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Pacific Biosciences of California, Inc.

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FY2023 Annual Report · Pacific Biosciences of California, Inc.
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2023 ANNUAL REPORT

Fellow Stockholders, 

Reflecting on the milestones achieved in 2023 and the opportunities in 2024 and beyond, I’d like to share some thoughts 
on our progress, challenges, and strategic direction for the coming year. 

Positioned for Long-Term, Sustainable Growth  

In 2023, we achieved record revenues surpassing $200 million and 56% top-line growth, driven by the successful launch 
of Revio, our revolutionary long-read sequencer. With Revio, customers were empowered to break the $1,000 long-read 
genome barrier and make significant progress in transforming how researchers analyze the genome. We shipped 173 
Revio systems during the year, a significant milestone that underscores the interest and impact of our platforms and the 
value of our differentiated technologies in providing deeper insights into biology. Nearly 40% of our Revio system orders in 
2023 were from new customers, further demonstrating Revio’s ability to take market share from existing sequencing 
technologies.  

We also began shipping Onso, our short-read sequencer, which has set new standards for accuracy and performance in 
the industry. With the successful launch of these two platforms, we can now address a multibillion-dollar short-read 
sequencing market. As we enter 2024, we’re focused on continuing to drive the adoption of both the Revio and Onso 
platforms and are encouraged by the significant traction we have seen with large-scale projects thus far. 

We ended the year with over $631 million in cash and investments, a testament to our robust financial position and ability 
to fund future growth initiatives. Additionally, we exchanged $441 million of our Convertible Senior Notes due 2028 for 
Convertible Senior Notes due 2030, further improving our capital structure.   

Customers Love Our Technology and Continue to Move Toward Long-Read Sequencing 

Revio has enabled customers to scale with PacBio and long reads like never before. Earlier this year, we shared that 
Revio will be used for large-scale programs like the Estonia Biobank initiative to sequence 10,000 whole human genomes 
using Revio exclusively and The GREGoR Consortium’s project to sequence thousands of samples for rare disease 
research and by customers such as Children’s Mercy Kansas City and Bioscientia who intend to use long reads as first-
line analysis in rare disease cases and replace multiple genetic tests with one HiFi whole genome.  

Today, we believe the sequencing market represents an opportunity of over $7 billion and growing and that PacBio long-
read sequencing, which provides a different level of insight than other available genomic technologies, will enable us to 
secure a large share of the available market opportunity. As we continue to grow and enhance our short-read technology, 
including Onso and our in-development high-throughput sequencer, we believe PacBio is positioned to meet demand 
across all segments of the sequencing market.  

Refocusing Efforts to Address Macro Challenges  

Although we successfully executed our goals for 2023, we have not been immune to the persistent macroeconomic 
challenges that have impacted our entire industry. Preliminary results for the first quarter of 2024 came in below 
expectations.  

While these results and expectations are lower than our initial projections, we remain optimistic about PacBio's long-term 
growth potential. Notably, approximately 57% of Revio placements in the quarter were from new customers, and 
continued adoption in large-scale, multi-thousand sample projects signaled ongoing interest in our platform despite near-
term headwinds. PacBio has some of the world's most advanced sequencing technology, and we are confident in our 
ability to unlock new insights into the genome that will benefit human health and scientific discovery. 

With our revised growth expectations, we are centering around four key strategic priorities: 

Improving commercial execution to drive adoption of both the Revio and Onso platforms. 

1. 
2.  Continuing the development of our benchtop long-read and high throughput short-read platforms. 
3. 
4.  Reducing certain annualized run-rate operating expenses.  

Implementing projects to improve our gross margin and drive manufacturing efficiencies. 

Looking Ahead 

Our performance in 2023 and our ability to capture the opportunities ahead are based on our differentiated sequencing 
technologies and our people. We are grateful for the hard work and dedication of the PacBio team, who continue to 
innovate and develop new products and technologies that provide scientists and researchers with even greater insights to 

better understand biology and health. Together, we are realizing our mission to enable the promise of genomics to better 
human health. 

Thank you for your continued support and trust in PacBio. I look forward to updating you on our progress over the course 
of the year. 

Sincerely,  

Christian Henry 

This letter contains “forward-looking statements” within the meaning of Section 21E of the Securities Exchange Act of 
1934, as amended, and the U.S. Private Securities Litigation Reform Act of 1995. All statements other than statements of 
historical fact are forward-looking statements, including our expectations for future operating results, revenue and 
guidance; our cost-saving plans and initiatives as well as the expected financial impact and timing of these plans and 
initiatives; and our potential market opportunity and total addressable market. You should not place undue reliance on 
forward-looking statements because they are subject to assumptions, risks, and uncertainties that could cause actual 
outcomes and results to differ materially from currently anticipated results. These risks include, but are not limited to, 
challenges inherent in developing, manufacturing, launching, marketing and selling new products, and achieving 
anticipated new sales; potential cancellation of existing instrument orders; assumptions, risks and uncertainties related to 
the ability to attract new customers and retain and grow sales from existing customers; risks related to lengthening sales 
cycles; risks related to PacBio’s ability to successfully execute and realize the benefits of acquisitions; the estimated size 
and estimated growth for the markets for PacBio’s products may be smaller than expected; the impact of U.S. export 
restrictions on the shipment of PacBio products to certain countries; rapidly changing technologies and extensive 
competition in genomic sequencing; unanticipated increases in costs or expenses; interruptions or delays in the supply of 
components or materials for, or manufacturing of, PacBio products and products under development; potential product 
performance and quality issues and potential delays in development and commercialization timelines; the possible loss of 
key employees, customers, or suppliers; customers and prospective customers curtailing or suspending activities using 
PacBio’s products; third-party claims alleging infringement of patents and proprietary rights or seeking to invalidate 
PacBio’s patents or proprietary rights; risks associated with international operations; and other risks associated with 
general macroeconomic conditions and geopolitical instability. Additional factors that could materially affect actual 
results can be found in PacBio’s most recent filings with the Securities and Exchange Commission, including PacBio’s most 
recent reports on Forms 8-K, 10-K, and 10-Q, and include those listed under the caption "Risk Factors." These forward-
looking statements are based on current expectations and speak only as of the date hereof; except as required by law, we 
disclaim any obligation to revise or update these forward-looking statements to reflect events or circumstances in the 
future, even if new information becomes available. 

 
 
 
 
 
 
 
 
 
 
 
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UNITED STATES
SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549 
________________________________________________________________________________
Form 10-K
________________________________________________________________________________
(Mark One)
x	ANNUAL REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT OF 
1934
For the fiscal year ended December 31, 2023

Or
o	TRANSITION REPORT PURSUANT TO SECTION 13 OR 15(d) OF THE SECURITIES EXCHANGE ACT 
OF 1934
For the transition period from         to
Commission File Number 001-34899
________________________________________________________________________________

Pacific Biosciences of California, Inc.
(Exact name of registrant as specified in its charter)
________________________________________________________________________________

Delaware

(State or other jurisdiction of
incorporation or organization)

1305 O’Brien Drive 
Menlo Park, CA 94025

(Address of principal executive offices)

16-1590339

(I.R.S. Employer
Identification No.)

94025

(Zip Code)

(Registrant’s telephone number, including area code)
(650) 521-8000
Securities registered pursuant to Section 12(b) of the Act: 

Title of each class 

Trading Symbol(s)

Name of each exchange on which registered 

Common Stock, par value $0.001 per share

PACB

The NASDAQ Stock Market LLC

Securities registered pursuant to Section 12(g) of the Act: 
None
________________________________________________________________________________

Indicate by check mark if the registrant is a well-known seasoned issuer, as defined in Rule 405 of the Securities Act. Yes x No o

Indicate by check mark if the registrant is not required to file reports pursuant to Section 13 or Section 15(d) of the Act. Yes o No 

x

Indicate by check mark whether the registrant (1) has filed all reports required to be filed by Section 13 or 15(d) of the Securities 
Exchange Act of 1934 during the preceding 12 months (or for such shorter period that the registrant was required to file such reports), 
and (2) has been subject to such filing requirements for the past 90 days. Yes x No o

Indicate  by  check  mark  whether  the  registrant  has  submitted  electronically  every  Interactive  Data  File  required  to  be  submitted 
pursuant to Rule 405 of Regulation S-T (§232.405 of this chapter) during the preceding 12 months (or for such shorter period that the 
registrant was required to submit such files). Yes x No o

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Indicate  by  check  mark  whether  the  registrant  is  a  large  accelerated  filer,  an  accelerated  filer,  a  non-accelerated  filer,  a  smaller 
reporting  company  or  an  emerging  growth  company.  See  the  definitions  of  “large  accelerated  filer,”  “accelerated  filer,”  “smaller 
reporting company” and “emerging growth company” in Rule 12b-2 of the Exchange Act. (Check one): 

Large accelerated filer

Non-accelerated filer

x

o

Accelerated filer

Smaller reporting company

Emerging growth company

o

o

o

If an emerging growth company, indicate by check mark if the registrant has elected not to use the extended transition period for 

complying with any new or revised financial accounting standards provided pursuant to Section 13(a) of the Exchange Act. o

Indicate  by  check  mark  whether  the  registrant  has  filed  a  report  on  and  attestation  to  its  management’s  assessment  of  the 
effectiveness of its internal control over financial reporting under Section 404(b) of the Sarbanes-Oxley Act (15 U.S.C. 7262(b)) by the 
registered public accounting firm that prepared or issued its audit report. Yes x No o

If  securities  are  registered  pursuant  to  Section  12(b)  of  the  Act,  indicate  by  check  mark  whether  the  financial  statements  of  the 

registrant included in the filing reflect the correction of an error to previously issued financial statements. o

Indicate by check mark whether any of those error corrections are restatements that required a recovery analysis of incentive-based 

compensation received by any of the registrant’s executive officers during the relevant recovery period pursuant to §240.10D-1(b). o

Indicate by check mark whether the registrant is a shell company (as defined in Rule 12b-2 of the Exchange Act). Yes o No x

Aggregate  market  value  of  registrant’s  common  stock  held  by  non-affiliates  of  the  registrant  on  June  30,  2023,  based  upon  the 
closing price of Common Stock on such date as reported by NASDAQ Global Select Market, was approximately $3,323,701,933. Shares 
of  voting  stock  held  by  each  officer  and  director  have  been  excluded  in  that  such  persons  may  be  deemed  to  be  affiliates.  This 
assumption regarding affiliate status is not necessarily a conclusive determination for other purposes. 

Number of shares outstanding of the registrant’s common stock as of January 31, 2024: 267,951,880

DOCUMENTS INCORPORATED BY REFERENCE:

Portions  of  the  registrant’s  definitive  Proxy  Statement  relating  to  its  2024  Annual  Meeting  of  Stockholders  are  incorporated  by 
reference into Part III of this Annual Report on Form 10-K where indicated. Such Proxy Statement will be filed with the U.S. Securities 
and Exchange Commission within 120 days after the end of the fiscal year to which this report relates. 

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Pacific Biosciences of California, Inc.

Annual Report on Form 10-K 

For the Fiscal Year Ended December 31, 2023

Table of Contents

PART I

Item 1.

Business

Item 1A. Risk Factors

Item 1B. Unresolved Staff Comments

Item 1C. Cybersecurity

Item 2.

Properties

Item 3.

Legal Proceedings

Item 4. Mine Safety Disclosures

PART II

Item 5. Market for Registrant’s Common Equity, Related Stockholder Matters and Issuer 

Purchases of Equity Securities

Item 6.

[Reserved]

Item 7. Management’s Discussion and Analysis of Financial Condition and Results of 

Operations

Item 7A. Quantitative and Qualitative Disclosures about Market Risk

Item 8.

Financial Statements and Supplementary Data

Item 9.

Changes in and Disagreements with Accountants on Accounting and Financial 
Disclosure

Item 9A. Controls and Procedures

Item 9B. Other Information

Item 9C Disclosure Regarding Foreign Jurisdictions that Prevent Inspections

PART III

Item 10. Directors, Executive Officers and Corporate Governance

Item 11. Executive Compensation

Item 12. Security Ownership of Certain Beneficial Owners and Management and Related 

Stockholder Matters

Item 13. Certain Relationships and Related Transactions, and Director Independence

Item 14. Principal Accountant Fees and Services

PART IV

Item 15. Exhibits, Financial Statement Schedules

Item 16. Form 10-K Summary

Signatures

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SPECIAL NOTE REGARDING FORWARD-LOOKING STATEMENTS

This  Annual  Report  on  Form  10-K,  including  the  sections  titled  “Business,”  “Risk  Factors”  and  “Management’s 
Discussion  and  Analysis  of  Financial  Condition  and  Results  of  Operations”  contain  or  may  contain  forward-
looking statements that are based on the beliefs and assumptions of the management of Pacific Biosciences of 
California, Inc. (the “Company,” “we,” “us,” or “our”) and on information currently available to our management. 
The statements contained in this Annual Report on Form 10-K that are not purely historical are forward-looking 
statements within the meaning of Section 27A of the Securities Act of 1933, as amended, and Section 21E of 
the Securities Exchange Act of 1934, as amended (the “Exchange Act”), and include, but are not limited to:

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the  availability,  uses,  accuracy,  sensitivity,  advantages,  compatibility,  pricing,  specifications,  quality,  or 
performance of, or benefits or expected benefits or using, our products or technologies, including the 
RevioTM and OnsoTM systems;

our current and future products; 

our strategic and commercial plans, including our expectations for Revio and Onso systems; 

our market opportunity, including market size and expected market growth;

our expectations regarding the conversion of backlog to revenue and the pricing and gross margin for 
products;

our  manufacturing  plans  including  developing  and  scaling  of  manufacturing  and  delivery  of  our 
products; 

our research and development plans; 

the anticipated impact of catastrophic events, including health epidemics or pandemics and military or 
other armed conflicts, on our business, business plans and results of operations;

our  product  development  plans,  roadmaps,  and  objectives,  including,  among  other  things,  statements 
relating  to  future  uses,  quality,  or  performance  of,  or  benefits  of  using,  products  or  technologies, 
updates, or improvements of our products; 

our intentions regarding seeking regulatory approval for our products; 

our competitive landscape, including competition in the short- and long-read sequencing technologies 
markets;

our expectations regarding collaborations and partnerships; 

our expectations regarding unrecognized income tax benefits; 

our  expectations  regarding  market  risk,  including  interest  rate  changes  and  general  macroeconomic 
conditions; 

the sufficiency of cash, cash equivalents, and investments to fund projected operating requirements;

the effects of recent accounting pronouncements on our financial statements; and 

other future events. 

Forward-looking  statements  can  be  identified  by  words  such  as:  “anticipates,”  “believes,”  “could,”  “estimates,” 
“expects,” “intends,” “may,” “plans,” “potential,” “predicts,” “projects,” “seeks,” “should,” “target,” “will,” “would,” or 
similar expressions and the negatives of those terms. Forward-looking statements involve known and unknown 
risks,  uncertainties,  and  other  factors  that  may  cause  our  actual  results,  performance,  or  achievements  to  be 
materially different from any future results, performance, or achievements expressed or implied by the forward-
looking statements. 

Factors that could cause or contribute to such differences include, but are not limited to, those discussed under 
the  heading  “Risk  Factors”  in  this  report  and  in  other  documents  we  file  with  the  Securities  and  Exchange 
Commission (SEC). Given these risks and uncertainties, you should not place undue reliance on forward-looking 
statements. Also, forward-looking statements represent management’s beliefs and assumptions as of the date 
of  this  report.  Except  as  required  by  law,  we  assume  no  obligation  to  update  forward-looking  statements 
publicly, or to update the reasons actual results could differ materially from those anticipated in these forward-
looking statements, even if new information becomes available in the future.

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This  Annual  Report  on  Form  10-K  also  contains  estimates,  projections,  and  other  information  concerning  our 
industry,  our  business,  and  the  markets  for  our  products,  including  data  regarding  the  estimated  size  and 
estimated  growth  for  those  markets.  Information  that  is  based  on  estimates,  forecasts,  projections,  market 
research,  or  similar  methodologies  is  inherently  subject  to  uncertainties  and  actual  events  or  circumstances 
may differ materially from events and circumstances reflected in this information. Unless otherwise expressly 
stated, we obtained this industry, business, market, and other data from reports, research surveys, studies, and 
similar  data  prepared  by  market  research  firms  and  other  third  parties,  industry,  medical  and  general 
publications, government data, and similar sources.

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ITEM 1. BUSINESS

Overview 

PART I

We are a premier life science technology company that is designing, developing, and manufacturing advanced 
sequencing  solutions  that  enable  scientists  and  clinical  researchers  to  improve  their  understanding  of  the 
genome and ultimately, resolve genetically complex problems. 

Our products and technology under development stem from two highly differentiated core technologies focused 
on  accuracy,  quality,  and  completeness,  which  include  our  HiFi  long-read  sequencing  technology  and  our 
Sequencing  by  Binding  (SBB®)  short-read  sequencing  technology.  Our  products  address  solutions  across  a 
broad  set  of  applications  including  human  genetics,  plant  and  animal  sciences,  infectious  disease  and 
microbiology, oncology, and other emerging applications. Long-read sequencing was recognized by the journal 
Nature Methods as its “method of the year” for 2022 for its contributions to biological understanding and future 
potential.

Our focus is on creating some of the world`s most advanced sequencing systems to provide our customers the 
most complete and accurate view of genomes, transcriptomes, and epigenomes.

Our  customers  include  academic  and  governmental  research  institutions,  commercial  testing  and  service 
laboratories,  genome  centers,  public  health  labs,  hospitals  and  clinical  research  institutes,  contract  research 
organizations (CROs), pharmaceutical companies, and agricultural companies.

Recent Developments

In 2023, we commercially released two new sequencing platforms, RevioTM and OnsoTM.

Revio is a new long-read sequencing system designed to enable the use of HiFi sequencing for large studies in 
human genetics, cancer research, and agricultural genomics. Commercial shipments for Revio commenced in 
the  first  quarter  of  2023,  and  173  systems  have  since  been  shipped  to  more  than  twenty  countries  as  of 
December  31,  2023.  We  released  SMRT  Link  13.0  software  on  the  Revio  system  which  includes  the  adaptive 
loading  feature  for  consistent  run  performance,  run  preview  for  improved  lab  efficiency,  and  expanded 
application support with functionality to sequence shorter and longer fragments of DNA.

Onso, a short-read DNA sequencing system, is designed to deliver industry-leading sensitivity and specificity for 
novel  insights  in  oncology,  disease  research,  and  other  applications.  We  commenced  customer  shipments  of 
Onso in the third quarter of 2023.

In  August  2023,  we  acquired  Apton  Biosystems  to  accelerate  the  development  of  a  high-throughput  platform 
based on the SBB chemistry utilized by Onso. Through acquiring Apton, we obtained expertise in state-of-the-art 
optics and image processing and a novel clustering method and chemistry designed to enable the sequencing 
of billions of clusters of DNA on one flow cell. 

We also develop products used to prepare molecules for sequencing on our platforms. In the fourth quarter of 
2023,  we  commenced  commercial  shipments  for  a  new  KinnexTM  line  of  products  used  to  concatenate  small 
molecules  into  larger  fragments  to  optimize  the  output  on  HiFi  sequencing  systems  like  Revio.  The  Kinnex 
products are designed for various applications in RNA and amplicon sequencing. As of December 31, 2023, we 
received orders from over 70 customers.

Our Mission and Impact

Our  mission  is  to  enable  the  promise  of  genomics  to  better  human  health.  Genomics  is  core  to  all  biological 
processes,  and  our  advanced  genomics  tools  provide  scientists  and  clinical  researchers  with  the  insights  to 
better understand biology and health. The “promise of genomics” postulates that medicine, agriculture, public 
health,  drug  development,  and  other  disciplines  will  be  transformed  by  incorporating  routine  genomic 
information over the coming decades. We see early progress toward this transformation in the applied use of 
genomics  in  areas  such  as  genetic  disease,  oncology,  and  sustainable  food  production.  However,  legacy 
genomics  technologies  have  fundamental  limitations  in  progressing  these  fields  toward  the  promise  of 
genomics.  We  believe  that  unleashing  the  full  potential  of  genomics  will  require  a  level  of  accuracy  and 
completeness  inaccessible  to  legacy  technologies.  Accuracy  and  completeness  are  central  to  our  product 
development strategy; thus, we have created some of the most innovative and high-quality genomics solutions 
on  the  market.  Our  products  also  have  enhanced  multi-omic  capabilities  to  look  beyond  the  genome  to  the 
transcriptome and epigenome, which we believe is key to understanding a full picture of biology.

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The Underlying Science

Genetic  inheritance  in  living  organisms  is  conveyed  through  a  naturally  occurring  information  storage  system 
known  as  deoxyribonucleic  acid,  or  DNA.  DNA  stores  information  in  linear  chains  of  the  chemical  bases 
adenine, cytosine, guanine, and thymine, represented by the symbols A, C, G, and T respectively.

In humans, the genome is comprised of approximately three billion DNA base pairs, which are divided into 23 
chromosomes  ranging  in  size  from  50  million  to  250  million  bases.  A  human  carries  two  copies  of  the 
inherited  from  each  parent.  Approximately  23,000  smaller  regions  within  these 
chromosomes,  one 
chromosomes, called genes, contain the blueprints for protein production. The proteins synthesized from these 
blueprints essentially underlie the operation of all biological systems.

Genome  sequencing  reads  the  bases  of  long  fragments  of  nucleic  acids.  Initial  genome  sequencing  studies 
have  shown  that  mutations  in  these  DNA  base  pairs  play  a  critical  role  in  human  disease,  contributing  to  the 
burgeoning  field  of  genomics.  Since  then,  recent  discoveries  have  highlighted  additional  complexities  of  DNA 
and RNA. These discoveries include the presence of chemical modifications to the bases, such as methylation, 
and  post-translational  modification,  or  the  processing  of  RNA  molecules  after  they  are  transcribed  from  the 
genome, both of which can affect protein synthesis. 

Our Principal Markets 

Researchers  utilize  our  solutions  in  human  genomics,  plant  and  animal  sciences,  infectious  disease  and 
microbiology, oncology, and other emerging applications.

Human Genomics: Improving rare disease research and understanding

According  to  a  World  Health  Organization  publication,  it  is  estimated  that  400  million  people  worldwide  are 
affected by up to 8,000 distinct rare diseases, with 80% of these believed to be genetic in nature. These genetic 
diseases  are  DNA  differences,  called  variants,  in  the  affected  individuals.  Variants  range  in  size  from  single 
nucleotide  substitutions  to  large  losses  or  gains  of  entire  chromosomes.  Other  sequencing  technologies 
applied to rare disease diagnosis are technologically limited to interrogating small variants, representing only a 
subset  of  possible  genomic  variation.  Consequently,  most  genetic  disease  cases  are  undiagnosed,  leaving 
families  on  multi-year  diagnostic  odysseys.  Sequencing  the  human  genome  with  long  and  accurate  reads 
enables  the  potential  detection  of  all  known  classes  of  disease-causing  variation.  In  addition,  the  ability  of 
PacBio’s  long-read  sequencing  technology  to  detect  5-Methylcytosine  DNA  methylation,  an  epigenetic 
modification  shown  to  alter  gene  behavior,  may  enable  further  advances  in  research  and  development  in 
genetic disease diagnosis.

Infectious Disease and Microbiology: Understanding and tracking microbes and pathogens in support of global 
public health

Our  technology  has  increased  the  scientific  community’s  understanding  of  microorganisms  and  viruses  and 
their malignancy, transmission, and potential resistance to antibiotics or vaccines. Our sequencing technology 
delivers highly comprehensive and complete genomes, enabling federal agencies, public health organizations, 
and healthcare providers to conduct wide-ranging research and surveillance activities to:

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generate  high  quality,  complete  genome  assemblies,  revealing  variants  of  all  known  types,  to  gain  a 
deeper understanding of community-acquired and hospital-associated infections and transmissions;

identify and characterize pathogens to inform regional, national, and global public health agencies for 
preparation and response to rapidly evolving microorganisms; and

characterize  complex  microbial  communities  to  understand  their  role  in  human,  animal,  and 
environmental health.

Oncology: Enable the discoveries of underlying causes of cancer, progression, and relapse

Understanding  tumor  cells`  cellular  and  molecular  complexity  is  critical  in  developing  more  effective  targeted 
cancer  therapies.  Single-cell  transcriptomics  is  particularly  impactful  in  defining  cellular  identity  and  function; 
however, other technologies miss critical information by only sequencing a portion of RNA. Our long-read RNA 
sequencing method, single-cell Iso-Seq (scIso-Seq), accurately detects molecular events such as RNA isoforms 
and  expressed  mutations  and  provides  gene  expression  information  at  the  single-cell  level.  We  believe  scIso-
Seq  is  uniquely  positioned  to  enable  discoveries  by  researchers  of  the  underlying  causes  of  cancer  initiation, 
progression, and relapse, as well as the discovery by researchers of novel diagnostic, prognostic, and predictive 
biomarkers that may inform future clinical tests. 

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As novel discoveries continue to be made using our long-read sequencing technology, we believe our SBB short-
read sequencing technology will enable us to meet customers` demands in the expanding non-invasive testing 
market in oncology. Due to the small amounts of circulating tumor DNA (ctDNA) present in the blood of early-
stage  cancer  patients  and  those  with  minimal  residual  disease  (MRD),  the  presence  of  cancer  often  goes 
undetected, and a more sensitive assay will be required. Based on testing internally and at beta customer sites, 
we  believe  our  SBB  technology  has  the  potential  to  offer  higher  accuracy  than  competitor  sequencing 
technologies,  which  may  in  the  future  support  our  customers’  development  of  more  sensitive  tests  for  the 
purpose of earlier detection and more robust monitoring of cancer.

Plant and Animal Sciences: Helping scientists answer biological questions for a healthier world 

In  Plant  and  Animal  Sciences,  academic,  government  and  corporate  researchers  are  using  our  technology  to 
explore  and  catalog  the  genetic  and  biological  diversity  of  organisms  for  the  breeding,  propagation,  and 
production  of  crops  and  livestock  while  conserving  the  planet’s  natural  resources.  PacBio  HiFi  sequencing 
enables  researchers  to  build  high  quality  de  novo  reference  genomes  and  transcriptomes  to  study  variations 
across  species  enabling  improvements  to  global  conservation  initiatives  and  support  the  breeding  and 
production of resilient and higher yielding crops to meet the world's growing population and demand.   

Our Technology, Products, and Solutions

We have developed HiFi long-read sequencing based on Single-Molecule Real-Time (SMRT) technology, which 
accurately  detects  the  nucleotide  sequence  and  epigenetic  status  of  individual  DNA  molecules.  We  are  also 
expanding  our  genomic  solutions  with  our  short-read  SBB  chemistry,  which  offers  sensitive  sequencing  for 
short-read applications.

Our sales consist of sequencing instruments, nanofluidic chips (SMRT Cells), and reagents for preparing DNA 
and  performing  sequencing  based  on  our  SMRT  technology;  flow  cells  and  reagents  for  preparing  DNA  and 
performing  sequencing  based  on  our  SBB  technology,  reagents  for  DNA  extraction  based  on  our  Nanobind 
technology; and the services we perform for customers. 

HiFi Long-Read Sequencing

Our HiFi long-read sequencing protocol was built upon our SMRT sequencing systems, including consumables 
and  software,  and  offers  customized  end-to-end  workflows  for  different  sequencing  applications.  Highly 
accurate,  long  sequence  reads  simplify  and  accelerate  data  analysis  algorithms,  reducing  the  need  for  error 
correction steps and/or assembly aspects, depending on the application.

Customers use our HiFi long-read sequencing platforms in a wide range of sequencing applications, including 
whole genome sequencing and de novo genome assembly, long-range phasing, targeted sequencing, full-length 
RNA and single-cell sequencing, characterization of metagenomic communities and other mixed DNA samples, 
viral  genome  sequencing,  and  others.  Our  technology  is  also  capable  of  detecting  epigenetic  markers 
simultaneously  by  analyzing  the  kinetics  of  DNA  polymerization  that  is  affected,  and  thereby  detectable,  by 
epigenetic markers such as 5-methylcytosine or N6-methyladenine. 

SMRT Technology

Our  proprietary  SMRT  Technology  enables  the  observation  of  DNA  synthesis  as  it  occurs  in  real-time  by 
harnessing  the  natural  process  of  DNA  replication,  which  in  nature  is  a  highly  efficient  and  accurate  process 
actuated  by  DNA  polymerases.  DNA  polymerases  attach  to  a  strand  of  DNA  to  be  replicated,  examine  the 
individual base at the point it is attached, and then determine which of the four building blocks, or nucleotides 
(A, C, G, or T), is required to complement that individual base. After determining which nucleotide is required, the 
polymerases incorporate that nucleotide into the growing strand being produced.

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SMRT  Sequencing  is  based  on  following  the  activity  of  DNA  polymerase  on  individual  DNA  molecules  in  real 
time  that  occurs  on  our  SMRT  Cells  that  are  monitored  and  analyzed  within  our  HiFi  long-read  sequencing 
systems: the Revio system, Sequel II system, Sequel IIe system, and Sequel system. Carried out on SMRTbell® 
templates,  which  attach  hairpin  adapters  to  the  ends  of  double-stranded  DNA  molecules  to  be  sequenced, 
SMRT  sequencing  allows  for  the  successive  sequencing  of  both  the  forward  and  reverse  strands  of  the 
individual DNA molecule occurring multiple times, thereby allowing for the same base of the same molecule to 
be sequenced more than once in a sequencing run. The base calls from the serial observation of the molecule 
can  be  processed  to  generate  the  final  base  call  in  an  analytical  procedure  called  circular  consensus 
sequencing,  leading  to  what  we  have  defined  as  our  HiFi  sequence  reads,  which  have  high  accuracy  typically 
being  defined  as  having  greater  than  99%  read  accuracy,  but  often  exceeding  greater  than  99.9%  accuracy, 
according to research we performed in collaboration with other researchers, subsequently published in Nature 
Biotechnology  in  2019.  HiFi  reads  typically  are  15-20  kilobases  in  size,  depending  on  the  input  fragments, 
providing sufficient read length with our accuracy to support a multitude of applications across human health, 
plant and animal, and microbiology, according to research we performed in collaboration with other researchers, 
subsequently published in Scientific Data in 2020. The ability to generate single-DNA molecule sequence reads 
that are both long and highly accurate allows researchers to obtain more contiguous, complete, and accurate 
genomic data, thereby allowing for greater insights into the complexity of biological systems.

HiFi Long-Read Sequencing Instruments: Revio system + Sequel systems 

Our  Revio,  Sequel,  Sequel  II,  and  Sequel  IIe  instruments  conduct,  monitor,  and  analyze  single-molecule 
biochemical reactions in real time. The instruments use extremely sensitive imaging systems to collect the light 
pulses emitted by fluorescent reagents allowing the observation of biological processes. Computer algorithms 
are used to translate the information that is captured by the optics system. Using the recorded information, light 
pulses are converted into either an A, C, G, or T base call with associated quality metrics. Once sequencing is 
started, the real-time data is delivered to the system’s primary analysis pipeline, which outputs base identity and 
quality values.

HiFi Consumables

Customers  purchase  proprietary  consumable  products  to  run  their  PacBio  systems,  including  our  SMRT  Cells 
and reagent kits. One SMRT Cell is consumed per sequencing reaction, and scientists can choose the number 
of SMRT Cells they use per experiment. 

We  offer  several  reagent  kits,  each  designed  to  address  a  specific  step  in  the  core  sequencing  workflow.  A 
library preparation kit is used to convert DNA into SMRTbell double-stranded DNA library formats and includes 
typical  molecular  biology  reagents,  such  as  ligase,  buffers,  and  exonucleases.  Our  binding/polymerase  kits 
include our modified DNA polymerase and are used to bind SMRTbell libraries to the polymerase in preparation 
for  sequencing.  Our  core  sequencing  kits  contain  reagents  required  for  on-instrument,  real-time  sequencing, 
including phospholinked nucleotides.

We have developed and offered a new line of Kinnex kits with companion SMRT Link software to enable high-
throughput,  scalable,  cost-effective  RNA  applications  including  bulk  RNA,  single-cell  RNA,  and  16S  rRNA 
sequencing.  The  Kinnex  kits  are  built  upon  the  Multiplexed  Array  Sequencing  (MAS-Seq)  method  for 
concatenating  smaller  amplicons  into  larger  fragments  to  sequence  on  PacBio's  long  read  sequencers, 
increasing the molecular yield by up to 16-fold. Throughput increase in these key RNA applications where the 
dynamic range of different RNA isoforms (bulk and single-cell) or microbial species (16S) can vary by orders of 
magnitude,  enables  characterization  of  these  complex  RNA  samples  while  drastically  reducing  sequencing 
need.

SBB Short-Read Sequencing

In contrast to SMRT sequencing, SBB reads short fragments of DNA (hundreds of bases instead of kilobases) in 
a  massively  parallel  manner.  Current  short-read  next  generation  sequencing  technologies  available  in  the 
market  incur  various  rates  of  errors  in  results.  Researchers  deploy  multiple  tactics  to  try  to  mitigate  these 
effects, including oversampling or implementing complex library preparation methods, yet still face challenges, 
including missing rare variants.

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We believe our proprietary SBB approach will enable researchers to address the gap in detecting rare variants, 
especially in complex heterogenous samples. Employing a two-phase sequencing chemistry, the SBB approach 
binds a dye-labeled nucleotide without incorporation into the DNA chain, then removes that base, then blocks 
and  extends  with  a  terminated  nucleotide.  Using  nucleotides  with  single  modifications,  we  incorporate  more 
native bases, avoiding potential scarring due to fluorescent linker presence. This design helps avoid raw errors 
and we believe can help us develop a product with substantially greater accuracy than currently marketed short-
read sequencing products. SBB enables simplified upfront library preparation, redefines coverage requirements, 
and reduces bioinformatic workload for downstream analysis. The accuracy of our novel sequencing approach 
has  the  potential  to  advance  translational  cancer  research,  drive  higher  fidelity  single-cell  applications,  and 
broadly enable clinical sequencing—even in regions of the genome prone to sequencing errors with other short-
read sequencing technologies.

SBB Short-Read Sequencing Instrument: Onso system 

Our  Onso  instrument  conducts,  monitors,  and  analyzes  SBB  biochemical  reactions.  The  instrument  uses 
extremely  sensitive  imaging  systems  to  collect  the  light  emitted  by  fluorescent  reagents  allowing  the 
observation of biological processes. Computer algorithms are used to translate the information that is captured 
by the optics system. Using the recorded information, light pulses are converted into either an A, C, G, or T base 
call with associated quality metrics. Once sequencing is started, the imaging data is delivered to the system’s 
primary analysis pipeline, which outputs base identity and quality values.

SBB Consumables

To  complement  our  Onso  instrument,  we  also  sell  a  range  of  SBB  consumable  products  including  flow  cells, 
clustering,  and  sequencing  reagent  kits.  One  flow  cell  and  associated  sequencing  reagent  pack  is  consumed 
per  sequencing  reaction.  Each  flow  cell  contains  two  lanes  and  scientists  can  choose  to  sequence  different 
samples in each lane while additionally combining any number of flow cells needed per experiment. 

We  offer  several  reagent  kits,  each  designed  to  address  a  specific  step  in  the  core  sequencing  workflow.  A 
library  preparation  kit  is  used  to  convert  DNA  into  SBB  compatible,  double-stranded  DNA  library  formats  and 
includes typical molecular biology reagents, such as ligase, buffers, and exonucleases. Additionally for library 
preparation,  our  conversion  kits  include  reagents  to  enable  scientists  to  convert  existing  sequencing  libraries 
into  an  SBB  compatible  format.  Finally,  our  clustering  and  sequencing  kits  contain  all  reagents  required  for 
generating  sequence  ready  clusters  on  flow  cell  and  performing  SBB  sequencing  reactions  on  instrument, 
respectively.

Our Strategy for Growth

To enable the promise of genomics, our strategy includes the following key elements:

•

•

•

•

Increase  technology  adoption  by  increasing  market  share  via  new  customer  acquisition,  continue 
Sequel II conversions to Revio, and scale Onso production; 

Leverage  innovation  to  complete  development  of  new  sequencing  platforms  and  launch  on-market 
system improvements;

Build  upon  clinical  momentum  by  expanding  HiFi  usage  in  large-scale  programs  and  translational 
research projects; and

Drive  towards  positive  cash  flow  through  gross  margin  expansion,  disciplined  operating  expense 
management, and a focus on working capital.

Marketing, Sales, Service, and Support 

We  market  our  products  through  a  global  sales  force  and  through  distribution  partners  in  Asia  and  Australia, 
certain parts of Europe, the Middle East and Africa, and Latin America. We plan to continue to invest in growing 
our  marketing,  sales,  service,  and  support  resources  as  we  drive  continued  adoption  of  products,  launch  new 
products, and expand our customer base.

Our business is subject to seasonal trends. See the Risk Factors section, specifically the risk factor titled Our 
operating results fluctuate from quarter to quarter and year over year, which makes our future results difficult to 
predict and could negatively impact the market price of our common stock, for additional information.

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Customers 

Our  customers  include  academic  and  governmental  research  institutions,  commercial  testing  and  service 
laboratories,  genome  centers,  public  health  labs,  hospitals  and  clinical  research  institutes,  contract  research 
organizations  (CROs),  pharmaceutical  companies,  and  agricultural  companies.  In  general,  our  customers  will 
isolate,  prepare,  and  analyze  genetic  samples  using  PacBio  sequencing  systems  in  their  own  laboratories,  or 
they will send their genetic samples to third-party service providers who in turn will sequence the samples with 
PacBio  systems  and  provide  the  sequence  data  back  to  the  customer  for  further  analysis.  For  example, 
customers in academic research institutions may have bacteria, animal, or human DNA samples isolated from 
various sources while agricultural biology companies may have DNA samples isolated from different strains of 
rice, corn, or other crops. For the year ended December 31, 2023, no single customer accounted for 10% or more 
of  our  total  revenue.  For  the  years  ended  December  31,  2022,  and  2021,  one  customer  accounted  for 
approximately 12% and 13% of our total revenue, respectively.

We believe that the majority of our current customers are early adopters of sequencing technology. By focusing 
our  efforts  on  high-value  applications,  and  developing  whole  product  solutions  around  these  applications,  we 
seek  to  drive  the  adoption  of  our  products  across  a  broader  customer  base  and  into  numerous  large-scale 
projects. In general, the broader adoption of new technologies by mainstream customers can take a number of 
years. 

Backlog 

As of December 31, 2023, our product backlog was approximately $18.7 million, compared to $51.5 million as 
of December 31, 2022. We define backlog as purchase orders or signed contracts from our customers, which 
we believe are firm and for which we have not yet recognized revenue. We expect to convert the majority of this 
backlog to revenue during 2024; however, our ability to do so is subject to customers who may seek to cancel or 
delay their orders even if we are prepared to fulfill them. 

Manufacturing 

We manufacture sequencing instruments, SMRT Cells, and reagents. Our key manufacturing and service facility 
in  Menlo  Park,  California  has  received  ISO  13485  and  ISO  9001  certifications  for  the  design,  development, 
manufacture,  distribution,  installation,  and  servicing  of  its  nucleic  acid  sequencing  platforms.  We  utilize 
subcontract manufacturers for components of the manufacturing process. We purchase both custom and off-
the-shelf components from a large number of suppliers and subject them to significant quality specifications. 
We  periodically  conduct  quality  audits  of  most  of  our  critical  suppliers  and  have  established  a  supplier 
qualification  program.  Some  of  the  components  required  in  our  products  are  currently  either  sole  sourced  or 
single sourced. 

Research and Development 

We have historically made and plan to continue to make significant investments in research and development. 
Our  research  and  development  efforts  focus  on  programs  to  develop  new  and  existing  platforms,  as  well  as 
increasing  throughput  and  decreasing  costs  on  behalf  of  our  customers.  We  are  currently  developing  higher 
throughput  platforms  that  utilize  our  HiFi  long-read  sequencing  and  SBB  short-read  sequencing  technologies. 
We also are developing a benchtop HiFi platform intended to improve accessibility for smaller laboratories, and 
we continue to extend our on-market platforms, Revio and Onso. In addition to platform development, we also 
innovate  across  end-to-end  workflows  to  improve  usability,  as  well  as  develop  new  applications  for  the 
advancement of human health. 

Intellectual Property 

Developing and maintaining a strong intellectual property portfolio is an important element of our business. We 
have sought, and will continue to seek, patent protection for our SMRT and SBB technology, for improvements 
to  our  SMRT  kit  and  SBB  technology,  as  well  as  for  any  of  our  other  technologies  where  we  believe  such 
protection will be advantageous. 

Our  current  patent  portfolio,  including  patents  exclusively  licensed  to  us,  is  directed  to  various  technologies, 
including  SMRT  nucleic  acid  sequencing  and  other  methods  for  analyzing  biological  samples,  zero-mode 
waveguide (ZMW) arrays, surface treatments, phospholinked nucleotides and other reagents for use in nucleic 
acid  sequencing,  optical  short-read  nucleic  acid  sequencing,  nucleic  acid  preparation,  and  purification 
components and systems, processes for identifying nucleotides within nucleic acid sequences, and processes 
for analysis and comparison of nucleic acid sequence data. Some of the patents and applications that we own, 
as well as some of the patents and applications that we have licensed from other parties, are subject to U.S. 

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government march-in rights, whereby the U.S. government may disregard our exclusive patent rights on its own 
behalf or on behalf of third parties by imposing licenses in certain circumstances, such as if we fail to achieve 
practical application of the U.S. government funded technology, because action is necessary to alleviate health 
or safety needs, to meet requirements of federal regulations, or to give preference to U.S. industry. In addition, 
U.S. government funded inventions must be reported to the government and U.S. government funding must be 
disclosed in any resulting patent applications. 

As of December 31, 2023, we own or hold exclusive licenses to 437 issued U.S. patents, 89 pending U.S. patent 
applications, 6 pending Patent Cooperation Treaty ("PCT") patent applications, 441 issued foreign patents, and 
166 pending foreign patent applications. The full term of the issued U.S. patents will expire between 2024 and 
2041. We also have non-exclusive patent licenses with various third parties to supplement our own large and 
robust patent portfolio.

Other Sequencing Solutions 

There  are  a  significant  number  of  companies  offering  nucleic  acid  sequencing  equipment  or  consumables. 
These include, but are not limited to, Illumina, Inc. (“Illumina”), BGI Genomics (also known as MGI or Complete 
Genomics),  Thermo  Fisher  Scientific  Inc.  (“Thermo”),  Oxford  Nanopore  Technologies  Ltd.  (“ONT  Ltd.”),  Roche 
Holding  AG  (“Roche”),  Qiagen  N.V.  (“Qiagen”),  Element  Biosciences,  Inc.  (“Element”),  Bionano  Genomics,  Inc. 
(“Bionano”),  Ultima  Genomics,  Inc.  (“Ultima”)  and  Singular  Genomics  Systems,  Inc.  (“Singular”).  These 
companies  may  have  different  levels  of  financial,  technical,  manufacturing,  administrative,  and  support 
resources available to them. We expect the competition to intensify within the overall nucleic acid sequencing 
market  as  there  are  also  several  companies  developing  new  sequencing  technologies,  products  and/or 
services.  Increased  competition  may  result  in  pricing  pressures,  which  could  harm  our  sales,  profitability,  or 
share of supply.

In  order  for  us  to  maintain  and  increase  our  sales,  we  will  need  to  demonstrate  that  our  products  deliver 
superior performance and value as a result of our key differentiators. Our HiFi long-read sequencing will need to 
continue to deliver very high consensus accuracy and long-read lengths and include single-molecule, real-time 
resolution,  with  the  ability  to  detect  real-time  kinetic  information,  fast  time-to-result  and  flexibility,  as  well  as 
support the breadth and depth of current and future applications.

Government Regulation

The  development,  testing,  manufacturing,  marketing,  post-market  surveillance,  distribution,  advertising,  and 
labeling  of  certain  medical  devices,  including  in  vitro  diagnostic  products  and  laboratory-developed  tests,  are 
subject to regulation in the United States by the Center for Devices and Radiological Health of the U.S. Food and 
Drug Administration (FDA) under the Federal Food, Drug, and Cosmetic Act (FDCA) and comparable state and 
foreign  regulatory  agencies.  FDA  defines  a  medical  device  as  an  instrument,  apparatus,  implement,  machine, 
contrivance,  implant,  in  vitro  reagent,  or  other  similar  or  related  article,  including  any  component  part  or 
accessory, which is (i) intended for use in the diagnosis of disease or other conditions, or in the cure, mitigation, 
treatment,  or  prevention  of  disease,  in  man  or  other  animals,  or  (ii)  intended  to  affect  the  structure  or  any 
function of the body of man or other animals and which does not achieve any of its primary intended purposes 
through chemical action within or on the body of man or other animals and which is not dependent upon being 
metabolized for the achievement of any of its primary intended purposes. Medical devices to be commercially 
distributed in the United States must receive from the FDA either clearance of a pre-market notification, known 
as 510(k), or pre-market approval pursuant to the FDC Act prior to marketing, unless subject to an exemption.

We  intend  to  label  and  sell  our  products  for  research  use  only  (“RUO”)  and  expect  to  sell  them  to  research 
customers  in  various  settings,  including  academic  institutions,  life  sciences  and  research  laboratories  that 
conduct  research,  and  biopharmaceutical  and  biotechnology  companies  for  non-diagnostic  and  non-clinical 
purposes. Our current RUO products are not intended or promoted for use in clinical practice in the diagnosis of 
disease  or  other  conditions,  and  they  are  labeled  for  research  use  only,  not  for  use  in  diagnostic  procedures. 
Accordingly,  we  believe  our  products,  as  we  intend  to  market  them,  are  not  subject  to  regulation  by  the  FDA. 
Rather,  while  FDA  regulations  require  that  RUO  products  be  labeled  for  research  use  only  and  to  market  and 
distribute RUO products in accordance with the FDA RUO guidance, the regulations do not subject RUO products 
to  the  FDA’s  jurisdiction  or  the  broader  pre-  and  post-market  controls  for  medical  devices.  However,  in  the 
future,  certain  of  our  products  or  related  applications,  such  as  those  that  may  be  developed  for  clinical  uses, 
could  be  subject  to  FDA  regulation,  or  the  FDA’s  regulatory  jurisdiction  could  be  expanded  to  include  our 
products.  If  we  wish  to  label  and  expand  product  lines  to  address  the  diagnosis  of  disease,  regulation  by 
governmental authorities in the United States and other countries will become an increasingly significant factor 
in development, testing, production, and marketing. In the future, products that we may develop in the molecular 

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diagnostic markets, depending on their intended use, may be regulated as medical devices or in vitro diagnostic 
products (“IVDs”) by the FDA and comparable agencies in other countries. In the U.S., if we market our products 
for use in performing clinical diagnostics, such products would be subject to regulation by the FDA under pre-
market and post-market control as medical devices, unless an exemption applies, and we would be required to 
obtain  either  prior  510(k)  clearance  or  prior  pre-market  approval  from  the  FDA  before  commercializing  the 
product.  Obtaining  the  requisite  regulatory  approvals  can  be  expensive  and  may  involve  considerable  delay. 
Some countries have regulatory review processes that are substantially longer than U.S. processes. Failure to 
obtain  regulatory  approval  in  a  timely  manner  and  meet  all  of  the  local  regulatory  requirements  including 
language and specific safety standards in any foreign country in which we plan to market our products could 
prevent  us  from  marketing  products  in  such  countries  or  subject  us  to  sanctions  and  fines.  Changes  to  the 
current regulatory framework, including the imposition of additional or new regulations, could arise at any time 
during the development or marketing of our products. 

In  November  2013,  the  FDA  issued  a  final  guidance  on  products  labeled  for  research  use  only,  which,  among 
other things, reaffirmed that a company may not make any clinical or diagnostic claims about an RUO product, 
stating  that  merely  including  a  labeling  statement  that  the  product  is  for  research  purposes  only  will  not 
necessarily render the device exempt from the FDA’s clearance, approval, or other regulatory requirements if the 
totality of circumstances surrounding the distribution of the product indicates that the manufacturer knows its 
product  is  being  used  by  customers  for  diagnostic  uses  or  the  manufacturer  intends  such  a  use.  These 
circumstances  may  include,  among  other  things,  written  or  verbal  marketing  claims  regarding  a  product’s 
performance  in  clinical  diagnostic  applications  and  a  manufacturer’s  provision  of  technical  support  for  such 
activities.  If  FDA  were  to  determine,  based  on  the  totality  of  circumstances,  that  our  products  labeled  and 
marketed  for  RUO  are  intended  for  diagnostic  purposes,  they  would  be  considered  medical  devices  that  will 
require clearance or approval prior to commercialization. Further, sales of devices for diagnostic purposes may 
subject  us  to  additional  healthcare  regulation.  We  continue  to  monitor  the  changing  legal  and  regulatory 
landscape to ensure our compliance with any applicable rules, laws and regulations. 

The FDA classifies medical devices into one of three classes. Devices deemed to pose lower risk to the patient 
are placed in either class I or II, which, unless an exemption applies, requires the manufacturer to submit a pre-
market  notification  requesting  FDA  clearance  for  commercial  distribution  pursuant  to  Section  510(k)  of  the 
FDCA. This process, known as 510(k) clearance, requires that the manufacturer demonstrate that the device is 
substantially equivalent to a previously cleared and legally marketed 510(k) device or a “pre-amendment” class 
III  device  for  which  pre-market  approval  applications  (“PMAs”)  have  not  been  required  by  the  FDA.  This  FDA 
review process typically takes from four to twelve months, although it can take longer. Most Class I devices are 
exempted  from  this  510(k)  pre-market  submission  requirement.  If  no  legally  marketed  predicate  can  be 
identified for a new device to enable the use of the 510(k) pathway, the device is automatically classified under 
the  FDCA  as  Class  III,  which  generally  requires  pre-market  approval,  or  PMA  approval.  However,  the  FDA  can 
reclassify  or  use  “de  novo  classification”  for  a  device  that  meets  the  FDCA  standards  for  a  Class  II  device, 
permitting  the  device  to  be  marketed  without  PMA  approval.  To  grant  such  a  reclassification,  FDA  must 
determine  that  the  FDCA’s  general  controls  alone,  or  general  controls  and  special  controls  together,  are 
sufficient  to  provide  a  reasonable  assurance  of  the  device’s  safety  and  effectiveness.  The  de  novo 
classification route is generally less burdensome than the PMA approval process.

Devices  deemed  by  the  FDA  to  pose  the  greatest  risk,  such  as  life-sustaining,  life-supporting,  or  implantable 
devices, or those deemed not substantially equivalent to a legally marketed predicate device, are placed in class 
III. Class III devices typically require PMA approval. To obtain PMA approval, an applicant must demonstrate the 
reasonable safety and effectiveness of the device based, in part, on data obtained in clinical studies. All clinical 
studies  of  investigational  medical  devices  to  determine  safety  and  effectiveness  must  be  conducted  in 
accordance  with  FDA’s  investigational  device  exemption  (“IDE”)  regulations,  including  the  requirement  for  the 
study  sponsor  to  submit  an  IDE  application  to  FDA,  unless  exempt,  which  must  become  effective  prior  to 
commencing human clinical studies. PMA reviews generally last between one and two years, although they can 
take  longer.  Both  the  510(k)  and  the  PMA  processes  can  be  expensive  and  lengthy  and  may  not  result  in 
clearance or approval. If we are required to submit our products for pre-market review by the FDA, we may be 
required to delay marketing and commercialization while we obtain pre-market clearance or approval from the 
FDA. There would be no assurance that we could ever obtain such clearance or approval.

All  medical  devices,  including  IVDs,  that  are  regulated  by  the  FDA  are  also  subject  to  the  quality  system 
regulation. Obtaining the requisite regulatory approvals, including the FDA quality system inspections that are 
required  for  PMA  approval,  can  be  expensive  and  may  involve  considerable  delay.  The  regulatory  approval 
process for such products may be significantly delayed, may be significantly more expensive than anticipated, 

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and  may  conclude  without  such  products  being  approved  by  the  FDA.  Without  timely  regulatory  approval,  we 
will  not  be  able  to  launch  or  successfully  commercialize  such  diagnostic  products.  Changes  to  the  current 
regulatory framework, including the imposition of additional or new regulations, could arise at any time during 
the development or marketing of our products. This may negatively affect our ability to obtain or maintain FDA 
or  comparable  regulatory  clearance  or  approval  of  our  products  in  the  future.  In  addition,  regulatory  agencies 
may introduce new requirements that may change the regulatory requirements for us or our customers, or both.

As noted above, although our products are currently labeled and sold for research purposes only, the regulatory 
requirements related to marketing, selling, and supporting such products could be uncertain and depend on the 
totality of circumstances. This uncertainty exists even if such use by our customers occurs without our consent. 
If the FDA or other regulatory authorities assert that any of our RUO products are subject to regulatory clearance 
or approval, our business, financial condition, or results of operations could be adversely affected.

For  example,  in  some  cases,  our  customers,  including  laboratories  that  offer  services  as  part  of  our  certified 
service provider program, may use our RUO products in their own laboratory-developed tests (“LDTs”) or in other 
FDA-regulated products for clinical diagnostic use. The FDA has historically exercised enforcement discretion in 
not  enforcing  the  medical  device  regulations  against  LDTs  and  LDT  manufacturers.  However,  on  October  3, 
2014, the FDA issued two draft guidance documents that set forth the FDA’s proposed risk-based framework for 
regulating LDTs, which are designed, manufactured, and used within a single laboratory. In January 2017, the 
FDA  announced  that  it  would  not  issue  final  guidance  on  the  oversight  of  LDTs  and  LDT  manufacturers  but 
would seek further public discussion on an appropriate oversight approach and give Congress an opportunity to 
develop  a  legislative  solution.  The  FDA  has  issued  warning  letters  to  genomics  labs  for  illegally  marketing 
genetic  tests  that  claim  to  predict  patients’  responses  to  specific  medications,  noting  that  the  FDA  has  not 
created  a  legal  “carve-out”  for  LDTs  and  retains  discretion  to  take  action  when  appropriate,  such  as  when 
certain genomic tests raise significant public health concerns. As laboratories and manufacturers develop more 
complex  genetic  tests  and  diagnostic  software,  the  FDA  may  increase  its  regulation  of  LDTs.  Legislative  and 
administrative proposals to amend the FDA's oversight of LDTs have been introduced in recent years, including 
the  Verifying  Accurate  Leading-edge  IVCT  Development  Act  of  2021  (the  “VALID  Act”).  In  September  2022, 
Congress passed the FDA user fee reauthorization legislation without substantive FDA policy riders, including 
the  VALID  Act,  but  Congress  may  revisit  the  policy  riders  and  enact  other  FDA  programmatic  reforms  in  the 
future. In October 2023, through rulemaking, the FDA proposed to amend the definition of “in vitro diagnostic 
products” in FDA regulations to include laboratories that manufacture such products and to phase out the FDA’s 
general  enforcement  discretion  approach  for  LDTs  so  that  IVDs  manufactured  by  a  laboratory  would  be 
regulated  similarly  to  IVDs.  It  is  unclear  when  the  FDA  will  finalize  and  begin  enforcing  such  rule.  Any  future 
legislative or administrative rule making or oversight of LDTs and LDT manufacturers, if and when finalized, may 
impact  the  sales  of  our  products  and  how  customers  use  our  products,  and  may  require  us  to  change  our 
business model in order to maintain compliance with these laws. We would become subject to additional FDA 
requirements if our products are determined to be medical devices or if we elect to seek 510(k) clearance or 
pre-market approval. If our products become subject to FDA regulation as medical devices, we would need to 
invest  significant  time  and  resources  to  ensure  ongoing  compliance  with  FDA  quality  system  regulations  and 
other post-market regulatory requirements.

