agreements  or  other  legislative  provisions  to  reduce  the  rate  of
withholding or similar taxes with the object of paying the sums due under deduction of a reduced rate of withholding tax or on a gross
basis. In the event there is no double taxation agreement or other legislative provision or the reduced rate of withholding tax under
the relevant double taxation agreement is greater than zero per cent., Autolus (or its agent) shall promptly pay such withholding or
similar tax by deducting the relevant amount from the payment due to UCLB, and send to UCLB proof of such withholding or similar
tax in a form in accordance with the relevant taxation authority as evidence of such payments. Similarly, in so far as withholding or
similar taxes are payable on sums ultimately due hereunder but are required to be made by Autolus’s Affiliates or Sub-Licensees,
such withholding may be made and Autolus shall work with UCLB to obtain from Autolus’s Affiliates and Sub-Licensees proof that
such withholding has been properly accounted for to the relevant tax authority and such documents as are reasonably necessary to
allow UCLB to take advantage of any double taxation agreement, other legislative provision or reduced rate as may be available to it.

Royalty Audits

UCLB shall have the right to appoint, [***] on at least [***] prior written notice to Autolus, an independent certificated accountant
reasonably acceptable to Autolus to undertake an audit of Autolus’s accounts and records relevant to the sales of Royalty
Products, Net Sales and Net Receipts to verify the accuracy of any payments due in respect of Royalties and Net Receipts.
The  independent  certified  accountant  shall  spend  no  more  than  [***]  at  the  premises  of  Autolus  for  the  purpose  of
undertaking  the  audit.  Thereafter,  Autolus  shall  within  [***]  of  receiving  a  written  request  from  the  independent  accountant
provide any additional information that is reasonable and reasonably requested for the purpose of assisting with the audit,
provided that the foregoing obligation shall expire [***] after the audit. The independent auditor shall be required to enter into
a  confidentiality  agreement  on  reasonable  and  standard  terms  with  Autolus  and  shall  not  be  entitled  to  disclose  any
confidential information of Autolus from the audit but shall be able to disclose whether or not Autolus is in compliance with its

f.

i..

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

ii..

reporting  obligations  and  the  levels  of  Royalty  and  Sublicence  Or  Patent  Sale  Payments  declared  and  paid,  and  any
discrepancy in the amount of Royalties and Sublicence Or Patent Sale Payments declared as against those calculated to be
due. To  comply  with  its  obligations  under  this  Clause  16.6.1,  Autolus  shall  include  obligations  in  its  Sublicences  and  any
assignment of the Assigned Patents to obtain and make available to the auditor appropriate information from Sub-Licensees
and  any  Assignee  Entity  of  an  Assigned  Patent  to  enable  the  independent  auditor  to  verify  the  accuracy  of  Royalties,  Net
Receipts and Sublicence Or Patent Sale Payments.

If,  as  a  result  of  an  audit  being  undertaken,  any  additional  amount  is  found  to  be  owed  by  Autolus  to  UCLB,  such  additional
amount shall be paid within [***] after receipt of the accountant’s report, along with interest at the annual interest rate of [***]
from  the  date  that  such  additional  amount  should  have  first  been  paid  until  paid  in  full.  If  the  amount  underreported  as
Royalties or Sublicence Or Patent Sale Payments for the relevant periods that are the subject of the audit, are in excess of
[***].  in  the  relevant  audit,  then  Autolus  shall  in  full  and  final  settlement  of  any  claim  of  breach  reimburse  UCLB  for  those
reasonable  and  customary  costs  charged  by  the  independent  auditor  for  conducting  such  audit  (upon  production  of
accompanying receipted invoices in respect of the same). If the accountant determines that there has been an overpayment
by Autolus, the amount of such overpayment shall be refunded to Autolus within [***] after receipt of the accountant’s report,
or at Autolus’s discretion, set-off against a future payment of Royalties or Sublicence Or Patent Sale Payments.

g.

Fair Market Value

Any disagreement between the Parties as to the fair market value for the purpose of calculating any Net Sales pursuant to Part A of
Schedule 8 of this Agreement shall be referred to an expert for resolution in accordance with the provisions of Part C of Schedule 8.
The value of such Net Sales in dispute shall (i) not be included in the calculation of the percentage of underreported royalties referred
to in Clause 16.6.2 for the purposes of determining responsibility for the auditor's fees; and (ii) be excluded from any late payment
charges or allegations of breach for non-payment until such time as the dispute is resolved, a value attributed and at least [***] has
passed from such final determination. Notwithstanding the foregoing provision, if the expert determines that the fair market value is
such that UCLB is entitled to additional sums, UCLB shall be entitled to charge interest on any outstanding amount on a daily basis
at a rate equivalent of [***], such interest shall be payable from the date UCLB issues a notice disputing the fair market value until the
date the UCLB receives such additional payment.
BUY-OUT OPTION

On  a  Royalty  Product  by  Royalty  Product  basis,  once  the  aggregate  Net  Sales  for  a  Royalty  Product  have  exceeded  GBP  £[***],
Autolus shall thereafter have a right, exercisable on written notice at any time, to negotiate with UCLB to buy out UCLB’s rights to
Royalties, Milestone Payments and Sublicence Or Patent Sale Payments Sales Milestone Payments on such Royalty Product (for
each  Royalty  Product  a  “Buy-Out  Option”).  The  reference  to  “buy  out”  in  this  Clause  shall  mean  that  UCLB  shall  cease  to  be
entitled to Royalties in exchange for some other cash consideration.

17.
a.

b.

Upon exercising the Buy-Out Option by way of Autolus serving a written notice on UCLB, the following shall apply until expiry of [***]

after the date Autolus’s notice is deemed served (unless extended by agreement between the Parties):

Autolus and UCLB shall promptly and actively negotiate throughout the [***] period, in good faith and acting reasonably, fair and

reasonable terms for, and the, conclusive agreement upon which the buy-out may be exercised;

in so far as UCLB does not actively and properly participate in such negotiations or does not act reasonably or in good faith, the
[***] period shall be extended by a period equal to, or otherwise fairly calculated to, compensate for any delay in or absence
from a negotiation by UCLB in accordance with the principles under Clause 17.2.1.

INTELLECTUAL PROPERTY PROSECUTION AND MAINTENANCE

Ownership

i..

ii..

18.
a.

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

i..
ii..

iii..

b.

i..

ii..
c.

i..

Nothing in this Agreement shall assign or purport to assign any Intellectual Property rights owned by one Party to the other Party.
With  the  exception  of  the  Assigned  Patents,  ownership  of  which  has  been  assigned  to  Autolus  with  effect  from  the  Third
Amendment Date, UCLB is and shall at all other times remain the sole and exclusive owner of all right, title and interest in
and to any and all Program IP. UCLB shall not assign, mortgage, encumber or otherwise gift or provide an option over any of
the Licensed Patents or Program IP without the prior written consent of Autolus.

Autolus is and shall at all times remain the sole and exclusive owner of all right, title and interest in and to any and all Intellectual
Property that it owns or Controls (other than by virtue of the licences granted hereunder) as of or after the Effective Date.

Patent Prosecution

In respect of the Retained Patents:
xxv.
xxvi.

UCLB shall not Surrender any of them without the prior written consent of Autolus;
UCLB shall ensure that all documents and correspondence that it, or its agents or other licensees receive in connection
with any of such Retained Patents shall be promptly and in any event within seven (7) days forwarded to Autolus, and
without limiting the foregoing, UCLB shall keep Autolus promptly informed in advance of any steps taken regarding the
RQR8 Patent Rights and the [***] Existing Patent;
UCLB shall promptly notify Autolus of any threatened or actual claim of invalidity or revocation or opposition of any of
the Retained Patents and shall provide full details and all such information available to it regarding such threatened or
actual claim;
in respect of the RQR8 Patent Rights, if the validity of any of them is challenged and UCLB (or its other licensees) does
not defend such challenge, then Autolus shall have the right (but not the obligation) to control, direct and conduct such
proceedings.  UCLB  shall  do  (or  not  do)  all  such  things  as  are  reasonably  directed  by  Autolus  to  enable  Autolus  to
control,  direct  and  conduct  such  proceedings,  including  allowing  Autolus’s  legal  representatives  to  conduct  such
litigation in UCLB’s name where required or beneficial provided that Autolus indemnifies UCLB and/or its Affiliates for
any Third Party costs, damages, expenses or liability incurred by UCLB and/or its Affiliates as a direct result of assisting
Autolus subject to Clause 18.8. Autolus shall pay UCLB’s and/or its Affiliates for any reasonable (economy) travel and
reasonable subsistence costs incurred by UCLB and/or its Affiliates as a result of assisting Autolus under this Clause
18.2.1(iv).  Autolus  shall  consult  and  co-operate  with  UCLB  and  its  licensees  outside  of  the  RQR8  Field  if  it  elects  to
defend such challenge;
UCLB shall provide assistance to and co-operate with Autolus in accordance with this Clause 18 without any further cost
to  Autolus,  save  that  (i)  if  UCLB  personnel  are  required  to  participate  in  any  opposition  proceeding  (or  comparable
proceeding  before  patent  offices  and  courts)  which  requires  full  time  involvement  for  more  than  [***]  per  annum  per
Program IP under any Program Licence, then for such excess co-operation beyond the [***] for that Program IP Autolus
shall  reimburse  UCLB  its  reasonable  costs,  and  (ii)  this  provision  shall  be  without  prejudice  to  the  indemnity  given  in
Clause 18.2.1(iv); and,
any enforcement of the Licensed Patents shall be subject to Clause 19.

xxvii.

xxviii.

xxix.

xxx.
[***]

In respect of the RQR8 Patent Rights:

Autolus  shall  be  responsible  for  [***].  of  those  Patent  Prosecution  Costs  properly  incurred  by  UCLB  in  the  prosecution  and
maintenance of the RQR8 Patent Rights provided that UCLB’s Third Party licensee outside the RQR8 Field and/or UCLB is
responsible for and pays [***] of those Patent Prosecution Costs;

ii..

UCLB  shall  keep  Autolus  informed  of  developments  in  the  prosecution  and  maintenance  of  the  RQR8  Patent  Rights  and  shall

provide Autolus with copies of

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

iii..

iv..

v..

d.
e.

f.

i..

ii..

iii..

g.

i..

ii..

all material correspondence to and from its patent attorneys or patent offices in relation to the RQR8 Patent Rights and shall
provide Autolus reasonable notice of and the opportunity at its own cost to participate in any conference calls or meetings
with UCLB’s patent attorneys in relation to the drafting, filing, prosecution, and maintenance of the RQR8 Patent Rights;
UCLB  shall  consult  with  Autolus  in  connection  with  UCLB’s  and  its  Third  Party  licensee’s  strategy  for  the  prosecution  and

maintenance of the RQR8 Patent Rights;

UCLB  shall  take  into  account  any  reasonable  comments  and  suggestions  of  Autolus  in  relation  to  the  prosecution  and

maintenance of the RQR8 Patent Rights; and

UCLB shall notify Autolus in advance of any step(s) UCLB proposes be taken which would change the specification or reduce the
scope  of  the  claims  of  the  RQR8  Patent  Rights  in  the  RQR8  Field,  and  having  done  so  shall  take  into  account  any
reasonable comments and suggestions promptly proposed by Autolus in relation to such steps.

Section intentionally left blank.
Autolus  and  UCLB  shall,  promptly  after  the  Effective  Date,  and  thereafter  throughout  the  Term  appoint  a  designated  and  named
member of its respective personnel, experienced in and responsible for Intellectual Property matters, which person shall act as the
liaison between Autolus and UCLB (and UCLB’s other licensees as necessary) with respect to the Licensed Patents and obligations
thereto  under  this  Agreement  and  shall  make  themselves  available  at  reasonable  times  and  on  reasonable  notice  to  address  any
matters concerning the Licensed Patents.
Validation and Maintenance of Assigned Patents

Autolus shall have the sole discretion to determine, on a reasonable basis and following its notification to UCLB, in which countries to
maintain  or  Surrender  the  Assigned  Patents.  Notwithstanding  the  foregoing  discretion,  if  Autolus  wishes  to  Surrender  any  of  the
Assigned Patents in any of the Core Countries then the following shall apply:

prior to taking any steps to Surrender an Assigned Patent in a Core Country, Autolus shall first provide UCLB with at least [***]

notice of its intention identifying the Assigned Patent and applicable Core Countries;

UCLB  shall  have  a  right  of  step-in  (to  be  exercised  within  [***]  of  notice  from  Autolus  under  Clause  18.6.1)  to  take  over  such
Assigned Patent in the applicable Core Country and if it exercises such right (i) UCLB shall thereafter be responsible for all
costs  and  expenses  associated  with  such  Assigned  Patent  for  that  applicable  Core  Country;  (ii)  Autolus’s  licence  to  that
Assigned Patent for that applicable Core Country shall continue in such Core Country and the Assigned Patent concerned
shall continue to be a Licensed Patent in respect of such Core Country; and,

if UCLB does not exercise its step-in right in accordance with Clause 18.6.2, then Autolus shall be entitled without breach of this

Agreement to Surrender such Assigned Patent in such Core Countries.