If our products become subject to FDA regulation as medical devices, the regulatory clearance or approval and 
the maintenance of continued and post-market regulatory compliance for such products will be expensive, time-
consuming, and uncertain both in timing and in outcome. Commercialization of such regulated medical devices 
can  increase  our  exposure  under  additional  laws.  For  example,  medical  device  companies  are  subject  to 
additional healthcare regulation and enforcement by the federal government and by authorities in the states and 
foreign  jurisdictions  in  which  they  conduct  their  business  and  may  constrain  the  financial  arrangements  and 
relationships through which we research, as well as sell, market and distribute any medical products for which 
we obtain marketing authorization. Such laws include, without limitation, state and federal anti-kickback, fraud 
and abuse, false claims, data privacy and security, and transparency laws and regulations related to payments 
and other transfers of value made to physicians and other healthcare providers. If our operations are found to 
be  in  violation  of  any  of  such  laws  or  any  other  governmental  regulations  that  apply,  we  may  be  subject  to 
penalties, 
limitation,  administrative,  civil,  and  criminal  penalties,  damages,  fines, 
disgorgement,  the  curtailment  or  restructuring  of  operations,  integrity  oversight  and  reporting  obligations, 
exclusion from participation in federal and state healthcare programs and imprisonment.

including,  without 

In the future, to the extent we develop any clinical diagnostic assays, we may pursue payment for such products 
through  a  diverse  and  broad  range  of  channels  and  seek  coverage  and  reimbursement  by  government  health 
insurance  programs  and  commercial  third-party  payors  for  such  products.  In  the  United  States,  there  is  no 

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uniform coverage for clinical laboratory tests. The extent of coverage and rate of payment for covered services 
or items vary from payor to payor. Obtaining coverage and reimbursement for such products can be uncertain, 
time-consuming,  and  expensive,  and,  even  if  favorable  coverage  and  reimbursement  status  were  attained  for 
our  tests,  to  the  extent  applicable,  less  favorable  coverage  policies  and  reimbursement  rates  may  be 
implemented in the future. Changes in healthcare regulatory policies could also increase our costs and subject 
us  to  additional  regulatory  requirements  that  may  interrupt  commercialization  of  our  products,  decrease  our 
revenue and adversely impact sales of, and pricing of and reimbursement for, our products.

International sales of medical devices are subject to foreign government regulations, which vary substantially 
from country to country. In the future, if we decide to distribute or market our diagnostic products as IVDs in 
Europe, such products are subject to regulation under the European Union (“EU”) IVD Medical Device Regulation 
(“IVDR”) EU 2017/746. Outside of the EU, regulatory approval needs to be sought on a country-by-country basis 
in  order  to  market  medical  devices.  Although  there  is  a  trend  towards  harmonization  of  a  quality  system, 
standards and regulations in each country may vary substantially, which can affect timelines of introduction.

We are committed to the protection of our employees and the environment. Our operations require the use of 
hazardous  materials  that  subject  us  to  various  federal,  state,  and  local  environmental  and  safety  laws  and 
regulations.  We  believe  that  we  are  in  material  compliance  with  current  applicable  laws  and  regulations. 
However, we could be held liable for damages and fines should contamination of the environment or individual 
exposures  to  hazardous  substances  occur.  In  addition,  we  cannot  predict  how  changes  in  these  laws  and 
regulations, or the development of new laws and regulations, will affect our business operations or the cost of 
compliance.

Additionally,  we  must  comply  with  complex  foreign  and  U.S.  laws  and  regulations,  such  as  the  U.S.  Foreign 
Corrupt Practices Act, the U.K. Bribery Act, and other local laws prohibiting corrupt payments to governmental 
officials, anti-competition regulations and sanctions imposed by the U.S. Office of Foreign Assets Control, and 
other similar laws and regulations. Violations of these laws and regulations could result in fines and penalties, 
criminal sanctions, restrictions on our business conduct, and on our ability to offer our products in one or more 
countries,  and  could  also  materially  affect  our  brand,  our  ability  to  attract  and  retain  employees,  our 
international operations, our business, and our operating results. Although we have implemented policies and 
procedures designed to ensure compliance with these laws and regulations, there can be no assurance that our 
employees, contractors, or agents will not violate our policies.

As  we  continue  to  expand  our  business  into  multiple  international  markets,  our  success  will  depend,  in  large 
part, on our ability to anticipate and effectively manage these and other risks associated with our international 
operations.  Any  of  these  risks  could  harm  our  international  operations  and  negatively  impact  our  sales, 
adversely affecting our business, results of operations, financial condition, and growth prospects.

Human Capital 

As  of  December  31,  2023,  we  had  796  full-time  employees.  Of  these  employees,  317  were  in  research  and 
development,  91  were  in  operations,  45  were  in  service,  226  were  in  marketing,  sales,  and  customer  support, 
and 117 were in general and administration. With the exception of our field-based sales, marketing, and service 
teams, the majority of our employees are in California. None of our employees are represented by labor unions 
or are covered by a collective bargaining agreement with respect to their employment. We have not experienced 
any work stoppages, and we consider our relationship with our employees to be good. 

Talent Acquisition and Retention

We recognize that our employees largely contribute to our success. To this end, we support business growth by 
seeking  to  attract  and  retain  best-in-class  talent.  Our  talent  acquisition  team  uses  internal  and  external 
resources to recruit highly skilled candidates globally. 

Total Rewards

Our total rewards philosophy has been to invest in our workforce by offering competitive and fair compensation 
and  benefits  packages.  We  provide  employees  with  compensation  packages  that  include  base  salary,  short-
term  incentives  such  as  annual  bonuses  and  commissions,  and  long-term  equity  awards.  We  also  offer 
comprehensive  employee  benefits,  which  vary  by  country  and  region,  such  as  life,  disability,  and  health 
insurance, health savings and flexible spending accounts, time off benefits, paid parental leave, Employee Stock 
Purchase Program, and a 401(k) plan. It is our expressed intent to be an employer of choice in our industry by 
providing market-competitive compensation and benefits packages.

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Health, Safety, and Wellness

The  health,  safety,  and  wellness  of  our  employees  is  a  priority  in  which  we  have  always  invested  and  will 
continue to do so. We provide our employees and their families with access to a variety of innovative, flexible, 
and  convenient  health  and  wellness  programs.  Program  benefits  are  intended  to  provide  protection  and 
security, so employees can have peace of mind concerning events that may require time away from work or that 
may  impact  their  financial  well-being.  These  programs  are  highlighted  and  updated  regularly  on  our  internal 
benefits platform. 

Diversity, Equity, and Inclusion

We  believe  a  diverse  workforce  is  critical  to  our  success.  Our  mission  is  to  value  differences  in  races, 
ethnicities,  religions,  nationalities,  genders,  ages,  sexual  orientations,  as  well  as  education,  skill  sets  and 
experience.  We  offer  training  programs  on  diversity  awareness  to  help  employees  understand,  recognize, 
respond, and prevent bias throughout the employee lifecycle. We are focused on inclusive hiring practices, fair 
and equitable treatment, organizational flexibility, and training and resources.

Training and Development

We  believe  in  encouraging  employees  to  becoming  lifelong  learners  by  providing  ongoing  learning  and 
leadership  training  opportunities.  We  provide  a  scaled  learning  platform  of  on-demand  and  virtual  classroom 
learning focused on personal and professional development. While we strive to provide real-time recognition of 
employee  performance,  we  have  a  formal  annual  review  process  not  only  to  determine  pay  and  equity 
adjustments  tied  to  individual  contributions,  but  to  identify  areas  where  training  and  development  may  be 
needed.

Available Information 

Our  website  is  located  at  www.pacb.com.  The  information  posted  on,  or  that  can  be  accessed  through,  our 
website is not incorporated by reference into this Annual Report on Form 10-K, and the inclusion of our website 
address is an inactive textual reference only. Our Annual Report on Form 10-K, Quarterly Reports on Form 10-Q, 
Current  Reports  on  Form  8-K,  and  amendments  to  reports  filed  or  furnished  pursuant  to  Sections  13(a)  and 
15(d)  of  the  Securities  Exchange  Act  of  1934,  as  amended,  are  available  free  of  charge  through  the  “Investor 
Relations”  section  of  our  website  as  soon  as  reasonably  practicable  after  we  electronically  file  such  material 
with, or furnish it to, the SEC. The SEC also maintains a website that contains our SEC filings. The address of the 
site is www.sec.gov.

Additionally, we use our website (including the blog section of our website) as well as our X (formerly Twitter) 
account  (@pacbio)  as  a  channel  of  distribution  for  important  company  information  and  to  comply  with  our 
disclosure  obligations  under  Regulation  FD. 
including  press  releases,  analyst 
Important 
presentations, and financial information regarding us, as well as corporate governance information, is routinely 
posted and accessible on the “Investor Relations” section of the website, which is accessible by clicking on the 
tab  labeled  “Company  -  Investors”  on  our  website  home  page.  In  addition,  important  information  is  routinely 
posted  and  accessible  on  the  blog  section  of  our  website,  which  is  accessible  through  our  website  at 
www.pacb.com/blog, as well as our X account (@pacbio). The contents of our website and our X account are 
not incorporated by reference into this Annual Report on Form 10-K or in any other report or document we file 
with the SEC, and any references to our website or X account are intended to be inactive textual references only.

information, 

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ITEM 1A. 

RISK FACTORS

You  should  carefully  consider  the  risks  and  uncertainties  described  below,  together  with  all  of  the  other 
information  in  our  public  filings  with  the  SEC,  which  could  materially  affect  our  business,  financial  condition, 
results  of  operations  and  prospects.  The  risks  described  below  are  not  the  only  risks  facing  us.  Risks  and 
uncertainties not currently known to us or that we currently deem to be immaterial also may materially affect our 
business,  financial  condition,  results  of  operations  and  prospects.  In  addition,  any  worsening  of  the  economic 
environment  may  exacerbate  the  risks  described  below,  any  of  which  could  have  a  material  impact  on  us.  This 
situation is changing rapidly, and additional impacts may arise that we are not aware of currently.

Summary Risk Factors

The  following  is  a  summary  of  the  principal  risks  that  could  adversely  affect  our  business,  operations,  and 
financial results. Such risks are discussed more fully below and include, but are not limited to, risks related to:

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our  ability  to  successfully  market,  commercialize,  and  sell  current  and  future  products  and  related 
maintenance services;

our ability to achieve profitability for our business;

our ability to repay our debt and fund our long-term operations;

our ability to successfully leverage and integrate our acquisitions and future acquisitions;

our ability to successfully research, develop and timely manufacture our current and future products;

• management of new product introductions and transitions, resultant costs, and ability of new products 

to generate promised performance;

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recent significant changes to our leadership team and resultant disruptions to our business;

retention, recruitment, and training of senior management, key personnel, scientists and engineers;

our ability to further penetrate nucleic acid sequencing applications, as well as grow product demand;

our  reliance  on  outsourcing  to  other  companies  for  manufacturing  certain  components  and  sub-
assemblies, some of which are sole-sourced;

our  ability  to  consistently  manufacture  our  instruments  and  consumables  to  meet  customers’ 
specifications, quantity, cost, or performance requirements;

the high amount of competition we face in our industry;

our ability to attract customers and increase sales of current and future products;

reliance on a limited number of customers for a significant portion of our revenues, including academic, 
research and government institutions;

the complexity of our products giving rise to defects or errors;

our unpredictable and lengthy sales cycles;

adverse  effects  resulting  from  political  and  economic  tensions  between  the  United  States  and  other 
countries, including China and Russia, and other geopolitical uncertainties;

securing  and  maintaining  patent  or  other  intellectual  property  protection  for  our  products  and  related 
improvements;

current and future legal proceedings filed against us claiming intellectual property infringement;

the potential adverse impact of health epidemics, including any resurgence of COVID-19 cases or other 
similar outbreaks;

governmental regulations that burden operations or narrow the market for our products;

evolving ethical, legal, privacy, social, and regulatory concerns regarding genetic testing;

volatility of the price of our common stock; and

our stock price falling as a result of future offerings or sales of securities.

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Our  risk  factors  are  not  guarantees  that  no  such  conditions  exist  as  of  the  date  hereof  and  should  not  be 
interpreted as an affirmative statement that such risks or conditions have not materialized, in whole or in part.

Risks Related to Our Business

The commercialization and sales of our current or future products may be unsuccessful or less successful than 
anticipated.  While  we  plan  to  continue  pursuing  new  products  and  expand  into  adjacent  markets,  we  have 
limited  experience  in  managing  and  selling  multiple  products  and,  as  a  result,  may  face  challenges  selling  in 
new  markets  and  fail  to  successfully  carry  out  these  initiatives,  which  may  adversely  impact  our  business, 
financial condition or results of operation.

We have made and expect to continue making substantial investments to develop new products and enhance 
our  existing  products  through  our  acquisitions  and  research  and  development  efforts.  For  example,  we 
commenced commercial shipments of Revio, our new long-read sequencing system in the first quarter of 2023, 
and commenced commercial shipments of Onso, our new SBB short-read platform, in the third quarter of 2023. 
Our  future  success  is  substantially  dependent  on  our  ability  to  successfully  develop  and  commercialize  our 
products,  including  Revio  and  Onso,  as  well  as  acquired  technologies,  which  are  anticipated  to  be  used  in 
demanding  scientific  research  that  requires  substantial  levels  of  accuracy  and  precision.  In  addition,  we  may 
not be successful in transitioning our prior generation products to our Revio product, or transitioning users of 
other third party short-read sequencing platforms to Onso, and could incur related obsolete inventory charges 
and losses on firm purchase commitments. Customers may also be slower than we anticipate in making new 
capital  equipment  acquisitions,  especially  in  the  current  economic  environment.  Due  to  challenges  we  may 
experience in developing and marketing our existing products and launching new products, we may not be able 
to effectively:

• manage  the  timeliness  of  our  new  product  introductions  and  the  rate  at  which  sales  of  our  new 
products  may  cannibalize  sales  of  our  older  products  or  manage  sales  and  marketing  of  multiple 
sequencing platforms;

•

drive adoption of our current and future products, including the Sequel II/IIe, Revio and Onso systems, 
and products under development;

• maintain our competitive position by continuing to attract and retain customers for our products;

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provide appropriate levels of customer training and support for our products;

implement an effective marketing strategy to promote awareness of our products;

develop and implement an effective sales and distribution strategy for our current and future products;

develop,  manufacture  and  commercialize  new  products  or  achieve  an  acceptable  return  on  our 
manufacturing or research and development efforts and expenses;

comply with regulatory requirements applicable to our products;

anticipate and adapt to changes in our market;

accommodate  customer  expectations  and  demands  with  respect  to  our  products,  increase  product 
adoption by our existing customers or develop new customer relationships;

deliver  our  beta  systems  to  our  external  beta  testing  sites  or  complete  our  external  beta  testing 
program on our currently expected timelines;

overcome unexpected challenges discovered during beta testing;

complete the scientific and technical validation of new products on our currently expected timeline or at 
all;

deliver our future products in a timely manner to our customers;

grow our market share by marketing and selling our products for new and additional applications;

• manage  the  significant  burdens  that  expanding  our  existing  or  future  products  into  current  and  new 
markets may impose on marketing, compliance, and other administrative and managerial resources;

• maintain and develop strategic relationships with vendors, manufacturers, and other industry partners 
to acquire necessary materials for the production of, and to develop, manufacture and commercialize, 
our existing or future products;

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adapt or scale our manufacturing activities to meet performance specifications and potential demand 
at a reasonable cost;

avoid infringement and misappropriation of third-party intellectual property;

obtain  and  maintain  any  necessary  licenses  to  third-party  intellectual  property  on  commercially 
reasonable terms;

obtain valid and enforceable patents that give us a competitive advantage or enforce existing patents;

protect our proprietary technology; and

attract, retain, and motivate qualified personnel.

The risks noted above, especially with respect to the marketing, sales, and commercialization of our products, 
may  be  heightened  by  the  impact  of  current  uncertain  market  and  other  conditions.  In  addition,  a  high 
percentage of our expenses is and will continue to be fixed. Accordingly, if we do not generate revenue as and 
when  anticipated,  we  could  suffer  a  material  adverse  effect  on  our  business,  financial  conditions,  results  of 
operations and prospects.

We  have  incurred  losses  to  date,  and  we  expect  to  continue  to  incur  significant  losses  as  we  develop  our 
business and may never achieve profitability.

We have generally incurred net losses each quarter since inception, and we cannot be certain if or when we will 
produce  sufficient  revenue  from  our  operations  to  support  our  costs.  Even  if  profitability  is  achieved  in  the 
future,  we  may  not  be  able  to  sustain  profitability  on  a  consistent  basis.  We  expect  to  continue  to  incur 
substantial losses and negative cash flow from operations for the foreseeable future.

Our net losses since inception and our expectation of incurring substantial losses and negative cash flow for 
the foreseeable future could:

• make it more difficult for us to satisfy our obligations;

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increase our vulnerability to general adverse economic and industry conditions;

limit our ability to fund future working capital, capital expenditures, research and development and other 
business opportunities;

increase the volatility of the price of our common stock;

limit our flexibility to react to changes in our business and the industry in which we operate;

place  us  at  a  disadvantage  to  other  companies  that  offer  nucleic  acid  sequencing  equipment  or 
consumables; and

limit our ability to borrow additional funds.

In addition, inflationary pressure, including as a result of supply shortages, has adversely impacted and could 
continue  to  adversely  impact  our  financial  results,  and  our  operating  costs  may  increase.  We  may  not  fully 
offset  these  cost  increases  by  raising  prices  for  our  products  and  services,  which  could  result  in  downward 
pressure on our margins. Further, our customers may choose to reduce their business with us if we increase our 
pricing.

Any or all of the foregoing may have a material adverse effect on our business, operations, financial condition, 
and prospects. An impairment in value of our tangible or intangible assets could also be recorded as a result of 
weaker economic conditions.

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We are not cash flow positive and may not have sufficient cash to make required payments under the terms of 
our debt or fund our long-term planned operations.

Our operations have consumed substantial amounts of cash since inception, and we expect to continue to incur 
substantial losses and negative cash flow from operations for the foreseeable future. Additional funds may not 
be available on terms acceptable to us or at all. We have incurred significant debt, and we may incur additional 
debt,  in  the  future.  As  of  December  31,  2023,  we  had  outstanding  approximately  $459.0  million  aggregate 
principal amount of 1.50% convertible senior notes due 2028 (the “2028 Notes”) and $441.0 million aggregate 
principal  amount  of  our  1.375%  convertible  senior  notes  due  2030  (the  “2030  Notes”  and,  together  with  the 
2028 Notes, the “Notes”). We may not have sufficient cash to make required payments under the terms of this 
debt,  and  should  this  occur,  debt  holders  have  rights  senior  to  common  stockholders  to  make  claims  on  our 
assets.  We  may  not  be  able  to  issue  equity  securities  due  to  unacceptable  terms  and  conditions  to  us  in  the 
capital markets. To the extent that we intend to raise additional funds through the sale of our common stock, 
downward  fluctuations  in  our  stock  price  could  adversely  affect  such  fundraising  efforts.  Furthermore,  equity 
financings normally involve shares sold at a discount to the current market price and fundraising through sales 
of  additional  shares  of  common  stock  or  other  equity  securities  will  have  a  dilutive  effect  on  our  existing 
investors. We may be required to seek equity financing at a time when the market price for our common stock is 
low, which will further contribute to dilution for existing holders.

We  believe  that  our  growth  will  depend,  in  part,  on  our  ability  to  fund  our  commercialization  efforts  and  our 
efforts  to  develop  new  products,  including  any  improvements  to  our  existing  products.  To  the  extent  our 
existing  resources  are  not  sufficient,  it  may  require  us  to  delay,  or  even  not  allow  us  to  conduct  any  or  all  of 
these activities that we believe would be beneficial for our future growth. We may need to raise additional funds 
through  public  or  private  debt  or  equity  financing  or  alternative  financing  arrangements,  which  may  include 
collaborations or licensing arrangements. If we are unable to raise funds on favorable terms, or at all, we may 
have to reduce our cash burn rate and may not be able to support our commercialization efforts, launching of 
new products, or operations, or to increase or maintain the level of our research and development activities.

If  we  are  unable  to  generate  sufficient  cash  flows  or  to  raise  adequate  funds  to  finance  our  forecasted 
expenditures,  we  may  have  to  make  significant  changes  to  our  operations,  including  delaying  or  reducing  the 
scope  of,  or  eliminating  some  or  all  of,  our  development  programs.  We  also  may  have  to  reduce  sales, 
marketing,  engineering,  customer  support  or  other  resources  devoted  to  our  existing  or  new  products,  or  we 
may need to cease operations. Any of these actions could materially impede our ability to achieve our business 
objectives and could materially harm our operating results. If our cash, cash equivalents and investments are 
insufficient  to  fund  our  projected  operating  requirements  and  we  are  unable  to  raise  capital,  it  could  have  a 
material adverse effect on our business, financial condition and results of operations and prospects.

We have made acquisitions and, in the future, may continue to acquire businesses, technologies or assets, form 
joint  ventures  or  make  other  strategic  investments  with  companies  that  could  adversely  affect  our  operating 
results, dilute our stockholders’ ownership, or cause us to incur debt or significant expense.

As  part  of  our  business  strategy,  we  have  acquired  and  expect  to  continue  to  pursue  acquisitions  of 
complementary  businesses,  technologies,  or  assets.  We  may  also  pursue  technology  license  arrangements, 
strategic alliances or investments that complement our business as we have previously with our acquisitions of 
Circulomics, Omniome and Apton in July 2021, September 2021 and August 2023, respectively.

Acquisitions  and  strategic  transactions  involve  numerous  risks,  any  of  which  could  harm  our  business  and 
negatively affect our financial condition and results of operations, including:

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intense  competition  for  suitable  acquisition  targets,  which  could  increase  prices  and  adversely  affect 
our ability to consummate deals on favorable or acceptable terms;

failure or material delay in closing a transaction;

transaction-related lawsuits or claims;

difficulties in integrating the technologies, operations, existing contracts, and personnel of an acquired 
company;

difficulties in retaining key employees or business partners of an acquired company;

difficulties in retaining suppliers, partners, or customers of an acquired company;

challenges with integrating the brand identity of an acquired company with our own;

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diversion  of  financial  and  management  resources  from  existing  operations  or  alternative  acquisition 
opportunities;

failure to realize the anticipated benefits or synergies of a transaction;

difficulties in developing technology post-acquisition;

failure to identify the problems, liabilities, or other shortcomings or challenges of an acquired company 
or  technology,  including  issues  related  to  intellectual  property,  regulatory  compliance  practices, 
litigation, revenue recognition or other accounting practices, or employee or user issues;

risks that regulatory bodies may enact new laws or promulgate new regulations that are adverse to an 
acquired company or business;

risks  that  regulatory  bodies  do  not  approve  our  acquisitions  or  business  combinations  or  delay  such 
approvals;

theft of our trade secrets or confidential information that we share with potential acquisition candidates 
or other potential strategic partners;

risk  that  an  acquired  company  or  investment  in  new  services  cannibalizes  a  portion  of  our  existing 
business; and

adverse market reaction to an acquisition or other strategic transaction.

To  finance  any  acquisitions  or  other  strategic  investments,  we  may  raise  additional  funds,  which  could 
adversely affect our existing stockholders and our business. If the price of our common stock is low or volatile, 
we  may  not  be  able  to  acquire  other  companies  for  stock.  In  addition,  our  stockholders  may  experience 
substantial  dilution  as  a  result  of  additional  securities  we  may  issue  for  acquisitions.  Open  market  sales  of 
substantial amounts of our common stock issued to stockholders of companies we acquire could also depress 
our stock price. Additional funds may not be available on terms that are favorable to us, or at all.

If  we  fail  to  address  the  foregoing  risks  or  other  problems  encountered  in  connection  with  past  or  future 
acquisitions of businesses, new technologies, services, and other assets and strategic investments, or if we fail 
to  successfully  integrate  such  acquisitions  or  investments,  our  business,  financial  condition,  and  results  of 
operations could be adversely affected, including potential impairments of goodwill and intangible assets.

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If we are unable to successfully develop and timely manufacture our current and future products, including with 
respect to SMRT Cells, Sequel II/IIe Systems, Revio, Onso, and other SMRT Cell, HiFi, and SBB products under 
development, and related products, our business may be adversely affected.

In light of the highly complex technologies involved in our products, there can be no assurance that we will be 
able to manufacture and commercialize our current and future products on a timely basis or continue providing 
adequate  support  for  our  existing  products.  The  commercial  success  of  our  products,  including  the  Sequel, 
Sequel  II/IIe,  Revio  and  Onso  Systems,  and  the  products  under  development,  including  acquired  technologies, 
depends  on  a  number  of  factors,  including  performance  and  reliability  of  the  systems,  our  anticipating  and 
effectively  addressing  customer  preferences  and  demands,  the  success  of  our  sales  and  marketing  efforts, 
effective  forecasting  and  management  of  product  demand,  purchase  commitments  and  inventory  levels, 
effective  management  of  manufacturing  and  supply  costs,  and  the  quality  of  our  products,  including 
consumables  such  as  SMRT  Cells  and  reagents.  Should  we  face  delays  in  or  discover  unexpected  defects 
during  the  further  development  or  manufacturing  process  of  instruments  or  consumables  related  to  our 
products,  including  with  respect  to  SMRT  Cells,  reagents,  Sequel  II/IIe  Systems,  Revio,  Onso,  and  other  SMRT 
Cell,  HiFi,  and  SBB  products  under  development,  including  acquired  technologies,  and  including  any  delays  or 
defects in software development or product functionality, the timing and success of the continued rollout and 
scaling of our products may be significantly impacted, which may materially and negatively impact our revenue 
and  gross  margin.  The  ability  of  our  customers  to  successfully  utilize  our  products  will  also  depend  on  our 
ability to deliver high quality SMRT Cells and reagents. We have designed SMRT Cells and other consumables 
specifically  for  the  Sequel,  Sequel  II/IIe,  and  Revio  Systems,  and  may  need  to  develop  in  the  future,  other 
customized  SMRT  Cells  and  consumables  for  our  future  products.  Our  production  of  the  SMRT  Cells  for  the 
Sequel and Sequel II/IIe Systems has been and may in the future, including with respect to the Revio system, be 
below  desired  levels  and  yields,  and  we  have  experienced  and  may  experience  in  the  future  manufacturing 
delays, product or quality defects, SMRT Cell variability, and other issues. For example, the COVID–19 pandemic 
has impacted and, any resurgence of COVID-19, or the outbreak of other similar health epidemics could result in 
more  pronounced  impacts  to  our  manufacturing  and  our  ability  to  supply  products.  The  performance  of  our 
consumables  is  critical  to  our  customers’  successful  utilization  of  our  products,  and  any  defects  or 
performance issues with our consumables would adversely affect our business. All of the foregoing could have 
a  material  adverse  effect  on  our  ability  to  sell  our  products  or  result  in  other  material  adverse  effects  on  our 
business, operations, financial condition, operations and prospects.

The development of our products is complex and costly. Problems in the design or quality of our products may 
have a material and adverse effect on our brand, business, financial condition, and operating results, and could 
result in us losing our certifications from the International Organization for Standardization (“ISO”). If we were to 
lose  ISO  certification,  then  our  customers  might  choose  not  to  purchase  products  from  us  and  this  could 
adversely  impact  our  ability  to  develop  products  approved  for  clinical  uses.  Unanticipated  problems  with  our 
products  could  divert  substantial  resources,  which  may  impair  our  ability  to  support  our  new  and  existing 
products and could substantially increase our costs. If we encounter development challenges or discover errors 
in  our  products  late  in  our  development  cycle,  including  during  external  beta  testing,  we  may  be  forced  to 
undertake  design  and/or  production  changes,  delay  product  shipments  or  the  scaling  of  manufacturing  or 
supply. The completion of the production and external testing of our beta systems may also take longer than 
currently  planned,  cost  more  than  currently  expected  and  the  scientific  and  technical  validation  may  not  be 
completed on our currently expected timelines or at all. Such testing may also expose fundamental flaws in our 
products that may cause us to abandon the further development of such products.

If the continued rollout of our current and future products, including with respect to the SMRT Cell, the Sequel II/
IIe, Revio, and Onso Systems, is delayed or is not successful or less successful than anticipated, then we may 
not be able to achieve an acceptable return, if any, on our substantial research and development efforts, and our 
business  may  be  materially  and  adversely  affected.  The  expenses  or  losses  associated  with  delayed  or 
unsuccessful  product  development  or  lack  of  market  acceptance  of  our  existing  and  new  products,  including 
the  SMRT  Cell,  Sequel  II/IIe  Systems,  Revio,  and  Onso,  could  materially  and  adversely  affect  our  business, 
operations, financial condition, and prospects.

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Our  research  and  development  efforts  may  not  result  in  the  benefits  that  we  anticipate,  and  our  failure  to 
successfully  market,  sell,  and  commercialize  our  current  and  future  products  could  have  a  material  adverse 
effect on our business, financial condition and results of operations.

We have dedicated significant resources to developing our current products, including sequencing systems and 
consumables based on our proprietary SMRT sequencing technology and our Sequel and Sequel II/IIe Systems. 
We are also engaged in substantial and complex research and development efforts, which, if successful, may 
result in the introduction of new products in the future, including in connection with the SMRT Cell, the Sequel II/
IIe  Systems,  Revio  and  Onso,  in  addition  to  other  products  currently  under  development,  including  acquired 
technologies. Our research and development efforts are complex and require us to incur substantial expenses 
and we may not be able to develop, manufacture and commercialize new products or obtain regulatory approval 
if necessary. We may divert significant resources to research and development initiatives that do not result in 
commercialized  products,  and  even  if  these  efforts  do  result  in  commercialized  products,  there  can  be  no 
assurance that such products will compete successfully in the market or achieve an acceptable return, if any, on 
our research and development efforts and expenses. Moreover, our joint research and development efforts with 
partners require significant management attention and operational resources. If we are unable to successfully 
manage  such  joint  research  and  development  efforts,  our  future  results  may  be  adversely  impacted. 
Furthermore,  we  will  need  to  continue  to  expand  our  internal  capabilities  or  seek  new  partnerships  or 
collaborations, or both, in order to successfully develop, market, sell and commercialize our products for and in 
the markets we seek to reach. If we are unable to do so or are delayed, then this could materially and adversely 
affect our business, operations, financial condition, and prospects.

We  must  successfully  manage  new  product  introductions  and  transitions,  including  with  respect  to  the  Revio 
and Onso Systems and each of their related consumables, and the development of acquired technologies, and 
we may incur significant costs during these transitions and development, and these efforts may not result in the 
benefits we anticipate.

If our products and services fail to deliver the performance, scalability or results expected by our current and 
future  customers,  or  are  not  delivered  on  a  timely  basis,  our  reputation  and  credibility  may  suffer,  our  current 
and future sales and revenue may be materially harmed and our business may not succeed. For instance, if we 
are not able to successfully execute on the commercialization of the Revio HiFi long-read sequencing system, 
and  the  Onso  SBB  short-read  sequencing  system,  and  each  of  their  related  consumables,  and  any  future 
products  that  may  be  developed  for  research,  medical  and  clinical  uses,  including  acquired  technologies,  it 
could have a material adverse effect on our business, financial condition and results of operations. In addition, 
the  introduction  of  future  products,  including  with  respect  to  future  long-read  and  short-read  products,  and 
related  consumables,  has  and  may  in  the  future  lead  to  our  limiting  or  ceasing  development  of  further 
enhancements to our existing products as we focus our resources on new products, and has resulted and could 
in  the  future  result  in  reduced  marketplace  acceptance  and  loss  of  sales  of  our  existing  products,  materially 
adversely  affecting  our  revenue  and  operating  results.  The  introduction  of  new  products,  including  the  recent 
commercialization of our Revio and Onso systems, has had and may in the future also have a negative impact 
on our revenue in the near-term as our current and future customers have delayed or cancelled and may in the 
future delay or cancel orders of existing products in anticipation of new products and we may also be pressured 
to decrease prices for our existing products. Our experience in managing product transitions is limited, and we 
have experienced, and may in the future experience, difficulty in managing or forecasting customer reactions, 
purchasing decisions or transition requirements with respect to newly launched products. We have incurred and 
may  continue  to  incur  significant  costs  in  completing  these  transitions,  including  costs  of  write-downs  of  our 
products,  as  current  or  future  customers  transition  to  new  products.  If  we  do  not  successfully  manage  these 
product  transitions,  including  with  respect  to  the  Revio  and  Onso  Systems  and  each  of  their  related 
consumables,  and  any  future  long-read  and  short-read  products,  our  business,  operations,  financial  condition, 
and prospects may be materially and adversely affected.

Our business may be  adversely affected by health epidemics, including any resurgence of COVID-19 cases or 
other similar outbreaks.

Our business has been and could be further adversely impacted by the effects of COVID-19 or other epidemics 
or pandemics. Although it is not possible at this time to estimate the impact that health epidemics, including the 
results  of  the  COVID-19  pandemic  and  any  future  resurgence  of  COVID-19  cases  or  other  similar  outbreaks, 
could have on our business, any pandemic or public health outbreaks or related disruptions and the measures 
taken  by  the  governments  of  countries  affected  could  disrupt  the  supply  chain  and  the  manufacture  of  our 
products.

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Our manufacturing partners and suppliers have been and could continue to be disrupted by conditions related to 
COVID-19 or other epidemics or pandemics, possibly resulting in disruption to the production of our products. If 
our manufacturing partners or suppliers are unable or fail to fulfill their obligations to us for any reason, we may 
not  be  able  to  manufacture  our  products  and  satisfy  customer  demand  or  our  obligations  under  sales 
agreements in a timely manner, and our business could be harmed as a result. There is significant uncertainty 
relating to the long-term effect of COVID-19 or other epidemics or pandemics on our business. Infections may 
resurge or become more widespread and any ensuing disruptions to business activities or supply chains could 
have a negative impact on our business, financial condition, and operating results. Because our semiconductor 
manufacturers are located in a region where immunization rates in certain communities may be low, new and 
emerging  variants  of  COVID-19  or  other  epidemics  or  pandemics  could  impact  workforce  availability  at  those 
locations  and  disrupt  supply.  For  example,  the  Chinese  government  may  re-impose  lockdowns  or  similar 
measures  to  combat  the  spread  of  health  epidemics  such  as  COVID-19  or  other  similar  outbreaks  and  such 
measures have had, and may continue to have in the future, a negative impact on manufacturing and/or supply 
chains, as well as customer demand for our products and demand through certain distributors.

For example, the COVID-19 pandemic caused us to modify our business practices, including limiting certain of 
our commercial operations and limiting certain employees from working in the office. We offered, and if there is 
a  resurgence  of  COVID-19  or  other  similar  outbreaks,  we  may  again  offer  a  significant  percentage  of  our 
employees flexibility in the amount of time they work in an office, which could adversely impact the productivity 
of certain employees and harm our business, including our future operating results. This may also present risks 
for  our  strategy  and  may  present  operational,  cybersecurity,  and  workplace  culture  challenges  that  may 
adversely affect our business.

Even after the COVID-19 pandemic has further subsided, we may continue to experience an adverse impact to 
our  business  as  a  result  of  its  global  economic  impact,  including  recessionary  effects  and  inflationary 
pressures.  Specifically,  difficult  macroeconomic  conditions,  such  as  decreases  in  discretionary  capital 
expenditure  spending,  changes  to  the  government  funding  environment,  a  reduction  in  or  the  lapsing  of 
COVID-19-related  governmental  stimulus  measures,  increased  and  prolonged  unemployment  or  a  decline  in 
consumer confidence as a result of the COVID-19 pandemic or other epidemics or pandemics, as well as limited 
or significantly reduced points of access of our products, could have a continuing adverse effect on the demand 
for some of our products and, consequently, related maintenance and support services. The degree of impact of 
COVID-19 or other epidemics or pandemics on our business will depend on several factors, such as the duration 
and the extent of the pandemic, the risk of waning immunity among persons already vaccinated and an increase 
in fatigue or skepticism with respect to initial or booster vaccinations, as well as actions taken by governments, 
businesses, and consumers in response to the pandemic, all of which continue to evolve and remain uncertain 
at this time.

Significant  changes  to  our  leadership  team  and  the  resulting  management  transitions  might  harm  our  future 
operating results.

We have experienced significant changes to our leadership team since 2020, including the appointment of our 
current President and Chief Executive Officer, Chief Financial Officer, Chief Operating Officer, Chief Commercial 
Officer, Vice President and Chief Accounting Officer, and Chair of the Board.

Although we believe these leadership transitions are in the best interest of our stakeholders, these transitions 
may result in the loss of personnel with deep institutional or technical knowledge. Further, the transition could 
potentially disrupt our operations and relationships with employees, suppliers, partners, and customers due to 
added  costs,  operational  inefficiencies,  decreased  employee  morale  and  productivity  and  increased  turnover. 
We  must  successfully  recruit  and  integrate  our  new  leadership  team  members  within  our  organization  to 
achieve  our  operating  objectives;  as  such,  the  leadership  transition  may  temporarily  affect  our  business 
performance and results of operations while the new members of our leadership team become familiar with our 
business.  In  addition,  our  competitors  may  seek  to  use  this  transition  and  the  related  potential  disruptions  to 
gain a competitive advantage over us. Furthermore, these changes may increase our dependency on the other 
members of our leadership team that remain with us, who are not contractually obligated to remain employed 
with us and may leave at any time. Any such departure could be particularly disruptive given that we are already 
experiencing  leadership  transitions  and,  to  the  extent  we  experience  additional  management  turnover, 
competition  for  top  management  is  high  such  that  it  may  take  some  time  to  find  a  candidate  that  meets  our 
requirements. Our future operating results depend substantially upon the continued service of our key personnel 
and in significant part upon our ability to attract and retain qualified management personnel. If we are unable to 
mitigate  these  or  other  similar  risks,  our  business,  results  of  operations  and  financial  condition  may  be 
materially and adversely affected.

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We  depend  on  the  continuing  efforts  of  our  senior  management  team  and  other  key  personnel.  If  we  lose 
members  of  our  senior  management  team  or  other  key  personnel  or  are  unable  to  successfully  retain,  recruit 
and train qualified scientists, engineers, sales personnel and other employees, our ability to maintain, develop 
and commercialize our products could be harmed and we may be unable to achieve our goals.

Our success depends upon the continuing services of members of our senior management team and scientific 
and  engineering  personnel.  In  particular,  our  scientists  and  engineers  are  critical  to  our  technological  and 
product  innovations,  and  we  will  need  to  hire  additional  qualified  personnel.  Our  industry,  is  characterized  by 
high demand and intense competition for talent, and the turnover rate has been and may continue to be high. 
We  compete  for  qualified  management  and  scientific  personnel  with  other  life  science  companies,  academic 
institutions  and  research  institutions,  particularly  those  focusing  on  genomics.  This  competition  had  been 
exacerbated by the increase in employee resignations in 2021 and 2022 that had been experienced by us and 
reported  by  employers  nationwide.  In  addition,  we  have  experienced  significant  turnover  in  our  senior 
management  team  in  recent  periods.  To  induce  valuable  employees  to  remain  at  our  company,  in  addition  to 
salary  and  cash  incentives,  we  have  issued  stock  options  and  restricted  stock  units  that  vest  over  time.  The 
value to employees of stock options and restricted stock units that vest over time may be significantly affected 
by movements in our stock price that are beyond our control and may at any time be insufficient to counteract 
more  lucrative  offers  from  other  companies.  We  may  face  challenges  in  retaining  and  recruiting  such 
individuals due to sustained declines in our stock price that could reduce the retention value of equity awards. 
The loss of qualified employees, or an inability to attract, retain, and motivate employees, could prevent us from 
pursuing  collaborations  and  materially  and  adversely  affect  our  support  of  existing  products,  product 
development  and  launches,  business  growth  prospects,  results  of  operations  and  financial  condition.  In 
addition, we will need to continue to recruit, hire and retain sales personnel to support the commercialization of 
our existing and new products. Our employees could leave our company with little or no prior notice and would 
be free to work for a competitor. In addition, changes to U.S. immigration policies, particularly to H-1B and other 
visa  programs,  could  restrain  the  flow  of  technical  and  professional  talent  into  the  U.S.  and  may  inhibit  our 
ability to hire qualified personnel. If one or more of our senior executives or other key personnel were unable or 
unwilling to continue in their present positions, we may not be able to replace them easily or at all, and other 
senior management may be required to divert attention from other aspects of the business. In addition, we do 
not  have  “key  person”  life  insurance  policies  covering  any  member  of  our  management  team  or  other  key 
personnel.  Further,  our  vaccination  and  return  to  office  protocols  related  to  COVID-19  or  other  epidemics  or 
pandemics  may  also  impact  the  recruitment  and  retention  of  key  employees.  The  loss  of  any  of  these 
individuals  or  any  inability  to  attract  or  retain  qualified  personnel,  including  scientists,  engineers,  sales 
personnel  and  others,  could  prevent  us  from  pursuing  collaborations  and  materially  and  adversely  affect  our 
support  of  existing  products,  product  development  and  introductions,  business  growth  prospects,  results  of 
operations and financial condition.

Our success is highly dependent on our ability to further penetrate nucleic acid sequencing applications as well 
as  on  the  growth  and  expansion  of  the  demand  for  our  products.  If  our  products  fail  to  achieve  and  sustain 
sufficient market acceptance, we will not generate expected revenue and our business may not succeed.

Although nucleic acid sequencing technology is well-established, our SMRT Sequencing technology is relatively 
new and evolving. We cannot be sure that our current or future products will gain acceptance in the marketplace 
at levels sufficient to support our costs. Our success depends, in part, on our ability to expand overall demand 
for nucleic acid sequencing to include new applications that are not practicable with other current technologies 
and  to  introduce  new  products  that  capture  a  larger  share  of  growing  overall  demand  for  sequencing.  To 
accomplish  this,  we  must  successfully  commercialize,  and  continue  development  of,  our  proprietary  SMRT 
Sequencing  technology  for  use  in  a  variety  of  life  science  and  other  research  applications,  including  uses  by 
academic,  government  and  clinical  laboratories,  as  well  as  pharmaceutical,  diagnostic,  biotechnology,  and 
agriculture  companies,  among  others.  However,  we  may  be  unsuccessful  in  these  efforts  and  the  sale  and 
commercialization of the SMRT Cell, Sequel II/IIe, Revio and Onso Systems, and related products may not grow 
sufficiently to cover our costs.

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There can be no assurance that we will be successful in adding new products or securing additional customers 
for  our  current  and  future  products,  including  with  respect  to  the  SMRT  Cell,  Sequel  II/IIe  Systems,  Revio  and 
Onso. If we are unable to successfully develop acquired technologies and sell acquired technology products, we 
may fail to achieve our strategic commercial initiatives in connection with the planned release of new products 
and  anticipated  entry  into  new  markets.  Our  ability  to  further  penetrate  existing  applications  and  any  new 
applications depends on a number of factors, including the cost, performance and perceived value associated 
with our products, as well as customers’ willingness to adopt a different approach to nucleic acid sequencing. 
Potential customers may have already made significant investments in other sequencing technologies and may 
be  unwilling  to  invest  in  new  technologies.  We  are  experiencing  pricing  pressures  caused  by  industry 
competition and increased demand for lower-priced instruments and lower operational costs. We have limited 
experience commercializing and selling products outside of the academic and research settings, and we cannot 
guarantee success in acquiring additional customers. Furthermore, we cannot guarantee that our products will 
be  satisfactory  to  potential  customers  or  that  our  products  will  perform  in  accordance  with  customer 
expectations.

Nucleic  acid  sequencing  applications  are  new  and  dynamic,  and  there  can  be  no  assurance  that  they  will 
develop  as  quickly  as  we  anticipate,  that  they  will  reach  their  full  potential  or  that  our  products  will  be 
appropriate  or  competitive  for  these  applications.  As  a  result,  we  may  be  required  to  refocus  our  marketing 
efforts, and we may have to make changes to the specifications of our products to enhance our ability to enter 
particular  applications  more  quickly.  We  may  also  need  to  delay  full-scale  commercial  deployment  of  new 
products as we develop them in order to perform quality control and early access user testing. We also need to 
maintain reliable supply chains for the various components in our new products and consumables to support 
large-scale  commercial  production.  Even  if  we  are  able  to  implement  our  technology  successfully,  we  and/or 
our  sales  and  distribution  partners  may  fail  to  achieve  or  sustain  market  acceptance  of  our  current  or  future 
products  across  the  full  range  of  our  intended  life  science  and  other  applications.  We  need  to  continue  to 
expand  and  update  our  internal  capabilities  or  to  collaborate  with  other  partners,  or  both,  in  order  to 
successfully expand sales of our products in the applications that we seek to reach, which we may be unable to 
do at the scale required to support our business.

If  the  demand  for  our  products  grows  more  slowly  than  anticipated,  if  we  are  unable  to  successfully  scale  or 
otherwise  ensure  sufficient  manufacturing  capacity  for  new  products  to  meet  demand,  if  we  are  not  able  to 
successfully market and sell our products, if competitors develop better or more cost-effective products, if our 
product launches and commercialization are not successful, or if we are unable to further grow our customer 
base or do not realize the growth with existing customers that we are expecting, our current and future sales 
and  revenue  may  be  materially  and  adversely  harmed,  or  we  may  recognize  an  impairment  loss,  and  our 
business may not succeed.

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We  rely  on  other  companies  for  the  manufacture  of  certain  components  and  sub-assemblies  and  intend  to 
outsource  additional  sub-assemblies  in  the  future,  some  of  which  are  sole  sources.  We  may  not  be  able  to 
successfully  scale  the  manufacturing  process  necessary  to  build  and  test  multiple  products  on  a  full 
commercial basis, which could materially harm our business.

Our  products  are  complex  and  involve  a  large  number  of  unique  components,  many  of  which  require  precise 
manufacturing.  The  nature  of  our  products  requires  customized  components  that  are  currently  available  only 
from a limited number of sources, and in some cases, single sources. We have chosen to source certain critical 
components from a single source, including suppliers for our SMRT Cells, reagents, and instruments. We cannot 
assure  you  that  product  supplies  will  not  be  limited  or  interrupted,  especially  with  respect  to  our  sole  source 
third-party  manufacturing  and  supply  collaborators,  or  that  product  supplies  will  be  of  satisfactory  quality  or 
continue to be available at acceptable prices. In particular, any replacement of our manufacturers could require 
significant effort and expertise because there may be a limited number of qualified replacements. We may be 
unable to negotiate binding agreements with our current and future sole source third-party manufacturing and 
supply collaborators or, in the event that such collaborators’ services become interrupted for any reason, find 
replacement manufacturers to support our development and commercial activities at commercially reasonable 
terms.  We  do  not  always  have  arrangements  in  place  for  a  redundant  or  second-source  supply  for  our  sole 
source vendors in the event they cease to provide their products or services to us or fail to provide sufficient 
quantities  in  a  timely  manner.  If  we  are  required  to  purchase  these  components  from  alternative  sources,  it 
could take several months or longer to qualify the alternative sources. If we are unable to source these product 
components  from  sole-source  third-party  manufacturing  and  supply  collaborators  for  any  reason,  including  in 
connection  with  acts  of  terrorism,  hostilities,  military  conflict  and  acts  of  war,  including  between  China  and 
Taiwan, or secure a sufficient supply of these product components on a timely basis, or if these components do 
not  meet  our  expectations  or  specifications  for  quality  and  functionality,  our  operations  and  manufacturing 
would  be  materially  and  adversely  affected,  we  could  be  unable  to  meet  customer  demand  and  our  business 
and results of operations may be materially and adversely affected.

The  operations  of  our  third-party  manufacturing  partners  and  suppliers  have  had  and  may  in  the  future  be 
disrupted by conditions unrelated to our business or operations or that are beyond our control, including but not 
limited to international trade restrictions, inflation, supply chain disruptions, and conditions related to COVID-19 
or  other  epidemics  or  pandemics.  If  our  manufacturing  partners  or  suppliers  are  unable  or  fail  to  fulfill  their 
obligations to us for any reason, we may not be able to manufacture our products and satisfy customer demand 
or our obligations under sales agreements in a timely manner, and our business could be harmed as a result. 
For  example,  the  global  shortage  of  semiconductors,  which  has  been  reported  since  early  2021,  has  caused 
challenges  for  us  in  our  supply  chain  and  resulted  in  some  cost  increases  that  have  and  may  continue  to 
adversely impact margins. During these periods of shortages or delays, the price of components may increase, 
or the components may not be available at all. Our suppliers have raised their prices and may continue to raise 
prices  that  we  may  not  be  able  to  pass  on  to  our  customers,  which  could  adversely  affect  our  business, 
including our competitive position, market share, revenues, and profit margins in material ways. We may not be 
able  to  secure  enough  components  at  reasonable  prices  or  of  acceptable  quality  to  build  new  products  in  a 
timely manner in the quantities or configurations needed. For example, the Chinese government may re-impose 
lockdowns  or  similar  measures  to  combat  the  spread  of  COVID-19  or  other  similar  outbreaks  and  these 
measures have had, and may continue to have in the future, a negative impact on manufacturing and/or supply 
chains, in addition to customer demand for our products and demand through certain distributors. If as a result 
of global economic or political instability, such as the political uncertainty in the Middle East associated with the 
Israel  and  Hamas  conflict,  an  escalation  of  the  war  in  Ukraine,  potential  uncertainty  related  to  Taiwan  and  its 
relationship  with  China,  other  disease  outbreaks,  or  supply  issues,  we  or  our  contractors  could  experience 
shortages, business disruptions or delays for materials sourced or manufactured in the affected countries, and 
their ability to supply us with instruments or product components may be affected. From time to time, certain 
components  of  our  systems  and  reagents  may  reach  the  end  of  their  life  cycles  or  become  obsoleted  by  our 
suppliers,  and  we  would  have  to  procure  alternative  sources  for  these  end-of-life  products.  If  we  encounter 
delays  or  difficulties  in  securing  the  quality  and  quantity  of  materials  we  require  for  our  products,  our  supply 
chain would be interrupted, which would adversely affect sales. If any of these events occur, our business and 
operating results could be harmed. Accordingly, if any of the foregoing occurs, our ability to commercialize our 
products,  revenue  and  gross  margins  could  suffer  until  lockdowns  from  COVID-19  or  other  epidemics  or 
pandemics infections are reduced, supply issues or business disruptions are resolved and/or other sources can 
be developed.

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In  addition,  because  our  semiconductor  suppliers  are  in  regions  that  may  have  communities  with  low 
vaccination rates, any variants of COVID-19 that evolve in the future or other outbreaks could lead to increased 
infections  among  workers  that  could  further  disrupt  the  supply  chain.  Our  current  manufacturing  process  is 
characterized by long lead times between the placement of orders for and delivery of our products. If we do not 
accurately  anticipate  our  needs  or  if  we  receive  insufficient  components  to  manufacture  our  products  on  a 
timely  basis  to  meet  customer  demand,  our  sales  and  our  gross  margin  may  be  adversely  affected,  and  our 
business  could  be  materially  harmed.  If  we  are  unable  to  reduce  our  manufacturing  costs  and  establish  and 
maintain  reliable,  high-volume  manufacturing  suppliers  as  we  scale  our  operations  and  expand  our  product 
offerings,  our  business,  operations,  financial  condition,  and  prospects  could  be  materially  and  adversely 
harmed.

We  may  be  unable  to  consistently  manufacture  our  instruments  and  consumables,  including  SMRT  Cells  and 
reagents, to the necessary specifications or in quantities necessary to meet demand at an acceptable cost or at 
an acceptable performance level.

In order to successfully generate revenue from our products, we need to supply our customers with products 
that  meet  their  expectations  for  quality  and  functionality  in  accordance  with  established  specifications.  Our 
customers  have  experienced  variability  in  the  performance  of  our  products.  We  have  experienced  and  may 
continue  to  experience  delays,  quality  issues  or  other  difficulties  leading  to  customer  dissatisfaction  with  our 
products.  Our  production  of  SMRT  Cells,  flow  cells  and  of  reagents  for  both  our-long  and  short-read 
technologies, involve a long and complex manufacturing process and has been and may in the future be below 
desired yields and resulting output levels. We have experienced and may experience in the future manufacturing 
delays, product defects, variability in the performance of SMRT Cells, flow cells and other products, inadequate 
reserves for inventory, or other issues.

There is no assurance that we will be able to manufacture our products so that they consistently achieve the 
product  specifications  and  quality  that  our  customers  expect,  including  any  products  developed  for  clinical 
uses. Problems in the design or quality of our products, including low manufacturing yields of SMRT Cells, flow 
cells,  or  sub-performing  reagent  lots  may  have  a  material  adverse  effect  on  our  brand,  business,  financial 
condition, and operating results, and could result in us losing our ISO certifications. If we were to lose our ISO 
certifications, then our customers might choose not to purchase products from us. There is also no assurance 
that we will be able to increase manufacturing yields and decrease costs, particularly if high rates of inflation 
continue, or that we will be successful in forecasting customer demand or manufacturing and supply costs, or 
that  product  supplies,  including  reagents  or  integrated  chips,  will  not  be  limited  or  interrupted,  or  will  be  of 
satisfactory  quality  or  continue  to  be  available  at  acceptable  prices.  Furthermore,  while  we  are  undertaking 
efforts  to  increase  our  manufacturing  scale  and  capability,  we  may  not  be  able  to  increase  manufacturing  to 
meet anticipated demand or may experience downtime in our manufacturing facilities, including, for example, if  
our suppliers are unable to meet our increased demand at a time when the supply chain is under duress due to 
potential dislocations and disruptions in product and employee availability (whether due to health epidemics or 
pandemics  or  otherwise).  An  inability  to  manufacture  products  and  components  that  consistently  meet 
specifications, in necessary quantities and at commercially acceptable costs, will have a negative impact, and 
may  have  a  material  adverse  effect  on  our  business,  product  development  timelines,  financial  condition  and 
results of operations.

Rapidly changing technology in life sciences and research diagnostics could make our products obsolete unless 
we  continue  to  develop,  manufacture  and  commercialize  new  and  improved  products  and  pursue  new 
opportunities.

industry 

Our 
is  characterized  by  rapid  and  significant  technological  changes,  frequent  new  product 
introductions and enhancements and evolving industry standards. These new and evolving technologies may be 
superior to, impair, or render obsolete the products we currently offer or the technologies currently underlying 
our  products.  Our  future  success  depends  on  our  ability  to  continually  improve  our  products,  to  develop  and 
introduce new products that address the evolving needs of our customers on a timely and cost-effective basis 
and to pursue new opportunities. These new opportunities may be outside the scope of our proven expertise or 
in  areas  where  demand  is  unproven,  and  new  products  and  services  developed  by  us  may  not  gain  market 
acceptance or may not adequately perform to capture market share. Our inability to develop and introduce new 
products  and  to  gain  market  acceptance  of  our  existing  and  new  products  could  harm  our  future  operating 
results.  Unanticipated  difficulties  or  delays 
in 
commercializing  our  existing  or  new  products  in  sufficient  quantities  and  of  acceptable  quality  to  meet 
customer  demand,  including  with  respect  to  the  SMRT  Cell,  Sequel  II/IIe  Systems,  Revio  and  Onso,  could 
diminish future demand for our products and may materially and adversely harm our future operating results.

in  replacing  existing  products  with  new  products  or 

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The  size  of  the  markets  for  our  products,  including  our  Revio  and  Onso  instruments,  may  be  smaller  than 
estimated, and new market opportunities may not develop as quickly as we expect, or at all, limiting our ability 
to successfully sell our products.

The  market  for  sequencing  systems  and  consumables  products  is  evolving,  making  it  difficult  to  accurately 
predict the size of the markets for our current and future products, including our Revio and Onso instruments. 
Our  estimates  of  the  total  addressable  market  for  our  current  and  future  products  are  based  on  a  number  of 
internal  and  third-party  estimates  and  assumptions  that  may  be  incorrect,  including  the  assumptions  that 
academic,  governmental,  corporate,  or  other  sources  of  funding  will  continue  to  be  available  to  life  sciences 
researchers at times and  in amounts necessary to allow them to purchase our products. In addition, sales of 
new products may take time to develop and mature and we cannot be certain that these market opportunities 
will develop as we expect. While we believe our assumptions and the data underlying our estimates of the total 
addressable market for our products are reasonable, these assumptions and estimates may not be correct and 
the conditions supporting our assumptions or estimates, or those underlying the third-party data we have used, 
may change at any time, thereby reducing the accuracy of our estimates. As a result, our estimates of the total 
addressable market and growth opportunities for our products may be incorrect.