SPCs and Patent Notifications

Without the prior written consent of Autolus, UCLB shall not file any supplementary protection certificate or patent term extension
right (“SPC”)  under  any  Retained  Patents  with  respect  to  the  issue  of  any  Regulatory  Approval  (including  any  Marketing
Approval)  for  any  product.  Upon  Autolus’s  request,  UCLB  shall  file  and,  at  Autolus’s  direction,  control  and  expense,
prosecute an application for an SPC against any of the Retained Patents with respect to any product.

Where any country in the Territory requires the holder of a Regulatory Approval with respect to a medicinal product or medical
device to designate one or more Patent Rights as being Patent Rights that protect such medicinal product or medical device
(including the purple book listing required by the FDA) (an “Purple Book Reference”), then Autolus shall have the sole right
to specify which (if any) Patent Rights should be listed in such references and UCLB shall list any of the Retained Patents if
Autolus wishes to do so.

h.

Indemnity Conditions

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

i..

ii..

iii..
iv..

19.
a.

Autolus’s obligation to continue to indemnify UCLB pursuant to Clauses 18.2.1(iv) and 19.2.2(ii) is conditional upon:

UCLB taking those steps, doing those things or refraining from doing those things requested of it by Autolus for the duration of the

indemnification;

UCLB  not  making  any  admission  or  settlement  (or  taking  steps  to  do  so)  concerning  the  proceedings  without  the  prior  written

consent of Autolus;

Autolus having sole conduct of the applicable proceedings for the duration of the indemnification;
any damages, account of profits, financial remedy or costs recovered from Third Parties (whether in UCLB’s name or otherwise)

in respect of the applicable proceedings being for the sole account of Autolus.

INTELLECTUAL PROPERTY ENFORCEMENT

A Party shall notify the other of any information it has regarding any Third Party infringement of (i) the Intellectual Property licensed
under or pursuant to this Agreement; and/or (ii) the Assigned Patents, in each case in so far as such infringements are related to any
products, services or processes.

b.

In  respect  of  any  alleged,  threatened  or  actual  infringement  of  the  Intellectual  Property  licensed  or  sub-licensed  hereunder

(“Enforcement Action”) the following, subject to Clause 19.3, shall apply:

i..

With respect to the Assigned Patents, Autolus shall have the sole right to determine whether or not it wishes to bring proceedings
for the Enforcement Action. With respect to other Licensed Patents, Autolus shall have the first right to determine whether or
not to bring such proceedings, and only if Autolus elects not to bring proceedings itself shall UCLB have the right to decide
whether or not to bring proceedings for the Enforcement Action (but in doing so UCLB shall have regard to the advice and
recommendations of Autolus);

ii..

where  Autolus,  in  exercising  its  right  under  Clause  19.2.1,  decides  to  enforce  any  of  the  Licensed  Patents  or  other  Intellectual

Property licensed hereunder, then:

xxxi.
xxxii.

at Autolus’s expense, Autolus shall have the right to control, direct and conduct such proceedings;
UCLB shall allow Autolus’s legal representatives to conduct any litigation in UCLB’s name (i) where required by law in
the country of the Enforcement Action or (ii) to the extent beneficial to the enforcement or relief sought; and (iii) in doing
so UCLB shall do (or not do) all such things as are directed by Autolus to enable Autolus to control, direct and conduct
such  proceedings  provided  that  Autolus  indemnifies  UCLB  and/or  its  Affiliates  for  any  Third  Party  costs,  damages,
expenses  or  liability  incurred  by  UCLB  and/or  its  Affiliates  directly  as  a  result  of  assisting  Autolus  control,  direct  and
conduct  such  proceedings  subject  to  Clause  18.8  (it  being  acknowledged  that  UCLB  shall  have  the  right  to  be
separately advised (but not represented before the proceedings) by its own counsel at UCLB’s own expense). Autolus
shall  pay  UCLB’s  and/or  its  Affiliates’  costs  for  any  reasonable  (economy)  travel  and  reasonable  subsistence  costs
incurred by UCLB and/or its Affiliates as a result of assisting Autolus under this Clause (ii);
UCLB  shall  use  its  reasonable  endeavours  to  procure  that  UCL  and  MP  shall  do  all  such  things  as  are  reasonably
directed by Autolus to assist or enable Autolus to control, direct and conduct such proceedings;
Autolus shall have the right to nominate, change or amend any Purple Book Reference and UCLB shall co-operate in
such nomination, change or amendment to list any of the Licensed Patents if Autolus wishes to do so; and,
Autolus  shall  keep  UCLB  promptly  and  fully  informed  of  any  and  all  steps  and  events  in  any  proceedings  (including
promptly  responding  to  any  requests  for  information  and  allowing  UCLB  to  attend  any  meetings)  and  shall  give  due
consideration to any reasonable comments and suggestions of UCLB with respect to such Enforcement Action;
UCLB  shall  keep  Autolus  promptly  and  fully  informed  of  any  and  all  steps  and  events  in  any  proceedings  (including  promptly

xxxv.

xxxiii.

xxxiv.

responding to any requests for information and allowing Autolus to attend any meetings) which are not being

iii..

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

iv..

directed or controlled by Autolus relating to any of the Licensed Patents or other Intellectual Property licensed hereunder and
shall give due consideration to any reasonable comments and suggestions of Autolus with respect to such action;

any  recovery  of  damages  or  other  financial  remedy  obtained  in  respect  of  the  Enforcement  Action  shall,  after  deduction  of  all
litigation costs (comprising attorney fees, expert fees, taxes, charges, disbursements, court fees and other costs incurred in
connection with proceedings), be (i) in the case of an Enforcement Action in respect of a Competitive Product be treated as
Net Sales, and (ii) in all other cases be for the account of Autolus; and,

v..

any  defence  of  the  validity  of  the  Licensed  Patents,  where  validity  is  put  in  issue  after  commencement  of  proceedings  for  the

Enforcement Action shall, notwithstanding the provisions of Clause 18 shall be subject to this Clause 19.

c.

For the avoidance of doubt, an Enforcement Action and Autolus’s right to conduct such action where the RQR8 Patent Right has been

infringed, shall only be in so far as the infringement is within the RQR8 Field.

d. Where either Party becomes aware of an infringement or potential infringement of the RQR8 Patent Rights, the Parties shall consult
with  each  other  and  with  UCLB’s  Third  Party  licensee  outside  of  the  RQR8  Field  to  decide  the  best  way  to  respond  to  such
infringement. Where Autolus pursues any Enforcement Action of the RQR8 Patent Rights in accordance with Clause 19.3 and if the
alleged infringement is both within and outside the RQR8 Field or there is any challenge to the validity of the RQR8 Patent Rights,
Autolus shall co-operate with UCLB’s Third Party licensee in relation to the conduct of such action and its settlement.
CONFIDENTIALITY

The Parties acknowledge that in connection with this Agreement, either Party may disclose or may have disclosed itself or on its behalf
(a  “Disclosing  Party”)  to  the  other  Party  (each  a  “Recipient  Party”)  information  belonging  to  such  Party  which  information  is
marked or stated in writing to be “confidential” or “trade secret” information or where the circumstances of the disclosure and/or the
nature  of  the  information  otherwise  reasonably  give  notice  of  the  confidential  character  of  the  information  (“Confidential
Information”). All such Confidential Information of a Disclosing Party shall, subject to Clause 20.3, be maintained in confidence by
each Recipient Party and shall not be used by the Recipient Party for any purpose except for its proper execution of its obligations
under  this  Agreement  and  the  Exploitation  of  any  Product  or  as  otherwise  expressly  authorised  (including,  in  respect  of  any
confidential  Know-How  to  the  extent  such  Know-How  is  licensed  to  the  Receiving  Party)  under  this  Agreement  or  to  the  extent
otherwise  agreed  in  writing  by  the  Disclosing  Party  provided  that  the  Recipient  Party  may  disclose  any  Confidential  Information
disclosed to it by the Disclosing Party to the extent that such disclosure by the Recipient Party is:

to  its  employees,  directors,  consultants  or  sub-contractors  but  only  on  a  “need  to  know”  basis  provided  each  such  employee,
director, consultant or sub-contractor is subject to obligations of confidentiality consistent with the obligations of confidentiality
in this Clause 20;

to its sub-licensees in respect of confidential Know-How that is licensed to the Recipient Party, but only on a “need to know” basis
provided each such sub-licensee is subject to obligations of confidentiality consistent with the obligations of confidentiality in
this Clause 20;

to an Ethics Committee or Regulatory Authority in connection with any Ethics Committee Application or seeking or maintaining
any Regulatory Approval for any product or therapy in accordance with this Agreement; provided, however, that reasonable
measures shall be taken to assure confidential treatment of such Information;

on  a  “need  to  know”  and  confidential  basis  to  its,  or  its  Affiliates’,  legal  and  financial  advisors  to  the  extent  such  disclosure  is
reasonably necessary in connection with such Party’s activities as expressly permitted by this Agreement or for the conduct
of its, or such Affiliates’, business;

to a prospective acquirer or licensee and such Third Party’s employees, advisors and representatives in each case on a “need to

know” confidential basis for the sole purpose of considering such transaction provided that such persons are

20.
a.

i..

ii..

iii..

iv..

v..

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

under substantially similar obligations of confidentiality and non-use as the Recipient Party is pursuant to this Clause 20.

b.

Throughout the Term of this Agreement and thereafter, each Recipient Party shall exercise a reasonable degree of care being at least
the same degree of care as it uses to protect its own Confidential Information of similar nature to preserve the confidentiality of all
Confidential Information of the Disclosing Party. Each Recipient Party shall safeguard Confidential Information against disclosure to
third parties, including Affiliates, employees and persons working or consulting for such Party that do not have an established current
need to know such Confidential Information for purposes in connection with this Agreement or to whom the Recipient Party is not
entitled to disclose the same pursuant to this Clause 20.

c.

The obligation of confidentiality contained in this Clause 20 shall not apply to any part of any Confidential Information of the Disclosing

Party:

that was in the possession of the Recipient Party, without any restriction on use or disclosure, prior to receipt from the Disclosing

Party;

that was at the time of disclosure by or on behalf of the Disclosing Party, in the public domain by public use, publication or general

knowledge;

that became general or public knowledge through no fault of a Recipient Party following disclosure hereunder;
that was properly obtained, without confidentiality or non-use restrictions, by the Recipient Party from a Third Party who was not

under a confidentiality or non-use obligation to the Disclosing Party;

that was documented to have been independently developed by or on behalf of the Recipient Party without the assistance of the

Confidential Information of the Disclosing Party.

The foregoing obligations of confidentiality and non-use shall not be breached by a Recipient Party disclosing Confidential Information
of the Disclosing Party to the extent the same is required to be disclosed by order of any court, governmental authority, Regulatory
Authority or other regulatory body (including any listing authority or financial regulator) provided, however, that the Recipient Party
should give the Disclosing Party prior notice of any such disclosure so as to afford the Disclosing Party a reasonable opportunity to
seek, at the expense of the Disclosing Party such protective orders or other relief as may be available in the circumstances.

Except for any press release agreed by the Parties, neither party shall during the Term, disclose any financial terms of this Agreement
without  the  prior  written  consent  of  the  other  Party  except  for  such  disclosure  as  may  be  reasonably  necessary  to  either  Party's
bankers, investors, attorneys or other professional advisors or in connection with any actual or proposed merger, sale or acquisition
or as may be required by law in the offering of securities or in securities or regulatory filings or otherwise.