The  future  growth  of  the  market  for  our  current  and  future  products  depends  on  many  factors  beyond  our 
control, including recognition and acceptance of our products by the research and scientific communities, the 
growth, prevalence and costs of competing products and solutions and the development of robust ecosystems 
supporting our products and their methodologies. For example, the market acceptance and growth of long-read 
sequencing technologies, like our Revio system, depends on a variety of factors, including the availability and 
cost-effectiveness  of  related  tools  for  high  quality  sample  collection  and  preparation  and  advanced 
bioinformatic  tools  to  process  results;  as  well  as  the  perceived  advantages  and  disadvantages  of  long-read 
sequencing  compared  to  short-read  or  other  sequencing  technologies:  consequently,  if  potential  customers 
conclude  the  costs  of  adopting  long-read  sequencing  technologies  outweigh  the  benefits,  the  market  for  our 
Revio systems may be negatively impaired. There can be no assurance that our current or future products will 
gain traction in the market. If the markets for our current and future products are smaller than estimated or do 
not develop as we expect, our growth may be limited, and it could materially and adversely affect our business, 
operations, financial condition and prospects.

Increased market adoption of our products by customers may depend on the availability of sample preparation 
and informatics tools, some of which may be developed by third parties.

Our  commercial  success  may  depend  in  part  upon  the  development  of  sample  preparation  and  software  and 
informatics  tools  by  third  parties  for  use  with  our  products.  We  cannot  guarantee  that  product  supplies, 
including reagents, will not be limited or interrupted, or will be of satisfactory quality or continue to be available 
at acceptable prices, or that third parties will develop tools that our current and future customers will find useful 
with  our  products,  or  that  customers  will  adopt  such  third-party  tools  on  a  timely  basis  or  at  all.  A  lack  of 
complementary  sample  preparation  and  informatics  tools,  or  delayed  updates  of  such  tools,  may  impede  the 
adoption of our products and may materially and adversely impact our business.

We  operate  in  a  highly  competitive  industry  and  if  we  are  not  able  to  compete  effectively,  our  business  and 
operating results will likely be harmed.

There  are  a  significant  number  of  companies  offering  nucleic  acid  sequencing  products  and/or  services, 
including  Illumina,  BGI  Genomics  (also  known  as  MGI  or  Complete  Genomics),  Thermo,  ONT  Ltd.,  Roche, 
Bionano, and Qiagen. Other companies recently entering the market include Ultima, Element and Singular. Many 
of these companies currently have greater name recognition, more substantial intellectual property portfolios, 
longer  operating  histories,  significantly  greater  financial,  technical,  research  and/or  other  resources,  more 
experience in new product development, larger and more established manufacturing capabilities and marketing, 
sales,  and  support  functions,  and/or  more  established  distribution  channels  to  deliver  products  to  customers 
than  we  do.  These  companies  may  be  able  to  respond  more  quickly  and  effectively  than  we  can  to  new  or 
changing opportunities, technologies, standards, or customer requirements.

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There  are  also  several  companies  that  are  in  the  process  of  developing  or  have  already  developed  and 
commercialized  new,  competing  or  potentially  competing  technologies,  products  and/or  services,  including 
ONT Ltd. and its subsidiaries, against whom we have filed complaints for patent infringement in the U.S. District 
Court  for  the  District  of  Delaware  and,  previously,  with  the  U.S.  International  Trade  Commission,  in  the  High 
Court of England and Wales and in the District Court of Mannheim, Germany. ONT Ltd. previously filed claims 
against us in the High Court of England and Wales and the District Court of Mannheim, Germany, also for patent 
infringement, and its subsidiary, Oxford Nanopore Technologies, Inc. (“ONT Inc.”), filed counterclaims against us 
in  the  U.S.  District  Court  for  the  District  of  Delaware  seeking  declaratory  judgements  of  non-infringement, 
invalidity  and  unenforceability  of  the  asserted  patents,  as  well  as  antitrust,  false  advertising  and  unfair 
competition  counterclaims  that  were  subsequently  dismissed  by  that  court.  Roche  is  developing  potentially 
competing sequencing products. Increased competition may result in pricing pressures, which could harm our 
sales,  profitability  or  market  share.  Our  failure  to  further  enhance  our  existing  products  and  to  introduce  new 
products  to  compete  effectively  could  materially  and  adversely  affect  our  business,  operations,  financial 
condition, and prospects.

We may be unable to successfully increase sales of our current products or market and sell our future products.

Our ability to achieve profitability depends, in part, on our ability to attract customers for our current and future 
products  including  Revio  and  Onso,  and  we  may  be  unable  to  effectively  market  or  sell  our  products  or  find 
appropriate  partners  to  do  so.  To  perform  sales,  marketing,  distribution,  and  customer  support  functions 
successfully, we face a number of risks, including:

•

•

•

•

•

our ability to attract, retain and manage qualified sales, marketing, and service personnel necessary to 
expand market acceptance for our technologies;

the performance and commercial availability expectations of our existing and potential customers with 
respect to new and existing products;

availability of potential sales and distribution partners to sell our technologies, and our ability to attract 
and retain such sales and distribution partners;

the  time  and  cost  of  maintaining  and  growing  a  specialized  sales,  marketing  and  service  force  for  a 
particular application, which may be difficult to justify in light of the revenue generated; and

our sales, marketing and service force may be unable to execute successful commercial activities.

We  have  enlisted  and  may  continue  to  enlist  third  parties  to  assist  with  sales,  distribution  and  customer 
support. There is no guarantee that we will be successful in attracting desirable sales and distribution partners, 
that we will be able to enter into arrangements with such partners on terms favorable to us or that we will be 
able to retain such partners on a going-forward basis. If our sales and marketing efforts, or those of any of our 
third-party  sales  and  distribution  partners,  are  not  successful,  or  our  products  do  not  perform  in  accordance 
with  customer  expectations,  our  technologies  and  products  may  not  gain  market  acceptance,  which  could 
materially and adversely impact our business, operations, financial condition, and prospects.

Large purchases by a limited number of customers represent a significant portion of our revenue, and any loss 
or  delay  of  expected  purchases  has  resulted,  and  in  the  future  could  result,  in  material  quarter-to-quarter 
fluctuations of our revenue or otherwise adversely affect our results of operations.

We receive a significant portion of our revenue from a limited number of customers. For example, for the years 
ended December 31, 2022 and 2021, one of our customers, who is our primary distributor in China, accounted 
for  approximately  12%  and  13%  of  our  total  revenue,  respectively.  For  the  year  ended  December  31,  2023,  no 
single  customer  accounted  for  10%  or  greater  of  our  total  revenue.  Many  of  these  customers  make  large 
purchases  on  a  purchase-order  basis  rather  than  pursuant  to  long-term  contracts.  As  a  consequence  of  the 
concentrated nature of our customer base and their purchasing behavior, our quarterly revenue and results of 
operations have fluctuated, and may fluctuate in the future, from quarter to quarter and are difficult to forecast. 
For  example,  the  cancellation  of  orders  or  acceleration  or  delay  in  anticipated  product  purchases  or  the 
acceptance  of  shipped  products  by  our  larger  customers  has  materially  affected,  and  in  the  future  could 
materially affect, our revenue and results of operations in any quarterly period. We have been, and may in the 
future be, unable to sustain or increase our revenue from our larger customers, or offset any discontinuation or 
decrease  of  purchases  by  our  larger  customers  with  purchases  by  new  or  other  existing  customers.  To  the 
extent one or more of our larger customers experience significant financial difficulty, bankruptcy or insolvency, 
this  could  have  a  material  adverse  effect  on  our  sales  and  our  ability  to  collect  on  receivables,  which  could 
materially and adversely harm our financial condition and results of operations.

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In  addition,  many  of  our  customers,  including  some  of  our  larger  customers,  have  negotiated,  or  may  in  the 
future negotiate, volume-based discounts or other more favorable terms from us or our sales and distribution 
partners, which can and have had a negative effect on our gross margins or revenue.

We  expect  that  such  concentrated  purchases  will  continue  to  contribute  materially  to  our  revenue  for  the 
foreseeable  future  and  that  our  results  of  operations  may  fluctuate  materially  as  a  result  of  such  larger 
customers’ buying patterns. In addition, we may see consolidation of our customer base. The loss of one of our 
larger customers, a significant delay or reduction in its purchases, or any volume-based discount or other more 
favorable terms that we or our sales and distribution partner(s) may agree to provide, in light of the aggregated 
purchase volume or buying power resulting from such consolidation, has harmed, and in the future could harm, 
our business, financial condition, results of operations and prospects.

Our  products  are  highly  complex,  have  recurring  support  requirements  and  could  have  unknown  defects  or 
errors, which may give rise to claims against us or divert application of our resources from other purposes.

Products  using  our  SMRT  sequencing  and  SBB  technology  are  highly  complex  and  may  develop  or  contain 
undetected  defects  or  errors.  Our  customers  have  previously  experienced  reliability  issues  with  our  existing 
products, including the Sequel System and the Sequel II/IIe Systems. In addition, it is possible our customers 
could  experience  reliability  issues  with  current  or  future  products,  including  the  Sequel  II/IIe,  Revio  and  Onso 
Systems. Despite internal and external testing, defects, or errors may arise in our products, which could result in 
a  failure  to  obtain,  maintain,  or  increase  market  acceptance  of  our  products,  diversion  of  development 
resources,  injury  to  our  reputation  and  increased  warranty,  service,  and  maintenance  costs.  New  products, 
including Revio and Onso, or enhancements to our existing products, including the SMRT Cell and Sequel II/IIe 
Systems,  in  particular  may  contain  undetected  errors  or  performance  problems  that  are  discovered  only  after 
delivery  to  customers.  If  our  products  have  reliability  or  other  quality  issues  or  require  unexpected  levels  of 
support in the future, the market acceptance and utilization of our products may not grow to levels sufficient to 
support our costs and our reputation and business could be harmed. Low utilization rates of our products could 
cause  our  revenue  and  gross  margins  to  be  adversely  affected.  We  provide  a  warranty  for  our  sequencing 
instruments and consumables, which is generally limited to replacing, repairing, or at our option, giving credit for 
any sequencing instrument or consumable with defects in material or workmanship. Service contracts for our 
sequencing instruments may be separately purchased. Defects or errors in our products may also discourage 
customers  from  purchasing  our  products.  The  costs  incurred  in  correcting  any  defects  or  errors  may  be 
substantial  and  could  materially  and  adversely  affect  our  operating  margins.  If  our  service  and  support  costs 
increase, our business and operations may be materially and adversely affected.

In addition, such defects or errors could lead to the filing of product liability claims against us or against third 
parties  whom  we  may  have  an  obligation  to  indemnify  against  such  claims,  which  could  be  costly  and  time-
consuming  to  defend  and  result  in  substantial  damages.  Although  we  have  product  liability  insurance,  any 
product liability insurance that we have or procure in the future may not protect our business from the financial 
impact  of  a  product  liability  claim.  Moreover,  we  may  not  be  able  to  obtain  adequate  insurance  coverage  on 
acceptable  terms.  Any  insurance  that  we  have  or  obtain  will  be  subject  to  deductibles  and  coverage  limits.  A 
product liability claim could have a material adverse effect on our business, financial condition, and results of 
operations.

A significant portion of our sales depends on customers’ spending budgets that may be subject to significant 
and unexpected variation which could have a negative effect on the demand for our products.

Our instruments represent significant capital expenditures for our customers in research applications. Current 
and  potential  customers  for  our  current  or  future  products  include  academic  and  government  institutions, 
genome centers, medical research institutions, clinical laboratories, pharmaceutical, agricultural, biotechnology, 
diagnostic and chemical companies. Their spending budgets can have a significant effect on the demand for 
our  products.  Spending  budgets  are  based  on  a  wide  variety  of  factors,  including  the  allocation  of  available 
resources  to  make  purchases,  funding  from  government  sources  which  is  highly  uncertain  and  subject  to 
change,  the  spending  priorities  among  various  types  of  research  equipment,  policies  regarding  capital 
expenditures  during  economically  uncertain  periods  and  the  potential  impacts  from  health  epidemics  or 
pandemics.  Any  decrease  in  capital  spending  or  change  in  spending  priorities  of  our  current  and  potential 
customers  could  significantly  reduce  the  demand  for  our  products.  Any  delay  or  reduction  in  purchases  by 
current or potential customers or our inability to forecast fluctuations in demand could materially and adversely 
harm our future operating results.

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We may not be able to convert our orders in backlog into revenue.

Our backlog represents product orders from our customers that we have confirmed but have not been able to 
fulfill, and, accordingly, for which we have not yet recognized revenue. We may not receive revenue from these 
orders, and any order backlog we report may not be indicative of our future revenue.

Many events can cause an order to be delayed or not completed at all, some of which may be out of our control, 
including  the  potential  impacts  from  health  epidemics  or  pandemics  and  our  suppliers,  especially  our  sole 
source  suppliers,  not  being  able  to  provide  us  with  products  or  components.  If  we  delay  fulfilling  customer 
orders or if customers reconsider their orders, those customers may seek to cancel or modify their orders with 
us. Customers may otherwise seek to cancel or delay their orders even if we are prepared to fulfill them. If our 
orders in backlog do not result in sales, our operating results may suffer.

Our sales cycles are unpredictable and lengthy, which makes it difficult to forecast revenue and may increase 
the magnitude of quarterly or annual fluctuations in our operating results.

The sales cycles for our sequencing instruments are lengthy because they represent a major capital expenditure 
and  generally  require  the  approval  of  our  customers’  senior  management.  This  may  contribute  to  substantial 
fluctuations in our quarterly or annual operating results, particularly during periods in which our sales volume is 
low. Because of these fluctuations, it is likely that in some future quarters our operating results will fall below 
the expectations of securities analysts or investors. If that happens, the market price of our stock would likely 
decrease. Past fluctuations in our quarterly and annual operating results have resulted in decreases in our stock 
price.  Such  fluctuations  also  mean  that  investors  may  not  be  able  to  rely  on  our  operating  results  in  any 
particular period as an indication of future performance. Sales to existing customers and the establishment of a 
business relationship with other potential customers is a lengthy process, generally taking several months and 
sometimes  longer.  Following  the  establishment  of  the  relationship,  the  negotiation  of  purchase  terms  can  be 
time-consuming, including as a result of seasonal factors, as discussed below, and a potential customer may 
require an extended evaluation and testing period. Our sales cycles may also lengthen, and those sales cycles 
may result in lower units sold per cycle, as we continue to introduce our Revio and Onso instruments and their 
associated consumables to the market, as our customers may have additional administrative, technical or other 
requirements associated with transitioning to new products and technologies. In anticipation of product orders, 
we  may  incur  substantial  costs  before  the  sales  cycle  is  complete  and  before  we  receive  any  customer 
payments. As a result, if a sale is not completed or is canceled or delayed, we may have incurred substantial 
expenses, making it more difficult for us to become profitable or otherwise negatively impacting our financial 
results.  Even  if  our  selling  efforts  are  successful,  the  realization  of  revenue  may  be  substantially  delayed,  our 
ability  to  forecast  our  future  revenue  may  be  more  limited  and  our  revenue  may  fluctuate  significantly  from 
quarter to quarter and year over year. For more information on the impact of these fluctuations on our results 
and stock price, see “—Our operating results fluctuate from quarter to quarter and year over year, which makes 
our future results difficult to predict and could negatively impact the market price of our common stock,” below.

Because some of our customers and suppliers are based in China, our business, financial condition and results 
of operations could be adversely affected by the political and economic tensions between the United States and 
China.

We are subject to risks associated with political conflicts between the U.S. and China. A significant portion of 
our revenue is generated from China. For example, for the years ended December 31, 2022 and 2021, one of our 
customers,  who  is  our  primary  distributor  in  China,  accounted  for  approximately  12%  and  13%  of  our  total 
revenue, respectively. For the year ended December 31, 2023, no single customer accounted for 10% or greater 
of our total revenue. In addition, certain components, some of which are critical components, of our products 
are manufactured in China. These components are either sourced directly from companies in China or indirectly 
from third parties that source from companies in China.

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Consequently, we are subject to significant risks associated with the trading relationship between the U.S. and 
China,  which  is  currently  characterized  by  significant  uncertainty.  Tariffs  imposed  by  the  U.S.  and  China  have 
increased,  and  may  continue  to  increase,  our  costs.  Additionally,  export  restrictions  imposed  by  the  U.S.  may 
impact our ability to export certain products to customers or distributors in China and restrict our ability to use 
certain integrated circuits in our products, and it is possible that additional restrictions will be put in place that 
could  impact  our  ability  to  provide  our  products  to  customers  or  distributors  in  China  or  source  components 
from China. Moreover, the Chinese government may retaliate against U.S. trade restrictions in ways that could 
impact  our  business.  Given  the  relatively  fluid  regulatory  environment  in  China  and  the  United  States  and 
uncertainty  how  the  U.S.  or  foreign  governments  will  act  with  respect  to  export  controls,  tariffs,  international 
trade agreements and policies, there could be additional import, export, tax, or other regulatory changes in the 
future. Any such changes could directly and adversely impact our financial results and results of operations. For 
more  information,  see  “—Enhanced  trade  tariffs,  import  restrictions,  export  restrictions,  Chinese  regulations  or 
other trade barriers may materially harm our business.”

Other risks could include:

•

•

•

interruptions  to  operations  in  China  as  a  result  of  potential  disease  outbreaks,  including  any  further 
COVID-19 related outbreaks, and natural catastrophic events, which have in the past and can result in 
the future in business closures, transportation restrictions, import and export complications and cause 
shortages in the supply of raw materials or disruptions in manufacturing;

product  supply  disruptions  and  increased  costs  as  a  result  of  heightened  exposure  to  changes  in  the 
policies of the Chinese government, political unrest or unstable economic conditions in China; and

the nationalization or other expropriation of private enterprises or intellectual property by the Chinese 
government.

Difficulties  in  this  relationship  may  require  us  to  take  actions  adverse  to  our  business  to  comply  with 
governmental restrictions on business and trade with China.

We  face  significant  risks  associated  with  doing  business  with  Taiwanese  suppliers  and  manufacturers  due  to 
the tense relationship between Taiwan and mainland China.

Substantially all of our consumable chips are partly manufactured by a company based in Taiwan. Our supply of 
consumables  chips  and  other  critical  components  may  be  materially  and  adversely  affected  by  diplomatic, 
geopolitical,  military  and  other  developments  affecting  the  relationship  between  China  and  Taiwan.  Recent 
military  exercises  in  the  Taiwan  Strait  have  contributed  to  geopolitical  uncertainty  regarding  the  future  of  the 
relationship  between  China  and  Taiwan.  Current  or  future  diplomatic,  geopolitical,  military  or  other  tensions 
between  China  and  Taiwan  may  lead  to  circumstances  that  negatively  affect  the  availability  of  such 
consumable chips and other critical components to us, which could limit or prohibit our ability to manufacture 
consumable chips and other critical components or lead to an increase in our supply costs if we cannot find a 
similar  cost  alternative  supplier,  which  could  materially  and  adversely  impact  our  business,  operations, 
prospects, financial condition and results, and results of operations.

Our  operating  results  fluctuate  from  quarter  to  quarter  and  year  over  year,  which  makes  our  future  results 
difficult to predict and could negatively impact the market price of our common stock.

We  operate  on  a  December  31st  year-end  and  believe  that  there  are  significant  seasonal  factors  which  may 
cause sales of our products, and particularly our sequencing instruments, to vary on a quarterly or yearly basis, 
contribute to lengthy sales cycles for our sequencing instruments, and increase the magnitude of quarterly or 
annual fluctuations in our operating results. We believe that this seasonality results from a number of factors, 
including  the  procurement  and  budgeting  cycles  of  many  of  our  customers,  especially  government-funded 
customers,  which  often  coincide  with  government  fiscal  year  ends.  For  example,  the  U.S.  government’s  fiscal 
year-end  occurs  in  our  third  quarter  and  may  result  in  increased  sales  of  our  products  during  this  quarter  if 
government-funded customers have unused funds that may be forfeited, or future budgets that may be reduced 
if funds remain unspent at fiscal year-end. Furthermore, Lunar New Year celebrations, which occur during our 
first quarter, and may last for a week or longer, resulting in closure of many of our customers’ offices in China 
and  across  the  Asia-Pacific  region  have  caused,  and  may  in  the  future  cause,  decreased  sales  of  our 
consumables  during  our  first  quarter.  These  factors  have  contributed,  and  in  the  future  may  contribute,  to 
substantial fluctuations in our quarterly operating results.

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Our  operating  results  during  any  given  period  can  also  be  impacted  by  numerous  other  factors,  including  the 
following:

• market acceptance for our products;

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our ability to attract new customers;

the length of our sales cycles, as discussed above;

our ability to achieve economies of scale and other manufacturing efficiencies at the rate we anticipate;

publications of studies by us, our competitors or third parties;

the timing and success of new product introductions by us or our competitors or other changes in the 
competitive dynamics of our industry, such as consolidation;

the amount and timing of our costs and expenses;

changes in our pricing policies or those of our competitors;

general economic, industry and market conditions;

the impact of catastrophic events, including health epidemics or pandemics and military or other armed 
conflicts;

the regulatory environment in which we operate;

expenses associated with warranty obligations or unforeseen product quality issues;

the hiring, training, and retention of key employees, including our ability to grow our sales organization;

litigation or other claims against us for intellectual property infringement or otherwise;

our ability to obtain additional financing as necessary; and

changes or trends in new technologies and industry standards.

Consequently,  it  is  possible  that  in  some  quarters  our  operating  results  will  fall  below  the  expectations  of 
securities analysts or investors. If that happens, the market price of our common stock would likely decrease. 
These fluctuations, among other factors, also mean that our operating results in any particular period may not 
be relied upon as an indication of future performance. Seasonal or cyclical variations in our sales have in the 
past,  and  may  in  the  future,  become  more  or  less  pronounced  over  time,  and  have  in  the  past  materially 
affected,  and  may  in  the  future  materially  affect,  our  business,  financial  condition,  results  of  operations,  and 
prospects.

Our ability to use net operating losses to offset future taxable income may be subject to substantial limitations, 
and changes to U.S. tax laws may cause us to make adjustments to our financial statements.

Under Section 382 of the Internal Revenue Code, a corporation that undergoes an “ownership change” is subject 
to limitations on its ability to utilize its pre-change net operating losses (“NOLs”) to offset future taxable income. 
We believe that we have had one or more ownership changes, and as a result our existing NOLs are currently 
subject  to  limitation.  Future  changes  in  our  stock  ownership  could  result  in  additional  ownership  changes, 
including potentially material changes, under Section 382. Consequently, we may not be able to utilize some or 
all of our NOLs even if we attain profitability.

Our  facilities  in  California  are  located  near  earthquake  faults,  and  the  occurrence  of  an  earthquake  or  other 
catastrophic  disaster  could  cause  damage  to  our  facilities  and  equipment,  which  could  require  us  to  cease  or 
curtail operations.

Our  facilities  in  California  are  located  near  earthquake  fault  zones  and  are  vulnerable  to  damage  from 
earthquakes. We are also vulnerable to damage from other types of disasters, including fire, floods, power loss, 
communications failures and similar events. If any disaster were to occur, our ability to operate our business at 
our facilities would be seriously, or potentially completely, impaired. In addition, the nature of our activities could 
cause  significant  delays  in  our  research  programs  and  commercial  activities  and  make  it  difficult  for  us  to 
recover  from  a  disaster.  The  insurance  we  maintain  may  not  be  adequate  to  cover  our  losses  resulting  from 
disasters  or  other  business  interruptions.  Accordingly,  an  earthquake  or  other  disaster  could  materially  and 
adversely harm our ability to conduct business.

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Risks Related to Our Intellectual Property

Failure  to  secure  patent  or  other  intellectual  property  protection  for  our  products  and  improvements  to  our 
products  may  reduce  our  ability  to  maintain  any  technological  or  competitive  advantage  over  our  current  and 
potential competitors.

Our ability to protect and enforce our intellectual property rights is uncertain and depends on complex legal and 
factual  questions.  Our  ability  to  establish  or  maintain  a  technological  or  competitive  advantage  over  our 
competitors may be diminished because of these uncertainties. For example:

• we or our licensors might not have been the first to make the inventions covered by each of our pending 

patent applications or issued patents;

• we or our licensors might not have been the first to file patent applications for these inventions;

•

•

•

•

it  is  possible  that  neither  our  pending  patent  applications  nor  the  pending  patent  applications  of  our 
licensors will result in issued patents;

the scope of the patent protection we or our licensors obtain may not be sufficiently broad to prevent 
others  from  practicing  our  technologies,  developing  competing  products,  designing  around  our 
patented technologies or independently developing similar or alternative technologies;

our and our licensors’ patent applications or patents have been, are and may in the future be, subject to 
interference,  opposition  or  similar  administrative  proceedings,  which  could  result  in  those  patent 
applications failing to issue as patents, those patents being held invalid or the scope of those patents 
being substantially reduced;

our  enforcement  of  patents  and  proprietary  rights  in  other  countries  may  be  problematic  or 
unpredictable;

• we may not be able to prevent third parties from practicing our inventions in all countries outside the 
United  States,  or  from  selling  or  importing  products  made  using  our  inventions  in  and  into  the  United 
States or other jurisdictions;

• we or our partners may not adequately protect our trade secrets;

• we may not develop additional proprietary technologies that are patentable; or

•

the  patents  of  others  may  limit  our  freedom  to  operate  and  prevent  us  from  commercializing  our 
technology in accordance with our plans.

The occurrence of any of these events could impair our ability to operate without infringing upon the proprietary 
rights of others or prevent us from establishing or maintaining a competitive advantage over our competitors.

Variability in intellectual property laws may adversely affect our intellectual property position.

Intellectual  property  laws,  and  patent  laws  and  regulations  in  particular,  have  been  subject  to  significant 
variability  either  through  administrative  or  legislative  changes  to  such  laws  or  regulations  or  changes  or 
differences in judicial interpretation, and it is expected that such variability will continue to occur. Additionally, 
intellectual property laws and regulations differ by country. Variations in the patent laws and regulations or in 
interpretations of patent laws and regulations in the United States and other countries may diminish the value of 
our intellectual property and may change the impact of third-party intellectual property on us. Accordingly, we 
cannot predict the scope of the patents that may be granted to us with certainty, the extent to which we will be 
able to enforce our patents against third parties or the extent to which third parties may be able to enforce their 
patents against us.

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Some of the intellectual property that is important to our business is owned by other companies or institutions 
and licensed to us, and changes to the rights we have licensed may adversely impact our business.

We  license  from  third  parties  some  of  the  intellectual  property  that  is  important  to  our  business.  If  the  third 
parties who license intellectual property to us fail to maintain the intellectual property that we have licensed, or 
lose rights to that intellectual property, the rights we have licensed may be reduced or eliminated, which would 
eliminate  barriers  against  our  competition.  Termination  of  these  licenses  or  reduction  or  elimination  of  our 
licensed  rights  may  result  in  our  having  to  negotiate  new  or  reinstated  licenses  with  less  favorable  terms,  or 
could  subject  us  to  claims  of  intellectual  property  infringement  or  contract  breach  in  litigation  or  other 
administrative proceedings that could result in damage awards against us and injunctions that could prohibit us 
from selling our products. In addition, some of our licenses from third parties limit the field in which we can use 
the  licensed  technology.  Therefore,  in  order  for  us  to  use  such  licensed  technology  in  potential  future 
applications that are outside the licensed field of use, we may be required to negotiate new licenses with our 
licensors or expand our rights under our existing licenses. We cannot be certain that we will be able to obtain 
such licenses or expanded rights on reasonable terms or at all. In the event a dispute with our licensors were to 
occur, our licensors may seek to renegotiate the terms of our licenses, increase the royalty rates that we pay to 
obtain and maintain those licenses, limit the field or scope of the licenses, or terminate the license agreements. 
In addition, we have limited rights to participate in the prosecution and enforcement of the patents and patent 
applications  that  we  have  licensed.  If  we  fail  to  meet  our  obligations  under  these  licenses,  or  if  we  have  a 
dispute regarding the terms of the licenses, these third parties could terminate the licenses, which could subject 
us  to  claims  of  intellectual  property  infringement.  As  a  result,  we  cannot  be  certain  that  these  patents  and 
applications  will  be  prosecuted  and  enforced  in  a  manner  consistent  with  the  best  interests  of  our  business. 
Further,  because  of  the  rapid  pace  of  technological  change  in  our  industry,  we  may  need  to  rely  on  key 
technologies developed or licensed by third parties, and we may not be able to obtain licenses and technologies 
from  these  third  parties  at  all  or  on  reasonable  terms.  The  occurrence  of  these  events  may  have  a  material 
adverse effect on our business, financial condition or results of operations.

The measures that we use to protect the security of and enforce our intellectual property and other proprietary 
rights may not be adequate, which could result in the loss of legal protection for, and thereby diminish the value 
of, such intellectual property and other rights.

In addition to patents, we also rely upon trademarks, trade secrets, copyrights, and unfair competition laws, as 
well  as  license  agreements  and  other  contractual  provisions,  to  protect  our  intellectual  property  and  other 
proprietary  rights.  Despite  these  measures,  any  of  our  intellectual  property  rights  could  be  challenged, 
invalidated,  circumvented,  or  misappropriated.  In  addition,  we  attempt  to  protect  our  intellectual  property  and 
proprietary information by requiring our employees and consultants to enter into confidentiality and assignment 
of  inventions  agreements,  and  by  entering  into  confidentiality  agreements  with  our  third-party  development, 
manufacturing,  sales,  and  distribution  partners,  who  may  also  acquire,  develop  and/or  commercialize 
alternative  or  competing  products  or  provide  services  to  our  competitors.  For  example,  Roche  had  certain 
access  to  our  trade  secrets  and  other  proprietary  information  pursuant  to  an  agreement  we  had  entered  into 
with Roche, subject to the confidentiality provisions thereof (certain of which provisions survive the termination 
of the agreement); however, Roche is developing potentially competing sequencing products. There can be no 
assurance that our measures have provided or will provide adequate protection for our intellectual property and 
proprietary information. These agreements may be breached, and we may not have adequate remedies for any 
such  breach.  In  addition,  our  trade  secrets  and  other  proprietary  information  may  be  disclosed  to  others,  or 
others  may  gain  access  to  or  disclose  our  trade  secrets  and  other  proprietary  information.  Enforcing  a  claim 
that  a  third  party  illegally  obtained  and  is  using  our  trade  secrets  is  expensive  and  time  consuming,  and  the 
outcome  is  unpredictable.  Additionally,  others  may  independently  develop  proprietary  information  and 
techniques  that  are  substantially  equivalent  to  ours.  The  occurrence  of  these  events  may  have  a  material 
adverse effect on our business, financial condition, or results of operations.

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Our  intellectual  property  may  be  subject  to  challenges  in  the  United  States  or  foreign  jurisdictions  that  could 
adversely affect our intellectual property position.

Our pending, issued and granted U.S. and foreign patents and patent applications have been, are and may in the 
future be, subject to challenges by ONT Ltd., ONT Inc. and Metrichor, Ltd. (“Metrichor” and, together with ONT 
Ltd. and ONT Inc., “ONT”) in addition to other parties asserting prior invention by others or invalidity on various 
grounds,  through  proceedings,  such  as  interferences,  reexaminations,  or  opposition  proceedings.  Addressing 
these  challenges  to  our  intellectual  property  has  been,  and  any  future  challenges  can  be,  costly  and  distract 
management’s  attention  and  resources.  For  example,  we  previously  incurred  significant  legal  expenses  to 
litigate and settle a complaint seeking review of a patent interference decision of the U.S. Patent and Trademark 
Office.  Additionally,  ONT  previously  requested  that  the  U.S.  Patent  and  Trademark  Office  institute  inter  partes 
reviews  of  certain  patents  that  we  have  asserted  against  ONT  Inc.  and  ONT  Ltd.  in  litigation  proceedings  for 
patent infringement. While none of the inter partes reviews requested by ONT were instituted by the U.S. Patent 
and Trademark Office, challenges of this nature before the Patent Trial and Appeal Board (“PTAB”) in the future 
could  result  in  determinations  that  our  patents  or  pending  patent  applications  are  unpatentable  to  us,  or  are 
invalidated or unenforceable in whole or in part and could require us to expend significant time, funds, and other 
resources  in  litigating  such  challenges.  Accordingly,  adverse  rulings  in  such  proceedings  could  negatively 
impact the scope of our intellectual property protection for our products and technology and could materially 
and  adversely  affect  our  business.  Similar  mechanisms  for  challenging  the  validity  and  enforceability  of  a 
patent exist in foreign patent offices and courts and may result in the revocation, cancellation, or amendment of 
any  foreign  patents  we  hold  now  or  in  the  future.  The  outcome  following  legal  assertions  of  invalidity  and 
unenforceability is unpredictable, and prior art could render our patents invalid. If a defendant were to prevail on 
a legal assertion of invalidity and/or unenforceability, we would lose at least part, and perhaps all, of the patent 
protection  on  such  products.  Such  a  loss  of  patent  protection  would  have  a  material  adverse  impact  on  our 
business.

Some of our technology is subject to “march-in” rights by the U.S. government.

Some of our patented technology was developed with U.S. federal government funding. When new technologies 
are  developed  with  U.S.  government  funding,  the  government  obtains  certain  rights  in  any  resulting  patents, 
including a nonexclusive license authorizing the government to use the invention for non-commercial purposes. 
These rights may permit the government to disclose our confidential information to third parties and to exercise 
“march-in” rights to use or allow third parties to use our patented technology. The government can exercise its 
march-in  rights  if  it  determines  that  such  action  is  necessary  to  (i)  achieve  practical  application  of  the  U.S. 
government-funded  technology,  (ii)  alleviate  health  or  safety  needs,  (iii)  meet  requirements  of  federal 
regulations,  or  (iv)  give  preference  to  U.S.  industry.  In  addition,  U.S.  government-funded  inventions  must  be 
reported  to  the  government  and  such  government  funding  must  be  disclosed  in  any  resulting  patent 
applications.  Furthermore,  our  rights  in  such  inventions  are  subject  to  government  license  rights  and  foreign 
manufacturing restrictions. The U.S. government has generally denied requests to exercise its march-in rights, 
even to provide access to potentially life-saving medications; however, if the U.S. government were to exercise 
its march-in rights to our patent technologies funded by the U.S. government, particularly for the benefit of one 
of more of our competitors, that may have a material adverse effect on our business.

We are involved in legal proceedings to enforce our intellectual property rights.

Our intellectual property rights involve complex factual, scientific, and legal questions. We operate in an industry 
characterized  by  significant  intellectual  property  litigation.  Even  though  we  may  believe  that  we  have  a  valid 
patent on a particular technology, other companies have from time to time taken, and may in the future take, 
actions that we believe violate our patent rights. For example, we were previously involved in legal proceedings 
with  ONT  and  Harvard  University  in  several  United  States  and  European  jurisdictions.  We  have  in  the  past 
received  adverse  rulings  against  us  with  respect  to  our  complaint  with  the  United  States  International  Trade 
Commission  for  one  of  these  proceedings.  Legal  actions  to  enforce  our  patent  rights  have  been,  and  will 
continue  to  be,  expensive,  and  may  divert  significant  management  time  and  resources.  Adverse  parties  from 
previous legal actions have brought, and they and others may in the future bring, claims against us and/or our 
intellectual property. Litigation is a significant ongoing expense, recognized in sales, general and administrative 
expense,  with  an  uncertain  outcome,  and  has  been,  and  may  in  the  future  be,  a  material  expense  for  us.  Our 
enforcement actions may not be successful, have given rise to legal claims against us and could result in some 
of  our  intellectual  property  rights  being  determined  to  be  invalid  or  not  enforceable.  Furthermore,  an  adverse 
determination  or  judgement  could  lead  to  an  award  of  damages  against  us,  or  the  issuance  of  an  injunction 
against us or our products that could prevent us from selling any products found to be infringing the intellectual 
property rights of another party.

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We have been, are currently, and could in the future be, subject to legal proceedings with third parties who may 
claim that our products infringe or misappropriate their intellectual property rights.

Our  products  are  based  on  complex,  rapidly  developing  technologies.  We  may  not  be  aware  of  issued  or 
previously filed patent applications that belong to third parties that mature into issued patents that cover some 
aspect of our products or their use. In addition, because patent litigation is complex and the outcome inherently 
uncertain,  our  belief  that  our  products  do  not  infringe  third-party  patents  of  which  we  are  aware  or  that  such 
third-party  patents  are  invalid  and  unenforceable  may  be  determined  to  be  incorrect.  As  a  result,  third  parties 
have  claimed,  and  may  in  the  future  claim,  that  we  infringe  their  patent  rights  and  have  filed,  and  may  in  the 
future file lawsuits or engage in other proceedings against us to enforce their patent rights. For example, we are 
involved  in  legal  proceedings  for  alleged  patent  infringement  and  related  matters  in  the  United  States  with 
Personal Genomics of Taiwan, Inc. (“PGI”), Take2 Technologies, Ltd., and the Chinese University of Hong Kong. 
In  addition,  ONT  Ltd.  and  Harvard  University  have,  in  the  past,  filed  claims  against  us  in  the  High  Court  of 
England  and  Wales  and  the  District  Court  of  Mannheim,  Germany  for  patent  infringement,  and  PGI  has  filed 
claims against us in the U.S. District Court for the District of Delaware and in the Wuhan People’s Court in China. 
We are aware of other issued patents and patent applications owned by third parties that could be construed to 
read  on  our  products,  and  related  maintenance  and  support  services.  Although  we  do  not  believe  that  our 
products or services infringe any valid issued patents, the third-party owners of these patents and applications 
may in the future claim that we infringe their patent rights and file lawsuits against us. In addition, as we enter 
new  markets,  our  competitors  and  other  third  parties  may  claim  that  our  products  infringe  their  intellectual 
property rights as part of a business strategy to impede our successful entry into those markets. Furthermore, 
parties making claims against us may be able to obtain injunctive or other relief, which effectively could block 
our ability to further develop or commercialize products or services and could result in the award of substantial 
damages  against  us.  Patent  litigation  between  competitors  in  our  industry  is  common.  Additionally,  we  have 
certain  obligations  to  many  of  our  customers  and  suppliers  to  indemnify  and  defend  them  against  claims  by 
third parties that our products or their use infringe any intellectual property of these third parties. In defending 
ourselves  against  any  of  these  claims,  we  have  in  the  past  incurred,  and  could  in  the  future  incur,  to  defend 
ourselves  or  our  customers,  substantial  costs,  and  the  attention  of  our  management  and  technical  personnel 
could  be  diverted.  For  example,  we  previously  incurred  significant  legal  expenses  to  litigate  and  settle  a 
complaint alleging patent infringement. Even if we have an agreement that indemnifies us against such costs, 
the  indemnifying  party  may  be  unable  to  uphold  its  contractual  obligations.  To  avoid  or  settle  legal  claims,  it 
may  be  necessary  or  desirable  in  the  future  to  obtain  licenses  relating  to  one  or  more  products  or  relating  to 
current or future technologies, which could negatively affect our gross margins. We may not be able to obtain 
these licenses on commercially reasonable terms, or at all. We may be unable to modify our products so that 
they do not infringe the intellectual property rights of third parties. In some situations, the results of litigation or 
settlement of claims may require us to cease allegedly infringing activities which could prevent us from selling 
some  or  all  of  our  products.  The  occurrence  of  these  events  may  have  a  material  adverse  effect  on  our 
business, financial condition, or results of operations.

In addition, in the course of our business, we may from time to time have access or be alleged to have access to 
confidential or proprietary information of others, which, though not patented, may be protected as trade secrets. 
Others  could  bring  claims  against  us  asserting  that  we  improperly  used  their  confidential  or  proprietary 
information,  or  that  we  misappropriated  their  technologies  and  incorporated  those  technologies  into  our 
products. A determination that we illegally used the confidential or proprietary information or misappropriated 
technologies of others in our products could result in us paying substantial damage awards or being prevented 
from further developing or selling some or all of our products, which could materially and adversely affect our 
business.

We have not yet registered some of our trademarks in all of our potential markets, and failure to secure those 
registrations could adversely affect our business.

Some of our trademark applications may not be allowed for registration, and our registered trademarks may not 
be maintained or enforced. In addition, in the U.S. Patent and Trademark Office and in comparable agencies in 
many foreign jurisdictions, third parties are given an opportunity to oppose pending trademark applications and 
to  seek  to  cancel  registered  trademarks.  Opposition  or  cancellation  proceedings  may  be  filed  against  our 
trademarks, and our trademarks may not survive such proceedings.

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Our  use  of  “open  source”  software  could  adversely  affect  our  ability  to  sell  our  products  and  subject  us  to 
possible litigation.

A  portion  of  the  products  or  technologies  developed  and/or  distributed  by  us  incorporate  “open  source” 
software, and we may incorporate open source software into other products or technologies in the future. Some 
open  source  software  licenses  require  that  we  disclose  the  source  code  for  any  modifications  to  such  open 
source software that we make and distribute to one or more third parties, and that we license the source code 
for such modifications to third parties, including our competitors, at no cost. We monitor the use of open source 
software in our products to avoid uses in a manner that would require us to disclose or grant licenses under our 
source code that we wish to maintain as proprietary; however, there can be no assurance that such efforts have 
been or will be successful. In some circumstances, distribution of our software that includes or is linked with 
open source software could require that we disclose and license some or all of our proprietary source code in 
that software, which could include permitting the use of such software and source code at no cost to the user. 
Open source license terms are often ambiguous and there is little legal precedent governing the interpretation 
of these licenses. Successful claims made by the licensors of open source software that we have violated the 
terms  of  these  licenses  could  result  in  unanticipated  obligations,  including  being  subject  to  significant 
damages, being enjoined from distributing products that incorporate open source software and being required 
to  make  available  our  proprietary  source  code  pursuant  to  an  open  source  license,  which  could  substantially 
help  our  competitors  develop  products  that  are  similar  to  or  better  than  ours  or  otherwise  materially  and 
adversely affect our business.

Risks Related to Regulation

We are, and may become, subject to governmental regulations that may impose burdens on our operations, and 
the markets for our products may be narrowed.

We are subject, both directly and indirectly, to the adverse impact of government regulation of our operations 
and markets. For example, export of our instruments may be subject to strict regulatory control in a number of 
jurisdictions, and we could experience disruption in our supply chain as a result of certain geopolitical events 
and conflicts and any related political or economic responses and counter-responses or otherwise by various 
global  actors.  Following  Russia’s  invasion  of  Ukraine  in  February  2022,  the  United  States  and  other  countries 
imposed certain economic sanctions and severe export control restrictions against Russia and Belarus as well 
as  certain  Russian  nationals  and  individuals  and  entities  with  ties  to  Russia,  Belarus,  and  this  conflict.  These 
sanctions and restrictions have continued to increase as the conflict has further escalated and now cover the 
export of our products to Russia, and the United States and other countries could impose even wider sanctions 
and export restrictions and take other actions in the future that could further limit our ability to provide products 
in certain locations. Additionally, restrictions on the ability to send certain products and technology related to 
semiconductors,  semiconductor  manufacturing,  and  supercomputing  to  China  without  an  export  license  may 
impact  our  ability  to  provide  products  to  customers  or  distributors  in  China.  We  have  expanded  and  are 
continuing  to  expand  the  international  jurisdictions  into  which  we  supply  products,  which  increases  the  risks 
surrounding governmental regulations relating to our business. The need to or failure to satisfy export control 
criteria or to obtain necessary clearances could delay or prevent shipment of products, which could materially 
and  adversely  affect  our  revenue  and  profitability.  Moreover,  the  life  sciences  industry,  which  is  expected  to 
continue to be one of the primary markets for our technology, has historically been heavily regulated. There are, 
for  example,  laws  in  several  jurisdictions  restricting  research  in  genetic  engineering,  which  may  narrow  our 
markets.  Given  the  evolving  nature  of  this  industry,  legislative  bodies  or  regulatory  authorities  may  adopt 
additional  regulations  that  may  adversely  affect  our  market  opportunities.  Additionally,  if  ethical  and  other 
concerns surrounding the use of genetic information, diagnostics or therapies become widespread, there may 
be less demand for our products.

Our  business  is  also  directly  affected  by  a  wide  variety  of  government  regulations  applicable  to  business 
enterprises generally and to companies operating in the life science industry in particular. Failure to comply with 
government regulations or obtain or maintain necessary permits and licenses could result in a variety of fines or 
other censures or an interruption in our business operations which may have a negative impact on our ability to 
generate  revenue  and  the  cost  of  operating  our  business.  In  addition,  changes  to  laws  and  government 
regulations  could  cause  a  material  adverse  effect  on  our  business  as  we  will  need  to  adapt  our  business  to 
comply with such changes. For example, a governmental prohibition on the use of human in vitro diagnostics or 
other  regulations  that  negatively  impact  the  research  and  development  activities  of  our  customers  would 
adversely  impact  our  commercialization  of  products  on  which  we  have  expended  significant  research  and 
development resources, which would in turn have a material adverse impact on our business and prospects.

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Our  products  could  become  subject  to  government  regulation  as  medical  devices  by  the  U.S.  Food  and  Drug 
Administration  or  other  domestic  and  international  regulatory  agencies  even  if  we  do  not  elect  to  seek 
regulatory clearance or approval to market our products for diagnostic purposes, which could increase our costs 
and impede or delay our commercialization efforts, thereby materially and adversely affecting our business and 
results of operations.

Our products are currently labeled and promoted as research use only (“RUO”) products, and are not currently 
designed,  or  intended  to  be  used,  for  clinical  diagnostic  tests  or  as  medical  devices.  However,  in  the  future, 
certain of our products or related applications, such as those that may be developed for clinical uses, could be 
subject to regulation by the U.S. Food and Drug Administration (“FDA”), or the FDA’s regulatory jurisdiction could 
be expanded to include our products. Also, even if our products are labeled, promoted, and intended as RUO, the 
FDA  or  comparable  agencies  of  other  countries  could  disagree  with  our  conclusion  that  our  products  are 
intended for research use only or deem our sales, marketing and promotional efforts as being inconsistent with 
the  FDA’s  guidance  on  RUO  products.  For  example,  our  customers  may  independently  elect  to  use  our  RUO 
labeled  products  in  their  own  laboratory  developed  tests  (“LDTs”)  for  clinical  diagnostic  use,  which  could 
subject  our  products  to  government  regulation,  and  the  regulatory  clearance  or  approval  and  maintenance 
process for such products may be uncertain, expensive, and time-consuming. Regulatory requirements related 
to marketing, selling, and distribution of RUO products could change or be uncertain, even if clinical uses of our 
RUO products by our customers were done without our consent. If the FDA or other regulatory authorities assert 
that any of our RUO products are subject to regulatory clearance or approval, our business, financial condition, 
or results of operations could be adversely affected. In the event that we fail to obtain and maintain necessary 
regulatory clearances or approvals for products that we develop for clinical uses, or if clearances or approvals 
for  future  products  and  indications  are  delayed  or  not  issued,  our  commercial  operations  may  be  materially 
harmed.  Furthermore,  even  if  we  are  granted  regulatory  clearances  or  approvals,  they  may  include  significant 
limitations on the indicated uses for the product, which may limit the market for the product. We do not have 
experience  in  obtaining  FDA  approvals  and  no  assurance  can  be  given  that  we  will  be  able  to  obtain  or  to 
maintain such approvals. Furthermore, any approvals that we may obtain can be revoked if safety or efficacy 
problems develop.

The  FDA  has  historically  exercised  enforcement  discretion  in  not  enforcing  the  medical  device  regulations 
against  laboratories  developing  and  offering  LDTs.  In  2017,  the  FDA  announced  that  it  would  not  issue  final 
guidance on the oversight of LDTs, and manufacturers of products used for LDTs, but would seek further public 
discussion  on  an  appropriate  oversight  approach,  and  give  Congress  an  opportunity  to  develop  a  legislative 
solution. The FDA has issued warning letters to certain genomics labs for illegally marketing genetic tests that 
claim to predict patients’ responses to specific medications, noting that the FDA has not created a legal “carve-
out” for LDTs and retains discretion to take action when appropriate, such as when certain genomic tests raise 
significant public health concerns.

As  manufacturers  develop  more  complex  diagnostic  tests  and  diagnostic  software,  the  FDA  may  increase  its 
regulation  of  LDTs.  Any  future  legislative  or  administrative  rule  making  or  oversight  of  LDTs,  if  and  when 
finalized,  may  impact  the  sales  of  our  products  and  how  customers  use  our  products,  and  may  require  us  to 
change  our  business  model  in  order  to  maintain  compliance  with  these  laws.  We  cannot  predict  how  these 
various efforts will be resolved, how Congress or the FDA will regulate LDTs in the future, or how that regulatory 
system  will  impact  our  business.  Changes  to  the  current  regulatory  framework,  including  the  imposition  of 
additional  or  new  regulations,  including  regulation  of  our  products,  could  arise  at  any  time  during  the 
development or marketing of our products, which may negatively affect our ability to obtain or maintain FDA or 
comparable regulatory approval of our products, if required. Further, sales of devices for diagnostic purposes 
may  subject  us  to  additional  healthcare  regulation  and  enforcement  by  the  applicable  government  agencies. 
Such laws include, without limitation, state and federal anti-kickback or anti-referral laws, healthcare fraud and 
abuse  laws,  false  claims  laws,  privacy  and  security  laws,  Physician  Payments  Sunshine  Act  and  related 
transparency  and  manufacturer  reporting  laws,  and  other  laws  and  regulations  applicable  to  medical  device 
manufacturers.

Additionally,  in  2013,  the  FDA  issued  Final  Guidance  “Distribution  of  In  Vitro  Diagnostic  Products  Labeled  for 
Research Use Only.” The guidance emphasizes that the FDA will review the totality of the circumstances when it 
comes  to  evaluating  whether  equipment  and  testing  components  are  properly  labeled  as  RUO.  The  final 
guidance states that merely including a labeling statement that the product is for research purposes only will 
not necessarily render the device exempt from the FDA’s clearance, approval, and other regulatory requirements 
if  the  circumstances  surrounding  the  distribution,  marketing  and  promotional  practices  indicate  that  the 
manufacturer  knows  its  products  are,  or  intends  for  its  products  to  be,  used  for  clinical  diagnostic  purposes. 

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These circumstances may include written or verbal sales and marketing claims or links to articles regarding a 
product’s  performance  in  clinical  applications  and  a  manufacturer’s  provision  of  technical  support  for  clinical 
applications.

In August 2020, the Department of Health and Human Services ("HHS") announced rescission of guidance and 
other  informal  issuances  of  FDA  regarding  pre-market  review  of  LDT  absent  notice-and-comment  rulemaking, 
stating  that,  absent  notice-and-comment  rulemaking  LDTs  are  not  required  to  obtain  FDA  pre-market 
authorization.  In  November  2021,  HHS  under  the  Biden  administration  issued  a  statement  that  withdrew  the 
August 2020 policy announcement stating that HHS does not have a policy on LDTs that is separate from FDA’s 
longstanding approach.

Legislative and administrative proposals to amend the FDA's oversight of LDTs have been introduced in recent 
years,  including  the  Verifying  Accurate  Leading-edge  IVCT  Development  Act  of  2021  (the  "VALID  Act"),  which 
aims to create a new category of medical products separate from medical devices called “in vitro clinical tests,” 
or IVCTs, and bring all such products within the scope of the FDA’s oversight. To date, Congress has not passed 
the VALID Act, but may revisit the VALID Act or similar policy riders and enact other FDA programmatic reforms 
in  the  future.  In  October  2023,  through  rulemaking,  the  FDA  proposed  to  amend  the  definition  of  "in  vitro 
diagnostic products" in FDA regulations to include laboratories that manufacture such products and to phase 
out  the  FDA's  general  enforcement  discretion  approach  for  LDTs  so  that  IVDs  manufactured  by  a  laboratory 
would be regulated similarly to IVDs. It is unclear when the FDA will finalize and begin enforcing such rule and 
how future legislation by federal and state governments and FDA regulation will impact the industry, including 
our business and that of our customers.

If the FDA determines our products or related applications should be subject to additional regulation as in vitro 
diagnostic devices based upon customers’ use of our products for clinical diagnostic or therapeutic decision-
making  purposes,  our  ability  to  market  and  sell  our  products  could  be  impeded  and  our  business,  prospects, 
results of operations and financial condition may be adversely affected. In addition, the FDA could consider our 
products to be misbranded or adulterated under the Federal Food, Drug, and Cosmetic Act and subject to recall 
and/or other enforcement action.

To the extent we elect to label and promote any of our products as medical devices, we would be required to 
obtain  prior  approval  or  clearance  by  the  FDA  or  comparable  foreign  regulatory  authority,  which  could  take 
significant  time  and  expense  and  could  fail  to  result  in  a  marketing  authorization  for  the  intended  uses  we 
believe are commercially attractive. Obtaining marketing authorization in one jurisdiction does not mean that we 
will be successful in obtaining marketing authorization in other jurisdictions where we conduct business.

If  we  elect  to  label  and  market  our  products  for  use  as,  or  in  the  performance  of,  clinical  diagnostics  in  the 
United  States,  thereby  subjecting  them  to  FDA  regulation  as  medical  devices,  we  would  be  required  to  obtain 
pre-market 510(k) clearance or pre-market approval from the FDA, unless an exception applies. It is possible, in 
the  event  we  elect  to  submit  510(k)  applications  for  certain  of  our  products,  that  the  FDA  would  take  the 
position that a more burdensome pre-market application, such as a PMA or a de novo application is required for 
some  of  our  products.  If  such  applications  were  required,  greater  time  and  investment  would  be  required  to 
obtain  FDA  approval.  Even  if  the  FDA  agreed  that  a  510(k)  was  appropriate,  FDA  clearance  can  be  expensive 
and time consuming. It generally takes a significant amount of time to prepare a 510(k), including conducting 
appropriate  testing  on  our  products,  and  several  months  to  years  for  the  FDA  to  review  a  submission. 
Notwithstanding  the  effort  and  expense,  FDA  clearance  or  approval  could  be  denied  for  some  or  all  of  our 
products for which we choose to market as a medical device or a clinical diagnostic device. Even if we were to 
seek  and  obtain  regulatory  approval  or  clearance,  it  may  not  be  for  the  intended  uses  we  request  or  that  we 
believe are important or commercially attractive. There can be no assurance that future products for which we 
may  seek  pre-market  clearance  or  approval  will  be  approved  or  cleared  by  FDA  or  a  comparable  foreign 
regulatory authority on a timely basis, if at all, nor can there be assurance that labeling claims will be consistent 
with our anticipated claims or adequate to support continued adoption of such products. Compliance with FDA 
or  comparable  foreign  regulatory  authority  regulations  will  require  substantial  costs,  and  subject  us  to 
heightened scrutiny by regulators and substantial penalties for failure to comply with such requirements or the 
inability  to  market  our  products.  The  lengthy  and  unpredictable  pre-market  clearance  or  approval  process,  as 
well  as  the  unpredictability  of  the  results  of  any  required  clinical  studies,  may  result  in  our  failing  to  obtain 
regulatory clearance or approval to market such products, which would significantly harm our business, results 
of operations, reputation, and prospects.

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If we sought and received regulatory clearance or approval for certain of our products, we would be subject to 
ongoing  FDA  obligations  and  continued  regulatory  oversight  and  review,  including  the  general  controls  listed 
above  and  the  FDA’s  QSRs  for  our  development  and  manufacturing  operations.  In  addition,  we  would  be 
required  to  obtain  a  new  510(k)  clearance  before  we  could  introduce  subsequent  material  modifications  or 
improvements  to  such  products.  We  could  also  be  subject  to  additional  FDA  post-marketing  obligations  for 
such products, any or all of which would increase our costs and divert resources away from other projects. If we 
sought and received regulatory clearance or approval and are not able to maintain regulatory compliance with 
applicable  laws,  we  could  be  prohibited  from  marketing  our  products  for  use  as,  or  in  the  performance  of, 
clinical  diagnostics  and/or  could  be  subject  to  enforcement  actions,  including  warning  letters  and  adverse 
publicity, fines, injunctions, and civil penalties; recall or seizure of products; operating restrictions; and criminal 
prosecution.