The  Parties  acknowledge  that  confidential  information  may  have  been  disclosed  pursuant  to  the  CDA  to  employees,  partners  and
representatives  of  Syncona  LLP  who  themselves  may  provide  services  or  advice  to  or  sit  on  the  board  of  Autolus  UCLB  hereby
agrees that notwithstanding the terms of the CDA employees, partners and representatives of Syncona Management LLP, Syncona
Partners  LLP  and  Syncona  LLP  who  received  confidential  information  from  UCLB  under  the  CDA  shall  be  entitled  to  disclose  the
same to Autolus and its employees, directors, consultants or sub-contractors subject to the terms of this Clause 20.
WARRANTIES AND COVENANTS

Autolus and UCLB each respectively represent and warrant to the other at the Effective Date that each of the warranties at Part A of
Schedule 9 in respect of itself, its Affiliates, its assets, its knowledge or its Intellectual Property is accurate as at the Effective Date.
UCLB represents and warrants to Autolus at the Effective Date that except as disclosed in a Disclosure Letter dated as of the Effective

Date each of the warranties at Part B of Schedule 9 is accurate at the Effective Date.

UCLB  represents  and  warrants  to  Autolus  at  the  Amendment  Date  that  except  as  disclosed  in  a  Disclosure  Letter  dated  as  of  the

Amendment Date each of the warranties at Part C of Schedule 9 is accurate at the Amendment Date.

i..

ii..

iii..
iv..

v..

d.

e.

f.

21.
a.

b.

c.

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

d.

e.

f.

g.

h.

i..

ii..
22.
a.

UCLB represents and warrants to Autolus at the Third Amendment Date that except as disclosed in a Disclosure Letter dated as of the

Third Amendment Date each of the warranties at Part E of Schedule 9 is accurate at the Third Amendment Date.

For warranties given by UCLB in respect of its knowledge or awareness, such knowledge or awareness shall be limited to the actual
knowledge  or  awareness  at  the  Effective  Date  or  the  Amendment  Date,  as  applicable,  (without  having  made  any  searches  or
enquiries, other than of UCLB’s Biopharm marked database) of the senior management team of UCLB (director status and above)
and [***].

For  warranties  given  by  UCLB  at  the  Second  Amendment  Date  in  respect  of  its  knowledge  or  awareness,  such  knowledge  or
awareness  shall  be  limited  to  the  actual  knowledge  or  awareness  at  the  Second  Amendment  Date,  (without  having  made  any
searches or enquiries, other than of UCLB’s Biopharm market database) of the senior management team of UCLB (director status
and above) and [***].

For warranties given by UCLB at the Third Amendment Date in respect of its knowledge or awareness, such knowledge or awareness
shall  be  limited  to  the  actual  knowledge  or  awareness  at  the  Third  Amendment  Date,  (without  having  made  any  searches  or
enquiries, other than of UCLB’s Biopharm market database) of the senior management team of UCLB (director status and above)
and [***].

Save  for  the  warranties  and  representations  expressly  set  forth  above  by  reference  to  Schedule  9,  (i)  the  Parties  exclude  all  other
warranties and representations of any kind, whether express or implied in connection with this Agreement, save that the foregoing
shall not exclude or limit any liability for fraud or fraudulent misrepresentation and (ii) without prejudice to the above, UCLB does not
give any warranty, representation or undertaking:

as to the efficacy, usefulness, fitness for purpose, quality, safety or commercial or technical viability of the Technology and/or any

Royalty Products;

that any of the Licensed Patents are or will be valid or will proceed to grant.

LIMITATION OF LIABILITY
Special, Indirect and Other Losses

In no event shall any Party or any of their respective Affiliates be liable for breach of contract, statutory duty, negligence or in any
other  way  for  special,  indirect,  incidental,  punitive  or  consequential  damages  or  for  any  indirect  economic  loss  or  indirect  loss  of
profits suffered by any other Party or their respective Affiliates.

b.

No Exclusion

UCLB’s total aggregate liability to Autolus for any and all loss or damage suffered by Autolus as a result of breach of or otherwise in
connection  with  this  Agreement  and  the  Patent  Assignment  in  respect  to  any  and  all  claims  arising  under  this  Agreement  or  the
Patent Assignment shall be limited to GBP £[***], provided that in the event that any breach of warranty 1.5 and/or 1.6 of Schedule 9
gives rise to loss suffered by Autolus in excess of this cap, the cap shall be increased to the sum of GBP £[***] such that UCLB’s total
aggregate liability for any and all claims arising under or in connection with this Agreement shall be limited to the sum of GBP £[***].
Nothing in this Agreement shall limit or be construed to limit in any way any liability a Party (or its respective Affiliates) may have to the
other Party (or its Affiliates) under this Agreement in respect of (i) death or personal injury caused by that Party’s (or its respective
Affiliates’)  negligence;  (ii)  any  fraud  or  fraudulent  misrepresentation  or  (iii)  any  other  liability  which,  by  rule  of  law,  may  not  be
excluded or limited by contract between parties.
INDEMNITY AND INSURANCE

Subject to Clause 23.2, Autolus shall indemnify and hold harmless:

the CRUK Funders and their respective officers and employees as well as those researchers and contributors who participated in
the  conduct  of  the  CRUK  Study,  including  Great  Ormond  Street  Hospital  NHS  Foundation  Trust  (“CRUK  Indemnified
Parties”), from and against any and all Third Party (excluding any of the Indemnified Parties) claims, proceedings, liabilities,
damages  and  expenses  (including,  reasonable  legal  fees)  arising  from  or  in  connection  with  Autolus’s  and/or  its
sublicensees’ exercise of the CRUK Rights granted to Autolus hereunder;

c.

23.
a.

i..

ii..

UCLB  and/or  its  Affiliates  and  any  officers,  employees,  contractors  and/or  consultants  of  UCLB  and/or  its  Affiliates  (“UCLB

Indemnified Parties”), from and

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

iii..

against  any  and  all  Third  Party  (excluding  any  of  the  Indemnified  Parties)  claims,  proceedings,  liabilities,  damages  and
expenses (including, reasonable legal fees) arising from or in connection with Autolus’s and/or its sublicensees’ exercise of
any of the rights (i) granted to Autolus hereunder or (ii) assigned to Autolus under the Patent Assignment;

each of the foregoing Third Party claims, proceedings, liabilities, damages and expenses (including, reasonable legal fees) being
an  “Indemnity  Claim”  and  CRUK  Indemnified  Parties  and  UCLB  Indemnified  Parties  collectively  being  the  “Indemnified
Parties” or individually an “Indemnified Party”. For the purposes of this Clause, “CRUK Rights” shall mean those parts of
the licences hereunder to Autolus where (i) CRT is the head licensor to UCLB which includes the GD2 Clinical Study Results
and  the  [***];  or  (ii)  the  CRUK  Funders  have  directly  funded  the  development  of  the  licensed  Intellectual  Property  being
certain of the BCMA Program IP.

b.

Autolus’s  obligation  to  indemnify  the  Indemnified  Parties  in  respect  of  an  Indemnity  Claim  is  dependent  upon  compliance  with  the

following provisions:

i..

ii..

iii..

promptly  after  receipt  by  an  Indemnified  Party  of  any  claim  or  alleged  claim  or  notice  of  the  commencement  of  any  action,
administrative or legal proceeding, or investigation to which the indemnity provided for in Clause 23.1 may apply, UCLB or
the Indemnified Party shall give written notice to Autolus of such fact and provide all information available to it and relevant to
the Indemnity Claim to Autolus;

the Indemnified Party shall permit Autolus to have sole control, conduct, defence and settlement of the Indemnity Claim and shall
not  make  any  admission  or  reach  any  settlement  with  the  Third  Party  other  than  at  Autolus’s  written  direction  or  with
Autolus’s prior written consent;

the Indemnified Party shall co-operate in good faith with Autolus in the conduct of any defence or settlement and shall provide
reasonable assistance and do all things as may be reasonably required to enable any Indemnified Claim to be defended and
shall provide promptly to Autolus (i) copies (or originals where available) of all correspondence and documents relevant to
the  Indemnified  Claim;  (ii)  reasonable  access  to  all  personnel  of  the  Indemnified  Party  (including  its  consultants)  to  assist
with defence of the Indemnified Claim and (iii) all other information, documents or assistance as may be reasonably required;

iv..

Autolus shall have the right at its sole discretion to bring any counterclaim in the name of:

xxxvi.

xxxvii.

any CRUK Indemnified Parties provided it receives the prior written consent of the applicable CRUK Indemnified Parties
(such consent not to be unreasonably withheld or delayed) to bring such counterclaim; and/or,
any UCLB Indemnified Parties provided it first notifies the applicable UCLB Indemnified Parties of its intention to bring
such counterclaim.

v..

Autolus shall have the right at its sole discretion to settle or compromise any Indemnity Claim except that Autolus shall not without

the prior written consent of the Indemnified Party:

xxxviii.
xxxix.

admit any liability on the part of any Indemnified Party; or,
in respect of any product liability claims the subject of the Indemnity Claim, not make any public statement that amounts
to any admission of wrongdoing on the part of the Indemnified Party.

Should any damages, financial remedy, costs or other recovery be made in favour of the Indemnified Party or Autolus, such sums

shall be for the sole account of Autolus.

Autolus  shall  consult  with  the  Indemnified  Party  on  the  defence  and/or  settlement  of  any  Indemnified  Claim  and  in  so  far  as  is
reasonable, Autolus shall consider any reasonable suggestions of the Indemnified Party in the conduct of the defence or settlement
of the Indemnity Claim.

vi..

c.

d.

Should Autolus assume conduct of the defence the Indemnified Party may retain separate legal advisers at its sole cost and expense,

save that if Autolus denies the applicability of the indemnity or reserves its position in relation to the same, the indemnity

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

e.

f.

24.
a.

b.

c.

d.

i..

in  Clause  23.1  shall  extend  to  the  Indemnified  Party’s  costs  and  expenses  so  incurred  if  Autolus’s  position  is  established  to  be
substantively incorrect.

Upon  termination  or  expiry  of  this  Agreement,  Autolus’s  obligation  to  provide  an  indemnity  to  the  Indemnified  Parties  pursuant  to
Clause 23.1 for any actions or proceedings shall expire [***] after the termination or expiry of the Agreement, save in respect of any
product liability actions or proceedings in which case no limit of time shall apply.

Autolus shall maintain, at its own cost, comprehensive and customary insurance including product liability insurance in an amount and
for a period sufficient to cover Autolus’s liabilities under this Agreement. [***] Autolus shall upon UCLB’s request, provide UCLB with
a copy of the latest certificate evidencing the coverage required hereby, and the amount thereof. UCLB shall be entitled to provide a
copy of such certificate to CRT. Such insurance shall be with a reputable insurance company.
TERMINATION

This Agreement shall take effect on the Effective Date and shall continue thereafter unless and until terminated in accordance with this
Clause 24 or if earlier until such time as the Royalty Term in each country in the Territory has expired and no further Sublicence Or
Patent Sale Payments or Milestone Payments are due, in which case all Licences, the Manufacturing Licence and UCL Background
Licence granted hereunder shall automatically convert to a perpetual, irrevocable, royalty free licence (the “Term”).

The Parties may, by mutual written agreement, agree that this Agreement be terminated in whole or on a Program Licence by Program

Licence basis.

Autolus may terminate this Agreement upon thirty (30) days prior written notice to UCLB on a (i) Program Licence by Program Licence

basis; or (ii) in respect of all Licences.

Either  Party  (a  “Non-Defaulting  Party”)  may  terminate  this  Agreement  (without  prejudice  to  its  other  rights  and  remedies)  with

immediate effect by written notice to the other Party (the “Defaulting Party”) if:

the Defaulting Party commits a material breach of its material obligations under this Agreement (it being acknowledged that UCLB
may not terminate under this Clause for any breach of Clause 11) and, if the breach is capable of remedy, fails to remedy it
during the longer period of (i) [***] or (ii) such other period as the Parties may, acting in good faith having regard to the nature
of the breach and the time required to remedy the same, agree in writing (the “Notice Period”), in each case starting on the
date of receipt of notice from the Non-Defaulting Party which specifies the breach in reasonable detail and requires it to be
remedied. If the Defaulting Party in good faith disputes that it has committed a material breach under this Agreement, or that
it has not cured the claimed breach within the Notice Period, it may refer the matter to the dispute resolution procedure under
Clause 33 provided that the termination shall not be effective until conclusion of all dispute resolution procedures pursued by
any Party including any proceedings before a court to determine the validity of the termination notice; or

the Defaulting Party suffers an Insolvency Event.

ii..
e. Without  prejudice  to  Clause  24.4,  UCLB  may  (unless  the  non-payment  is  remedied  in  the  [***])  terminate  this  Agreement  upon  [***]
prior written notice if Autolus has not paid sums in excess of £[***] which are properly due under this Agreement, provided that the
sums are not subject to a bona fide dispute between the Parties. In the event of a payment dispute:

each Party shall provide the other with written reasons as to why it believes any disputed sums are either not payable or payable

(as applicable):

the  Parties  shall  attempt  to  resolve  the  payment  dispute  by  following  the  escalation  process  for  dispute  resolution  set  out  in

Clause 33.2; and

where the Parties are unable to resolve the payment dispute by way of the escalation process set out in Clause 33.2, the Parties

shall seek to resolve the dispute by following the dispute resolution procedure set out in Clauses 33.3 and/or 33.9.
Following resolution of any payment dispute, Autolus shall pay UCLB any amount agreed or adjudged to be due, together with interest
thereon, such interest shall be payable at a rate of [***], for the period from when such amount was originally due until the date that
UCLB receives the agreed or adjudged sums. If Autolus fails to pay UCLB the requisite payment within [***] of the date of receipt of
invoice from UCLB requesting the agreed or

i..

ii..

iii..

f.

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

g.

h.

i.

25.
a.

i..
ii..
iii..

adjudged sums, UCLB shall be entitled to terminate this Agreement with immediate effect by written notice at the end of the [***].

If the disputed amount is a part of a larger payment, Autolus shall pay UCLB the non-disputed amount no later than [***] after receipt of
the  invoice  from  UCLB  requesting  the  non-disputed  amount,  failing  which  UCLB  shall  be  entitled  to  terminate  the  Agreement
pursuant to the provisions of Clause 24.5.

The  Parties  shall  continue  to  perform  their  obligations  under  this  Agreement,  notwithstanding  any  dispute  between  the  Parties  with

respect to payment.

Save as provided under this Clause 24 (but without prejudice to the rights to terminate individual Program Licences under Clause 11),

the Parties shall have no other right to terminate this Agreement including under any right according to common law.
CONSEQUENCES OF TERMINATION

Upon termination of a Program Licence under Clause 24, other than the Background Licence granted pursuant to Clause 2.1.2 and the

Manufacturing Licence granted pursuant to Clause 2.3.2:

the applicable Program Licence shall automatically terminate;
Autolus shall, at its expense, assign to UCLB any Assigned Patents relating solely to such Program Licence; and,
Autolus shall cease to have rights under this Agreement to Exploit the Royalty Product applicable to such Program Licence and
all of its rights and obligations under this Agreement concerning such Royalty Product shall cease including the rights under
Clauses 2.1.1 and 2.3.1 applicable to such Program Licence.

b.

Upon termination of a Program Licence under Clause 11.1.5, 11.2.3, or 11.3.2 other than the Background Licence granted pursuant to

i..
ii..
iii..

iv..

c.

d.

i..

ii..
iii..

iv..

Clause 2.1.2 and the Manufacturing Licence granted pursuant to Clause 2.3.2:

the applicable Program Licence shall automatically terminate;
Autolus shall, at its expense, assign to UCLB any Assigned Patents relating solely to such Program Licence;
Autolus shall cease to have rights under this Agreement to Exploit the Royalty Products applicable to such Program Licence and
all of its rights and obligations under this Agreement concerning such Royalty Products shall cease including the rights under
Clauses 2.1.1 and 2.3.1 applicable to such Program Licence; and,

subject to Clause 25.3, UCLB shall have the option to negotiate with Autolus to agree terms for the grant of an exclusive licence
to those Autolus Improvements Controlled by Autolus and free of any restriction or encumbrance, such licence to be limited
to the Exploitation of those Autolus Improvements relating to the specific Program Licence relating to Program IP that has
been terminated, provided that those Autolus Improvements are only used together with that same applicable Program IP.

Upon termination of a Program Licence under Clause 11.4.3, UCLB and Autolus shall negotiate the terms for any licence pursuant to
Clause  25.2.3  in  good  faith  and  seeking  fair  and  reasonable  commercial  terms  and  UCLB’s  right  to  negotiate  with  Autolus  shall
expire [***] following notice of termination.

Upon termination of this Agreement as a whole under Clause 24:

all  Licences  other  than  the  Background  Licence  granted  pursuant  to  Clause  2.1.2  and  the  Manufacturing  Licence  granted

pursuant to Clause 2.3.2 shall automatically terminate;

Autolus shall, at its expense, assign to UCLB any Assigned Patents;
Autolus shall cease to have rights under this Agreement to Exploit any Royalty Products and all of its rights and obligations under

this Agreement concerning Royalty Products shall cease; and,

where termination is effected by UCLB for Autolus’s breach, UCLB shall have the option to negotiate with Autolus to agree terms
for  the  grant  of  an  exclusive  licence  to  those  Autolus  Improvements  Controlled  by  Autolus  and  free  of  any  restriction  or
encumbrance, such licence to be limited to the Exploitation of those Improvements together with the Technology, such option
and negotiation rights to expire [***] following notice of termination served under Clause 24.

e.

The termination of any Licence hereunder shall be without prejudice to the survival of any sub-licence novated to UCLB pursuant to the

conditions under Clause 3.3.3.

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

f.

26.
a.

b.

c.

27.
a.

b.

c.

Termination  or  expiry  of  this  Agreement  for  whatever  reason  shall  not  affect  the  accrued  rights  (including  those  relating  to  any
payments due or payable hereunder) of any Party arising under or out of this Agreement at the date of termination or expiry and all
provisions which are expressed to survive this Agreement or continue after the Term and the provisions of Clauses, 3.3.3, 4.1, 18.9,
22, 23, 25, 30, 31 and 33 shall survive termination or expiry and remain in full force and effect
FORCE MAJEURE

In this Agreement “force majeure” shall mean any cause preventing a Party from performing any or all of its obligations (other than an
obligation to pay sums due) which arises from or is attributable to acts, events, omissions or accidents beyond the reasonable control
of the Party so prevented including to the extent that these are beyond such control industrial disputes, nuclear accident or acts of
God, war or terrorist activity, riot, civil commotion, malicious damage, accident, fire, flood, storm.

If a Party is prevented from performance of any of its obligations under this Agreement by force majeure, that Party shall as soon as
reasonably possible serve notice in writing on the other Parties specifying the nature and extent of the circumstances giving rise to
force  majeure,  and  shall  subject  to  service  of  such  notice  have  no  liability  in  respect  of  any  delay  in  performance  or  any  non-
performance  of  any  such  obligation  save  for  any  payment  obligation  which  shall  continue  in  full  force  and  effect  (and  the  time  for
performance shall be extended accordingly) to the extent that the delay or non-performance is due to force majeure.

If a Party is prevented from performance of substantially all or all of its obligations by force majeure for a continuous period of more
than [***] in total, the other Party may terminate this Agreement forthwith on service of written notice upon the Party so prevented, in
which case the Parties shall not have any liability to the other except that rights and liabilities which accrued prior to such termination
shall continue to subsist.
FURTHER ASSURANCE

During the Term, UCLB shall at its own cost execute all such documents and do or cause to be done all such other things as Autolus
may from time to time require in order to enable and provide Autolus with the benefit of the Licences granted to it hereunder and
otherwise to give full effect to this Agreement.

During  the  Term  UCLB  shall  comply  with  its  obligations  under  Clauses  9.1  to  9.5  so  it  may  facilitate  Autolus  using  its  commercially

reasonable endeavours in accordance with its obligations under Clause 11.

 Without limiting its obligations under Clause 27.1, UCLB shall complete (or procure the completion of) such documents and take such
other steps as shall be necessary or desirable to enable Autolus to be recorded on any registry as the licensee of the Intellectual
Property licensed to it hereunder.

d.

UCLB  shall  procure  the  assistance  of  UCL  and  require  UCL  to  do  or  refrain  from  doing  things  which  would  otherwise  constitute  a

28.
a.

breach of the terms of this Agreement.
PUBLICITY

Upon  execution  of  this  Agreement,  the  Parties  shall  agree  the  content  and  timing  for  a  joint  public  statement  release.  Until  the  [***]
anniversary of the Effective Date, if either Party wishes to make any formal press release regarding the development of any Royalty
Products, the Parties will, acting reasonably and in good faith, agree the terms of the publicity statement. Thereafter should Autolus
wish to include UCL’s or UCLB’s name in a press release or if a press release by Autolus concerns the launch of a Royalty Product,
again the Parties will, acting reasonably and in good faith, agree the terms of the publicity statement. Notwithstanding anything in this
Agreement to the contrary, a Party shall not be prevented from complying with its obligations to make public statements regarding
this  Agreement,  its  subject  matter  or  developments  under  this  Agreement  pursuant  to  the  rules  of  any  stock  market  or  other  laws
applicable to it.

b.

In order to enable UCLB and UCL to monitor the benefit that they are providing, and to enable UCL to demonstrate the impact of its
research activities, to society and the economy, as reasonably requested by UCLB, Autolus shall provide to UCLB non confidential
information on how it has used the Technology and the societal and economic benefits generated therefrom.

c.

Autolus acknowledges that UCLB and UCL shall be entitled to make use of any information received from Autolus (and the information

contained therein) pursuant to

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

29.
a.

b.

Clause  28.2  in  applications  for  research  or  other  granted  related  funding  and  in  submissions  to  Higher  Education  funding  bodies
such as HEFCE and/or HEIF (or any replacements for either of those entities) and like entities, and to use Autolus’s name in their
general publicity materials subject to Autolus’s prior written approval, which approval shall not be unreasonably withheld.
ASSIGNMENT

Save  as  provided  in  Clause  29,  neither  Party  shall  without  the  prior  written  consent  of  the  other  Party  assign  any  of  its  rights  or
obligations under this Agreement, or purport to do any of the same. Any purported assignment in breach of this clause shall confer
no rights on the purported assignee.

Subject to Clause 29.3 and 29.4, Autolus shall be entitled to assign its rights together with its obligations under this Agreement to any
Affiliate of Autolus or to any acquirer of all or substantially all of Autolus’s business provided that such assignee agrees in writing to
be bound by all of the terms and conditions of this Agreement and provided also that the provisions of Clause 3.1 and 3.2 shall apply
with  respect  to  any  proposed  assignment  as  if  it  were  a  sub-licence.  No  assignment  shall  be  valid  or  effective  unless  or  until  the
assignee shall agree, in writing, to be bound by the provisions of this Agreement.

c. Without prejudice to UCLB’s right to terminate the Agreement pursuant to Clause 24.4.2 (where Autolus suffers an Insolvency Event),
Autolus may grant security over or assign by way of security any of its rights and obligations under this Agreement provided that any
such assignment shall comply with the provisions of Clause 29.2.

d.

e.

f.

30.

Autolus shall not be entitled to assign the Agreement during the grace periods (20 or 30 days) referred to in the definition of Insolvency

Event and any assignment of the Agreement during this period shall not be valid or effective.

UCLB  shall  not  assign  any  of  the  Technology  to  any  Third  Party  nor  grant  any  mortgage,  charge  or  other  encumbrance  over  the

Technology.