Further, if we decide to seek regulatory clearance or approval for certain of our products in countries outside of 
the  United  States  or  if  a  foreign  regulatory  authority  determines  that  our  products  are  regulated  as  medical 
devices,  we  would  be  subject  to  extensive  medical  device  laws  and  regulations  outside  of  the  United  States. 
Sales of such products outside the United States will likely be subject to foreign regulatory requirements, which 
can vary greatly from country to country. As a result, the time required to obtain clearances or approvals outside 
the United States may differ from that required to obtain FDA clearance or approval and we may not be able to 
obtain  foreign  regulatory  approvals  on  a  timely  basis  or  at  all.  In  Europe,  we  would  need  to  comply  with  the 
Medical Device Regulation 2017/745 and In Vitro Diagnostic Regulation 2017/746, which became effective May 
26, 2017, with application dates of May 26, 2021 (postponed from 2020) and May 26, 2022, respectively. This 
will increase the difficulty of regulatory approvals in Europe in the future. In addition, the FDA regulates exports 
of  medical  devices.  The  number  and  scope  of  these  requirements  are  increasing.  Unlike  many  of  the  other 
companies offering nucleic acid sequencing equipment or consumables, this is an area where we do not have 
expertise.  We,  or  our  other  third-party  sales  and  distribution  partners,  may  not  be  able  to  obtain  regulatory 
approvals  in  such  countries  or  may  incur  significant  costs  in  obtaining  or  maintaining  our  foreign  regulatory 
approvals. In addition, the export by us of certain of our products, which have not yet been cleared for domestic 
commercial  distribution,  may  be  subject  to  FDA  or  other  export  restrictions.  Failure  to  comply  with  these 
regulatory requirements or obtain and maintain required approvals, clearances and certifications could impair 
our  ability  to  commercialize  our  products  for  diagnostic  use  outside  of  the  United  States.  Any  action  brought 
against  us  for  violations  of  these  laws  or  regulations,  even  if  successfully  defended,  could  cause  us  to  incur 
significant legal expenses and divert our management’s attention from the operation of our business.

Enhanced trade tariffs, import restrictions, export restrictions, Chinese regulations, or other trade barriers may 
materially harm our business.

We are continuing to expand our international operations as part of our growth strategy and have experienced 
an  increasing  concentration  of  sales  in  certain  regions  outside  the  United  States,  especially  the  Asia-Pacific 
region, as discussed above. There is currently significant uncertainty about the future relationship between the 
United  States  and  various  other  countries,  most  significantly  China,  with  respect  to  trade  policies,  treaties, 
government  regulations  and  tariffs.  Starting  in  September  2018,  the  U.S.  Trade  Representative  (the  “USTR”) 
enacted  various  tariffs  of  7.5%,  10%,  15%,  and  25%  on  the  import  of  Chinese  products,  including  non-U.S. 
components  and  materials  that  may  be  used  in  our  products.  Additionally,  China  also  has  imposed  tariffs  on 
imports  into  China  from  the  United  States.  These  tariffs  have  and  could  continue  to  raise  our  costs. 
Furthermore,  tariffs,  trade  restrictions,  or  trade  barriers  that  have  been,  and  may  in  the  future  be,  placed  on 
products such as ours by foreign governments, especially China, have raised, and could further raise, amounts 
paid  for  some  or  all  of  our  products,  which  may  result  in  the  loss  of  customers  and  our  business,  and  our 
financial condition and results of operations may be harmed. Further tariffs may be imposed that could cover 
imports  of  additional  components  and  materials  used  in  our  products,  or  our  business  may  be  adversely 
impacted  by  retaliatory  trade  measures  taken  by  China  or  other  countries,  including  restricted  access  to 
components or materials used in our products or increased amounts that must be paid for our products, which 
could materially harm our business, financial condition, and results of operations.

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Additionally,  the  U.S.  government  imposed  controls  restricting  the  ability  to  send  certain  products  and 
technology related to semiconductors, semiconductor manufacturing, and supercomputing to China without an 
export license. These controls also apply to certain hardware containing these specified integrated circuits. In 
many cases, these licenses are subject to a policy of denial and will not be issued. It is possible that additional 
restrictions  will  be  put  in  place.  These  existing  and  future  controls  may  impact  our  ability  to  export  certain 
products to customers or distributors in China or other locations and restrict our ability to use certain integrated 
circuits  in  our  products.  The  U.S.  government  also  continues  to  add  additional  entities  in  China  to  restricted 
party  lists  impacting  the  ability  of  U.S.  companies  to  provide  items  to  these  entities.  Moreover,  in  November 
2018, the U.S. Commerce Department’s Bureau of Industry and Security (“BIS”) released an advance notice of 
proposed  rulemaking  to  control  the  export  of  emerging  technologies.  This  notice  included  “[b]iotechnology, 
including nanobiology; synthetic biology; genomic and genetic engineering; or neurotech” as possible areas of 
increased  export  controls.  The  Biden  Administration  has  continued  to  provide  updated  lists  of  emerging 
technologies  subject  to  national  security  consents.  These  lists  continue  to  include  biotechnologies  including 
“[g]enome  and  protein  engineering  including  design  tools”  and  “[b]iomanufacturing  and  bioprocessing 
technologies.” Therefore, it is possible that our ability to export our products to customers or distributors may 
be further restricted in the future.

It is possible that the Chinese government will retaliate in response to existing or future U.S. export controls or 
trade restrictions in ways that could impact our business. It also is possible that additional restrictions will be 
put in place that could impact our ability to provide our products to customers or distributors in China or source 
components  from  China.  The  continued  threats  of  tariffs,  trade  restrictions  and  trade  barriers  could  have  a 
generally  disruptive  impact  on  the  global  economy  and,  therefore,  negatively  impact  our  sales.  Given  the 
relatively  fluid  regulatory  environment  in  China  and  the  United  States  and  uncertainty  how  the  U.S.  or  foreign 
governments will act with respect to export controls, tariffs, international trade agreements and policies, there 
could be additional tax or other regulatory changes in the future. Any such changes could directly and adversely 
impact our financial results and results of operations.

Our  international  business  could  expose  us  to  business,  regulatory,  political,  operational,  financial,  and 
economic risks associated with doing business outside of the United States.

Engaging in international business inherently involves a number of difficulties and risks, including:

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required  compliance  with  existing  and  changing  foreign  regulatory  requirements  and  laws  that  are  or 
may be applicable to our business in the future, such as the European Union’s General Data Protection 
Regulation  (“GDPR”)  and  other  data  privacy  requirements,  labor  and  employment  regulations,  anti-
competition regulations, the U.K. Bribery Act of 2010 and other anti-corruption laws, regulations relating 
to  the  use  of  certain  hazardous  substances  or  chemicals  in  commercial  products,  and  require  the 
collection, reuse, and recycling of waste from products we manufacture;

required  compliance  with  U.S.  laws  such  as  the  Foreign  Corrupt  Practices  Act,  and  other  U.S.  federal 
laws  and  trade  and  economic  sanctions  and  other  regulations  established  by  the  Office  of  Foreign 
Asset Control;

export requirements and import or trade restrictions;

laws and business practices favoring local companies;

restrictions on both inbound and outbound cross-border investment;

foreign  currency  exchange,  longer  payment  cycles  and  difficulties  in  enforcing  agreements  and 
collecting receivables through certain foreign legal systems;

changes  in  social,  economic,  and  political  conditions  or  in  laws,  regulations  and  policies  governing 
foreign trade, manufacturing, research and development, and investment both domestically as well as 
in  the  other  countries  and  jurisdictions  in  which  we  operate  and  into  which  we  may  sell  our  products 
including  as  a  result  of  the  separation  of  the  United  Kingdom  from  the  European  Union  (“Brexit”)  and 
ongoing  geopolitical  tensions  related  to  the  political  uncertainty  and  military  actions  associated  with 
the war in Ukraine, resulting sanctions imposed by the U.S. and other countries, and retaliatory actions 
taken by Russia in response to such sanctions;

potentially  adverse  tax  consequences,  tariffs,  customs  charges,  bureaucratic  requirements,  and  other 
trade barriers;

difficulties and costs of staffing and managing foreign operations; and

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difficulties  protecting,  maintaining,  enforcing,  or  procuring  intellectual  property  rights  and  defending 
against intellectual property claims under the law and judicial systems of other countries.

If  one  or  more  of  these  risks  occurs,  it  could  require  us  to  dedicate  significant  resources  to  remedy  such 
occurrence, and if we are unsuccessful in finding a solution, our financial results will suffer.

Our  operations  involve  the  use  of  hazardous  materials,  and  we  must  comply  with  environmental,  health  and 
safety  laws,  which  can  be  expensive  and  may  adversely  affect  our  business,  operating  results  and  financial 
condition.

Our research and development and manufacturing activities involve the use of hazardous materials, including 
chemicals and biological materials, and some of our products include hazardous materials. Accordingly, we are 
subject to federal, state, local and foreign laws, regulations, and permits relating to environmental, health and 
safety matters, including, among others, those governing the use, storage, handling, exposure to and disposal of 
hazardous materials and wastes, the health and safety of our employees, and the shipment, labeling, collection, 
recycling,  treatment,  and  disposal  of  products  containing  hazardous  materials.  Liability  under  environmental 
laws  and  regulations  can  be  joint  and  several  and  without  regard  to  fault  or  negligence.  For  example,  under 
certain  circumstances  and  under  certain  environmental  laws,  we  could  be  held  liable  for  costs  relating  to 
contamination at our or our predecessors’ past or present facilities and at third-party waste disposal sites. We 
could also be held liable for damages arising out of human exposure to hazardous materials. There can be no 
assurance  that  violations  of  environmental,  health  and  safety  laws  will  not  occur  as  a  result  of  human  error, 
accident, equipment failure or other causes. The failure to comply with past, present or future laws could result 
in  the  imposition  of  substantial  fines  and  penalties,  remediation  costs,  property  damage  and  personal  injury 
claims,  investigations,  the  suspension  of  production  or  product  sales,  loss  of  permits  or  a  cessation  of 
operations.  Any  of  these  events  could  harm  our  business,  operating  results,  and  financial  condition.  We  also 
expect that our operations will be affected by new environmental, health and safety laws and regulations on an 
ongoing basis, or more stringent enforcement of existing laws and regulations. New laws or changes to existing 
laws  may  result  in  additional  costs  and  may  increase  penalties  associated  with  violations  or  require  us  to 
change the content of our products or how we manufacture them, which could have a material adverse effect 
on our business, operating results, and financial condition.

Ethical, legal, privacy, data protection and social concerns or governmental restrictions surrounding the use of 
genetic information could reduce demand for our technology.

Our  products  may  be  used  to  provide  genetic  information  about  humans,  agricultural  crops  and  other  living 
organisms. The information obtained from our products could be used in a variety of applications which may 
have underlying ethical, legal, privacy, data protection and social concerns, including the genetic engineering or 
modification  of  agricultural  products  or  testing  for  genetic  predisposition  for  certain  medical  conditions. 
Governmental authorities could, for safety, social or other purposes, call for limits on or regulation of the use of 
genetic  testing,  and  may  consider  or  adopt  such  regulations  or  other  restrictions.  Such  concerns  or 
governmental restrictions could limit the use of our products or be costly and burdensome to comply with, and 
actual  or  perceived  violations  of  any  such  restrictions  may  lead  to  the  imposition  of  substantial  fines  and 
penalties, remediation costs, claims and litigation, regulatory investigations and proceedings, and other liability, 
any  of  which  could  have  a  material  adverse  effect  on  our  business,  financial  condition,  and  results  of 
operations.

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Regulations related to conflict minerals has caused us to incur, and will continue to cause us to incur, additional 
expenses and could limit the supply and increase the costs of certain materials used in the manufacture of our 
products.

We are subject to requirements under the Dodd-Frank Wall Street Reform and Consumer Protection Act of 2010 
that require us to  conduct  diligence and report on whether or not our products contain conflict minerals. The 
implementation  of  these  requirements  could  adversely  affect  the  sourcing,  availability  and  pricing  of  the 
materials used in the manufacture of components used in our products. Furthermore, the complex nature of our 
products requires components and materials that may be available only from a limited number of sources and, 
in  some  cases,  from  only  a  single  source.  We  have  incurred,  and  will  continue  to  incur,  additional  costs  to 
comply  with  the  disclosure  requirements,  including  costs  related  to  conducting  diligence  procedures  to 
determine  the  sources  of  conflict  minerals  that  may  be  used  or  necessary  to  the  production  of  our  products 
and, if applicable, potential changes to components, processes, or sources of supply as a consequence of such 
verification  activities.  We  may  face  reputational  harm  if  we  determine  that  certain  of  our  products  contain 
minerals  that  are  not  determined  to  be  conflict  free  or  if  we  are  unable  to  alter  our  processes  or  sources  of 
supply  to  avoid  using  such  materials.  In  such  circumstances,  the  reputational  harm  could  materially  and 
adversely affect our business, financial condition, or results of operations.

Risks Related to Owning Our Common Stock

The price of our common stock has been, is, and may continue to be, highly volatile, and you may be unable to 
sell your shares at or above the price you paid to acquire them.

The  market  price  of  our  common  stock  is  highly  volatile,  and  we  expect  it  to  continue  to  be  volatile  for  the 
foreseeable  future  in  response  to  many  risk  factors  listed  in  this  section,  and  others  beyond  our  control, 
including:

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actual or anticipated fluctuations in our financial condition and operating results;

announcements  of  new  products,  technological  innovations  or  strategic  partnerships  by  us  or  our 
competitors;

announcements  by  us,  our  customers,  partners,  or  suppliers  relating  directly  or  indirectly  to  our 
products, services or technologies;

overall conditions in our industry and market;

addition or loss of significant customers;

changes in laws or regulations applicable to our products;

actual or anticipated changes in our growth rate relative to our competitors;

announcements  by  us  or  our  competitors  of  significant  acquisitions,  strategic  partnerships,  joint 
ventures, capital commitments or achievement of significant milestones;

additions or departures of key personnel;

competition from existing products or new products that may emerge;

issuance of new or updated research or reports by securities analysts;

fluctuations in the valuation of companies perceived by investors to be comparable to us;

disputes or other developments related to proprietary rights, including patents, litigation matters or our 
ability to obtain intellectual property protection for our technologies;

announcement or expectation of additional financing efforts;

sales of our common stock by us or our stockholders;

stock price and volume fluctuations attributable to inconsistent trading volume levels of our shares;

reports, guidance and ratings issued by securities or industry analysts;

operating results below the expectations of securities analysts or investors; and

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general economic and market conditions, which could be impacted by various events including health 
epidemics  or  pandemics,  interest  rate  fluctuations,  increases  in  fuel  prices,  foreign  currency 
fluctuations,  international  tariffs,  acts  of  terrorism,  hostilities  or  the  perception  that  hostilities  may  be 
imminent,  military  conflict  and  acts  of  war,  including  further  political  uncertainty  and  military  actions 
associated  with  the  war  in  Ukraine  and  the  related  response,  including  sanctions  or  other  restrictive 
actions, by the United States and/or other countries.

If  any  of  the  forgoing  occurs,  it  would  cause  our  stock  price  or  trading  volume  to  decline.  Stock  markets  in 
general  and  the  market  for  companies  in  our  industry  in  particular  have  experienced  price  and  volume 
fluctuations,  which  were  exacerbated  by  the  COVID-19  pandemic,  and  current  macroeconomic  trends  and 
geopolitical  events,  and  have  affected  and  continue  to  affect  the  market  prices  of  equity  securities  of  many 
companies. These fluctuations often have been unrelated or disproportionate to the operating performance of 
those  companies.  These  broad  market  and  industry  fluctuations,  as  well  as  general  economic,  political  and 
market  conditions  such  as  recessions,  interest  rate  changes  or  international  currency  fluctuations,  may 
negatively impact the market price of our common stock. You may not realize any return on your investment in 
us and may lose some or all of your investment. In the past, companies that have experienced volatility in the 
market price of their stock have been subject to securities class action litigation. We have been a party to this 
type  of  litigation  in  the  past  and  may  be  the  target  of  this  type  of  litigation  again  in  the  future.  Securities 
litigation  against  us  could  result  in  substantial  costs  and  divert  our  management’s  attention  from  other 
business concerns, which could seriously harm our business.

Sales  of  substantial  amounts  of  our  common  stock  in  the  public  markets,  or  the  perception  that  such  sales 
might occur, could reduce the market price that our common stock might otherwise attain and may dilute your 
voting power and your ownership interest in us.

Sales of a substantial number of shares of our common stock in the public market, or the perception that such 
sales could occur, could adversely affect the market price of our common stock and may make it more difficult 
for existing stockholders to sell their common stock at a time and price that they deem appropriate and may 
dilute their voting power and ownership interest in us.

In addition, if our stockholders sell, or indicate an intent to sell, a large number of shares of our common stock 
in  the  public  market,  it  could  cause  our  stock  price  to  fall,  particularly  if  such  sales  occur  over  a  short  period 
time (for example, following delivery of shares upon achievement of milestones in our acquisition agreements). 
We may also issue shares of common stock or securities convertible into our common stock in connection with 
a financing, acquisition, our equity incentive plans, or otherwise. Any such issuances would result in dilution to 
our existing stockholders and the market price of our common stock may be adversely affected.

Concentration  of  ownership  by  our  principal  stockholders  may  result  in  control  by  such  stockholders  of  the 
composition of our board of directors.

Our  existing  principal  stockholders,  executive  officers,  directors,  and  their  affiliates  beneficially  own  a 
significant number of our outstanding shares of common stock. In addition, such parties may acquire additional 
control by purchasing stock that we issue in connection with our future fundraising efforts. These parties may 
now  and  in  the  future  be  able  to  exercise  a  significant  level  of  control  over  all  matters  requiring  stockholder 
approval,  including  the  election  of  directors.  This  control  could  have  the  effect  of  delaying  or  preventing  a 
change  of  control  of  our  company  or  changes  in  management  and  will  make  the  approval  of  certain 
transactions difficult or impossible without the support of these stockholders.

Anti-takeover  provisions  in  our  charter  documents  and  under  Delaware  law  could  make  an  acquisition  of  us, 
which  may  be  beneficial  to  our  stockholders,  more  difficult  and  may  prevent  attempts  by  our  stockholders  to 
replace or remove our current management and limit the market price of our common stock.

Provisions  in  our  amended  and  restated  certificate  of  incorporation  and  amended  and  restated  bylaws  may 
have the effect of delaying or preventing a change of control or changes in our management. Our amended and 
restated certificate of incorporation and amended and restated bylaws include provisions that:

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authorize our board of directors to issue, without further action by the stockholders, up to 50,000,000 
shares of undesignated preferred stock and up to approximately 1,000,000,000 shares of authorized but 
unissued shares of common stock;

require  that  any  action  to  be  taken  by  our  stockholders  be  effected  at  a  duly  called  annual  or  special 
meeting and not by written consent;

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specify that special meetings of our stockholders can be called only by our board of directors, the Chair 
of the Board, the Chief Executive Officer or the President;

establish advance notice procedures for stockholder approvals to be brought before an annual meeting 
of our stockholders, including proposed nominations of persons for election to our board of directors;

establish  that  our  board  of  directors  is  divided  into  three  classes,  Class  I,  Class  II  and  Class  III,  with 
each class serving staggered terms;

provide that our directors may be removed only for cause; and

provide  that  vacancies  on  our  board  of  directors  may  be  filled  only  by  a  majority  of  directors  then  in 
office, even though less than a quorum.

These provisions may frustrate or prevent any attempts by our stockholders to replace or remove our current 
management by making it more difficult for stockholders to replace members of our board of directors, which is 
responsible  for  appointing  the  members  of  our  management.  In  addition,  because  we  are  incorporated  in 
Delaware,  we  are  governed  by  the  provisions  of  Section  203  of  the  Delaware  General  Corporation  Law,  which 
limits the ability of stockholders owning in excess of 15% of our outstanding voting stock to merge or combine 
with us.

Our amended and restated bylaws designate a state or federal court located within the State of Delaware as the 
exclusive forum for certain stockholder litigation matters, and also provide that the federal district courts will be 
the exclusive forum for resolving any complaint asserting a cause of action arising under the Securities Act of 
1933, as amended, each of which could limit our stockholders’ ability to choose the judicial forum for disputes 
with us or our directors, officers, stockholders, or employees.

Our amended and restated bylaws provide that unless we consent in writing to the selection of an alternative 
forum, the Court of Chancery of the State of Delaware (or, if the Court of Chancery does not have jurisdiction, 
another State court in Delaware or the federal district court for the District of Delaware) will, to the fullest extent 
permitted  by  law,  be  the  sole  and  exclusive  forum  for:  (i)  any  derivative  action  or  proceeding  brought  on  our 
behalf;  (ii)  any  action  asserting  a  claim  of  breach  of  a  fiduciary  duty  owed  by  any  of  our  current  or  former 
directors, stockholders, officers, or other employees to us or our stockholders; (iii) any action arising pursuant 
to  any  provision  of  the  Delaware  General  Corporation  Law;  (iv)  any  action  to  interpret,  apply,  enforce  or 
determine  the  validity  of  our  amended  and  restated  certificate  of  incorporation  or  our  amended  and  restated 
bylaws;  or  (v)  any  action  asserting  a  claim  governed  by  the  internal  affairs  doctrine,  except  as  to  each  of  (i) 
through  (v)  above,  for  any  claim  as  to  which  such  court  determines  that  there  is  an  indispensable  party  not 
subject to the jurisdiction of such court.

Section  22  of  the  Securities  Act  creates  concurrent  jurisdiction  for  federal  and  state  courts  over  all  such 
Securities Act actions. Accordingly, both state and federal courts have jurisdiction to entertain such claims. To 
prevent  having  to  litigate  claims  in  multiple  jurisdictions  and  the  threat  of  inconsistent  or  contrary  rulings  by 
different  courts,  among  other  considerations,  our  amended  and  restated  bylaws  also  provide  that,  unless  we 
consent  in  writing  to  the  selection  of  an  alternative  forum,  the  federal  district  courts  of  the  United  States  of 
America will be the exclusive forum for resolving any complaint asserting a cause of action arising under the 
Securities  Act  including,  without  limitation  and  for  the  avoidance  of  doubt,  any  auditor,  underwriter,  expert, 
control person or other defendant.

Any  person  or  entity  purchasing,  holding  or  otherwise  acquiring  any  interest  in  any  of  our  securities  shall  be 
deemed to have notice of and consented to the foregoing bylaw provisions. Although we believe these exclusive 
forum provisions benefit us by providing increased consistency in the application of Delaware law and federal 
securities  laws  in  the  types  of  lawsuits  to  which  each  applies,  the  exclusive  forum  provisions  may  limit  a 
stockholder’s  ability  to  bring  a  claim  in  a  judicial  forum  of  its  choosing  for  disputes  with  us  or  any  of  our 
directors, stockholders, officers or other employees, which may discourage lawsuits with respect to such claims 
against  us  and  our  current  and  former  directors,  stockholders,  officers  or  other  employees.  In  addition,  a 
stockholder  that  is  unable  to  bring  a  claim  in  the  judicial  forum  of  its  choosing  may  be  required  to  incur 
additional costs in the pursuit of actions which are subject to the exclusive forum provisions described above. 
Our  stockholders  will  not  be  deemed  to  have  waived  our  compliance  with  the  federal  securities  laws  and  the 
rules and regulations thereunder as a result of our exclusive forum provisions. Further, in the event a court finds 
either  exclusive  forum  provision  contained  in  our  bylaws  to  be  unenforceable  or  inapplicable  in  an  action,  we 
may  incur  additional  costs  associated  with  resolving  such  action  in  other  jurisdictions,  which  could  harm  our 
results of operations.

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Our  large  number  of  authorized  but  unissued  shares  of  common  stock  may  potentially  dilute  existing 
stockholders’ stockholdings.

We have a significant number of authorized but unissued shares of common stock. Our board of directors may 
issue shares of common stock from this authorized but unissued pool from time to time without stockholder 
approval, resulting in the dilution of our existing stockholders.

We do not intend to pay dividends for the foreseeable future.

We have never declared or paid any dividends on our common stock and do not intend to pay any dividends in 
the foreseeable future. We anticipate that we will retain all of our future earnings for use in the operation of our 
business  and  for  general  corporate  purposes.  Any  determination  to  pay  dividends  in  the  future  will  be  at  the 
discretion of our board of directors. Accordingly, investors must rely on sales of their common stock after price 
appreciation, which may never occur, as the only way to realize any future gains on their investments.

Risks Related to Our Notes

We  may  not  have  the  ability  to  raise  the  funds  necessary  to  settle  conversions  of  the  Notes  in  cash  or  to 
repurchase the Notes upon a fundamental change, and our future debt may contain limitations on our ability to 
pay cash upon conversion or repurchase of the Notes.

As of December 31, 2023, we had outstanding approximately $459.0 million aggregate principal amount of our 
2028 Notes and $441.0 million aggregate principal amount of our 2030 Notes. The 2028 Notes will mature on 
February  15,  2028,  subject  to  earlier  conversion,  redemption  or  repurchase,  including  upon  a  fundamental 
change.  The  2030  Notes  will  mature  on  December  15,  2030,  subject  to  earlier  conversion,  redemption  or 
repurchase, including upon a fundamental change. The 2030 Notes and 2028 Notes are collectively referred to 
as the Notes.

Holders  of  the  Notes  will  have  the  right  to  require  us  to  repurchase  all  or  a  portion  of  their  Notes  upon  the 
occurrence  of  a  fundamental  change  before  the  maturity  date  at  a  repurchase  price  equal  to  100%  of  the 
principal  amount  of  the  Notes  to  be  repurchased,  plus  unpaid  interest  to,  but  excluding,  the  maturity  date.  In 
addition, upon conversion of the Notes, unless we elect to deliver solely shares of our common stock to settle 
such conversion (other than paying cash in lieu of delivering any fractional share), we will be required to settle a 
portion or all of our conversion obligation in cash in respect of the Notes being converted. Moreover, we will be 
required  to  repay  the  Notes  in  cash  at  their  maturity  unless  earlier  converted,  redeemed,  or  repurchased. 
However, we may not have enough available cash or be able to obtain financing at the time we are required to 
make repurchases of Notes surrendered therefor or pay cash with respect to Notes being converted or at their 
maturity.

In addition, our ability to repurchase Notes or to pay cash upon conversions of Notes or at their maturity may be 
limited by law, regulatory authority or agreements governing our future indebtedness. Our failure to repurchase 
Notes at a time when the repurchase is required by the indenture or to pay cash upon conversions of Notes or 
at their maturity as required by the indenture would constitute a default under the indenture. A default under the 
indenture or the fundamental change itself could also lead to a default under agreements governing our future 
indebtedness. Moreover, the occurrence of a fundamental change under the indenture could constitute an event 
of default under any such agreement. If the payment of the related indebtedness were to be accelerated after 
any applicable notice or grace periods, we may not have sufficient funds to repay the indebtedness or to pay 
cash amounts due upon conversion, upon required repurchase or at maturity of the Notes.

If the Notes are converted, it may adversely affect our financial condition and operating results.

Holders of the Notes are entitled to convert their Notes at any time at their option. If one or more holders elect 
to  convert  their  Notes,  unless  we  elect  to  satisfy  our  conversion  obligation  by  delivering  solely  shares  of  our 
common stock (other than paying cash in lieu of delivering any fractional share), we would be required to settle 
a  portion  or  all  of  our  conversion  obligation  in  cash,  which  could  adversely  affect  our  liquidity.  In  addition, 
issuances  of  shares  of  common  stock  upon  conversion  of  our  Notes  could  depress  the  market  price  of  our 
common  stock  and  impair  our  ability  to  raise  capital  through  the  sale  of  additional  equity  securities.  The 
existence of the Notes may encourage short selling by market participants because the conversion of the Notes 
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General Risk Factors

Unfavorable global economic or political conditions could adversely affect our business, financial condition or 
results of operations.

General conditions in the global economy and in the global financial markets could adversely affect our results 
of  operations,  including  the  potential  effects  from  the  COVID-19  pandemic  or  other  similar  outbreaks  as 
discussed above, and the overall demand for nucleic acid sequencing products may be particularly vulnerable to 
unfavorable economic conditions. A global financial crisis, inflation or a global or regional political disruption, as 
well as acts of terrorism, hostilities, military conflict and acts of war, including any further political uncertainty 
and military actions in the Middle East associated with the Israel and Hamas conflict and the war in Ukraine, as 
well  as  the  related  responses,  could  cause  extreme  volatility  in  the  capital  and  credit  markets.  A  severe  or 
prolonged economic downturn or political disruption could result in a variety of risks to our business, including 
weakened demand for our products and our ability to raise additional capital when needed on acceptable terms, 
if at all. A weak  or  declining economy or political disruption could also strain our manufacturers or suppliers, 
possibly resulting in supply disruption, or cause our customers to delay making payments for our product and 
services.  An  impairment  in  value  of  our  tangible  or  intangible  assets  could  also  be  recorded  as  a  result  of 
weaker economic conditions. Any of the foregoing could harm our business and we cannot anticipate all of the 
ways  in  which  the  political  or  economic  climate  and  financial  market  conditions  could  adversely  impact  our 
business.

Delivery  of  our  products  could  be  delayed  or  disrupted  by  factors  beyond  our  control,  and  we  could  lose 
customers as a result.

We  rely  on  third-party  carriers  for  the  timely  delivery  of  our  products.  As  a  result,  we  are  subject  to  carrier 
disruptions and increased costs that are beyond our control. Any failure to deliver products to our customers in 
a safe and timely manner may damage our reputation and brand and could cause us to lose customers. If our 
relationship with any of these third-party carriers is terminated or impaired or if any of these carriers are unable 
to deliver our products, the delivery of our products by our customers may be delayed, which could harm our 
business  and  financial  results.  The  failure  to  deliver  our  products  in  a  safe  and  timely  manner  may  harm  our 
relationship with our customers, increase our costs and otherwise disrupt our operations.

Doing business internationally creates operational and financial risks for our business.

We  currently  conduct  operations  in  various  countries  and  jurisdictions,  and  continue  to  expand  to  new 
international jurisdictions as part of our growth strategy and have experienced an increasing concentration of 
sales  in  certain  regions  outside  the  U.S.  We  sell  directly  and  through  distribution  partners  throughout  Europe, 
the  Asia-Pacific  region,  Mexico,  Brazil,  and  South  Africa  and  have  a  significant  portion  of  our  sales  and 
customer support personnel in Europe and the Asia-Pacific region. As a result, we or our distribution partners 
may be subject to additional regulations and increased diversion of management time and efforts. Conducting 
and  launching  operations  on  an  international  scale  requires  close  coordination  of  activities  across  multiple 
jurisdictions  and  time  zones  and  consumes  significant  management  resources.  If  we  fail  to  coordinate  and 
manage these activities effectively, our business, financial condition or results of operations could be materially 
and  adversely  affected  and  failure  to  comply  with  laws  and  regulations  applicable  to  business  operations  in 
foreign  jurisdictions  may  also  subject  us  to  significant  liabilities  and  other  penalties.  International  operations 
entail a variety of other risks, including, without limitation:

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potential limits to travel as a result of COVID-19 or other epidemics or pandemics;

challenges in staffing and managing foreign operations;

potentially  longer  sales  cycles  and  more  time  required  to  engage  and  educate  customers  on  the 
benefits of our platform outside of the United States;

the potential need for localized software and documentation;

reduced  protection  for  intellectual  property  rights  in  some  countries  and  practical  difficulties  of 
enforcing intellectual property and contract rights abroad;

defending against intellectual property claims in other countries;

restrictions  on  both  inbound  and  outbound  cross-border  investment,  including  enhanced  oversight  by 
the  Committee  on  Foreign  Investment  in  the  United  States  (“CFIUS”)  and  substantial  restrictions  on 
investment from China;

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U.S.  and  foreign  government  trade  restrictions,  including  those  which  may  impose  restrictions  on  the 
importation,  exportation,  re-exportation,  sale,  shipment  or  other  transfer  of  programming,  technology, 
components, and/or services to foreign persons;

changes in diplomatic and trade relationships, including new tariffs, trade protection measures, import 
or export licensing requirements, trade embargoes, sanctions, and other trade barriers;

tariffs  imposed  by  the  U.S.  on  goods  from  other  countries  and  tariffs  imposed  by  other  countries  on 
U.S. goods, including the tariffs by the U.S. government on various imports from China, Canada, Mexico, 
and  the  European  Union  (“E.U.”)  and  by  the  governments  of  these  jurisdictions  on  certain  U.S.  goods, 
and any other possible tariffs that may be imposed on products such as ours, the scope and duration of 
which, if implemented, remains uncertain;

deterioration of political relations between the U.S. and Russia, China, Japan, Korea, Canada, the United 
Kingdom (“U.K.”), and the E.U., which could have a material adverse effect on our sales and operations 
in these countries;

changes  in  social,  political,  and  economic  conditions  or  in  laws,  regulations  and  policies  governing 
foreign  trade,  manufacturing,  development,  and  investment  both  domestically  as  well  as  in  the  other 
countries and jurisdictions into which we sell our products, including as a result of the withdrawal of the 
U.K. from the E.U.;

difficulties in obtaining export licenses or in overcoming other trade barriers and restrictions resulting in 
delivery delays;

fluctuations in currency exchange rates and the related effect on our results of operations;

increased financial accounting and reporting burdens and complexities;

disruptions to global trade due to disease outbreaks or conflicts;

potential increases on tariffs or restrictions on trade generally; and

significant taxes or other burdens of complying with a variety of foreign laws and regulations, including 
laws  and  regulations  relating  to  privacy  and  data  protection  such  as  the  E.U.  General  Data  Protection 
Regulation which took effect in the E.U. in 2018.

In  conducting  our  international  operations,  we  are  subject  to  U.S.  laws  relating  to  our  international  activities, 
such  as  the  Foreign  Corrupt  Practices  Act  of  1977,  as  well  as  foreign  laws  relating  to  our  activities  in  other 
countries,  such  as  the  United  Kingdom  Bribery  Act  of  2010.  Additionally,  the  inclusion  of  one  of  our  foreign 
customers on any U.S. Government sanctioned persons list, including but not limited to the U.S. Department of 
Commerce’s  List  of  Denied  Persons  and  the  U.S.  Department  of  Treasury’s  List  of  Specially  Designated 
Nationals and Blocked Persons List, could be material to our earnings. Failure to comply with these laws may 
subject us to claims or financial and/or other penalties in the United States and/or foreign countries that could 
materially  and  adversely  impact  our  operations  or  financial  condition.  These  risks  have  become  increasingly 
prevalent as we have expanded our sales into countries that are generally recognized as having a higher risk of 
corruption.

We face risks related to the current global economic environment, which could delay or prevent our customers 
from  purchasing  our  products,  which  could  in  turn  harm  our  business,  financial  condition,  and  results  of 
operations. The state of the global economy continues to be uncertain. The current global economic conditions 
and  uncertain  credit  markets  and  concerns  regarding  the  availability  of  credit  pose  a  risk  that  could  impact 
customer demand for our products, as well as our ability to manage normal commercial relationships with our 
customers, suppliers, and creditors, including financial institutions. If the current global economic environment 
deteriorates, our business could be negatively affected.

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Moreover, changes in the value of the relevant currencies may affect the cost of certain items required in our 
operations. Changes in currency exchange rates may also affect the relative prices at which we are able to sell 
products  in  the  same  market.  Our  revenue  from  international  customers  may  be  negatively  impacted  as 
increases  in  the  U.S.  dollar  relative  to  our  international  customers’  local  currencies  could  make  our  products 
more expensive, impacting our ability to compete or as a result of financial or other instability in such locations 
which could result in decreased sales of our products. Our costs of materials from international suppliers may 
also  increase  as  the  value  of  the  U.S.  dollar  decreases  relative  to  their  local  currency.  Foreign  policies  and 
actions regarding currency valuation could result in actions by the United States and other countries to offset 
the effects of such fluctuations. Such actions may materially and adversely impact our financial condition and 
results of operations.

Violations  of  complex  foreign  and  U.S.  laws  and  regulations  could  result  in  fines  and  penalties,  criminal 
sanctions  against  us,  our  officers,  or  our  employees,  prohibitions  on  the  conduct  of  our  business  and  on  our 
ability to offer our products and services in one or more countries, and could also materially affect our brand, 
our  international  growth  efforts,  our  ability  to  attract  and  retain  employees,  our  business,  and  our  operating 
results.  Even  if  we  implement  policies  or  procedures  designed  to  ensure  compliance  with  these  laws  and 
regulations, there can be no assurance that our distribution partners, our employees, contractors, or agents will 
not violate our policies and subject us to potential claims or penalties.

If we fail to maintain proper and effective internal controls, our ability to produce accurate financial statements 
on a timely basis could be impaired, which would adversely affect our business and our stock price.

Ensuring that we have adequate internal financial and accounting controls and procedures in place to produce 
accurate  financial  statements  on  a  timely  basis  is  a  costly  and  time-consuming  effort  that  needs  to  be 
evaluated frequently. We may in the future discover areas of our internal financial and accounting controls and 
procedures  that  need  improvement.  Operating  as  a  public  company  requires  sufficient  resources  within  the 
accounting  and  finance  functions  in  order  to  produce  timely  financial  information,  ensure  the  level  of 
segregation  of  duties,  and  maintain  adequate  internal  control  over  financial  reporting  customary  for  a  U.S. 
public company.

Our  management  is  responsible  for  establishing  and  maintaining  adequate  internal  control  over  financial 
reporting to provide reasonable assurance regarding the reliability of our financial reporting and the preparation 
of financial statements for external purposes in accordance with U.S. GAAP. Our management does not expect 
that our internal control over financial reporting will prevent or detect all errors and all fraud. A control system, 
no  matter  how  well  designed  and  operated,  can  provide  only  reasonable,  not  absolute,  assurance  that  the 
control system’s objectives will be met. Because of the inherent limitations in all control systems, no evaluation 
of controls can provide absolute assurance that misstatements due to error or fraud will not occur or that all 
control issues and instances of fraud, if any, within our company will have been detected.

Pursuant to Section 404 of the Sarbanes-Oxley Act, we perform periodic evaluations of our internal control over 
financial reporting. While we have in the past performed this evaluation and concluded that our internal control 
over  financial  reporting  was  operating  effectively,  there  can  be  no  assurance  that  in  the  future  material 
weaknesses or significant deficiencies will not exist or otherwise be discovered. In addition, if we are unable to 
produce accurate financial statements on a timely basis, investors could lose confidence in the reliability of our 
financial  statements,  which  could  cause  the  market  price  of  our  common  stock  to  decline  and  make  it  more 
difficult for us to finance our operations and growth.

Our business could be negatively impacted by changes in the United States political environment.

There is significant ongoing uncertainty with respect to potential legislation, regulation and government policy 
at  the  federal  level,  as  well  as  the  state  and  local  levels.  Any  such  changes  could  significantly  impact  our 
business as well as the markets in which we compete. Specific legislative and regulatory proposals discussed 
during  election  campaigns  and  more  recently  that  might  materially  impact  us  include,  but  are  not  limited  to, 
changes to spending priorities and potential reductions in research funding. Uncertainty about U.S. government 
funding has posed, and may continue to pose, a risk as customers may choose to postpone or reduce spending 
in  response  to  actual  or  anticipated  restraints  on  funding.  To  the  extent  changes  in  the  political  environment 
have a negative impact on us or on our markets, our business, results of operation and financial condition could 
be materially and adversely impacted in the future.

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Disruption of critical information technology systems or material breaches in the security of our systems could 
harm our business, customer relations and financial condition.

in  our  products,  may  be  vulnerable  to  damage  from  a  variety  of  sources, 

Information  technology  (“IT”)  helps  us  to  operate  efficiently,  interface  with  customers,  maintain  financial 
accuracy  and  efficiently  and  accurately  produce  our  financial  statements.  IT  systems  are  used  extensively  in 
virtually  all  aspects  of  our  business,  including  in  our  products,  sales  forecast,  order  fulfillment  and  billing, 
customer  service,  logistics,  and  management  of  data  from  running  samples  on  our  products.  Our  success 
depends, in part, on the continued and uninterrupted performance of our IT systems. Our IT systems, including 
those  used 
including 
telecommunications  or  network  failures,  power  loss,  natural  disasters,  human  acts,  computer  viruses, 
ransomware,  computer  denial-of-service  attacks,  unauthorized  access  to  customer  or  employee  data  or 
company trade secrets, and other attempts to harm our systems. Furthermore, there may be a heightened risk 
of potential cybersecurity incidents and security breaches to which we could be vulnerable by state-sponsored 
or affiliated actors or others in connection with the political uncertainty and military actions in the Middle East 
associated with the Israel and Hamas conflict and the war in Ukraine. Certain of our systems are not redundant, 
and our disaster recovery planning is not sufficient for every eventuality. Despite any precautions we may take, 
such problems could result in, among other consequences, disruption of our operations, which could harm our 
reputation and financial results.

If  we  do  not  allocate  and  effectively  manage  the  resources  necessary  to  build  and  sustain  the  proper  IT 
infrastructure,  including  those  used  in  our  products,  we  could  be  subject  to  transaction  errors,  processing 
inefficiencies, loss of customers, business disruptions or loss of or damage to intellectual property. If our data 
management systems do not effectively collect, store, process and report relevant data for the operation of our 
business,  whether  due  to  equipment  malfunction  or  constraints,  software  deficiencies  or  human  error,  our 
ability  to  effectively  plan,  forecast  and  execute  our  business  plan  and  comply  with  applicable  laws  and 
regulations will be impaired, perhaps materially. Any such impairment could materially and adversely affect our 
reputation,  financial  condition,  results  of  operations,  cash  flows  and  the  timeliness  with  which  we  report  our 
internal and external operating results.

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Security  breaches  and  other  disruptions  could  compromise  our  information  and  expose  us  to  liability,  which 
would cause our business and reputation to suffer.

In the ordinary course of  our business, we collect and store sensitive data, including intellectual property, our 
proprietary  business  information  and  that  of  our  customers,  suppliers  and  business  partners,  and  personal 
information of our customers and employees, in our data centers and on our networks. The secure processing, 
maintenance and transmission of this information is critical to our operations. Despite our security measures, 
our IT infrastructure may be vulnerable to attacks by hackers, computer viruses, malicious codes, ransomware, 
unauthorized  access  attempts,  and  cyber-  or  phishing-attacks,  or  breached  or  otherwise  disrupted  due  to 
employee error, malfeasance, faulty password management or other disruptions. Third parties may attempt to 
fraudulently  induce  employees  or  other  persons  into  disclosing  usernames,  passwords  or  other  sensitive 
information,  which  may  in  turn  be  used  to  access  our  IT  systems,  commit  identity  theft  or  carry  out  other 
unauthorized or illegal activities. Any such breach or incident could compromise our systems and networks and 
the  information  stored  or  otherwise  processed  there  could  be  accessed,  publicly  disclosed,  lost,  stolen  or 
otherwise processed in an unauthorized manner. We engage third-party vendors and service providers to store 
and otherwise process some of our data, including sensitive and personal information. Our vendors and service 
providers  may  also  be  the  targets  of  the  risks  described  above,  including  cyber-attacks,  malicious  software, 
ransomware,  phishing  schemes,  and  fraud.  Our  ability  to  monitor  our  vendors  and  service  providers’  data 
security is limited, and, in any event, third parties may be able to circumvent those security measures, resulting 
in  the  unauthorized  access  to,  misuse,  disclosure,  loss  or  destruction  of  our  data,  including  sensitive  and 
personal  information,  and  disruption  of  our  or  third-party  service  providers’  systems.  We  and  our  third-party 
service providers may face difficulties in identifying, or promptly responding to, potential security breaches and 
other  instances  of  unauthorized  access  to,  or  disclosure,  other  processing,  or  loss  or  unavailability  of, 
information.  Any  hacking  or  other  attack  on  our  or  our  third-party  service  providers’  or  vendors’  systems,  and 
any unauthorized access to, or disclosure, other processing, or loss or unavailability of, information suffered by 
us or our third-party service providers or vendors, or the perception that any of these have occurred, could result 
in  legal  claims  or  proceedings,  loss  of  intellectual  property,  liability  under  laws  that  protect  the  privacy  of 
personal  information,  negative  publicity,  disruption  of  our  operations  and  damage  to  our  reputation,  and  data 
integrity  issues,  which  could  divert  our  management’s  attention  from  the  operation  of  our  business  and 
materially  and  adversely  affect  our  business,  revenues  and  competitive  position.  Moreover,  we  may  need  to 
increase  our  efforts  to  train  our  personnel  to  detect  and  defend  against  cyber-  or  phishing-attacks,  which  are 
becoming more sophisticated and frequent, and we may need to implement additional protective measures to 
reduce the risk of potential security breaches and security incidents, which could cause us to incur significant 
additional  expenses.  Retaliatory  acts  by  Russia  in  response  to  Western  sanctions  or  otherwise  in  connection 
with the war in Ukraine could include cyber-attacks that could disrupt the economy generally or that may either 
directly or indirectly impact our operations specifically.

In  addition,  our  insurance  may  be  insufficient  to  cover  our  losses  resulting  from  cyber-attacks,  breaches,  or 
other  interruptions,  and  any  incidents  may  result  in  loss  of,  or  increased  costs  of,  such  insurance.  The 
successful  assertion  of  one  or  more  large  claims  against  us  that  exceed  available  insurance  coverage,  the 
occurrence  of  changes  in  our  insurance  policies,  including  premium  increases  or  the  imposition  of  large 
deductible  or  co-insurance  requirements,  or  denials  of  coverage,  could  have  a  material  adverse  effect  on  our 
business, including our financial condition, results of operations and reputation.

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Our use of artificial intelligence and machine learning technologies may result in reputational harm or liability.

We have incorporated and may continue to incorporate additional artificial intelligence and machine learning, or 
AIML, technologies into our sequencing platforms, marketing programs, and analysis software, including Revio 
and otherwise within our business, and these solutions and features are advantageous to describing, enhancing, 
and maximizing the capabilities of our differentiated technologies and to our future growth over time. We rely 
and  expect  to  rely  on  AIML  technologies  such  as  basecalling,  variant  calling,  epigenetic  analysis  and  tertiary 
analysis, but there can be no assurance that we will realize the desired or anticipated benefits from AIML or any 
at all. We may also fail to properly implement or utilize AIML technologies. Our competitors or other third parties 
may incorporate AIML into their products, platforms, software and services or otherwise within their business 
more quickly or more successfully than us, which could impair our ability to compete effectively and adversely 
affect our results of operations. Additionally, our use of AIML technologies may expose us to additional claims, 
demands and proceedings by private parties and regulatory authorities and subject us to legal liability as well as 
brand and reputational harm. For example, if output from AIML technologies or that they assist in producing are 
or  are  alleged  to  be  deficient,  inaccurate,  or  biased,  or  for  such  output,  or  such  technologies  or  their 
development  or  deployment,  including  the  collection,  use,  or  other  processing  of  data  used  to  train  or  create 
such AIML technologies, to alleged to infringe upon or to have misappropriated third-party intellectual property 
rights or to violate applicable laws, regulations, or other actual or asserted legal obligations to which we are or 
may  become  subject,  then  our  business,  financial  condition,  and  results  of  operations  may  be  adversely 
affected. The legal, regulatory, and policy environments around AIML are evolving rapidly, and we may become 
subject to new and evolving legal and other obligations. These and other developments may require us to make 
significant changes to our use of AIML, including by limiting or restricting our use of AIML, and may require us 
to  make  significant  changes  to  our  policies  and  practices,  which  may  necessitate  expenditure  of  significant 
time,  expense,  and  other  resources,  AIML  also  presents  emerging  ethical  issues,  and  if  our  use  of  AIML 
becomes controversial, we may experience brand or reputational harm.

We are currently subject to, and may in the future become subject to additional, U.S. federal and state laws and 
regulations  imposing  obligations  on  how  we  collect,  store  and  process  personal  information.  Our  actual  or 
perceived failure to comply with such obligations could harm our business. Ensuring compliance with such laws 
could  also  impair  our  efforts  to  maintain  and  expand  our  future  customer  base,  and  thereby  decrease  our 
revenue.

In the ordinary course of our business, we currently, and in the future will, collect, store, transfer, use or process 
sensitive data, including personal information of employees, and intellectual property and proprietary business 
information owned or controlled by ourselves and other parties. The secure processing, storage, maintenance, 
and transmission of this critical information are vital to our operations and business strategy. We are, and may 
increasingly become, subject to various laws and regulations, as well as contractual obligations, relating to data 
privacy and security in the jurisdictions in which we operate. The regulatory environment related to data privacy 
and  security  is  increasingly  rigorous,  with  new  and  constantly  changing  requirements  applicable  to  our 
business, and enforcement practices are likely to remain uncertain for the foreseeable future. These laws and 
regulations  may  be  interpreted  and  applied  differently  over  time  and  from  jurisdiction  to  jurisdiction,  and  it  is 
possible  that  they  will  be  interpreted  and  applied  in  ways  that  may  have  a  material  adverse  effect  on  our 
business, financial condition, results of operations and prospects.

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In the United States, various federal and state regulators, including governmental agencies like the Consumer 
Financial  Protection  Bureau  and  the  Federal  Trade  Commission,  have  adopted,  or  are  considering  adopting, 
laws  and  regulations  concerning  personal  information  and  data  security.  Certain  state  laws  may  be  more 
stringent or broader in scope, or offer greater individual rights, with respect to personal information than federal, 
international  or  other  state  laws,  and  such  laws  may  differ  from  each  other,  all  of  which  may  complicate 
compliance efforts. For example, the California Consumer Privacy Act (“CCPA”), which increases privacy rights 
for California residents and imposes obligations on companies that process their personal information, came 
into  effect  on  January  1,  2020.  Among  other  things,  the  CCPA  requires  covered  companies  to  provide  new 
disclosures  to  California  consumers  and  provide  such  consumers  new  data  protection  and  privacy  rights, 
including the ability to opt-out of certain sales of personal information. The CCPA provides for civil penalties for 
violations,  as  well  as  a  private  right  of  action  for  certain  data  breaches  that  result  in  the  loss  of  personal 
information. This private right of action may increase the likelihood of, and risks associated with, data breach 
litigation.  In  November  2020,  California  also  passed  the  California  Privacy  Rights  Act,  or  (“CPRA”),  which 
significantly expanded the CCPA as of January 1, 2023, including by introducing additional obligations such as 
data  minimization  and  storage  limitations  and  granting  additional  rights  to  consumers,  among  others.  The 
enactment  of  the  CCPA  has  prompted  similar  legislative  developments  in  other  states,  and  numerous  other 
states  have  proposed,  and  in  certain  cases  enacted,  legislation  relating  to  privacy  and  data  security,  many  of 
which  are  similar  to  the  CCPA  and  CPRA.  Similar  laws  are  being  considered  by  other  state  legislatures.  In 
addition,  laws  in  all  50  U.S.  states  require  businesses  to  provide  notice  to  consumers  whose  personal 
information  has  been  disclosed  as  a  result  of  a  data  breach.  State  laws  are  changing  rapidly  and  there  is 
discussion in the U.S. Congress of a new comprehensive federal data privacy law. These and future laws and 
regulations may increase our compliance costs and potential liability.

Furthermore,  regulations  promulgated  pursuant  to  the  Health  Insurance  Portability  and  Accountability  Act  of 
1996  (“HIPAA”),  establish  privacy  and  security  standards  that  limit  the  use  and  disclosure  of  individually 
identifiable  health  information  (known  as  “protected  health  information”)  and  require  the  implementation  of 
administrative, physical and technological safeguards to protect the privacy of protected health information and 
ensure  the  confidentiality,  integrity  and  availability  of  electronic  protected  health  information.  Determining 
whether  protected  health  information  has  been  handled  in  compliance  with  applicable  privacy  standards  and 
our contractual obligations can require complex factual and statistical analyses and may be subject to changing 
interpretation.  Although  we  take  measures  to  protect  sensitive  data  from  unauthorized  access,  use  or 
disclosure, our information technology and infrastructure may be vulnerable to attacks by hackers or viruses or 
disrupted,  breached  or  otherwise  compromised  due  to  employee  error,  malfeasance  or  other  malicious  or 
inadvertent  disruptions.  Any  such  breach  or  disruption  could  compromise  our  networks  and  the  information 
stored  there  could  be  accessed,  manipulated,  publicly  disclosed,  lost,  stolen,  made  unavailable,  or  otherwise 
processed  without  authorization.  Any  such  disruption,  access,  breach,  unavailability,  theft,  loss  or  other 
unauthorized processing of information, or the perception that any of these has occurred could result in legal 
claims or proceedings, and liability under federal or state laws that protect the privacy of personal information, 
such  as  the  HIPAA,  the  Health  Information  Technology  for  Economic  and  Clinical  Health  Act,  and  regulatory 
penalties. Notice of breaches must be made to affected individuals, the Secretary of the Department of Health 
and Human Services, and for extensive breaches, notice may need to be made to the media or state attorneys 
general. Such a notice could harm our reputation and our ability to compete.

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While  we  have  in  place  formal  policies  and  procedures  related  to  the  storage,  collection,  and  processing  of 
information, and have conducted data privacy audits, we continue to evaluate our compliance needs, including 
the  need  to  conduct  additional  internal  and  external  data  privacy  audits  or  adopt  additional  policies  and 
procedures, to ensure our compliance with all applicable data protection laws and regulations. Additionally, we 
do not currently have policies and procedures in place for assessing our third-party vendors’ compliance with 
applicable data protection laws and regulations. All of these evolving compliance and operational requirements 
impose significant costs, such as costs related to organizational changes, implementing additional protection 
technologies, training employees and engaging consultants, which are likely to increase over time. In addition, 
such requirements may require us to modify our data processing practices and policies, distract management 
or divert resources from other initiatives and projects, all of which could have a material adverse effect on our 
business, financial condition, results of operations and prospects. Any failure or perceived failure by us or our 
third-party  vendors,  collaborators,  contractors  and  consultants  to  comply  with  any  applicable  federal,  state  or 
similar  foreign  laws  and  regulations  relating  to  data  privacy  and  security,  could  result  in  damage  to  our 
reputation, as well as proceedings or litigation by governmental agencies or other third parties, including class 
action privacy litigation in certain jurisdictions, which would subject us to significant fines, sanctions, awards, 
penalties or judgments, all of which could have a material adverse effect on our business, financial condition, 
results of operations and prospects.

Increased scrutiny of our environmental, social or governance responsibilities may result in additional costs and 
risks, and may adversely impact our reputation, employee retention, and willingness of customers and suppliers 
to do business with us.

Investor advocacy groups, institutional investors, investment funds, proxy advisory services, stockholders, and 
customers are increasingly focused on environmental, social, and governance (“ESG”) practices of companies. 
Additionally,  public  interest  and  legislative  pressure  related  to  public  companies’  ESG  practices  continues  to 
grow.  If  our  ESG  practices  fail  to  meet  regulatory  requirements  or  investor  or  other  industry  stakeholders' 
evolving  expectations  and  standards  for  responsible  corporate  citizenship  in  areas  including  environmental 
stewardship,  support  for  local  communities,  board  and  employee  diversity,  human  capital  management, 
employee  health  and  safety  practices,  product  quality,  supply  chain  management,  corporate  governance  and 
transparency,  and  employing  ESG  strategies  in  our  operations,  our  brand,  reputation  and  employee  retention 
may be negatively impacted and customers and suppliers may be unwilling to do business with us. In addition, 
as we work to align our ESG practices with industry standards, we will likely continue to expand our disclosures 
in these areas and doing so may result in additional costs and require additional resources to monitor, report, 
and  comply  with  our  various  ESG  practices.  If  we  fail  to  adopt  ESG  standards  or  practices  as  quickly  as 
stakeholders  desire,  report  on  our  ESG  efforts  or  practices  accurately,  or  satisfy  the  expectations  of 
stakeholders, our reputation, business, financial performance, and growth may be adversely impacted.

ITEM 1B. 

UNRESOLVED STAFF COMMENTS

None.

ITEM 1C. 

CYBERSECURITY

Risk Management and Strategy

We  have  established  policies  and  processes  for  assessing,  identifying,  and  managing  material  risk  from 
cybersecurity  threats,  and  have  integrated  these  processes  into  our  overall  risk  management  systems  and 
processes.  We routinely assess material risks from cybersecurity threats, including any potential unauthorized 
occurrence  on  or  conducted  through  our  information  systems  that  may  result  in  adverse  effects  on  the 
confidentiality, integrity, or availability of our information systems or any information residing therein.