In the event of termination of the agreement pursuant to which UCLB is granted (i) a sub-licensable licence with respect to the [***] in
relation  to  the  TRBC1/2  Product  and  the  CRUK  Study  Results;  (ii)  a  right  to  grant  an  option  to  a  licence  for  certain  TRBC1/2
improvements made by a student funded by CRUK; and (iii) a right to grant an option to a licence to a [***] project funded by [***],
provided that Autolus is not in breach of its obligations under this Agreement and provided Autolus agrees to pay CRUK the sums
equivalent to those sums otherwise payable by UCLB to CRUK for the aforementioned licences and rights, Autolus shall receive a
direct  licence  from  CRUK  with  respect  to  the  [***]  and  the  CRUK  Study  Results  and  the  options  or  licences  with  respect  to  the
TRBC1/2 improvements and the [***] project (“Direct Licence”). The scope of such Direct Licence shall be the same as the scope of
the corresponding licences and rights under this Agreement and shall not impose more onerous obligations on Autolus. On receipt of
the Direct Licence from CRUK, Autolus shall be entitled to deduct an amount equivalent to any sums Autolus pays CRUK under the
Direct Licence from any amount Autolus is due to pay UCLB with respect to the GD2 Product, the TRBC1/2 Product (including the
[***]) and the BCMA Product. Nothing in this Clause shall limit or exclude or operate to waive any liability of UCLB should UCLB's
licences from CRUK terminate or expire.
NOTICES
All notices required to be served by the Parties to this Agreement under the terms hereof shall be sufficiently served if dispatched by
first  class  post  or  commercial  courier  to  the  addresses  of  each  of  the  Parties  set  out  below.  All  such  notices  shall  be  deemed
received within five (5) days after such dispatch.

If to:

Autolus

UCLB

Forest House, 58 Wood Lane, London, W12 7RZ
Attn. General Counsel

The Network Building, 97 Tottenham Court Road, London, W1T 4TP

and  any  modification  or  amendment  to  such  address  must  itself  be  notified  in  writing  to  the  other  Parties  in  accordance  with  the
terms of this Clause.
MISCELLANEOUS PROVISIONS

Entire Agreement

31.
a.

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

i..

ii..

iii..

iv..
b.

i..

ii..

This  Agreement  and  any  variations,  amendments  or  other  modifications  in  relation  to  this  Agreement  constitutes  the  entire
agreement  between  the  Parties  relating  to  its  subject  matter  and  save  for  the  CDA  supersedes  all  prior  agreements  and
understandings, both written and oral, between the Parties with respect to the Programs, the Program IP, the Manufacturing
Know-How and the UCL Background IP.

Each  Party  acknowledges  that  in  entering  into  this  Agreement  it  does  not  do  so  on  the  basis  of  and  does  not  rely  on  any
representation,  warranty,  or  other  provision  except  as  expressly  provided  in  this  Agreement  and  all  conditions,  warranties
and other terms implied by statute or common law are hereby excluded to the fullest extent permitted by law provided that
nothing in this Clause should be construed as limiting or excluding liability for fraud.

Except as otherwise provided in this Agreement, the only remedy available to a Party for breach of this Agreement shall be for
breach  of  contract  under  the  terms  of  this  Agreement  and  no  Party  shall  be  liable  in  tort  or  otherwise  arising  from  such
breach.  The  rights  and  remedies  provided  by  this  Agreement  are  cumulative  and  (except  as  otherwise  provided  in  this
Agreement) are not exclusive of any rights or remedies provided by law.

Nothing in this Clause 31 shall limit or exclude any liability for fraud or fraudulent misrepresentation.

Amendment and Waiver

Any  agreement  to  amend,  vary  or  modify  the  terms  of  this  Agreement  in  any  manner  shall  be  valid  only  if  the  amendment,
variation or modification is effected in writing and signed by duly authorised representatives of each of the Parties hereto.
No  delay  by  any  Party  in  enforcing  any  of  the  provisions  of  this  Agreement  shall  be  deemed  a  waiver  of  that  Party’s  right

subsequently to enforce such provision.

c.

Severability

i..

ii..

iii..

If  any  term  or  provision  of  any  part  thereof  contained  herein  shall  be  declared  or  become  unenforceable  invalid  or  illegal  in  any
respect under the law of any relevant jurisdiction:

such term or provision or part thereof shall be deemed to have been severed from the remaining terms of this Agreement and the
terms and conditions hereof shall remain in full force and effect as if this Agreement had been executed without the offending
provision appearing herein; and

the Parties shall endeavour to agree an amendment which to the fullest extent possible will give lawful effect to their intentions as

expressed in any term or provision severed under Clause 31.3.1;

If any restriction in this Agreement is held by any court or other competent authority to be invalid or unenforceable, then the Party
against whom such restriction was intended to apply agrees to be bound by a restriction the same as the terms of the most
onerous restriction which the court or other competent authority would have allowed in place of the affected restriction.

d.

Status of the Parties

Except as otherwise provided, each Party shall bear its own costs and expenses in connection with the preparation, negotiation,

execution and performance of this Agreement and the documents referred to in it.

No Party is authorised to act as the agent of the other for any purpose whatsoever and no Party shall on behalf of the other(s)
enter  into,  or  make,  or  purport  to  enter  into  or  make  or  represent  that  it  has  any  authority  to  enter  into  or  make  any
representation or warranty.

Nothing in this Agreement shall be deemed to constitute a partnership or joint venture company between any or all of the Parties
and  none  of  the  Parties  shall  do  or  suffer  to  be  done  anything  whereby  it  might  be  represented  as  a  partner  of  the  other
Parties.

Each Party shall be directly responsible to the other Parties for all actions or omissions of its respective Affiliates, agents and sub-
contractors  relating  to  the  subject  matter  of  this  Agreement  and  shall  be  responsible  for  and  liable  for  the  fulfilment  and
observance by itself and its Affiliates, agents and sub-contractors of the applicable obligations and restrictions on it and its
Affiliates, agents and sub-

i..

ii..

iii..

iv..

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

v..

32.

33.
a.

b.

c.

contractors hereunder (or to be imposed on them pursuant to the terms hereunder).

A person who is not a Party to this Agreement has no right under the Contracts (Rights of Third Parties) Act 1999 to enforce any
term of this Agreement but this does not affect any right or remedy of a Third Party which exists or is available apart from that
Act. Notwithstanding the above, (i) an Indemnified Party which is not a Party to this Agreement may enforce the provisions of
Clause 23.1 where it has the benefit of the indemnity provided in Clause 23.1; and (ii) the Wellcome Trust, the Chairman of
the  Wellcome  Trust  and  the  Appointed  Expert  may  enforce  the  provisions  of  Clause  3.2.3.  The  rights  of  the  Parties  to
terminate, rescind or agree any variation, waiver or settlement under this Agreement are not subject to the consent of any
person  that  is  not  a  Party  to  this  Agreement,  including  any  Indemnified  Party  or  the  Wellcome  Trust,  the  Chairman  of  the
Wellcome Trust or the Appointed Expert, provided that the Parties may not vary or waive the rights of the Wellcome Trust,
the Chairman of the Wellcome Trust or the Appointed Expert under Clause 3.2.3 without their prior written consent.

COUNTERPARTS
This Agreement may be executed in any number of counterparts and by the Parties to it on separate counterparts, each of which
shall be an original but all of which together shall constitute one and the same instrument, and shall not be effective until each of the
Parties has executed at least one counterpart.
DISPUTE RESOLUTION, GOVERNING LAW AND JURISDICTION

All  controversies  or  claims  of  whatever  nature  arising  out  of  or  relating  in  any  manner  whatsoever  to  this  Agreement  or  any  of  the
documents  referred  to  in  this  Agreement,  including  but  not  limited  to  a  controversy  or  claim  involving  the  validity,  enforceability,
interpretation  or  construction  of  this  Agreement  or  any  of  the  documents  referred  to  in  this  Agreement,  shall  be  governed  by  and
construed in all respects in accordance with the laws of England.

In  the  event  of  any  dispute,  difference  or  question  arising  in  connection  with  this  Agreement,  either  Party  shall  be  entitled  but  not
obliged to escalate the matter to the Parties’ Executive Officers by serving a written notice on the other Party’s Executive Officer, in
which  case  the  Parties’  Executive  Officers  shall  make  themselves  available  to  discuss  the  dispute,  difference  or  question,  as  the
case may be (the “Unresolved Matter”),  and  use  good  faith  efforts  to  resolve  such  Unresolved  Matter  within  the  thirty  (30)  days
following the delivery of such notice.

If the Parties agree to submit, they shall submit to non-binding mediation by a neutral mediator (with the understanding that the role of
the mediator shall not be to render a decision but to assist the Parties in reaching a mutually acceptable resolution) who shall be
accredited  by  the  Centre  of  Dispute  Resolution  (“CEDR”)  or  otherwise  appropriately  qualified,  and  the  mediation  regarding  the
Unresolved  Matter  shall  take  place  in  London  UK  (or  such  other  location  as  may  be  mutually  agreed  upon  by  the  Parties).  The
mediator shall be chosen by agreement of the Parties, or if they are unable to agree on a mediator within fourteen (14) days of a
request from one Party to the other or if the agreed mediator is unable or unwilling to act, either Party may apply to CEDR to appoint
a mediator.

d. Within fourteen (14) days of the mediator being appointed, the Parties shall seek guidance from the mediator on a programme for the
exchange of information and the structure to be adopted for negotiations. Either Party may request a preliminary meeting with the
mediator for this purpose which shall be attended by both Parties.

Unless otherwise agreed, all negotiations concerning the dispute shall be conducted in confidence and shall be without prejudice to the
rights of the parties in any future proceedings. The mediation is non-binding and Parties shall not be obliged to accept or follow any
recommendation of the mediator.

If the Parties reach agreement on the resolution of the dispute, the agreement shall be reduced to writing and shall be binding on the

Parties once it is signed by their duly authorised representatives.

If the Unresolved Matter is not resolved by mediation within sixty (60) days of appointment of the mediator, either Party may, subject to

Clause 33.9, make any claim or application before the court as it sees fit.

e.

f.

g.

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

h.

i.

Notwithstanding the provisions of Clause 33.2 or of Clause 33.3, subject to Clause 33.9, each Party shall be free to seek temporary

injunctive relief in court as the situation may necessitate based upon any irreparable harm which may ensue.

Each Party acknowledges and agrees that the courts of England shall have exclusive jurisdiction to resolve any controversy or claim of
whatsoever  nature  arising  out  of  or  relating  in  any  manner  to  this  Agreement,  any  terms  of  this  Agreement,  or  any  breach  of  this
Agreement or any such terms.

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

Schedule 1.

The Programs

[***]

[***]

[***]

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[***]

[***]

[***]

[***]

[***]

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[***]

[***]

[***]

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

Schedule 2.

Program IP

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

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[***]

[***]

[***]

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[***]

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

[***]

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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UCL Background[***]

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    96

a.

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    97

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    98

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Schedule 3.

Manufacturing Know-How

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Disclosure Process[***]

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Part A: Existing Licences[***]

Schedule 4.

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[***][***][***][***][***][***][***]

(L)

: Commercial Agreements

    101

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permitted Studies[***]

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

Schedule 5.

Part A:Net Sales Definition

◦

◦

◦

◦

◦

Net Sales shall mean, subject to the remainder of Part A of this Schedule, the gross sum received by Autolus or its Sub-Licensees
(excluding  Net  Receipts)  or  Assignee  Entities  from  the  supply  of  a  Royalty  Product  by  Autolus  or  its  Sub-Licensees  or  Assignee
Entities to a Third Party in a bona fide, arm’s length transaction, less the following:
i.
ii.

normal and customary trade and quantity discounts actually granted;
amounts repaid or credited by reasons of defects, rejection, recalls, returns, rebates and allowances of goods or because of
retroactive price reductions;
chargebacks and other amounts paid on sale of the Royalty Product;
amounts  payable  resulting  from  government/regulator-mandated  rebate  programs  including  pursuant  to  indigent  patient
programs and patient discount programs;
tariffs, duties, excise, sales, value-added and other taxes, identified in the relevant invoice;
retroactive price reductions that are actually allowed or granted;
cash discounts or credits for timely payment;
delayed ship order credits; and,
all freight, postage, storage, shipping and insurance, identified in the relevant invoice.

iii.
iv.

v.
vi.
vii.
viii.
ix.

Net Sales will not include:

x.

xi.
xii.

any  transfers  between  Autolus,  its  Affiliates  and  any  Sub-Licensees  or  Assignee  Entities,  or  for  the  supply  of  any  Royalty
Products for clinical trial activities, research purposes, charitable donations or compassionate use;
for the purposes of calculating sales thresholds or triggering any Milestone, any Complementary Diagnostic Products;
any sums for any products, services or processes that are not Royalty Products.

Where  Autolus  or  any  Sub-Licensee  or  Assignee  Entity  sells  any  Royalty  Products  other  than  on  normal  arms-length  commercial
terms  exclusively  for  money,  the  Net  Sales  of  the  Royalty  Product  supplied  shall  be  determined  as  the  fair  market  value  of  such
Royalty Product.
Where Autolus or any Sub-Licensee or Assignee Entity sells a product that consists of a Royalty Product in combination with, co-
formulated with, or co-packaged with a product that contains one or more non-cellular therapeutically active agents(s) (which are not
expressed  or  produced  by  the  Royalty  Product  itself)  (“Combination Product”),  then  the  Net  Sales  of  the  Royalty  Product  in  the
country of sale shall be calculated as follows:

xiii.

xiv.

if  the  Royalty  Product(s)  (on  the  one  hand)  and  the  non-cellular  therapy  product(s)  (together  or  separately)  each  are  sold
separately in commercially reasonable quantities in such country, Net Sales will be calculated by multiplying the total sales of
the Combination Product by the fraction A/(A+B), where A is the average gross selling price in such country of the Royalty
Product(s)  contained  in  the  Combination  Product  sold  separately  during  the  calendar  quarter  in  the  same  formulation  and
dosage, and B is the sum of the gross selling price in such country of the non-cellular therapeutic product(s) contained in the
Combination Product sold separately during the calendar quarter in the same formulation and dosage; and
if the Royalty Product(s) (on the one hand) and/or the non-cellular therapeutic product(s) (together or separately) are not sold
separately in commercially relevant quantities in a country during a particular payment period, or if they are sold separately
but  the  average  gross  selling  price  of  either  the  Royalty  Product(s)  (on  the  one  hand)  or  the  non-cellular  therapeutic
product(s) (together or separately) cannot be determined, in such country, then the Parties will meet and negotiate in good
faith an appropriate mechanism for determining the royalty payable on such Combination Product. If the Parties are unable
to  agree  an  appropriate  mechanism,  the  Parties  shall  refer  to  an  expert  for  determination  of  the  agreed  mechanism  by
following the provisions of Part C of this Schedule.

Where Autolus or any Sub-Licensee or Assignee Entity sells a Royalty Product as part of a package that includes services that relate
to the manipulation of, administration of, or

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delivery of the applicable Royalty Product then the Net Sales of the Royalty Product as part of the package shall be calculated as a
proportion of the package price fairly attributed to the Royalty Product alone.
Where Autolus or any Sub-Licensee or Assignee Entity sells a Royalty Product as a Combination Product, and such Combination
Product  is  sold  as  part  of  a  package  that  includes  services  that  relate  to  the  manipulation  of,  administration  of,  or  delivery  of  the
applicable Combination Product, then the Net Sales of the Royalty Product as part of the Combination Product and as part of the
package shall be calculated in the following order (i) firstly the value attributable to the Combination Product shall be calculated as a
proportion of the package price fairly attributed to the Combination Product alone; and (ii) secondly the value of the Royalty Product
as part of the Combination Product shall be determined in accordance with paragraph 4.
Where  Autolus  or  any  Sub-Licensee  or  Assignee  Entity  sells  a  product  that  is  not  a  Combination  Product  but  which  consists  of  a
Royalty  Product  in  combination  with,  co-formulated  with,  or  co-packaged  with  any  other  product  or  components  (not  covered  by
paragraphs  4,  5  or  6  above),  then  no  apportionment  of  the  Net  Sales  between  such  Royalty  Product  and  the  other  product  or
components  shall  apply  save  that  where  a  Royalty  Product  is  sold  in  combination  with,  co-formulated  with,  or  co-packaged  with
another  cellular  therapeutically  active  agent  which  does  not  use  the  Technology,  for  the  purpose  of  calculating  the  Net  Sales  with
respect to the Royalty Product, the Parties shall in good faith discuss and agree the apportionment.
Any dispute as to the determination of a fair market value that cannot be resolved through discussion between the Parties shall be
referred to an expert for resolution in accordance with the provisions of part C of this Schedule 8.

◦

◦

◦

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Schedule 6.

: Net Receipts Definition

◦

◦

Net Receipts shall mean, subject to the remainder of Part B of this Schedule, the gross sums received by Autolus pursuant to (i) a
Sublicence in so far as they are attributable to the Technology so licensed, or (ii) a Patent Sale in so far as they are attributable to the
Assigned Patents so assigned less all tariffs, duties, taxes, excise, sales and value added taxes.
The calculation of Net Receipts shall be subject to the following:
i.

Net  Receipts  will  include  (i)  payments  for  the  grant  of  a  sublicence,  or  in  the  case  of  the  Assigned  Patent,  a  licence  or
assignment of such Assigned Patent, including up front signing fees, stage payments and milestone payments (ii) payments
for options for a sublicence, or in the case of the Assigned Patent, an option for a licence or an assignment of such Assigned
Patent  or  for  the  exercise  of  such  options;  and  (iii)  in  return  for  the  grant  of  a  sublicence,  or  in  the  case  of  an  Assigned
Patent, the grant of a licence or an assignment of such Assigned Patent, the cash paid by a sublicensee (or its Affiliate) or
Assignee Entity (or its Affiliate) for purchasing shares, options or other securities in Autolus but only to the extent that the
price paid for such shares, options or other securities exceeds the fair market value calculated at the time of subscription; If
Autolus grants a sub-licence, or in the case of an Assigned Patent, a licence or an assignment of such Assigned Patent in
return for non-cash consideration, Autolus shall agree with UCLB what (if any) cash value will be attributed to such non-cash
consideration from which cash value the Net Receipts will be calculated and when any such Net Receipts shall be paid, such
agreement to be reached within [***] of closing the deal and provided that either party shall be entitled to refer the matter to
the Expert for determination in the event that agreement cannot be reached by the parties.
Net receipts will exclude (i) any damages or account of profits due, paid or recoverable; (ii) any settlement fees or costs; (iii)
payments  under  or  by  virtue  of  any  indemnity;  (iii)  any  royalty  payments,  profit  share  or  other  compensation  calculated,
directly or indirectly by reference to volume or units of Royalty Products sold, provided UCLB received Royalties in respect of
the  same;  (iv)  any  sums  received  by  way  of  transfer  pricing;  (v)  any  sums  received  or  bona  fide  sums  paid  for  Royalty
Products supplied by Autolus or its Affiliates or Assignee Entities, provided UCLB received Royalties in respect of the same;
(vi) litigation costs and fees, or other payments received in respect of the enforcement or defence of any Intellectual Property
rights; (vii) any sums received for any products, services or processes that are not Royalty Products; and (viii) any service
fees, FTE payments or other payments received to cover a contractual expense.

ii.

    105

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

Schedule 7.

: Expert Procedure
Any dispute arising out of or in connection with Clauses 15.1.3, 15.2.1, 16.7 of this Agreement and paragraphs 3, 4, 5, 6 or 7 of Part
A of Schedule 8 or paragraph 2a of Part B of Schedule 8 to this Agreement and/or its performance shall be referred to an expert by
either Party serving on the other Party notice (“Referral Notice”) that it wishes to refer the dispute to an expert. For the avoidance of
doubt, no reference shall be made to the expert as to what product or therapy constitutes a Royalty Product and any dispute in that
regard shall be determined in accordance with Clause 33.3 and/or Clause 33.9 of the Agreement. If either Party challenges whether
a  product  or  therapy  constitutes  a  Combination  Product  that  dispute  shall  be  determined  in  accordance  with  Clause  33.3  and/or
Clause 33.9 of the Agreement.
The dispute shall be determined by a single independent impartial expert who shall be agreed between the Parties or, in the absence
of agreement between the parties within [***] of the service of a Referral Notice, be appointed by the then President of the Institute of
Chartered Accountants or any successor organisatcion thereto.
The seat of the dispute resolution shall be the normal place of residence of the expert.
The language of the dispute resolution shall be English.
The expert shall not have power to alter, amend or add to the provisions of this Agreement, except that the expert shall have the
power to decide all procedural matters relating to the dispute, and may call for a one day hearing if desirable and appropriate.
The expert shall have the power to request copies of any documents in the possession and/or control of the parties which may be
relevant to the dispute. The parties shall forthwith provide to the expert and the other party copies of any documents so requested by
the expert.
The expert shall decide the dispute as an expert and not as an arbitrator.
The  decision  of  the  expert  shall  be  final  and  binding  upon  both  parties  except  in  the  case  of  manifest  error.  The  parties  hereby
exclude any rights of application or appeal to any court, to the extent that they may validly so agree, and in particular in connection
with any question of law arising in the course of the reference out of the award.
The expert shall determine the proportions in which the parties shall pay the costs of the expert's procedure. The expert shall have
the authority to order that all or a part of the legal or other costs of a party shall be paid by the other party. UCLB’s liability in this
regard shall not be subject to the cap on liability under Clause 22.
All documents and information disclosed in the course of the expert proceedings and the decision and award of the expert shall be
kept  strictly  confidential  by  the  recipient  and  shall  not  be  used  by  the  recipient  for  any  purpose  except  for  the  purposes  of  the
proceedings and/or the enforcement of the expert’s decision and award.

◦

◦

◦
◦
◦

◦

◦
◦

◦

◦

    106

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

Schedule 8.

Warranties and Covenants

Part A. : Mutual Warranties & Representations

In respect of each Party making the warranty and representation:
it is a company duly organised, validly existing, and in good standing under the laws of England;
it has full corporate power and authority to execute, deliver, and perform this Agreement and has taken all corporate action required
by  law  and  its  organisational  documents  to  authorise  the  execution  and  delivery  of  this  Agreement  and  the  consummation  of  the
transactions contemplated by this Agreement;
this  Agreement  constitutes  a  valid  and  binding  agreement  enforceable  against  it  in  accordance  with  its  terms  (except  as  the
enforceability thereof may be limited by bankruptcy, insolvency or similar laws affecting creditors’ rights generally and laws restricting
the  availability  of  equitable  remedies  and  may  be  subject  to  general  principles  of  equity  whether  or  not  such  enforceability  is
considered in a proceeding at law or in equity);
the  execution  and  delivery  of  this  Agreement  and  all  other  instruments  and  documents  required  to  be  executed  pursuant  to  this
Agreement and the consummation of the transactions contemplated hereby and thereby do not and shall not (i) conflict with or result
in  a  breach  of  any  provision  of  its  organisational  documents,  (ii)  result  in  a  breach  of  any  agreement  to  which  it  is  a  party;  or  (iii)
violate any law.

Part B. : UCLB Warranties & Covenants at the Effective Date

UCLB warrants and represents to Autolus that as at the Effective Date:
Material Information
the information set out in the Schedules at the Effective Date is accurate and so far as UCLB is aware is materially complete;
all the information which is contained in the Disclosure Letter dated as of the Effective Date and the documents (if any) annexed to it
is complete and accurate and not misleading;
each statement of opinion or belief which is attributed in the Disclosure Letter dated as of the Effective Date is honestly held by the
members of the UCLB senior management team;
each document if annexed to the Disclosure Letter dated as of the Effective Date is a complete and accurate copy of the original, and
no such document has been amended (orally or in writing) or superseded;
Intellectual Property
it is the sole and exclusive owner, free of all encumbrances, of all right, title and interest in and to the Original Licensed Patents;
in respect of the Original Program IP, Manufacturing Know-How and UCL Background IP it is either:
i.
ii.

the sole and exclusive owner, free of all encumbrances, of all right, title and interest in and to such Intellectual Property; or
a  licensee  with  the  right  to  grant  the  licences  granted  herein  on  the  terms  granted  herein  in  respect  of  such  Intellectual
Property;

it has not granted, or agreed to grant, any licences or entered into any agreements which may adversely affect or conflict with this
Agreement and/or with any of the Licences granted hereunder and/or options to licences granted hereunder as at the Effective Date;
it has not granted, or agreed to grant, any assurance or waiver not to enforce in respect of any of the Intellectual Property licensed
hereunder as at the Effective Date in so far as such consents, assurances or waivers would enable the Third Party to develop, free of
infringement, any product or therapy that is Covered by or has been developed using or uses any of the Intellectual Property licensed
hereunder as at the Effective Date;
there is no other Patent Right (beyond the Original Licensed Patents) owned by UCLB as at the Effective Date that is required for the
use and practise of any of the Original

i.

a.
b.

c.

d.

i.

e.
f.

g.

h.

i.
j.

k.

l.

m.