We conduct periodic risk assessments to identify cybersecurity threats, as well as assessments in the event of 
a  material  change  in  our  business  practices  that  may  affect  information  systems  that  are  vulnerable  to  such 
cybersecurity  threats.    These  risk  assessments  include  identification  of  reasonably  foreseeable  internal  and 
external  risks,  the  likelihood  and  potential  damage  that  could  result  from  such  risks,  and  the  sufficiency  of 
existing policies, procedures, systems, and safeguards in place to manage such risks.

Following  these  risk  assessments,  we  evaluate  whether  and  how  to  re-design,  implement,  and  maintain 
reasonable  safeguards  to  minimize  identified  risks;  reasonably  address  any  identified  gaps  in  existing 
safeguards;  and  regularly  monitor  the  effectiveness  of  our  safeguards.    We  devote  significant  resources  and 
designate high-level personnel, including our Senior Director and Head of Information Technology who reports 
to our Chief Operating Officer, to manage the risk assessment and mitigation process. 

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As part of our overall risk management system, we monitor and test our safeguards, including through annual 
third-party vulnerability assessments. We train our employees on these safeguards, in collaboration with human 
resources, IT, and departmental management.  Personnel at all levels and departments are made aware of our 
cybersecurity policies through trainings. 

We  engage  assessors,  consultants,  auditors,  or  other  third  parties  in  connection  with  our  risk  assessment 
processes.  These  service  providers  assist  us  to  design  and  implement  our  cybersecurity  policies  and 
procedures, as well as to monitor and test our safeguards.  

We  require  each  third-party  infrastructure  and  applications  service  provider  that  has  access  to  our  system  to 
certify  that  it  has  the  ability  to  implement  and  maintain  appropriate  security  measures,  consistent  with  all 
applicable laws, to implement and maintain reasonable security measures in connection with their work with us, 
and to promptly report any suspected breach of its security measures that may affect our company. 

We have not previously experienced a cybersecurity incident that was determined to be material. For additional 
information regarding whether any risks from cybersecurity threats are reasonably likely to materially affect our 
company, including our business strategy, results of operations, or financial condition, please refer to Item 1A, 
“Risk Factors,” in this annual report on Form 10-K.

Governance

One  of  the  key  functions  of  our  board  of  directors  is  informed  oversight  of  our  risk  management  process, 
including risks from cybersecurity threats. Our board of directors is responsible for monitoring and assessing 
strategic  risk  exposure,  and  our  executive  officers  are  responsible  for  the  day-to-day  management  of  the 
material risks we face. Our board of directors administers its cybersecurity risk oversight function directly as a 
whole, as well as through the audit committee. 

Our  Senior  Director  and  Head  of  Information  Technology  and  our  management  committee  on  cybersecurity, 
which includes Facilities, HR, IT, Legal, and Management, are primarily responsible to assess and manage our 
material  risks  from  cybersecurity  threats.    Our  cybersecurity  team  and  IT  management  have  years  of 
cybersecurity experience and expertise including holding numerous applicable cybersecurity certifications, such 
as ISC2 Certified Information Systems Security Professional (CISSP).

Our  Senior  Director  and  Head  of  Information  Technology  and  our  management  committee  on  cybersecurity 
oversee our cybersecurity policies and processes, including those described in “Risk Management and Strategy” 
above.  Our Senior Director and Head of Information Technology is informed about and monitors the prevention, 
detection,  mitigation,  and  remediation  of  cybersecurity 
implantation  of  the 
cybersecurity risk management program and working directly with our security team. The Senior Director and 
Head  of  Information  Technology  also  provides  appropriate  information  and  updates  to  our  management 
committee on cybersecurity.

incidents  by 

leading  the 

Our Senior Director and Head of Information Technology and representatives from our management committee 
on cybersecurity provide annual briefings to the audit committee regarding our company’s cybersecurity risks 
and  activities,  including  any  recent  cybersecurity  incidents  and  related  responses,  cybersecurity  systems 
testing,  activities  of  third  parties,  and  the  like.    Our  audit  committee  provides  regular  updates  to  the  board  of 
directors on such reports. 

ITEM 2. PROPERTIES 

Our  corporate  headquarters,  research  and  development  facilities,  and  manufacturing  and  distribution  centers 
are located in Menlo Park, California, where we lease approximately 180,200 square feet under a lease expiring 
on October 31, 2027. We operate additional research, development, and support functions in San Diego, where 
we  lease  approximately  73,500  square  feet  under  a  lease  expiring  on  September  30,  2027.  Additionally,  our 
European  headquarters  is  located  in  London,  where  we  lease  approximately  7,300  square  feet  under  a  lease 
expiring November 30, 2026. Including these leases, we lease approximately 270,000 square feet globally. 

We  believe  that  our  existing  facilities,  together  with  suitable  additional  or  alternative  space  available  on 
commercially reasonable terms, will be sufficient to meet our needs. 

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ITEM 3. LEGAL PROCEEDINGS 

U.S. District Court Proceedings

On September 26, 2019, Personal Genomics of Taiwan, Inc. (“PGI”) filed a complaint in the U.S. District Court for 
the  District  of  Delaware  against  us  for  patent  infringement  (C.A.  No.  19-cv-1810)  (the  “PGI  District  Court 
matter”).  The  matter  from  this  complaint  is  based  on  PGI’s  U.S.  Patent  No.  7,767,441  (the  “‘441  Patent”).  We 
plan to vigorously defend in this matter. On November 20, 2019, we filed our answer to the complaint, denying 
infringement and seeking a declaratory judgement of invalidity of the ‘441 Patent.

On June 22, 2020, we filed a petition requesting institution of an inter-partes review ("IPR") to the Patent Trial 
and Appeals Board (the “Board”) at the United States Patent Office requesting the Board to find a set of claims 
in the ‘441 Patent invalid. On June 27, 2020, we filed a second petition requesting institution of an IPR 
requesting the Board to find another set of claims in the ‘441 Patent invalid. The two petitions (the “PacBio IPR 
Petitions”) requesting IPRs assert that all of the claims relevant to the PGI complaint are invalid. On January 19, 
2021, the Board ordered that both PacBio IPR Petitions be instituted on all grounds presented. On January 18, 
2022, the Board issued decisions on the two IPRs. In one IPR, all challenged claims were found unpatentable, 
including PGI’s core device claims. In the second IPR, the Board did not find the disputed claims unpatentable. 
We appealed the decision in the second IPR to the U.S. Court of Appeals for the Federal Circuit, which held a 
hearing on December 7, 2023. On January 9, 2024, the U.S. Court of Appeals for the Federal Circuit affirmed the 
decisions of the Board.

On August 19, 2020, the court ordered a stay of the PGI District Court matter based on a joint stipulation by the 
parties pending a final written decision on the IPRs. Following the final decision on the IPRs described above, on 
February 2, 2022, the judge ordered that the PGI District Court matter be reopened. However, in a subsequent 
order dated September 15, 2022, the judge stayed the PGI District Court matter pending a final decision by the 
U.S.  Court  of  Appeals  for  the  Federal  Circuit  regarding  the  appeal  described  above.  We  plan  to  vigorously 
defend against the remaining claims.

In December 2022, Take2 Technologies, Ltd. (“Take2”) and the Chinese University of Hong Kong filed a 
complaint in the U.S. District Court for Delaware against us alleging infringement of U.S. Patent No. 11,091,794 
(the “’794 Patent”) (C.A. No. 22- cv-01595). The complaint alleges that our SequelTM II systems, Sequel IIe 
Systems, and RevioTM Systems that operate version 11.0 or later of the SMRTTM Link software, infringe the ‘794 
Patent. The complaint seeks unspecified monetary damages and an order enjoining us from infringing the ’794 
Patent. We believe the infringement allegations in the complaint lack merit and we intend to vigorously defend 
in this matter. We filed a motion to dismiss on February 14, 2023. We also filed a motion to transfer the case to 
the Northern District of California which was granted on August 2, 2023 and the case was transferred on August 
16, 2023 (C.A. No. 5:23-cv-04166). A hearing occurred on February 22, 2024 on the motion to dismiss the case 
and the motion was taken under submission for future decision. Take2 filed a motion to disqualify our in-house 
legal department from representing PacBio in the district court action on September 20, 2023. We opposed 
Take2’s disqualification motion on October 4, 2023. An oral hearing on the disqualification motion was held on 
October 26, 2023 and the court issued orders November 6 and December 4 of 2023 partially granting the 
motion.  While some members of the in-house legal department were disqualified, General Counsel for PacBio 
was not disqualified and continues to represent PacBio in the district court action. On October 17, 2023, we filed 
a petition requesting institution of an IPR requesting the Board to find all claims of the ’794 patent invalid. We 
anticipate a decision by the United States Patent and Trademark Office on whether to institute a trial on the 
validity of the claims of the '794 patent on or before April 26, 2024.

Proceedings in China

On May 12, 2020, PGI filed a complaint in the Wuhan Intermediate People’s Court in China alleging infringement 
of  one  or  more  claims  of  China  patent  No.  CN101743321B  (the  “CN321  Patent”),  which  is  related  to  the  ‘441 
Patent.  On  November  23,  2020,  we  filed  an  Invalidation  Petition  at  the  China  National  Intellectual  Property 
Administration  (CNIPA)  demonstrating  the  invalidity  of  the  claims  in  the  CN321  Patent  on  grounds  of 
insufficient disclosure, and the lack of support, essential technical features, clarity, novelty, and inventiveness. A 
hearing  in  the  invalidation  proceeding  at  the  CNIPA  was  held  on  April  29,  2021.  On  September  2,  2021,  the 
CNIPA issued its decision on the Invalidation Petition and determined that all claims (1-61) of the CN321 patent 
were invalid. On December 1, 2021, PGI filed an appeal with the Beijing IP Court, contesting the CNIPA decision. 
We filed a petition with the Wuhan Intermediate People’s court requesting dismissal of the infringement action 
based  on  the  CNIPA  invalidation  decision,  and  PGI  filed  a  petition  to  withdraw  its  complaint.  The  Wuhan 
Intermediate People’s court granted PGI’s petition and dismissed the infringement action in May 2022.

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Other Proceedings

From  time  to  time,  we  may  also  be  involved  in  a  variety  of  other  claims,  lawsuits,  investigations,  and 
proceedings  relating  to  securities  laws,  product  liability,  patent  infringement,  contract  disputes,  employment, 
and other matters that arise in the normal course of our business. In addition, third parties may, from time to 
time, assert claims against us in the form of letters and other communications.

We record a provision for contingent losses when it is both probable that a liability has been incurred and the 
amount of the loss can be reasonably estimated. We currently do not believe that the ultimate outcome of any 
of the matters described above is probable or reasonably estimable, or that these matters will have a material 
adverse  effect  on  our  business;  however,  the  results  of  litigation  and  claims  are  inherently  unpredictable. 
Regardless  of  the  outcome,  litigation  can  have  an  adverse  impact  on  us  because  of  litigation  and  settlement 
costs, diversion of management resources, and other factors.

ITEM 4. MINE SAFETY DISCLOSURES

Not applicable.

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PART II

ITEM  5. MARKET  FOR  REGISTRANT’S  COMMON  EQUITY,  RELATED  STOCKHOLDER  MATTERS  AND  ISSUER 
PURCHASES OF EQUITY SECURITIES 

Our common stock is traded on The Nasdaq Global Select Market under the symbol “PACB.” 

Holders of Record 

As of January 31, 2024, there were approximately 76 stockholders of record of our common stock, although we 
believe that there are a significantly larger number of beneficial owners of our common stock. 

Dividend Policy 

We have never declared or paid any cash dividend on our common stock and have no present plans to do so. 
We intend to retain earnings for use in the operation and expansion of our business.

Performance Graph 

The performance graph included in this Annual Report on Form 10-K shall not be deemed “filed” for purposes of 
Section  18  of  the  Securities  Exchange  Act  of  1934,  as  amended  (the  “Exchange  Act”),  or  incorporated  by 
reference into any filing of Pacific Biosciences under the Securities Act of 1933, as amended, or the Exchange 
Act, except as shall be expressly set forth by specific reference in such filing. 

The  following  graph  shows  a  comparison  from  December  31,  2018  through  December  31,  2023  of  the 
cumulative  total  return  for  our  common  stock,  the  Nasdaq  Composite  Index  and  the  Nasdaq  Biotechnology 
Index. Such returns are based on historical results and are not intended to suggest future performance. Data for 
The Nasdaq Composite Index and the Nasdaq Biotechnology Index assume reinvestment of dividends. 

COMPARISON OF 5 YEAR CUMULATIVE TOTAL RETURN*
Among Pacific Biosciences, Inc, the NASDAQ Composite Index
and the NASDAQ Biotechnology Index

*$100 invested on 12/31/2018 in stock or index, including reinvestment of dividends.
Fiscal year ending December 31.

Recent Sales of Unregistered Securities 

Other than as previously reported on our Current Reports on Form 8-K filed with the SEC on June 26, 2023 and 
August 2, 2023, there were no unregistered sales of equity securities by us during 2023.

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ITEM 6. [Reserved]

ITEM  7. MANAGEMENT’S  DISCUSSION  AND  ANALYSIS  OF  FINANCIAL  CONDITION  AND  RESULTS  OF 
OPERATIONS 

You  should  read  the  following  discussion  and  analysis  of  our  financial  condition  and  results  of  operations 
together with our consolidated financial statements and the related notes included in this Annual Report on Form 
10-K.  Some  of  the  information  contained  in  this  discussion  and  analysis  or  set  forth  elsewhere  in  this  Annual 
Report  on  Form  10-K,  including  information  with  respect  to  our  plans  and  strategy  for  our  business  and  related 
financing,  includes  forward-looking  statements  that  involve  risks  and  uncertainties.  You  should  read  the  “Risk 
Factors” section of this Annual Report on Form 10-K for a discussion of important factors that could cause actual 
results to differ materially from the results described in or implied by the forward-looking statements contained in 
the following discussion and analysis. 

Our Management’s Discussion and Analysis (MD&A) is organized in the following sections: 

•

•

•

•

•

•

•

•

Overview and Outlook 

Results of Operations 

Liquidity and Capital Resources 

Critical Accounting Policies and Estimates

Quantitative and Qualitative Disclosure of Market Risk

Recent Accounting Pronouncements

Contractual Obligations

Off Balance Sheet Arrangements

Overview and Outlook

About PacBio

We are a premier life science technology company that is designing, developing, and manufacturing advanced 
sequencing  solutions  that  enable  scientists  and  clinical  researchers  to  improve  their  understanding  of  the 
genome and ultimately, resolve genetically complex problems.

Our products and technology under development stem from two highly differentiated core technologies focused 
on  accuracy,  quality,  and  completeness,  which  include  our  HiFi  long-read  sequencing  technology  and  our 
Sequencing  by  Binding  (SBB®)  short-read  sequencing  technology.  Our  products  address  solutions  across  a 
broad set of research applications including human genetics, plant and animal sciences, infectious disease and 
microbiology, oncology, and other emerging applications. 

Our focus is on creating some of the world’s most advanced sequencing systems to provide our customers the 
most complete and accurate view of genomes, transcriptomes, and epigenomes. 

Our  customers  include  academic  and  governmental  research  institutions,  commercial  testing  and  service 
laboratories,  genome  centers,  public  health  labs,  hospitals  and  clinical  research  institutes,  contract  research 
organizations (CROs), pharmaceutical companies, and agricultural companies.

As  of  December  31,  2023,  our  commercial  team  is  comprised  of  approximately  215  employees,  including  71 
quota-carrying  representatives,  many  with  advanced  degrees  in  biology  and  significant  experience  in  the 
genomics industry.

Strategic Objectives

2023  exceeded  our  expectations  as  we  drove  adoption  of  our  advanced  sequencing  technologies, 
demonstrated  short-read  accuracy  with  our  Onso  platform,  and  progressed  development  of  our  pipeline 
technologies and products.

Our 2024 strategic objectives are to: 

•

•

Increase  technology  adoption  by  increasing  market  share  via  new  customer  acquisition,  continue 
Sequel II conversions to Revio, and scale Onso production; 

Leverage  innovation  to  complete  development  of  new  sequencing  platforms  and  launch  on-market 
system improvements;

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•

•

Build  upon  clinical  momentum  by  expanding  HiFi  usage  in  large-scale  programs  and  translational 
research projects; and

Drive  towards  positive  cash  flow  through  gross  margin  expansion,  disciplined  operating  expense 
management, and a focus on working capital. 

We will continue to leverage our commercial organization and make significant improvements in the efficiency 
and usability of our products in pursuit of a broader customer base. We believe the commercial investments we 
have recently made will further help drive growth in our business. 

To increase the adoption of HiFi sequencing, we have various development programs in progress to expand our 
product  portfolio  as  well  as  increase  the  throughput  and  improve  the  usability  of  our  existing  sequencing 
technologies. We continue to focus on programs to accelerate new platform launches in the near to mid-term 
as well as increase applications for our technologies. 

We  continue  to  believe  that  with  the  capabilities  of  our  HiFi  chemistry  and  SMRT  technology,  we  can  be  a 
market  leader  in  whole-genome  clinical  sequencing.  Leading  institutions  have  adopted  our  products  to  study 
rare  and  inherited  disease.  We  believe  the  market  opportunity  for  clinical  sequencing  is  significant  and  could 
drive substantial revenue growth for the company. We plan to continue to pursue partner collaborations where 
the  technologies  being  developed  or  applications  being  considered  extend  beyond  whole-genome  clinical 
sequencing. Collaborative arrangements add to the awareness of our products and service offerings and may 
drive new applications for use of our technology.

Financial Overview

Key highlights of our 2023 consolidated financial results include the following: 

•

•

•

•

Revenue  increased  $72.2  million,  or  56%,  to  $200.5  million  for  the  year  ended  December  31,  2023,  as 
compared to $128.3 million for the year ended December 31, 2022. Revenue was comprised of $120.5 
million  in  instrument  revenue,  $63.4  million  in  consumables  revenue  and  $16.6  million  in  service  and 
other revenue for the year ended December 31, 2023. The increase was primarily driven by the launch of 
Revio  in  the  first  quarter  of  2023,  which  is  sold  at  a  higher  average  selling  price  than  our  previous 
Sequel II and IIe platforms. We ended the year with an installed base of 173 Revio systems.

Gross  profit  as  a  percentage  of  revenue  (gross  margin)  was  26.3%  for  the  year  ended  December  31, 
2023, compared to 38.2% for the year ended December 31, 2022. Gross margin declined due primarily 
to the instrument mix, as Revio instruments sold during the period had a lower margin than the Sequel 
II/IIe  system,  primarily  due  to  loyalty  discounts  provided  and  higher  initial  manufacturing  costs, 
including  warranty  costs  as  well  as  charges  for  scrap  inventory.  In  addition,  we  recognized  an 
adjustment  during  the  year  ended  December  31,  2023  primarily  related  to  excess  instrument  and 
consumables inventory. The excess instrument adjustment was primarily due to a change in the Sequel 
II demand primarily from one customer. The consumables inventory adjustment was primarily resulting 
from  a  faster-than-expected  decline  in  demand  of  Sequel  II/IIe  consumables  due  primarily  to  a  faster 
than expected ramp on the Revio system.

Loss from operations increased $27.3 million or 9%, to $334.5 million for the year ended December 31, 
2023,  as  compared  to  $307.2  million  for  the  year  ended  December  31,  2022,  driven  primarily  by  an 
increase of $31.0 million of operating expenses, including a $12.7 million increase in the change in the 
fair value of the contingent consideration, a $9.0 million increase in non-recurring merger-related costs 
incurred, and a $9.0 million increase in sales, general, and administrative expenses, partially offset by a 
$5.8  million  decrease  in  research  and  development  expenses.  See  Note  2.  Business  Acquisitions  for 
further details. 

Cash, cash equivalents, and short-term investments were $631.4 million at December 31, 2023, which 
represents an 18% decrease compared to the balance at December 31, 2022.

Macroeconomic dynamics including inflation, exchange rates and concerns about an economic downturn, have 
impacted  both  the  Company  and  our  customers’  behavior.  For  example,  some  customers  and  potential 
customers are managing capital more conservatively, resulting in lengthened sales cycles, and capital funding 
in  China  is  being  deferred,  resulting  in  less  capital  equipment  purchases.  These  factors  could  continue  to 
impact our revenues and results of operations in 2024; however, as the size and duration of these impacts is 
uncertain, we cannot reasonably estimate the future impact to our operations and financial results. 

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See  the  Risk  Factors  section  for  further  discussion  of  the  possible  impact  of  macroeconomic  factors  on  our 
business.

Results of Operations 

A  detailed  discussion  of  our  consolidated  financial  results  comparison  between  2023  and  2022  is  presented 
below. A discussion of the changes in our results of operations between the years ended December 31, 2022 
and December 31, 2021, has been omitted from this Annual Report on Form 10-K but may be found in Item 7. 
Management’s Discussion and Analysis of Financial Condition and Results of Operations of our Annual Report on 
Form  10-K  for  the  year  ended  December  31,  2022,  filed  with  the  Securities  and  Exchange  Commission  on 
February  28,  2023,  which  is  incorporated  herein  by  reference,  and  is  available  free  of  charge  on  the  SEC’s 
website at www.sec.gov and our corporate website (www.pacb.com).

Comparison of the Years Ended December 31, 2023 and 2022

(in thousands, except per share amounts)

2023

2022

$ Change

% Change

Years Ended December 31,

Revenue:

Product revenue 

Service and other revenue 

Total revenue 

Cost of Revenue:

Cost of product revenue 

Cost of service and other revenue 

Amortization of acquired intangible assets

Loss on purchase commitment

Total cost of revenue 

Gross profit 

Operating Expense:

Research and development 

Sales, general and administrative 

Merger-related expenses

Amortization of acquired intangible assets

Change in fair value of contingent consideration

Total operating expense 

$ 

183,872  $ 

108,699  $ 

75,173 

16,649 

200,521 

127,568 

14,754 

1,983 

3,436 

147,741 

52,780 

187,170 

169,818 

9,042 

6,157 

15,060 

387,247 

19,605 

128,304 

60,932 

13,899 

733 

3,705 

79,269 

49,035 

193,000 

160,854 

— 

— 

2,377 

356,231 

(2,956) 

72,217 

66,636 

855 

1,250 

(269) 

68,472 

3,745 

(5,830) 

8,964 

9,042 

6,157 

12,683 

31,016 

Operating loss 

(334,467)   

(307,196)   

(27,271) 

Loss on extinguishment of debt

Interest expense 

Other income, net 

Loss before benefit from income taxes

Benefit from income taxes

Net loss

Revenue

(2,033)   

— 

(14,343)   

(14,690)   

32,684 

7,638 

(318,159)   

(314,248)   

(11,424)   

— 

(2,033) 

347 

25,046 

(3,911) 

(11,424) 

$ 

(306,735)  $ 

(314,248)  $ 

7,513 

 69% 

 (15%) 

 56% 

 109% 

 6% 

 171% 

 (7%) 

 86% 

 8% 

 (3%) 

 6% 

 — 

 — 

 534% 

 9% 

 9% 

 — 

 (2%) 

 328% 

 1% 

 — 

 (2%) 

The increase in product revenue resulted primarily from an increase of $71.7 million in instrument revenue, as 
well as an increase of $3.4 million in consumable revenue. 

The  increase  in  instrument  revenue  was  primarily  due  to  the  sale  of  173  Revio  systems  that  have  a  higher 
average  selling  price  as  compared  to  the  Sequel  II/IIe  platform.  We  expect  the  installed  base  of  Revio 
instruments to grow, reflecting customer demand for the new product. As a result of this new product launch, 
we anticipate the installed base and sales volumes of Sequel II/IIe to continue to decline compared to recent 

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quarters.    Additionally,  we  commenced  the  shipment  of  Onso  products  during  the  year  ended  December  31, 
2023 and expect the installed base to continue to grow. 

The  increase  in  consumable  sales  was  primarily  due  to  higher  Revio  consumables  sales  attributable  to  the 
growth in the Revio instrument installed base, partially offset by a decline in Sequel consumables as customers 
transition  to  the  new  platform.  As  the  Revio  installed  base  continues  to  grow,  we  anticipate  the  related 
consumable sales to continue to increase.

The  decrease  in  service  and  other  revenue  was  primarily  due  to  the  change  in  our  terms  of  the  warranty 
provided with the instrument during the first quarter of 2022 to remove the service component.  As a result, the 
warranty  is  no  longer  a  separate  performance  obligation  and,  accordingly,  we  accrue  for  the  cost  of  the 
assurance warranty when revenue of the instrument is recognized, and no longer recognize a component of the 
instrument  revenue  in  service  and  other  revenue  over  the  warranty  period.  Service  revenue  also  declined  as 
customers transition to Revio, which includes a first-year warranty, and opt to not renew their Sequel II/IIe plans. 
As the Revio installed base begins to surpass the warranty period in 2024 and customers transition to service 
plans,  we  expect  the  retrospective  decline  in  growth  trend  to  reverse  within  the  year  and  service  revenues  to 
potentially exceed 2023 levels.

Cost of Revenue, Gross Profit, and Gross Margin

The  increase  in  the  cost  of  product  revenue  was  driven  primarily  by  an  increase  in  system  placements  and 
higher  overall  product  costs  on  the  Revio  platform,  including  warranty  costs,  as  well  as  an  increase  in 
adjustments  of  approximately  $4.6  million  as  compared  to  the  prior  year  primarily  relating  to  excess 
consumables inventory resulting from faster-than-expected decline in demand of Sequel II/IIe consumables due 
primarily  to  a  faster  than  expected  ramp  on  the  Revio  system.  Cost  of  revenue  included  share-based 
compensation  expense  of  $5.4  million  and  $4.8  million  during  the  years  ended  December  31,  2023  and  2022 
respectively.

The loss on purchase commitment was $3.4 million and $3.7 million for the years ended December 31, 2023 
and  2022,  respectively.  The  purchase  commitment  loss  is  based  on  an  estimate  of  future  excess  inventory 
related to supply agreements, for which we do not expect to have related sales.

Gross  profit  increased  $3.7  million,  or  8%.  Gross  margin  was  26.3%  for  the  year  ended  December  31,  2023 
compared  to  38.2%  for  the  year  ended  December  31,  2022.  The  decrease  in  gross  margin  percentage  was 
primarily  due  to  instrument  mix,  in  addition  to  charges  for  scrap  inventory  and  an  increase  in  adjustments 
primarily relating to excess consumables inventory resulting from a faster-than-expected decline in demand of 
Sequel II/IIe consumables due primarily to a faster than expected ramp on the Revio system. Revio instruments 
sold  during  the  period  had  a  lower  margin  primarily  due  to  loyalty  discounts  provided  and  higher  initial 
manufacturing costs, including warranty costs. Gross margin could fluctuate depending on the pace at which 
Sequel II/IIe revenue declines, Revio consumable revenue ramps, manufacturing efficiencies and warranty costs 
improve, as well as fluctuation in average selling prices.

Research and Development Expense

The  decrease  in  research  and  development  expense  was  primarily  driven  by  the  transition  of  Revio  from 
development to commercialization. Research and development expense included share-based compensation of 
$22.4 million and $30.7 million during the years ended December 31, 2023 and 2022, respectively.

Sales, General, and Administrative Expense

The  increase  in  sales,  general,  and  administrative  expense  was  primarily  driven  by  an  increase  in  sales  and 
marketing  headcount  as  we  continue  to  grow  our  commercial  footprint.  Sales,  general,  and  administrative 
expense  included  share-based  compensation  expenses  of  $44.3  million  and  $43.1  million  during  the  years 
ended December 31, 2023 and 2022, respectively.

Merger-Related Expenses

Merger-related  expenses  of  $9.0  million  during  the  year  ended  December  31,  2023,  consist  of  $4.9  million  of 
transaction  costs  arising  from  the  acquisition  of  Apton,  $2.8  million  of  compensation  expense  resulting  from 
the  liquidity  event  bonus  plan  in  connection  with  the  Apton  acquisition,  and  $1.3  million  of  share-based 
compensation  expense  resulting  from  the  acceleration  of  certain  equity  awards  in  connection  with  the  Apton 
acquisition. We recognized $1.3 million of share-based compensation expense for the acceleration that was not 
attributable to pre-combination services.

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Amortization of Acquired Intangible Assets

Amortization of acquired intangible assets of $6.2 million during the year ended December 31, 2023 consists of 
amortization expense attributable to acquired intangible assets that are not directly related to sales generating 
activities.

Change in Fair Value of Contingent Consideration

The change in fair value of contingent consideration during the year ended December 31, 2023, represents the 
remeasurement impact of the Omniome and Apton contingent consideration due upon the achievement of the 
respective milestone. The increase in the change in fair value of contingent consideration during the year was 
primarily due to the Omniome contingent consideration and was primarily attributable to the passage of time, 
changes in the discount rates and probabilities of milestone achievement. 

The  contingent  consideration  milestone  for  the  Omniome  acquisition  was  defined  as  the  first  commercial 
shipment to a customer of both an instrument and related consumables, utilizing SBB technology. As a result of 
the  milestone  achievement  in  September  2023,  former  Omniome  securityholders  received  as  milestone 
consideration, among other things, an aggregate of approximately $100.9 million in cash and approximately 9.0 
million shares of our common stock. 

Loss on Extinguishment of Debt

Loss on extinguishment of debt of $2.0 million during the year ended December 31, 2023, represents the loss 
resulting from the difference in the fair value of the 2030 Notes and the principal, in addition to the write-off of 
the unamortized debt issuance costs on the portion of the 2028 Notes that were exchanged as part of the debt 
modification during the year ended December 31, 2023.

Interest Expense

Interest  expense  for  the  year  ended  December  31,  2023  was  $14.3  million  compared  to  $14.7  million  for  the 
year ended December 31, 2022 and was primarily comprised of interest on the Convertible Senior Notes.

Other Income, Net

The increase in other income, net was primarily driven by investment income. 

Benefit from Income Taxes

A deferred income tax benefit of $11.4 million for the year ended December 31, 2023, is related to the release of 
the  valuation  allowance  for  deferred  tax  assets  due  to  the  recognition  of  deferred  tax  liabilities  in  connection 
with the Apton acquisition. We maintain a valuation allowance on the net deferred tax assets of our U.S. entities 
as we have concluded that it is more likely than not that we will not realize our deferred tax assets. Accordingly, 
this benefit from income taxes is reflected on our consolidated statements of operations and comprehensive 
loss for the year ended December 31, 2023.

Liquidity and Capital Resources 

Our  primary  sources  of  liquidity,  other  than  our  holdings  of  cash,  cash  equivalents,  and  investments,  has 
primarily  been  through  the  issuance  of  debt  or  equity  securities,  together  with  cash  flow  from  operating 
activities. For example, in January 2023, as discussed above, we issued and sold an aggregate of 20,125,000 
shares  of  our  common  stock  in  a  follow-on  public  offering  for  aggregate  gross  proceeds  of  approximately 
$201.3  million.  We  have  historically  incurred,  and  expect  to  continue  to  incur,  operating  losses  and  generate 
negative cash flows from operations on an annual basis due to the investments we intend to make as described 
in  Results  of  Operations  above,  and  as  a  result,  we  may  require  additional  capital  resources  to  execute  our 
strategic initiatives to grow our business.

Cash, Cash Equivalents, and Investments

As  of  December  31,  2023,  we  had  $631.4  million  in  cash,  cash  equivalents,  and  investments,  compared  to 
$772.3  million  at  December  31,  2022.  The  decrease  was  primarily  attributable  to  $259.2  million  cash  used  in 
operating activities for the twelve months ended December 31, 2023.

Convertible Senior Notes

At December 31, 2022, we had $900 million of principal Convertible Senior Notes outstanding resulting from our 
February  9,  2021,  issuance  of  convertible  notes  due  2028  (the  “2028  Notes”)  with  an  aggregate  principal  of 
$900 million. The Notes bear interest at a rate of 1.50% per annum. Interest on the 2028 Notes is payable semi-

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annually in arrears on February 15 and August 15 commencing on August 15, 2021. The 2028 Notes will mature 
on February 15, 2028, subject to earlier conversion, redemption, or repurchase. The proceeds from the issuance 
of the convertible notes are being used to fund operations, strategic investments, and capital requirements. 

The 2028 Notes are convertible at the option of the holder at any time until the second scheduled trading day 
prior to the maturity date, including in connection with a redemption by us. The 2028 Notes are convertible into 
shares of our common stock based on an initial conversion rate of 22.9885 shares of common stock per $1,000 
principal amount of the 2028 Notes (which is equal to an initial conversion price of $43.50 per share), in each 
case subject to customary anti-dilution and other adjustments as a result of certain extraordinary transactions. 
Upon  conversion  of  the  2028  Notes,  we  may  elect  to  settle  such  conversion  obligation  in  shares,  cash  or  a 
combination of shares and cash. 

In  June  2023,  we  entered  into  a  privately  negotiated  exchange  agreement  with  the  holder  of  our  outstanding 
2028  Notes,  pursuant  to  which  we  issued  $441.0  million  in  aggregate  principal  amount  of  our  1.375% 
Convertible Senior Notes due in 2030 (the "2030 Notes") in exchange for $441.0 million principal amount of the 
2028  Notes.  Interest  on  the  2030  Notes  is  payable  semi-annually  in  arrears  on  June  15  and  December  15 
commencing  on  December  15,  2023.  The  2030  Notes  will  mature  on  December  15,  2030,  subject  to  earlier 
conversion, redemption, or repurchase. 

The 2030 Notes are convertible at the option of the holder at any time until the second scheduled trading day 
prior  to  the  maturity  date,  including  in  connection  with  a  redemption  by  the  Company.  The  2030  Notes  are 
convertible into shares of our common stock based on an initial conversion rate of 46.5116 shares of common 
stock per $1,000 principal amount of the 2030 Notes (which is equal to an initial conversion price of $21.50 per 
share),  in  each  case  subject  to  customary  anti-dilution  and  other  adjustments  as  a  result  of  certain 
extraordinary  transactions.  Upon  conversion  of  the  2030  Notes,  we  may  elect  to  settle  such  conversion 
obligation in shares, cash or a combination of shares and cash. 

With  certain  exceptions,  upon  a  change  of  control  of  our  company  or  the  failure  of  our  common  stock  to  be 
listed  on  certain  stock  exchanges,  the  holders  of  the  2028  Notes  and  2030  notes  may  require  that  we 
repurchase all or part of the principal amount of those Notes at a purchase price of par plus unpaid interest up 
to, but excluding, the maturity date.

The Indenture includes customary “events of default,” which may result in the acceleration of the maturity of the 
Notes under the Indenture. The Indenture also includes customary covenants for convertible notes of this type.

See Note 7. Convertible Senior Notes for further details.

Additional Capital Requirements 

We  believe  that  our  existing  cash,  cash  equivalents,  and  investments  will  be  sufficient  to  fund  our  projected 
operating and capital requirements for at least the next 12 months from the date of filing of this Annual Report 
on  Form  10-K  for  the  year  ended  December  31,  2023.  Operating  needs  include  planned  costs  to  operate  our 
business, including costs to fund working capital and capital expenditures. Recent and expected working and 
other capital requirements, in addition to the above matters, include: 

•

•

Our  purchase  orders  and  contractual  obligations  of  approximately  $109.9  million  as  of  December  31, 
2023,  which  consist  of  open  purchase  orders  and  contractual  obligations  in  the  ordinary  course  of 
business,  including  commitments  with  contract  manufacturers  and  suppliers  for  which  we  have  not 
received the goods or services. A majority of these purchase obligations are due within a year. Although 
open purchase orders are considered enforceable and legally binding, the terms generally allow us the 
option  to  cancel,  reschedule  and  adjust  our  requirements  based  on  our  business  needs  prior  to  the 
delivery of goods or performance of services.

As  described  in  more  detail  in  Note  8  -  Commitments  and  Contingencies  in  the  Notes  to  the 
Consolidated  Financial  Statements,  we  signed  a  Supply  Agreement,  which  was  amended  in  October 
2022, with a supplier for the purchase of certain products over the period of 2023 through 2026. As part 
of  the  Supply  Agreement,  we  made  a  $9.0  million  deposit  during  the  year  ended  December  31,  2022, 
and an additional deposit of $6.0 million in 2023, to secure the supply of certain products through the 
term  of  the  contract.  If  we  breach  the  minimum  volume  purchase  commitment  during  any  applicable 
year,  the  supplier  is  entitled  to  retain  a  portion  or  all  of  the  deposit  corresponding  to  that  year.  If  we 
terminate the Supply Agreement before January 15, 2027, Supplier will refund the remaining balance of 
the  Deposit.  If  the  supplier  breaches  its  minimum  volume  supply  commitment  during  any  applicable 
year  or  portions  thereof,  our  remedies  include  termination,  pursuit  of  damages,  or  pursuit  of  specific 

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performance.  If,  on  or  before  October  31,  2024,  we  commit  to  purchasing  a  set  amount  of  additional 
products during the calendar year 2026, Supplier will increase its maximum capacity guarantee to meet 
the  additional  demand.  Should  we  exercise  this  option,  we  will  be  required  to  make  an  additional 
deposit of $5.0 million to Supplier within 30 days of the exercise.

Our research and development expenditures of $187.2 million in 2023 and $193.0 million in 2022. While 
we expect to continue our investment in research and development in 2024, including enhancements of 
our  existing  products,  and  continued  development  of  other  new  technology  and  products,  we  expect 
research  and  development  expenses  to  decline  slightly  in  2024  as  compared  to  the  year  ended 
December 31, 2023 due to recent product transitions. 

Cash outflows for capital expenditures of $8.8 million in 2023 and $16.8 million in 2022. We expect to 
continue  to  invest  in  capital  expenditures  in  fiscal  2024  to  continue  to  support  manufacturing  and 
expansion  of  our  business,  and  anticipate  a  slight  increase  in  2024  as  compared  to  the  year  ended 
December 31, 2023. 

Amounts  related  to  future  lease  payments  for  operating  lease  obligations  at  December  31,  2023, 
totaling $46.7 million, with $12.1 million expected to be paid within the next 12 months.

Amounts due under the term loan acquired in connection with Omniome at December 31, 2023, totaling 
$0.5 million, the remainder of which is expected to be paid within the next 12 months. Please see Note 
6. Balance Sheet Components for additional information.

Payments made to third party collaborators to help advance our technologies and the capabilities of our 
products.  We  may  also  choose  to  drive  investments  to  help  create  an  ecosystem  of  customers, 
partners,  and  collaborators  whose  expertise  and  offerings  complement  and  enhance  the  capabilities 
and utility of our technology and increase genomic data available on our platforms.

Payments related to licensing and other arrangements, which are cancellable license agreements with 
third parties for certain patent rights and technology. Under the terms of these agreements, we may be 
obligated to pay royalties based on revenue from the sales of licensed products, or minimum royalties, 
whichever is greater, and license maintenance fees. The future license maintenance fees and minimum 
royalty payments under the license agreements are not deemed to be material.

•

•

•

•

•

•

Our future capital requirements and the adequacy of our available funds will depend on many factors, including: 

•

•

•

•

•

our  ability  to  successfully  commercialize  and  develop  products  and  solutions  that  address  customer 
needs;

the pace of adoption of our products and our ability to obtain new customers in markets; 

the progress of our research and development programs and our ability to initiate or expand research 
programs; 

our  ability  to  manage  manufacturing  and  production  costs,  including  purchase  obligations,  and 
litigation  costs,  including  the  costs  involved  in  preparing,  filing,  prosecuting,  defending  and  enforcing 
intellectual property rights; and

the  extent  to  which  we  engage  in  collaborations  with  partners  and  acquire  other  businesses  or 
technologies.

If  economic,  financial,  business,  or  other  factors  adversely  affect  our  ability  to  fund  our  projected  operating 
cash requirements, we may be required to obtain funding through traditional or alternative sources of financing. 
We cannot be certain that funds will be available on favorable terms, or at all. If we are required and unable to 
raise additional capital when desired, our business, operating results, and financial condition may be adversely 
affected. See our risk factor captioned “We are not cash flow positive and may not have sufficient cash to make 
required payments under the terms of our debt or fund our long-term planned operations” for more information.

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Cash Flow Summary

(in thousands)

Cash used in operating activities

Cash provided by investing activities

Cash provided by financing activities

Years Ended December 31,

2023

2022

$ 

(259,173)  $ 

(263,211) 

4,604 

108,891 

116,083 

9,622 

Net decrease in cash, cash equivalents and restricted cash

$ 

(145,678)  $ 

(137,506) 

Operating Activities

Our  primary  uses  of  cash  in  operating  activities  include  the  development  of  future  products  and  product 
enhancements, manufacturing, and support functions related to our sales, general, and administrative activities. 

Cash used in operating activities for the year ended December 31, 2023, of $259.2 million was due primarily to a 
$306.7 million net loss that was partially offset by non-cash items such as share-based compensation of $72.1 
million, a change in the estimated fair value of contingent consideration of $15.1 million, depreciation of $11.5 
million, inventory provision of $10.6 million, amortization of intangible assets of $8.3 million, and amortization 
of  right-of-use  assets  of  $6.8  million,  offset  by  accretion  of  discount  and  amortization  of  premium  on 
marketable securities, net of $12.8 million, and deferred income taxes of $11.4 million. Cash flow impact from 
changes in net operating assets and liabilities of $59.0 million, was primarily attributable to increases of $17.8 
million  in  accounts  receivable,  net,  $13.8  million  in  inventory,  net,  $9.0  million  in  prepaid  expenses  and  other 
assets,  and  decreases  of  $14.9  million  in  contingent  consideration  liability,  $10.4  million  in  deferred  revenue, 
and $8.8 million in operating lease liabilities, partially offset by increases of $13.1 million in accrued expenses, 
and $2.4 million in other liabilities.

Cash used in operating activities for the year ended December 31, 2022, of $263.2 million was due primarily to a 
$314.2 million net loss that was partially offset by non-cash items such as share-based compensation of $78.6 
million,  depreciation  of  $9.5  million,  amortization  of  right-of-use  assets  of  $6.9  million,  inventory  provision  of 
$6.0  million,  and  a  change  in  the  estimated  fair  value  of  contingent  consideration  of  $2.4  million.  Cash  flow 
impact  from  changes  in  net  operating  assets  and  liabilities  of  $54.0  million,  was  primarily  attributable  to 
increases of $33.9 million in inventory, net, $12.3 million in prepaid expenses and other assets, and decreases 
of  $7.7  million  in  operating  lease  liabilities,  $3.7  million  in  accrued  expenses,  and  $3.7  million  in  deferred 
revenue, partially offset by a decrease of $5.5 million in accounts receivable, net, an increase of $1.0 million in 
accounts payable, and $0.9 million in other liabilities.

Investing Activities 

Our investing activities consist primarily of capital expenditures and investment purchases and maturities. Cash 
provided by investing activities for the year ended December 31, 2023, was due primarily to capital expenditures 
of $8.8 million and purchases  of investments of  $756.6 million offset by maturities of investments of $769.5 
million.

Cash  used  in  investing  activities  for  the  year  ended  December  31,  2022,  was  due  primarily  to  capital 
expenditures  of  $16.8  million  and  purchases  of  investments  of  $442.8  million  offset  by  maturities  of 
investments of $575.8 million.

Financing Activities 

Cash  provided  by  financing  activities  during  the  year  ended  December  31,  2023,  resulted  from  net  proceeds 
from issuance of common stock under equity offerings of $189.2 million, net proceeds of $15.3 million from the 
issuance of common stock through our equity compensation plans, partially offset by $86.4 million due to the 
payment of contingent consideration, $7.4 million due to the payment of debt issuance costs, and $1.8 million 
due to the payment of notes payable. 

Cash provided by financing activities during the year ended December 31, 2022, resulted from net proceeds of 
$11.2  million  from  the  issuance  of  common  stock  through  our  equity  compensation  plans,  partially  offset  by 
$1.6 million due to the payment of notes payable. 

Off-Balance Sheet Arrangements

As of December 31, 2023, we did not have any off-balance sheet arrangements.

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In  the  ordinary  course  of  business,  we  enter  into  standard  indemnification  arrangements.  Pursuant  to  these 
arrangements, we indemnify, hold harmless, and agree to reimburse the indemnified parties for losses suffered 
or incurred by the indemnified party in connection with any trade secret, copyright, patent or other intellectual 
property  infringement  claim  by  any  third  party  with  respect  to  its  technology,  or  from  claims  relating  to  our 
performance  or  non-performance  under  a  contract,  any  defective  products  supplied  by  us,  or  any  acts  or 
omissions, or willful misconduct, committed by us or any of our employees, agents or representatives. The term 
of these indemnification agreements is generally perpetual after the execution of the agreement. The maximum 
potential amount of future payments we could be required to make under these agreements is not determinable 
because it involves claims that may be made against us in future periods but have not yet been made. To date, 
we have not incurred costs to defend lawsuits or settle claims related to these indemnification agreements.

We  also  enter  and  have  entered  into  indemnification  agreements  with  our  directors  and  officers  that  may 
require  us  to  indemnify  them  against  liabilities  that  arise  by  reason  of  their  status  or  service  as  directors  or 
officers,  except  as  prohibited  by  applicable  law.  In  addition,  we  may  have  obligations  to  hold  harmless  and 
indemnify  third  parties  involved  with  our  fundraising  efforts  and  their  respective  affiliates,  directors,  officers, 
employees, agents or other representatives against any and all losses, claims, damages and liabilities related to 
claims arising against such parties pursuant to the terms of agreements entered into between us and such third 
parties in connection with such fundraising efforts. To the extent that such indemnification obligations apply to 
the lawsuits described in Legal Proceedings in Part I, Item 3 of this Annual Report on Form 10-K, any associated 
expenses  incurred  are  included  within  the  related  accrued  litigation  expense  amounts.  No  additional  liability 
associated with such indemnification agreements has been recorded as of December 31, 2023.

Critical Accounting Policies and Estimates 

Management’s  Discussion  and  Analysis  of  Financial  Condition  and  Results  of  Operations  is  based  upon  our 
Consolidated Financial Statements, which we have prepared in accordance with U.S. GAAP. The preparation of 
these financial statements requires management to make estimates and assumptions that affect the reported 
amounts  of  assets,  liabilities,  revenue,  cost  of  revenue,  and  operating  expenses,  and  related  disclosure  of 
contingent assets and liabilities. Management based its estimates on historical experience and on various other 
assumptions that it believes to be reasonable under the circumstances, the results of which form the basis for 
making judgements about the carrying values of assets and liabilities that are not readily apparent from other 
sources. Actual results may differ materially from these estimates under different assumptions or conditions.

An  accounting  policy  is  deemed  to  be  critical  if  it  requires  an  accounting  estimate  to  be  made  based  on 
assumptions  about  matters  that  are  highly  uncertain  at  the  time  the  estimate  is  made,  if  different  estimates 
reasonably  could  have  been  used,  or  if  changes  in  the  estimate  that  are  reasonably  likely  to  occur  could 
materially impact the financial statements. 

Revenue Recognition 

Our revenue is generated primarily from the sale of products and services. Product revenue primarily consists of 
sales  of  our  instruments  and  related  consumables;  service  and  other  revenue  consist  primarily  of  revenue 
earned from product maintenance agreements.

We  account  for  a  contract  with  a  customer  when  there  is  a  legally  enforceable  contract  between  us  and  the 
customer, the rights of the parties are identified, the contract has commercial substance, and collectability of 
the  contract  consideration  is  probable.  Revenues  are  recognized  when  control  of  the  promised  goods  are 
transferred to our customers, or services are performed, in an amount that reflects the consideration we expect 
to be entitled to in exchange for those goods or services. 

We  may  enter  into  contracts  with  customers  that  include  a  combination  of  promised  products  and  services, 
resulting in arrangements containing multiple performance obligations. We determine whether each product or 
service  is  distinct,  in  order  to  identify  the  performance  obligations  in  the  contract  and  allocate  the  contract 
transaction price among the distinct performance obligations. A performance obligation is considered distinct 
from other obligations in a contract when it provides a benefit to the customer either on its own or together with 
other  resources  that  are  readily  available  to  the  customer  and  is  separately  identified  in  the  contract.  We 
consider  a  performance  obligation  satisfied  once  we  have  transferred  control  of  a  good  or  service  to  the 
customer, meaning the customer has the ability to use and obtain the benefit of the good or service. Therefore, 
instrument revenue is recognized upon transfer of control of the asset to the customer, which is generally upon 
delivery for sales made to our non-distributor customers and upon shipment for sales made to our distributor 
customers. 

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The  consideration  for  contracts  with  multiple  performance  obligations  is  allocated  between  separate 
performance obligations based on their individual standalone selling price. We determine the best estimate of 
standalone selling price using average selling prices over a 12-month period combined with an assessment of 
current  market  conditions.  If  the  standalone  selling  price  is  not  directly  observable,  we  rely  on  estimates  by 
considering  multiple  factors  including,  but  not  limited  to,  overall  market  conditions,  including  geographic  or 
regional  specific  factors,  internal  costs,  profit  objectives,  pricing  practices,  and  other  observable  inputs.  We 
recognize revenues as performance obligations are satisfied by transferring control of the product or service to 
the  customer  or  over  the  term  of  a  product  maintenance  agreement  with  a  customer.  Our  revenue 
arrangements generally do not provide a right of return.

Certain of our agreements provide options to customers which can be exercised at a future date, such as the 
option  to  purchase  our  product  at  discounted  prices,  among  others.  In  accounting  for  customer  options,  we 
determine  whether  an  option  is  a  material  right  and  this  requires  us  to  exercise  significant  judgment.  If  a 
contract provides the customer an option to acquire additional goods or services at a discount that exceeds the 
range  of  discounts  that  we  typically  give  for  that  product  or  service  for  the  same  class  of  customer,  or  if  the 
option  provides  the  customer  certain  additional  goods  or  services  for  free,  the  option  may  be  considered  a 
material  right.  If  the  contract  gives  the  customer  the  option  to  acquire  additional  goods  or  services  at  their 
normal standalone selling prices, we would likely determine that the option is not a material right and, therefore, 
account for it when the customer exercises the option. If the standalone selling price of the option is not directly 
observable, an estimated standalone selling price is utilized which considers adjustments for discounts that the 
customer  could  receive  without  exercising  the  option  and  the  likelihood  that  the  option  will  be  exercised.  We 
may  also  utilize  the  alternative  approach  to  estimate  the  standalone  selling  price,  available  pursuant  to  the 
applicable  accounting  guidance,  to  the  extent  we  conclude  the  applicable  criteria  for  using  the  alternative 
approach has been met. We update the transaction price for expected consideration, subject to constraint, each 
reporting period if our estimate of future goods to be ordered by customers change.

Inventories 

Inventories are stated at the lower of cost or net realizable value. Cost is determined using the first-in, first-out 
(“FIFO”) method. Adjustments to reduce the cost of inventory to its net realizable value, if required, are made for 
estimated  excess  or  obsolete  balances.  Cost  includes  depreciation,  labor,  material,  and  overhead  costs, 
including  product  and  process  technology  costs.  Determining  net  realizable  value  of  inventories  involves 
numerous judgements, including projecting future average selling prices, sales volumes, and costs to complete 
products in work in process inventories. 

We  make  inventory  purchases  and  commitments  to  meet  future  shipment  schedules  based  on  forecasted 
demand  for  our  products.  The  business  environment  in  which  we  operate  is  subject  to  rapid  changes  in 
technology and customer demand. We perform a detailed assessment of inventory each period, which includes 
a review of, among other factors, demand requirements, product life cycle and development plans, component 
cost  trends,  product  pricing,  product  expiration,  and  quality  issues.  Based  on  our  analysis,  we  record 
adjustments  to  inventory  for  potentially  excess,  obsolete,  or  impaired  goods,  when  appropriate,  to  report 
inventory  at  net  realizable  value.  Inventory  adjustments  may  be  required  if  actual  demand,  component  costs, 
supplier arrangements, or product life cycles differ from our estimates. Any such adjustments would result in a 
charge to our results of operations.

Business Combinations

Under the acquisition method of accounting, we allocate the fair value of the total consideration transferred to 
the  tangible  and  identifiable  intangible  assets  acquired  and  liabilities  assumed  based  on  their  estimated  fair 
values on the date of acquisition. The fair values assigned, defined as the price that would be received to sell an 
asset or paid to transfer a liability in an orderly transaction between willing market participants, are based on 
estimates  and  assumptions  determined  by  management.  These  valuations  require  us  to  make  estimates  and 
assumptions,  especially  with  respect  to  intangible  assets.  We  record  the  excess  consideration  over  the 
aggregate  fair  value  of  tangible  and  intangible  assets,  net  of  liabilities  assumed,  as  goodwill.  Costs  that  we 
incur to complete the business combination, such as legal and other professional fees, are expensed as they 
are incurred.

In connection with certain acquisitions, contingent consideration can be earned by the sellers upon completion 
of certain future performance milestones. In these cases, a liability is recorded on the acquisition date for an 
estimate  of  the  acquisition  date  fair  value  of  the  contingent  consideration.  Changes  in  the  fair  value  of 
contingent  consideration  subsequent  to  the  acquisition  date  are  recognized  in  operating  expenses  in  our 
consolidated statements of operations and comprehensive loss.

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We typically use the discounted cash flow method to value our acquired intangible assets. This method requires 
significant management judgment to forecast future operating results and utilizes significant assumptions such 
as assumed revenue projections, discount rates and obsolescence factors. The estimates we use to value and 
amortize intangible assets are consistent with the plans and estimates that we use to manage our business and 
are  based  on  available  historical  information  and  industry  estimates  and  averages.  If  the  subsequent  actual 
results and updated projections of the underlying business activity change compared with the assumptions and 
projections  used  to  develop  these  values,  we  could  experience  impairment  charges.  In  addition,  we  have 
estimated the economic lives of certain acquired assets and these lives are used to calculate depreciation and 
amortization  expense.  If  our  estimates  of  the  economic  lives  change,  depreciation  or  amortization  expense 
could  be  accelerated  or  extended.  We  capitalize  in-process  research  and  development  (IPR&D),  which  is 
considered  indefinite  lived  until  the  completion  or  abandonment  of  the  associated  research  and  development 
efforts. Upon reaching the end of the relevant research and development project (i.e., upon commercialization), 
the IPR&D asset is amortized over its estimated useful life. If the relevant research and development project is 
abandoned, the IPR&D asset is expensed in the period of abandonment.

If  the  initial  accounting  for  a  business  combination  is  incomplete  by  the  end  of  a  reporting  period  that  falls 
within  the  measurement  period,  we  report  provisional  amounts  in  our  financial  statements.  During  the 
measurement  period,  we  adjust  the  provisional  amounts  recognized  at  the  acquisition  date  to  reflect  new 
information  obtained  about  facts  and  circumstances  that  existed  as  of  the  acquisition  date  that,  if  known, 
would have affected the measurement of the amounts recognized as of that date. We record these adjustments 
to  the  provisional  amounts  with  a  corresponding  offset  to  goodwill.  Any  adjustments  identified  after  the 
measurement period are recorded in the consolidated statements of operations and comprehensive loss.

We acquired $55.0 million of IPR&D, and $52.3 million of goodwill in connection with the acquisition of Apton 
Biosystems, Inc. in the third quarter of 2023. 

Goodwill and Intangible Assets with Indefinite Lives — Impairment Assessment

Goodwill and other intangible assets with indefinite useful lives (i.e., IPR&D) are not amortized, however they are 
tested annually for impairment, in the second and fourth quarter of our fiscal year, respectively, and whenever 
events  or  changes  in  circumstances  indicate  that  it  is  more  likely  than  not  that  the  fair  value  is  less  than  the 
carrying value. Events that would indicate impairment and trigger an interim impairment test include, but are not 
limited to, unexpected adverse business conditions, economic factors, unanticipated technological changes or 
competitive activities, loss of key personnel and acts by governments or courts.

We  perform  our  goodwill  impairment  analysis  at  the  reporting  unit  level.  We  have  one  reporting  unit,  which 
aligns  with  our  reporting  structure  and  availability  of  discrete  financial  information.  During  the  goodwill 
impairment  review,  we  assess  qualitative  factors  to  determine  whether  it  is  more  likely  than  not  that  the  fair 
value of our reporting unit is less than the carrying amount, including goodwill. The qualitative factors include, 
but are not limited to, macroeconomic conditions, industry and market considerations, and our overall financial 
performance. If, after assessing the totality of these qualitative factors, we determine that it is not more likely 
than not that the fair value of our reporting unit is less than the carrying amount, then no additional assessment 
is deemed necessary. Otherwise, we proceed to compare the estimated fair value of the reporting unit with the 
carrying value, including goodwill. If the carrying amount of the reporting unit exceeds the fair value, we record 
an impairment loss based on the difference. If a quantitative assessment is performed, the evaluation includes 
management  estimates  of  cash  flow  projections  based  on  internal  future  projections  and/or  use  of  a  market 
approach by looking at market values of comparable companies. Key assumptions include, but are not limited 
to,  revenue  and  operating  income  growth  rates,  discount  rates  and  other  factors.  We  consider  peer  revenues 
and  earnings  trading  multiples  from  companies  that  have  operational  and  financial  characteristics  that  are 
similar  to  the  asset  under  measurement  and  estimated  weighted-average  costs  of  capital.  Different 
assumptions from those made in our analysis could materially affect projected cash flows and the evaluation of 
assets  for  impairment.  We  also  consider  our  market  capitalization  as  a  part  of  our  analysis.  We  may  elect  to 
bypass  the  qualitative  assessment  in  a  period  and  proceed  to  perform  the  quantitative  goodwill  impairment 
test.

During the IPR&D impairment review, we assess qualitative factors to determine whether it is more likely than 
not that the fair value of the IPR&D is less than the carrying amount. The qualitative factors include, but are not 
limited  to,  macroeconomic  conditions,  industry-specific  conditions,  and  company-specific  conditions.  If,  after 
assessing the totality of these qualitative factors, we determine that it is not more likely than not that the fair 
value  of  the  IPR&D  is  less  than  the  carrying  amount,  then  no  additional  assessment  is  deemed  necessary. 
Otherwise, we proceed to compare the estimated fair value of the IPR&D with the carrying value. If the carrying 

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amount  of  the  IPR&D  exceeds  the  fair  value,  we  record  an  impairment  loss  based  on  the  difference.  If  a 
quantitative assessment is performed, the evaluation includes management estimates of cash flow projections 
based  on  internal  future  projections.  Key  assumptions  include,  but  are  not  limited  to,  revenue  projections, 
revenue  growth  rates,  discount  rates  and  other  factors.  We  consider  peer  revenues  and  earnings  trading 
multiples from companies that have operational and financial characteristics that are similar to the asset under 
measurement and estimated weighted-average costs of capital. Different assumptions from those made in our 
analysis could materially affect projected cash flows and the evaluation of assets for impairment. We may elect 
to bypass the qualitative assessment in a period and proceed to perform the quantitative impairment test.

Intangible Assets and Other Long-Lived Assets — Impairment Assessment

We perform regular reviews to determine if any event has occurred that may indicate that the carrying values of 
our intangible assets with finite lives and other long-lived assets are impaired. If indicators of impairment exist, 
we assess the recoverability of the affected assets by determining whether their carrying amounts exceed their 
undiscounted expected future cash flows. If the affected assets are not recoverable, we estimate the fair value 
of  the  assets  and  record  an  impairment  loss  if  the  carrying  value  exceeds  the  fair  value.  Factors  that  may 
indicate  potential  impairment  include  a  significant  decline  in  our  stock  price  and  market  capitalization 
compared to net book value, significant changes in the ability of an asset to generate positive cash flows and 
the pattern of utilization of a particular asset.

In order to estimate the fair values of identifiable intangible assets with finite lives and other long-lived assets, 
we estimate the present value of future cash flows from those assets. The key assumptions that we use in our 
discounted cash flow model are the amount and timing of estimated future cash flows to be generated by the 
asset over an extended period of time and a rate of return that considers the relative risk of achieving the cash 
flows, the time value of money, and other factors that a willing market participant would consider. Significant 
judgment is required to estimate the amount and timing of future cash flows and the relative risk of achieving 
those cash flows.

Assumptions and estimates about future values and remaining useful lives are complex and often subjective. 
They can be affected by a variety of factors, including external factors such as industry and economic trends, 
and internal factors such as changes in our business strategy and our internal forecasts. 

Contingent Consideration

In  connection  with  the  acquisition  of  Omniome  in  the  third  quarter  of  2021,  we  entered  into  an  arrangement 
where  we  were  obligated  to  pay  $200  million  in  cash  and  equity  dependent  upon  the  achievement  of  a 
milestone  event  upon  the  first  commercial  shipment  of  products  developed  from  our  acquired  sequencing 
technology.  In  the  third  quarter  of  2023,  we  commenced  customer  shipments  of  the  Onso  short-read 
sequencing  instrument.  The  milestone  payment  associated  with  PacBio’s  acquisition  of  Omniome  was 
triggered in September 2023 once both the Onso instrument and related consumables had been shipped to one 
customer.  Consequently,  we  paid  the  former  Omniome  security  holders  milestone  consideration  of  an 
aggregate of approximately $100.9 million in cash and approximately 9.0 million shares of our common stock in 
October 2023.

In  connection  with  the  acquisition  of  Apton,  we  entered  into  an  arrangement  where  we  are  obligated  to  pay 
former  holders  of  Apton's  outstanding  equity  interests  $25.0  million  upon  the  achievement  of  $50  million  in 
revenue  associated  with  a  high  throughput  sequencer  using  Apton's  technology,  provided  that  the  milestone 
event occurs prior to the 5-year anniversary of the closing date of the acquisition, which we may elect to pay in 
cash,  shares  of  our  common  stock  or  a  combination  of  cash  and  shares  of  our  common  stock.  See  Note  2. 
Business Acquisitions for further information.

The contingent consideration liability was measured at fair value as of the acquisition date and is remeasured 
periodically  at  each  reporting  date,  with  changes  in  fair  value  recorded  as  change  in  fair  value  of  contingent 
consideration  in  the  consolidated  statements  of  operations  and  comprehensive  loss.  For  the  Omniome 
contingent consideration, the initial measurement and post-acquisition remeasurement required estimates and 
assumptions  using  a  scenario-based  method  that  considers  a  range  of  potential  outcomes  of  milestone 
achievement dates and assigned probabilities of occurrence for each outcome. Outcomes were discounted to 
present value, which was then weighted by the probability of each scenario to determine the total fair value of 
the  contingent  consideration  payment  as  of  each  reporting  period.  This  method  requires  significant 
management judgment, including the probability of achieving certain future milestones and discount rates. For 
the  Apton  contingent  consideration,  the  initial  measurement  and  post-acquisition  remeasurement  required 
estimates  and  assumptions  using  a  Monte  Carlo  Simulation  to  estimate  the  volatility  and  systematic  relative 

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risk of revenues subject to sales milestone payments and discounting the associated cash payment amounts to 
their present values using a credit-risk-adjusted interest rate to determine the total fair value of the contingent 
consideration  payment  as  of  each  reporting  period.  This  method  requires  significant  management  judgment, 
including  risk-adjusted  forecasted  revenues  for  products  and  services  leveraging  Apton's  technology  and  an 
estimated  credit  spread.  Future  changes  in  our  estimates  could  result  in  expenses  or  gains.  Refer  to  Note  5. 
Financial Instruments for further discussion on valuation assumptions.

Recent Accounting Pronouncements

Please  see  Note  1.  Organization  and  Significant  Accounting  Policies,  subsection  titled  “Recent  Accounting 
Pronouncements”,  in  Part  II,  Item  8  of  this  Annual  Report  on  Form  10-K  for  information  regarding  applicable 
recent accounting pronouncements.

ITEM 7A.  QUANTITATIVE AND QUALITATIVE DISCLOSURES ABOUT MARKET RISK 

Interest Rate and Market Risk

Our investment portfolio is exposed to market risk from changes in interest rates. The goals of our investment 
policy  are  preservation  of  capital,  fulfillment  of  liquidity  needs  and  fiduciary  control  of  cash  and  cash 
equivalents  and  investments.  We  also  seek  to  maximize  income  from  our  investments  without  assuming 
significant risk. To achieve our goals, we maintain a portfolio of cash equivalents and investments in a variety of 
securities  of  high  credit  quality.  The  securities  in  our  investment  portfolio  are  not  leveraged,  are  classified  as 
available for sale and are, due to their short-term nature, subject to minimal interest rate risk. The fair market 
value of our fixed rate securities may be adversely impacted by increases in interest rates while income earned 
may  decline  as  a  result  of  decreases  in  interest  rates.  A  hypothetical  100  basis-point  (one  percentage  point) 
increase  or  decrease  in  interest  rates  compared  to  rates  on  December  31,  2023  would  have  affected  the  fair 
value of our investment portfolio by approximately $2.7 million.

The  2030  Notes  were  recorded  at  fair  value  as  of  the  closing  date  of  the  Exchange  Transaction,  less  debt 
issuance  costs,  on  our  consolidated  balance  sheets.  We  carry  our  remaining  2028  Notes  at  the  principal 
amount, less unamortized debt issuance costs, on our consolidated balance sheets. Because the 2030 Notes 
and  2028  Notes  have  fixed  annual  interest  rates  of  1.375%  and  1.50%,  respectively,  we  do  not  have  any 
economic interest rate exposure or financial statement risk associated with changes in interest rates. The fair 
value of the notes, however, may fluctuate when interest rates and the market price of our stock changes. See 
Note 7. Convertible Senior Notes in Part II, Item 8 of this Annual Form 10-K for additional information. 

Foreign Exchange Risk

Our  revenue,  expense,  and  capital  purchasing  activities  are  primarily  transacted  in  U.S.  dollars;  however,  a 
portion of our operations is conducted in foreign currencies. As a result, we have foreign exchange exposures 
relating to non-U.S. dollar denominated cash flows and monetary assets and liabilities that are denominated in 
currencies other than U.S. dollars. The value of the amounts is exposed to changes in currency exchange rates 
from the time the transactions are originated, until the time the cash settlement is converted into U.S. dollars. 
Our  foreign  currency  exposure  is  primarily  concentrated  in  the  Euro.  A  10%  strengthening  of  the  U.S.  dollar 
exchange rate against all currencies with which we have exposure, after taking into account offsetting positions 
at  December  31,  2023  would  have  resulted  in  a  $0.9  million  decrease  in  the  carrying  amounts  of  those  net 
assets.  Actual  gains  and  losses  in  the  future  may  differ  materially  from  these  hypothetical  gains  and  losses 
based  on  changes  in  the  timing  and  amount  of  foreign  currency  exchange  rate  movements  and  our  actual 
exposure. Our international operations are subject to risks typical of international operations, including, but not 
limited to, differing economic conditions, changes in political climate, differing tax structures, other regulations 
and restrictions, and foreign exchange rate volatility.

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ITEM 8. FINANCIAL STATEMENTS AND SUPPLEMENTARY DATA

PACIFIC BIOSCIENCES OF CALIFORNIA, INC. 
Index to Consolidated Financial Statements 

Report of Independent Registered Public Accounting Firm (PCAOB ID: 42)

Consolidated Financial Statements

Consolidated Balance Sheets

Consolidated Statements of Operations and Comprehensive Loss

Consolidated Statements of Stockholders’ Equity

Consolidated Statements of Cash Flows

Notes to Consolidated Financial Statements

Page(s)

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To the Stockholders and the Board of Directors of Pacific Biosciences of California, Inc.

Report of Independent Registered Public Accounting Firm

Opinion on the Financial Statements

We have audited the accompanying consolidated balance sheets of Pacific Biosciences of California, Inc. (the 
Company)  as  of  December  31,  2023  and  2022,  the  related  consolidated  statements  of  operations  and 
comprehensive  loss,  stockholders’  equity  and  cash  flows  for  each  of  the  three  years  in  the  period  ended 
December 31, 2023, and the related notes (collectively referred to as the “consolidated financial statements”). In 
our opinion, the consolidated financial statements present fairly, in all material respects, the financial position 
of the Company at December 31, 2023 and 2022, and the results of its operations and its cash flows for each of 
the three years in the period ended December 31, 2023, in conformity with U.S. generally accepted accounting 
principles.

We  also  have  audited,  in  accordance  with  the  standards  of  the  Public  Company  Accounting  Oversight  Board 
(United  States)  (PCAOB),  the  Company's  internal  control  over  financial  reporting  as  of  December  31,  2023, 
based on criteria established in Internal Control–Integrated Framework issued by the Committee of Sponsoring 
Organizations  of  the  Treadway  Commission  (2013  framework)  and  our  report  dated  February  28,  2024, 
expressed an unqualified opinion thereon.

Basis for Opinion

These  financial  statements  are  the  responsibility  of  the  Company's  management.  Our  responsibility  is  to 
express an opinion on the Company’s financial statements based on our audits. We are a public accounting firm 
registered with the PCAOB and are required to be independent with respect to the Company in accordance with 
the  U.S.  federal  securities  laws  and  the  applicable  rules  and  regulations  of  the  Securities  and  Exchange 
Commission and the PCAOB.

We conducted our audits in accordance with the standards of the PCAOB. Those standards require that we plan 
and  perform  the  audit  to  obtain  reasonable  assurance  about  whether  the  financial  statements  are  free  of 
material misstatement, whether due to error or fraud. Our audits included performing procedures to assess the 
risks  of  material  misstatement  of  the  financial  statements,  whether  due  to  error  or  fraud,  and  performing 
procedures  that  respond  to  those  risks.  Such  procedures  included  examining,  on  a  test  basis,  evidence 
regarding  the  amounts  and  disclosures  in  the  financial  statements.  Our  audits  also  included  evaluating  the 
accounting  principles  used  and  significant  estimates  made  by  management,  as  well  as  evaluating  the  overall 
presentation of the financial statements. We believe that our audits provide a reasonable basis for our opinion. 

Critical Audit Matters 

The critical audit matters communicated below are matters arising from the current period audit of the financial 
statements  that  were  communicated  or  required  to  be  communicated  to  the  audit  committee  and  that:  (1) 
relate  to  accounts  or  disclosures  that  are  material  to  the  financial  statements  and  (2)  involved  our  especially 
challenging, subjective or complex judgments. The communication of critical audit matters does not alter in any 
way our opinion on the consolidated financial statements, taken as a whole, and we are not, by communicating 
the critical audit matters below, providing separate opinions on the critical audit matters or on the accounts or 
disclosures to which they relate. 

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Description 
Matter 

of 

the 

Revenue recognition - Identification of performance obligations and 
allocation of contract consideration

For  the  year  ended  December  31,  2023,  the  Company  recognized 
revenue  of  $200.5  million,  including  $183.9  million  of  product 
revenue,  which  consists  primarily  of  instrument  sales  and  related 
consumables.  As  described  in  Note  1  to  the  consolidated  financial 
statements,  the  Company  may  enter  into  contracts  with  customers 
that  includes  a  combination  of  promised  products  and  services, 
resulting 
in  arrangements  containing  multiple  performance 
obligations.  The  Company  identifies  a  performance  obligation  for 
each promise to transfer to the customer, a product or service that is 
distinct.  The  consideration  is  allocated  between  each  performance 
obligation  based  on  its  individual  standalone  selling  price,  which  is 
estimated  by  the  Company,  using  historical  sales  data,  as  well  as 
management judgment.

The Company enters into, or periodically modifies, revenue contracts 
with non-standard terms, requiring management to evaluate whether 
these  non-standard  terms  represent  a  performance  obligation.  For 
example,  the  Company  may  offer  specified  discounts  on  current 
products within an arrangement and on future purchase options, for 
which  historical  information  may  not  be  available.  As  part  of  the 
Company's 
identification  of  performance  obligations  and  the 
resulting  determination  of  the  allocation  of  contract  consideration, 
the  Company  considers  if  these  specified  discounts  represent  a 
material  right  when  compared  to  the  estimated  standalone  selling 
price  and,  therefore,  a  performance  obligation  to  be  included  in  the 
allocation of the contract value.

identification  of 

Auditing  management’s 
the  performance 
obligations  and  the  resulting  determination  of  the  allocation  of 
contract consideration in certain contracts involved a higher degree 
of  judgment  due  to  the  subjective  nature  of  identifying  certain 
related  determination  of 
performance  obligations  and 
standalone  selling  price  when 
is  not  based  on  historical 
information.

the 
it 

How  We  Addressed  the 
Matter in Our Audit 

We obtained an understanding, evaluated the design, and tested the 
operating  effectiveness  of 
internal  controls 
addressing management’s identification of performance obligations 
including  standalone 
and  allocation  of  contract  consideration, 
selling price determination.

the  Company’s 

Our  audit  procedures  included,  among  others,  reading  executed 
contracts  for  a  sample  of  arrangements  and  evaluating  whether 
terms of the contracts (including specified discounts on current and 
future  purchase  options)  resulted 
in  additional  performance 
obligations.  Additionally,  we  tested  the  completeness  and  accuracy 
of  the  information  used  in  management’s  allocation  of  contract 
consideration, 
in  underlying 
calculations to determine standalone selling price.

incorporated 

the  data 

including 

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Description 
Matter 

of 

the 

How  We  Addressed  the 
Matter in Our Audit 

Business combination - Valuation of intangible asset

As described in Note 2 to the consolidated financial statements, the 
Company completed its acquisition of Apton Biosystems, Inc. during 
for  as  a  business 
2023.  The 
combination,  and 
indefinite-lived 
intangible asset of $55.0 million.

transaction  was  accounted 
the  Company 

recorded  an 

Auditing  the  Company’s  accounting  for  the  acquisition  was 
challenging  because  the  determination  of  the  fair  value  of  the 
identified  intangible  asset,  which  consisted  of  in-process  research 
and  development  (IPR&D),  required  management  to  make  certain 
subjective  estimates  and  assumptions.  The  Company  used  an 
income approach to measure the intangible asset. The valuation of 
the  intangible  asset  is  subject  to  higher  estimation  uncertainty  due 
to management’s judgments in determining significant assumptions, 
which  included  certain  components  of  the  revenue  projection  and 
the  discount  rate.  A  change  in  these  significant  assumptions  could 
have a significant effect on the fair value of the intangible asset.

We obtained an understanding, evaluated the design and tested the 
operating  effectiveness  of  controls  addressing  the  identified  audit 
risk.  For  example,  we  tested  controls  over  management’s  review  of 
the significant assumptions used to develop the fair value estimate 
of  the  intangible  asset.  We  also  tested  management’s  controls  to 
validate that data used in the fair value estimate was complete and 
accurate.

To  test  the  estimated  fair  value  of  the  intangible  asset,  we 
performed audit procedures that included, among others, evaluating 
the  Company’s  valuation  model  with  the  assistance  of  valuation 
specialists,  performing  sensitivity  analyses  to  determine  which 
assumptions had the greatest impact on the overall determination of 
value, and testing the completeness and accuracy of the underlying 
data  used  to  develop  the  assumptions.  We  also  evaluated  the 
assumptions  by  comparing  them  to  market  and  economic  trends, 
historical  results  of  the  Company’s  business  and  other  guideline 
companies within the same industry.

/s/ Ernst & Young LLP

We have served as the Company’s auditor since 2011.

San Mateo, California

February 28, 2024

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PACIFIC BIOSCIENCES OF CALIFORNIA, INC.

Consolidated Balance Sheets

(in thousands, except per share amounts)

Assets

Current assets 

Cash and cash equivalents 

Investments 

Accounts receivable, net

Inventory, net

Prepaid expenses and other current assets 

Short-term restricted cash

Total current assets 

Property and equipment, net 

Operating lease right-of-use assets, net 

Long-term restricted cash

Intangible assets, net

Goodwill

Other long-term assets 

Total assets 

Liabilities and Stockholders’ Equity 

Current liabilities

Accounts payable 

Accrued expenses 

Deferred revenue, current 

Operating lease liabilities, current

Other liabilities, current 

Contingent consideration liability, current

Total current liabilities 

Deferred revenue, non-current 

Contingent consideration liability, non-current

Operating lease liabilities, non-current 

Convertible senior notes, net, non-current

Other liabilities, non-current 

Total liabilities 

Commitments and contingencies 

Stockholders’ equity

Preferred stock, $0.001 par value:

December 31,

2023

2022

$ 

179,911  $ 

451,505 

36,615 

56,676 

17,040 

300 

325,089 

447,229 

18,786 

50,381 

10,289 

300 

742,047 

852,074 

36,432 

32,593 

2,422 

456,984 

462,261 

13,274 

41,580 

39,763 

2,922 

410,245 

409,974 

10,528 

$ 

1,746,013  $ 

1,767,086 

$ 

15,062  $ 

45,708 

16,342 

9,591 

8,326 

— 

95,029 

5,530 

19,550 

31,606 

892,243 

751 

12,028 

32,596 

30,498 

8,886 

7,233 

172,094 

263,335 

1,794 

— 

41,070 

896,683 

1,300 

1,044,709 

1,204,182 

Authorized 50,000 shares; No shares issued or outstanding 

— 

— 

Common stock, $0.001 par value:

Authorized 1,000,000 shares; issued and outstanding 267,744 and 226,505 shares at December 31, 
2023 and December 31, 2022, respectively 

Additional paid-in capital 

Accumulated other comprehensive income (loss)

Accumulated deficit 

Total stockholders’ equity 

268 

227 

2,539,892 

2,099,782 

219 

(4,765) 

(1,839,075) 

(1,532,340) 

701,304 

562,904 

Total liabilities and stockholders’ equity 

$ 

1,746,013  $ 

1,767,086 

See accompanying notes to the consolidated financial statements. 

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PACIFIC BIOSCIENCES OF CALIFORNIA, INC.

Consolidated Statements of Operations and Comprehensive Loss

(in thousands, except per share amounts)

Revenue:

Product revenue 

Service and other revenue 

Total revenue 

Cost of Revenue:

Cost of product revenue 

Cost of service and other revenue 

Amortization of acquired intangible assets

Loss on purchase commitment

Total cost of revenue 

Gross profit 

Operating Expense:

Research and development 

Sales, general and administrative 

Merger-related expenses

Change in fair value of contingent consideration

Amortization of acquired intangible assets

Total operating expense 

Operating loss 

Loss from Continuation Advances from Illumina

Loss on extinguishment of debt

Interest expense 

Other income, net 

Loss before benefit from income taxes

Benefit from income taxes

Net loss

Other comprehensive income (loss):

Unrealized gain (loss) on investments 

Comprehensive loss

Net loss per share:

Basic 

Diluted 

Years Ended December 31,

2023

2022

2021

$ 

183,872  $ 

108,699  $ 

113,505 

16,649 

200,521 

127,568 

14,754 

1,983 

3,436 

147,741 

52,780 

187,170 

169,818 

9,042 

15,060 

6,157 

19,605 

128,304 

17,008 

130,513 

60,932 

13,899 

733 

3,705 

79,269 

49,035 

193,000 

160,854 

— 

2,377 

— 

56,358 

14,989 

306 

— 

71,653 

58,860 

112,899 

124,124 

31,129 

1,143 

— 

387,247 

356,231 

269,295 

(334,467) 

(307,196) 

(210,435) 

— 

(2,033) 

(14,343) 

32,684 

— 

— 

(14,690) 

7,638 

(52,000) 

— 

(12,530) 

93 

(318,159) 

(314,248) 

(274,872) 

(11,424) 

— 

(93,649) 

(306,735) 

(314,248) 

(181,223) 

4,984 

(3,678) 

(1,172) 

(301,751)  $ 

(317,926)  $ 

(182,395) 

(1.21)  $ 

(1.21)  $ 

(1.40)  $ 

(1.40)  $ 

(0.89) 

(0.89) 

$ 

$ 

$ 

Weighted average shares outstanding used in calculating ͏net loss per share

Basic 

Diluted 

253,629 

253,629 

224,550 

224,550 

204,136 

204,136 

See accompanying notes to the consolidated financial statements.

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PACIFIC BIOSCIENCES OF CALIFORNIA, INC. 

Consolidated Statements of Stockholders’ Equity

(in thousands)

Shares

Amount

Common Stock 

Additional 
Paid-in
Capital

Accumulated
Other
Comprehensive 
Income (Loss)

Accumulated 
Deficit

Total
Stockholders'
Equity

Balance at December 31, 2020

192,294 $ 

192  $ 

1,372,083  $ 

85  $ 

(1,036,869)  $ 

335,491 

Net loss

Other comprehensive loss

Issuance of common stock in 
conjunction with equity plans

Issuance of common stock in 
Private Placement, net of 
issuance costs

Issuance of common stock in 
acquisition of Omniome

Share-based compensation 
expense

—  

—  

8,557  

11,215  

8,912  

—  

— 

— 

9 

11 

9 

— 

— 

— 

31,797 

294,834 

237,876 

73,355 

— 

(181,223) 

(181,223) 

(1,172) 

— 

— 

— 

— 

— 

— 

— 

— 

— 

(1,172) 

31,806 

294,845 

237,885 

73,355 

Balance at December 31, 2021

220,978 $ 

221  $ 

2,009,945  $ 

(1,087)  $ 

(1,218,092)  $ 

790,987 

Net loss

Other comprehensive loss

Issuance of common stock in 
conjunction with equity plans

Share-based compensation 
expense

—  

—  

5,527  

—  

— 

— 

6 

— 

— 

— 

11,224 

78,613 

— 

(314,248) 

(314,248) 

(3,678) 

— 

— 

— 

— 

— 

(3,678) 

11,230 

78,613 

Balance at December 31, 2022

226,505 $ 

227  $ 

2,099,782  $ 

(4,765)  $ 

(1,532,340)  $ 

562,904 

Net loss

Other comprehensive income

Issuance of common stock 
following milestone achievement

Issuance of common stock in 
acquisition of Apton

Issuance of common stock in 
connection with Apton liquidity 
event bonus plan

Issuance of common stock from 
Underwritten Public Equity 
Offering, net of issuance costs

Issuance of common stock in 
conjunction with equity plans

Share-based compensation 
expense

—  

—  

8,988  

6,121  

169  

20,125  

5,836  

—  

— 

— 

9 

6 

— 

20 

6 

— 

— 

— 

84,752 

76,636 

2,111 

189,180 

15,313 

72,118 

— 

(306,735) 

(306,735) 

4,984 

— 

— 

— 

— 

— 

— 

— 

— 

— 

— 

— 

— 

— 

4,984 

84,761 

76,642 

2,111 

189,200 

15,319 

72,118 

Balance at December 31, 2023

267,744 $ 

268  $ 

2,539,892  $ 

219  $ 

(1,839,075)  $ 

701,304 

See accompanying notes to the consolidated financial statements.

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PACIFIC BIOSCIENCES OF CALIFORNIA, INC.

Consolidated Statements of Cash Flows

(in thousands)
Cash flows from operating activities
Net loss
Adjustments to reconcile net loss to net cash used in operating activities

Loss (gain) from Continuation Advances 
Depreciation 
Amortization of intangible assets
Amortization of right-of-use assets
Share-based compensation 
Merger-related compensation expense
Loss on extinguishment of debt
Amortization of premium and accretion of discount on marketable securities, net
Change in the estimated fair value of contingent consideration
Inventory provision
Deferred income taxes
Other
Changes in assets and liabilities 

Accounts receivable, net
Inventory, net
Prepaid expenses and other assets 
Accounts payable 
Accrued expenses 
Deferred revenue 
Operating lease liabilities
Contingent consideration liability
Other liabilities 

Net cash used in operating activities 

Cash flows from investing activities
Purchase of property and equipment 
Purchase of intangible assets
Cash paid for purchases of acquired entities, net of cash acquired
Purchase of investments 
Sales of investments 
Maturities of investments 

Net cash provided by (used in) in investing activities 

Cash flows from financing activities
Continuation Advances
Proceeds from issuance of Convertible Senior Notes, net of issuance costs
Proceeds from issuance of common stock under equity offerings, net of issuance costs
Proceeds from issuance of common stock from equity plans
Payment of debt issuance costs
Payment of contingent consideration
Notes payable principal payoff 
Other

Net cash provided by financing activities 

Net (decrease) increase in cash, cash equivalents, and restricted cash
Cash, cash equivalents, and restricted cash at beginning of period 
Cash, cash equivalents, and restricted cash at end of period 
Cash and cash equivalents at end of period 
Restricted cash at end of period 
Cash, cash equivalents, and restricted cash at end of period 

Supplemental disclosure of cash flow information
Interest paid
Supplemental disclosure of non-cash investing and financing activities
Inventory transferred to property and equipment 

Property and equipment transferred to inventory

Right-of-use asset and liability additions and modifications

Issuance of common stock in acquisition of Apton and Omniome

Issuance of common stock in connection with Apton liquidity event bonus plan

Convertible Senior Notes exchange

Issuance of common stock following milestone achievement

$ 

$ 

$ 

$ 

$ 

$ 

$ 

$ 

$ 

$ 

Years Ended December 31,
2022

2021

2023

$ 

(306,735)  $ 

(314,248)  $ 

(181,223) 

— 
11,463 
8,261 
6,810 
72,118 
3,395 
2,033 
(12,840) 
15,060 
10,584 
(11,424) 
1,059 

(17,829) 
(13,841) 
(8,984) 
206 
13,103 
(10,420) 
(8,759) 
(14,882) 
2,449 
(259,173) 

(8,843) 
— 
(102) 
(756,567) 
595 
769,521 
4,604 

— 
9,480 
913 
6,925 
78,613 
— 
— 
(244) 
2,377 
6,027 
— 
918 

5,455 
(33,906) 
(12,324) 
1,025 
(3,651) 
(3,734) 
(7,724) 
— 
887 
(263,211) 

(16,750) 
(179) 
— 
(442,788) 
— 
575,800 
116,083 

— 
— 
189,200 
15,319 
(7,375) 
(86,411) 
(1,842) 
— 
108,891 
(145,678) 
328,311 
182,633  $ 
179,911 
2,722 
182,633  $ 

— 
— 
— 
11,230 
— 
— 
(1,608) 
— 
9,622 
(137,506) 
465,817 
328,311  $ 
325,089 
3,222 
328,311  $ 

52,000 
7,199 
381 
4,005 
73,355 
— 
— 
4,011 
1,143 
678 
(93,649) 
593 

(7,166) 
(13,109) 
(1,024) 
6,363 
15,320 
25,736 
(4,990) 
— 
(803) 
(111,180) 

(5,931) 
— 
(319,793) 
(988,046) 
212,734 
422,505 
(678,531) 

(52,000) 
895,536 
294,845 
31,806 
— 
— 
(361) 
(245) 
1,169,581 
379,870 
85,947 
465,817 
460,725 
5,092 
465,817 

15,687  $ 

14,049  $ 

6,928 

3,984  $ 

(7,022)  $ 

—  $ 

76,642  $ 

2,111  $ 

441,000  $ 

84,761  $ 

2,812  $ 

(715)  $ 

—  $ 

—  $ 

—  $ 

—  $ 

—  $ 

2,586 

(383) 

2,576 

237,885 

— 

— 

— 

See accompanying notes to the consolidated financial statements. 

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PACIFIC BIOSCIENCES OF CALIFORNIA, INC.

Notes to Consolidated Financial Statements

NOTE 1. ORGANIZATION AND SIGNIFICANT ACCOUNTING POLICIES

Business Overview

We  are  a  life  science  technology  company  that  is  designing,  developing,  and  manufacturing  advanced 
sequencing  solutions  that  enable  scientists  and  clinical  researchers  to  improve  their  understanding  of  the 
genome and ultimately, resolve genetically complex problems. Our products and technology under development 
stem from two highly differentiated core technologies focused on accuracy, quality, and completeness, which 
include  our  HiFi  long-read  sequencing  technology  and  our  Sequencing  by  Binding  (SBB®)  technology.  Our 
products  address  solutions  across  a  broad  set  of  applications  including  human  genetics,  plant  and  animal 
sciences,  infectious  disease  and  microbiology,  oncology,  and  other  emerging  applications.  Our  focus  is  on 
creating some of the world's most advanced sequencing systems to provide our customers the most complete 
and  accurate  view  of  genomes,  transcriptomes,  and  epigenomes.  Our  customers  include  academic  and 
governmental research institutions, commercial testing and service laboratories, genome centers, public health 
labs,  hospitals  and  clinical  research  institutes,  contract  research  organizations  (CROs),  pharmaceutical 
companies, and agricultural companies.

References  in  this  report  to  “PacBio,”  “we,”  “us,”  the  “Company,”  and  “our”  refer  to  Pacific  Biosciences  of 
California, Inc. and its consolidated subsidiaries.

Basis of Presentation and Consolidation

Our  consolidated  financial  statements  have  been  prepared  in  conformity  with  accounting  principles  generally 
accepted in the United States, or U.S. GAAP, as set forth in the Financial Accounting Standards Board, or FASB, 
Accounting  Standards  Codification,  or  ASC.  The  consolidated  financial  statements  include  the  accounts  of 
Pacific Biosciences and our wholly owned subsidiaries. All intercompany transactions and balances have been 
eliminated. Certain prior period amounts have been reclassified to conform to the current period presentation.

Use of Estimates 

The  preparation  of  financial  statements  in  conformity  with  GAAP  requires  us  to  make  estimates  and 
assumptions  that  affect  the  amounts  reported  in  the  financial  statements  and  accompanying  notes  to  the 
financial statements. On an ongoing basis, we evaluate our significant estimates including, but not limited to, 
the valuation of inventory, the fair value of contingent consideration, the valuation of acquired intangible assets, 
the  useful  lives  assigned  to  long-lived  assets,  the  computation  of  provisions  for  income  taxes,  and  the 
valuations  related  to  our  convertible  senior  notes.  While  the  extent  of  the  potential  impact  of  the  current 
macroeconomic conditions on our business is highly uncertain, we considered information available related to 
assumptions  and  estimates  used  to  determine  the  results  reported  and  asset  valuations  as  of  December  31, 
2023. Actual results could differ materially from these estimates. 

Functional Currency

The  U.S.  dollar  is  the  functional  currency  of  our  international  operations.  We  remeasure  foreign  subsidiaries 
monetary assets and liabilities to the U.S. dollar and record net gains or losses from remeasurement in other 
income, net, in the consolidated statements of operations and comprehensive loss.

Cash, Cash Equivalents, Restricted Cash, and Investments 

We  consider  all  highly  liquid  investments  purchased  with  an  original  maturity  of  90  days  or  less  to  be  cash 
equivalents.  Cash  equivalents  may  be  comprised  of  money  market  funds,  certificates  of  deposit,  commercial 
paper, corporate bonds and notes, and government agencies’ securities.

We classify our investments in debt securities as available-for-sale and report the investments at fair value in 
current assets. We evaluate our available-for-sale investments in unrealized loss positions and assess whether 
the  unrealized  loss  is  credit-related.  Unrealized  gains  and  losses  that  are  not  credit-related  are  recognized  in 
accumulated other comprehensive income (loss) in stockholders’ equity. Realized gains and losses, expected 
credit losses, as well as interest income, on available-for-sale securities are also reported in other income, net. 
The cost used in the determination of gains and losses of securities sold is based on the specific identification 
method.  The  cost  of  marketable  securities  is  adjusted  for  the  amortization  of  premiums  and  discounts  to 
expected maturity. Premium and discount amortization is recorded in other income, net. 

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Our  investment  portfolio  at  any  point  in  time  contains  investments  in  cash  deposits,  money  market  funds, 
commercial  paper,  corporate  debt  securities,  and  U.S.  government  and  agency  securities  with  high  credit 
ratings.  We  have  established  guidelines  regarding  diversification  and  maturities  of  investments  with  the 
objectives of maintaining safety and liquidity, while maximizing yield.

Restricted  cash  includes  cash  that  is  not  readily  available  for  use  in  the  Company’s  operating  activities. 
Restricted cash is primarily comprised of cash pledged under letters of credit.

Concentration and Other Risks

Financial instruments that potentially subject us to credit risk consist principally of interest-bearing investments 
and  trade  receivables.  We  maintain  cash,  cash  equivalents,  and  investments  with  various  major  financial 
institutions. The counterparties to the agreements relating to our investment securities consist of various major 
corporations,  financial  institutions,  municipalities,  and  government  agencies  of  high  credit  standing.  At 
December  31,  2023,  most  of  our  cash  was  deposited  with  U.S.  financial  institutions.  Our  investment  policy 
generally restricts the amount of credit exposure to any one issuer. There is no limit to the percentage of the 
portfolio  that  may  be  maintained  in  securities  issued  by  the  U.S.  Treasury  and  U.S.  Government  Agencies,  or 
other  securities  fully  backed  by  U.S.  Treasury  or  Government  agencies.  We  have  not  experienced  significant 
credit losses from financial institutions.

Our trade receivables are derived from revenue to customers and distributors located in the United States and 
other countries. We perform credit evaluations of our customers’ financial condition and, generally, require no 
collateral from our customers. The allowance for credit losses is based on our assessment of the collectability 
of  customer  accounts.  We  regularly  review  our  trade  receivables  including  consideration  of  factors  such  as 
historical  experience,  the  age  of  the  accounts  receivable  balances,  customer  creditworthiness,  customer 
industry, and current and forecasted economic conditions that may affect a customer’s ability to pay. We have 
not experienced any significant credit losses to date. 

Although  we  have  historically  not  experienced  significant  credit  losses,  our  exposure  to  credit  losses  may 
increase if our customers are adversely affected by changes in economic pressures or uncertainty associated 
with local or global economic recessions, or other customer-specific factors. 

For the year ended December 31, 2023, no single customer accounted for 10% or greater of our total revenue. 
For the years ended December 31, 2022 and 2021, one customer accounted for approximately 12%, and 13% of 
our total revenue, respectively. 

As of December 31, 2023 and 2022, 49% and 57% of our accounts receivable were from domestic customers, 
respectively.  As  of  December  31,  2023,  one  customer  represented  approximately  10%  of  our  net  accounts 
receivable.  As  of  December  31,  2022,  one  customer  represented  approximately  10%  of  our  net  accounts 
receivable. 

We  currently  purchase  several  key  parts  and  components  used  in  the  manufacture  of  our  products  from  a 
limited  number  of  suppliers.  Generally,  we  have  been  able  to  obtain  an  adequate  supply  of  such  parts  and 
components but in certain instances have incurred additional costs to secure a supply of constrained material. 
An  extended  interruption  in  the  supply  of  parts  and  components  currently  obtained  from  our  suppliers  could 
adversely affect our business and consolidated financial statements. 

Inventory, Net 

Inventories  are  stated  at  the  lower  of  cost  or  net  realizable  value  on  a  first-in,  first-out  (“FIFO”)  method. 
Adjustments  to  reduce  the  cost  of  inventory  to  its  net  realizable  value,  if  required,  are  made  for  estimated 
excess or obsolete balances. Cost includes depreciation, labor, material, and overhead costs, including product 
and process technology costs. Determining net realizable value of inventories involves numerous judgements, 
including  projecting  future  average  selling  prices,  sales  volumes,  and  costs  to  complete  products  in  work  in 
process inventories. 

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We  make  inventory  purchases  and  commitments  to  meet  future  shipment  schedules  based  on  forecasted 
demand  for  our  products.  The  business  environment  in  which  we  operate  is  subject  to  rapid  changes  in 
technology and customer demand. We perform a detailed assessment of inventory each period, which includes 
a review of, among other factors, demand requirements, product life cycle and development plans, component 
cost  trends,  product  pricing,  product  expiration,  and  quality  issues.  Based  on  our  analysis,  we  record 
adjustments  to  inventory  for  potentially  excess,  obsolete,  or  impaired  goods,  when  appropriate,  to  report 
inventory  at  net  realizable  value.  Inventory  adjustments  may  be  required  if  actual  demand,  component  costs, 
supplier arrangements, or product life cycles differ from our estimates. Any such adjustments would result in a 
charge to our results of operations.

Property and Equipment, Net 

Property  and  equipment  are  stated  at  cost,  reviewed  regularly  for  impairment,  and  depreciated  over  the 
estimated useful lives of the assets, using the straight-line method. Leasehold improvements are depreciated 
over  the  shorter  of  the  lease  term  or  the  estimated  useful  life  of  the  related  asset.  Major  improvements  are 
capitalized, while maintenance and repairs are expensed as incurred. Transfers of assets between property and 
equipment, net, and inventory are transferred at standard cost and recognized at carrying value.

Estimated useful lives of the major classes of property and equipment are as follows:

Leasehold improvements

Lab equipment

Computer equipment

Computer software

Furniture and fixtures

Operating Leases 

Estimated Useful 
Lives

3 to 10 years

3 to 5 years

3 to 5 years

3 years

3 to 5 years

We record operating lease right-of-use assets and liabilities on our consolidated balance sheets for all leases 
with a term of more than 12 months. The operating lease right-of-use assets and liabilities are calculated as the 
present  value  of  remaining  minimum  lease  payments  over  the  remaining  lease  term  using  our  estimated 
in  the 
secured 
measurement of the lease liability comprise the fixed rent per the term of the Lease. Operating lease expense is 
recognized  on  a  straight-line  basis  over  the  lease  term,  with  variable  lease  payments,  such  as  common  area 
maintenance fees, recognized in the period incurred. 

incremental  borrowing  rates  at  the  commencement  date.  Lease  payments 

included 

Business Combinations

Under the acquisition method of accounting, we allocate the fair value of the total consideration transferred to 
the  tangible  and  identifiable  intangible  assets  acquired  and  liabilities  assumed  based  on  their  estimated  fair 
values on the date of acquisition. These valuations require us to make estimates and assumptions, especially 
with respect to intangible assets. We record the excess consideration over the aggregate fair value of tangible 
and  intangible  assets,  net  of  liabilities  assumed,  as  goodwill.  Costs  that  we  incur  to  complete  the  business 
combination, such as legal and other professional fees, are expensed as they are incurred.

In connection with certain acquisitions, contingent consideration can be earned by the sellers upon completion 
of certain future performance milestones. In these cases, a liability is recorded on the acquisition date for an 
estimate of the acquisition date fair value of the contingent consideration. These estimates require significant 
management judgment, including probabilities of achieving certain future milestones. Changes in the fair value 
of the contingent consideration subsequent to the acquisition date are recognized in operating expense in our 
consolidated statements of operations and comprehensive loss.

If  the  initial  accounting  for  a  business  combination  is  incomplete  by  the  end  of  a  reporting  period  that  falls 
within  the  measurement  period,  we  report  provisional  amounts  in  our  financial  statements.  During  the 
measurement  period,  we  adjust  the  provisional  amounts  recognized  at  the  acquisition  date  to  reflect  new 
information  obtained  about  facts  and  circumstances  that  existed  as  of  the  acquisition  date  that,  if  known, 
would have affected the measurement of the amounts recognized as of that date. We record these adjustments 
to  the  provisional  amounts  with  a  corresponding  offset  to  goodwill.  Any  adjustments  identified  after  the 
measurement period are recorded in the consolidated statements of operations and comprehensive loss.

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Goodwill, Intangible Assets, and Other Long-Lived Assets

Assets  acquired,  including  intangible  assets  and  capitalized  in-process  research  and  development  (“IPR&D”), 
and liabilities assumed are measured at fair value as of the acquisition date. Goodwill, which has an indefinite 
useful life, represents the excess of cost over fair value of the net assets acquired. Intangible assets acquired in 
a business combination that are used for IPR&D activities are considered indefinite lived until the completion or 
abandonment  of  the  associated  research  and  development  efforts.  Upon  reaching  the  end  of  the  relevant 
research and development project (i.e., upon commercialization), the IPR&D asset is assessed for impairment 
and  then  amortized  over  its  estimated  useful  life.  If  the  relevant  research  and  development  project  is 
abandoned, the IPR&D asset is expensed in the period of abandonment.

Goodwill and IPR&D are not amortized; however, they are reviewed for impairment at least annually. We perform 
annual  impairment  testing  of  goodwill  in  the  second  quarter  of  each  year,  or  more  frequently  if  indicators  of 
potential  impairment  exist.  We  generally  perform  annual  impairment  testing  of  IPR&D  in  the  fourth  quarter  of 
each year, or more frequently if indicators of potential impairment exist.

We  perform  our  goodwill  impairment  analysis  at  the  reporting  unit  level.  We  have  one  reporting  unit,  which 
aligns  with  our  reporting  structure  and  availability  of  discrete  financial  information.  During  the  goodwill 
impairment  review,  we  assess  qualitative  factors  to  determine  whether  it  is  more  likely  than  not  that  the  fair 
values of our reporting unit is less than the carrying amount, including goodwill. The qualitative factors include, 
but are not limited to, macroeconomic conditions, industry and market considerations, and our overall financial 
performance. If, after assessing the totality of these qualitative factors, we determine that it is not more likely 
than not that the fair value of our reporting unit is less than the carrying amount, then no additional assessment 
is deemed necessary. Otherwise, we proceed to compare the estimated fair value of the reporting unit with the 
carrying value, including goodwill. If the carrying amount of the reporting unit exceeds the fair value, we record 
an impairment loss based on the difference. We may elect to bypass the qualitative assessment in a period and 
proceed to perform the quantitative goodwill impairment test.

During the IPR&D impairment review, we assess qualitative factors to determine whether it is more likely than 
not that the fair value of the IPR&D is less than the carrying amount. The qualitative factors include, but are not 
limited  to,  macroeconomic  conditions,  industry-specific  conditions,  and  company-specific  conditions.  If,  after 
assessing the totality of these qualitative factors, we determine that it is not more likely than not that the fair 
value  of  the  IPR&D  is  less  than  the  carrying  amount,  then  no  additional  assessment  is  deemed  necessary. 
Otherwise, we proceed to compare the estimated fair value of the IPR&D with the carrying value. If the carrying 
amount  of  the  IPR&D  exceeds  the  fair  value,  we  record  an  impairment  loss  based  on  the  difference.  We  may 
elect to bypass the qualitative assessment in a period and proceed to perform the quantitative impairment test.

Finite-lived  intangibles  assets  include  our  acquired  developed  technology  and  customer  relationships.  We 
capitalize finite-lived intangibles assets and generally amortize them on a straight-line basis over the estimated 
useful  lives.  We  regularly  review  the  carrying  amount  and  useful  lives  of  our  finite-lived  assets  to  determine 
whether indicators of impairment may exist which warrant adjustments to carrying values or estimated useful 
lives.  If  indicators  of  impairment  exist,  an  impairment  test  is  performed  to  assess  the  recoverability  of  the 
affected  assets  by  determining  whether  the  carrying  amount  of  such  assets  exceeds  the  undiscounted 
expected future cash flows. If the affected assets are not recoverable, we estimate the fair value of the assets 
and  record  an  impairment  loss  if  the  carrying  value  of  the  assets  exceeds  the  fair  value.  Factors  that  may 
indicate  potential  impairment  include  a  significant  decline  in  our  stock  price  and  market  capitalization 
compared to the net book value, significant changes in the ability of a particular asset to generate positive cash 
flows for our strategic business objectives, and the pattern of utilization of a particular asset.

Revenue Recognition 

Our revenue is generated primarily from the sale of products and services. Product revenue primarily consists of 
sales  of  our  instruments  and  related  consumables;  service  and  other  revenue  consist  primarily  of  revenue 
earned from product maintenance agreements.

We  account  for  a  contract  with  a  customer  when  there  is  a  legally  enforceable  contract  between  us  and  the 
customer, the rights of the parties are identified, the contract has commercial substance, and collectability of 
the  contract  consideration  is  probable.  Revenues  are  recognized  when  control  of  the  promised  goods  are 
transferred to our customers, or services are performed, in an amount that reflects the consideration we expect 
to be entitled to in exchange for those goods or services. 

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We  may  enter  into  contracts  with  customers  that  include  a  combination  of  promised  products  and  services, 
resulting in arrangements containing multiple performance obligations. We determine whether each product or 
service  is  distinct,  in  order  to  identify  the  performance  obligations  in  the  contract  and  allocate  the  contract 
transaction price among the distinct performance obligations. A performance obligation is considered distinct 
from other obligations in a contract when it provides a benefit to the customer either on its own or together with 
other  resources  that  are  readily  available  to  the  customer  and  is  separately  identified  in  the  contract.  We 
consider  a  performance  obligation  satisfied  once  we  have  transferred  control  of  a  good  or  service  to  the 
customer, meaning the customer has the ability to use and obtain the benefit of the good or service. Therefore, 
instrument revenue is recognized upon transfer of control of the asset to the customer, which is generally upon 
delivery for sales made to our non-distributor customers and upon shipment for sales made to our distributor 
customers. 

The  consideration  for  contracts  with  multiple  performance  obligations  is  allocated  between  separate 
performance obligations based on their individual standalone selling price. We determine the best estimate of 
standalone selling price using average selling prices over a 12-month period combined with an assessment of 
current  market  conditions.  If  the  standalone  selling  price  is  not  directly  observable,  we  rely  on  estimates  by 
considering  multiple  factors  including,  but  not  limited  to,  overall  market  conditions,  including  geographic  or 
regional  specific  factors,  internal  costs,  profit  objectives,  pricing  practices,  and  other  observable  inputs.  We 
recognize revenues as performance obligations are satisfied by transferring control of the product or service to 
the  customer  or  over  the  term  of  a  product  maintenance  agreement  with  a  customer.  Our  revenue 
arrangements generally do not provide a right of return. Revenue is recorded net of discounts and sales taxes 
collected on behalf of governmental authorities.

Certain of our agreements provide options to customers which can be exercised at a future date, such as the 
option  to  purchase  our  product  at  discounted  prices,  among  others.  In  accounting  for  customer  options,  we 
determine  whether  an  option  is  a  material  right  and  this  requires  us  to  exercise  significant  judgment.  If  a 
contract provides the customer an option to acquire additional goods or services at a discount that exceeds the 
range  of  discounts  that  we  typically  give  for  that  product  or  service  for  the  same  class  of  customer,  or  if  the 
option  provides  the  customer  certain  additional  goods  or  services  for  free,  the  option  may  be  considered  a 
material  right.  If  the  contract  gives  the  customer  the  option  to  acquire  additional  goods  or  services  at  their 
normal standalone selling prices, we would likely determine that the option is not a material right and, therefore, 
account for it when the customer exercises the option. If the standalone selling price of the option is not directly 
observable, an estimated standalone selling price is utilized which considers adjustments for discounts that the 
customer  could  receive  without  exercising  the  option  and  the  likelihood  that  the  option  will  be  exercised.  We 
may  also  utilize  the  alternative  approach  to  estimate  the  standalone  selling  price,  available  pursuant  to  the 
applicable  accounting  guidance,  to  the  extent  we  conclude  the  applicable  criteria  for  using  the  alternative 
approach has been met. We update the transaction price for expected consideration, subject to constraint, each 
reporting period if our estimate of future goods to be ordered by customers change. 

Additionally,  we  generally  provide  a  one-year  warranty  on  instruments.  We  accrue  the  cost  of  the  assurance 
warranty when revenue of the instrument is recognized. Employee sales commissions are generally recorded as 
selling,  general,  and  administrative  expense  when  incurred  as  the  amortization  period  for  such  costs,  if 
capitalized, would have been one year or less.

Cost of Revenue

Cost  of  revenue  reflects  the  direct  cost  of  product  components,  third-party  manufacturing  services,  and  our 
internal  manufacturing  overhead  and  customer  service  infrastructure  costs  incurred  to  produce,  deliver, 
maintain, and support our instruments, consumables, and services.

Manufacturing  overhead  is  predominantly  comprised  of  labor  and  facility  costs.  We  capitalize  manufacturing 
overhead into inventory based on a standard cost model that approximates actual costs. 

Service  costs  include  the  direct  costs  of  components  used  in  support,  repair  and  maintenance  of  customer 
instruments  as  well  as  the  cost  of  personnel,  materials,  shipping  and  support  infrastructure  necessary  to 
support our installed customer base. 

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Research and Development 

Research and development expense consists primarily of expenses for personnel engaged in the development 
of our core technology, the design and development of our future products and current product enhancements. 
These  expenses  also  include  prototype-related  expenditures,  development  equipment  and  supplies,  partner 
development costs, facilities costs, and other related overhead. We expense research and development costs 
during  the  period  in  which  the  costs  are  incurred.  We  defer  and  capitalize  non-refundable  advance  payments 
made  for  research  and  development  activities  until  the  related  goods  are  received  or  the  related  services  are 
rendered.

Credit Losses

Trade accounts receivable

The  allowance  for  credit  losses  is  based  on  our  assessment  of  the  collectability  of  customer  accounts.  We 
regularly  review  the  allowance  by  considering  factors  such  as  the  age  of  the  accounts  receivable  balances, 
customer creditworthiness, customer industry, and current and forecasted economic conditions that may affect 
a customer’s ability to pay. Credit loss expense was immaterial for the years ended December 31, 2023, 2022, 
and 2021.

Available-for-sale debt securities

Our  investment  portfolio  contains  investments  in  cash  deposits,  money  market  funds,  commercial  paper, 
corporate  debt  securities  and  U.S.  government  and  agency  securities.  We  regularly  assess  whether  our 
securities in an unrealized loss position are credit related. The credit-related portion of unrealized losses, and 
any subsequent improvements, are recorded in interest income. Unrealized losses that are not credit related are 
included  in  accumulated  other  comprehensive  income  (loss).  The  unrealized  losses  on  our  investments  are 
mainly  attributable  to  government  securities,  including  U.S.  government  and  U.S.  agency  bond  securities, 
impacted by movements in market rates and not due to issuer credit risk. We have the ability to hold and do not 
intend to sell the investments in unrealized loss positions before the recovery of their amortized cost bases.

Although  we  have  historically  not  experienced  significant  credit  losses,  our  exposure  to  credit  losses  may 
increase if our customers are adversely affected by changes in economic pressures or uncertainty associated 
with  local  or  global  economic  recessions,  disruptions  associated  with  the  evolution  of  COVID-19  or  other 
epidemics or pandemics, or other customer-specific factors. 

Income Taxes 

We  account  for  income  taxes  under  the  asset  and  liability  method,  which  requires,  among  other  things,  that 
deferred income taxes be provided for temporary differences between the tax bases of our assets and liabilities 
and  the  amounts  reported  in  the  financial  statements.  In  addition,  deferred  tax  assets  are  recorded  for  the 
future benefit of utilizing net operating losses and research and development credit carryforwards. The effect of 
a change in tax rates on the deferred tax assets and liabilities is recognized in the provision for income taxes in 
the period that includes the enactment date. A full valuation allowance is provided against our net deferred tax 
assets as it is more likely than not that the deferred tax assets will not be fully realized. 

We regularly review our positions taken relative to income taxes. To the extent our tax positions are more likely 
than  not  going  to  result  in  additional  taxes,  we  accrue  the  estimated  amount  of  tax  related  to  such  uncertain 
positions. 

Share-based Compensation 

We  recognize  share-based  compensation  expense  for  share-based  payments,  including  stock  options, 
restricted  stock  units,  performance  stock  units  and  stock  issued  under  our  employee  stock  purchase  plan 
("ESPP")    based  on  the  grant-date  fair  value.  We  estimate  the  fair  value  of  stock  options  and  ESPP  using  an 
option-pricing  model.  See  Note  10.  Stockholders’  Equity  for  further  information  regarding  share-based 
compensation.

Other Comprehensive Income (Loss)

Other comprehensive income (loss) is comprised of unrealized gains (losses) on our investment securities. 

Shipping and Handling

Costs related to shipping and handling are included in cost of revenues for all periods presented.

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Earnings per Share

Basic  net  loss  per  share  is  computed  by  dividing  net  loss  by  the  weighted-average  number  of  shares  of 
common  stock  outstanding  during  the  period.  Diluted  net  loss  per  share  is  computed  using  the  weighted-
average number of shares of common stock outstanding and potential shares assuming the dilutive effect of 
outstanding stock options, restricted stock units, and common stock issuable pursuant to our ESPP, using the 
treasury stock method.

Recent Accounting Pronouncements

Recently Adopted Accounting Standards

In  October  2021,  the  FASB  issued  Accounting  Standards  Update  ("ASU")  No.  2021-08,  Business  Combinations 
(Topic  805):  Accounting  for  Contract  Assets  and  Contract  Liabilities  from  Contracts  with  Customers.  This  ASU 
provides  specific  guidance  on  how  to  recognize  contract  assets  and  contract  liabilities  related  to  revenue 
contracts  with  customers  acquired  in  a  business  combination.  This  amendment  improves  comparability  for 
both the recognition and measurement of acquired revenue contracts with customers at the date of and after a 
business combination. We adopted this ASU on January 1, 2023. The adoption of this guidance did not have a 
material effect on our consolidated financial statements.