    107

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

Program IP, UCL Background IP, having regard to UCLB’s understanding of Autolus’s development plan for the period of [***] after
the Effective Date;
as  at  the  Effective  Date  no  invention  disclosure  forms  have  been  logged  in  UCLB’s  database  and  categorised  as  “Biopharm”  and
with  the  status  “being  assessed”,  disclosing  patentable  inventions  in  respect  of  which  (if  patent  applications  were  filed  for  such
inventions within [***] of the Effective Date) to the best of UCLB’s knowledge and belief a licence would be required for the use and
practise of any of the Original Program IP, UCL Background IP, having regard to UCLB’s understanding of Autolus’s development
plan for the period of [***] after the Effective Date;
it has provided Autolus with details of Third Party Patent Rights of which it is aware, and which to the best of its knowledge and belief
not  having  (i)  conducted  any  professional  freedom  to  operate  searches  or  (ii)  sought  advice  from  a  qualified  patent  attorney  or
solicitor,  may  be  relevant  to  the  development  of  any  Original  Royalty  Product  on  the  basis  of  UCLB’s  understanding  of  Autolus’s
development plan for the period of [***] after the Effective Date;
it is not aware that the disclosure to Autolus of Know-How forming part of the Original Program IP, Manufacturing Know-How or UCL
Background IP will amount to a breach of any obligation of confidentiality owed by UCL, MP or UCLB to any Third Party;
it is not aware of any material breach by UCLB, UCL or MP of any Third Party contracts set out in Schedule 6;
it has not received any negative opinion from any patent office as to the validity of the Original Licensed Patents;
there is no on-going litigation to which UCL or UCLB is a party concerning any of the Original Program IP, Manufacturing Know-How
or UCL Background IP;
Clinical Studies
there has been no clinical use of any of the Materials.

n.

o.

p.

q.
r.
s.

t.

Part C. : UCLB Warranties & Covenants at the Amendment Date

UCLB warrants and represents to Autolus that as at the Amendment Date:
Material Information
the information included as at the Amendment Date to the Schedules is accurate and so far as UCLB is aware is materially complete;
all the information which is contained in the Disclosure Letter dated as of the Amendment Date and the documents (if any) annexed
to it is complete and accurate and not misleading;
each statement of opinion or belief which is attributed in the Disclosure Letter dated as of the Amendment Date is honestly held by
the members of the UCLB senior management team;
each document if annexed to the Disclosure Letter dated as of the Amendment Date is a complete and accurate copy of the original,
and no such document has been amended (orally or in writing) or superseded;
Intellectual Property
it is the sole and exclusive owner, free of all encumbrances, of all right, title and interest in and to the Additional Licensed Patents;
it  has  not  granted,  or  agreed  to  grant,  any  licences  or  entered  into  any  agreements  in  respect  of  the  Additional  Licensed  Patents
which may adversely affect or conflict with this Agreement and/or with any of the Licences granted hereunder;
it has not granted, or agreed to grant, any assurance or waiver not to enforce in respect of any of the Additional Licensed Patents in
so far as such consents, assurances or waivers would enable the Third Party to develop, free of infringement, any product or therapy
that is Covered by any of the Additional Licensed Patents licensed hereunder;
there is no other Patent Right (beyond the Licensed Patents) owned by UCLB as at the Effective Date that is required for the use and
practise  of  any  of  the  Additional  Licensed  Patents  having  regard  to  UCLB’s  understanding  of  Autolus’s  development  plan  for  the
period of [***] after the Effective Date;
as  at  the  Effective  Date  no  invention  disclosure  forms  have  been  logged  in  UCLB’s  database  and  categorised  as  “Biopharm”  and
with the status “being assessed”, disclosing patentable inventions in respect of which (if patent applications were filed for such

i.

1.
2.

3.

4.

5.
6.

7.

8.

9.

    108

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

inventions within [***] of the Effective Date) to the best of UCLB’s knowledge and belief a licence would be required for the use and
practise  of  any  of  the  Additional  Licensed  Patents,  having  regard  to  UCLB’s  understanding  of  Autolus’s  development  plan  for  the
period of [***] after the Effective Date;
it has provided Autolus with details of Third Party Patent Rights of which it is aware, and which to the best of its knowledge and belief
not  having  (i)  conducted  any  professional  freedom  to  operate  searches  or  (ii)  sought  advice  from  a  qualified  patent  attorney  or
solicitor,  may  be  relevant  to  the  development  of  any  Additional  Royalty  Product  (except  for  a  TRBC2  Dx  Product)  on  the  basis  of
UCLB’s understanding of Autolus’s development plan for the period of [***] after the Effective Date;
it has not received any negative opinion from any patent office as to the validity of the Additional Licensed Patents;
there is no on-going litigation to which UCL or UCLB is a party concerning any of the Additional Licensed Patents.

10.

11.
12.

Part D. UCLB Warranties & Covenants at the Second Amendment Date

UCLB warrants and represents to Autolus that as at the Second Amendment Date:
Material Information
except  as  disclosed  in  a  Disclosure  Letter  dated  as  of  the  Second  Amendment  Date,  the  information  included  as  at  the  Second
Amendment Date to the Schedules is accurate and so far as UCLB is aware is materially complete;
all the information which is contained in the Disclosure Letter dated as of the Second Amendment Date and the documents (if any)
annexed to it is complete and accurate and not misleading;
each statement of opinion or belief which is attributed in the Disclosure Letter dated as of the Second Amendment Date is honestly
held by the members of the UCLB senior management team;
each document if annexed to the Disclosure Letter dated as of the Second Amendment Date is a complete and accurate copy of the
original, and no such document has been amended (orally or in writing) or superseded;
Intellectual Property
it is the sole and exclusive owner, free of all encumbrances, of all right, title and interest in and to the CAT19 Program IP;
it has not granted, or agreed to grant, any licences or entered into any agreements in respect of the CAT19 Program IP which may
adversely affect or conflict with this Agreement and/or with any of the Licences granted hereunder;
it has not granted, or agreed to grant, any assurance or waiver not to enforce in respect of any of the CAT19 Program IP in so far as
such consents, assurances or waivers would enable the Third Party to develop, free of infringement, any product or therapy that is
Covered by any of the CAT19 Program IP licensed hereunder;
there is no other Patent Right (beyond the Licensed Patents) owned by UCLB as at the Effective Date that is required for the use and
practise of any of the CAT19 Program IP having regard to UCLB’s understanding of Autolus’s development plan for the period of [***]
after the Effective Date;
as  at  the  Effective  Date  no  invention  disclosure  forms  have  been  logged  in  UCLB’s  database  and  categorised  as  “Biopharm”  and
with  the  status  “being  assessed”,  disclosing  patentable  inventions  in  respect  of  which  (if  patent  applications  were  filed  for  such
inventions within [***] of the Effective Date) to the best of UCLB’s knowledge and belief a licence would be required for the use and
practise of any of the CAT19 Program IP, having regard to UCLB’s understanding of Autolus’s development plan for the period of [***]
after the Effective Date;
it has provided Autolus with details of Third Party Patent Rights of which it is aware, and which to the best of its knowledge and belief
not  having  (i)  conducted  any  professional  freedom  to  operate  searches  or  (ii)  sought  advice  from  a  qualified  patent  attorney  or
solicitor, may be relevant to the development of any CAT19 Product on the basis of UCLB’s understanding of Autolus’s development
plan for the period of [***] after the Effective Date;

i.

1.

2.

3.

4.

5.
6.

7.

8.

9.

10.

    109

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

11.
12.

it has not received any negative opinion from any patent office as to the validity of the CAT19 Patent Rights;
there is no on-going litigation to which UCL or UCLB is a party concerning any of the CAT19 Program IP.

Part E. UCLB Warranties & Covenants at the Third Amendment Date

UCLB warrants and represents to Autolus that as at the Third Amendment Date:
Material Information
the information included as at the Third Amendment Date to Schedule 1 and Schedule 2 under the titles TRBC2 Dx Program and
CAROUSEL Study is accurate and so far as UCLB is aware is materially complete;
Intellectual Property
Immediately prior to the assignment of the Assigned Patents pursuant to the Patent Assignment, with the exception of the TRBC2 Dx
Patent Rights which are jointly owned by UCLB and Autolus, it is the sole and exclusive owner, free of all encumbrances, of all right,
title and interest in and to the Assigned Patents;
it has not granted, or agreed to grant, any licences or entered into any agreements in respect of the TRBC2 Dx Patent Rights which
may adversely affect or conflict with this Agreement and/or with any of the Licences granted hereunder;
it has not granted, or agreed to grant, any assurance or waiver not to enforce in respect of any of the TRBC2 Dx Patent Rights in so
far as such consents, assurances or waivers would enable the Third Party to develop, free of infringement, any product or therapy
that is Covered by any of the TRBC2 Dx Patent Rights;
there is no other Patent Right (beyond the Retained Patents) owned by UCLB as at the Third Amendment Date that is required for
the  use  and  practise  of  the  TRBC2  Dx  Program  IP  or  the  CAROUSEL  Program  IP,  having  regard  to  UCLB’s  understanding  of
Autolus’s development plan for the period of [***] after the Effective Date;
as  at  the  Third  Amendment  Date  no  invention  disclosure  forms  have  been  logged  in  UCLB’s  database  and  categorised  as
“Biopharm”  and  with  the  status  “being  assessed”,  disclosing  patentable  inventions  in  respect  of  which  (if  patent  applications  were
filed for such inventions within [***] of the Third Amendment Date) to the best of UCLB’s knowledge and belief a licence would be
required  for  the  use  and  practise  of  the  TRBC2  Dx  Program  IP  or  the  CAROUSEL  Program  IP,  having  regard  to  UCLB’s
understanding of Autolus’s development plan for the period of [***] after the Effective Date;
it has provided Autolus with details of Third Party Patent Rights of which it is aware, and which to the best of its knowledge and belief
not  having  (i)  conducted  any  professional  freedom  to  operate  searches  or  (ii)  sought  advice  from  a  qualified  patent  attorney  or
solicitor,  may  be  relevant  to  the  development  of  the  TRBC2  Dx  Product  or  the  CAT19  CNS  Product  on  the  basis  of  UCLB’s
understanding of Autolus’s development plan for the period of [***] after the Effective Date;
there is no on-going litigation to which UCL or UCLB is a party concerning any of the TRBC2 Dx Patent Rights.

i.

1.

2.

3.

4.

5.

6.

7.

8.

    110

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

1.

Royalty and Net Receipts Statements

1.

In respect of each country where Royalty Products were supplied during that Quarter:

a.

the  Net  Sales  of  each  type  of  Royalty  Product  supplied  expressed  both  in  local  currency  and  in  British  pounds  sterling
together with conversion rates used;
the royalty rate applicable to each type of Royalty Product supplied in that country;
the calculation of the royalties payable in respect of each type of Royalty Product; and
the total amount of royalties payable in respect of that country;

b.
c.
d.
For the world as a whole:

a.
b.
c.
In respect of any Royalty Products supplied to which the provisions of paragraph 3 of Part A of Schedule 8 are applicable:

the total amount of royalties payable under Clause 14.1;
the amount of any reduction or deduction made pursuant to Clauses 14.4 to 14.10, inclusive; and
the amount of any withholding tax deducted pursuant to Clause 16.5.

2.

3.

a.
b.

the amount of each type of Royalty Product supplied; and
the actual price at which the Royalty Products were supplied and the nature and value of any other consideration provided
for the Royalty Products.

4.

In respect of any Royalty Products supplied to which the provisions of paragraphs 4, 5, 6 or 7 of Part A of Schedule 8 are applicable:

a.
b.

the amount and description of any Combination Product or any packages of products or services supplied; and
the  actual  price  at  which  the  package  of  any  Combination  Product  or  any  products  or  services  were  supplied  and  the
proportion of the sales price attributed to the Royalty Product in the relevant supply contract.

5.

In respect of any Net Receipts, the calculation of sums payable in accordance with Clause 15 and Part B of Schedule 8.

    111

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

6.