Accounting Pronouncements Pending Adoption

In  December  2023,  the  FASB  issued  ASU  2023-09,  Income  Taxes  (Topic  740):  Improvements  to  Income  Tax 
Disclosures. This ASU requires entities to expand its existing income tax disclosures, specifically related to the 
rate reconciliation and income taxes paid. This authoritative guidance will be effective for us in fiscal year 2025, 
with early adoption permitted. The Company is currently evaluating the impact of the ASU but does not expect 
any material impacts upon adoption.

NOTE 2. BUSINESS ACQUISITIONS

Apton Biosystems

On  August  2,  2023,  we  acquired  Apton  Biosystems,  Inc.  (“Apton”),  a  California-based  genomics  company 
focused  on  developing  a  high  throughput  short-read  sequencer  using  highly  differentiated  optics  and  image 
processing, paired with novel clustering and chemistry (the “Apton acquisition”). 

In connection with the Apton acquisition, all outstanding equity securities of Apton were cancelled in exchange 
for  shares  of  our  common  stock  with  a  fair  value  of  $76.6  million,  cash  of  $0.2  million,  and  contingent 
consideration with a preliminary estimated fair value of $18.5 million. Excluded from consideration transferred 
was $1.3 million attributable to accelerated share-based compensation expense. The fair value of the 6,121,571 
common  shares  issued  was  determined  based  on  the  closing  market  price  of  our  common  stock  on  the 
acquisition date.

In connection with the Apton acquisition, contingent consideration of $25.0 million, which we may elect to pay 
in cash, shares of our common stock or a combination of cash and shares of our common stock, is due upon 
the achievement of a milestone, defined as the achievement of $50.0 million in revenue associated with Apton's 
technology, provided that the milestone event occurs prior to the 5-year anniversary of the closing date of the 
acquisition. At this time, the number of shares, if any, to be issued in connection with the achievement of the 
specified  milestone  is  not  known  and  will  be  calculated  based  on  the  daily  volume-weighted  average  price  of 
our common stock for the twenty trading days ending on and including the fifth trading day immediately prior to 
the occurrence of the specified milestone. Upon achievement of the milestone, we may pay cash in lieu of our 
common  stock  to  ensure  that  the  issuance  of  our  common  stock  does  not  exceed  19.9%  of  our  outstanding 
shares of common stock then outstanding.

The  contingent  consideration  is  accounted  for  as  a  liability  at  fair  value,  with  changes  during  each  reporting 
period recognized in our consolidated statements of operations and comprehensive loss. The fair value of the 
contingent  consideration  liability  is  calculated,  with  the  assistance  from  a  third-party  valuation  firm,  using  a 
Monte  Carlo  Simulation  to  estimate  the  volatility  and  systematic  relative  risk  of  revenues  subject  to  sales 
milestone  payments  and  discounting  the  associated  cash  payment  amounts  to  their  present  values  using  a 
credit-risk-adjusted interest rate.

We allocated the consideration transferred to the identifiable assets acquired and liabilities assumed based on 
preliminary estimates of their respective fair values at the date of the completion of the Apton acquisition, and 
such  allocation  is  subject  to  adjustment  for  up  to  one  year  after  the  close  of  the  acquisition  as  additional 

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information  is  obtained.  The  major  classes  of  assets  and  liabilities  to  which  we  have  allocated  the  total  fair 
value  of  the  consideration  transferred,  based  on  the  preliminary  estimated  fair  values  were  as  follows  (in 
thousands):

Cash and cash equivalents

In-process research and development

Goodwill

Other assets, current

Deferred income tax liability

Liabilities assumed

Total consideration transferred

$ 

97 

55,000 

52,287 

153 

(11,338) 

(2,191) 

$ 

94,008 

The  purchase  price  allocation  is  preliminary,  primarily  due  to  the  pending  finalization  of  review  of  various  tax 
attributes. We continue to collect information regarding certain estimates and assumptions, including potential 
liabilities  and  contingencies.  We  will  record  adjustments  to  the  fair  value  of  the  assets  acquired,  liabilities 
assumed  and  goodwill  within  the  twelve  months  measurement  period,  if  necessary.  During  the  year  ended 
December 31, 2023, we recorded a measurement period adjustment of $1.6 million to decrease goodwill, and a 
corresponding $2.0 million increase in intangible assets and $0.4 million decrease in the deferred tax liability on 
the  consolidated  balance  sheets,  and  a  $0.7  million  increase  to  our  benefit  from  income  taxes  on  the 
consolidated statements of operations and comprehensive loss. The measurement period adjustment was due 
to  new  information  that  became  available  to  us  upon  the  completion  of  the  valuation  assessment  of  the  in-
process research and development and the tax provision.

We  incurred  costs  related  to  the  Apton  acquisition  of  approximately  $9.0  million  during  the  year  ended 
December  31,  2023,  which  are  included  in  merger-related  expenses  on  the  consolidated  statement  of 
operations  and  comprehensive  loss.  Merger-related  expenses  include  $2.8  million  relating  to  a  liquidity  event 
bonus plan that was treated as a separate transaction and included the issuance of 168,621 shares of common 
stock that were issued with a fair value of $2.1 million based on the closing market price of our common stock 
on  the  acquisition  date.  As  a  result,  the  total  shares  issued  in  connection  with  the  Apton  acquisition  were 
6.3 million shares of common stock. 

The  excess  of  the  value  of  consideration  paid  over  the  aggregate  fair  value  of  those  net  assets  has  been 
recorded  as  goodwill.  We  recognized  goodwill  of  $52.3  million,  based  on  preliminary  estimates,  which  is 
primarily attributable to the synergies expected to occur from the integration of Apton and is not deductible for 
income  tax  purposes.  We  preliminarily  allocated  $55.0  million  of  the  purchase  price  to  acquired  in-process 
research and development ("IPR&D"). The fair value of the IPR&D was determined, with the assistance of a third-
party  valuation  firm,  using  an  income  approach  based  on  a  forecast  of  expected  future  cash  flows.  Expected 
future cash flows utilize significant assumptions such as assumed revenue projections and discount rate. 

Omniome, Inc.

On  September  20,  2021,  we  completed  our  acquisition  of  Omniome,  Inc.  (“Omniome”),  a  San  Diego-based 
company,  to  obtain  their  proprietary  short-read  DNA  sequencing  platform  capable  of  delivering  high  accuracy 
(the “Omniome acquisition”). 

In  connection  with  the  Omniome  acquisition,  all  outstanding  equity  securities  of  Omniome  were  cancelled  in 
exchange for approximately $315.7 million in cash, 8,911,580 shares of our common stock with a fair value of 
$249.4 million and contingent consideration with a fair value of $168.6 million. The fair value of the 8,911,580 
common shares issued was determined based on the closing market price of PacBio’s common shares on the 
acquisition date.

In  addition,  approximately  $18.9  million,  comprised  of  $7.4  million  of  cash,  226,811  shares  of  our  common 
stock with a fair value of $6.3 million, and $5.2 million related to contingent consideration, was accounted for as 
a one-time post-acquisition share-based compensation expense. This share-based compensation expense was 
due to accelerated vesting of Omniome stock awards in connection with the acquisition.

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In  connection  with  the  acquisition  the  contingent  consideration  of  $200  million  (composed  of  $100  million  in 
cash and $100 million in shares of our common stock) is due upon the achievement of a milestone, defined as 
the  first  commercial  shipment  to  a  customer  of  a  nucleotide  sequencing  platform,  comprising  both  an 
instrument and related consumables, that utilizes SBB technology. The number of shares of stock to be issued 
will be determined using the volume-weighted average of the trading prices of our common stock for the twenty 
trading days ending with and including the trading day that is two days immediately prior to the achievement of 
the  milestone.  Of  the  $100  million  in  shares  of  our  common  stock  to  be  issued  as  part  of  the  milestone, 
$4.1 million was attributable to stock options issued by PacBio in replacement of Omniome’s unvested options 
as part of the transaction. Upon achievement of the milestone, shares will be issued not in excess of an amount 
equal to 19.9% of our outstanding shares of common stock on the date of closing (prior to the issuance of any 
shares issued in connection with the transaction or the related private placement), less 11,500,000 shares.

The  contingent  consideration  is  accounted  for  as  a  liability  at  fair  value,  with  changes  during  each  reporting 
period recognized in our Consolidated Statements of Operations and Comprehensive Loss. The fair value of the 
contingent  consideration  liability  is  calculated,  with  the  assistance  from  a  third-party  valuation  firm,  using  a 
scenario-based  method  that  considers  a  range  of  possible  outcomes  and  their  assigned  probabilities  of 
occurrence. The potential outcomes are discounted to present value at a discount rate equal to the sum of the 
term-matched  risk-free-interest  rate  plus  PacBio’s  credit  spread.  On  September  20,  2023,  we  achieved  the 
commercial  milestone  in  connection  with  the  acquisition  of  Omniome.  See  Note  5.  Financial  Instruments  for 
additional information on amounts paid and shares issued to former Omniome securityholders during the year 
ended December 31, 2023.

Total consideration transferred for the acquisition is as follows (in thousands):

Total cash paid

Fair value of share consideration

Fair value of contingent consideration

Less: Share-based compensation expense excluded from consideration transferred

Total consideration transferred

$ 

315,703 

249,435 

168,574 

(18,923) 

$ 

714,789 

The  acquisition  was  accounted  for  as  a  business  combination  and,  accordingly,  the  total  fair  value  of  the 
consideration transferred was allocated to the tangible and intangible assets acquired and liabilities assumed 
based on their fair values on the acquisition date. As of December 31, 2021, the major classes of assets and 
liabilities  to  which  we  have  allocated  the  total  fair  value  of  the  consideration  transferred  were  as  follows  (in 
thousands):

Cash and cash equivalents

Property and equipment, net

Operating lease right-of-use assets, net

In-process research and development

Goodwill

Other assets, non-current

Deferred income tax liability

Liabilities assumed

Total consideration transferred

$ 

15,338 

6,123 

18,095 

400,000 

390,665 

3,203 

(91,814) 

(26,821) 

$ 

714,789 

During  the  year  ended  December  31,  2021,  we  recorded  a  measurement  period  adjustment  of  $1.6  million  to 
decrease goodwill and a corresponding $0.4 million to decrease the deferred tax liability on the Consolidated 
Balance Sheet, and a $1.2 million decrease to our benefit from income taxes on the Consolidated Statements of 
Operations  and  Comprehensive  (Loss)  Income.  The  measurement  period  adjustment  was  due  to  new 
information  that  became  available  to  us  upon  the  completion  of  the  IRC  Section  382  Tax  Study,  where  we 
identified  additional  net  operating  losses  that  are  available  to  us  from  acquired  assets.  Refer  to  Note  9  – 
Income Taxes, in Part II, Item 8 of the Annual Report on Form 10-K for the year ended December 31, 2021 for 
more information. There were no measurement period adjustments recorded in the year ended December 31, 
2022.

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The goodwill recognized was primarily attributable to the assembled workforce and synergies that are expected 
to occur from the integration of Omniome and is not deductible for income tax purposes. 

We incurred costs related to the Omniome acquisition of approximately $12.0 million during the twelve months 
ended  December  31,  2021,  which  are  included  in  merger-related  costs  on  the  Consolidated  Statement  of 
Operations  and  Comprehensive  (Loss)  Income.  No  significant  merger-related  costs  were  incurred  during  the 
twelve months ended December 31, 2022.

The  following  unaudited  pro  forma  financial  information  presents  combined  results  of  operations  for  each  of 
the  periods  presented  as  if  Omniome  had  been  acquired  as  of  the  beginning  of  2020,  giving  effect  on  a  pro 
forma basis to the purchase accounting adjustments such as $12.0 million of PacBio acquisition-related costs, 
$18.9 million of share-based compensation expense related to acceleration of certain Omniome stock options 
not attributable to pre-combination service, and a $91.0 million one-time income tax benefit from the reduction 
of our deferred tax asset valuation allowance resulting from the Omniome acquisition, as well as a pro forma 
adjustment to reflect $16.7 million of Omniome’s acquisition-related costs. 

The unaudited pro forma information presented below is for informational purposes only and is not necessarily 
indicative  of  the  consolidated  results  of  the  combined  business  had  the  acquisition  actually  occurred  at  the 
beginning of 2020 or the results of future operations of the combined business.

The following table summarizes the unaudited pro forma financial information:

(in thousands, except per share amounts)

Pro forma total revenue

Pro forma net (loss) income

Pro forma net (loss) income per share - basic and diluted

Years Ended December 31,

2021

2020

$ 

$ 

$ 

130,513  $ 

(278,451)  $ 

(1.27)  $ 

78,893 

17,510 

0.09 

Our consolidated financial statements include the results of operations for Omniome beginning September 20, 
2021. Revenues of $0 and a net loss of $15.6 million from the acquired Omniome business have been included 
in our Consolidated Statement of Operations and Comprehensive (Loss) Income for the twelve months ended 
December 31, 2021.

Circulomics, Inc.

On  July  20,  2021,  we  acquired  Circulomics  Inc.  (“Circulomics”),  a  Maryland-based  biotechnology  company 
focused  on  delivering  highly  differentiated  sample  preparation  products  that  enable  genomic  workflows  (the 
“Circulomics acquisition”). 

We  paid  $29.5  million  in  cash  in  exchange  for  all  outstanding  shares  of  common  stock  of  Circulomics.  We 
allocated the consideration transferred to the identifiable assets acquired and liabilities assumed based on their 
respective fair values at the date of the completion of the Circulomics acquisition. The major classes of assets 
and liabilities to which we have allocated the total fair value of the consideration transferred were as follows (in 
thousands):

Cash and cash equivalents

Property and equipment, net

Intangible assets

Goodwill

Other assets, non-current

Deferred income tax liability

Liabilities assumed

Total consideration transferred

$ 

987 

214 

11,360 

19,309 

467 

(2,672) 

(118) 

$ 

29,547 

The  excess  of  the  value  of  consideration  paid  over  the  aggregate  fair  value  of  those  net  assets  has  been 
recorded as goodwill. We recognized goodwill of $19.3 million, which is primarily attributable to the synergies 
expected from capabilities in extraction and sample preparation and is not deductible for income tax purposes. 
We  recorded  $11.4  million  for  the  fair  value  of  acquired  intangible  assets,  which  consists  of  developed 
technology and customer relationships.

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NOTE 3. INVITAE COLLABORATION

On June 24, 2022, we entered into an Amended and Restated Development and Commercialization Agreement 
(the  “Amended  and  Restated  Agreement”)  with  Invitae  Corporation  (“Invitae”).  The  Amended  and  Restated 
Agreement amended and restated the existing Development and Commercialization Agreement, effective as of 
January  12,  2021,  as  amended  by  Amendment  No.  1  to  Development  and  Commercialization  Agreement, 
entered  into  on  June  3,  2021,  by  and  between  us  and  Invitae  (together,  the  “Original  Agreement”).  Unless 
otherwise agreed in writing or terminated in accordance with the Amended and Restated Agreement, the term 
of the Amended and Restated Agreement shall continue until June 30, 2028 (“Term”). 

Pursuant  to  the  Original  Agreement,  Invitae  provided  certain  funding  to  us  to  develop  products  relating  to 
production-scale  high-throughput  sequencing  (“Program  Products”).  If  Program  Products  were  to  become 
commercially available, Invitae had the right to purchase the Program Products at preferred pricing.

Under  the  Amended  and  Restated  Agreement,  we  will  continue  to  receive  feedback,  input  and  insight  from 
Invitae in connection with the intended development of our new sequencing systems; however, such feedback 
will not be contractually required, and Invitae has no contractual right to participate in decisions regarding the 
development  program  for  such  new  sequencing  systems.  Our  development  plans  for  such  new  sequencing 
systems will be at our discretion and pursuant to our own internal processes and programs. Invitae will not be 
contractually  obligated  to  reimburse  us  for  development  costs  under  the  Amended  and  Restated  Agreement. 
There  can  be  no  assurances  that  the  in-development  sequencing  systems  will  continue  to  be  developed,  be 
successfully developed or become available for commercial sale.

In  consideration  of  the  non-refundable  payments  received  from  Invitae  pursuant  to  the  Original  Agreement  of 
$23.5  million,  we  will  provide  Invitae  with  credits  in  connection  with  Invitae’s  anticipated  purchase  of  certain 
currently  sequencing  systems  (instruments,  consumables  and  service  contracts).  The  credits  will  expire  on 
June  30,  2025  (“Credit  Expiration  Date”).  Subject  to  certain  conditions,  Invitae  will  also  be  entitled  to  most 
favored pricing for the Company’s Sequel IIe systems and certain in-development systems through the Term. 

We  and  Invitae  may  terminate  the  Amended  and  Restated  Agreement  if  the  other  party  remains  in  material 
breach of the Amended and Restated Agreement following a cure period to remedy the material breach.

The  Amended  and  Restated  Agreement  was  deemed  a  contract  modification  and  accounted  for  on  a 
prospective  basis  in  accordance  with  ASC  Topic  606.  We  will  recognize  proportionate  amounts  of  the 
transaction price, including payments made by Invitae to us pursuant to the Original Agreement, in revenue as 
the remaining performance obligations are satisfied, which is when Invitae places purchase orders for certain 
sequencing platforms and the associated goods or services are delivered. Any remaining unused credits will be 
recognized when they expire.

Invitae  purchased  certain  instruments  and  consumables  under  the  terms  of  the  Amended  and  Restated 
Agreement, for which $10.5 million and $3.7 million of revenue was recognized as product revenue during the 
years  ended  December  31,  2023  and  2022,  respectively,  on  the  consolidated  statements  of  operations  and 
comprehensive loss.

As of December 31, 2023, $8.0 million of deferred revenue, current, and $2.9 million of deferred revenue, non-
current, is recorded on the consolidated balance sheet relating to all future performance obligations under the 
Amended and Restated Agreement.

NOTE 4. TERMINATION OF MERGER WITH ILLUMINA

On  November  1,  2018,  we  entered  into  an  Agreement  and  Plan  of  Merger  (as  amended,  the  “Illumina  Merger 
Agreement”) with Illumina, Inc. (“Illumina”) and FC Ops Corp., a wholly owned subsidiary of Illumina (“Illumina 
Merger Sub”). On January 2, 2020, we, Illumina and Illumina Merger Sub, entered into an agreement to terminate 
the Merger Agreement (the “Termination Agreement”). 

Continuation Advances from Illumina

As part of the Termination Agreement, Illumina paid us cash payments (“Continuation Advances”) totaling $52 
million.  Up  to  the  full  $52.0  million  of  Continuation  Advances  paid  to  us  were  repayable  without  interest  to 
Illumina  if,  within  two  years  of  March  31,  2020,  we  entered  into,  or  consummated  a  Change  of  Control 
Transaction or raised at least $100 million in a single equity or debt financing (that may have multiple closings), 
with the amount repayable dependent on the amount raised by us. 

Resulting from the issuance and sale of $900 million of 1.50% Convertible Senior Notes due February 15, 2028, 
$52.0 million of Continuation Advances were paid without interest to Illumina in February 2021 and recorded a 

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non-operating  expense  in  the  consolidated  statements  of  operations  and  comprehensive  loss  for  the  year 
ended December 31, 2021.

NOTE 5. FINANCIAL INSTRUMENTS

Fair Value of Financial Instruments

Fair value is the exchange price that would be received to sell an asset or paid to transfer a liability in an orderly 
transaction between market participants at the measurement date. 

The  fair  value  hierarchy  established  under  GAAP  requires  an  entity  to  maximize  the  use  of  observable  inputs 
and minimize the use of unobservable inputs when measuring fair value. The three levels of inputs that may be 
used to measure fair value are as follows: 

•

•

•

Level 1: quoted prices in active markets for identical assets or liabilities;

Level 2: inputs other than Level 1 that are observable, either directly or indirectly, such as quoted prices 
in active markets for similar assets or liabilities, quoted prices for identical or similar assets or liabilities 
in markets that are not active, or other inputs that are observable or can be corroborated by observable 
market data for substantially the full term of the assets or liabilities; and

Level 3: unobservable inputs that are supported by little or no market activity and that are significant to 
the fair value of the assets or liabilities.

We  consider  an  active  market  as  one  in  which  transactions  for  the  asset  or  liability  occurs  with  sufficient 
frequency  and  volume  to  provide  pricing  information  on  an  ongoing  basis.  Conversely,  we  view  an  inactive 
market as one in which there are few transactions for the asset or liability, the prices are not current, or price 
quotations  vary  substantially  either  over  time  or  among  market  makers.  Where  appropriate,  our  non-
performance  risk,  or  that  of  our  counterparty,  is  considered  in  determining  the  fair  values  of  liabilities  and 
assets, respectively. 

We classify our cash deposits and money market funds within Level 1 of the fair value hierarchy because they 
are valued using bank balances or quoted market prices. We classify our investments as Level 2 instruments 
based on market pricing and other observable inputs. We did not classify any of our investments within Level 3 
of the fair value hierarchy.

Assets and liabilities measured at fair value are classified in their entirety based on the lowest level input that is 
significant to the fair value measurement. Our assessment of the significance of a particular input to the entire 
fair value measurement requires management to make judgments and consider factors specific to the asset or 
liability. 

The  carrying  amount  of  our  accounts  receivable,  prepaid  expenses,  other  current  assets,  accounts  payable, 
accrued expenses and other liabilities, current, approximate fair value due to their short maturities. 

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U.S. government & agency 
securities

Total cash and cash 
equivalents 

Investments:

Commercial paper 

Corporate debt securities 

U.S. government & agency 
securities

Total investments 

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Assets and Liabilities Measured at Fair Value on a Recurring Basis

The  following  table  sets  forth  the  fair  value  of  our  financial  assets  and  liabilities  that  were  measured  on  a 
recurring basis as of December 31, 2023 and December 31, 2022, respectively:

(in thousands)

Assets

Cash and cash 
equivalents:

Cash and money market 
funds 

December 31, 2023

December 31, 2022

Level 1

Level 2

Level 3

Total

Level 1

Level 2

Level 3

Total

$  70,172  $ 

—  $ 

—  $  70,172  $ 137,636  $ 

—  $ 

—  $  137,636 

Commercial paper 

— 

— 

— 

— 

— 

  166,453 

— 

  166,453 

— 

  109,739 

— 

  109,739 

— 

  21,000 

— 

21,000 

  70,172 

  109,739 

— 

  179,911 

  137,636 

  187,453 

— 

  325,089 

— 

— 

— 

— 

9,947 

  88,579 

  352,979 

  451,505 

— 

— 

— 

— 

— 

— 

9,947 

  88,579 

  352,979 

  451,505 

— 

— 

— 

— 

  127,302 

  49,491 

  270,436 

  447,229 

300 

2,422 

300 

2,922 

— 

— 

— 

— 

— 

— 

— 

— 

  127,302 

49,491 

  270,436 

  447,229 

300 

2,922 

Short-term restricted cash  

300 

Long-term restricted cash  

2,422 

— 

— 

Total assets measured at 
fair value 

$  72,894  $ 561,244  $ 

—  $ 634,138  $ 140,858  $ 634,682  $ 

—  $  775,540 

Liabilities

Contingent consideration 
- Omniome acquisition

Contingent consideration 
- Apton acquisition

Total liabilities measured 
at fair value 

$ 

$ 

$ 

—  $ 

—  $ 

—  $ 

—  $ 

—  $ 

—  $ 172,094  $  172,094 

—  $ 

—  $  19,550  $  19,550  $ 

—  $ 

—  $ 

—  $ 

— 

—  $ 

—  $  19,550  $  19,550  $ 

—  $ 

—  $ 172,094  $  172,094 

We  classify  contingent  consideration,  which  was  incurred  in  connection  with  the  acquisition  of  Apton,  within 
Level 3 as factors used to develop the estimate of fair value include unobservable inputs that are not supported 
by  market  activity  and  are  significant  to  the  fair  value.  Estimates  and  assumptions  used  in  the  Monte  Carlo 
simulation include risk-adjusted forecasted revenues for products and services leveraging Apton's technology 
and an estimated credit spread. 

We  estimate  the  fair  value  of  the  contingent  consideration  liability  based  on  the  simulated  revenue  of  the 
Company through the 5-year anniversary of the closing date of the acquisition. As of December 31, 2023, the 
key input used in the determination of the fair value included projected revenues of the Company relating to the 
high-throughput  short-read  products  and  services  leveraging  Apton's  technology.  A  decrease  in  the  projected 
revenues would result in a decrease in the fair value of the liability. The discount rates used are the sum of the 
U.S. risk-free rate and the estimated subordinated credit spread for B- credit rating, which ranges from 7.8% to 
8.2%.  Changes  in  our  estimated  subordinated  credit  spread  can  result  in  changes  in  the  fair  value  of  the 
contingent consideration liability, where a lower credit spread may result in an increased liability valuation.

On September 20, 2023, we achieved the commercial milestone in connection with the acquisition of Omniome. 
Consequently,  former  Omniome  securityholders  were  entitled  to  receive  as  milestone  consideration,  among 
other things, an aggregate of approximately $100.9 million in cash and approximately 9.0 million shares of our 
common stock, representing $95.9 million divided by the volume-weighted average of the trading prices of our 
common  stock  for  the  twenty  trading  days  ending  with  and  including  the  trading  day  that  was  two  days 

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immediately prior to the achievement of the milestone. The $95.9 million represents the $100.0 million that was 
to be paid in shares of our common stock offset by $4.1 million attributable to stock options issued by PacBio 
in  replacement  of  Omniome’s  unvested  options  as  part  of  the  transaction,  pursuant  to  the  terms  of  the 
Omniome merger agreement. 

Following the achievement of the commercial milestone, $101.3 million of the contingent consideration, which 
includes  certain  payroll  taxes,  was  paid  during  the  year  ended  December  31,  2023.  Additionally,  8,988,391 
shares were issued at a value of $84.8 million to the former Omniome securityholders. 

As  a  result  of  the  achievement  of  the  milestone,  the  contingent  consideration  liability  incurred  in  connection 
with the acquisition of Omniome was no longer considered a Level 3 liability at December 31, 2023. There were 
no transfers between Level 1, Level 2, or Level 3 assets or liabilities reported at fair value on a recurring basis 
for the year ended December 31, 2023, and our valuation techniques did not change compared to the prior year. 

Changes  in  the  estimated  fair  value  of  the  contingent  consideration  liability  for  the  year  ended  December  31, 
2023 were as follows:

(in thousands)

Beginning balance as of December 31, 2022

Additions

Change in estimated fair value

Achievement of milestone

Ending balance as of December 31, 2023

Level 3

$ 

172,094 

18,450 

15,060 

(186,054) 

$ 

19,550 

Changes  to  the  fair  value  are  recorded  as  the  Change  in  fair  value  of  contingent  consideration  in  the 
consolidated statement of operations and comprehensive loss.

For the year ended December 31, 2023, there were no transfers between Level 1, Level 2, or Level 3 assets or 
liabilities reported at fair value on a recurring basis and our valuation techniques did not change compared to 
the prior year.

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Cash, Cash Equivalents, Restricted Cash, and Investments

The following table summarizes our cash, cash equivalents, restricted cash, and investments:

(in thousands)

Cash and cash equivalents:

Cash and money market funds

U.S. government & agency securities

Total cash and cash equivalents 

Investments:

Commercial paper 

Corporate debt securities 

U.S. government & agency securities

Total investments 

Total cash, cash equivalents, and investments 

Short-term restricted cash

Long-term restricted cash

(in thousands)

Cash and cash equivalents:

Cash and money market funds

Commercial paper 

U.S. government & agency securities

Total cash and cash equivalents 

Investments:

Commercial paper 

Corporate debt securities 

U.S. government & agency securities

Total investments 

Total cash, cash equivalents, and investments 

Short-term restricted cash

Long-term restricted cash

December 31, 2023

Gross
unrealized
gains 

Gross
unrealized
losses 

Fair
Value 

Amortized
Cost 

$ 

70,172  $ 

—  $ 

—  $ 

70,172 

109,786 

179,958 

9,947 

88,263 

353,029 

451,239 

13 

13 

— 

373 

478 

851 

(60)   

(60)   

109,739 

179,911 

— 

(57)   

(528)   

(585)   

9,947 

88,579 

352,979 

451,505 

631,197  $ 

864  $ 

(645)  $ 

631,416 

300  $ 

2,422  $ 

—  $ 

—  $ 

—  $ 

—  $ 

300 

2,422 

$ 

$ 

$ 

December 31, 2022

Gross
unrealized
gains

Gross
unrealized
losses

Fair
Value

Amortized
Cost

$ 

137,636  $ 

—  $ 

—  $ 

137,636 

166,514 

20,994 

325,144 

127,626 

49,998 

274,315 

451,939 

— 

6 

6 

9 

— 

1 

10 

(61)   

166,453 

— 

21,000 

(61)   

325,089 

(333)   

(507)   

(3,880)   

(4,720)   

127,302 

49,491 

270,436 

447,229 

$ 

$ 

$ 

777,083  $ 

16  $ 

(4,781)  $ 

772,318 

300  $ 

2,922  $ 

—  $ 

—  $ 

—  $ 

—  $ 

300 

2,922 

The  following  table  summarizes  the  contractual  maturities  of  our  cash  equivalents  and  available-for-sale 
investments, excluding money market funds, as of December 31, 2023:

(in thousands)

Due in one year or less 

Due after one year through 5 years 

Total investments 

Fair Value

$ 

$ 

465,181 

96,063 

561,244 

Actual maturities may differ from contractual maturities because issuers may have the right to call or prepay 
obligations without call or prepayment penalties.

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Investment  income  included  in  other  income,  net  on  the  consolidated  statement  of  operations  and 
comprehensive  loss  was  $32.8  million  and  $9.2  million  for  the  years  ended  December  31,  2023  and  2022, 
respectively.

NOTE 6. BALANCE SHEET COMPONENTS 

Inventory, Net

Inventory, net, consisted of the following components:

(in thousands)

Purchased materials

Work in process

Finished goods

Inventory, net

Property and Equipment, Net

Property and equipment, net, consisted of the following components: 

(in thousands)

Laboratory equipment and machinery 

Leasehold improvements

Computer equipment

Software

Furniture and fixtures

Construction in progress

Total

Less: Accumulated depreciation 

Property and equipment, net 

December 31,

2023

2022

$ 

20,168  $ 

23,436 

13,072 

$ 

56,676  $ 

24,139 

14,062 

12,180 

50,381 

December 31,

2023

2022

$ 

44,907  $ 

35,226 

19,528 

6,628 

3,594 

1,343 

38,998 

34,129 

18,438 

6,879 

3,426 

4,698 

111,226 

106,568 

(74,794)   

(64,988) 

$ 

36,432  $ 

41,580 

Construction  in  progress  consists  of  capitalizable  costs  that  have  been  incurred  for  the  construction  of  long-
lived assets and is primarily comprised of amounts that will be classified as lab equipment. 

Depreciation  expense  during  the  years  ended  December  31,  2023,  2022,  and  2021  was  $11.5  million,  $9.5 
million, and $7.2 million, respectively. 

Goodwill and intangible Assets

Goodwill

As of December 31, 2023 and 2022, the goodwill balance was $462.3 million and $410.0 million, respectively. 
Goodwill preliminarily increased by $52.3 million, due to the Apton acquisition, of which $11.3 million relates to 
a deferred income tax liability. Goodwill is reviewed for impairment at least annually during the second quarter, 
or  more  frequently  if  an  event  occurs  indicating  the  potential  for  impairment.  We  performed  our  annual 
assessment for goodwill impairment in the second quarter of 2023, noting no impairment.

Acquired Intangible Assets

Intangible assets include acquired IPR&D of $55.0 million as a result of the Apton acquisition in August 2023. 
As  of  December  31,  2023,  the  research  and  development  project  had  not  been  completed  or  abandoned  and, 
therefore,  the  IPR&D  intangible  asset  is  not  currently  subject  to  amortization.  During  the  year  ended 
December  31,  2023,  acquired  IPR&D  of  $400.0  million  as  a  result  of  the  Omniome  acquisition  in  September 
2021 was completed and became subject to amortization. IPR&D is reviewed for impairment at least annually, 
or  more  frequently  if  an  event  occurs  indicating  the  potential  for  impairment.  We  performed  our  annual 
assessment for IPR&D impairment in the third quarter of 2023, noting no impairment.

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In addition to IPR&D, we had the following acquired definite-lived intangible assets as of December 31, 2023 (in 
thousands, except years):

Developed technology

Customer relationships

Total

Estimated 
Useful Life
(in years)

Gross
Carrying
Amount

Accumulated
Amortization

Net
Carrying
Amount

15

2

$ 

$ 

411,179  $ 

(9,195)  $ 

401,984 

360 

(360)   

— 

411,539  $ 

(9,555)  $ 

401,984 

Amortization  expense  of  intangibles  was  $8.3  million,  $0.9  million  and  $0.4  million  for  the  years  ended 
December  31,  2023,  2022,  and  2021,  respectively.  For  the  years  ended  December  31,  2023,    2022,  and  2021 
amortization  expense  of  intangibles  in  cost  of  revenue  was  $2.0  million,  $0.7  million,  and  $0.3  million, 
respectively.  For the years ended December 31, 2023, 2022, and 2021, amortization expense of intangibles in 
operating expenses was $6.3 million, $0.2 million, and $0.1 million, respectively.

Amortization of intangible assets is included within our cost of revenue if the costs and expenses related to the 
intangible  assets  are  attributable  to  revenue  generating  activities.  Amortization  expense  for  intangible  assets 
that are not directly related to sales generating activities are amortized to operating expenses. For developed 
technology  intangible  assets  that  are  utilized  in  both  revenue  generating  activities  and  in  research  and 
development activities, we allocate the amortization expense between cost of revenue and operating expenses. 
The  definite-lived  intangible  assets  are  amortized  using  the  straight-line  method  over  their  estimated  useful 
lives.

The  estimated  future  amortization  expense  of  acquisition-related  intangible  assets  with  definite  lives  is 
estimated as follows (in thousands):

2024

2025

2026

2027

2028

2029 and thereafter

Total

Accrued Expenses

Accrued expenses consisted of the following components: 

(in thousands)

Salaries and benefits

Accrued interest payable

Accrued purchase commitments

Accrued product development costs

Accrued professional services and legal fees

Inventory accrual

Warranty accrual

Other 

Accrued expenses 

Product Warranties

$ 

27,412 

27,412 

27,412 

27,412 

27,412 

264,924 

$ 

401,984 

December 31,

2023

2022

$ 

29,337  $ 

17,432 

2,834 

2,613 

1,033 

2,641 

353 

4,681 

2,216 

5,100 

3,705 

2,326 

1,005 

332 

1,651 

1,045 

$ 

45,708  $ 

32,596 

We  generally  provide  a  one-year  warranty  on  instruments.  In  addition,  we  provide  a  limited  warranty  on 
consumables. At the time revenue is recognized, an accrual is established for estimated warranty costs based 
on  historical  experience  as  well  as  anticipated  product  performance.  We  periodically  review  the  warranty 
reserve for adequacy and adjust the warranty accrual, if necessary, based on actual experience and estimated 

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costs to be incurred. Warranty expense is recorded as a component of cost of product revenue. There were no 
material changes in estimates for the periods presented below. 

Changes in the reserve for product warranties were as follows:

(in thousands)

Balance at beginning of period

Additions charged to cost of product revenue

Repairs and replacements

Balance at end of period

Deferred Revenue

Years Ended December 31,

2023

2022

$ 

$ 

1,651  $ 

8,227 

(5,197)   

4,681  $ 

594 

3,199 

(2,142) 

1,651 

As  of  December  31,  2023,  we  had  a  total  of  $21.9  million  of  deferred  revenue,  $16.3  million  of  which  was 
recorded  as  deferred  revenue,  current  and  primarily  relates  to  future  performance  obligations  under  the 
Amended and Restated Agreement with Invitae, as described in Note 3. Invitae Collaboration, as well as deferred 
service contract revenues. The deferred revenue, non-current balance of $5.6 million primarily relates to future 
performance  obligations  under  the  Amended  and  Restated  Agreement  with  Invitae  and  deferred  service 
contract  revenues  and  is  scheduled  to  be  recognized  in  the  next  5  years.  The  deferred  revenue,  non-current 
balance  includes  $2.9  million  that  was  reclassified  from  deferred  revenue,  current  to  deferred  revenue,  non-
current following receipt of a non-cancellable order from Invitae during the year ended December 31, 2023 for 
partial utilization of the available credits, which is expected to be recognized in revenue after 12 months from 
December  31,  2023.  Revenue  recorded  in  the  year  ended  December  31,  2023  includes  $18.9  million  of 
previously deferred revenue that was included in deferred revenue, current as of December 31, 2022.

Term Loans

In connection with the acquisition of Omniome, we acquired $1.3 million in short-term debt and $3.0 million in 
long-term debt relating to a term loan facility that Omniome obtained in April 2020. Borrowings on the term loan 
facility were used to fund Omniome’s purchases of equipment, which serves as collateral. Each term loan has a 
term of 43 months and bears a fixed interest rate of approximately 17% annually. The fee for the elective option 
to prepay all, but not less than all, of the borrowed amounts at any time after the 24th month and before the 43rd 
month  after  the  commencement  date,  is  4%  of  the  outstanding  loan  balance.  Payments  are  made  in  equal 
monthly installments including principal and interest. 

As  of  December  31,  2023,  the  carrying  value  of  term  loans  outstanding  was  $0.5  million,  recorded  as  part  of 
other liabilities, current on the consolidated balance sheet. The interest expense was $0.3 million for the year 
ended December 31,  2023, which was included as part of interest expense in the consolidated statements of 
operations and comprehensive loss.

As of December 31, 2023, the future principal payments remaining on term loans was the following:

(in thousands)

2024

Total

Other Liabilities, Current

Other liabilities, current, consisted of the following components: 

(in thousands)

Accrued Employee Stock Purchase Plan

Short-term loan

Other 

Other liabilities, current

$ 

$ 

490 

490 

December 31,

2023

2022

$ 

$ 

3,715  $ 

490 

4,121 

8,326  $ 

3,638 

1,842 

1,753 

7,233 

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NOTE 7. CONVERTIBLE SENIOR NOTES

2030 Convertible Senior Notes

In  June  2023,  we  entered  into  a  privately  negotiated  exchange  agreement  with  a  holder  of  our  outstanding 
1.50%  Convertible  Senior  Notes  due  2028  (the  “2028  Notes”),  pursuant  to  which  we  issued  $441.0  million  in 
aggregate principal amount of our 1.375% Convertible Senior Notes due 2030 (the “2030 Notes”) in exchange 
for  $441.0  million  principal  amount  of  the  2028  Notes  (the  “Exchange  Transaction”),  pursuant  to  exemptions 
from registration under the Securities Act of 1933, as amended, and the rules and regulations thereunder. The 
2030 Notes were issued on June 30, 2023.

The 2030 Notes are governed by an indenture (the “2030 Indenture”) between the Company and U.S. Bank Trust 
Company,  National  Association,  as  trustee.  The  2030  Notes  bear  interest  at  a  rate  of  1.375%  per  annum. 
Interest on the 2030 Notes is payable semi-annually in arrears on June 15 and December 15, commencing on 
December  15,  2023.  The  2030  Notes  will  mature  on  December  15,  2030,  subject  to  earlier  conversion, 
redemption or repurchase.

The 2030 Notes are convertible at the option of the holder at any time until the second scheduled trading day 
prior  to  the  maturity  date,  including  in  connection  with  a  redemption  by  the  Company.  The  2030  Notes  are 
convertible into shares of our common stock based on an initial conversion rate of 46.5116 shares of common 
stock per $1,000 principal amount of the 2030 Notes (which is equal to an initial conversion price of $21.50 per 
share of common stock), in each case subject to customary anti-dilution and other adjustments as a result of 
certain extraordinary transactions. Upon conversion of the 2030 Notes, we may elect to settle such conversion 
obligation in shares of our common stock, cash or a combination of shares of our common stock and cash.

On or after June 20, 2028, the 2030 Notes will be redeemable by the Company in the event that the closing sale 
price of our common stock has been at least 150% of the conversion price then in effect for at least 20 trading 
days (whether or not consecutive) during any 30 consecutive trading day period (including the last trading day 
of such period) ending on, and including, the trading day immediately preceding the date on which we provide 
the redemption notice at a redemption price of 100% of the principal amount of such 2030 Notes, plus accrued 
and unpaid interest up to, but excluding, the redemption date.

Upon  the  occurrence  of  a  Fundamental  Change  (as  defined  in  the  2030  Indenture),  the  holders  of  the  2030 
Notes may require that we repurchase all or part of the principal amount of the 2030 Notes at a purchase price 
equal to 100% of the principal amount of the notes to be repurchased, plus any accrued and unpaid interest up 
to,  but  excluding,  the  fundamental  change  repurchase  date,  and  all  unpaid  interest  from  the  fundamental 
change repurchase date thereon, but excluding, the maturity date.

The 2030 Indenture includes customary “events of default,” which may result in the acceleration of the maturity 
of  the  2030  Notes  under  the  2030  Indenture.  The  2030  Indenture  also  includes  customary  covenants  for 
convertible notes of this type.

To the extent we elect, the sole remedy for an event of default relating to our failure to comply with certain of 
our reporting obligations shall, for the first 360 calendar days after the occurrence of such an event of default, 
consist exclusively of the right to receive additional interest on the 2030 Notes at a rate equal to (i) 0.25% per 
annum of the principal amount of the 2030 Notes outstanding for each day during the first 180 calendar days of 
the  360-day  period  after  the  occurrence  of  such  an  event  of  default  during  which  such  event  of  default  is 
continuing (or, if earlier, the date on which such event of default is cured or waived) and (ii) 0.50% per annum of 
the principal amount of the 2030 Notes outstanding for each day from, and including, the 181st calendar day to, 
and including, the 360th calendar day after the occurrence of such an event of default during which such event 
of default is continuing (or, if earlier, the date on which such event of default is cured or waived as provided for 
in the 2030 Indenture). On the 361st day after such event of default (if the event of default relating to our failure 
to comply with its obligations is not cured or waived prior to such 361st day), the 2030 Notes shall be subject to 
acceleration as provided for in the 2030 Indenture.

The  2030  Notes  are  accounted  for  in  accordance  with  the  authoritative  guidance  for  convertible  debt 
instruments that may be settled in cash upon conversion. Under ASU 2020-06, the guidance requires that debt 
with an embedded conversion feature is accounted for in its entirety as a liability and no portion of the proceeds 
from the issuance of the convertible debt instrument is accounted for as attributable to the conversion feature 
unless  the  conversion  feature  is  required  to  be  accounted  for  separately  as  an  embedded  derivative  or  the 
conversion feature results in a substantial premium. The conversion feature of the 2030 Notes is not accounted 
for as an embedded derivative because it is considered to be indexed to our common stock, and the 2030 Notes 
were  not  issued  at  a  substantial  premium;  therefore,  the  2030  Notes  are  accounted  for  in  their  entirety  as  a 

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liability. Because we may elect to settle any conversions entirely in shares, and because settlement in shares is 
the default settlement method, the liability is classified as non-current. 

The requirement to repurchase the 2030 Notes, including unpaid interest to the maturity date in the event of a 
Fundamental  Change,  is  considered  a  put  option  for  certain  periods  requiring  bifurcation  under  ASC  815  – 
Derivatives and Hedging. However, given the low probability of such a Fundamental Change occurring during the 
applicable periods, the value of the embedded derivative is immaterial.

The  additional  interest  feature  in  the  event  of  our  failure  to  comply  with  certain  reporting  obligations  is  also 
considered an embedded derivative requiring bifurcation under ASC 815. However, due to the nature and terms 
of the reporting obligations, the value of the embedded derivative is immaterial.

The  Exchange  Transaction  was  accounted  for  as  an  extinguishment  driven  by  the  change  in  fair  value  of  the 
embedded  conversion  option.  We  recorded  a  loss  on  extinguishment  of  debt  of  approximately  $2.0  million  in 
connection  with  the  Exchange  Transaction  during  the  year  ended  December  31,  2023,  which  represents  the 
difference  between  the  fair  value  and  the  principal  amount  of  the  2030  Notes  of  the  debt  at  the  modification 
date, plus unamortized debt issuance costs of $1.5 million related to the respective portion of the 2028 Notes.

We  incurred  issuance  costs  related  to  the  2030  Notes  of  approximately  $7.3  million,  which  were  recorded  as 
debt issuance costs and are presented as a reduction to the 2030 Notes on our consolidated balance sheets 
and  are  amortized  to  interest  expense  using  the  effective  interest  method  over  the  term  of  the  2030  Notes, 
resulting in an effective interest rate of 1.6%. We also paid accrued but unpaid interest of $2.5 million on the 
2028 Notes in connection with the Exchange Transaction on June 30, 2023.

We did not receive any cash proceeds from the Exchange Transaction. In exchange for issuing the 2030 Notes 
pursuant  to  the  Exchange  Transaction,  we  received  and  cancelled  the  exchanged  2028  Notes.  Following  the 
closing  of  the  Exchange  Transaction,  $459.0  million  in  aggregate  principal  amount  of  2028  Notes  remained 
outstanding with terms unchanged.

The  net  carrying  amount  of  the  liability  for  the  2030  Notes  is  included  as  convertible  senior  notes,  net,  non-
current in the consolidated balance sheets as follows (in thousands):

Principal amount

Unamortized debt premium

Unamortized debt issuance costs

Net carrying amount

December 31,

2023

2022

$ 

$ 

$ 

$ 

441,000  $ 

524  $ 

(6,907)  $ 

434,617  $ 

Interest expense for the 2030 Notes for the years ended December 31, 2023, 2022, and 2021 was as follows:

(in thousands)

Contractual interest expense

Amortization of debt issuance costs

Total interest expense

Years Ended December 31,

2023

2022

2021

$ 

$ 

3,032  $ 

463 

3,495  $ 

—  $ 

— 

—  $ 

— 

— 

— 

— 

— 

— 

— 

As of December 31, 2023, the estimated fair value (Level 2) of the 2030 Notes was $410.5 million. The fair value 
of the 2030 Notes is estimated using a binomial lattice model that is primarily affected by the trading price of 
our common stock, market interest rates and volatility.

2028 Convertible Senior Notes

On February 9, 2021, we entered into an investment agreement (the “Investment Agreement”) with SB Northstar 
LP (the “Purchaser”), a subsidiary of SoftBank Group Corp., relating to the issuance and sale to the Purchaser of 
$900.0 million in aggregate principal amount of the 2028 Notes. The 2028 Notes were issued on February 16, 
2021.  As  discussed  above,  in  June  2023  we  completed  an  exchange  of  $441.0  million  in  aggregate  principal 
amount  of  our  2028  Notes  for  $441.0  million  aggregate  principal  amount  of  the  2030  Notes,  leaving 
approximately $459.0 million in aggregate principal amount of 2028 Notes outstanding.

The 2028 Notes are governed by an indenture (the “2028 Indenture”) between the Company and U.S. Bank 
National Association, as trustee. The 2028 Notes bear interest at a rate of 1.50% per annum. Interest on the 

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2028 Notes is payable semi-annually in arrears on February 15 and August 15 and commenced on August 15, 
2021. The 2028 Notes will mature on February 15, 2028, subject to earlier conversion, redemption, or 
repurchase.

The 2028 Notes are convertible at the option of the holder at any time until the second scheduled trading day 
prior  to  the  maturity  date,  including  in  connection  with  a  redemption  by  the  Company.  The  2028  Notes  are 
convertible into shares of our common stock based on an initial conversion rate of 22.9885 shares of common 
stock per $1,000 principal amount of the 2028 Notes (which is equal to an initial conversion price of $43.50 per 
share of common stock), in each case subject to customary anti-dilution and other adjustments as a result of 
certain extraordinary transactions. Upon conversion of the 2028 Notes, we may elect to settle such conversion 
obligation in shares of our common stock, cash or a combination of shares of our common stock and cash.

On or after February 20, 2026, the 2028 Notes will be redeemable by the Company in the event that the closing 
sale  price  of  our  common  stock  has  been  at  least  150%  of  the  conversion  price  then  in  effect  for  at  least  20 
trading  days  (whether  or  not  consecutive)  during  any  30  consecutive  trading  day  period  (including  the  last 
trading day of such period) ending on, and including, the trading day immediately preceding the date on which 
we provide the redemption notice at a redemption price of 100% of the principal amount of such 2028 Notes, 
plus accrued and unpaid interest up to, but excluding, the redemption date.

Upon  the  occurrence  of  a  Fundamental  Change  (as  defined  in  the  2028  Indenture),  the  holders  of  the  2028 
Notes may require that we repurchase all or part of the principal amount of the 2028 Notes at a purchase price 
of par plus unpaid interest up to, but excluding, the maturity date.

The 2028 Indenture includes customary “events of default,” which may result in the acceleration of the maturity 
of  the  2028  Notes  under  the  2028  Indenture.  The  2028  Indenture  also  includes  customary  covenants  for 
convertible notes of this type.

To the extent we elect, the sole remedy for an event of default relating to our failure to comply with certain of 
our reporting obligations shall, for the first 360 calendar days after the occurrence of such an event of default, 
consist exclusively of the right to receive additional interest on the 2028 Notes at a rate equal to (i) 0.25% per 
annum of the principal amount of the 2028 Notes outstanding for each day during the first 180 calendar days of 
the  360-day  period  after  the  occurrence  of  such  an  event  of  default  during  which  such  event  of  default  is 
continuing (or, if earlier, the date on which such event of default is cured or waived) and (ii) 0.50% per annum of 
the principal amount of the 2028 Notes outstanding for each day from, and including, the 181st calendar day to, 
and including, the 360th calendar day after the occurrence of such an event of default during which such event 
of default is continuing (or, if earlier, the date on which such event of default is cured or waived as provided for 
in the 2028 Indenture). On the 361st day after such event of default (if the event of default relating to our failure 
to comply with its obligations is not cured or waived prior to such 361st day), the 2028 Notes shall be subject to 
acceleration as provided for in the 2028 Indenture.

The  2028  Notes  are  accounted  for  in  accordance  with  the  authoritative  guidance  for  convertible  debt 
instruments that may be settled in cash upon conversion. Under ASU 2020-06, the guidance requires that debt 
with an embedded conversion feature is accounted for in its entirety as a liability and no portion of the proceeds 
from the issuance of the convertible debt instrument is accounted for as attributable to the conversion feature 
unless  the  conversion  feature  is  required  to  be  accounted  for  separately  as  an  embedded  derivative  or  the 
conversion feature results in a substantial premium. The conversion feature of the 2028 Notes is not accounted 
for as an embedded derivative because it is considered to be indexed to our common stock, and the 2028 Notes 
were  not  issued  at  a  premium;  therefore,  the  2028  Notes  are  accounted  for  in  their  entirety  as  a  liability. 
Because  we  may  elect  to  settle  any  conversions  entirely  in  shares,  and  because  settlement  in  shares  is  the 
default settlement method, the liability is classified as non-current. 

The requirement to repurchase the 2028 Notes including unpaid interest to the maturity date in the event of a 
Fundamental  Change  is  considered  a  put  option  for  certain  periods  requiring  bifurcation  under  ASC  815  – 
Derivatives  and  Hedging.  However,  given  the  low  probability  of  a  Fundamental  Change  occurring  during  the 
applicable periods, the value of the embedded derivative is immaterial.

The  additional  interest  feature  in  the  event  of  our  failure  to  comply  with  certain  reporting  obligations  is  also 
considered an embedded derivative requiring bifurcation under ASC 815. However, due to the nature and terms 
of the reporting obligations, the value of the embedded derivative is immaterial.

We  incurred  issuance  costs  related  to  the  2028  Notes  of  approximately  $4.5  million,  which  were  recorded  as 
debt issuance cost and are presented as a reduction to the 2028 Notes on our consolidated balance sheets and 

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are amortized to interest expense using the effective interest method over the term of the 2028 Notes, resulting 
in an effective interest rate of 1.6%. 

The  net  carrying  amount  of  the  liability  for  the  2028  Notes  is  included  as  convertible  senior  notes,  net,  non-
current in the consolidated balance sheets as follows (in thousands):

Principal amount

Unamortized debt issuance costs

Net carrying amount

December 31,

2023

2022

$ 

$ 

459,000  $ 

900,000 

(1,374)   

(3,317) 

457,626  $ 

896,683 

Interest expense for the 2028 Notes was as follows for the years ended December 31, 2023, 2022, and 2021:

(in thousands)

Contractual interest expense

Amortization of debt issuance costs

Total interest expense

Years Ended December 31,

2023

2022

2021

$ 

$ 

10,133  $ 

13,500  $ 

11,812 

472 

617 

532 

10,605  $ 

14,117  $ 

12,344 

As of December 31, 2023, the estimated fair value (Level 2) of the 2028 Notes was $395.4 million. The fair value 
of the Notes is estimated using a pricing model that is primarily affected by the trading price of our common 
stock and market interest rates.

NOTE 8. COMMITMENTS AND CONTINGENCIES 

Leases

We record operating lease right-of-use assets and liabilities on our consolidated balance sheets for all leases 
with a term of more than 12 months. In connection with the acquisition of Omniome, we acquired $18.1 million 
in right-of-use assets and liabilities on our consolidated balance sheets. The operating lease right-of-use assets 
and  liabilities  are  calculated  as  the  present  value  of  remaining  minimum  lease  payments  over  the  remaining 
lease  term  using  our  estimated  secured  incremental  borrowing  rates  at  the  commencement  date.  Lease 
payments included in the measurement of the lease liability comprise the fixed rent per the term of the Lease. 
All of our leases are operating leases. Lease payments comprise the base rent per the term of the Lease. Lease 
expense  for  these  leases  is  recognized  on  a  straight-line  basis  over  the  lease  term,  with  variable  lease 
payments, such as common area maintenance fees, recognized in the period those payments are incurred.

We  often  have  options  to  renew  lease  terms  for  buildings.  In  addition,  certain  lease  arrangements  may  be 
terminated prior to their original expiration date at our discretion. We evaluate renewal and termination options 
at the lease commencement date to determine if we are reasonably certain to exercise the option on the basis 
of economic factors.

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As of December 31, 2023, the maturities of our operating lease liabilities were as follows:

(in thousands)

2024

2025

2026

2027

2028

Thereafter

Total undiscounted operating lease payments

Less: imputed interest

Present value of operating lease liabilities 

Balance Sheet Classification

Operating lease liabilities, current

Operating lease liabilities, non-current

Total operating lease liabilities

$ 

12,082 

12,307 

12,412 

9,930 

— 

— 

46,731 

(5,534) 

$ 

41,197 

$ 

$ 

9,591 

31,606 

41,197 

We use our incremental borrowing rate to determine the present value of lease payments, as the implicit rates in 
our  leases  are  not  readily  determinable.  The  weighted-average  discount  rate  used  to  measure  our  operating 
lease  liabilities  was  6.8%.  The  weighted-average  remaining  lease  term  for  our  operating  leases  as  of 
December 31, 2023 was 3.8 years.

Cash Flows

Cash  paid  for  amounts  included  in  the  present  value  of  operating  lease  liabilities  was  $12.1  million  and 
$11.2  million  for  the  years  ended  December  31,  2023  and  2022,  respectively,  and  were  included  in  operating 
cash flows. 

Operating Lease Costs

Operating lease costs were $10.4 million and $10.5 million for the years ended December 31, 2023 and 2022, 
respectively.

Contingencies 

We  may  become  involved  in  legal  proceedings,  claims  and  assessments  from  time  to  time  in  the  ordinary 
course of business. We accrue liabilities for such matters when it is probable that future expenditures will be 
made and such expenditures can be reasonably estimated. 

We do not believe that the ultimate outcome of any such pending matters is probable or reasonably estimable, 
or that these matters will have a material adverse effect on our business; however, the results of litigation and 
claims  are  inherently  unpredictable.  Regardless  of  the  outcome,  litigation  can  have  an  adverse  impact  on  us 
because of litigation and settlement costs, diversion of management resources, and other factors.

Please see subsection titled Legal Proceedings, in Part I, Item 3 of this Annual Report on Form 10-K.

Indemnification

Pursuant  to  Delaware  law  and  agreements  entered  into  with  each  of  our  directors  and  officers,  we  may  have 
obligations,  under  certain  circumstances,  to  hold  harmless  and  indemnify  each  of  our  directors  and  officers 
against  losses  suffered  or  incurred  by  the  indemnified  party  in  connection  with  their  service  to  us,  and 
judgements, fines, settlements and expenses related to claims arising against such directors and officers to the 
fullest extent permitted under Delaware law, our bylaws and our certificate of incorporation. We also enter and 
have entered into indemnification agreements with our directors and officers that may require us to indemnify 
them  against  liabilities  that  arise  by  reason  of  their  status  or  service  as  directors  or  officers,  except  as 
prohibited by applicable law. In addition, we may have obligations to hold harmless and indemnify third parties 
involved  with  our  fundraising  efforts  and  their  respective  affiliates,  directors,  officers,  employees,  agents  or 
other  representatives  against  any  and  all  losses,  claims,  damages  and  liabilities  related  to  claims  arising 
against  such  parties  pursuant  to  the  terms  of  agreements  entered  into  between  such  third  parties  and  us  in 

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connection with such fundraising efforts. To the extent that any such indemnification obligations apply to the 
lawsuits  described  above,  any  associated  expenses  incurred  are  included  within  the  related  accrued  litigation 
expense amounts. No additional liability associated with such indemnification obligations has been recorded as 
of December 31, 2023.

Purchase Commitments

In the normal course of business, we enter into agreements to purchase goods or services or license intellectual 
property, certain of which are not cancellable without penalty. For those agreements with variable terms, we do 
not estimate the total obligation beyond any minimum quantities or pricing as of the reporting date. Licensing 
agreements  under  which  we  commit  to  ongoing  minimum  royalty  payments,  some  of  which  are  subject  to 
adjustment, may be terminated under certain circumstances.

Our  purchase  orders  and  contractual  obligations  are  approximately  $109.9  million  as  of  December  31,  2023, 
which consist of open purchase orders and contractual obligations in the ordinary course of business, including 
commitments with contract manufacturers and suppliers for which we have not received the goods or services. 
A majority of these purchase obligations are due within a year. Although open purchase orders are considered 
enforceable  and  legally  binding,  the  terms  generally  allow  us  the  option  to  cancel,  reschedule  and  adjust  our 
requirements based on our business needs prior to the delivery of goods or performance of services.

We recognized a loss on purchase commitment of $3.4 million for the year ended December 31, 2023, which 
was  recorded  as  part  of  accrued  expenses  on  the  consolidated  balance  sheet  and  is  included  in  the 
aforementioned purchase orders and contractual obligations amount. The purchase commitment loss is based 
on an estimate of future excess inventory related to a supply agreement with a third-party vendor, for which we 
do not expect to have related sales.

We have a long-term supply agreement, which was amended in October 2022 (the “Supply Agreement”), for the 
purchase of certain products with a semiconductor manufacturer (“Supplier”). The Supply Agreement provides 
for minimum purchase commitments through 2026 on our part in exchange for guaranteed capacity at Supplier. 
We  are  responsible  for  providing  certain  materials  to  allow  our  Supplier  to  perform  its  obligations  under  the 
contract. 

We  paid  our  Supplier  a  deposit  of  $9.0  million  in  November  2022  and  an  additional  deposit  of  $6.0  million  in 
2023, for a total of $15.0 million (the “Deposit”). The Deposit is fully refundable to us, in accordance with the 
Supply  Agreement,  if  we  meet  the  minimum  volume  purchase  commitment  for  the  applicable  year.  As  of 
December  31,  2023,  $3.0  million  related  to  the  Deposit  was  included  in  prepaid  expenses  and  other  current 
assets in the consolidated balance sheets and $12.0 million related to the Deposit was included in other long-
term  assets  in  the  consolidated  balance  sheets,  as  we  believe  it  is  probable  the  minimum  volume  purchase 
commitment level will be achieved.

NOTE 9. INCOME TAXES

We are subject to income taxes both in the United States and certain foreign jurisdictions in which we operate, 
and  we  use  estimates  in  determining  our  provisions  for  income  taxes.  Significant  management  judgement  is 
required  in  determining  our  provision  for  income  taxes,  deferred  tax  assets  and  liabilities,  and  valuation 
allowances  recorded  against  net  deferred  tax  assets  in  accordance  with  U.S.  GAAP.  These  estimates  and 
judgements occur in the calculation of tax credits, benefits, and deductions, and in the calculation of certain tax 
assets and liabilities, which arise from differences in the timing of recognition of revenue and expense for tax 
and  financial  statement  purposes,  as  well  as  the  interest  and  penalties  related  to  uncertain  tax  positions. 
Significant changes to these estimates may result in an increase or decrease to our tax provision in the current 
or subsequent period.

We assess all material positions taken in any income tax return, including all significant uncertain positions, in 
all  tax  years  that  are  still  subject  to  assessment  or  challenge  by  relevant  taxing  authorities.  Assessing  an 
uncertain tax position begins with the initial determination of the position’s sustainability and is measured at the 
largest amount of benefit that is greater than 50% likely of being realized upon ultimate settlement. As of each 
balance sheet date, unresolved uncertain tax positions must be reassessed, and we will determine whether the 
factors  underlying  the  sustainability  assertion  have  changed  and  the  amount  of  the  recognized  tax  benefit  is 
still appropriate.

We account for Global Intangible Low-taxed Income as a period cost.

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During  the  years  ended  December  31,  2023,  2022,  and  2021  income/(loss)  before  taxes  from  U.S.  operations 
were ($318.9) million, ($315.7) million, and ($275.4) million, respectively, and income/(loss) before taxes from 
foreign operations was $0.7 million, $1.8 million, and $0.8 million, respectively. 

Provision (Benefit) for Income Taxes

The provision (benefit) for income taxes consists of the following (in thousands):

Total Current

Deferred:

Federal

State

Foreign

Total Deferred

Years ended December 31,

2023

2022

2021

$ 

—  $ 

—  $ 

— 

(9,956)   

(1,468)   

— 

(11,424)   

— 

— 

— 

— 

(83,742) 

(9,907) 

— 

(93,649) 

(Benefit) Provision for Income Taxes

$ 

(11,424)  $ 

—  $ 

(93,649) 

Income tax provision (benefit) related to continuing operations differ from the amounts computed by applying 
the statutory income tax rate of 21% to pretax loss as follows: 

Statutory tax rate 

State tax rate, net of federal benefit

Change in valuation allowance 

Tax credits

Share-based compensation

Merger Expenses

Other

Total

Years ended December 31,

2023

2022

2021

 21.0 %

 3.0 

 (20.0) 

 2.0 

 (2.1) 

 (0.1) 

 (0.2) 

 3.6 %

 21.0 %

 4.4 

 (25.1) 

 2.2 

 (2.2) 

 — 

 (0.4) 

 (0.1) %

 21.0 %

 5.5 

 (4.9) 

 2.5 

 10.9 

 (0.9) 

 (0.1) 

 34.0 %

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Deferred  income  taxes  reflect  the  net  tax  effects  of  loss  and  credit  carryforwards  and  temporary  differences 
between the carrying amounts of assets and liabilities for financial reporting purposes and the amounts used 
for income tax purposes. Significant components of our deferred tax assets for federal and state income taxes 
are as follows (in thousands): 

Deferred tax assets:

Net operating loss carryforwards

Research and development credits 

Capitalized research and experimental expenses

Accruals and reserves

Cancellation of indebtedness income and interest expense

Share-based compensation

Operating lease liability

Total deferred tax assets 

Less: Valuation allowance

Total deferred tax assets:

Intangibles

Fixed assets

Operating lease right-of-use assets

Total deferred tax liabilities

Deferred tax liabilities, net

December 31,

2023

2022

$ 

435,488  $ 

400,629 

83,922 

63,196 

16,872 

14,907 

18,584 

9,510 

71,526 

34,863 

13,830 

4,587 

17,117 

11,537 

642,479 

554,089 

(525,703)   

(445,574) 

116,776 

108,515 

(109,488)   

(98,931) 

(548)   

(7,491)   

(1,262) 

(9,157) 

(117,527)   

(109,350) 

$ 

(751)  $ 

(835) 

At December 31, 2023, we maintained a valuation allowance against our net deferred tax assets which totaled 
$525.7  million,  including  net  operating  loss  carryforwards  and  research  and  development  credits  of  $435.5 
million and $83.9 million, respectively.

A valuation allowance is recorded when it is more likely than not that all or some portion of the deferred income 
tax  assets  will  not  be  realized.  We  regularly  assess  the  need  for  a  valuation  allowance  against  our  deferred 
income tax assets by considering both positive and negative evidence related to whether it is more likely than 
not that our deferred income tax assets will be realized. In evaluating our ability to recover our deferred income 
tax assets within the jurisdiction from which they arise, we consider all available positive and negative evidence, 
including  scheduled  reversals  of  deferred  income  tax  liabilities,  projected  future  taxable  income,  tax-planning 
strategies, and results of recent operations. A deferred income tax benefit of $11.4 million for the year ended 
December  31,  2023,  is  related  to  the  release  of  the  valuation  allowance  for  deferred  tax  assets  due  to  the 
recognition  of  deferred  tax  liabilities  in  connection  with  the  Apton  acquisition.  We  maintain  a  valuation 
allowance on the net deferred tax assets of our U.S. entities as we have concluded that it is more likely than not 
that we will not realize our deferred tax assets. Accordingly, this benefit from income taxes is reflected on our 
consolidated statements of operations and comprehensive loss for the year ended December 31, 2023. 

For  the  year  ended  December  31,  2023,  the  Company's  valuation  allowance  increased  to  $525.7  million, 
primarily because of an increase in our net operating losses, credits, and capitalized research and experimental 
expenses that were fully offset by a valuation allowance. For the year ended December 31, 2022, the Company's 
valuation allowance increased to $445.6 million, primarily because of an increase in our net operating losses, 
credits, and capitalized research and experimental expenses that were fully offset by a valuation allowance. 

As  of  December  31,  2023,  we  had  a  net  operating  loss  carryforward  for  federal  income  tax  purposes  of 
approximately  $1,709.3  million,  of  which  $788.1  million  is  subject  to  expiration  beginning  in  2024.  We  had  a 
total  state  net  operating  loss  carryforward  of  approximately  $1,171.8  million,  which  are  subject  to  annual 
expirations. Utilization of some of the federal and state net operating loss and credit carryforwards are subject 
to  annual  limitations  due  to  the  “change  in  ownership”  provisions  of  the  Internal  Revenue  Code  of  1986  and 
similar state provisions. The annual limitations may result in the expiration of net operating losses and credits 
before utilization. 

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We  have  federal  credits  of  approximately  $57.7  million,  which  will  begin  to  expire  in  2024  if  not  utilized  and 
state  research  credits  of  approximately  $49.8  million,  which  have  no  expiration  date.  These  tax  credits  are 
subject to the same limitations discussed above. 

As of December 31, 2023, our total unrecognized tax benefit was $14.6 million. A reconciliation of the beginning 
and ending unrecognized tax benefit balance is as follows (in thousands): 

Balance as of December 31, 2020

$ 

Increase in balance related to tax positions taken in prior year 

Increase in balance related to tax positions taken during current year 

Balance as of December 31, 2021

Decrease in balance related to tax positions taken in prior year 

Increase in balance related to tax positions taken during current year 

Balance as of December 31, 2022

Increase in balance related to tax positions taken in prior year 

Increase in balance related to tax positions taken during current year 

5,954 

189 

2,192 

8,335 

(10) 

2,085 

10,410 

2,044 

2,100 

Balance as of December 31, 2023

$ 

14,554 

Our practice is to recognize interest and/or penalties related to income tax matters in income tax expense. As 
of  December  31,  2023  and  2022,  we  had  no  accrued  interest  or  penalties  due  to  our  net  operating  losses 
available to offset any tax adjustment. If total unrecognized tax benefits were realized in the future, it would not 
result in any tax benefit as we currently have a full valuation allowance. We file U.S. federal and various state 
income tax returns. For U.S. federal and state income tax purposes, the statute of limitations currently remains 
open  for  the  years  ending  December  31,  2020  to  present  and  December  31,  2019  to  present,  respectively.  In 
addition, all the net operating losses and research and development credit carryforwards that may be utilized in 
future years may be subject to examination. We are not currently under examination by income tax authorities in 
any jurisdiction.

NOTE 10. STOCKHOLDERS’ EQUITY

Preferred Stock 

Our Certificate of Incorporation, as amended and restated in October 2010 in connection with the closing of our 
initial  public  offering,  authorizes  us  to  issue  1,000,000,000  shares  of  $0.001  par  value  common  stock  and 
50,000,000  shares  of  $0.001  par  value  preferred  stock.  As  of  December  31,  2023  and  2022,  there  were  no 
shares of preferred stock issued or outstanding.

Common Stock 

Common stockholders are entitled to dividends when and if declared by our board of directors. There have been 
no dividends declared to date. The holder of each share of common stock is entitled to one vote. 

Underwritten Public Equity Offerings

In August 2020, we entered into an underwriting agreement, relating to the public offering of 19,430,000 shares 
of our common stock, $0.001 par value per share, at a price to the public of $4.47 per share. Under the terms of 
the underwriting agreement, we also granted the underwriters a 30-day option to purchase up to an additional 
2,914,500 shares of our common stock, which was subsequently exercised in full, and the offering including the 
sale  of  shares  of  common  stock  subject  to  the  underwriters’  option,  closed  in  August  2020.  In  total,  we  sold 
22.3 million shares of our common stock. We paid a commission equal to 6% of the gross proceeds from the 
sale  of  shares  of  our  common  stock.  The  total  net  proceeds  to  us  from  the  offering  after  deducting  the 
underwriting  discount  were  approximately  $93.9  million,  excluding  approximately  $0.3  million  of  offering 
expenses. 

In  November  2020,  we  entered  into  an  underwriting  agreement,  relating  to  the  public  offering  of  6,096,112 
shares of our common stock, $0.001 par value per share, at a price to the public of $14.25 per share. Under the 
terms  of  the  underwriting  agreement,  we  also  granted  the  underwriters  a  30-day  option  to  purchase  up  to  an 
additional  914,416  shares  of  our  common  stock,  which  was  subsequently  exercised  in  full,  and  the  offering 
including the sale of shares of common stock subject to the underwriters’ option, closed in November 2020. In 
total,  we  sold  7.0  million  shares  of  our  common  stock.  We  paid  a  commission  equal  to  6%  of  the  gross 
proceeds  from  the  sale  of  shares  of  our  common  stock.  The  total  net  proceeds  to  us  from  the  offering  after 

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deducting the underwriting discount were approximately $93.9 million, excluding approximately $0.3 million of 
offering expenses. 

In total, for the year ended December 31, 2021, we issued 29.4 million shares of our common stock through our 
two underwritten public offerings with an average offering price of $6.40. The total net proceeds to us from the 
two offerings, after deducting the underwriting commission and offering expenses, were approximately $187.2 
million. 

In  January  2023,  we  entered  into  an  underwriting  agreement,  relating  to  the  public  offering  of  17.5  million 
shares of our common stock, $0.001 par value per share, at a price to the public of $10.00 per share. Under the 
terms  of  the  underwriting  agreement,  we  also  granted  the  underwriters  a  30-day  option  to  purchase  up  to  an 
additional 2.6 million shares of our common stock, which was subsequently exercised in full, and the offering, 
including  the  sale  of  shares  of  common  stock  subject  to  the  underwriters'  option,  closed  in  January  2023.  In 
total,  we  sold  20.1  million  shares  of  our  common  stock.  We  paid  a  commission  equal  to  5.75%  of  the  gross 
proceeds  from  the  sale  of  shares  of  our  common  stock.  The  total  net  proceeds  to  us  from  the  offering  after 
deducting the underwriting discount were approximately $189.7 million, excluding approximately $0.5 million of 
offering expenses.

Private Placement of Common Stock

On  July  19,  2021,  in  connection  with  the  Omniome  acquisition,  we  entered  into  a  purchase  agreement  with 
certain qualified institutional buyers and institutional accredited investors, pursuant to which we agreed to sell 
an  aggregate  of  11,214,953  shares  of  common  stock,  at  a  price  of  $26.75  per  share,  for  aggregate  gross 
proceeds  of  approximately  $300  million.  The  transaction  closed  on  September  20,  2021.  We  registered  the 
private placement shares for resale following the closing of the merger.

Equity Plans 

The 2020 Equity Incentive Plan (the “2020 Plan”), the 2020 Inducement Equity Incentive Plan (the “Inducement 
Plan”),  and  the  2021  adopted  Omniome  Equity  Incentive  Plan  of  Pacific  Biosciences  of  California,  Inc.  (the 
“Omniome Plan”) allow for the issuance of stock options, restricted units and awards, and performance-based 
awards.

On December 2, 2020, the Board of Directors (the “Board”) adopted the Inducement Plan and reserved 2,500,000 
shares  of  our  common  stock  for  issuance  pursuant  to  equity  awards  granted  under  the  Inducement  Plan.  On 
April  18,  2021  and  November  22,  2021,  the  Board  amended  the  Inducement  Plan  to  reserve  an  additional 
750,000 and 360,000 shares, respectively. 

On  September  20,  2021,  in  connection  with  the  acquisition  of  Omniome,  we  adopted  the  Omniome  Equity 
Incentive  Plan  of  Pacific  Biosciences  of  California,  Inc.  (the  “Omniome  Plan”).  Under  the  Omniome  Merger 
Agreement, each unvested option to purchase Omniome common stock, granted under the Omniome Plan held 
by employees continuing with us, was assumed by PacBio and converted into an option to purchase shares of 
our  common  stock.  The  terms  and  conditions  of  the  converted  options  are  substantially  the  same  (including 
vesting  and  exercisability),  except  that  (A)  the  assumed  options  cover  shares  of  PacBio’s  common  stock;  (B) 
the  number  of  shares  of  our  common  stock  subject  to  the  assumed  option  is  equal  to  the  product  of  (i)  the 
number of shares of Omniome common stock subject to the corresponding unvested option, multiplied by (ii) 
the exchange ratio (as defined below), with any resulting fractional share rounded down to the nearest whole 
share; and (C) the exercise price per share of the assumed options is equal to the quotient of (i) the exercise 
price per share of the corresponding unvested option to purchase shares of Omniome common stock, divided 
by (ii) the exchange ratio (as defined below), with any resulting fractional cent rounded up to the nearest whole 
cent. The exchange ratio was equal to 0.259204639. We reserved 2,494,128 shares of our common stock for 
issuance pursuant to equity awards under the Omniome Plan. 

On  May  25,  2022,  stockholders  approved  an  amendment  to  the  2020  Plan,  and  we  reserved  an  additional 
18,000,000 shares of our common stock for issuance pursuant to equity awards granted under the 2020 Plan.

As  of  December  31,  2023,  we  had  12.8  million  shares  remaining  and  available  for  future  issuance  under  the 
2020 Plan, Inducement Plan, and the Omniome Plan.

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Stock Options

Time-based stock options

The  following  table  summarizes  time-based  stock  option  activity  for  all  of  our  equity  compensation  plans  for 
the year ended December 31, 2023 (in thousands, except per share amounts):

Outstanding at December 31, 2022

Granted 

Exercised 

Canceled 

Outstanding at December 31, 2023

Performance-based stock options

Number
of shares

Weighted-
average
exercise price

14,618 $ 

419 $ 

(1,119) $ 

(910) $ 

13,008 $ 

10.60 

11.26 

4.74 

17.71 

10.63 

The  following  table  summarizes  performance-based  stock  option  activity  for  all  of  our  equity  compensation 
plans for the year ended December 31, 2023 (in thousands, except per share amounts):

Outstanding at December 31, 2022

Granted 

Exercised 

Canceled 

Outstanding at December 31, 2023

Number
of shares

Weighted-
average
exercise price

258 $ 

— $ 

(251) $ 

(4) $ 

3 $ 

4.71 

— 

4.71 

4.71 

4.74 

The performance condition was achieved during the year ended December 31, 2023.

The  aggregate  intrinsic  value  of  the  outstanding  options  presented  in  the  tables  above  as  of  December  31, 
2023, totaled $31.8 million, and had a weighted-average remaining contractual life of 5.9 years.

The  aggregate  intrinsic  value  of  outstanding  options  represents  the  total  pre-tax  intrinsic  value  (i.e.  the 
difference between $9.81, our closing stock price on the last trading day of our fourth quarter of 2023 and the 
option exercise price multiplied by the number of in-the-money options) that would have been received by the 
option  holders  had  all  option  holders  exercised  their  options  on  December  31,  2023.  The  aggregate  intrinsic 
value changes at each reporting date based on the fair market value of our common stock.

The  vested  and  exercisable  options  as  of  December  31,  2023,  totaled  10,039,742  shares,  had  an  aggregate 
intrinsic value of $30.0 million, a weighted-average exercise price per share of $9.70, and a weighted-average 
remaining contractual life of 5.2 years. 

The vested and expected to vest options as of December 31, 2023, totaled 13,026,464 shares, had an aggregate 
intrinsic value of $31.6 million, a weighted-average exercise price per share of $10.81, and a weighted-average 
remaining contractual life of 5.8 years. 

The total intrinsic value of stock options exercised during the years ended December 31, 2023, 2022, and 2021 
was  $8.8  million,  $5.0  million,  and  $146.1  million,  respectively.  The  total  intrinsic  value  of  options  exercised 
represents  the  difference  between  our  closing  stock  price  on  the  exercise  date  and  the  option  exercise  price, 
multiplied by the number of in-the-money options exercised.

The weighted-average grant-date fair value of all options granted was $7.32 in 2023, $5.93 in 2022, and $18.36 
in 2021, each determined by the Black-Scholes option valuation method.

Restricted Stock Units ("RSU") and Performance Stock Units ("PSU")

Each  Restricted  Stock  Unit  (RSU)  represents  one  equivalent  share  of  our  common  stock  to  be  issued  after 
satisfying the applicable continued service-based vesting criteria over a specified period. These RSUs are time-
based and vest over four years at a rate of 25% annually. The RSUs do not entitle participants to the rights of 
holders  of  common  stock,  such  as  voting  rights,  until  the  shares  are  issued.  The  fair  value  of  these  RSUs  is 

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based on the closing price of our common stock on the date of grant. We measure compensation expense for 
these RSUs at fair value on the date of grant and recognize the expense over the expected vesting period on a 
straight-line basis.  RSUs that are expected to vest are net of estimated future forfeitures. 

We issue PSUs for which the number of shares issuable in the third year of the performance period is based on 
performance  relative  to  specified  revenue  targets  and  continued  employment  through  the  vesting  period.  
Maximum  achievement  of  the  revenue  goal  under  the  PSUs  will  result  in  up  to  200%  of  the  target  number  of 
shares  subject  to  the  PSUs  to  become  eligible  to  vest,  while  not  meeting  the  minimum  achievement  of  the 
revenue goal under PSUs will result in no shares subject to the PSUs becoming eligible to vest.

The following table summarizes the time-based RSUs and PSUs activity for the year ended December 31, 2023 
(shares in thousands): 

Outstanding at December 31, 2022

Granted

Vested

Forfeited

Outstanding at December 31, 2023

Weighted-average grant date fair 
value

Restricted 
Stock Units 
(RSUs)

Performance 
Stock Units 
(PSUs)

RSU

PSU

8,535

7,141

(2,730)

(1,638)

11,308

— $ 

15.16  $ 

564  

—  

(23)

9.67 

14.41 

13.86 

541 $ 

12.06  $ 

— 

9.43 

— 

9.43 

9.43 

The  total  fair  value  of  shares  vested  related  to  RSUs  and  PSUs  during  the  years  ended  December  31,  2023, 
2022, and 2021 was $39.3 million, $39.2 million, and $9.2 million, respectively. 

The  weighted-average  grant-date  fair  value  of  all  RSUs  and  PSUs  granted  was  $9.65  in  2023,  $10.15  in  2022, 
and $35.33 in 2021. 

Employee Stock Purchase Plan 

As of December 31, 2023, a total of 29.5 million shares of our common stock have been reserved for issuance 
under our 2010 Employee Stock Purchase Plan (the “ESPP”). The ESPP permits eligible employees to purchase 
common stock at a discount through payroll deductions during defined offering periods. Each offering period 
will generally consist of four purchase periods, each purchase period being approximately six months. The price 
at which the stock is purchased is equal to the lower of 85% of the fair market value of the common stock at the 
beginning of an offering period or at the end of a purchase period. If the stock price at the end of the purchase 
period  is  lower  than  the  stock  price  at  the  beginning  of  the  offering  period,  that  offering  period  will  be 
terminated  and  a  new  offering  period  will  come  into  place.  The  ESPP  provides  for  an  annual  increase  to  the 
shares  available  for  issuance  at  the  beginning  of  each  fiscal  year  equal  to  the  lesser  of  2%  of  the  common 
shares then outstanding, 4,000,000 shares, or an amount determined by the ESPP’s administrator. 

For  the  years  ended  December  31,  2023,  2022,  and  2021,  1,735,058  shares,  1,878,168  shares,  and  1,913,968 
shares of common stock were purchased under the ESPP, respectively. As of December 31, 2023, 12,197,447 
shares of our common stock remain available for issuance under our ESPP.

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Share-based Compensation

Total share-based compensation expense consists of the following (in thousands): 

Cost of revenue

Research and development

Sales, general and administrative

Merger-related expenses - stock-settled

Merger-related expenses - milestone

Share-based compensation

Merger-related expenses - cash-settled

Years Ended December 31,

2023

2022

2021

$ 

5,399  $ 

4,802  $ 

22,435 

44,284 

— 

— 

30,676 

43,135 

— 

— 

72,118 

78,613 

— 

— 

6,126 

20,275 

35,403 

6,349 

5,202 

73,355 

7,373 

Total share-based compensation expense

$ 

72,118  $ 

78,613  $ 

80,728 

As  of  December  31,  2023  and  2022,  $0.6  million  and  $0.7  million  of  share-based  compensation  cost  was 
capitalized in inventory, net, on our consolidated balance sheets, respectively. 

The tax benefit of share-based compensation expense was immaterial for the years ended December 31, 2023, 
2022, and 2021. 

Determining Fair Value

We  estimate  the  fair  value  of  share  options  granted  using  the  Black-Scholes  valuation  method  and  a  single 
option  award  approach.  This  fair  value  is  then  amortized  on  a  straight-line  basis  over  the  requisite  service 
periods of the awards, which is generally the vesting period. The fair market value of RSU awards granted is the 
closing  price  of  our  shares  on  the  date  of  grant  and  is  generally  recognized  as  compensation  expense  on  a 
straight-line basis over the respective vesting period. For shares purchased under our Employee Stock Purchase 
Plan,  or  ESPP,  we  estimate  the  grant-date  fair  value,  and  the  resulting  share-based  compensation  expense, 
using the Black-Scholes option-pricing model.

Expected Term – The expected term used in the Black-Scholes valuation method represents the period that the 
stock  options  are  expected  to  be  outstanding  and  is  determined  based  on  historical  experience  of  similar 
awards, giving consideration to the contractual terms of the stock options and vesting schedules.

Expected  Volatility  –  The  expected  volatility  used  in  the  Black-Scholes  valuation  method  is  derived  from  the 
implied volatility related to our share price over the expected term.

Expected  Dividend  –  We  have  never  paid  dividends  on  our  shares  and,  accordingly,  the  dividend  yield 
percentage is zero for all periods.

Risk-Free Interest Rate – The risk-free interest rate used in the Black-Scholes valuation method is the implied 
yield  currently  available  on  U.S.  Treasury  constant  maturities  issued  with  a  term  equivalent  to  the  expected 
terms.

Stock Options

We estimated the fair value of employee stock options using the Black-Scholes option pricing model. The fair 
value  of  employee  stock  options  is  amortized  on  a  straight-line  basis  over  the  requisite  service  period  of  the 
awards. 

When determining the current share prices underlying the stock options for calculating the grant-date fair value, 
we reference observable market prices of similar or identical instruments in active markets. 

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The fair value of employee stock options was estimated using the following weighted-average assumptions:

Expected term in years

Expected volatility 

Risk-free interest rate 

Dividend yield 

Years Ended December 31,

2023

4.9

2022

4.6

2021

2.1 - 4.6

77% - 78%

70% - 76%

67% - 80%

3.73% – 4.60% 0.41% – 3.66% 0.05% – 1.10%

—

—

—

Weighted-average grant date fair value per share

$ 

7.32  $ 

5.93  $ 

15.53 

Cash received from option exercises for the years ended December 31, 2023, 2022, and 2021 was $6.5 million, 
$3.4 million and $25.4 million, respectively. 

ESPP 

We estimate the fair value of shares to be issued under the ESPP using the Black-Scholes option pricing model. 
The fair value of shares to be issued under the ESPP was estimated using the following assumptions:

Expected term in years

Expected volatility 

Risk-free interest rate 

Dividend yield 

Years Ended December 31,

2023

0.5 - 2.0

2022

0.5 - 2.0

2021

0.5 - 2.0

79% - 97%

70% - 97%

67% - 68%

4.9% - 5.5%

0.6% - 3.5%

0.1% - 0.2%

—

—

—

Weighted-average grant date fair value per share

$ 

5.34  $ 

4.28  $ 

25.07 

Cash received through the ESPP for the years ended December 31, 2023, 2022, and 2021 was $8.8 million, $7.8 
million, and $6.4 million, respectively. 

As of December 31, 2023, $101.9 million of total unrecognized compensation expense related to stock options, 
restricted stock, and ESPP shares was expected to be recognized over a weighted-average period of 2.2 years.

NOTE 11. NET LOSS PER SHARE

Basic net loss per share is computed by dividing net loss by the weighted average number of shares of common 
stock outstanding during the period. Diluted net loss per share is computed using the weighted-average number 
of  shares  of  common  stock  outstanding  and  potential  shares  assuming  the  dilutive  effect  of  the  convertible 
senior notes, using the if-converted method, and outstanding equity awards using the treasury stock method.

The following table presents the calculation of the basic and diluted net loss per share amounts presented in 
the consolidated statements of operations and comprehensive loss (in thousands, except per share amounts): 

Numerator:

Net loss

Denominator:

Basic

Years Ended December 31,

2023

2022

2021

$ 

(306,735)  $ 

(314,248)  $ 

(181,223) 

Weighted-average shares used in computing basic net loss per share

253,629 

224,550 

204,136 

Basic net loss per share

Diluted

$ 

(1.21)  $ 

(1.40)  $ 

(0.89) 

Weighted-average shares used in computing diluted net loss per 
share

Diluted net loss per share

253,629

224,550

204,136

$ 

(1.21)  $ 

(1.40)  $ 

(0.89) 

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The following shares issuable upon conversion of the convertible senior notes and outstanding equity awards 
were excluded from the computation of diluted net loss per share for the periods presented because the effect 
of including such shares would have been antidilutive (in thousands): 

(in thousands)

Shares issuable upon conversion of convertible senior notes

Equity awards

Years Ended December 31,

2023

2022

2021

31,063

27,246

20,690

27,291

20,690

21,419

As described in Note 2. Business Acquisitions, the contingently issuable shares would be due upon the 
achievement of a milestone. See Note 10. Stockholders’ Equity for detailed information on RSUs with time-based 
vesting and RSUs with performance-based vesting.

NOTE 12. SEGMENT AND GEOGRAPHIC INFORMATION

We are organized as, and operate in, one reportable segment: the development, manufacturing, and marketing 
of an integrated platform for genetic analysis. Our chief operating decision-maker is our Chief Executive Officer. 
The Chief Executive Officer reviews financial information presented on a consolidated basis for the purposes of 
evaluating  financial  performance  and  allocating  resources,  accompanied  by  information  about  revenue  by 
geographic regions. Our assets are primarily located in the United States of America and not allocated to any 
specific  region,  and  we  do  not  measure  the  performance  of  geographic  regions  based  upon  asset-based 
metrics. Therefore, geographic information is presented only for revenue.

A summary of our revenue by geographic location is as follows:

(in thousands)

Americas

Europe, Middle East, and Africa

Asia-Pacific

Total 

A summary of our revenue by category is as follows:

(in thousands)

Instrument revenue

Consumable revenue

Product revenue

Service and other revenue

Total revenue

Years Ended December 31,

2023

2022

2021

$ 

105,410  $ 

69,561  $ 

40,658 

54,453 

22,598 

36,145 

64,521 

30,271 

35,721 

$ 

200,521  $ 

128,304  $ 

130,513 

Years Ended December 31,

2023

2022

2021

$ 

120,451  $ 

48,719  $ 

63,421 

183,872 

16,649 

59,980 

108,699 

19,605 

61,324 

52,181 

113,505 

17,008 

$ 

200,521  $ 

128,304  $ 

130,513 

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ITEM  9. CHANGES  IN  AND  DISAGREEMENTS  WITH  ACCOUNTANTS  ON  ACCOUNTING  AND  FINANCIAL 
DISCLOSURE

None.

ITEM 9A. 

CONTROLS AND PROCEDURES

Evaluation of Disclosure Controls and Procedures 

Our  management,  with  the  participation  of  our  chief  executive  officer,  our  chief  financial  officer,  and  our 
principal accounting officer, evaluated the effectiveness of our disclosure controls and procedures (as defined 
in  Rules  13a–15(e)  and  15d–15(e)  of  the  Exchange  Act)  as  of  the  end  of  the  period  covered  by  this  Annual 
Report on Form 10–K. Disclosure controls and procedures include, without limitation, controls and procedures 
designed to ensure that information required to be disclosed by a company in the reports that it files or submits 
under  the  Exchange  Act  is  accumulated  and  communicated  to  our  management,  including  its  principal 
executive and principal financial officers, as appropriate to allow timely decisions regarding required disclosure. 
Management  recognizes  that  any  controls  and  procedures,  no  matter  how  well  designed  and  operated,  can 
provide  only  reasonable  assurance  of  achieving  their  objectives,  and  management  necessarily  applies  its 
judgement  in  evaluating  the  cost-benefit  relationship  of  possible  controls  and  procedures.  Based  on  this 
evaluation, our chief executive officer, chief financial officer and our principal accounting officer concluded that 
our disclosure controls and procedures were effective as of the end of the period covered by this report. 

Management’s Report on Internal Control Over Financial Reporting 

Our  management  is  responsible  for  establishing  and  maintaining  adequate  internal  control  over  financial 
reporting,  as  such  term  is  defined  in  Exchange  Act  Rules  13a-15(f).  Pacific  Biosciences  of  California,  Inc’s 
internal  control  over  financial  reporting  is  designed  to  provide  reasonable  assurance  to  the  Company’s 
management  and  board  of  directors  regarding  the  preparation  and  fair  presentation  of  published  financial 
statements.  Because  of  its  inherent  limitations,  internal  control  over  financial  reporting  may  not  prevent  or 
detect misstatements. Therefore, even those systems determined to be effective can provide only reasonable 
assurance  with  respect  to  financial  statement  preparation  and  presentation.  Management  assessed  the 
effectiveness of the Company’s internal control over financial reporting as of December 31, 2023. In making this 
assessment,  management  used  the  criteria  set  forth  by  the  Committee  of  Sponsoring  Organizations  of  the 
Treadway Commission (2013 framework) (the COSO criteria). Based on our assessment, we concluded that, as 
of  December  31,  2023,  the  Company’s  internal  control  over  financial  reporting  was  effective  based  on  those 
criteria.

The Company’s internal control over financial reporting as of December 31, 2023 has been audited by Ernst & 
Young  LLP,  the  independent  registered  public  accounting  firm  who  also  audited  the  Company’s  financial 
statements.  Ernst  &  Young’s  attestation  report  on  the  Company’s  internal  control  over  financial  reporting 
appears on page 113 hereof.

Changes in Internal Control Over Financial Reporting 

There were no material changes in our internal control over financial reporting identified in connection with the 
evaluation required by Rule 13a-15(d) and 15d-15(d) of the Exchange Act that occurred during the year ended 
December  31,  2023,  that  have  materially  affected,  or  were  reasonably  likely  to  materially  affect,  our  internal 
control over financial reporting.

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To the Stockholders and the Board of Directors of Pacific Biosciences of California, Inc.

Report of Independent Registered Public Accounting Firm

Opinion on Internal Control over Financial Reporting

We  have  audited  Pacific  Biosciences  of  California,  Inc.’s  internal  control  over  financial  reporting  as  of 
December  31,  2023,  based  on  criteria  established  in  Internal  Control—Integrated  Framework  issued  by  the 
Committee of Sponsoring Organizations of the Treadway Commission (2013 framework) (the COSO criteria). In 
our opinion, Pacific Biosciences of California, Inc. (the Company) maintained, in all material respects, effective 
internal control over financial reporting as of December 31, 2023, based on the COSO criteria.

We  also  have  audited,  in  accordance  with  the  standards  of  the  Public  Company  Accounting  Oversight  Board 
(United  States)  (PCAOB),  the  2023  consolidated  financial  statements  of  the  Company  and  our  report  dated 
February 28, 2024, expressed an unqualified opinion thereon.

Basis for Opinion

The  Company’s  management  is  responsible  for  maintaining  effective  internal  control  over  financial  reporting 
and  for  its  assessment  of  the  effectiveness  of  internal  control  over  financial  reporting  included  in  the 
accompanying  Management’s  Report  on  Internal  Control  Over  Financial  Reporting.  Our  responsibility  is  to 
express  an  opinion  on  the  Company’s  internal  control  over  financial  reporting  based  on  our  audit.  We  are  a 
public  accounting  firm  registered  with  the  PCAOB  and  are  required  to  be  independent  with  respect  to  the 
Company  in  accordance  with  the  U.S.  federal  securities  laws  and  the  applicable  rules  and  regulations  of  the 
Securities and Exchange Commission and the PCAOB.

We conducted our audit in accordance with the standards of the PCAOB. Those standards require that we plan 
and  perform  the  audit  to  obtain  reasonable  assurance  about  whether  effective  internal  control  over  financial 
reporting was maintained in all material respects.

Our audit included obtaining an understanding of internal control over financial reporting, assessing the risk that 
a  material  weakness  exists,  testing  and  evaluating  the  design  and  operating  effectiveness  of  internal  control 
based  on  the  assessed  risk,  and  performing  such  other  procedures  as  we  considered  necessary  in  the 
circumstances. We believe that our audit provides a reasonable basis for our opinion. 

Definition and Limitations of Internal Control Over Financial Reporting

A  company’s  internal  control  over  financial  reporting  is  a  process  designed  to  provide  reasonable  assurance 
regarding the reliability of financial reporting and the preparation of financial statements for external purposes 
in  accordance  with  generally  accepted  accounting  principles.  A  company’s  internal  control  over  financial 
reporting  includes  those  policies  and  procedures  that  (1)  pertain  to  the  maintenance  of  records  that,  in 
reasonable detail, accurately and fairly reflect the transactions and dispositions of the assets of the company; 
(2) provide reasonable assurance that transactions are recorded as necessary to permit preparation of financial 
statements in accordance with generally accepted accounting principles, and that receipts and expenditures of 
the  company  are  being  made  only  in  accordance  with  authorizations  of  management  and  directors  of  the 
company;  and  (3)  provide  reasonable  assurance  regarding  prevention  or  timely  detection  of  unauthorized 
acquisition,  use,  or  disposition  of  the  company’s  assets  that  could  have  a  material  effect  on  the  financial 
statements.

Because  of  its  inherent  limitations,  internal  control  over  financial  reporting  may  not  prevent  or  detect 
misstatements. Also, projections of any evaluation of effectiveness to future periods are subject to the risk that 
controls may become inadequate because of changes in conditions, or that the degree of compliance with the 
policies or procedures may deteriorate.

/s/ Ernst & Young LLP

San Mateo, California 
February 28, 2024

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ITEM 9B. 

OTHER INFORMATION

Securities Trading Plans of Directors and Executive Officers.

During our last fiscal quarter, no director or officer, as defined in Rule 16a-1(f), adopted or terminated a “Rule 
10b5-1  trading  arrangement”  or  a  “non-Rule  10b5-1  trading  arrangement,”  each  as  defined  in  Item  408  of 
Regulation S-K.

ITEM 9C. 

DISCLOSURE REGARDING FOREIGN JURISDICTIONS THAT PREVENT INSPECTIONS

Not applicable.

PART III

ITEM 10. 

DIRECTORS, EXECUTIVE OFFICERS AND CORPORATE GOVERNANCE 

Information  responsive  to  this  item  is  incorporated  herein  by  reference  to  our  definitive  proxy  statement  with 
respect to our 2024 Annual Meeting of Stockholders to be filed with the SEC within 120 days after the end of the 
fiscal year covered by this Annual Report on Form 10-K. 

ITEM 11. 

EXECUTIVE COMPENSATION 

Information  responsive  to  this  item  is  incorporated  herein  by  reference  to  our  definitive  proxy  statement  with 
respect to our 2024 Annual Meeting of Stockholder to be filed with the SEC within 120 days after the end of the 
fiscal year covered by this Annual Report on Form 10-K. 

ITEM  12. 

SECURITY  OWNERSHIP  OF  CERTAIN  BENEFICIAL  OWNERS  AND  MANAGEMENT  AND 
RELATED STOCKHOLDER MATTERS 

Information  responsive  to  this  item  is  incorporated  herein  by  reference  to  our  definitive  proxy  statement  with 
respect to our 2024 Annual Meeting of Stockholders to be filed with the SEC within 120 days after the end of the 
fiscal year covered by this Annual Report on Form 10-K. 

ITEM 13. 

CERTAIN RELATIONSHIPS AND RELATED TRANSACTIONS, AND DIRECTOR INDEPENDENCE

Information  responsive  to  this  item  is  incorporated  herein  by  reference  to  our  definitive  proxy  statement  with 
respect to our 2024 Annual Meeting of Stockholders to be filed with the SEC within 120 days after the end of the 
fiscal year covered by this Annual Report on Form 10-K. 

ITEM 14. 

PRINCIPAL ACCOUNTANT FEES AND SERVICES 

Information  responsive  to  this  item  is  incorporated  herein  by  reference  to  our  definitive  proxy  statement  with 
respect to our 2024 Annual Meeting of Stockholders to be filed with the SEC within 120 days after the end of the 
fiscal year covered by this Annual Report on Form 10-K. 

ITEM 15. 

EXHIBITS, FINANCIAL STATEMENT SCHEDULES. 

PART IV

(a)

The  following  documents  are  filed  as  part  of,  or  incorporated  by  reference  into,  this  Annual 
Report on Form 10-K: 

1. Financial Statements: See Index to Consolidated Financial Statements under Item 8 of this 

Annual Report on Form 10-K. 

2. Financial Statement Schedules: All schedules are omitted because they are not required, are 
not  applicable  or  the  information  is  included  in  the  consolidated  financial  statements  or 
notes thereto. 

3. Exhibits: We have filed or incorporated by reference into this Annual Report on Form 10-K, 

the exhibits listed on the accompanying Exhibit Index immediately below. 

(b)

(c)

Financial Statement Schedules: See Item 15(a)(2), above. 

Exhibits: Refer to the Exhibit Index that follows. 

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Exhibit Index

Exhibit
͏Number
3.1

3.2

4.1

4.2

4.3

4.4

4.5

4.6

10.1+

10.2+

10.3+

10.4+

10.5+

10.6+

10.7+

10.8+

10.9+

10.10+

10.11+

10.12+

10.13+

10.14+

10.15+

10.16+

10.17+

10.18†

10.19

10.20††

10.21+

Description

Amended and Restated Certificate of Incorporation

Third Amended and Restated Bylaws of Pacific Biosciences of 
California, Inc. 

Specimen Common Stock Certificate

Incorporated by reference herein

Form

10-K

8-K

S-1/A

Exhibit 
No.

Filing Date

3.1

March 23, 2011

3.1

4.1

November 7, 2022

October 1, 2010

Description of Registrant’s securities registered under Section 12 of the 
Exchange Act

10-K

4.2

February 28, 2020

Indenture, dated February 16, 2021, between Pacific Biosciences of 
California, Inc., and U.S. Bank National Association, as Trustee

10-Q

4.1

August 4, 2023

Form of 1.50% Convertible Senior Notes due 2028 (included in Exhibit 
4.3)

10-Q

4.1

August 4, 2023

Indenture, dated as of June 30, 2023, by and between Pacific 
Biosciences of California, Inc. and U.S. Bank Trust Company, National 
Association, as Trustee

Form of 1.375% Convertible Senior Note due 2030

Form of Director and Executive Officer Indemnification Agreement

2010 Equity Incentive Plan

8-K

8-K

S-1

S-1

4.1

4.2

10.1

10.4

June 30, 2023

June 30, 2023

August 16, 2010

August 16, 2010

2010 Equity Incentive Plan forms of agreement

10-Q

10.1 May 2, 2018

2010 Employee Stock Purchase Plan and forms of agreement 
thereunder

2010 Outside Director Equity Incentive Plan 

2010 Outside Director Equity Incentive Plan forms of agreement

2020 Equity Incentive Plan, as amended 

Form of Global Stock Option Agreement under the Pacific Biosciences 
of California, Inc., 2020 Equity Incentive Plan, as amended

Form of Global Restricted Stock Unit Agreement under the Pacific 
Biosciences of California, Inc. 2020 Equity Incentive Plan, as amended

S-1

S-1

10-Q

8-K

8-K

8-K

10.5

10.6

August 16, 2010

August 16, 2010

10.2 May 2, 2018

10.1 May 26, 2022

10.2 May 26, 2022

10.3 May 26, 2022

Omniome Equity Incentive Plan of Pacific Biosciences of California, 
Inc., and related forms of agreement thereunder

10-Q

10.4

November 5, 2021

Pacific Biosciences of California, Inc. 2020 Inducement Equity Incentive 
Plan, as amended, and forms of agreement thereunder

8-K

10.1

November 19, 
2021

Letter Relating to Employment Terms by and between the Registrant 
and Susan G. Kim effective September 28, 2020

10-Q

10.2

November 3, 2020

Form of Change in Control and Severance Agreement for executive 
officers

10-K

10.14

February 26, 2021

Letter Relating to Employment Terms by and between the Registrant 
and Christian O. Henry effective September 14, 2020

10-K

10.15

February 26, 2021

Amended Change in Control and Severance Agreement by and between 
the Registrant and Christian O. Henry dated February 3, 2021

10-K

10.17

February 26, 2021

Letter Relating to Employment Terms by and between the Registrant 
and Mark Van Oene effective January 8, 2021

10-K

10.18

February 26, 2021

Lease Agreement by and between the Registrant and Menlo Park 
Portfolio II, LLC, dated July 22, 2015

10-Q

10.2

August 5, 2015

First Amendment to Lease Agreement by and between the Registrant 
and Menlo Park Portfolio II, LLC, dated December 23, 2016

10-K

10.50 March 6, 2017

Investment Agreement, dated as of February 9, 2021, between Pacific 
Biosciences of California, Inc. and SB Northstar LP.

8-K

10.1

February 10, 2021

Exclusive License Agreement by and between the Registrant and 
Cornell Research Foundation, Inc., dated as of February 1, 2004

S-1/A

10.8

October 22, 2010

Letter Relating to Employment Terms by and between the Registrant 
and Michele Farmer effective May 17, 2021

10-Q

10.2

August 6, 2021

115

 
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10.22

10.23

10.24

10.25+

10.26+

10.27

10.28+

21.1

23.1

31.1

31.2

32.1*

32.2*

97.1

101.INS

Agreement and Plan of Merger of Reorganization among Pacific 
Biosciences of California, Inc., Apollo Acquisition Corp., Apollo 
Acquisition Sub, LLC, Omniome, Inc. and Shareholder Representative 
Services, LLC, as securityholder representative, dated as of July 19, 
2021

Securities Purchase Agreement, dated as of July 19, 2021, by and 
between Pacific Biosciences of California, Inc., and each of the 
Investors

Registration Rights Agreement, dated as of July 19, 2021, by and 
between Pacific Biosciences of California, Inc., and each of the 
Investors

Letter Relating to Employment Terms by and between the Registrant 
and Jeff Eidel effective August 16, 2022

Change in Control and Severance Agreement by and between the 
Registrant and Susan G. Kim effective February 3, 2021

Letter Agreement, dated June 23, 2023, between the Company and 
Chimera Investment LLC

Form of Performance-Based Restricted Stock Unit Award Agreement 
under the Pacific Biosciences of California, Inc. 2020 Equity Incentive 
Plan, as amended

List of Subsidiaries of the Registrant

Consent of Independent Registered Public Accounting Firm

Certification of Chief Executive Officer pursuant to Exchange Act Rules 
13a-14(a) and 15d-14(a), as adopted pursuant to Section 302 of the 
Sarbanes-Oxley Act of 2002

Certification of Chief Financial Officer pursuant to Exchange Act Rules 
13a-14(a) and 15d-14(a), as adopted pursuant to Section 302 of the 
Sarbanes-Oxley Act of 2002

Certification of Chief Executive Officer pursuant to 18 U.S.C. Section 
1350, as adopted pursuant to Section 906 of the Sarbanes-Oxley Act of 
2002

Certification of Chief Financial Officer pursuant to 18 U.S.C. Section 
1350, as adopted pursuant to Section 906 of the Sarbanes-Oxley Act of 
2002

Compensation Recovery Policy

XBRL Instance Document (the instance document does not appear in 
the Interactive Data File because its XBRL tags are embedded within 
the Inline XBRL document)

101.SCH XBRL Taxonomy Extension Schema Document

101.CAL XBRL Taxonomy Extension Calculation Linkbase Document

101.DEF

XBRL Taxonomy Extension Definition Linkbase Document

101.LAB XBRL Taxonomy Extension Labels Linkbase Document

101.PRE XBRL Taxonomy Extension Presentation Linkbase Document

104

Cover Page Interactive File (formatted as inline XBRL and contained in 
Exhibit 101)

_____________________

+

Indicates management contract or compensatory plan. 

8-K

10.1

July 20, 2021

8-K

10.2

July 20, 2021

8-K

8-K

8-K

8-K

10.3

July 20, 2021

99.3

January 24, 2023

99.4

January 24, 2023

10.1

June 23, 2023

Filed herewith

Filed herewith

Filed herewith

Filed herewith

Filed herewith

Furnished 
herewith

Furnished 
herewith

Filed herewith

Filed herewith

Filed herewith

Filed herewith

Filed herewith

Filed herewith

Filed herewith

Filed herewith

† Confidential  treatment  has  been  requested  for  portions  of  this  exhibit.  These  portions  have  been  omitted  and  have  been  filed 

separately with the Securities and Exchange Commission.

†† Certain confidential information contained in this Exhibit was omitted by means of marking such portions with brackets because the 

identified confidential information (i) is not material and (ii) would be competitively harmful if publicly disclosed.

*

The certifications attached as Exhibit 32.1 and 32.2 that accompany this Annual Report on Form 10-K are deemed furnished and not 
filed with the Securities and Exchange Commission and are not to be incorporated by reference into any filing of Pacific Biosciences 
of California, Inc. under the Securities Act of 1933, as amended, or the Securities Exchange Act of 1934, as amended, whether made 
before or after the date of this Annual Report on Form 10-K, irrespective of any general incorporation language contained in such filing.

116

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ITEM 16. 

FORM 10-K SUMMARY

None.

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Pursuant to the requirements of Section 13 or 15(d) of the Securities Exchange Act of 1934, as amended, the 
Registrant  has  duly  caused  this  Annual  Report  on  Form  10-K  to  be  signed  on  its  behalf  by  the  undersigned, 
thereunto duly authorized. 

Signatures 

Date: February 28, 2024

Pacific Biosciences of California, Inc.

By:

/s/ Christian O. Henry 

Christian O. Henry

President and Chief Executive Officer

Date: February 28, 2024

By:

/s/ Susan G. Kim 

Susan G. Kim

Chief Financial Officer 

Date: February 28, 2024

By:

/s/ Michele Farmer 

Michele Farmer

Vice President and Chief Accounting 
Officer 

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Table of Contents

POWER OF ATTORNEY 

KNOW  ALL  PERSONS  BY  THESE  PRESENTS,  that  each  person  whose  signature  appears  below  hereby 
constitutes and appoints Christian O. Henry, Susan G. Kim, Brett Atkins, and Michele Farmer, and each of them, 
as  his  or  her  true  and  lawful  attorney-in-fact  and  agent,  with  full  power  of  substitution  and  resubstitution,  for 
each individual in any and all capacities, to sign any and all amendments to this Annual Report on Form 10-K, 
and to file the same, with all exhibits thereto and other documents in connection therewith, with the Securities 
and Exchange Commission, granting unto said attorneys-in-fact and agents, and each of them, full power and 
authority  to  do  and  perform  each  and  every  act  and  thing  requisite  and  necessary  to  be  done  in  connection 
therewith,  as  fully  for  all  intents  and  purposes  as  he  or  she  might  or  could  do  in  person,  hereby  ratifying  and 
confirming all that said attorneys-in-fact and agents, or any of them, or the individual’s substitute, may lawfully 
do or cause to be done by virtue hereof.

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Table of Contents

Pursuant to the requirements of the Securities Exchange Act of 1934, as amended, this report has been signed 
by the following persons on behalf of the Registrant in the capacities and on the dates indicated.

Signature

Title

Date

/s/ Christian O. Henry  Director, President and Chief Executive 

February 28, 2024

Christian O. Henry

Officer
(Principal Executive Officer)

/s/ Susan G. Kim

Susan G. Kim

Chief Financial Officer 
(Principal Financial Officer)

February 28, 2024

/s/ Michele Farmer

Michele Farmer

Vice President and Chief Accounting Officer 
(Principal Accounting Officer)

February 28, 2024

/s/ John F. Milligan

Chairman of the Board of Directors

February 28, 2024

February 28, 2024

February 28, 2024

February 28, 2024

February 28, 2024

February 28, 2024

February 28, 2024

February 28, 2024

John F. Milligan

/s/ William W. Ericson Director

William W. Ericson

/s/ Hannah A. 
Valantine

Director

Hannah A. Valantine

/s/ Randall S. 
Livingston

Director

Randall S. Livingston

/s/ Marshall L. Mohr Director

Marshall L. Mohr

/s/ Kathy Ordoñez

Director

Kathy Ordoñez

/s/ Lucy Shapiro

Director

Lucy Shapiro

/s/ David Meline

Director 

David Meline 

120

EXECUTIVE OFFICERS

Christian Henry
President and Chief Executive Officer

Jeff Eidel
Chief Commercial Officer

BOARD OF DIRECTORS

Mark Van Oene
Chief Operating Officer

Susan G. Kim
Chief Financial Officer

Christian Henry
President and Chief Executive 
Officer

John Milligan, PhD (chair)
Chairman of the Board of 
Directors

William Ericson
Founding Partner at Wildcat 
Venture Partners

Randy Livingston
VP for Business Affairs and 
Chief Financial Officer at 
Stanford University

Kathy Ordoñez
Director and former Chief 
Commercial Officer

David Meline
Former Chief Financial Officer
at Moderna, Inc.

Marshall Mohr
EVP, Global Business Services
at Intuitive Surgical, Inc.

Lucy Shapiro, PhD
Professor of Cancer Research 
and Director of Molecular and 
Genetic Medicine at Stanford 
University’s School of Medicine

Hannah A. Valantine, MD
Professor of Medicine 
(Cardiovascular) at the Stanford 
University Medical Center

For additional biographical information on our directors and executive officers, see the sections of our 
proxy statement captioned “Corporate Governance — Board of Directors and Committees of the Board” and 
“Executive Officers.” A copy of our proxy statement is included with this annual report to stockholders.

CORPORATE COUNSEL

TRANSFER AGENT AND REGISTRAR

Wilson Sonsini Goodrich & Rosati
650 Page Mill Road
Palo Alto, California 94304

COMMON STOCK LISTING

The Nasdaq Global Select Market
Ticker symbol: PACB

CORPORATE HEADQUARTERS

Computershare
By mail:
c/o Shareholder Services
P.O. Box 43078
Providence, RI 02940-3078

By courier:
c/o Shareholder Services
150 Royall Street, Suite 101
Canton, MA 02021

Phone: 866.401.4874
Foreign shareholders: 201.680.6578
www.computershare.com/investor

1305 O’Brien Drive, Menlo Park, CA 94025  |  650.521.8000  |  www.pacb.com

Research use only. Not for use in diagnostic procedures. © 2024 Pacific Biosciences of California, Inc. 
(“PacBio”). All rights reserved. Information in this document is subject to change without notice. PacBio 
assumes no responsibility for any errors or omissions in this document. Certain notices, terms, conditions 
and/or use restrictions may pertain to your use of PacBio products and/or third-party products. Refer to the 
applicable PacBio terms and conditions of sale and to the applicable license terms at pacb.com/license. 
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Pacific Biosciences
1305 O’Brien Drive Menlo Park, CA 94025
650.521.8000  |  www.pacb.com