CARPALL and ALLCAR19 Patient Clinical Data generated before the Second Amendment Date:

Release of Patient Clinical Data

1. CARPALL and ALLCAR19 Patient Clinical Data shall be released by sponsor once the sponsor has received confirmation from

the trials sites that [***].

CARPALL and ALLCAR19 Patient Clinical Data generated after the Second Amendment Date:

2. Trial  management  reports  produced  for  the  benefit  of  the  trial  management  groups  for  the  CARPALL  Study  and  the
ALLCAR19  Study  shall  be  released  by  the  sponsor  to  Autolus  at  the  same  time  as  their  formal  release  to  relevant  trial
management  group.  For  information  only,  at  the  Second  Amendment  Date  such  reports  are  produced  at  a  frequency  of
approximately [***]. These trial management reports shall include a final report at the end of [***] of each study accompanied
by  release  of  all  supporting  clinical  data.  The  next  trial  management  group  data  releases  are  scheduled  for  the  CARPALL
Study on [***], and for the ALLCAR19 Study [***]. UCLB accepts no liability if the frequency of such reporting shall change for
any reason and shall not be obliged to make available such reports before they are received by the relevant trial management
group.

3. Raw  data  comprising  the  CARPALL  and  ALLCAR19  Patient  Clinical  Data  shall  be  released  by  the  sponsor  to  Autolus  (i)
following  the  treatment  of  [***]  in  the  relevant  study,  or  (ii)  [***],  whichever  is  the  shorter  period  of  time.  As  a  minimum
requirement, the raw data comprising [***] will be verified by the sponsor prior to such release Autolus acknowledges the data
is subject to change until database lock at the end of the respective study.

4. During the long term follow up phase of the study the CARPALL and ALLCAR19 Patient Clinical Data shall be released by the
sponsor to Autolus [***], or at the same time as their formal release to the trial management group provided that occurs [***].
5. Autolus shall promptly reimburse UCL for any sums (including internal costs) that UCL incurs in order to comply with UCL’s
obligations to Autolus as set out in Paragraphs 1.2 to 1.4 above (inclusive) and which would not have otherwise been incurred
by UCL in the absence of these obligations. Such reimbursement shall be subject to further terms agreed in any collaboration
agreement entered into in accordance with Clause 4.12.

CAROUSEL Patient Clinical Data

6. Trial management reports produced for the benefit of the trial management groups for the CAROUSEL Study shall be released
by  the  sponsor  to  Autolus  at  the  same  time  as  their  formal  release  to  relevant  trial  management  group.  These  trial
management  reports  shall  include  a  final  report  at  the  end  of  [***]  of  each  study  accompanied  by  release  of  all  supporting
clinical data. UCLB accepts no liability if the frequency of such reporting shall change for any reason and shall not be obliged
to make available such reports before they are received by the relevant trial management group.

7. Raw  data  comprising  the  CAROUSEL  Patient  Clinical  Data  shall  be  released  by  the  sponsor  to  Autolus  (i)  following  the
treatment of [***] in the relevant study, or (ii) [***], whichever is the shorter period of time. As a minimum requirement, the raw
data  comprising  the  [***]  will  be  verified  by  the  sponsor  prior  to  such  release  Autolus  acknowledges  the  data  is  subject  to
change until database lock at the end of the respective study.

8. During the long term follow up phase of the study the CAROUSEL Patient Clinical Data shall be released by the sponsor to

Autolus [***], or at the same time as their formal release to the trial management group provided that occurs [***].

9. Autolus shall promptly reimburse UCL for any sums (including internal costs) that UCL incurs in order to comply with UCL’s
obligations to Autolus as set out in Paragraphs 1.6 and 1.7 above and which would not have otherwise been incurred by UCL
in the absence of these obligations.

    112

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

    113

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

7.

CD19 SEQUENCE[***]

    114

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

8.

(i)

[***]

9.

    115

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IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

ASSIGNED PATENTS[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

[***]

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

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[***]

[***]

[***]

[***]

[***]

[***]

[***]

CERTAIN CONFIDENTIAL INFORMATION CONTAINED IN THIS DOCUMENTS, MARKED BY [***], HAS BEEN OMITTED BECAUSE IT
IS NOT MATERIAL AND WOULD LIKELY CAUSE COMPETITIVE HARM TO THE COMPANY IF PUBLICLY DISCLOSED.

IN WITNESS WHEREOF, the Parties hereto have caused their duly authorised officers to execute and acknowledge this Agreement as of
the date first written above.

SIGNED by a director on behalf of AUTOLUS

LIMITED

SIGNED by a director on behalf of UCL

BUSINESS LTD

)

)

)
)

)
)

Signature……………………………………………..

Print Name…………………………………………...

Signature……………………………………………..

Print Name…………………………………………...

I, Martin Pule, of [***], have read, understand and accept the provisions of this Agreement and how it relates to my research and the MP
Laboratory.

Signed ___________________________________    Date : _____________________

    124

Certification by the Principal Executive Officer pursuant to
Securities Exchange Act Rules 13a-14(a) and 15d-14(a)
as adopted pursuant to Section 302 of the Sarbanes-Oxley Act of 2002

I, Christian Itin, certify that:

1.

2.

3.

4.

(a)

(b)

(c)

(d)

I have reviewed this annual report on Form 20-F of Autolus Therapeutics plc (the “Company”);

Based on my knowledge, this report does not contain any untrue statement of a material fact or omit to state a material fact necessary to
make the statements made, in light of the circumstances under which such statements were made, not misleading with respect to the
period covered by this report;

Based on my knowledge, the financial statements, and other financial information included in this report, fairly present in all material
respects the financial condition, results of operations and cash flows of the Company as of, and for, the periods presented in this report;

The Company’s other certifying officer(s) and I are responsible for establishing and maintaining disclosure controls and procedures (as
defined in Exchange Act Rules 13a-15(e) and 15d-15(e)) and internal control over financial reporting (as defined in Exchange Act
Rules 13a-15(f) and 15d-15(f)) for the Company and have:

Designed such disclosure controls and procedures, or caused such disclosure controls and procedures to be designed under
our supervision, to ensure that material information relating to the Company, including its consolidated subsidiaries, is
made known to us by others within those entities, particularly during the period in which this report is being prepared;

Designed such internal control over financial reporting, or caused such internal control over financial reporting to be
designed under our supervision, to provide reasonable assurance regarding the reliability of financial reporting and the
preparation of financial statements for external purposes in accordance with generally accepted accounting principles;

Evaluated the effectiveness of the Company’s disclosure controls and procedures and presented in this report our
conclusions about the effectiveness of the disclosure controls and procedures, as of the end of the period covered by this
report based on such evaluation; and

Disclosed in this report any change in the Company’s internal control over financial reporting that occurred during the
period covered by the annual report that has materially affected, or is reasonably likely to materially affect, the Company’s
internal control over financial reporting; and

5.

(a)

(b)

The Company’s other certifying officer(s) and I have disclosed, based on our most recent evaluation of internal control over financial
reporting, to the Company’s auditors and the audit committee of the Company’s board of directors (or persons performing the
equivalent functions):

All significant deficiencies and material weaknesses in the design or operation of internal control over financial reporting
which are reasonably likely to adversely affect the Company’s ability to record, process, summarize and report financial
information; and

Any fraud, whether or not material, that involves management or other employees who have a significant role in the
Company’s internal control over financial reporting.

Date: March 4, 2021

/s/ Christian Itin, Ph.D.
Name: Christian Itin, Ph.D.
Title: Chief Executive Officer and Chairman of the Board of Directors
(Principal Executive Officer)

Certification by the Principal Financial Officer pursuant to
Securities Exchange Act Rules 13a-14(a) and 15d-14(a)
as adopted pursuant to Section 302 of the Sarbanes-Oxley Act of 2002

I, Andrew J. Oakley, certify that:

1.

2.

3.

4.

(a)

(b)

(c)

(d)

I have reviewed this annual report on Form 20-F of Autolus Therapeutics plc (the “Company”);

Based on my knowledge, this report does not contain any untrue statement of a material fact or omit to state a material fact necessary to
make the statements made, in light of the circumstances under which such statements were made, not misleading with respect to the
period covered by this report;

Based on my knowledge, the financial statements, and other financial information included in this report, fairly present in all material
respects the financial condition, results of operations and cash flows of the Company as of, and for, the periods presented in this report;

The Company’s other certifying officer(s) and I are responsible for establishing and maintaining disclosure controls and procedures (as
defined in Exchange Act Rules 13a-15(e) and 15d-15(e)) and internal control over financial reporting (as defined in Exchange Act
Rules 13a-15(f) and 15d-15(f)) for the Company and have:

Designed such disclosure controls and procedures, or caused such disclosure controls and procedures to be designed under
our supervision, to ensure that material information relating to the Company, including its consolidated subsidiaries, is
made known to us by others within those entities, particularly during the period in which this report is being prepared;

Designed such internal control over financial reporting, or caused such internal control over financial reporting to be
designed under our supervision, to provide reasonable assurance regarding the reliability of financial reporting and the
preparation of financial statements for external purposes in accordance with generally accepted accounting principles;

Evaluated the effectiveness of the Company’s disclosure controls and procedures and presented in this report our
conclusions about the effectiveness of the disclosure controls and procedures, as of the end of the period covered by this
report based on such evaluation; and

Disclosed in this report any change in the Company’s internal control over financial reporting that occurred during the
period covered by the annual report that has materially affected, or is reasonably likely to materially affect, the Company’s
internal control over financial reporting; and

5.

(a)

(b)

The Company’s other certifying officer(s) and I have disclosed, based on our most recent evaluation of internal control over financial
reporting, to the Company’s auditors and the audit committee of the Company’s board of directors (or persons performing the
equivalent functions):

All significant deficiencies and material weaknesses in the design or operation of internal control over financial reporting
which are reasonably likely to adversely affect the Company’s ability to record, process, summarize and report financial
information; and

Any fraud, whether or not material, that involves management or other employees who have a significant role in the
Company’s internal control over financial reporting.

Date: March 4, 2021

/s/ Andrew J. Oakley
Name: Andrew J. Oakley
Title: Senior Vice President and Chief Financial Officer
(Principal Financial Officer)

Certification by the Principal Executive Officer and Principal Financial Officer pursuant to
18 U.S.C. Section 1350, as adopted pursuant to
Section 906 of the Sarbanes-Oxley Act of 2002

Pursuant to the requirement set forth in Rule 13a-14(b) of the Securities Exchange Act of 1934, as amended (the “Exchange Act”) and Section 1350 of
Chapter 63 of Title 18 of the United States Code (18 U.S.C. §1350), Christian Itin, Chief Executive Officer of Autolus Therapeutics plc (the “Company”),
and Andrew J. Oakley, Senior Vice President and Chief Financial Officer of the Company, each hereby certifies that, to the best of his knowledge:

(1)

(2)

The Company’s Annual Report on Form 20-F for the year ended December 31, 2020, to which this Certification is attached as Exhibit 13.1
(the “Report”) fully complies with the requirements of Section 13(a) or 15(d) of the Securities Exchange Act of 1934, as amended; and

The  information  contained  in  the  Report  fairly  presents,  in  all  material  respects,  the  financial  condition  and  results  of  operations  of  the
Company.

Date: March 4, 2021

/s/ Christian Itin, Ph.D.
Name:  Christian Itin, Ph.D.
Title:    Chief Executive Officer and Chairman of the Board of Directors
(Principal Executive Officer)

/s/ Andrew J. Oakley
Name:   Andrew J. Oakley
Title:    Senior Vice President and Chief Financial Officer
(Principal Financial Officer)    

        
We consent to the incorporation by reference in the following Registration Statements:

Consent of Independent Registered Public Accounting Firm

(1) Registration Statement (Form S-8 No. 333-226457) pertaining to the Autolus Limited 2017 Share Option Plan and Autolus

Therapeutics plc 2018 Equity Incentive Plan, and

(2) Registration Statement (Form F-3 No. 333-232690) of Autolus Therapeutics plc and in the related Prospectus

of our report dated March 4, 2021, with respect to the consolidated financial statements of Autolus Therapeutics plc included in this Annual
Report (Form 20-F) of Autolus Therapeutics plc for the year ended December 31, 2020.

/s/ Ernst & Young LLP
Reading, United Kingdom
March 4, 2021